treatments-xml/data/E9/3D/87/E93D8780802A8E4C75A7FDD7FB9DF94F.xml

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<mods:title id="46456088DB034BC69B878D816C983C4B">New species of Colyphus Spinola (Coleoptera: Cleridae: Clerinae) from Central America, with taxonomic and distributional notes on some others</mods:title>
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<mods:namePart id="FE13649D12E3B5CEB0423BDE5CD74CAD">Rifkind, Jacques</mods:namePart>
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<mods:namePart id="A90D49F2E1C045DF325AB7B5C54022F0">Barr, William F.</mods:namePart>
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<mods:date id="4D44C8AF0F93303E36DE7F1C206B5AE9">2011</mods:date>
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<emphasis id="53E0EA84802A8E4E75A7FDD7FDDAFDEB" bold="true" box="[151,573,511,536]" pageId="1" pageNumber="56">
<taxonomicName id="A6944D15802A8E4E75A7FDD7FE14FDEB" authority="Rifkind &amp; Barr" authorityName="Rifkind &amp; Barr" box="[151,499,511,536]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="ska" status="sp. nov.">
<emphasis id="53E0EA84802A8E4E75A7FDD7FED5FDEB" bold="true" box="[151,306,511,536]" italics="true" pageId="1" pageNumber="56">Colyphus ska</emphasis>
Rifkind &amp; Barr
</taxonomicName>
, 
<taxonomicNameLabel id="48D357FF802A8E4E74CFFE28FDDAFDEB" box="[511,573,512,536]" pageId="1" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
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(
<figureCitation id="F9AF2A13802A8E4E75AFFE0CFF07FDCF" box="[159,224,548,572]" captionStart="FIGURE 1" captionStartId="2.[151,250,1051,1074]" captionText="FIGURE 1. Habitus of Colyphus ska Rifkind &amp; Barr, n. sp. (holotype male)." pageId="1" pageNumber="56">Fig. 1</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="298E651D802A8E4E75A7FE44FEA5FA33" pageId="1" pageNumber="56" type="materials_examined">
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<emphasis id="53E0EA84802A8E4E75A7FE44FE87FD77" bold="true" box="[151,352,619,644]" pageId="1" pageNumber="56">
<typeStatus id="BE2F8834802A8E4E75A7FE44FF35FD77" box="[151,210,620,644]" pageId="1" pageNumber="56">Type</typeStatus>
specimens.
</emphasis>
<typeStatus id="BE2F8834802A8E4E745EFE44FE31FD77" box="[366,470,620,644]" pageId="1" pageNumber="56" type="holotype">Holotype</typeStatus>
male: 
<collectingCountry id="19837606802A8E4E7701FE44FD52FD77" box="[561,693,620,644]" name="Costa Rica" pageId="1" pageNumber="56">Costa Rica</collectingCountry>
, Puntarenas, Monteverde, 
<date id="152A1056802A8E4E76C5FE44FB98FD77" box="[1013,1151,620,644]" pageId="1" pageNumber="56" value="1912-06-13">12–13 June</date>
, 2002, F. Hovore, coll. (
<collectionCode id="0785AE53802A8E4E75AFFEA7FF09FD54" LSID="urn:lsid:biocol.org:col:33156" box="[159,238,655,679]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33156" name="California State Collection of Arthropods" pageId="1" pageNumber="56">CSCA</collectionCode>
). 
<typeStatus id="BE2F8834802A8E4E7433FEB8FE93FD5B" box="[259,372,656,680]" pageId="1" pageNumber="56" type="paratype">Paratypes</typeStatus>
: 
<collectingCountry id="19837606802A8E4E74B0FEA7FDCEFD54" box="[384,553,655,679]" name="Costa Rica" pageId="1" pageNumber="56">COSTA RICA</collectingCountry>
, PUNTARENAS PROVINCE: 1, Monteverde, 
<quantity id="A66C9B73802A8E4E7177FEA7FB47FD54" box="[1095,1184,655,679]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="56" unit="m" value="1500.0">1500 m</quantity>
, 
<date id="152A1056802A8E4E719EFEA7FAD6FD54" box="[1198,1329,655,679]" pageId="1" pageNumber="56" value="1991-02-23">23–ii–1991</date>
. H. &amp; A. Howden; 7, Monteverde, 
