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<document ID-CLB-Dataset="72036" ID-DOI="http://dx.doi.org/10.3897/dez.69.83840" ID-GBIF-Dataset="eafc7326-93e2-4d55-b708-2da4b77e2e5e" ID-Pensoft-Pub="1860-1324-1-71" ID-Pensoft-UUID="DF59A16543B05799A4FD7B66E721FC76" ID-ZooBank="7DEB0587A3734B06BAADCA6102EC6581" ModsDocID="1860-1324-69-1-71" checkinTime="1655205803791" checkinUser="pensoft" docAuthor="Gueorguiev, Borislav, Wrase, David W., Assmann, Thorsten, Muilwijk, Jan &amp; Machard, Patrice" docDate="2022" docId="D816CC852981571EA5B1E555A3DEC960" docLanguage="en" docName="DeutEntomolZeit 69(1): 71-122" docOrigin="Deutsche Entomologische Zeitschrift 69 (1)" docPubDate="2022-06-13" docSource="http://dx.doi.org/10.3897/dez.69.83840" docTitle="Platyderus (Eremoderus) brunki Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard 2022, sp. nov." docType="treatment" docUuid="C5B5E518-02FF-4E24-A1C1-70F560699170" docUuidSource="ZooBank" docVersion="6" id="DF59A16543B05799A4FD7B66E721FC76" lastPageNumber="71" masterDocId="DF59A16543B05799A4FD7B66E721FC76" masterDocTitle="Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)" masterLastPageNumber="122" masterPageNumber="71" pageNumber="71" updateTime="1732679132351" updateUser="ExternalLinkService">
<mods:mods id="647AC23741F8960C9B2CB9A65EE37863" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="59CC2B095388BAFCBE5D6B67B3362ACF">Taxonomic revision of the African and Southwest Asian species of Platyderus Stephens, subg. Eremoderus Jeanne (Coleoptera, Carabidae, Sphodrini)</mods:title>
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<mods:namePart id="530F540E5D0C7C76E8B413562DBCC363">Gueorguiev, Borislav</mods:namePart>
<mods:nameIdentifier id="6DC3E880C6C2422F97E685D4610F6582" type="ORCID">https://orcid.org/0000-0002-8532-0657</mods:nameIdentifier>
<mods:affiliation id="053BF1C090EFD6C3FBDDE853314042EF">National Museum of Natural History, Bulgarian Academy of Sciences, 1 Blvd. Tsar Osvoboditel, 1000 Sofia, Bulgaria</mods:affiliation>
<mods:nameIdentifier id="E3DC072A74A5020E6BEDF7384A236967" type="email">gueorguiev@nmnhs.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2CDE947E5CCF933ADC4A8F9E2D0457C1" type="personal">
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<mods:namePart id="B108149B4F77352770217E8C3DF6BEFA">Wrase, David W.</mods:namePart>
<mods:nameIdentifier id="326BD9A1531F65C7E91B003502AB8402" type="ORCID">https://orcid.org/0000-0003-1779-2684</mods:nameIdentifier>
<mods:affiliation id="B57FD42D357EDA7722735917C66DFE99">Oderstr. 2, 15306, Gusow-Platkow, Germany</mods:affiliation>
</mods:name>
<mods:name id="CCB2549795F0A194C7D59FD6E883B1AD" type="personal">
<mods:role id="27BF1E57637DC9B20DCC9EBE79E52883">
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<mods:namePart id="45E8F70E5F5D03236500D019FA11DCDB">Assmann, Thorsten</mods:namePart>
<mods:nameIdentifier id="E096E2DA9CD1FC74E724707F8523D5C7" type="ORCID">https://orcid.org/0000-0002-9203-769X</mods:nameIdentifier>
<mods:affiliation id="486A61365B9E0206A62AAB8100F78FC7">Leuphana University Lueneburg, Institute of Ecology, Universitaetsallee 1, 21335 Lueneburg, Germany</mods:affiliation>
</mods:name>
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<mods:namePart id="10EF851D45A5F8BC917BF285EEE0B9DC">Muilwijk, Jan</mods:namePart>
<mods:affiliation id="7853DAB4BA757207EC4F87D168220A2D">Department of Entomology, Naturalis Biodiversity Centre, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
</mods:name>
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<mods:namePart id="F06461571D6F9BD955BDF5574C0FB686">Machard, Patrice</mods:namePart>
<mods:affiliation id="717A8217855E5C67486886FED69D249C">29 Chemin de Champigny, Molineuf, F- 41190 Valencisse, France</mods:affiliation>
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<mods:title id="AB7CE6A6D38018A42C7C468110B9F878">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:date id="2F00A863A942E2D1FF49242E88F61BB2">2022</mods:date>
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<treatment id="D816CC852981571EA5B1E555A3DEC960" ID-GBIF-Taxon="195868741" LSID="urn:lsid:zoobank.org:act:C5B5E518-02FF-4E24-A1C1-70F560699170" httpUri="http://treatment.plazi.org/id/D816CC852981571EA5B1E555A3DEC960" lastPageNumber="71" pageId="0" pageNumber="71" scope_family="Carabidae" scope_order="Coleoptera" scope_subGenus="Platyderus" scope_tribe="Sphodrini">
<subSubSection id="A9500031E3AAC7E510D4617ECE38A389" pageId="0" pageNumber="71" type="nomenclature">
<paragraph id="03E037DCCB7F7330AEF0CEE3F3A24E47" pageId="0" pageNumber="71">
14.
