663 lines
117 KiB
XML
663 lines
117 KiB
XML
<document id="8088808F27A886816A62AF34CE0E0E62" ID-CLB-Dataset="9408" ID-DOI="10.11646/zootaxa.4890.4.2" ID-GBIF-Dataset="c6cc29e2-db8c-4f62-bf0d-a52db92197a6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4306459" ID-ZooBank="CAA4F713-3765-4619-A1DD-D6E00BDECA45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1607102189326" checkinUser="plazi" docAuthor="Dow, Rory A." docDate="2020" docId="AE5F560FFFA0A14976BBFC82FB94FE6A" docLanguage="en" docName="zootaxa.4890.4.2.pdf" docOrigin="Zootaxa 4890 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeliccia junis Dow 2020, sp. nov." docType="treatment" docVersion="9" lastPageNumber="478" masterDocId="52662E77FFA1A14C762CFF9CFFE6FFE3" masterDocTitle="Revision of the genus Coeliccia Kirby in Borneo part III: Two new species from the C. membranipes-group from Sarawak and Brunei (Odonata: Zygoptera: Platycnemididae)" masterLastPageNumber="492" masterPageNumber="473" pageNumber="474" updateTime="1698847158219" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="7C9174FA4F368E905E9084B602ADE349" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8D7C2A6A443EBA5A60F5F31D7B5DE04B">
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<mods:title id="0B403A57D12C7165E9F094F4553C2AE1">Revision of the genus Coeliccia Kirby in Borneo part III: Two new species from the C. membranipes-group from Sarawak and Brunei (Odonata: Zygoptera: Platycnemididae)</mods:title>
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</mods:titleInfo>
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<mods:name id="88C483541690B5DBCD217D3CE122FFC4" type="personal">
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<mods:role id="AC8E06BB617B132017A07098BD1F33BB">
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<mods:roleTerm id="DB527767A0B59359D0E98BA51974DB9E">Author</mods:roleTerm>
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<mods:namePart id="E6E8FA3C976F1F752FA8E14E5754E643">Dow, Rory A.</mods:namePart>
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<mods:typeOfResource id="FB6A2F76654BCD990ED05653FCD95745">text</mods:typeOfResource>
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<mods:titleInfo id="949D20C96A9F363878DD20DD7F0AF704">
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<mods:title id="3124124E8119D4A1412B8C07E0630752">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D9F1CD01952992CF391D5A89044DDB41">
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<mods:date id="1296A821BA7D9206F66BE5EE344D8B55">2020</mods:date>
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<mods:number id="60302DEA31BE7B3D7CB39BE59DC54CD2">2020-12-04</mods:number>
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<mods:detail id="D6C8B2C8ECB748AE09B720024FF21424" type="volume">
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<mods:number id="5F09C9C4F5D9B3779F32DBBBCA25C547">4890</mods:number>
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<mods:detail id="AFF7A25FB263CA643B1CA2FD6DC851CF" type="issue">
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<mods:number id="B44D9FD3F48A11B15F02A67AAEC6476C">4</mods:number>
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<mods:start id="192EB122549D89B35006BDA0C1CD716B">473</mods:start>
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<mods:classification id="348E80324F190D4D42D18C3B4B14D79A">journal article</mods:classification>
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<mods:identifier id="6F5C2C27E43D7EDC6071CCE6FA2510B2" type="DOI">10.11646/zootaxa.4890.4.2</mods:identifier>
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<mods:identifier id="D8A2F39909518E171D37EDAD2E418C9B" type="ISSN">1175-5326</mods:identifier>
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<treatment id="AE5F560FFFA0A14976BBFC82FB94FE6A" ID-DOI="http://doi.org/10.5281/zenodo.4323680" ID-GBIF-Taxon="170712176" ID-Zenodo-Dep="4323680" LSID="urn:lsid:plazi:treatment:AE5F560FFFA0A14976BBFC82FB94FE6A" httpUri="http://treatment.plazi.org/id/AE5F560FFFA0A14976BBFC82FB94FE6A" lastPageId="5" lastPageNumber="478" pageId="1" pageNumber="474">
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<subSubSection id="6EECB492FFA0A14D76BBFC82FE56FCDA" box="[151,432,798,825]" pageId="1" pageNumber="474" type="nomenclature">
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<paragraph id="2649E719FFA0A14D76BBFC82FE56FCDA" blockId="1.[151,938,798,861]" box="[151,432,798,825]" pageId="1" pageNumber="474">
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<heading id="7D015075FFA0A14D76BBFC82FE56FCDA" bold="true" box="[151,432,798,825]" fontSize="11" level="1" pageId="1" pageNumber="474" reason="1">
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<emphasis id="14823B0BFFA0A14D76BBFC82FE56FCDA" bold="true" box="[151,432,798,825]" pageId="1" pageNumber="474">
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<taxonomicName id="E1F69C9AFFA0A14D76BBFC82FEADFCDA" ID-CoL="9X2BN" authority="Dow, 2020" authorityName="Dow" authorityYear="2020" box="[151,331,798,825]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="474" phylum="Arthropoda" rank="species" species="junis" status="sp. nov.">
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<emphasis id="14823B0BFFA0A14D76BBFC82FEADFCDA" bold="true" box="[151,331,798,825]" italics="true" pageId="1" pageNumber="474">Coeliccia junis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="0FB18670FFA0A14D777FFC83FE56FCDA" box="[339,432,799,825]" pageId="1" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="6EECB492FFA0A14D76BBFCDFFC4CFCBE" box="[151,938,834,861]" pageId="1" pageNumber="474" type="description">
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<paragraph id="2649E719FFA0A14D76BBFCDFFC4CFCBE" blockId="1.[151,938,798,861]" box="[151,938,834,861]" pageId="1" pageNumber="474">
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(
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<figureCitation id="BECDFB9CFFA0A14D76B3FCDFFEE2FCBE" box="[159,260,835,861]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1903,1927]" captionTargetBox="[167,1421,414,1877]" captionTargetId="figure-153@2.[167,1421,414,1877]" captionTargetPageId="2" captionText="FIGURES 1–9. Heads of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view. Coeliccia roberti sp. nov. holotype male: (4) dorsal view; (5) frontal view; (6) ventral view. Coeliccia roberti sp. nov. female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="1" pageNumber="474">Figs 1–3</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D773CFCDFFEB1FCBE" box="[272,343,835,861]" captionStart="FIGURES 10–13" captionStartId="3.[151,264,1205,1229]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure-15@3.[151,1437,181,1181]" captionTargetPageId="3" captionText="FIGURES 10–13. Prothorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (10) dorsal view; (11) lateral view. Coeliccia roberti sp. nov. holotype male: (12) dorsal view; (13) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306463" httpUri="https://zenodo.org/record/4306463/files/figure.png" pageId="1" pageNumber="474">10–11</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D774FFCDFFE4DFCBE" box="[355,427,835,861]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-18@8.[151,1436,181,1680]" captionTargetPageId="8" captionText="FIGURES 16–21. Synthorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (16) dorsal view; (17) lateral view. Coeliccia junis sp. nov. paratype male (SAR18_PCD27): (18) dorsal view; (19) lateral view. Coeliccia roberti sp. nov. holotype male: (20) dorsal view; (21) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306469" httpUri="https://zenodo.org/record/4306469/files/figure.png" pageId="1" pageNumber="474">16–19</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D779BFCDFFE33FCBE" box="[439,469,835,861]" captionStart="FIGURES 24–25" captionStartId="9.[151,264,1486,1510]" captionTargetBox="[151,1436,881,1463]" captionTargetId="figure-96@9.[151,1436,881,1463]" captionTargetPageId="9" captionText="FIGURES 24–25. Genital ligula of Coeliccia species in ventral view: (24) Coeliccia junis paratype SAR18_PCD27. (25) Scan- ning Electron Microscope image of Coeliccia roberti SAR06-PCD77, not in type series, Lambir Hills National Park. All scale bars 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4306473" httpUri="https://zenodo.org/record/4306473/files/figure.png" pageId="1" pageNumber="474">24</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D77CDFCDFFDCFFCBE" box="[481,553,835,861]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1583,1607]" captionTargetBox="[151,1436,489,1560]" captionTargetId="figure-190@10.[151,1436,489,1560]" captionTargetPageId="10" captionText="FIGURES 26–31. Terminal abdominal segments of Coeliccia species. Coeliccia junis sp. nov. holotype male: (26) dorsal view; (27) lateral view. Coeliccia roberti sp. nov. holotype male: (28) dorsal view; (29) lateral view. Coeliccia roberti sp. nov., female described: (30) dorsal view; (31) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306475" httpUri="https://zenodo.org/record/4306475/files/figure.png" pageId="1" pageNumber="474">26–27</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D7419FCDFFD98FCBE" box="[565,638,835,861]" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1972,1996]" captionTargetBox="[415,1173,181,1948]" captionTargetId="figure-15@13.[415,1173,181,1948]" captionTargetPageId="13" captionText="FIGURES 32–34. Dorsal view of anal appendages of male Coeliccia species, all scale bars 0.5mm. (32) Coeliccia junis sp. nov. holotype. (33) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (34) Coeliccia roberti sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.4306477" httpUri="https://zenodo.org/record/4306477/files/figure.png" pageId="1" pageNumber="474">32–33</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D74A5FCDEFD41FCBE" box="[649,679,834,861]" captionStart="FIGURES 35–37" captionStartId="14.[151,264,791,815]" captionTargetBox="[157,1429,181,768]" captionTargetId="figure-18@14.[151,1436,181,768]" captionTargetPageId="14" captionText="FIGURES 35–37. Sketches of the epiproct of Coeliccia species in dorsal view, also showing inner basal parts of the cerci: (35) Coeliccia junis paratype SAR18_PCD27; (36) Coeliccia nigrohamata SAR06-PPCD83, Sabaat River, foot of Gunung Pueh, Kuching Division, Sarawak, leg. R.A. Dow, included for comparison; (37) Coeliccia roberti paratype BRU14_PCD17." figureDoi="http://doi.org/10.5281/zenodo.4306479" httpUri="https://zenodo.org/record/4306479/files/figure.png" pageId="1" pageNumber="474">35</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D749FFCDFFD1AFCBE" box="[691,764,835,861]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="1" pageNumber="474">38–39</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D752BFCDFFCA9FCBE" box="[775,847,835,861]" captionStart="FIGURES 42–45" captionStartId="15.[151,264,1132,1156]" captionTargetBox="[151,1435,181,1108]" captionTargetId="figure-15@15.[151,1435,181,1108]" captionTargetPageId="15" captionText="FIGURES 42–45. Ventral view of anal appendages of male Coeliccia species. Scale bars (where present) 0.2mm. (42) Coeliccia junis sp. nov. holotype. (43) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (44) Coeliccia roberti sp. nov. holotype. (45) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306483" httpUri="https://zenodo.org/record/4306483/files/figure.png" pageId="1" pageNumber="474">42–43</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D7577FCDFFC9CFCBE" box="[859,890,835,861]" captionStart="FIGURES 46–47" captionStartId="15.[151,264,1790,1814]" captionTargetBox="[151,1437,1252,1766]" captionTargetId="figure-60@15.[151,1437,1252,1766]" captionTargetPageId="15" captionText="FIGURES 46–47. Ventral view of cerci of male Coeliccia species. (46) Coeliccia junis sp. nov. paratype (SAR18_PCD28) with right paraproct removed to allow ventral view of the right cercus. (47) Detail of Coeliccia roberti sp. nov. (SAR06-PCD151, not in type series) with paraprocts removed to allow ventral view of the cerci. Scale bars 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4306485" httpUri="https://zenodo.org/record/4306485/files/figure.png" pageId="1" pageNumber="474">46</figureCitation>
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,
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<figureCitation id="BECDFB9CFFA0A14D75A9FCDEFC44FCBE" box="[901,930,834,861]" captionStart="FIGURE 52" captionStartId="17.[151,250,1693,1717]" captionTargetBox="[151,1436,273,1670]" captionTargetId="figure-72@17.[151,1436,273,1670]" captionTargetPageId="17" captionText="FIGURE 52. Map of Borneo showing the distributions of two Coeliccia species. Red triangle—Coeliccia junis sp. nov; white circle—Coeliccia roberti sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4306489" httpUri="https://zenodo.org/record/4306489/files/figure.png" pageId="1" pageNumber="474">52</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="6EECB492FFA0A14D76BBFC17FDFFFBBA" pageId="1" pageNumber="474" type="materials_examined">
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<paragraph id="2649E719FFA0A14D76BBFC17FA9EFC0E" blockId="1.[151,1437,906,2013]" pageId="1" pageNumber="474">
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<emphasis id="14823B0BFFA0A14D76BBFC17FEAFFC47" bold="true" box="[151,329,906,933]" pageId="1" pageNumber="474">Type material:</emphasis>
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<materialsCitation id="969EED44FFA0A14D777FFC17FA9EFC0E" ID-GBIF-Occurrence="2989990303" collectingDate="2018-08-10" collectionCode="RMNH" collectorName="Cheong Yi Wei" country="Malaysia" county="Division" elevation="100" latitude="2.7884" location="Bukit Nyegoh" longLatPrecision="7" longitude="113.3151" municipality="Bukit" pageId="1" pageNumber="474" specimenCount="1" typeStatus="holotype">
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<emphasis id="14823B0BFFA0A14D777FFC17FE2FFC46" bold="true" box="[339,457,907,933]" pageId="1" pageNumber="474">
|
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<typeStatus id="F94D59BBFFA0A14D777FFC17FE22FC46" box="[339,452,907,933]" pageId="1" pageNumber="474" type="holotype">Holotype</typeStatus>
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||
:
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</emphasis>
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Ƌ (SAR018_PCD101), near small brown water pool near main “Day 4” stream near
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||
<location id="2329B1C2FFA0A14D76BBFC33FED3FC2A" LSID="urn:lsid:plazi:treatment:AE5F560FFFA0A14976BBFC82FB94FE6A:2329B1C2FFA0A14D76BBFC33FED3FC2A" box="[151,309,943,969]" country="Malaysia" county="Division" latitude="2.7884" longLatPrecision="7" longitude="113.3151" municipality="Bukit" name="Bukit Nyegoh" pageId="1" pageNumber="474">Bukit Nyegoh</location>
|
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and
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<location id="2329B1C2FFA0A14D7746FC33FE1CFC2A" LSID="urn:lsid:plazi:treatment:AE5F560FFFA0A14976BBFC82FB94FE6A:2329B1C2FFA0A14D7746FC33FE1CFC2A" box="[362,506,943,969]" country="Malaysia" county="Division" latitude="2.7884" longLatPrecision="7" longitude="113.3151" municipality="Bukit" name="Bukit Jugam" pageId="1" pageNumber="474">Bukit Jugam</location>
|
||
,
|
||
<collectingMunicipality id="C62D7D63FFA0A14D7429FC33FDA5FC2A" box="[517,579,943,969]" pageId="1" pageNumber="474">Bukit</collectingMunicipality>
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<taxonomicName id="E1F69C9AFFA0A14D7465FC33FB27FC2A" authority="Wild Life Corridor, Planted Forest Project, Bintulu" authorityName="Wild Life Corridor, Planted Forest Project, Bintulu" box="[585,1217,943,969]" class="Magnoliopsida" family="Convolvulaceae" genus="Mina" higherTaxonomySource="CoL" kingdom="Plantae" order="Solanales" pageId="1" pageNumber="474" phylum="Tracheophyta" rank="genus">Mina Wild Life Corridor, Planted Forest Project, Bintulu</taxonomicName>
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<collectingCounty id="CF289F95FFA0A14D72EBFC33FACCFC2A" box="[1223,1322,943,969]" pageId="1" pageNumber="474">Division</collectingCounty>
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,
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<collectingCountry id="5EE1A789FFA0A14D7319FC33FA7EFC2A" box="[1333,1432,943,969]" name="Malaysia" pageId="1" pageNumber="474">Sarawak</collectingCountry>
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, approximate coordinates
|
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<geoCoordinate id="43C281DEFFA0A14D779EFC4FFDF1FC0E" box="[434,535,979,1005]" degrees="2.7884" direction="north" orientation="latitude" pageId="1" pageNumber="474" precision="5" value="2.7884">2.7884N</geoCoordinate>
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,
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<geoCoordinate id="43C281DEFFA0A14D740DFC4EFD78FC0E" box="[545,670,978,1005]" degrees="113.3151" direction="east" orientation="longitude" pageId="1" pageNumber="474" precision="5" value="113.3151">113.3151E</geoCoordinate>
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, ca.