<date id="152A1056802A8E4E74F0FE9CFD7FFD3F" box="[448,664,692,716]" pageId="1" pageNumber="56" value="1988-04-19" valueMax="1988-04-26" valueMin="1988-04-19">April 19–26, 1988</date>
, E. Giesbert, coll.; 2, Monteverde, 
<date id="152A1056802A8E4E7105FE9DFAE2FD3F" box="[1077,1285,692,717]" pageId="1" pageNumber="56" value="1987-05-23" valueMax="1987-05-27" valueMin="1987-05-23">May 23–27, 1987</date>
, E. Giesbert, coll.; 3, Monteverde, 
<date id="152A1056802A8E4E74BDFEF1FDBEFD1C" box="[397,601,727,753]" pageId="1" pageNumber="56" value="1996-05-11" valueMax="1996-05-13" valueMin="1996-05-11">May 11–13, 1996</date>
, E. Giesbert, coll.; 1, Monteverde, 4000', 
<geoCoordinate id="04A05051802A8E4E710AFEFFFB7EFD03" box="[1082,1177,727,752]" direction="north" orientation="latitude" pageId="1" pageNumber="56" precision="555" value="10.1">10.10 N</geoCoordinate>
, 
<geoCoordinate id="04A05051802A8E4E7196FEFFFAEEFD03" box="[1190,1289,727,752]" direction="west" orientation="longitude" pageId="1" pageNumber="56" precision="555" value="-83.43">83.43 W</geoCoordinate>
, 
<date id="152A1056802A8E4E7026FEFFFA7FFD1C" box="[1302,1432,727,751]" pageId="1" pageNumber="56" value="1983-07-09">9–vii–1983</date>
, Warren Colby; 1, Monteverde, 
<date id="152A1056802A8E4E7733FED4FD66FCE6" box="[515,641,764,789]" pageId="1" pageNumber="56" value="1917-05-18">17/18 May</date>
, 1984, F. Hovore, coll.;1, Monteverde, 
<date id="152A1056802A8E4E7160FED4FAE4FCE7" box="[1104,1283,764,788]" pageId="1" pageNumber="56" value="1984-05-26" valueMax="1984-06-03" valueMin="1984-05-26">v–26—vi–3–84</date>
, E. Riley, D. Rider, &amp; D. Le Doux; 2, Monteverde, 
<date id="152A1056802A8E4E7775FF37FD2FFCCB" box="[581,712,799,824]" pageId="1" pageNumber="56" value="1920-06-24">20–24 June</date>
, 1986, W. Hanson, G. Bohart; 4, Monteverde Forest Res., 
<date id="152A1056802A8E4E7066FF37FF1BFCAF" pageId="1" pageNumber="56" value="1988-05-19" valueMax="1988-05-21" valueMin="1988-05-19">v–19– 21–1988</date>
, 
<quantity id="A66C9B73802A8E4E7438FF6CFE87FCAF" box="[264,352,836,860]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="56" unit="m" value="1500.0">1500 m</quantity>
, B. Ratcliffe &amp; M. Jameson; 1, Res. Biol. Monteverde, Est. La Casona, 
<quantity id="A66C9B73802A8E4E71AAFF6CFB15FCAF" box="[1178,1266,836,860]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="56" unit="m" value="1520.0">1520 m</quantity>
, 
<date id="152A1056802A8E4E7030FF6CFA93FCAF" box="[1280,1396,836,860]" pageId="1" pageNumber="56" value="1992-02">Feb. 1992</date>
, N. Obando; 1, Res. Biol. Monteverde, 
<quantity id="A66C9B73802A8E4E7717FF4FFD98FC73" box="[551,639,871,896]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="56" unit="m" value="1520.0">1520 m</quantity>
, 
<date id="152A1056802A8E4E77BBFF4FFD1AFC8C" box="[651,765,871,895]" pageId="1" pageNumber="56" value="1991-04">Abr. 1991</date>
, N. Obando; 1, R. B. Monteverde, Est. La Casona, 
<quantity id="A66C9B73802A8E4E7070FF4FFA70FC73" box="[1344,1431,871,896]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="56" unit="m" value="1520.0">1520 m</quantity>
, 
<date id="152A1056802A8E4E75A7FFA4FEE7FC57" box="[151,256,908,932]" pageId="1" pageNumber="56" value="1912-07-05">5–12 Jul.</date>
, 1994, K. Martinez; 2, Res. Biol. Monteverde, Est. La Casona, 
<quantity id="A66C9B73802A8E4E76F3FFA4FBFEFC57" box="[963,1049,908,932]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="56" unit="m" value="1520.0">1520 m</quantity>
, 
<date id="152A1056802A8E4E7115FFA5FB7EFC57" box="[1061,1177,908,933]" pageId="1" pageNumber="56" value="1991-05">May 1991</date>
, N. Obando; 1, Monteverde, Est. La Casona, 
<quantity id="A66C9B73802A8E4E74A7FF87FE0AFC3B" box="[407,493,943,968]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="56" unit="m" value="1520.0">1520 m</quantity>