<taxonomicName id="0ECCB9A3FBECE250EA7C6ACE5492B027" LSID="http://zoobank.org/C5B5E518-02FF-4E24-A1C1-70F560699170" authority="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard, 2022" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki" status="sp. nov." subGenus="Eremoderus">Platyderus (Eremoderus) brunki</taxonomicName>
<taxonomicNameLabel id="79EE0F4E2E65C38F580F0200F7990736" pageId="0" pageNumber="71">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="EF4EF542AD41BDF96117896548AB61E1" pageId="0" pageNumber="71" type="description">
<paragraph id="419D06C5BBECED231E2AE34446C86E20" pageId="0" pageNumber="71">
<figureCitation id="C8F68E13A8A23E7301C52580441EE82B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">Figs 3C</figureCitation>
<figureCitation id="47B763F39EE6C89F5BF6069F18E874AE" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">, 14F</figureCitation>
<figureCitation id="0181E6B62C14CEB7101D21F8F9F5FB28" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="7E61355898DDC8C7C8F8206F0411E19F" pageId="0" pageNumber="71" type="type locality">
<paragraph id="F7E905F9019D4BF13C8D0E990743FE22" pageId="0" pageNumber="71">Type locality.</paragraph>
<paragraph id="190CCBE8E4E87B97260A329031A62D2A" pageId="0" pageNumber="71">Yemen, 'Amran Governorate, Thula District, between Kaukaban and Shibam, approximate GPS coordinates: 15.503, 43.901.</paragraph>
</subSubSection>
<subSubSection id="A198F7549074E449D563AC6175374E96" pageId="0" pageNumber="71" type="type material">
<paragraph id="3DD82EA3325B0ED9F2BF8E38854A59CE" pageId="0" pageNumber="71">Type material.</paragraph>
<paragraph id="B808AD9E4C0BE003FD1AE985737A2385" pageId="0" pageNumber="71">
<emphasis id="E6AB8ADCE3702774DCACE6C447536221" bold="true" italics="true" pageId="0" pageNumber="71">Holotype</emphasis>
♀, 'YEMEN, ~ 40 km NW Sanaa, / Kaukaban/Shibam, small waste- / waterstream, ruralic vegetation, / sand, 2700-2930 m Handaufsam. /
<geoCoordinate id="038E34B8604E81C2E793C600A4893621" degrees="15" direction="north" minutes="30" orientation="latitude" precision="1" seconds="9.98" value="15.502772">15°30'9.98&quot;N</geoCoordinate>
,
<geoCoordinate id="27201D6A6A3DF56C7C953FE0F8547D2B" degrees="43" direction="east" minutes="54" orientation="longitude" precision="1" seconds="2.66" value="43.90074">43°54'2.66&quot;E</geoCoordinate>
/ 02. Dez. 2009, legit. Ingo Brunk [p, w] // Coll. / I. Brunk / Dresden/Germ [h, pn]' (cBRU).