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<elevation id="ADDB002AFFA0A14D74E3FC4FFC93FC0E" box="[719,885,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="1" pageNumber="474" unit="m" value="100.0" valueMax="110.0" valueMin="90.0">
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<quantity id="E10E4AFCFFA0A14D74E3FC4FFCDEFC0E" box="[719,824,979,1005]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="1" pageNumber="474" unit="m" value="100.0" valueMax="110.0" valueMin="90.0">90–110m</quantity>
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a.s.l.
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</elevation>
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,
|
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<date id="5248C1D9FFA0A14D75ADFC4FFBEAFC0E" box="[897,1036,979,1005]" pageId="1" pageNumber="474" value="2018-08-10">
|
||
<collectingDate id="420C3831FFA0A14D75ADFC4FFBEAFC0E" box="[897,1036,979,1005]" pageId="1" pageNumber="474" value="2018-08-10">10 viii 2018</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="8B0382CFFFA0A14D726BFC4FFB12FC0E" box="[1095,1268,979,1005]" pageId="1" pageNumber="474">Cheong Yi Wei</collectorName>
|
||
, in
|
||
<collectionCode id="40E77FDCFFA0A14D7330FC4FFA92FC0E" box="[1308,1396,979,1005]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="474" type="Museum">RMNH</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA0A14D76EBFC6BFDFFFBBA" blockId="1.[151,1437,906,2013]" pageId="1" pageNumber="474">
|
||
<materialsCitation id="969EED44FFA0A14D76EBFC6BFDFFFBBA" ID-GBIF-Occurrence="2989990301" collectingDate="2018-01-20" collectionCode="DD" collectorName="Dow." country="Malaysia" county="Bintulu Division" elevation="30" latitude="2.931" location="Sungai Menaung" longLatPrecision="55" longitude="113.3903" municipality="Binyo Penyilam" pageId="1" pageNumber="474" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="14823B0BFFA0A14D76EBFC6BFEA0FBF2" bold="true" box="[199,326,1015,1041]" pageId="1" pageNumber="474">
|
||
<typeStatus id="F94D59BBFFA0A14D76EBFC6BFEA7FBF2" box="[199,321,1015,1041]" pageId="1" pageNumber="474" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
(2
|
||
<collectionCode id="40E77FDCFFA0A14D7747FC6BFE71FBF2" box="[363,407,1015,1041]" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="1" pageNumber="474" type="Herbarium">ƋƋ</collectionCode>
|
||
): 2 ƋƋ (SAR18_PCD27–28), in low pH swamp forest beside tributary to
|
||
<location id="2329B1C2FFA0A14D72FBFC6BFA7EFBF2" LSID="urn:lsid:plazi:treatment:AE5F560FFFA0A14976BBFC82FB94FE6A:2329B1C2FFA0A14D72FBFC6BFA7EFBF2" box="[1239,1432,1015,1041]" country="Malaysia" county="Bintulu Division" latitude="2.931" longLatPrecision="55" longitude="113.3903" municipality="Binyo Penyilam" name="Sungai Menaung" pageId="1" pageNumber="474">Sungai Menaung</location>
|
||
,
|
||
<collectingMunicipality id="C62D7D63FFA0A14D76BBFB87FEA9FBD6" box="[151,335,1051,1077]" pageId="1" pageNumber="474">Binyo Penyilam</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="CF289F95FFA0A14D7775FB87FDF0FBD6" box="[345,534,1051,1077]" pageId="1" pageNumber="474">Bintulu Division</collectingCounty>
|
||
,
|
||
<collectingCountry id="5EE1A789FFA0A14D740DFB87FD62FBD6" box="[545,644,1051,1077]" name="Malaysia" pageId="1" pageNumber="474">Sarawak</collectingCountry>
|
||
,
|
||
<collectingCountry id="5EE1A789FFA0A14D74A3FB87FD1EFBD6" box="[655,760,1051,1077]" name="Malaysia" pageId="1" pageNumber="474">Malaysia</collectingCountry>
|
||
, approximate coordinates
|
||
<geoCoordinate id="43C281DEFFA0A14D7230FB87FB94FBD6" box="[1052,1138,1051,1077]" degrees="2.931" direction="north" orientation="latitude" pageId="1" pageNumber="474" precision="55" value="2.931">2.931N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="43C281DEFFA0A14D7251FB87FB1EFBD6" box="[1149,1272,1051,1077]" degrees="113.3903" direction="east" orientation="longitude" pageId="1" pageNumber="474" precision="5" value="113.3903">113.3903E</geoCoordinate>
|
||
, ca.
|
||
<elevation id="ADDB002AFFA0A14D7305FB87FA71FBD6" box="[1321,1431,1051,1077]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="474" unit="m" value="30.0">
|
||
<quantity id="E10E4AFCFFA0A14D7305FB87FABDFBD6" box="[1321,1371,1051,1077]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="474" unit="m" value="30.0">30m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
,
|
||
<date id="5248C1D9FFA0A14D76BBFBA3FEE2FBBA" box="[151,260,1087,1113]" pageId="1" pageNumber="474" value="2018-01-20">
|
||
<collectingDate id="420C3831FFA0A14D76BBFBA3FEE2FBBA" box="[151,260,1087,1113]" pageId="1" pageNumber="474" value="2018-01-20">20 i 2018</collectingDate>
|
||
</date>
|
||
, leg. RAD, in coll.
|
||
<collectorName id="8B0382CFFFA0A14D77F1FBA3FDFFFBBA" box="[477,537,1087,1113]" pageId="1" pageNumber="474">Dow.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6EECB492FFA0A14D76EBFBFFFB03FB26" pageId="1" pageNumber="474" type="etymology">
|
||
<paragraph id="2649E719FFA0A14D76EBFBFFFB03FB26" blockId="1.[151,1437,906,2013]" pageId="1" pageNumber="474">
|
||
<emphasis id="14823B0BFFA0A14D76EBFBFFFEB4FB9E" bold="true" box="[199,338,1123,1149]" pageId="1" pageNumber="474">Etymology:</emphasis>
|
||
Junis, a noun in apposition, named for Joanes Unggang (date of birth
|
||
<date id="5248C1D9FFA0A14D7256FBFFFB1EFB9E" box="[1146,1272,1123,1149]" pageId="1" pageNumber="474" value="1978-06-16">16 vi 1978</date>
|
||
), friend of the author and facilitator of both the trips on which material of this new species was collected. Junis (“Joo-nis”) is the local pronunciation of Joanes’ name in the Sungai Katibas area of
|
||
<collectingCountry id="5EE1A789FFA0A14D7551FB37FC06FB26" box="[893,992,1195,1221]" name="Malaysia" pageId="1" pageNumber="474">Sarawak</collectingCountry>
|
||
, where he comes from.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6EECB492FFA0A14876EBFB52FA96FCDA" lastPageId="4" lastPageNumber="477" pageId="1" pageNumber="474" type="description">
|
||
<paragraph id="2649E719FFA0A14D76EBFB52FBC6F9EA" blockId="1.[151,1437,906,2013]" pageId="1" pageNumber="474">
|
||
<emphasis id="14823B0BFFA0A14D76EBFB52FDC3FB0A" bold="true" box="[199,549,1230,1257]" pageId="1" pageNumber="474">
|
||
Description of
|
||
<typeStatus id="F94D59BBFFA0A14D775BFB53FE38FB0A" box="[375,478,1231,1257]" pageId="1" pageNumber="474" type="holotype">holotype</typeStatus>
|
||
male.
|
||
</emphasis>
|
||
<emphasis id="14823B0BFFA0A14D7401FB53FD8CFB0A" box="[557,618,1231,1257]" italics="true" pageId="1" pageNumber="474">Head</emphasis>
|
||
(
|
||
<figureCitation id="BECDFB9CFFA0A14D7454FB53FD38FB0A" box="[632,734,1231,1257]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1903,1927]" captionTargetBox="[167,1421,414,1877]" captionTargetId="figure-153@2.[167,1421,414,1877]" captionTargetPageId="2" captionText="FIGURES 1–9. Heads of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view. Coeliccia roberti sp. nov. holotype male: (4) dorsal view; (5) frontal view; (6) ventral view. Coeliccia roberti sp. nov. female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="1" pageNumber="474">Figs 1–3</figureCitation>
|
||
): Labium cream except hooks of labial palps, which are black. Labrum shining black. Mandible bases shining black with grey area lower anterior corner. Clypeus black except faint grey central mark on anteclypeus. Genae shining black except immediately adjacent to mandible bases where very dark brown. Pale blue band running from eye margin to clypeus and narrowly above lateral ca. one quarter of clypeus on either side (
|
||
<figureCitation id="BECDFB9CFFA0A14D7790FAC3FDE4FA9A" box="[444,514,1375,1401]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1903,1927]" captionTargetBox="[167,1421,414,1877]" captionTargetId="figure-153@2.[167,1421,414,1877]" captionTargetPageId="2" captionText="FIGURES 1–9. Heads of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view. Coeliccia roberti sp. nov. holotype male: (4) dorsal view; (5) frontal view; (6) ventral view. Coeliccia roberti sp. nov. female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="1" pageNumber="474">Fig. 2</figureCitation>
|
||
). Antenna black with top part of scape and base of pedicel whitish, top of pedicel brown. Frons and vertex mostly black, with irregular, incomplete pale marking from lateral margin lateral ocellus running behind antennae to eye margin as shown in
|
||
<figureCitation id="BECDFB9CFFA0A14D74C2FA3BFCD4FA22" box="[750,818,1447,1473]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1903,1927]" captionTargetBox="[167,1421,414,1877]" captionTargetId="figure-153@2.[167,1421,414,1877]" captionTargetPageId="2" captionText="FIGURES 1–9. Heads of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view. Coeliccia roberti sp. nov. holotype male: (4) dorsal view; (5) frontal view; (6) ventral view. Coeliccia roberti sp. nov. female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="1" pageNumber="474">Fig. 1</figureCitation>
|
||
. Ocelli whitish. Pale yellow, elongate oval, transverse postocular spots. Underside of head (
|
||
<figureCitation id="BECDFB9CFFA0A14D741BFA57FD9FFA06" box="[567,633,1483,1509]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1903,1927]" captionTargetBox="[167,1421,414,1877]" captionTargetId="figure-153@2.[167,1421,414,1877]" captionTargetPageId="2" captionText="FIGURES 1–9. Heads of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view. Coeliccia roberti sp. nov. holotype male: (4) dorsal view; (5) frontal view; (6) ventral view. Coeliccia roberti sp. nov. female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="1" pageNumber="474">Fig. 3</figureCitation>
|
||
) black with pale marks at eye margin and behind mandible bases, these narrowly connected to small pale marks beside point of attachment of prothorax.