, 
<date id="152A1056802A8E4E74C9FF87FD4AFC3B" box="[505,685,943,968]" pageId="1" pageNumber="56">24 Jul.—6 Ago.</date>
, 1995, K. Martinez; 2, Monteverde, 
<date id="152A1056802A8E4E7172FF87FB51FC34" box="[1090,1206,943,967]" pageId="1" pageNumber="56" value="1985-05-07">v–7–1985</date>
. A. J. Gilbert, blacklight; 1, Monteverde, 
<date id="152A1056802A8E4E7496FFFCFDC0FC1F" box="[422,551,980,1004]" pageId="1" pageNumber="56" value="1985-05-23">v–23–1985</date>
, A. J. Gilbert, blacklight; 1, Monteverde, 
<date id="152A1056802A8E4E710CFFFCFB5AFC1F" box="[1084,1213,980,1004]" pageId="1" pageNumber="56" value="1985-05-24">v–24–1985</date>
, A. J. Gilbert; 1, Monteverde, 
<date id="152A1056802A8E4E741CFFDFFE15FBFC" box="[300,498,1015,1039]" pageId="1" pageNumber="56" value="1990-07-19" valueMax="1990-07-21" valueMin="1990-07-19">July 19–21, 1990</date>
, G. Snelling, coll.; 2, Monteverde, Cloud For. Res., 
<quantity id="A66C9B73802A8E4E7108FFDFFB77FBE3" box="[1080,1168,1015,1040]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="1" pageNumber="56" unit="m" value="1450.0">1450 m</quantity>
, 
<date id="152A1056802A8E4E71ACFFDFFB33FBFC" box="[1180,1236,1015,1039]" bridgedPair="-" pageId="1" pageNumber="56" value="1918-05">v–18</date>
/19–85, J. Doyen; 
<quantity id="A66C9B73802A8E4E75A7F834FF09FBC7" box="[151,238,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="56" unit="km" value="1.2">1, 2 km</quantity>
E Monteverde, 
<quantity id="A66C9B73802A8E4E7495F834FE1AFBC7" box="[421,509,1052,1076]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="1" pageNumber="56" unit="m" value="1450.0">1450 m</quantity>
, 
<date id="152A1056802A8E4E773AF834FDA5FBC7" box="[522,578,1052,1076]" bridgedPair="-" pageId="1" pageNumber="56" value="1918-05">v–18</date>
/20–85, J. Powell, coll.; 1, Monteverde, Campbell's Bullpen, 
<date id="152A1056802A8E4E71C6F835FA7FFBC7" box="[1270,1432,1052,1077]" pageId="1" pageNumber="56" value="1992-05-29">May 29, 1992</date>
, lighting, Lingafelter, Jameson, Ratcliffe; 2, Monteverde, 
<date id="152A1056802A8E4E7627F868FC3BFBAB" box="[791,988,1088,1112]" pageId="1" pageNumber="56" value="1990-07-19" valueMax="1990-07-21" valueMin="1990-07-19">July 19–21, 1990</date>
, G. Snelling; 3, Monteverde, 
<date id="152A1056802A8E4E7014F868FF35FB8F" pageId="1" pageNumber="56" value="1996-06-12" valueMax="1996-06-15" valueMin="1996-06-12">12–15–vi– 1996</date>
, F. Hovore, coll.; 1, Monteverde, cloud forest, 
<date id="152A1056802A8E4E77CCF84CFC49FB8F" box="[764,942,1124,1148]" pageId="1" pageNumber="56" value="2000-04-16" valueMax="2000-04-19" valueMin="2000-04-16">iv–16–19–2000</date>
, J. &amp; A. Rifkind, P. Gum, colls.; 1, 
<collectingCountry id="19837606802A8E4E706EF84CFF3EFB53" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="56">Santa Elena</collectingCountry>
, ca 
<quantity id="A66C9B73802A8E4E7438F8A0FE87FB53" box="[264,352,1160,1184]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="56" unit="m" value="1700.0">1700 m</quantity>
, 
<date id="152A1056802A8E4E745EF8A0FDC7FB53" box="[366,544,1160,1184]" pageId="1" pageNumber="56" value="2000-04-18" valueMax="2000-04-19" valueMin="2000-04-18">iv–18–19–2000</date>
, cloud forest, J. &amp; A. Rifkind, P. Gum, colls., beating roadside slash; 1, 
<collectingCountry id="19837606802A8E4E706EF8A0FF3EFB37" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="56">Santa Elena</collectingCountry>
, 
<date id="152A1056802A8E4E75D6F884FEBFFB37" box="[230,344,1196,1220]" pageId="1" pageNumber="56" value="1985-05-06">v–6–1985</date>
, A. J. Gilbert; SAN 
<collectionCode id="0785AE53802A8E4E7772F884FD65FB37" box="[578,642,1196,1220]" pageId="1" pageNumber="56">JOSÉ</collectionCode>