</paragraph>
</subSubSection>
<subSubSection id="0101859E89DD6A70FA0B77C4C1D985F2" pageId="0" pageNumber="71" type="materials_examined">
<paragraph id="7288BF644BF9AA45826C48D4F9529404" pageId="0" pageNumber="71">Other material examined.</paragraph>
<paragraph id="BCFB34D9E5AECFFD536C5700A718F636" pageId="0" pageNumber="71">
<materialsCitation id="AFF509186082773F1F7D638E8BB34A32" collectingDate="1938-02-03" collectorName="El Errein, B. M. Exp., S. W. Arabia., H. Scott, E. B. Britton., Reiche" country="YEMEN" county="Remains" determinerName="E. B. Britton" elevation="2835" location="Haz" municipality="El Errein" specimenCount="1" stateProvince="San'a">
<collectingCounty id="3778818EE4F536D38FFB2E7BB2930058">Remains</collectingCounty>
of a specimen of uncertain sex in bad condition (only pronotum and elytra preserved), 'UNDER A STONE //
<collectingCountry id="AD307D33B06856E4B8C121FE889DDAF5" name="Yemen">YEMEN</collectingCountry>
/
<collectorName id="F0D8EFC4A129C9E08395B0C9A27C938A">
<collectingMunicipality id="BE6FC64FC49BCC1064D67E13596FADE7">El Errein</collectingMunicipality>
</collectorName>
, nr.
<location id="3E358768E5D3A423AA89691B0268ECCE" LSID="urn:lsid:plazi:treatment:D816CC852981571EA5B1E555A3DEC960:3E358768E5D3A423AA89691B0268ECCE" country="YEMEN" county="Remains" municipality="El Errein" name="Haz" stateProvince="San'a">Haz</location>
, / about
<quantity id="32C5BDACCC15151EDD1DA48E364BC047" metricMagnitude="4" metricUnit="m" metricValue="2.5749504" unit="mi" value="16.0">16 miles</quantity>
N.W. / of
<normalizedToken id="9D4B708F120033C11771FF439A88A41B" originalValue="San’a">
<collectingRegion id="B99147F8F35C73EAAFF38E3642E94B3B" country="Yemen" name="San'a'">San'a</collectingRegion>
</normalizedToken>
, ca.
<quantity id="F188696F4162FFA38CF5B99D471D1587" metricMagnitude="3" metricUnit="m" metricValue="2.83464" unit="ft" value="9300.0">
<elevation id="5691CF37ECC0D8E069DD068F58CB79F1" metricMagnitude="3" metricUnit="m" metricValue="2.83464" unit="ft" value="9300.0">9,300ft.</elevation>
</quantity>
[=
<quantity id="2B6AB52C74B7534C4E8D70CCF5613BC5" metricMagnitude="3" metricUnit="m" metricValue="2.835" unit="m" value="2835.0">
<elevation id="D98AADB3E537F517D5725F9AFABC724F" metricMagnitude="3" metricUnit="m" metricValue="2.835" unit="m" value="2835.0">2835 m</elevation>
</quantity>
] /
<collectingDate id="D3F830DBE79DF770BD681521AB625C3D" value="1938-02-03">3.ii.1938</collectingDate>
.//
<collectorName id="9381707295BEE83F4CAF60A03714F0B7">B.M. Exp.</collectorName>
to /
<collectorName id="DF510613F65866AF742604936C828014">S.W. Arabia.</collectorName>
/
<collectorName id="99F812D16BF3643195F5386701B60254">H. Scott</collectorName>
&amp; /
<collectorName id="F4FE4B749CBDCC3790F10D9553589A20">E.B. Britton.</collectorName>
/ B.M. 1938-246, //
<taxonomicName id="8D34C6279A6257256A2705D0DFB5F7FE" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">Platyderus</taxonomicName>
/
<taxonomicName id="190BBEF500F297211B697DDEB11AD7CC" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">Platyderus Eremoderus languidus</taxonomicName>
/
<collectorName id="E7D03581FEB4410F2C629371EABEAEFF">Reiche</collectorName>
/
<determinerName id="99E438DE862692A7035A6C1F75F1CF91">E.B. Britton</determinerName>
det. / 1946' (NHMUK)
</materialsCitation>
.