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA0A14E76EBF98FFCF6FE6A" blockId="1.[151,1437,906,2013]" lastBlockId="2.[151,1436,151,393]" lastPageId="2" lastPageNumber="475" pageId="1" pageNumber="474">
|
||
<emphasis id="14823B0BFFA0A14D76EBF98FFEFEF9CE" box="[199,280,1555,1581]" italics="true" pageId="1" pageNumber="474">Thorax</emphasis>
|
||
: Prothorax (
|
||
<figureCitation id="BECDFB9CFFA0A14D7788F98FFDC4F9CE" box="[420,546,1555,1581]" captionStart="FIGURES 10–13" captionStartId="3.[151,264,1205,1229]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure-15@3.[151,1437,181,1181]" captionTargetPageId="3" captionText="FIGURES 10–13. Prothorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (10) dorsal view; (11) lateral view. Coeliccia roberti sp. nov. holotype male: (12) dorsal view; (13) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306463" httpUri="https://zenodo.org/record/4306463/files/figure.png" pageId="1" pageNumber="474">Figs 10–11</figureCitation>
|
||
) with propleuron yellow except small dark mark in lower rear corner and black adjacent to posterior pronotal lobe. Posterior and anterior pronotal lobes black, except narrowly laterally on anterior lobe, where yellow. Anterior carina of anterior pronotal lobe almost reaching same level as main part (
|
||
<figureCitation id="BECDFB9CFFA0A14D7327F9C7FABAF996" box="[1291,1372,1627,1653]" captionStart="FIGURES 10–13" captionStartId="3.[151,264,1205,1229]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure-15@3.[151,1437,181,1181]" captionTargetPageId="3" captionText="FIGURES 10–13. Prothorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (10) dorsal view; (11) lateral view. Coeliccia roberti sp. nov. holotype male: (12) dorsal view; (13) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306463" httpUri="https://zenodo.org/record/4306463/files/figure.png" pageId="1" pageNumber="474">Fig. 11</figureCitation>
|
||
). Upper part of notopleural projection just present as small rounded lump. Middle lobe yellow laterally, black dorsally posteriorly with hemispherical, dome-like, bulge on either side bearing many short tubercles (
|
||
<figureCitation id="BECDFB9CFFA0A14D72E0F93FFAC6F95E" box="[1228,1312,1699,1725]" captionStart="FIGURES 10–13" captionStartId="3.[151,264,1205,1229]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure-15@3.[151,1437,181,1181]" captionTargetPageId="3" captionText="FIGURES 10–13. Prothorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (10) dorsal view; (11) lateral view. Coeliccia roberti sp. nov. holotype male: (12) dorsal view; (13) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306463" httpUri="https://zenodo.org/record/4306463/files/figure.png" pageId="1" pageNumber="474">Fig. 11</figureCitation>
|
||
). Posterior pronotal lobe simple, collar like, free margin slightly raised relative to the rest. Synthorax (
|
||
<figureCitation id="BECDFB9CFFA0A14D72BDF95BFAF7F901" box="[1169,1297,1735,1762]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-18@8.[151,1436,181,1680]" captionTargetPageId="8" captionText="FIGURES 16–21. Synthorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (16) dorsal view; (17) lateral view. Coeliccia junis sp. nov. paratype male (SAR18_PCD27): (18) dorsal view; (19) lateral view. Coeliccia roberti sp. nov. holotype male: (20) dorsal view; (21) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306469" httpUri="https://zenodo.org/record/4306469/files/figure.png" pageId="1" pageNumber="474">Figs 16–17</figureCitation>
|
||
) with mesepisternum black with large blue antehumeral markings (
|
||
<figureCitation id="BECDFB9CFFA0A14D752CF977FCB7F8E5" box="[768,849,1771,1798]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-18@8.[151,1436,181,1680]" captionTargetPageId="8" captionText="FIGURES 16–21. Synthorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (16) dorsal view; (17) lateral view. Coeliccia junis sp. nov. paratype male (SAR18_PCD27): (18) dorsal view; (19) lateral view. Coeliccia roberti sp. nov. holotype male: (20) dorsal view; (21) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306469" httpUri="https://zenodo.org/record/4306469/files/figure.png" pageId="1" pageNumber="474">Fig. 16</figureCitation>
|
||
), extending from near mesostigmal plates to ca. apex of antealar triangle, outer margin following mesepleural suture for most of length, inner margin irregularly excised centrally. Mesepimeron largely black except narrowly blue above the interpleural suture for some distance and tiny pale mark in upper corner adjacent to antealar carina (
|
||
<figureCitation id="BECDFB9CFFA0A14D74C5F8CBFCDDF892" box="[745,827,1879,1905]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-18@8.[151,1436,181,1680]" captionTargetPageId="8" captionText="FIGURES 16–21. Synthorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (16) dorsal view; (17) lateral view. Coeliccia junis sp. nov. paratype male (SAR18_PCD27): (18) dorsal view; (19) lateral view. Coeliccia roberti sp. nov. holotype male: (20) dorsal view; (21) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306469" httpUri="https://zenodo.org/record/4306469/files/figure.png" pageId="1" pageNumber="474">Fig. 17</figureCitation>
|
||
). Metepisternum largely blue with black stripe running from antealar carina most of way towards spiracle and narrowly black adjacent to antealar carina. Metepimeron and venter of synthorax pale yellow. Mesokatepisternum black except lower corner adjacent to metepisternum where pale yellowish, Metakatepisternum entirely pale. Legs with coxae and trochanters almost entirely pale, femur most-ly pale with black stripe along extensor surface, dark spines and black stripe along outer flexor surface in distal ca. 2/3 of anterior femur only, tibia mostly pale, dark spines and dark brown on most of flexor surface, tarsi pale with obscure darker areas, claws brown, spines dark. Wings with arc situated slightly distal to Ax 2 (only very slightly distal in right Hw). Fw with 16 Px, Hw with 14 (left) or 15 (right) Px. Two to three post quadrilateral cells in all wings. R
|
||
<subScript id="BA72E55CFFA3A14E76D6FEA5FEE4FEA4" attach="left" box="[250,258,313,327]" fontSize="6" pageId="2" pageNumber="475">4</subScript>
|
||
arising very slightly distal (Fw) or very slightly proximal (Hw) to Sn. Pt black with narrow irregular white margin, broad, almost a rhombus, covering one and a half to two underlying cells except in right Fw where it covers ca. two thirds of one exceptionally large underlying cell.
|
||
</paragraph>
|
||
<caption id="7289B791FFA3A14E76BBF8F3FBD3F82C" ID-DOI="http://doi.org/10.5281/zenodo.4306461" ID-Zenodo-Dep="4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="2" pageNumber="475" startId="2.[151,264,1903,1927]" targetBox="[167,1421,414,1877]" targetPageId="2">
|
||
<paragraph id="2649E719FFA3A14E76BBF8F3FBD3F82C" blockId="2.[151,1436,1903,1999]" pageId="2" pageNumber="475">
|
||
<emphasis id="14823B0BFFA3A14E76BBF8F3FEDCF864" bold="true" box="[151,314,1903,1927]" pageId="2" pageNumber="475">FIGURES 1–9.</emphasis>
|
||
Heads of males of
|
||
<taxonomicName id="E1F69C9AFFA3A14E77D0F8F3FDBBF864" authorityName="Kirby in Borneo" authorityYear="1890" box="[508,605,1903,1927]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="475" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="14823B0BFFA3A14E77D0F8F3FDBBF864" box="[508,605,1903,1927]" italics="true" pageId="2" pageNumber="475">Coeliccia</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="E1F69C9AFFA3A14E7494F8F3FCB6F864" authority="Dow, 2020" authorityName="Dow" authorityYear="2020" box="[696,848,1903,1927]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="junis" status="sp. nov.">
|
||
<emphasis id="14823B0BFFA3A14E7494F8F3FCB6F864" box="[696,848,1903,1927]" italics="true" pageId="2" pageNumber="475">Coeliccia junis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14823B0BFFA3A14E757AF8F3FC41F864" bold="true" box="[854,935,1903,1927]" pageId="2" pageNumber="475">
|
||
<taxonomicNameLabel id="0FB18670FFA3A14E757AF8F3FC41F864" box="[854,935,1903,1927]" pageId="2" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view.
|
||
<taxonomicName id="E1F69C9AFFA3A14E77AAF80FFDD5F848" authority="Dow, 2020" authorityName="Dow" authorityYear="2020" box="[390,563,1939,1963]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="roberti" status="sp. nov.">
|
||
<emphasis id="14823B0BFFA3A14E77AAF80FFDD5F848" box="[390,563,1939,1963]" italics="true" pageId="2" pageNumber="475">Coeliccia roberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14823B0BFFA3A14E7417F80FFD6AF848" bold="true" box="[571,652,1939,1963]" pageId="2" pageNumber="475">
|
||
<taxonomicNameLabel id="0FB18670FFA3A14E7417F80FFD6AF848" box="[571,652,1939,1963]" pageId="2" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype male: (4) dorsal view; (5) frontal view; (6) ventral view.
|
||
<taxonomicName id="E1F69C9AFFA3A14E7317F80FFF3AF82C" authority="Dow, 2020" authorityName="Dow" authorityYear="2020" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="475" phylum="Arthropoda" rank="species" species="roberti" status="sp. nov.">
|
||
<emphasis id="14823B0BFFA3A14E7317F80FFF3AF82C" italics="true" pageId="2" pageNumber="475">Coeliccia roberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14823B0BFFA3A14E76CFF82BFED5F82C" bold="true" box="[227,307,1975,1999]" pageId="2" pageNumber="475">
|
||
<taxonomicNameLabel id="0FB18670FFA3A14E76CFF82BFED5F82C" box="[227,307,1975,1999]" pageId="2" pageNumber="475" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="7289B791FFA2A14F76BBFB29FC3AFB12" ID-DOI="http://doi.org/10.5281/zenodo.4306463" ID-Zenodo-Dep="4306463" httpUri="https://zenodo.org/record/4306463/files/figure.png" pageId="3" pageNumber="476" startId="3.[151,264,1205,1229]" targetBox="[151,1437,181,1181]" targetPageId="3">
|
||
<paragraph id="2649E719FFA2A14F76BBFB29FC3AFB12" blockId="3.[151,1436,1205,1265]" pageId="3" pageNumber="476">
|
||
<emphasis id="14823B0BFFA2A14F76BBFB29FEB5FB2E" bold="true" box="[151,339,1205,1229]" pageId="3" pageNumber="476">FIGURES 10–13.</emphasis>
|
||
Prothorax of males of
|
||
<taxonomicName id="E1F69C9AFFA2A14F741AFB29FD71FB2E" authorityName="Kirby in Borneo" authorityYear="1890" box="[566,663,1205,1229]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="476" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="14823B0BFFA2A14F741AFB29FD71FB2E" box="[566,663,1205,1229]" italics="true" pageId="3" pageNumber="476">Coeliccia</emphasis>
|
||
</taxonomicName>
|
||
species.
|
||
<taxonomicName id="E1F69C9AFFA2A14F74DDFB29FC6EFB2E" authority="Dow, 2020" authorityName="Dow" authorityYear="2020" box="[753,904,1205,1229]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="junis" status="sp. nov.">
|
||
<emphasis id="14823B0BFFA2A14F74DDFB29FC6EFB2E" box="[753,904,1205,1229]" italics="true" pageId="3" pageNumber="476">Coeliccia junis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14823B0BFFA2A14F75A2FB29FC3BFB2E" bold="true" box="[910,989,1205,1229]" pageId="3" pageNumber="476">
|
||
<taxonomicNameLabel id="0FB18670FFA2A14F75A2FB29FC3BFB2E" box="[910,989,1205,1229]" pageId="3" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype male: (10) dorsal view; (11) lateral view.