PROVINCE: 1, Zurqui de Moravia, 
<quantity id="A66C9B73802A8E4E7118F884FB66FB37" box="[1064,1153,1196,1220]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="56" unit="m" value="1600.0">1600 m</quantity>
, 
<date id="152A1056802A8E4E71BEF884FB0BFB37" box="[1166,1260,1196,1220]" bridgedPair="-" pageId="1" pageNumber="56" value="1992-04">iv–1992</date>
, col. Paul Hanson; 3, Zurqui de Moravia, 
<quantity id="A66C9B73802A8E4E74F7F8F8FDFAFB1B" box="[455,541,1232,1256]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="56" unit="m" value="1600.0">1600 m</quantity>
, 
<geoCoordinate id="04A05051802A8E4E7718F8F8FD73FB1B" box="[552,660,1232,1256]" direction="north" orientation="latitude" pageId="1" pageNumber="56" precision="925" value="10.05">10° 03' N</geoCoordinate>
, 
<geoCoordinate id="04A05051802A8E4E77AFF8F8FCE0FB1B" box="[671,775,1232,1256]" direction="west" orientation="longitude" pageId="1" pageNumber="56" precision="925" value="-84.01667">84° 01'W</geoCoordinate>
, P. Hanson, coll., 
<date id="152A1056802A8E4E76FEF8F8FBC3FB1B" box="[974,1060,1232,1256]" bridgedPair="-" pageId="1" pageNumber="56" value="1992-05">v–1992</date>
, Malaise trap; 2, Zurqui de Moravia, 
<quantity id="A66C9B73802A8E4E75F8F8DCFEC7FAFF" box="[200,288,1268,1292]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="56" unit="m" value="1600.0">1600 m</quantity>
, 
<geoCoordinate id="04A05051802A8E4E741DF8DCFE7AFAFF" box="[301,413,1268,1292]" direction="north" orientation="latitude" pageId="1" pageNumber="56" precision="925" value="10.05">10° 03' N</geoCoordinate>
, 
<geoCoordinate id="04A05051802A8E4E7499F8DCFDFAFAFF" box="[425,541,1268,1292]" direction="west" orientation="longitude" pageId="1" pageNumber="56" precision="925" value="-84.01667">84° 01' W</geoCoordinate>
, iv– 
<date id="152A1056802A8E4E777FF8DCFD43FAFF" box="[591,676,1268,1292]" bridgedPair="-" pageId="1" pageNumber="56" value="1993-05">v–1993</date>
, Malaise trap, cloud forest. P. Hanson, coll.; 1, Zurqui de Moravia, 
<quantity id="A66C9B73802A8E4E75A7F930FF0AFAC3" box="[151,237,1304,1328]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="56" unit="m" value="1600.0">1600 m</quantity>
, cloud forest, 
<date id="152A1056802A8E4E74BAF931FDCFFAC3" box="[394,552,1304,1329]" pageId="1" pageNumber="56" value="1994-05-22">May 22, 1994</date>
, beating melastomes, etc., J. Rifkind, P. Gum, colls.; 1, Coronado, 
<date id="152A1056802A8E4E703FF930FA9AFAC3" box="[1295,1405,1304,1328]" bridgedPair="-" pageId="1" pageNumber="56" value="1923-08">viii–1923</date>
, F. Nevermann; 1, Coronado, 
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, 
<date id="152A1056802A8E4E775DF914FD0DFAA7" box="[621,746,1340,1364]" pageId="1" pageNumber="56" value="1931-08-15">15–viii–31</date>
, F. Nevermann, an Blüten von &quot;plainillo&quot;; 1, La Palma, 
<quantity id="A66C9B73802A8E4E7054F914FF48FA8B" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="56" unit="m" value="1500.0">1500 m</quantity>
, 
<date id="152A1056802A8E4E758CF948FECDFA8B" box="[188,298,1376,1400]" pageId="1" pageNumber="56" value="1931-03-17">17–iii–31</date>
, Fidel 
<collectingCountry id="19837606802A8E4E7449F948FE2DFA8B" box="[377,458,1376,1400]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="1" pageNumber="56">Tristán</collectingCountry>
, leg.; CARTAGO PROVINCE: 5, 
<taxonomicName id="A6944D15802A8E4E7666F949FC74FA8B" box="[854,915,1377,1400]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="irazu">Irazu</taxonomicName>
, W. Slope, 
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, 
<date id="152A1056802A8E4E714CF948FB0FFA8B" box="[1148,1256,1376,1400]" pageId="1" pageNumber="56" value="1926-04-30">30–iv–26</date>
, F. Nevermann, colr. 