</paragraph>
<paragraph id="7F416094CF82CD27C52FAE5AEEA09C1C" pageId="0" pageNumber="71">
TME:
<specimenCount id="6CF7949C8494182953170400206750B2" type="generic">2 specimens</specimenCount>
. TGE:
<specimenCount id="5DAED92502806DA4345E3CECF5A43384" type="female">1♀</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C9B6EAE2A170F4A6D205726BBD830CE7" pageId="0" pageNumber="71" type="etymology">
<paragraph id="033B1F841E1F8C96E7E07C9800DE4B9E" pageId="0" pageNumber="71">Etymology.</paragraph>
<paragraph id="6D897BCF2543FB6C215123567970433F" pageId="0" pageNumber="71">Latinized patronym based on the surname of Ingo Brunk (Dresden, Germany), who collected the first well-preserved representative of the new species.</paragraph>
</subSubSection>
<subSubSection id="E61C8889D659B77EC38D5AE911C1A764" pageId="0" pageNumber="71" type="diagnosis">
<paragraph id="A53B3C620306195FE879D7ACBCF1AFE1" pageId="0" pageNumber="71">Diagnosis.</paragraph>
<paragraph id="B9377375612234005D1B11FC74EF6BA1" pageId="0" pageNumber="71">
<taxonomicName id="A2AC75101974B65D0271822B20AECF04" authorityName="Guéorguiev &amp; Wrase &amp; Assmann &amp; Muilwijk &amp; Machard" authorityYear="2022" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus brunki" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="brunki">
<emphasis id="5C83AD0E5AB6D40FC4BFFE0990FB2259" italics="true" pageId="0" pageNumber="71">Platyderus brunki</emphasis>
</taxonomicName>
sp. nov. readily differs from all other
<taxonomicName id="3B574C0DC2EA31078CC017900B56E76A" authorityName="Jeanne" authorityYear="1996" class="Insecta" family="Carabidae" genus="Eremoderus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Eremoderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="9E25AFE420F428512BF6B167F30D7892" italics="true" pageId="0" pageNumber="71">Eremoderus</emphasis>
</taxonomicName>
-taxa in its smallest value in ratio PW/PL (Table
<tableCitation id="74AFD47F8862DC5AE2E478A59F0CBC4D" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
). It is most closely related to
<taxonomicName id="AA1669BC90B6D0D178344B25EBCB6D39" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="B6B83DBA332E276B85E8F1DF6B3F59B6" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
; both species are the only two within &quot;
<taxonomicName id="43ACF063EF816521C633226FC3B26768" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="91097CFC095776D08B05B48496A95F06" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group with darkest coloration of integument and sides of pronotum not sinuate from widest point to base (
<taxonomicName id="83849AC37FB84B6AB9BD6AACD58E0E8F" class="Insecta" kingdom="Animalia" lsidName="P. irakensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="irakensis">
<emphasis id="C3E119084269F072B9A9E23436AC0263" italics="true" pageId="0" pageNumber="71">P. irakensis</emphasis>
</taxonomicName>
has also sides of pronotum not sinuate to base, but much lighter coloration of integument and higher value in ratio EL/PL). The new species is distinct from
<taxonomicName id="B2554F1FFBB1801BCB89CF20E6B553D3" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="FEBEC51BD7D05E5521E2C953C1BC6432" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov., in: (1) pronotum with anterior angles more rounded at tips (vs. pronotum with anterior angles more pointed at tips, Fig.
<figureCitation id="B6C4D1300B2262741A679C94EFB53994" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3B, C</figureCitation>
); (2) pronotum nearly as long as wide, PW/PL: 1.02-1.05 (vs. pronotum appreciably wider than long, PW/PL: 1.07-1.11); (3) pronotum subcordate, less constricted anteriorly, PW/PA: 1.38-1.42, with greater value in ratio PA/PB: 0.80-0.88 (vs. pronotum subquadrate, more constricted anteriorly, PW/PA: 1.44-1.48, with smaller value in ratio PA/PB: 0.76-0.79); (4) elytra compared to pronotum slightly shorter, EL/PL: 2.40-2.44 (vs. elytra compared to pronotum slightly longer, EL/PL: 2.47-2.56). Females are different from all other representatives of the &quot;
<taxonomicName id="7F15FE88B52EB3E766E3597C9D3AB5B7" baseAuthorityName="Reiche &amp; Saulcy" baseAuthorityYear="1855" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) languidus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="languidus" subGenus="Eremoderus">
<emphasis id="B25E456D7D85E87CCD37337819494F6B" italics="true" pageId="0" pageNumber="71">Platyderus Eremoderus languidus</emphasis>
</taxonomicName>
&quot; group by the presence of two, instead of one, dorsolateral ensiform setae on the apical gonocoxite (Fig.