|
||
<taxonomicName id="E1F69C9AFFA2A14F76FFFB45FE66FB12" authority="Dow, 2020" authorityName="Dow" authorityYear="2020" box="[211,384,1241,1265]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="476" phylum="Arthropoda" rank="species" species="roberti" status="sp. nov.">
|
||
<emphasis id="14823B0BFFA2A14F76FFFB45FE66FB12" box="[211,384,1241,1265]" italics="true" pageId="3" pageNumber="476">Coeliccia roberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14823B0BFFA2A14F77ABFB45FE31FB12" bold="true" box="[391,471,1241,1265]" pageId="3" pageNumber="476">
|
||
<taxonomicNameLabel id="0FB18670FFA2A14F77ABFB45FE31FB12" box="[391,471,1241,1265]" pageId="3" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
holotype male: (12) dorsal view; (13) lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2649E719FFA2A14876EBFA83FC31FF1A" blockId="3.[151,1437,1311,2022]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="477" pageId="3" pageNumber="476">
|
||
<emphasis id="14823B0BFFA2A14F76EBFA83FED7FADA" box="[199,305,1311,1337]" italics="true" pageId="3" pageNumber="476">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="BECDFB9CFFA2A14F776CFA83FE22FAD9" box="[320,452,1311,1338]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1583,1607]" captionTargetBox="[151,1436,489,1560]" captionTargetId="figure-190@10.[151,1436,489,1560]" captionTargetPageId="10" captionText="FIGURES 26–31. Terminal abdominal segments of Coeliccia species. Coeliccia junis sp. nov. holotype male: (26) dorsal view; (27) lateral view. Coeliccia roberti sp. nov. holotype male: (28) dorsal view; (29) lateral view. Coeliccia roberti sp. nov., female described: (30) dorsal view; (31) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306475" httpUri="https://zenodo.org/record/4306475/files/figure.png" pageId="3" pageNumber="476">Figs 26–27</figureCitation>
|
||
): S1 (visible in
|
||
<figureCitation id="BECDFB9CFFA2A14F745AFA83FD21FAD9" box="[630,711,1311,1338]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-18@8.[151,1436,181,1680]" captionTargetPageId="8" captionText="FIGURES 16–21. Synthorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (16) dorsal view; (17) lateral view. Coeliccia junis sp. nov. paratype male (SAR18_PCD27): (18) dorsal view; (19) lateral view. Coeliccia roberti sp. nov. holotype male: (20) dorsal view; (21) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306469" httpUri="https://zenodo.org/record/4306469/files/figure.png" pageId="3" pageNumber="476">Fig. 17</figureCitation>
|
||
) pale yellowish except black in narrow apical annulus including posterior carina and area behind, narrowly on dorsum. S2 pale yellowish laterally except small dark brown subtriangular area basally immediately above margin of tergite, becoming brown dorsally, this widest apically; narrow pale stripe centrally on dorsum subbasally. S3–7 pale lower laterally (e.g. immediately above lower margin of tergite), becoming brown dorsally, pale basal annulus, interrupted dorsally, and pair very faint pale subapical dorsal markings, on each segment. S8 pale lower laterally, this becoming blue apically, otherwise black (
|
||
<figureCitation id="BECDFB9CFFA2A14F72F9FA4FFACFFA0D" box="[1237,1321,1491,1518]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1583,1607]" captionTargetBox="[151,1436,489,1560]" captionTargetId="figure-190@10.[151,1436,489,1560]" captionTargetPageId="10" captionText="FIGURES 26–31. Terminal abdominal segments of Coeliccia species. Coeliccia junis sp. nov. holotype male: (26) dorsal view; (27) lateral view. Coeliccia roberti sp. nov. holotype male: (28) dorsal view; (29) lateral view. Coeliccia roberti sp. nov., female described: (30) dorsal view; (31) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306475" httpUri="https://zenodo.org/record/4306475/files/figure.png" pageId="3" pageNumber="476">Fig. 27</figureCitation>
|
||
). S9 black basally with obscure pale area laterally, remainder pale blue, black extending central dorsally to ca. half segment length, posterior carina black but area behind blue (
|
||
<figureCitation id="BECDFB9CFFA2A14F74FDF987FCB2F9D5" box="[721,852,1563,1590]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1583,1607]" captionTargetBox="[151,1436,489,1560]" captionTargetId="figure-190@10.[151,1436,489,1560]" captionTargetPageId="10" captionText="FIGURES 26–31. Terminal abdominal segments of Coeliccia species. Coeliccia junis sp. nov. holotype male: (26) dorsal view; (27) lateral view. Coeliccia roberti sp. nov. holotype male: (28) dorsal view; (29) lateral view. Coeliccia roberti sp. nov., female described: (30) dorsal view; (31) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306475" httpUri="https://zenodo.org/record/4306475/files/figure.png" pageId="3" pageNumber="476">Figs 26–27</figureCitation>
|
||
). S10 pale blue with narrow dark brown areas at apical margin (
|
||
<figureCitation id="BECDFB9CFFA2A14F7736F9A3FE7BF9B9" box="[282,413,1599,1626]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1583,1607]" captionTargetBox="[151,1436,489,1560]" captionTargetId="figure-190@10.[151,1436,489,1560]" captionTargetPageId="10" captionText="FIGURES 26–31. Terminal abdominal segments of Coeliccia species. Coeliccia junis sp. nov. holotype male: (26) dorsal view; (27) lateral view. Coeliccia roberti sp. nov. holotype male: (28) dorsal view; (29) lateral view. Coeliccia roberti sp. nov., female described: (30) dorsal view; (31) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306475" httpUri="https://zenodo.org/record/4306475/files/figure.png" pageId="3" pageNumber="476">Figs 26–27</figureCitation>
|
||
). Very broad and relatively long, black, slightly bi-lobed epiproct visible in dorsal view (
|
||
<figureCitation id="BECDFB9CFFA2A14F7343F9A3FF53F99D" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1972,1996]" captionTargetBox="[415,1173,181,1948]" captionTargetId="figure-15@13.[415,1173,181,1948]" captionTargetPageId="13" captionText="FIGURES 32–34. Dorsal view of anal appendages of male Coeliccia species, all scale bars 0.5mm. (32) Coeliccia junis sp. nov. holotype. (33) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (34) Coeliccia roberti sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.4306477" httpUri="https://zenodo.org/record/4306477/files/figure.png" pageId="3" pageNumber="476">Fig. 32</figureCitation>
|
||
, clearer in
|
||
<figureCitation id="BECDFB9CFFA2A14F771DF9FFFE67F99D" box="[305,385,1635,1662]" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1972,1996]" captionTargetBox="[415,1173,181,1948]" captionTargetId="figure-15@13.[415,1173,181,1948]" captionTargetPageId="13" captionText="FIGURES 32–34. Dorsal view of anal appendages of male Coeliccia species, all scale bars 0.5mm. (32) Coeliccia junis sp. nov. holotype. (33) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (34) Coeliccia roberti sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.4306477" httpUri="https://zenodo.org/record/4306477/files/figure.png" pageId="3" pageNumber="476">Fig. 33</figureCitation>
|
||
which shows a
|
||
<typeStatus id="F94D59BBFFA2A14F7418F9F8FD71F99D" box="[564,663,1636,1662]" pageId="3" pageNumber="476" type="paratype">paratype</typeStatus>
|
||
,
|
||
<figureCitation id="BECDFB9CFFA2A14F748FF9FFFD14F99D" box="[675,754,1635,1662]" captionStart="FIGURES 35–37" captionStartId="14.[151,264,791,815]" captionTargetBox="[157,1429,181,768]" captionTargetId="figure-18@14.[151,1436,181,768]" captionTargetPageId="14" captionText="FIGURES 35–37. Sketches of the epiproct of Coeliccia species in dorsal view, also showing inner basal parts of the cerci: (35) Coeliccia junis paratype SAR18_PCD27; (36) Coeliccia nigrohamata SAR06-PPCD83, Sabaat River, foot of Gunung Pueh, Kuching Division, Sarawak, leg. R.A. Dow, included for comparison; (37) Coeliccia roberti paratype BRU14_PCD17." figureDoi="http://doi.org/10.5281/zenodo.4306479" httpUri="https://zenodo.org/record/4306479/files/figure.png" pageId="3" pageNumber="476">Fig. 35</figureCitation>
|
||
shows a sketch of the epiproct in dorsal view made from the same
|
||
<typeStatus id="F94D59BBFFA2A14F76FAF914FEDEF941" box="[214,312,1672,1698]" pageId="3" pageNumber="476" type="paratype">paratype</typeStatus>
|
||
) and raised up so just visible in lateral view (indicated with a white line in
|
||
<figureCitation id="BECDFB9CFFA2A14F7246F91BFB5FF941" box="[1130,1209,1671,1698]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="3" pageNumber="476">Fig. 38</figureCitation>
|
||
) as well. Cerci (
|
||
<figureCitation id="BECDFB9CFFA2A14F7346F91BFF53F925" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1972,1996]" captionTargetBox="[415,1173,181,1948]" captionTargetId="figure-15@13.[415,1173,181,1948]" captionTargetPageId="13" captionText="FIGURES 32–34. Dorsal view of anal appendages of male Coeliccia species, all scale bars 0.5mm. (32) Coeliccia junis sp. nov. holotype. (33) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (34) Coeliccia roberti sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.4306477" httpUri="https://zenodo.org/record/4306477/files/figure.png" pageId="3" pageNumber="476">Figs 32</figureCitation>
|
||
,
|
||
<figureCitation id="BECDFB9CFFA2A14F76EFF930FF06F925" box="[195,224,1708,1734]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="3" pageNumber="476">38</figureCitation>
|
||
) mostly blue, dark interiorly, narrowly black above apically; basal tooth present but hidden under epiproct so that only visible in an end-on view, strong central ventral interior tooth with apex before half-length, visible in lateral view (
|
||
<figureCitation id="BECDFB9CFFA2A14F7705F96FFE98F8ED" box="[297,382,1779,1806]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="3" pageNumber="476">Fig. 38</figureCitation>
|
||
). In dorsal view (
|
||
<figureCitation id="BECDFB9CFFA2A14F746CF96FFD77F8ED" box="[576,657,1779,1806]" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1972,1996]" captionTargetBox="[415,1173,181,1948]" captionTargetId="figure-15@13.[415,1173,181,1948]" captionTargetPageId="13" captionText="FIGURES 32–34. Dorsal view of anal appendages of male Coeliccia species, all scale bars 0.5mm. (32) Coeliccia junis sp. nov. holotype. (33) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (34) Coeliccia roberti sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.4306477" httpUri="https://zenodo.org/record/4306477/files/figure.png" pageId="3" pageNumber="476">Fig. 32</figureCitation>
|
||
) each cercus subrectangular, apex at outer side, distinct ridge running diagonally forwards from apex part way to inner margin. In lateral view (
|
||
<figureCitation id="BECDFB9CFFA2A14F75E8F88BFBF3F8D1" box="[964,1045,1815,1842]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="3" pageNumber="476">Fig. 38</figureCitation>
|
||
) cercus tapering slightly from base to rounded apex, almost missile shaped. Paraprocts (
|
||
<figureCitation id="BECDFB9CFFA2A14F74CCF8A7FCD1F8B5" box="[736,823,1851,1878]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="3" pageNumber="476">Figs 38</figureCitation>
|
||
,
|
||
<figureCitation id="BECDFB9CFFA2A14F756EF8A0FCB9F8B5" box="[834,863,1852,1878]" captionStart="FIGURES 42–45" captionStartId="15.[151,264,1132,1156]" captionTargetBox="[151,1435,181,1108]" captionTargetId="figure-15@15.[151,1435,181,1108]" captionTargetPageId="15" captionText="FIGURES 42–45. Ventral view of anal appendages of male Coeliccia species. Scale bars (where present) 0.2mm. (42) Coeliccia junis sp. nov. holotype. (43) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (44) Coeliccia roberti sp. nov. holotype. (45) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306483" httpUri="https://zenodo.org/record/4306483/files/figure.png" pageId="3" pageNumber="476">42</figureCitation>
|
||
) mostly whitish in visible parts, tips reaching same level of those of cerci, turned inwards and slightly down apically; in ventral view running almost straight after base along outer margin before turning abruptly but smoothly inwards towards apices (
|
||
<figureCitation id="BECDFB9CFFA2A14F7218F81FFB6EF87D" box="[1076,1160,1923,1950]" captionStart="FIGURES 42–45" captionStartId="15.[151,264,1132,1156]" captionTargetBox="[151,1435,181,1108]" captionTargetId="figure-15@15.[151,1435,181,1108]" captionTargetPageId="15" captionText="FIGURES 42–45. Ventral view of anal appendages of male Coeliccia species. Scale bars (where present) 0.2mm. (42) Coeliccia junis sp. nov. holotype. (43) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (44) Coeliccia roberti sp. nov. holotype. (45) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306483" httpUri="https://zenodo.org/record/4306483/files/figure.png" pageId="3" pageNumber="476">Fig. 42</figureCitation>
|
||
), contracting to strongly in-turned apices along inner margin from ca. two-thirds length, so that the inner margin is a semicircle in this part, the apices almost touching (
|
||
<figureCitation id="BECDFB9CFFA2A14F77E2F857FDC4F805" box="[462,546,1995,2022]" captionStart="FIGURES 42–45" captionStartId="15.[151,264,1132,1156]" captionTargetBox="[151,1435,181,1108]" captionTargetId="figure-15@15.[151,1435,181,1108]" captionTargetPageId="15" captionText="FIGURES 42–45. Ventral view of anal appendages of male Coeliccia species. Scale bars (where present) 0.2mm. (42) Coeliccia junis sp. nov. holotype. (43) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (44) Coeliccia roberti sp. nov. holotype. (45) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306483" httpUri="https://zenodo.org/record/4306483/files/figure.png" pageId="3" pageNumber="476">Fig. 42</figureCitation>
|
||
). In lateral view paraprocts (
|
||
<figureCitation id="BECDFB9CFFA2A14F754CF857FC56F805" box="[864,944,1995,2022]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="3" pageNumber="476">Fig. 38</figureCitation>
|
||
) with lower margin running gently upwards from base to point of down turn, but along upper margin with an abrupt, step-like contraction just before one-third length, thereafter tapering gradually towards apices. In ventral view apical part of paraprocts appearing narrow, but actually rather flattened so that this is an artefact of the view (see
|
||
<figureCitation id="BECDFB9CFFA5A1487554FF43FC2DFF1A" box="[888,971,223,249]" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1972,1996]" captionTargetBox="[415,1173,181,1948]" captionTargetId="figure-15@13.[415,1173,181,1948]" captionTargetPageId="13" captionText="FIGURES 32–34. Dorsal view of anal appendages of male Coeliccia species, all scale bars 0.5mm. (32) Coeliccia junis sp. nov. holotype. (33) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (34) Coeliccia roberti sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.4306477" httpUri="https://zenodo.org/record/4306477/files/figure.png" pageId="4" pageNumber="477">Fig. 32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA5A14876EBFE9FFE0DFEA1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="477">
|
||
<emphasis id="14823B0BFFA5A14876EBFE9FFE83FEFE" box="[199,357,259,285]" italics="true" pageId="4" pageNumber="477">Genital ligula</emphasis>
|
||
: only examined in a
|
||
<typeStatus id="F94D59BBFFA5A1487477FE9FFD5DFEFE" box="[603,699,259,285]" pageId="4" pageNumber="477" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="BECDFB9CFFA5A14874E1FE9FFCC4FEFE" box="[717,802,259,285]" captionStart="FIGURES 24–25" captionStartId="9.[151,264,1486,1510]" captionTargetBox="[151,1436,881,1463]" captionTargetId="figure-96@9.[151,1436,881,1463]" captionTargetPageId="9" captionText="FIGURES 24–25. Genital ligula of Coeliccia species in ventral view: (24) Coeliccia junis paratype SAR18_PCD27. (25) Scan- ning Electron Microscope image of Coeliccia roberti SAR06-PCD77, not in type series, Lambir Hills National Park. All scale bars 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4306473" httpUri="https://zenodo.org/record/4306473/files/figure.png" pageId="4" pageNumber="477">Fig. 24</figureCitation>
|
||
), of typical form for the
|
||
<taxonomicName id="E1F69C9AFFA5A148726EFE9FFAABFEFE" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[1090,1357,259,285]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA5A148726EFE9FFAABFEFE" box="[1090,1357,259,285]" italics="true" pageId="4" pageNumber="477">Coeliccia membranipes</emphasis>
|
||
</taxonomicName>
|
||
–group (also see
|
||
<figureCitation id="BECDFB9CFFA5A14876D3FEBBFEA7FEA2" box="[255,321,295,321]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1903,1927]" captionTargetBox="[167,1421,414,1877]" captionTargetId="figure-153@2.[167,1421,414,1877]" captionTargetPageId="2" captionText="FIGURES 1–9. Heads of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view. Coeliccia roberti sp. nov. holotype male: (4) dorsal view; (5) frontal view; (6) ventral view. Coeliccia roberti sp. nov. female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="4" pageNumber="477">
|
||
Fig.