<typeStatus id="BE2F8834802A8E4E75E1F9ADFED9FA6E" box="[209,318,1413,1437]" pageId="1" pageNumber="56" type="paratype">Paratypes</typeStatus>
are deposited in 
<collectionCode id="0785AE53802A8E4E7735F9ACFDB6FA6F" box="[517,593,1412,1436]" pageId="1" pageNumber="56">CASC</collectionCode>
, 
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, 
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, 
<collectionCode id="0785AE53802A8E4E762AF9ACFC89FA6F" LSID="urn:lsid:biocol.org:col:34802" box="[794,878,1412,1436]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="56">LACM</collectionCode>
, 
<collectionCode id="0785AE53802A8E4E764BF9ACFC26FA6F" box="[891,961,1412,1436]" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="1" pageNumber="56">INBC</collectionCode>
, 
<collectionCode id="0785AE53802A8E4E76FEF9ACFBF0FA6F" box="[974,1047,1412,1436]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="56">FSCA</collectionCode>
, 
<collectionCode id="0785AE53802A8E4E7114F9ACFB92FA6F" LSID="urn:lsid:biocol.org:col:1013" box="[1060,1141,1412,1436]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="1" pageNumber="56">EMEC</collectionCode>
, 
<collectionCode id="0785AE53802A8E4E71B2F9ACFB35FA6F" box="[1154,1234,1412,1436]" httpUri="http://grbio.org/cool/m2jc-wmeq" name="University of California, Riverside" pageId="1" pageNumber="56">UCRC</collectionCode>
, 
<collectionCode id="0785AE53802A8E4E71EFF9ACFAD4FA6F" LSID="urn:lsid:biocol.org:col:34868" box="[1247,1331,1412,1436]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="1" pageNumber="56">UNSM</collectionCode>
, 
<collectionCode id="0785AE53802A8E4E700FF9ADFA70FA6F" box="[1343,1431,1413,1436]" httpUri="http://grbio.org/cool/brtj-zfcn" name="W.F. Barr Entomological Collection" pageId="1" pageNumber="56">WFBM</collectionCode>
, 
<collectionCode id="0785AE53802A8E4E75A7F980FF39FA33" box="[151,222,1448,1472]" pageId="1" pageNumber="56">JNRC</collectionCode>
, 
<collectionCode id="0785AE53802A8E4E75D9F980FED9FA33" box="[233,318,1448,1472]" pageId="1" pageNumber="56">WOPC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="298E651D802A8E4E75F7F9E3FCCBF8DB" pageId="1" pageNumber="56" type="diagnosis">
<paragraph id="612B3696802A8E4E75F7F9E3FCCBF8DB" blockId="1.[151,1437,619,2012]" pageId="1" pageNumber="56">
<emphasis id="53E0EA84802A8E4E75F7F9E3FEA6FA17" bold="true" box="[199,321,1483,1508]" pageId="1" pageNumber="56">Diagnosis.</emphasis>
Distinguishable from its congeners on the basis of color and elytral sculpturing. 
<taxonomicName id="A6944D15802A8E4E7032F9E4FA7CFA17" box="[1282,1435,1484,1508]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="ska">
<emphasis id="53E0EA84802A8E4E7032F9E4FA7CFA17" box="[1282,1435,1484,1508]" italics="true" pageId="1" pageNumber="56">Colyphus ska</emphasis>
</taxonomicName>
appears most similar to another Costa Rican species, 
<taxonomicName id="A6944D15802A8E4E77D7F9D8FBE3F9FB" authority="Rifkind, 1997" authorityName="Rifkind" authorityYear="1997" box="[743,1028,1520,1544]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="hansoni">
<emphasis id="53E0EA84802A8E4E77D7F9D8FC87F9FB" box="[743,864,1520,1544]" italics="true" pageId="1" pageNumber="56">C. hansoni</emphasis>
<bibRefCitation id="05054B67802A8E4E7656F9D8FBE3F9FB" author="Rifkind" box="[870,1028,1520,1544]" pageId="1" pageNumber="56" refString="Rifkind, J. (1997) Two new species of Cleridae (Coleoptera) from a Costa Rican cloudforest. Revista de Biologia Tropical, 45, 1117 - 1124." type="journal article" year="1997">Rifkind, 1997</bibRefCitation>
</taxonomicName>
, with which it is sympatric in part of its range. The species are easily separable based on color pattern: 
<taxonomicName id="A6944D15802A8E4E76B2FA3DFC1BF9DF" box="[898,1020,1556,1580]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="hansoni">
<emphasis id="53E0EA84802A8E4E76B2FA3DFC1BF9DF" box="[898,1020,1556,1580]" italics="true" pageId="1" pageNumber="56">C. hansoni</emphasis>
</taxonomicName>
is tricolorous (red, ivory and black), whereas the new species is strictly bicolorous (stramineous and black). 