<figureCitation id="4D6F55E42F2EB21FA4F1900FF3CEE496" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A42053743EC3926373B21B894DA16CEA" pageId="0" pageNumber="71" type="description">
<paragraph id="590A9A7E00303D9359AEB612CA33F186" pageId="0" pageNumber="71">Description.</paragraph>
<paragraph id="7F81BBA8564B7F37D8097B02D321E3E7" pageId="0" pageNumber="71">
<emphasis id="FFC4CD173C06101A67E1C78A61284961" bold="true" pageId="0" pageNumber="71">
<emphasis id="DDCF49BA781D94359A62FE2E3FAE8C2A" italics="true" pageId="0" pageNumber="71">Habitus</emphasis>
.
</emphasis>
Specimens of large size for
<taxonomicName id="FAB1C20C448EAE341FDF220342AE778A" authorityName="Stephens" authorityYear="1827" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="FEB66EC360187972794F7BCD7F22E36F" italics="true" pageId="0" pageNumber="71">Platyderus</emphasis>
</taxonomicName>
species (BL of holotype: 9.50 mm; BW: 3.30 mm), with elongate, moderately convex body (Fig.
<figureCitation id="BB675E84F5ACC07BB99F6AAA9A37B134" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal habitus. A. Platyderus (Eremoderus) afghanistanicus, sp. nov., holotype; B. P. (E.) arabicus, sp. nov., holotype; C. P. (E.) brunki, sp. nov., holotype; D. P. (E.) brunneus brunneus Karsch, 1881, lectotype and its labels; E. P. (E.) brunneus brunneus Karsch, 1881, male specimen, 50 km W of Ben Gardane, Medenine Governorate, Tunisia; F. P. (E.) brunneus ferrantei Reitter, 1909, lectotype and its labels. Scale bars: 1 mm." figureDoi="10.3897/dez.69.83840.figure3" httpUri="https://binary.pensoft.net/fig/695640" pageId="0" pageNumber="71">3C</figureCitation>
).
<emphasis id="47D5BD8718D8F2508DA9FB4B3BA090E1" bold="true" pageId="0" pageNumber="71">
<emphasis id="4D3CAC33A4640BAAAA33CFAA1E9144C2" italics="true" pageId="0" pageNumber="71">Measurements and ratios</emphasis>
.
</emphasis>
See Table
<tableCitation id="FA433765B7A6BED3ED3B22A66B20E690" captionStart="Table 3" captionStartId="T3" captionText="Table 3. Morphometric data for species of the ' Platyderus Eremoderus languidus ' group of Platyderus." httpUri="http://table.plazi.org/id/D5F1A345CA056C2DBA2597271787B1E8" pageId="0" pageNumber="71" tableUuid="D5F1A345CA056C2DBA2597271787B1E8">3</tableCitation>
.
<emphasis id="7B0454786CE415BEBF72C7D0EC2D74B2" bold="true" pageId="0" pageNumber="71">
<emphasis id="3BAE866FE8EBF06F6C305B2D78C1D83F" italics="true" pageId="0" pageNumber="71">Color and lustre</emphasis>
.
</emphasis>
Body and appendages uniformly reddish-brown, with palpi and antennae slightly lighter. Integument slightly to moderately shiny, head and pronotum shinier than elytra.
<emphasis id="A6B4BAAAA72CF4413D9B6DEA3104C141" bold="true" pageId="0" pageNumber="71">
<emphasis id="C4082DF76557135460015E01D87D7286" italics="true" pageId="0" pageNumber="71">Microsculpture and punctation</emphasis>
.
</emphasis>
Microreticulation of pronotum and elytra as in
<taxonomicName id="DE5B896C17951B344EACB3ECB8E09A74" class="Insecta" kingdom="Animalia" lsidName="P. arabicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="arabicus">
<emphasis id="940E9746A8D5B99618CC7737627BA0B2" italics="true" pageId="0" pageNumber="71">P. arabicus</emphasis>
</taxonomicName>
sp. nov. Ventral surface largely with well-impressed isodiametric or slight transverse sculpticells, epipleura, mesosternum and middle coxa with sculpticells less apparent. Head (holotype) nearly smooth, impunctate, wrinkles present only on clypeus and frons, including frontal foveae and area surrounding them. Pronotum surface largely smooth, only apical part medially in front of anterior transverse impression and basal area medially behind posterior transverse impression longitudinally wrinkled, as well as each side along lateral margin with a few shallow punctures not reaching anterior half. Elytral intervals without apparent punctation. Abdominal ventrites nearly smooth, impunctate, 3-5 with a few fine wrinkles at sides.