|
||
<quantity id="E10E4AFCFFA5A148771FFEBBFEA7FEA2" box="[307,321,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="477" unit="in" value="1.0">1</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="42679AE8FFA5A1487749FEBBFE39FEA1" author="Dow, R. A." box="[357,479,295,322]" pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA5A14876EBFED7FC41FE86" blockId="4.[151,1437,151,2013]" box="[199,935,330,357]" pageId="4" pageNumber="477">Measurements (mm): Abdomen with anal appendages 35, Hw 21.</paragraph>
|
||
<paragraph id="2649E719FFA5A14876EBFEF3FE44FE6A" blockId="4.[151,1437,151,2013]" box="[199,418,367,393]" pageId="4" pageNumber="477">
|
||
<emphasis id="14823B0BFFA5A14876EBFEF3FEC3FE6A" bold="true" box="[199,293,367,393]" pageId="4" pageNumber="477">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA5A14876EBFE0EFC2CFCF6" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="477">
|
||
<emphasis id="14823B0BFFA5A14876EBFE0EFE3AFE4E" bold="true" box="[199,476,402,429]" pageId="4" pageNumber="477">
|
||
Variation in
|
||
<typeStatus id="F94D59BBFFA5A1487772FE0FFE30FE4E" box="[350,470,403,429]" pageId="4" pageNumber="477" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Note that the two
|
||
<typeStatus id="F94D59BBFFA5A1487483FE0FFCFCFE4E" box="[687,794,403,429]" pageId="4" pageNumber="477" type="paratype">paratypes</typeStatus>
|
||
(SAR18_PCD27 is shown in
|
||
<figureCitation id="BECDFB9CFFA5A1487242FE0FFB08FE4E" box="[1134,1262,403,429]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-18@8.[151,1436,181,1680]" captionTargetPageId="8" captionText="FIGURES 16–21. Synthorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (16) dorsal view; (17) lateral view. Coeliccia junis sp. nov. paratype male (SAR18_PCD27): (18) dorsal view; (19) lateral view. Coeliccia roberti sp. nov. holotype male: (20) dorsal view; (21) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306469" httpUri="https://zenodo.org/record/4306469/files/figure.png" pageId="4" pageNumber="477">Figs 18–19</figureCitation>
|
||
,
|
||
<figureCitation id="BECDFB9CFFA5A14872D7FE0FFAFFFE4E" box="[1275,1305,403,429]" captionStart="FIGURES 24–25" captionStartId="9.[151,264,1486,1510]" captionTargetBox="[151,1436,881,1463]" captionTargetId="figure-96@9.[151,1436,881,1463]" captionTargetPageId="9" captionText="FIGURES 24–25. Genital ligula of Coeliccia species in ventral view: (24) Coeliccia junis paratype SAR18_PCD27. (25) Scan- ning Electron Microscope image of Coeliccia roberti SAR06-PCD77, not in type series, Lambir Hills National Park. All scale bars 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4306473" httpUri="https://zenodo.org/record/4306473/files/figure.png" pageId="4" pageNumber="477">24</figureCitation>
|
||
,
|
||
<figureCitation id="BECDFB9CFFA5A1487309FE0FFAA5FE4E" box="[1317,1347,403,429]" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1972,1996]" captionTargetBox="[415,1173,181,1948]" captionTargetId="figure-15@13.[415,1173,181,1948]" captionTargetPageId="13" captionText="FIGURES 32–34. Dorsal view of anal appendages of male Coeliccia species, all scale bars 0.5mm. (32) Coeliccia junis sp. nov. holotype. (33) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (34) Coeliccia roberti sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.4306477" httpUri="https://zenodo.org/record/4306477/files/figure.png" pageId="4" pageNumber="477">33</figureCitation>
|
||
,
|
||
<figureCitation id="BECDFB9CFFA5A148737CFE0EFA8AFE4E" box="[1360,1388,402,429]" captionStart="FIGURES 35–37" captionStartId="14.[151,264,791,815]" captionTargetBox="[157,1429,181,768]" captionTargetId="figure-18@14.[151,1436,181,768]" captionTargetPageId="14" captionText="FIGURES 35–37. Sketches of the epiproct of Coeliccia species in dorsal view, also showing inner basal parts of the cerci: (35) Coeliccia junis paratype SAR18_PCD27; (36) Coeliccia nigrohamata SAR06-PPCD83, Sabaat River, foot of Gunung Pueh, Kuching Division, Sarawak, leg. R.A. Dow, included for comparison; (37) Coeliccia roberti paratype BRU14_PCD17." figureDoi="http://doi.org/10.5281/zenodo.4306479" httpUri="https://zenodo.org/record/4306479/files/figure.png" pageId="4" pageNumber="477">35</figureCitation>
|
||
and
|
||
<figureCitation id="BECDFB9CFFA5A14876BBFE2BFF53FE32" box="[151,181,439,465]" captionStart="FIGURES 42–45" captionStartId="15.[151,264,1132,1156]" captionTargetBox="[151,1435,181,1108]" captionTargetId="figure-15@15.[151,1435,181,1108]" captionTargetPageId="15" captionText="FIGURES 42–45. Ventral view of anal appendages of male Coeliccia species. Scale bars (where present) 0.2mm. (42) Coeliccia junis sp. nov. holotype. (43) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (44) Coeliccia roberti sp. nov. holotype. (45) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306483" httpUri="https://zenodo.org/record/4306483/files/figure.png" pageId="4" pageNumber="477">43</figureCitation>
|
||
, SAR18_PCD
|
||
<quantity id="E10E4AFCFFA5A1487770FE2BFE71FE32" box="[348,407,439,465]" metricMagnitude="-1" metricUnit="m" metricValue="7.112" pageId="4" pageNumber="477" unit="in" value="28.0">28 in</quantity>
|
||
<figureCitation id="BECDFB9CFFA5A148778DFE2BFE13FE31" box="[417,501,439,466]" captionStart="FIGURES 46–47" captionStartId="15.[151,264,1790,1814]" captionTargetBox="[151,1437,1252,1766]" captionTargetId="figure-60@15.[151,1437,1252,1766]" captionTargetPageId="15" captionText="FIGURES 46–47. Ventral view of cerci of male Coeliccia species. (46) Coeliccia junis sp. nov. paratype (SAR18_PCD28) with right paraproct removed to allow ventral view of the right cercus. (47) Detail of Coeliccia roberti sp. nov. (SAR06-PCD151, not in type series) with paraprocts removed to allow ventral view of the cerci. Scale bars 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4306485" httpUri="https://zenodo.org/record/4306485/files/figure.png" pageId="4" pageNumber="477">Fig. 46</figureCitation>
|
||
) are somewhat discoloured after being stored in suboptimal conditions for some time and subsequently treated for fungus. The interior excision in the inner margin of the antehumeral stripes in both
|
||
<typeStatus id="F94D59BBFFA5A14876FCFE63FEDDFDFA" box="[208,315,511,537]" pageId="4" pageNumber="477" type="paratype">paratypes</typeStatus>
|
||
is a slightly different shape to that in the
|
||
<typeStatus id="F94D59BBFFA5A1487522FE63FC96FDFA" box="[782,880,511,537]" pageId="4" pageNumber="477" type="holotype">holotype</typeStatus>
|
||
(compare
|
||
<figureCitation id="BECDFB9CFFA5A14875C4FE63FB74FDFA" box="[1000,1170,511,537]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-18@8.[151,1436,181,1680]" captionTargetPageId="8" captionText="FIGURES 16–21. Synthorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (16) dorsal view; (17) lateral view. Coeliccia junis sp. nov. paratype male (SAR18_PCD27): (18) dorsal view; (19) lateral view. Coeliccia roberti sp. nov. holotype male: (20) dorsal view; (21) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306469" httpUri="https://zenodo.org/record/4306469/files/figure.png" pageId="4" pageNumber="477">Figs 16 and 18</figureCitation>
|
||
). There are a few other, insignificant, differences in markings that are not discussed further here. The pterostigma in one
|
||
<typeStatus id="F94D59BBFFA5A14872E2FDBFFAC8FDDE" box="[1230,1326,547,573]" pageId="4" pageNumber="477" type="paratype">paratype</typeStatus>
|
||
is shorter and deeper in all wings than in the
|
||
<typeStatus id="F94D59BBFFA5A148740AFDDBFD6DFD82" box="[550,651,583,609]" pageId="4" pageNumber="477" type="holotype">holotype</typeStatus>
|
||
, and brown rather than black, in the other
|
||
<typeStatus id="F94D59BBFFA5A1487247FDDBFB2DFD82" box="[1131,1227,583,609]" pageId="4" pageNumber="477" type="paratype">paratype</typeStatus>
|
||
the pterostigma is of similar shape to the
|
||
<typeStatus id="F94D59BBFFA5A14877B5FDF7FE1DFD66" box="[409,507,619,645]" pageId="4" pageNumber="477" type="holotype">holotype</typeStatus>
|
||
in all but one wing but is greyish brown. In both
|
||
<typeStatus id="F94D59BBFFA5A1487205FDF7FB72FD66" box="[1065,1172,619,645]" pageId="4" pageNumber="477" type="paratype">paratypes</typeStatus>
|
||
the epiproct is directed more strongly upwards than in the
|
||
<typeStatus id="F94D59BBFFA5A1487436FD13FD9AFD4A" box="[538,636,655,681]" pageId="4" pageNumber="477" type="holotype">holotype</typeStatus>
|
||
so that it is more visible in lateral view (compare
|
||
<figureCitation id="BECDFB9CFFA5A1487284FD13FAA9FD4A" box="[1192,1359,655,681]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="4" pageNumber="477">Figs 38 and 39</figureCitation>
|
||
), since it seems likely that the epiproct is capable of some movement this may merely be the result of its position at preservation rather than genuine variation. In one
|
||
<typeStatus id="F94D59BBFFA5A14874ACFD4BFD06FD12" box="[640,736,727,753]" pageId="4" pageNumber="477" type="paratype">paratype</typeStatus>
|
||
the epiproct is largely pale rather than black. The apices of the paraprocts are not so strongly in-turned in one (compare
|
||
<figureCitation id="BECDFB9CFFA5A1487538FD67FC58FCF6" box="[788,958,763,789]" captionStart="FIGURES 42–45" captionStartId="15.[151,264,1132,1156]" captionTargetBox="[151,1435,181,1108]" captionTargetId="figure-15@15.[151,1435,181,1108]" captionTargetPageId="15" captionText="FIGURES 42–45. Ventral view of anal appendages of male Coeliccia species. Scale bars (where present) 0.2mm. (42) Coeliccia junis sp. nov. holotype. (43) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (44) Coeliccia roberti sp. nov. holotype. (45) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306483" httpUri="https://zenodo.org/record/4306483/files/figure.png" pageId="4" pageNumber="477">Figs 42 and 43</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA5A14876EBFC83FA96FCDA" blockId="4.[151,1437,151,2013]" box="[199,1392,798,825]" pageId="4" pageNumber="477">