<bibRefCitation id="05054B67802A8E4E76F2FA10FB8DF9A3" author="Rifkind" box="[962,1130,1592,1616]" pageId="1" pageNumber="56" refString="Rifkind, J. (1997) Two new species of Cleridae (Coleoptera) from a Costa Rican cloudforest. Revista de Biologia Tropical, 45, 1117 - 1124." type="journal article" year="1997">Rifkind (1997)</bibRefCitation>
has discussed the relationship of these 
<taxonomicName id="A6944D15802A8E4E7402FA74FE7BF987" box="[306,412,1628,1652]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="53E0EA84802A8E4E7402FA74FE7BF987" box="[306,412,1628,1652]" italics="true" pageId="1" pageNumber="56">Colyphus</emphasis>
</taxonomicName>
species with the enigmatic Costa Rican species 
<taxonomicName id="A6944D15802A8E4E76FAFA74FA7FF987" authority="Gorham, 1882" authorityName="Gorham" authorityYear="1882" box="[970,1432,1628,1652]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="subviolaceus">
<emphasis id="53E0EA84802A8E4E76FAFA74FB31F987" box="[970,1238,1628,1652]" italics="true" pageId="1" pageNumber="56">Enoclerus subviolaceus</emphasis>
(
<bibRefCitation id="05054B67802A8E4E71D6FA74FA68F987" author="Gorham" box="[1254,1423,1628,1652]" pageId="1" pageNumber="56" refString="Gorham, H. S. (1882) Family Cleridae. In Goodman and Salvin (editors). Biologia Centrali-Americana. Insecta, Coleoptera 3 (2), 129 - 168. Taylor &amp; Francis, London." type="journal article" year="1882">Gorham, 1882</bibRefCitation>
)
</taxonomicName>
. They share a similar facies, sexually dimorphic tarsal claws and prominent subbasal tumescences, but 
<taxonomicName id="A6944D15802A8E4E7024FAA9FF2EF94F" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="subviolaceus">
<emphasis id="53E0EA84802A8E4E7024FAA9FF2EF94F" italics="true" pageId="1" pageNumber="56">E. subviolaceus</emphasis>
</taxonomicName>
has the elytral surface dotted with pustular elevations, and the male lacks the sagittate condition of the apical region of the tegmen. We mention these distinctions in detail, because we have examined two specimens of 
<taxonomicName id="A6944D15802A8E4E707DFAE1FEE6F8F7" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="subviolaceus">
<emphasis id="53E0EA84802A8E4E707DFAE1FEE6F8F7" italics="true" pageId="1" pageNumber="56">E. subviolaceus</emphasis>
</taxonomicName>
(
<figureCitation id="F9AF2A13802A8E4E743FFAC4FEB5F8F7" box="[271,338,1772,1796]" captionStart="FIGURE 2" captionStartId="2.[151,250,2010,2033]" captionTargetBox="[544,1024,1124,1988]" captionTargetId="figure@2.[544,1042,1124,1988]" captionTargetPageId="2" captionText="FIGURE 2. Habitus of Enoclerus subviolaceus (Gorham, 1882), from Monteverde, Puntarenas Province, Costa Rica." httpUri="https://zenodo.org/record/200831/files/figure.png" pageId="1" pageNumber="56">Fig. 2</figureCitation>
represents a specimen from Monteverde, Puntarenas Province, 
<collectingCountry id="19837606802A8E4E712AFAC4FB71F8F7" box="[1050,1174,1772,1796]" name="Costa Rica" pageId="1" pageNumber="56">Costa Rica</collectingCountry>
) that bear an uncannily identical color pattern to many specimens of 
<taxonomicName id="A6944D15802A8E4E77BFFB38FCC2F8DB" box="[655,805,1808,1832]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="ska">
<emphasis id="53E0EA84802A8E4E77BFFB38FCC2F8DB" box="[655,805,1808,1832]" italics="true" pageId="1" pageNumber="56">Colyphus ska</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="298E651D802A8E4C75F7FB1BFBE2FA8B" lastPageId="3" lastPageNumber="58" pageId="1" pageNumber="56" type="description">
<paragraph id="612B3696802A8E4C75F7FB1BFE2CFB53" blockId="1.[151,1437,619,2012]" lastBlockId="3.[151,1437,151,1725]" lastPageId="3" lastPageNumber="58" pageId="1" pageNumber="56">
<emphasis id="53E0EA84802A8E4E75F7FB1BFE05F8BF" bold="true" box="[199,482,1843,1868]" pageId="1" pageNumber="56">
Description (
<typeStatus id="BE2F8834802A8E4E7453FB1BFE32F8BF" box="[355,469,1843,1868]" pageId="1" pageNumber="56" type="holotype">Holotype</typeStatus>
).