<emphasis id="CB3B591A7FB68068CB1F17D34C2D328E" bold="true" italics="true" pageId="0" pageNumber="71">Head</emphasis>
(holotype). About two-thirds as wide as pronotum (PW/HW of holotype: 1.36). Eyes long, slightly convex. Labrum subrectangular, as long as clypeus, with anterior margin concave. Frontoclypeal suture distinct in middle, indistinct at sides. Frontal furrows shallow, subfoveolate. Paraorbital sulci straight, backward barely reaching posterior margin of eye, not reaching level of posterior supraorbital pore.
<emphasis id="58949ED928EB6EBBCB9774C71347214A" bold="true" pageId="0" pageNumber="71">
<emphasis id="77B2937838F3EA7274A34588FBC4B2A3" italics="true" pageId="0" pageNumber="71">Thorax</emphasis>
.
</emphasis>
Pronotum almost as wide as long (PW/PL= 1.02-1.05), with widest point at second quarter (holotype) or medial fifth (specimen from &quot;El Errein, nr. Haz&quot;). Anterior and posterior transverse impressions slightly distinct medially, indistinct laterally. Sides not sinuate, smoothly convex anteriorly and medially, straight (specimen from &quot;El Errein, nr. Haz&quot;) or convex posteriorly (holotype); anterior bead present laterally, lacking in medial 1/8 to 1/10; basal bead present laterally, reduced to absent in medial half; posterior angles rounded (holotype) or obtuse (specimen from &quot;El Errein, nr. Haz&quot;). Metepisternum as long as wide, MA/MI= 1.00.
<emphasis id="CB80510955717E2862FA7983FB9EF5C7" bold="true" pageId="0" pageNumber="71">
<emphasis id="6D285A1BB35D8D0C2274CCF33383817A" italics="true" pageId="0" pageNumber="71">Elytra</emphasis>
.
</emphasis>
Elongate, about one and a half times as long as elytra (EL/EW= 1.58-1.59), two and a half times as long as pronotum (EL/PL= 2.40-2.44), and a one and half as wide as pronotum (EW/PW= 1.48), with widest point at medial third. Parascutellar striole scarcely distinct, very shallow; striae more impressed than striole, barely punctate; parascutellar striole not joining stria 1; striae 1-5 and sometimes 6 reaching basal bead, 7 ending little before. Interval 3 with three discal setiferous punctures, medial one larger than others, in midst of interval 3 (holotype) or near stria 2 (specimen from &quot;El Errein, nr. Haz&quot;), anterior and posterior punctures smaller, adjoining stria 3 (posterior puncture lacking on right elytron of holotype). Umbilicate setiferous series with 16 or 17 punctures of each elytron.
<emphasis id="1465489AE9958CF6B62E1449AD5406D0" bold="true" pageId="0" pageNumber="71">
<emphasis id="8D4D34967AF1D07713BEC418EE6BEF0E" italics="true" pageId="0" pageNumber="71">Legs</emphasis>
.
</emphasis>
Posterior side of profemur with one seta in basal third and one in medial third. Left mesofemur with 5, right one with 4 setiferous punctures on anterior side ventrally. Anterior side of metafemur ventrally with two long setae, one in basal third and one in medial third.
<emphasis id="C4109A88AB27201C37A3C1A7849B37EB" bold="true" pageId="0" pageNumber="71">
<emphasis id="2DB31572FEFF39B2CFAB65B62CF00EF3" italics="true" pageId="0" pageNumber="71">Male genitalia</emphasis>
.
</emphasis>
Unknown.
<emphasis id="79EF9DA28FACA28BB1187B4F2EBC2759" bold="true" italics="true" pageId="0" pageNumber="71">Female genitalia</emphasis>
(Fig.