|
||
<emphasis id="14823B0BFFA5A14876EBFC83FE90FCDA" bold="true" box="[199,374,799,825]" pageId="4" pageNumber="477">Measurements</emphasis>
|
||
(mm): 16–18 Px in Fw,
|
||
<quantity id="E10E4AFCFFA5A14874A5FC82FD0AFCDA" box="[649,748,798,825]" metricMagnitude="-1" metricUnit="m" metricValue="3.937" metricValueMax="4.064" metricValueMin="3.81" pageId="4" pageNumber="477" unit="in" value="15.5" valueMax="16.0" valueMin="15.0">15–16 in</quantity>
|
||
Hw; abdomen without anal appendages 35; Hw 21.5–22.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6EECB492FFA5A14876EBFCDEFC2AFB41" pageId="4" pageNumber="477" type="diagnosis">
|
||
<paragraph id="2649E719FFA5A14876EBFCDEFC2AFB41" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="477">
|
||
<emphasis id="14823B0BFFA5A14876EBFCDEFEA4FCBF" bold="true" box="[199,322,834,860]" pageId="4" pageNumber="477">Diagnosis.</emphasis>
|
||
Within the
|
||
<taxonomicName id="E1F69C9AFFA5A14877E7FCDFFD31FCBE" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[459,727,835,861]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA5A14877E7FCDFFD31FCBE" box="[459,727,835,861]" italics="true" pageId="4" pageNumber="477">Coeliccia membranipes</emphasis>
|
||
</taxonomicName>
|
||
–group the male of
|
||
<taxonomicName id="E1F69C9AFFA5A1487596FCDFFBF5FCBF" authorityName="Dow" authorityYear="2020" box="[954,1043,835,861]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A1487596FCDFFBF5FCBF" box="[954,1043,835,861]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
is readily separated from those of
|
||
<taxonomicName id="E1F69C9AFFA5A14876BBFCFBFEA9FC62" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[151,335,871,897]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA5A14876BBFCFBFEA9FC62" box="[151,335,871,897]" italics="true" pageId="4" pageNumber="477">C. membranipes</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E1F69C9AFFA5A1487772FCFBFE19FC62" authorityName="Laidlaw" authorityYear="1912" box="[350,511,871,897]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="nemoricola">
|
||
<emphasis id="14823B0BFFA5A1487772FCFBFE19FC62" box="[350,511,871,897]" italics="true" pageId="4" pageNumber="477">C. nemoricola</emphasis>
|
||
</taxonomicName>
|
||
(and allied forms) and
|
||
<taxonomicName id="E1F69C9AFFA5A148752BFCFBFC26FC62" authorityName="Kimmins" authorityYear="1936" box="[775,960,871,897]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="cyaneothorax">
|
||
<emphasis id="14823B0BFFA5A148752BFCFBFC26FC62" box="[775,960,871,897]" italics="true" pageId="4" pageNumber="477">C. cyaneothorax</emphasis>
|
||
</taxonomicName>
|
||
by its excised antehumeral markings. It is also separated from the aforementioned taxa and from
|
||
<taxonomicName id="E1F69C9AFFA5A14874D4FC17FC4FFC46" authorityName="Laidlaw" authorityYear="1918" box="[760,937,907,933]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="nigrohamata">
|
||
<emphasis id="14823B0BFFA5A14874D4FC17FC4FFC46" box="[760,937,907,933]" italics="true" pageId="4" pageNumber="477">C. nigrohamata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E1F69C9AFFA5A148759AFC17FB78FC46" authority="Dow, 2016" authorityName="Dow" authorityYear="2016" box="[950,1182,907,933]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="matok">
|
||
<emphasis id="14823B0BFFA5A148759AFC17FBFCFC46" box="[950,1050,907,933]" italics="true" pageId="4" pageNumber="477">C. matok</emphasis>
|
||
<bibRefCitation id="42679AE8FFA5A148720DFC17FB78FC46" author="Dow, R. A." box="[1057,1182,907,933]" pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E1F69C9AFFA5A1487285FC17FF3DFC29" authority="(Selys, 1863)" baseAuthorityName="Selys" baseAuthorityYear="1863" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="octogesima">
|
||
<emphasis id="14823B0BFFA5A1487285FC17FAAFFC47" box="[1193,1353,907,933]" italics="true" pageId="4" pageNumber="477">C. octogesima</emphasis>
|
||
(Selys, 1863)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="E1F69C9AFFA5A14876CAFC33FE18FC2A" authority="Dow, 2016" authorityName="Dow" authorityYear="2016" box="[230,510,943,969]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="paludensis">
|
||
<emphasis id="14823B0BFFA5A14876CAFC33FE98FC2A" box="[230,382,943,969]" italics="true" pageId="4" pageNumber="477">C. paludensis</emphasis>
|
||
<bibRefCitation id="42679AE8FFA5A14877A8FC33FE18FC2A" author="Dow, R. A." box="[388,510,943,969]" pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E1F69C9AFFA5A148741FFC33FD44FC2B" authorityName="Lieftinck" authorityYear="1953" box="[563,674,943,969]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="resecta">
|
||
<emphasis id="14823B0BFFA5A148741FFC33FD44FC2B" box="[563,674,943,969]" italics="true" pageId="4" pageNumber="477">C. resecta</emphasis>
|
||
</taxonomicName>
|
||
by its broad epiproct. The size of the epiproct also separates
|
||
<taxonomicName id="E1F69C9AFFA5A148736AFC33FA7AFC2B" authorityName="Dow" authorityYear="2020" box="[1350,1436,943,969]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A148736AFC33FA7AFC2B" box="[1350,1436,943,969]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="E1F69C9AFFA5A14876F8FC4FFDCAFC0E" authority="Laidlaw, 1918" authorityName="Laidlaw" authorityYear="1918" box="[212,556,979,1005]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="macrostigma">
|
||
<emphasis id="14823B0BFFA5A14876F8FC4FFE60FC0F" box="[212,390,979,1005]" italics="true" pageId="4" pageNumber="477">C. macrostigma</emphasis>
|
||
<bibRefCitation id="42679AE8FFA5A14877A1FC4FFDCAFC0E" author="Laidlaw, F. F." box="[397,556,979,1005]" pageId="4" pageNumber="477" pagination="223 - 232" refId="ref12643" refString="Laidlaw, F. F. (1918) Some additions to the known dragonfly fauna of Borneo, with an account of new species of the genus Coeliccia. Proceedings of the Zoological Society of London, 1918, 223 - 232, figs. 2 - 9. https: // doi. org / 10.1111 / j. 1096 - 3642.1917. tb 02061. x" type="journal article" year="1918">Laidlaw, 1918</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E1F69C9AFFA5A148744EFC4FFD29FC0E" authority="Dow, 2020" authorityName="Dow" authorityYear="2020" box="[610,719,979,1005]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="roberti" status="sp. nov.">
|
||
<emphasis id="14823B0BFFA5A148744EFC4FFD29FC0E" box="[610,719,979,1005]" italics="true" pageId="4" pageNumber="477">C. roberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="14823B0BFFA5A14874FAFC4FFCC9FC0E" bold="true" box="[726,815,979,1005]" pageId="4" pageNumber="477">
|
||
<taxonomicNameLabel id="0FB18670FFA5A14874FAFC4FFCC9FC0E" box="[726,815,979,1005]" pageId="4" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but in these species the epiproct is also broad although nowhere near so large.
|
||
<taxonomicName id="E1F69C9AFFA5A14877B3FC6BFDAEFBF3" authorityName="Dow" authorityYear="2020" box="[415,584,1015,1041]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A14877B3FC6BFDAEFBF3" box="[415,584,1015,1041]" italics="true" pageId="4" pageNumber="477">Coeliccia junis</emphasis>
|
||
</taxonomicName>
|
||
is also separated from
|
||
<taxonomicName id="E1F69C9AFFA5A148757CFC6BFC58FBF2" authorityName="Dow" authorityYear="2020" box="[848,958,1015,1041]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="roberti">
|
||
<emphasis id="14823B0BFFA5A148757CFC6BFC58FBF2" box="[848,958,1015,1041]" italics="true" pageId="4" pageNumber="477">C. roberti</emphasis>
|
||
</taxonomicName>
|
||
by the position of the ventral tooth on the cercus and indeed the details and position of this tooth also separate it from most other species in the group. In
|
||
<taxonomicName id="E1F69C9AFFA5A14873AFFB87FECEFBBB" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="macrostigma">
|
||
<emphasis id="14823B0BFFA5A14873AFFB87FECEFBBB" italics="true" pageId="4" pageNumber="477">C. macrostigma</emphasis>
|
||
</taxonomicName>
|
||
the ventral tooth of the cercus is in a similar position to that of
|
||
<taxonomicName id="E1F69C9AFFA5A14875DCFBA3FBA1FBBB" authorityName="Dow" authorityYear="2020" box="[1008,1095,1087,1113]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A14875DCFBA3FBA1FBBB" box="[1008,1095,1087,1113]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
but shaped differently and the apex of the cercus is also a different shape (compare
|
||
<figureCitation id="BECDFB9CFFA5A14874F1FBFFFCBFFB9E" box="[733,857,1123,1149]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="4" pageNumber="477">Figs 38–39</figureCitation>
|
||
here with Fig.
|
||
<quantity id="E10E4AFCFFA5A14875D2FBFFFBD0FB9E" box="[1022,1078,1123,1149]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="477" unit="in" value="60.0">60 in</quantity>
|
||
<bibRefCitation id="42679AE8FFA5A1487217FBFFFB2AFB9D" author="Dow, R. A." box="[1083,1228,1123,1150]" pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow (2016))</bibRefCitation>
|
||
and the paraprocts are also different (compare
|
||
<figureCitation id="BECDFB9CFFA5A14877E0FB1BFDAFFB42" box="[460,585,1159,1185]" captionStart="FIGURES 42–45" captionStartId="15.[151,264,1132,1156]" captionTargetBox="[151,1435,181,1108]" captionTargetId="figure-15@15.[151,1435,181,1108]" captionTargetPageId="15" captionText="FIGURES 42–45. Ventral view of anal appendages of male Coeliccia species. Scale bars (where present) 0.2mm. (42) Coeliccia junis sp. nov. holotype. (43) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (44) Coeliccia roberti sp. nov. holotype. (45) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306483" httpUri="https://zenodo.org/record/4306483/files/figure.png" pageId="4" pageNumber="477">Figs 42–43</figureCitation>
|
||
here with Fig.