</emphasis>
Length: 9.70 mm. Form: medium sized, elongate. Color: integument black except antennomeres 
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part testaceous, basal 1/2 of elytra covered by a broad stramineous band, complete to lateral margins, sinuate posteriorly, narrowly extended posteriorly along suture; each elytron marked with a small, reddish-brown macula on the umbone, on the post-humeral tubercle, and again before the middle; a narrow transverse irregular stramineous fascia present at elytral posterior 7/9, narrowly interrupted at lateral margins and at suture; elytral apices marked with a vague stramineous spot. Head: surface shining, finely, densely punctate; setation sparse, composed of mostly short, rather robust white setae, with a thin intermingling of longer, finer black setae; front distinctly bi-impressed; eyes moderately convex, emargination shorter than length of antennal scape; antenna not quite attaining base of pronotum; antennomeres 8–11 forming a rather loose, gradually expanded club, 11 elongate, inner margin subsinuate, with apex subacute. Gular process rectangulate. Pronotum: convex, about as wide as head measured across eyes, narrower than base of elytra; surface shining, shallowly and densely punctate, feebly, transversely rugulose at sides; vestiture consisting of a rather sparse covering of long, erect black setae with a thin admixture of long, erect white setae laterally, posteriorly on the disk, and on the collar; anterior transverse impression deeply incised across disk, broadly U-shaped at middle; lateral foveae indistinct, anterior margin transverse, sides constricted at anterior transverse impression, convexly expanded to maximum width just before middle, then arcuately narrowed to moderately prolonged, subparallel sides of basal collar; posterior pronotal slope acute. Scutellum rounded apically, surface set with pale setae. Elytra: more than 
<date id="152A105680288E4C76E9FE0CFC13FDCF" box="[985,1012,548,572]" pageId="3" pageNumber="58">2x</date>
as long as wide, widest just posterior to middle, surface shining, rather densely and deeply punctate, punctures smaller and more shallow on apical 1/2; vestiture as follows: anterior stramineous area thinly covered with a mixture of mostly medium length, pale and black, suberect and erect setae; posterior black area a little more densely vested with mostly medium length, suberect setae, except subapical pale fascia and pale apices thinly set with reclinate whitish setae; disk subflattened above, subbasal tumescences pronounced, tuberculate, with crests smooth and slightly longitudinally expanded; umbones prominent, slightly roughened; sides subparallel at anterior 1/3, then slightly divergent to about apical 1/ 3, and thereafter arcuately convergent to dehiscent, subacute apices. Apical slope gradual. Legs: profemora considerably more expanded at middle than meso- and metafemora; tarsal pulvilli well developed; tarsal claws as follows: first and second pair with each claw bearing a rather long, recurved, chisel-shaped tooth; third pair with teeth stout and triangular. Mesosternum: posterior median process narrow, elongate, moderately elevated apically. Metasternum: convex; surface densely, minutely punctate and roughened, and moderately densely vested with reclinate silvery setae at sides, reflective and almost glabrous at middle; anterior medium process sulcate, elevated anteriorly. Abdomen: visible sternites 1–5 sparsely, irregularly punctate, sparsely set with scattered fine, short and long, silvery setae; sternite 5 with sides slightly convergent posteriorly and hind margin very feebly, broadly, arcuately emarginate; hind margin of sternite 6 deeply, rather narrowly, arcuately emarginate, with hind angles subacute; tergite 6 with apex broadly rounded, projecting well past hind margin of sternite 6. Genitalia: subapical region of the tegmen feebly sagittate.
</paragraph>
<caption id="35EB661E80298E4D75A7F833FC42FBC2" box="[151,933,1051,1074]" pageId="2" pageNumber="57">
<paragraph id="612B369680298E4D75A7F833FC42FBC2" blockId="2.[151,933,1051,1074]" box="[151,933,1051,1074]" pageId="2" pageNumber="57">
<emphasis id="53E0EA8480298E4D75A7F833FEF4FBC2" bold="true" box="[151,275,1051,1074]" pageId="2" pageNumber="57">FIGURE 1.</emphasis>
Habitus of 
<taxonomicName id="A6944D1580298E4D74B9F834FD53FBC1" authority="Rifkind &amp; Barr" authorityName="Rifkind &amp; Barr" box="[393,692,1051,1074]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="ska" status="sp. nov.">
<emphasis id="53E0EA8480298E4D74B9F834FDF6FBC1" box="[393,529,1052,1074]" italics="true" pageId="2" pageNumber="57">Colyphus ska</emphasis>
Rifkind &amp; Barr
</taxonomicName>
, 
<emphasis id="53E0EA8480298E4D778EF834FD10FBC2" bold="true" box="[702,759,1052,1073]" pageId="2" pageNumber="57">
<taxonomicNameLabel id="48D357FF80298E4D778EF834FD10FBC2" box="[702,759,1052,1073]" pageId="2" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(holotype male).
</paragraph>
</caption>
<caption id="35EB661E80298E4D75A7FBF2FAA7F802" box="[151,1344,2010,2034]" httpUri="https://zenodo.org/record/200831/files/figure.png" pageId="2" pageNumber="57" targetBox="[544,1024,1124,1988]" targetPageId="2">
<paragraph id="612B369680298E4D75A7FBF2FAA7F802" blockId="2.[151,1344,2010,2034]" box="[151,1344,2010,2034]" pageId="2" pageNumber="57">
<emphasis id="53E0EA8480298E4D75A7FBF2FEF4F803" bold="true" box="[151,275,2010,2033]" pageId="2" pageNumber="57">FIGURE 2.</emphasis>
Habitus of 
<taxonomicName id="A6944D1580298E4D74B9FBF3FCC3F801" authority="Gorham, 1882" authorityName="Gorham" authorityYear="1882" box="[393,804,2011,2034]" class="Insecta" family="Cleridae" genus="Enoclerus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="subviolaceus">
<emphasis id="53E0EA8480298E4D74B9FBF3FD9EF802" box="[393,633,2011,2033]" italics="true" pageId="2" pageNumber="57">Enoclerus subviolaceus</emphasis>
(Gorham, 1882)
</taxonomicName>
, from Monteverde, Puntarenas Province, Costa Rica.
</paragraph>
</caption>
<paragraph id="612B369680288E4C75F7F883FBE2FA8B" blockId="3.[151,1437,151,1725]" pageId="3" pageNumber="58">
<emphasis id="53E0EA8480288E4C75F7F883FEDCFB37" bold="true" box="[199,315,1195,1220]" pageId="3" pageNumber="58">Variation.</emphasis>
Length ranges from 7.50 to 11.75mm. The extent and delineation of the elytral stramineous fasciae and dark maculae are rather variable across the 
<typeStatus id="BE2F883480288E4C7798F8F9FD3FFB1A" box="[680,728,1233,1257]" pageId="3" pageNumber="58">type</typeStatus>
series, and one specimen has the dark elytral pigmentation radically reduced. In some individuals, the subapical stramineous band is reduced or entirely absent, with the darkened integument interrupted only by a fascia of pale setae. Unlike those of the male, the tarsal claws of the female all bear stout, triangular teeth. Sternite 
<quantity id="A66C9B7380288E4C771CF914FDBFFAA7" box="[556,600,1340,1364]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="58" unit="in" value="5.0">5 in</quantity>
the female has sides roundly tapering to a rather broad subtruncate posterior margin; sternite 6 and tergite 6 with posterior margins conjointly rounded.
</paragraph>
</subSubSection>
<subSubSection id="298E651D80288E4C75F7F9ACFC5CFA33" pageId="3" pageNumber="58" type="etymology">
<paragraph id="612B369680288E4C75F7F9ACFC5CFA33" blockId="3.[151,1437,151,1725]" pageId="3" pageNumber="58">
<emphasis id="53E0EA8480288E4C75F7F9ACFEA9FA6F" bold="true" box="[199,334,1412,1436]" pageId="3" pageNumber="58">Etymology.</emphasis>
The specific name (a noun in apposition) derives from the second 
<taxonomicName id="A6944D1580288E4C7101F9ACFBB1FA6F" box="[1073,1110,1412,1436]" class="Insecta" family="Cleridae" genus="Colyphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="ska">ska</taxonomicName>
wave (ca. 1978), also known as the &quot;2 Tone&quot; genre, a reference to the new species' bicolorous aspect.
</paragraph>
</subSubSection>
<subSubSection id="298E651D80288E4C75F7F9E3FE42F9FB" pageId="3" pageNumber="58" type="distribution">
<paragraph id="612B369680288E4C75F7F9E3FE42F9FB" blockId="3.[151,1437,151,1725]" pageId="3" pageNumber="58">
<emphasis id="53E0EA8480288E4C75F7F9E3FE87FA17" bold="true" box="[199,352,1483,1508]" pageId="3" pageNumber="58">Distribution.</emphasis>
Known from the provinces of San José and Cartago, and the Monteverde region of Puntarenas province, in 
<collectingCountry id="1983760680288E4C7413F9D8FE46F9FB" box="[291,417,1520,1544]" name="Costa Rica" pageId="3" pageNumber="58">Costa Rica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="298E651D80288E4C75F7FA3BFB9DF94F" pageId="3" pageNumber="58" type="biology_ecology">
<paragraph id="612B369680288E4C75F7FA3BFB9DF94F" blockId="3.[151,1437,151,1725]" pageId="3" pageNumber="58">
<emphasis id="53E0EA8480288E4C75F7FA3BFEC0F9DF" bold="true" box="[199,295,1555,1580]" pageId="3" pageNumber="58">Biology.</emphasis>
All specimens were collected at elevations above 
<quantity id="A66C9B7380288E4C7658FA3CFC5FF9DF" box="[872,952,1556,1580]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="58" unit="m" value="1300.0">1300m</quantity>
, mostly in cloud forest habitat. Specimens have been taken in Malaise traps, by beating living and cut vegetation, and at U.V. light. One individual was taken from a sapping tree. This is a relatively common, rather large sized clerid, with a moderately extensive geographical distribution: it is puzzling that it should have eluded discovery and description by the 19th and early 20th century coleopterists who made such great efforts to collect and name Costa Rican 
<taxonomicName id="A6944D1580288E4C7123FA8CFB91F94F" box="[1043,1142,1700,1724]" class="Insecta" family="Cleridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="family">Cleridae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>