<figureCitation id="8B5635DC3BC24F9F83AE40ACD7941576" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Spermathecal complex and gonocoxites, ventral view. A. Platyderus (Eremoderus) lassallei, sp. nov., female paratype, Iran, Mazandaran Province, E Qolqol; B. P. (E.) ledouxi Morvan, 1974, female specimen, 10 km SW Rudbarak, Mazandaran Province, Iran; C. P. (E.) taghizadehi Morvan, 1974, female specimen, Alburz Dizin, Tehran Province, Iran; D. P. (Platyderus) umbratus (Menetries), female specimen, Iran, Kohgiluyeh va Boyer-Ahmad Province, Sisakht; E. P. (E.) arabicus, sp. nov., holotype; F. P. (E.) brunki, sp. nov., holotype. Scale bars: 0.5 mm. For abbreviations see captions on Fig. 13." figureDoi="10.3897/dez.69.83840.figure14" httpUri="https://binary.pensoft.net/fig/695651" pageId="0" pageNumber="71">14F</figureCitation>
). Apical gonocoxite with pointed apex and two dorsolateral ensiform setae. Spermathecal canal connected in medial third of receptaculum.
</paragraph>
</subSubSection>
<subSubSection id="B15DDE8E00D8A72C276542EED704BDD8" pageId="0" pageNumber="71" type="habitat">
<paragraph id="50E3CE1A68BED9B65B5A3238EC7FA67D" pageId="0" pageNumber="71">Habitat.</paragraph>
<paragraph id="EA44654BF4ABCECDB91217643E930616" pageId="0" pageNumber="71">
The holotype was collected by sifting the waste-water vegetation and soil at a humid place where waste water falls down (Ingo Brunk, pers. comm.). The correct altitude of the type locality is ca. 2700 m a. s. l. The locality is situated approximately half-way on a very steep walking path starting from Kawkaban (town on the top of the Mount Kawkaban) downwards to Schibam Kawkaban (town on the bottom of the mountain). Together with the holotype, numerous specimens of
<taxonomicName id="C6EBA98C25791DE1D29B78A3B9C2DFB4" authorityName="Belousov" authorityYear="2017" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" lsidName="Trechus (Arabotrechus) rougemontiellus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="rougemontiellus" subGenus="Arabotrechus">Trechus (Arabotrechus) rougemontiellus</taxonomicName>
Belousov, 2017 and few
<taxonomicName id="8BF19F0A1BAAE3AFFFCFCC3A0608205D" class="Insecta" family="Carabidae" genus="Bembidion" kingdom="Animalia" lsidName="Bembidion" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis id="989544D5A05B78AB436CD4D41A541BA5" italics="true" pageId="0" pageNumber="71">Bembidion</emphasis>
</taxonomicName>
spp. were collected.
</paragraph>
<caption id="50408317C1DC681E2043924A449599DA" doi="10.3897/dez.69.83840.figure13" httpUri="https://binary.pensoft.net/fig/695650" pageId="0" pageNumber="71" start="Figure 13" startId="F13">
<paragraph id="186159E279C956FA2199A62EDCEC0693" pageId="0" pageNumber="71">
<emphasis id="FD006047F6F587C19F3286417AC8DC2A" bold="true" pageId="0" pageNumber="71">Figure 13.</emphasis>
Spermathecal complex and gonocoxites, ventral view.
<emphasis id="90E9A2B1F2F3A80079F1C0C8451235BC" bold="true" pageId="0" pageNumber="71">A.</emphasis>
<taxonomicName id="319AA356741370D0F2232F3F53C9CDD1" authorityName="Maran" authorityYear="1940" class="Insecta" family="Carabidae" genus="Platyderus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platyderus (Eremoderus) weiratheri" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="weiratheri" subGenus="Eremoderus">Platyderus (Eremoderus) weiratheri</taxonomicName>
<normalizedToken id="666869EF0EF24B55279A8DD919AA1FBF" originalValue="Mařan">Maran</normalizedToken>
, 1940, topotype female specimen;
<emphasis id="A698588B6022E4EDF6D8180138B67C76" bold="true" pageId="0" pageNumber="71">B.</emphasis>
<taxonomicName id="E6D0EE7A91B1DAD4FEBA088227E7578F" class="Insecta" kingdom="Animalia" lsidName="P. (E.) felixi" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="felixi" subGenus="E.">P. (E.) felixi</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis id="D40D3C2153B59BEF669D2F611DA0E709" bold="true" pageId="0" pageNumber="71">C.</emphasis>
<taxonomicName id="46B61937022E53A98B5C50D1548D112A" class="Insecta" kingdom="Animalia" lsidName="P. (E.) iranicus" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="iranicus" subGenus="E.">P. (E.) iranicus</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis id="2FB1B840AB56345FA8470F019295AC6E" bold="true" pageId="0" pageNumber="71">D.</emphasis>
<taxonomicName id="6B6977BAAE76F811C8A93DDC22B0BB02" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
, sp. nov., topotype female paratype;
<emphasis id="3F6E029497FC236124CE9737A641CA4F" bold="true" pageId="0" pageNumber="71">E.</emphasis>
<taxonomicName id="C11859DFDE13A0BF2CEFCB96F1B94EC4" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vanensis" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vanensis" subGenus="E.">P. (E.) vanensis</taxonomicName>
, sp. nov., female paratype, col de Buglan 1500 m, border area of
<normalizedToken id="8C3297C53F2DA6541D1D45863D4A0D90" originalValue="Bingöl">Bingoel</normalizedToken>
and
<normalizedToken id="862C4AD0C536DF16918DDF4C5829AD3E" originalValue="Muş">Mus</normalizedToken>
provinces, Turkey;
<emphasis id="52F7182B51DAEA63C094B0B6B7145F8E" bold="true" pageId="0" pageNumber="71">F.</emphasis>
<taxonomicName id="CE21516C59BE6FC4104C1473F4FE1253" class="Insecta" kingdom="Animalia" lsidName="P. (E.) vrabeci" order="Coleoptera" pageId="0" pageNumber="71" phylum="Arthropoda" rank="species" species="vrabeci" subGenus="E.">P. (E.) vrabeci</taxonomicName>
, sp. nov., topotype female paratype. Scale bars: 0.5 mm. Abbreviations:
<emphasis id="7844FC20C2ABF780549D2B1093B28612" bold="true" pageId="0" pageNumber="71">ag</emphasis>
- apical gonocoxite;
<emphasis id="AB357D7458224FB93904092B295424EE" bold="true" pageId="0" pageNumber="71">at</emphasis>
- atrium;
<emphasis id="FF076DE33BA125144CD405945D76AEE9" bold="true" pageId="0" pageNumber="71">bc</emphasis>
- bursa copulatrix;
<emphasis id="49646EF3B6D836F904FAFC7EDCCAC980" bold="true" pageId="0" pageNumber="71">bg</emphasis>
- basal gonocoxite;
<emphasis id="D53ECA72289A33EEC5ABCD61FB2115F2" bold="true" pageId="0" pageNumber="71">co</emphasis>
- common oviduct;
<emphasis id="DFD485FC261A99652A1DA5CCD5E31D19" bold="true" pageId="0" pageNumber="71">rc</emphasis>
- receptaculum;
<emphasis id="7D3A8FC1AADE6F7FD2D17F7616D124F9" bold="true" pageId="0" pageNumber="71">sg</emphasis>
- spermathecal gland;
<emphasis id="DEC34678D75960E5A275008C8DB0F27A" bold="true" pageId="0" pageNumber="71">smc</emphasis>
- seminal canal;
<emphasis id="0B212AC1F581B5617AD9FD9142CBA3DD" bold="true" pageId="0" pageNumber="71">spc</emphasis>
- spermathecal canal.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="95225C4D59ED43D19C69B46D378002D7" pageId="0" pageNumber="71" type="distribution">
<paragraph id="C351A19D3F428694C42B786FC52DE77C" pageId="0" pageNumber="71">Distribution.</paragraph>
<paragraph id="E7E91D35E8F29BA3053EEF5B6E38874B" pageId="0" pageNumber="71">
Yemen (
<normalizedToken id="480DCD26935B7025D09C49BB924E98F5" originalValue="Sana’a">Sana'a</normalizedToken>
Governorate), see Fig.
<figureCitation id="5DDF7F7631A71FDC24492FD7001DD606" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Distribution of species from subgenus Platyderus Eremoderus in the Levant and the Arabian Peninsula: Platyderus arabicus sp. nov. (yellow question marks), P. brunki sp. nov. (blue circle), P. irakensis sp. nov. (green circles), P. jordanensis sp. nov. (brown circles), and P. languidus (red circles)." figureDoi="10.3897/dez.69.83840.figure19" httpUri="https://binary.pensoft.net/fig/695656" pageId="0" pageNumber="71">19</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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