|
||
<quantity id="E10E4AFCFFA5A14874D8FB1AFCCBFB42" box="[756,813,1158,1185]" metricMagnitude="0" metricUnit="m" metricValue="1.6509999999999998" pageId="4" pageNumber="477" unit="in" value="65.0">65 in</quantity>
|
||
<bibRefCitation id="42679AE8FFA5A148751FFB1BFC2FFB41" author="Dow, R. A." box="[819,969,1159,1186]" pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow (2016))</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6EECB492FFA5A14976EBFB37FB94FE6A" lastPageId="5" lastPageNumber="478" pageId="4" pageNumber="477" type="discussion">
|
||
<paragraph id="2649E719FFA5A14876EBFB37FCDCFA22" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="477">
|
||
<emphasis id="14823B0BFFA5A14876EBFB37FEDDFB26" bold="true" box="[199,315,1195,1221]" pageId="4" pageNumber="477">Remarks.</emphasis>
|
||
To expand upon the characters mentioned in the diagnosis, the details of the male antehumeral markings of
|
||
<taxonomicName id="E1F69C9AFFA5A1487706FB53FE30FB0B" authorityName="Dow" authorityYear="2020" box="[298,470,1231,1257]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A1487706FB53FE30FB0B" box="[298,470,1231,1257]" italics="true" pageId="4" pageNumber="477">Coeliccia junis</emphasis>
|
||
</taxonomicName>
|
||
also clearly separate it from all named species in the
|
||
<taxonomicName id="E1F69C9AFFA5A1487261FB53FAE1FB0A" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[1101,1287,1231,1257]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA5A1487261FB53FAE1FB0A" box="[1101,1287,1231,1257]" italics="true" pageId="4" pageNumber="477">C. membranipes</emphasis>
|
||
</taxonomicName>
|
||
–group, even those where these markings are excised (for instance compare
|
||
<figureCitation id="BECDFB9CFFA5A1487578FB6FFC1CFAEE" box="[852,1018,1267,1293]" captionStart="FIGURES 16–21" captionStartId="8.[151,264,1704,1728]" captionTargetBox="[151,1436,181,1680]" captionTargetId="figure-18@8.[151,1436,181,1680]" captionTargetPageId="8" captionText="FIGURES 16–21. Synthorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (16) dorsal view; (17) lateral view. Coeliccia junis sp. nov. paratype male (SAR18_PCD27): (18) dorsal view; (19) lateral view. Coeliccia roberti sp. nov. holotype male: (20) dorsal view; (21) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306469" httpUri="https://zenodo.org/record/4306469/files/figure.png" pageId="4" pageNumber="477">Figs 16 and 18</figureCitation>
|
||
here with
|
||
<figureCitation id="BECDFB9CFFA5A148725EFB6FFB09FAEE" box="[1138,1263,1266,1293]" captionStart="FIGURES 35–37" captionStartId="14.[151,264,791,815]" captionTargetBox="[157,1429,181,768]" captionTargetId="figure-18@14.[151,1436,181,768]" captionTargetPageId="14" captionText="FIGURES 35–37. Sketches of the epiproct of Coeliccia species in dorsal view, also showing inner basal parts of the cerci: (35) Coeliccia junis paratype SAR18_PCD27; (36) Coeliccia nigrohamata SAR06-PPCD83, Sabaat River, foot of Gunung Pueh, Kuching Division, Sarawak, leg. R.A. Dow, included for comparison; (37) Coeliccia roberti paratype BRU14_PCD17." figureDoi="http://doi.org/10.5281/zenodo.4306479" httpUri="https://zenodo.org/record/4306479/files/figure.png" pageId="4" pageNumber="477">Figs 35, 37</figureCitation>
|
||
and
|
||
<quantity id="E10E4AFCFFA5A148730AFB6FFAB9FAEE" box="[1318,1375,1267,1293]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" pageId="4" pageNumber="477" unit="in" value="39.0">
|
||
<figureCitation id="BECDFB9CFFA5A148730AFB6FFAA4FAEE" box="[1318,1346,1267,1293]" captionStart="FIGURES 38–41" captionStartId="14.[151,264,1936,1960]" captionTargetBox="[151,1437,912,1912]" captionTargetId="figure-70@14.[151,1437,912,1912]" captionTargetPageId="14" captionText="FIGURES 38–41. Lateral view of anal appendages of male Coeliccia species, scale bars on Figs. 38–40 0.5mm. (38) Coeliccia junis sp. nov. holotype, right hand side flipped to left; epiproct indicated with a white line. (39) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (40) Coeliccia roberti sp. nov. holotype. (41) Coeliccia roberti sp. nov. paratype (SAR06-PCD103)." figureDoi="http://doi.org/10.5281/zenodo.4306481" httpUri="https://zenodo.org/record/4306481/files/figure.png" pageId="4" pageNumber="477">39</figureCitation>
|
||
in
|
||
</quantity>
|
||
<bibRefCitation id="42679AE8FFA5A148734AFB6FFF0BFAD1" author="Dow, R. A." pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow (2016))</bibRefCitation>
|
||
, however since there is clearly some variation in the markings of
|
||
<taxonomicName id="E1F69C9AFFA5A14875E8FA8BFBFCFAD3" authorityName="Dow" authorityYear="2020" box="[964,1050,1303,1329]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A14875E8FA8BFBFCFAD3" box="[964,1050,1303,1329]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
this character is not stressed in the diagnosis. Another character that appears to separate
|
||
<taxonomicName id="E1F69C9AFFA5A14874DEFAA7FCACFAB7" authorityName="Dow" authorityYear="2020" box="[754,842,1339,1365]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A14874DEFAA7FCACFAB7" box="[754,842,1339,1365]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
from both
|
||
<taxonomicName id="E1F69C9AFFA5A14875E7FAA7FB98FAB7" box="[971,1150,1339,1365]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="macrostigma">
|
||
<emphasis id="14823B0BFFA5A14875E7FAA7FB98FAB7" box="[971,1150,1339,1365]" italics="true" pageId="4" pageNumber="477">C. macrostigma</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E1F69C9AFFA5A148729BFAA7FAC3FAB6" authorityName="Dow" authorityYear="2020" box="[1207,1317,1339,1365]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="roberti">
|
||
<emphasis id="14823B0BFFA5A148729BFAA7FAC3FAB6" box="[1207,1317,1339,1365]" italics="true" pageId="4" pageNumber="477">C. roberti</emphasis>
|
||
</taxonomicName>
|
||
is that the underside of the head is less extensively pale than in those species (compare
|
||
<figureCitation id="BECDFB9CFFA5A14875DFFAC3FBD2FA9A" box="[1011,1076,1375,1401]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1903,1927]" captionTargetBox="[167,1421,414,1877]" captionTargetId="figure-153@2.[167,1421,414,1877]" captionTargetPageId="2" captionText="FIGURES 1–9. Heads of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view. Coeliccia roberti sp. nov. holotype male: (4) dorsal view; (5) frontal view; (6) ventral view. Coeliccia roberti sp. nov. female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="4" pageNumber="477">Fig. 3</figureCitation>
|
||
here with
|
||
<figureCitation id="BECDFB9CFFA5A1487287FAC3FB0AFA9A" box="[1195,1260,1375,1401]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1903,1927]" captionTargetBox="[167,1421,414,1877]" captionTargetId="figure-153@2.[167,1421,414,1877]" captionTargetPageId="2" captionText="FIGURES 1–9. Heads of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view. Coeliccia roberti sp. nov. holotype male: (4) dorsal view; (5) frontal view; (6) ventral view. Coeliccia roberti sp. nov. female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="4" pageNumber="477">Fig. 9</figureCitation>
|
||
and with
|
||
<figureCitation id="BECDFB9CFFA5A1487377FAC3FA7AFA9A" box="[1371,1436,1375,1401]" captionStart="FIGURES 1–9" captionStartId="2.[151,264,1903,1927]" captionTargetBox="[167,1421,414,1877]" captionTargetId="figure-153@2.[167,1421,414,1877]" captionTargetPageId="2" captionText="FIGURES 1–9. Heads of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (1) dorsal view; (2) dorsal-frontal view; (3) ventral view. Coeliccia roberti sp. nov. holotype male: (4) dorsal view; (5) frontal view; (6) ventral view. Coeliccia roberti sp. nov. female described: (7) dorsal view; (8) dorsal-frontal view; (9) ventral view." figureDoi="http://doi.org/10.5281/zenodo.4306461" httpUri="https://zenodo.org/record/4306461/files/figure.png" pageId="4" pageNumber="477">
|
||
Fig.
|
||
<quantity id="E10E4AFCFFA5A14873A2FAC3FA7AFA9A" box="[1422,1436,1375,1401]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="4" pageNumber="477" unit="in" value="7.0">7</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="42679AE8FFA5A1487698FA1FFEA1FA7D" author="Dow, R. A." box="[180,327,1411,1438]" pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow (2016))</bibRefCitation>
|
||
, however this character is not unique within the
|
||
<taxonomicName id="E1F69C9AFFA5A1487549FA1FFBFBFA7E" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[869,1053,1411,1437]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA5A1487549FA1FFBFBFA7E" box="[869,1053,1411,1437]" italics="true" pageId="4" pageNumber="477">C. membranipes</emphasis>
|
||
</taxonomicName>
|
||
–group (for instance see
|
||
<figureCitation id="BECDFB9CFFA5A148731CFA1FFA66FA7E" box="[1328,1408,1411,1437]" captionStart="FIGURES 10–13" captionStartId="3.[151,264,1205,1229]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure-15@3.[151,1437,181,1181]" captionTargetPageId="3" captionText="FIGURES 10–13. Prothorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (10) dorsal view; (11) lateral view. Coeliccia roberti sp. nov. holotype male: (12) dorsal view; (13) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306463" httpUri="https://zenodo.org/record/4306463/files/figure.png" pageId="4" pageNumber="477">
|
||
Fig.
|
||
<quantity id="E10E4AFCFFA5A1487348FA1FFA66FA7E" box="[1380,1408,1411,1437]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="4" pageNumber="477" unit="in" value="13.0">13</quantity>
|
||
</figureCitation>
|
||
in
|
||
<bibRefCitation id="42679AE8FFA5A14876BBFA3BFECFFA21" author="Dow, R. A." box="[151,297,1447,1474]" pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow (2016))</bibRefCitation>
|
||
and is therefore not more generally diagnostic.
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA5A14876EBFA57FA9DF979" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="477">
|
||
The shape of the anterior pronotal lobe viewed dorsally (
|
||
<figureCitation id="BECDFB9CFFA5A1487561FA57FC79FA06" box="[845,927,1483,1509]" captionStart="FIGURES 10–13" captionStartId="3.[151,264,1205,1229]" captionTargetBox="[151,1437,181,1181]" captionTargetId="figure-15@3.[151,1437,181,1181]" captionTargetPageId="3" captionText="FIGURES 10–13. Prothorax of males of Coeliccia species. Coeliccia junis sp. nov. holotype male: (10) dorsal view; (11) lateral view. Coeliccia roberti sp. nov. holotype male: (12) dorsal view; (13) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306463" httpUri="https://zenodo.org/record/4306463/files/figure.png" pageId="4" pageNumber="477">Fig. 10</figureCitation>
|
||
) is unusual in
|
||
<taxonomicName id="E1F69C9AFFA5A1487264FA57FB79FA07" authorityName="Dow" authorityYear="2020" box="[1096,1183,1483,1509]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A1487264FA57FB79FA07" box="[1096,1183,1483,1509]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
. In most species from the
|
||
<taxonomicName id="E1F69C9AFFA5A14876ECFA73FE9EF9EA" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[192,376,1519,1545]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA5A14876ECFA73FE9EF9EA" box="[192,376,1519,1545]" italics="true" pageId="4" pageNumber="477">C. membranipes</emphasis>
|
||
</taxonomicName>
|
||
–group the sides of the main raised part of the anterior pronotal lobe converge slightly after the anterior carina or remain approximately parallel (sometimes rounded) to the rear in dorsal view but in
|
||
<taxonomicName id="E1F69C9AFFA5A1487322F98FFA83F9CF" authorityName="Dow" authorityYear="2020" box="[1294,1381,1555,1581]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A1487322F98FFA83F9CF" box="[1294,1381,1555,1581]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
they diverge; this character is shared, at least among named species, with only (both sexes of)
|
||
<taxonomicName id="E1F69C9AFFA5A148725EF9ABFAECF9B2" authorityName="Dow" authorityYear="2016" box="[1138,1290,1591,1617]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="paludensis">
|
||
<emphasis id="14823B0BFFA5A148725EF9ABFAECF9B2" box="[1138,1290,1591,1617]" italics="true" pageId="4" pageNumber="477">C. paludensis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="BECDFB9CFFA5A148736FF9ABFA7AF9B2" box="[1347,1436,1591,1617]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1583,1607]" captionTargetBox="[151,1436,489,1560]" captionTargetId="figure-190@10.[151,1436,489,1560]" captionTargetPageId="10" captionText="FIGURES 26–31. Terminal abdominal segments of Coeliccia species. Coeliccia junis sp. nov. holotype male: (26) dorsal view; (27) lateral view. Coeliccia roberti sp. nov. holotype male: (28) dorsal view; (29) lateral view. Coeliccia roberti sp. nov., female described: (30) dorsal view; (31) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306475" httpUri="https://zenodo.org/record/4306475/files/figure.png" pageId="4" pageNumber="477">Figs. 27</figureCitation>
|
||
and
|
||
<quantity id="E10E4AFCFFA5A14876EAF9C7FF19F996" box="[198,255,1627,1653]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="4" pageNumber="477" unit="in" value="33.0">
|
||
<figureCitation id="BECDFB9CFFA5A14876EAF9C7FF04F996" box="[198,226,1627,1653]" captionStart="FIGURES 32–34" captionStartId="13.[151,264,1972,1996]" captionTargetBox="[415,1173,181,1948]" captionTargetId="figure-15@13.[415,1173,181,1948]" captionTargetPageId="13" captionText="FIGURES 32–34. Dorsal view of anal appendages of male Coeliccia species, all scale bars 0.5mm. (32) Coeliccia junis sp. nov. holotype. (33) Coeliccia junis sp. nov. paratype (SAR18_PCD27). (34) Coeliccia roberti sp. nov. holotype." figureDoi="http://doi.org/10.5281/zenodo.4306477" httpUri="https://zenodo.org/record/4306477/files/figure.png" pageId="4" pageNumber="477">33</figureCitation>
|
||
in
|
||
</quantity>
|
||
<bibRefCitation id="42679AE8FFA5A1487729F9C7FE7CF995" author="Dow, R. A." box="[261,410,1627,1654]" pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow (2016))</bibRefCitation>
|
||
. The dome-like bulges on the middle pronotal lobe are also unusual in the
|
||
<taxonomicName id="E1F69C9AFFA5A14872FBF9C7FA68F996" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[1239,1422,1627,1653]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA5A14872FBF9C7FA68F996" box="[1239,1422,1627,1653]" italics="true" pageId="4" pageNumber="477">C. membranipes</emphasis>
|
||
</taxonomicName>
|
||
group but are shared with
|
||
<taxonomicName id="E1F69C9AFFA5A1487790F9E3FD88F97B" box="[444,622,1663,1689]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="macrostigma">
|
||
<emphasis id="14823B0BFFA5A1487790F9E3FD88F97B" box="[444,622,1663,1689]" italics="true" pageId="4" pageNumber="477">C. macrostigma</emphasis>
|
||
</taxonomicName>
|
||
(both sexes) and
|
||
<taxonomicName id="E1F69C9AFFA5A1487518F9E3FC47F97A" authorityName="Dow" authorityYear="2020" box="[820,929,1663,1689]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="roberti">
|
||
<emphasis id="14823B0BFFA5A1487518F9E3FC47F97A" box="[820,929,1663,1689]" italics="true" pageId="4" pageNumber="477">C. roberti</emphasis>
|
||
</taxonomicName>
|
||
(where they are only present in the male).
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA5A14876EBF93FFE4FF892" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="477">
|
||
The very large epiproct of
|
||
<taxonomicName id="E1F69C9AFFA5A14877D0F93FFDB3F95F" authorityName="Dow" authorityYear="2020" box="[508,597,1699,1725]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A14877D0F93FFDB3F95F" box="[508,597,1699,1725]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
is unusual and highly distinctive. The only other
|
||
<taxonomicName id="E1F69C9AFFA5A14872BFF93FFAABF95E" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[1171,1357,1699,1725]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA5A14872BFF93FFAABF95E" box="[1171,1357,1699,1725]" italics="true" pageId="4" pageNumber="477">C. membranipes</emphasis>
|
||
</taxonomicName>
|
||
–group species with an epiproct that is both broad and often visible in dorsal view are
|
||
<taxonomicName id="E1F69C9AFFA5A148722AF95BFB5EF903" box="[1030,1208,1735,1761]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="macrostigma">
|
||
<emphasis id="14823B0BFFA5A148722AF95BFB5EF903" box="[1030,1208,1735,1761]" italics="true" pageId="4" pageNumber="477">C. macrostigma</emphasis>
|
||
</taxonomicName>
|
||
(see Fig.
|
||
<quantity id="E10E4AFCFFA5A148730AF95AFAB9F902" box="[1318,1375,1734,1761]" metricMagnitude="0" metricUnit="m" metricValue="1.4986" pageId="4" pageNumber="477" unit="in" value="59.0">59 in</quantity>
|
||
<bibRefCitation id="42679AE8FFA5A148734AF95BFF0DF8E5" author="Dow, R. A." pageId="4" pageNumber="477" pagination="79 - 103" refId="ref12360" refString="Dow, R. A. (2016) Revision of the genus Coeliccia Kirby in Borneo part II: Two new species from the membranipes - group, with a redescription of C. macrostigma Laidlaw (Odonata: Zygoptera: Platycnemididae). Zootaxa, 4184 (1), 79 - 103. https: // doi. org / 10.11646 / zootaxa. 4184.1.5" type="journal article" year="2016">Dow (2016))</bibRefCitation>
|
||
and
|
||
<taxonomicName id="E1F69C9AFFA5A148770DF977FE69F8E6" authorityName="Dow" authorityYear="2020" box="[289,399,1771,1797]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="roberti">
|
||
<emphasis id="14823B0BFFA5A148770DF977FE69F8E6" box="[289,399,1771,1797]" italics="true" pageId="4" pageNumber="477">C. roberti</emphasis>
|
||
</taxonomicName>
|
||
, but in these the epiproct is significantly smaller and less conspicuous than in
|
||
<taxonomicName id="E1F69C9AFFA5A14872D5F977FAB6F8E7" authorityName="Dow" authorityYear="2020" box="[1273,1360,1771,1797]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A14872D5F977FAB6F8E7" box="[1273,1360,1771,1797]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
. Since it is not very clear in the dorsal views of the anal appendages,
|
||
<figureCitation id="BECDFB9CFFA5A1487569F893FC27F8CA" box="[837,961,1806,1833]" captionStart="FIGURES 35–37" captionStartId="14.[151,264,791,815]" captionTargetBox="[157,1429,181,768]" captionTargetId="figure-18@14.[151,1436,181,768]" captionTargetPageId="14" captionText="FIGURES 35–37. Sketches of the epiproct of Coeliccia species in dorsal view, also showing inner basal parts of the cerci: (35) Coeliccia junis paratype SAR18_PCD27; (36) Coeliccia nigrohamata SAR06-PPCD83, Sabaat River, foot of Gunung Pueh, Kuching Division, Sarawak, leg. R.A. Dow, included for comparison; (37) Coeliccia roberti paratype BRU14_PCD17." figureDoi="http://doi.org/10.5281/zenodo.4306479" httpUri="https://zenodo.org/record/4306479/files/figure.png" pageId="4" pageNumber="477">Figs 35–37</figureCitation>
|
||
show sketches of the epiproct in
|
||
<typeStatus id="F94D59BBFFA5A148731DF893FA7AF8CA" box="[1329,1436,1807,1833]" pageId="4" pageNumber="477" type="paratype">paratypes</typeStatus>
|
||
of
|
||
<taxonomicName id="E1F69C9AFFA5A148769AF8AFFEEAF8AF" authorityName="Dow" authorityYear="2020" box="[182,268,1843,1869]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA5A148769AF8AFFEEAF8AF" box="[182,268,1843,1869]" italics="true" pageId="4" pageNumber="477">C. junis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="BECDFB9CFFA5A1487730F8AFFE89F8AE" box="[284,367,1842,1869]" captionStart="FIGURES 35–37" captionStartId="14.[151,264,791,815]" captionTargetBox="[157,1429,181,768]" captionTargetId="figure-18@14.[151,1436,181,768]" captionTargetPageId="14" captionText="FIGURES 35–37. Sketches of the epiproct of Coeliccia species in dorsal view, also showing inner basal parts of the cerci: (35) Coeliccia junis paratype SAR18_PCD27; (36) Coeliccia nigrohamata SAR06-PPCD83, Sabaat River, foot of Gunung Pueh, Kuching Division, Sarawak, leg. R.A. Dow, included for comparison; (37) Coeliccia roberti paratype BRU14_PCD17." figureDoi="http://doi.org/10.5281/zenodo.4306479" httpUri="https://zenodo.org/record/4306479/files/figure.png" pageId="4" pageNumber="477">Fig. 35</figureCitation>
|
||
),
|
||
<taxonomicName id="E1F69C9AFFA5A14877A8F8AFFE14F8AE" authorityName="Dow" authorityYear="2020" box="[388,498,1843,1869]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="roberti">
|
||
<emphasis id="14823B0BFFA5A14877A8F8AFFE14F8AE" box="[388,498,1843,1869]" italics="true" pageId="4" pageNumber="477">C. roberti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="BECDFB9CFFA5A148742DF8AFFDB4F8AE" box="[513,594,1843,1869]" captionStart="FIGURES 35–37" captionStartId="14.[151,264,791,815]" captionTargetBox="[157,1429,181,768]" captionTargetId="figure-18@14.[151,1436,181,768]" captionTargetPageId="14" captionText="FIGURES 35–37. Sketches of the epiproct of Coeliccia species in dorsal view, also showing inner basal parts of the cerci: (35) Coeliccia junis paratype SAR18_PCD27; (36) Coeliccia nigrohamata SAR06-PPCD83, Sabaat River, foot of Gunung Pueh, Kuching Division, Sarawak, leg. R.A. Dow, included for comparison; (37) Coeliccia roberti paratype BRU14_PCD17." figureDoi="http://doi.org/10.5281/zenodo.4306479" httpUri="https://zenodo.org/record/4306479/files/figure.png" pageId="4" pageNumber="477">Fig. 37</figureCitation>
|
||
) with
|
||
<taxonomicName id="E1F69C9AFFA5A14874B6F8AFFCADF8AE" authorityName="Laidlaw" authorityYear="1918" box="[666,843,1843,1869]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="nigrohamata">
|
||
<emphasis id="14823B0BFFA5A14874B6F8AFFCADF8AE" box="[666,843,1843,1869]" italics="true" pageId="4" pageNumber="477">C. nigrohamata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="BECDFB9CFFA5A1487576F8AFFC4BF8AD" box="[858,941,1843,1870]" captionStart="FIGURES 26–31" captionStartId="10.[151,264,1583,1607]" captionTargetBox="[151,1436,489,1560]" captionTargetId="figure-190@10.[151,1436,489,1560]" captionTargetPageId="10" captionText="FIGURES 26–31. Terminal abdominal segments of Coeliccia species. Coeliccia junis sp. nov. holotype male: (26) dorsal view; (27) lateral view. Coeliccia roberti sp. nov. holotype male: (28) dorsal view; (29) lateral view. Coeliccia roberti sp. nov., female described: (30) dorsal view; (31) lateral view." figureDoi="http://doi.org/10.5281/zenodo.4306475" httpUri="https://zenodo.org/record/4306475/files/figure.png" pageId="4" pageNumber="477">Fig. 26</figureCitation>
|
||
), which has a typical epiproct for the group, included for comparison
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA5A14976EBF8E7FC93FF1A" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1436,151,393]" lastPageId="5" lastPageNumber="478" pageId="4" pageNumber="477">
|
||
The ventral view of the cercus unobscured by the paraprocts shown in
|
||
<figureCitation id="BECDFB9CFFA5A14875C9F8E7FBD3F876" box="[997,1077,1915,1941]" captionStart="FIGURES 46–47" captionStartId="15.[151,264,1790,1814]" captionTargetBox="[151,1437,1252,1766]" captionTargetId="figure-60@15.[151,1437,1252,1766]" captionTargetPageId="15" captionText="FIGURES 46–47. Ventral view of cerci of male Coeliccia species. (46) Coeliccia junis sp. nov. paratype (SAR18_PCD28) with right paraproct removed to allow ventral view of the right cercus. (47) Detail of Coeliccia roberti sp. nov. (SAR06-PCD151, not in type series) with paraprocts removed to allow ventral view of the cerci. Scale bars 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.4306485" httpUri="https://zenodo.org/record/4306485/files/figure.png" pageId="4" pageNumber="477">Fig. 46</figureCitation>
|
||
is instructive as to the structure of the diagnostically important ventral expansion of the cercus, in this view it is revealed to be the most expanded part of a long ridge or fold. The general form is typical for the
|
||
<taxonomicName id="E1F69C9AFFA5A1487577F85FFBF2F83E" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[859,1044,1987,2013]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="477" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA5A1487577F85FFBF2F83E" box="[859,1044,1987,2013]" italics="true" pageId="4" pageNumber="477">C. membranipes</emphasis>
|
||
</taxonomicName>
|
||
–group at least, with the expansion bearing a tooth in its basal part, but the position of the expansion in
|
||
<taxonomicName id="E1F69C9AFFA4A149758CFF0BFC1EFF53" authorityName="Dow" authorityYear="2020" box="[928,1016,151,177]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="478" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA4A149758CFF0BFC1EFF53" box="[928,1016,151,177]" italics="true" pageId="5" pageNumber="478">C. junis</emphasis>
|
||
</taxonomicName>
|
||
is unusually basal compared to most species of the
|
||
<taxonomicName id="E1F69C9AFFA4A1497719FF27FE0AFF36" baseAuthorityName="Rambur" baseAuthorityYear="1842" box="[309,492,187,213]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="478" phylum="Arthropoda" rank="species" species="membranipes">
|
||
<emphasis id="14823B0BFFA4A1497719FF27FE0AFF36" box="[309,492,187,213]" italics="true" pageId="5" pageNumber="478">C. membranipes</emphasis>
|
||
</taxonomicName>
|
||
–group, more of it is typically visible in lateral view than in some species (often only the apex of the tooth is visible in this view) and the tooth is large.
|
||
</paragraph>
|
||
<paragraph id="2649E719FFA4A14976EBFE9FFB94FE6A" blockId="5.[151,1436,151,393]" pageId="5" pageNumber="478">
|
||
<taxonomicName id="E1F69C9AFFA4A14976EBFE9FFE97FEFF" authorityName="Dow" authorityYear="2020" box="[199,369,259,285]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="478" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA4A14976EBFE9FFE97FEFF" box="[199,369,259,285]" italics="true" pageId="5" pageNumber="478">Coeliccia junis</emphasis>
|
||
</taxonomicName>
|
||
is only known from two locations in Bintulu Division,
|
||
<collectingCountry id="5EE1A789FFA4A14975DEFE9FFBB3FEFE" box="[1010,1109,259,285]" name="Malaysia" pageId="5" pageNumber="478">Sarawak</collectingCountry>
|
||
, where it occurs at very low densities. At Binyo Penyilam it was found in low pH swamp forest beside a small stream;
|
||
<taxonomicName id="E1F69C9AFFA4A14972A8FEBBFAD2FEA2" authorityName="Laidlaw" authorityYear="1918" box="[1156,1332,295,321]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="478" phylum="Arthropoda" rank="species" species="nigrohamata">
|
||
<emphasis id="14823B0BFFA4A14972A8FEBBFAD2FEA2" box="[1156,1332,295,321]" italics="true" pageId="5" pageNumber="478">C. nigrohamata</emphasis>
|
||
</taxonomicName>
|
||
occurred on the stream in close proximity to the spot where
|
||
<taxonomicName id="E1F69C9AFFA4A14974F3FED7FCD1FE87" authorityName="Dow" authorityYear="2020" box="[735,823,331,357]" class="Insecta" family="Platycnemididae" genus="Coeliccia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="478" phylum="Arthropoda" rank="species" species="junis">
|
||
<emphasis id="14823B0BFFA4A14974F3FED7FCD1FE87" box="[735,823,331,357]" italics="true" pageId="5" pageNumber="478">C. junis</emphasis>
|
||
</taxonomicName>
|
||
was found. In the Bukit
|
||
<taxonomicName id="E1F69C9AFFA4A149727BFED7FA97FE86" authority="Wild Life Corridor" authorityName="Wild Life Corridor" box="[1111,1393,331,357]" class="Magnoliopsida" family="Convolvulaceae" genus="Mina" higherTaxonomySource="CoL" kingdom="Plantae" order="Solanales" pageId="5" pageNumber="478" phylum="Tracheophyta" rank="genus">Mina Wild Life Corridor</taxonomicName>
|
||
the
|
||
<typeStatus id="F94D59BBFFA4A14976BBFEF3FF1FFE6A" box="[151,249,367,393]" pageId="5" pageNumber="478" type="holotype">holotype</typeStatus>
|
||
was collected at a small pool in a section of swampy (low pH) kerangas forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |