288 lines
20 KiB
XML
288 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.67.6703" ID-PMC="PMC4972245" ID-Pensoft-Pub="1314-2003-67-1" ID-Pensoft-UUID="D846EB3F7746FFFE4A469751FFEF3B22" ID-PubMed="27536193" ID-Zenodo-Dep="133270" ModsDocID="1314-2003-67-1" checkinTime="1469054247116" checkinUser="pensoft" docAuthor="Kriebel, Ricardo" docDate="2016" docId="9BE05C4DBB54346593EA873B5E66ABAD" docLanguage="en" docName="PhytoKeys 67: 1-326" docOrigin="PhytoKeys 67" docPubDate="2016-07-20" docSource="http://dx.doi.org/10.3897/phytokeys.67.6703" docTitle="Conostegia dentata Triana" docType="treatment" docVersion="7" id="D846EB3F7746FFFE4A469751FFEF3B22" lastPageNumber="125" masterDocId="D846EB3F7746FFFE4A469751FFEF3B22" masterDocTitle="A Monograph of Conostegia (Melastomataceae, Miconieae)" masterLastPageNumber="326" masterPageNumber="1" pageNumber="124" updateTime="1668141747722" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A Monograph of Conostegia (Melastomataceae, Miconieae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kriebel, Ricardo</mods:namePart>
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<mods:affiliation>Department of Botany, University of Wisconsin-Madison, 430 Lincoln Drive Madison, Wisconsin 53706, USA</mods:affiliation>
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<mods:nameIdentifier type="email">kriebelr@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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||
<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-07-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>67</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>326</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.67.6703</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.67.6703</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-67-1</mods:identifier>
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||
<mods:identifier type="Pensoft-UUID">D846EB3F7746FFFE4A469751FFEF3B22</mods:identifier>
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<mods:identifier type="Zenodo-Dep">133270</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127858389" LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD" httpUri="http://treatment.plazi.org/id/9BE05C4DBB54346593EA873B5E66ABAD" lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124">
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<subSubSection pageId="123" pageNumber="124" type="nomenclature">
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<paragraph pageId="123" pageNumber="124">
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<taxonomicName LSID="9BE05C4D-BB54-3465-93EA-873B5E66ABAD" authority="Triana" authorityName="Triana" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia dentata" order="Myrtales" pageId="123" pageNumber="124" phylum="Tracheophyta" rank="species" species="dentata">Conostegia dentata Triana</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="123" pageNumber="124" type="reference_group">
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<paragraph pageId="123" pageNumber="124">
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia dentata" order="Myrtales" pageId="123" pageNumber="124" phylum="Tracheophyta" rank="species" species="dentata">Conostegia dentata</taxonomicName>
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Triana, Trans. Linn. Soc. London 28: 99. 1872. Type: Colombia.
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<normalizedToken originalValue="Chocó">Choco</normalizedToken>
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: J. Triana 4113 (not seen-see discussion for details on the type).
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</paragraph>
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<paragraph pageId="123" pageNumber="124">
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<taxonomicName authorityName="Gleason" authorityYear="1939" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia hispida" order="Myrtales" pageId="123" pageNumber="124" phylum="Tracheophyta" rank="species" species="hispida">Conostegia hispida</taxonomicName>
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Gleason, Bull. Torrey Bot. Club 66: 415. 1939. Type: Ecuador. Esmeraldas: Playa Rica, Parroquia de
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<normalizedToken originalValue="Concepción">Concepcion</normalizedToken>
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, 105 m, 10 December 1936, Y.
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<normalizedToken originalValue="Mexía">Mexia</normalizedToken>
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8430 (holotype: NY!, isotypes: BM, F, GH, MO!, NA, S, UC, US!).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="123" pageNumber="124" type="description">
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<paragraph pageId="123" pageNumber="124">Description.</paragraph>
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<paragraph pageId="123" pageNumber="124">
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Shrub to small tree 1.3-8 m with sub-terete stems that are densely setose with simple bristles up to 4 mm long; the nodal line hard to see and covered with setae as the rest of the node and internode. Leaves at a node equal to sub equal in size. Petiole 0.5-4.9 cm long. Leaves 11.9-35
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<normalizedToken originalValue="×">x</normalizedToken>
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5-12.9 cm, 3-5 plinerved, with the innermost pair of veins arising up to about 3 cm above the base and diverging mostly in opposite or sub opposite fashion from the mid vein, obovate to nearly elliptical, the base acute to decurrent on the petiole and with paired formicaria ca. 2-4 cm long on the leaf surface or extending to the petiole, the apex rounded to obtuse and abruptly acuminate, the margin dentate, undulate-dentate or denticulate, adaxially sparsely setose, abaxially setose. Inflorescence a terminal compact panicle 1.5-4(-7) cm long, inflorescence rachis obscured by the dense setose indument; bracts and bracteoles 1.5-4 mm. Pedicel ca. 2 mm long, obscured by the indument. Flowers (5-)6(-8) merous, calyptrate. Flower buds ca. 9-15
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<normalizedToken originalValue="×">x</normalizedToken>
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5-7 mm, narrowly ovate not constricted about the middle, rounded a the base, acute and long attenuate at the apex, not constricted in the middle, the calycine and hypanthial portions undifferentiated; the hypanthium 5-6.25
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<normalizedToken originalValue="×">x</normalizedToken>
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5.5-7.5 mm, densely hirsute with trichomes with swollen bases. Petals ca. 9-13
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<normalizedToken originalValue="×">x</normalizedToken>
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9-12 mm, white or pink, obtriangular, spreading, glabrous, the apex emarginate. Stamens 19-30, ca. 7-8 mm long, their posture at anthesis not seen, the filament 4-5 mm long, lacking a conspicuous geniculation, anther 2.8-3.5 mm, linear-oblong, recurved near the base, yellow, laterally compressed, the connective thickened and with a small bump dorsally, the pore ca. 0.1 mm, subterminal and slightly ventrally inclined. Ovary 6-10 locular, inferior, the apex glabrous and forming a collar around the style base. Style 6.5-7.5 mm long, gently curved, vertical distance from the anthers to the stigma ca. 1-2 mm, horizontal distance absent, the stigma sub capitate, ca. 1.3 mm wide. Berry ca. 7
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<normalizedToken originalValue="×">x</normalizedToken>
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7 mm when dry. Seeds not seen.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="123" pageNumber="124" type="distribution">
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<paragraph pageId="123" pageNumber="124">Distribution</paragraph>
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<paragraph pageId="123" pageNumber="124">
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(Fig.
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<figureCitation captionStart="Figure 119" captionStartId="F119" captionText="Figure 119. Distribution of Conostegia dentata." figureDoi="10.3897/phytokeys.67.6703.figure119" httpUri="https://binary.pensoft.net/fig/96477" pageId="123" pageNumber="124">119</figureCitation>
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). Ranging from Panama to Colombia and Ecuador on the Pacific coast, 0-680 m in elevation.
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</paragraph>
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<caption ID-Zenodo-Dep="973880" doi="10.3897/phytokeys.67.6703.figure119" httpUri="https://binary.pensoft.net/fig/96477" pageId="123" pageNumber="124" start="Figure 119" startId="F119">
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<paragraph pageId="123" pageNumber="124">
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<emphasis bold="true" pageId="123" pageNumber="124">Figure 119.</emphasis>
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Distribution of
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia dentata" order="Myrtales" pageId="123" pageNumber="124" phylum="Tracheophyta" rank="species" species="dentata">
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<emphasis italics="true" pageId="123" pageNumber="124">Conostegia dentata</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph pageId="123" pageNumber="124">
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia dentata" order="Myrtales" pageId="123" pageNumber="124" phylum="Tracheophyta" rank="species" species="dentata">
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<emphasis italics="true" pageId="123" pageNumber="124">Conostegia dentata</emphasis>
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</taxonomicName>
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is easily recognized because of its setose indument on stems, inflorescences and flower buds, leaves with formicaria at the base and dentate margins, and compact inflorescences. The study of flowers in herbarium specimens revealed an exserted style as is typical of species of section
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="" order="Myrtales" pageId="123" pageNumber="124" phylum="Tracheophyta" rank="section" section="Australis">
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<emphasis italics="true" pageId="123" pageNumber="124">Australis</emphasis>
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</taxonomicName>
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.
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<bibRefCitation author="Almeda, F" journalOrPublisher="Flora Mesoamericana" pageId="239" pageNumber="240" pagination="164 - 338" refId="B8" refString="Almeda, F, 2009. Melastomataceae. Flora Mesoamericana vol. 4: 164 - 338" title="Melastomataceae." volume="vol. 4" year="2009">Almeda (2009)</bibRefCitation>
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was unable to locate the type specimen
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<emphasis italics="true" pageId="123" pageNumber="124">Triana 4113</emphasis>
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of this species at BM and when checking
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<normalizedToken originalValue="Triana’s">Triana's</normalizedToken>
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herbarium list noted that that number is stated to be from Antioquia instead of
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<normalizedToken originalValue="Chocó">Choco</normalizedToken>
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, but no genus or species are stated. Almeda did locate
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<emphasis italics="true" pageId="123" pageNumber="124">Triana 4112</emphasis>
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which corresponds to
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<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia dentata" order="Myrtales" pageId="123" pageNumber="124" phylum="Tracheophyta" rank="species" species="dentata">
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<emphasis italics="true" pageId="123" pageNumber="124">Conostegia dentata</emphasis>
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</taxonomicName>
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but that specimen is from Barbacoas province instead of
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<normalizedToken originalValue="Chocó">Choco</normalizedToken>
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province where
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<emphasis italics="true" pageId="123" pageNumber="124">Triana 4113</emphasis>
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was cited to have been collected. I agree with
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<bibRefCitation author="Schnell, CE" journalOrPublisher="Harvard University" pageId="243" pageNumber="244" refId="B77" refString="Schnell, CE, 1996. The Genus Conostegia (Melastomataceae). Harvard University" title="The Genus Conostegia (Melastomataceae)" year="1996">Schnell (1996)</bibRefCitation>
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that the original description leaves little doubt on its identity namely because of the description of the setose indument, congested inflorescences and long petals. Because the setae of the fruiting hypanthia are white instead of brown as in the rest if the plant,
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<bibRefCitation author="Schnell, CE" journalOrPublisher="Harvard University" pageId="243" pageNumber="244" refId="B77" refString="Schnell, CE, 1996. The Genus Conostegia (Melastomataceae). Harvard University" title="The Genus Conostegia (Melastomataceae)" year="1996">Schnell (1996)</bibRefCitation>
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hypothesized that this color difference might function to attract dispersal agents.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124" type="materials_examined">
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<paragraph pageId="123" pageNumber="124">Specimens examined.</paragraph>
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<paragraph lastPageId="124" lastPageNumber="125" pageId="123" pageNumber="124">
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<emphasis bold="true" pageId="123" pageNumber="124">
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<collectingCountry name="Panama">PANAMA</collectingCountry>
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</emphasis>
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(fide Schnell).
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<emphasis bold="true" pageId="123" pageNumber="124">
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<normalizedToken originalValue="Darién">
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<collectingRegion country="Panama" name="Darien">Darien</collectingRegion>
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</normalizedToken>
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</emphasis>
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: Cocalita near the Colombian border on the Pacific side, Dwyer 4395 (BR, GH, UD); Atlantic slope NW
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<pageBreakToken pageId="124" pageNumber="125" start="start">of</pageBreakToken>
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Santa Fe,
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<quantity metricMagnitude="4" metricUnit="m" metricValue="1.1" unit="km" value="11.0">11 km</quantity>
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from Escuela Agricola Alto de Piedra, in valley of Rio Dos Bocas, Mori and Kallunki 3850 (
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<collectingCountry name="United States of America">US</collectingCountry>
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).
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</paragraph>
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<paragraph pageId="124" pageNumber="125">
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<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Caliche, Cuatrecasas" country="COLOMBIA" location="rio Micay" municipality="Costa del Pacifico" specimenCount="1" stateProvince="Cauca">
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<emphasis bold="true" pageId="124" pageNumber="125">
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<collectingCountry name="Colombia">COLOMBIA</collectingCountry>
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.
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<collectingRegion country="Colombia" name="Cauca">Cauca</collectingRegion>
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</emphasis>
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:
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<collectingMunicipality>
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Costa del
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<normalizedToken originalValue="Pacífico">Pacifico</normalizedToken>
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</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:1C30BC906E0358CCA8F3E328367F4ED5" country="COLOMBIA" municipality="Costa del Pacifico" name="rio Micay" stateProvince="Cauca">
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<normalizedToken originalValue="río">rio</normalizedToken>
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Micay
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</location>
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, orilla derecha, en
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<collectorName>Caliche</collectorName>
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,
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<collectorName>Cuatrecasas</collectorName>
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14190 (NY)
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</materialsCitation>
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.
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<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="de Buey, El Valle" country="US" county="Rio Mutata" location="Alto de Buey" municipality="Rio El Valle" specimenCount="1" stateProvince="Choco">
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<emphasis bold="true" pageId="124" pageNumber="125">
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<normalizedToken originalValue="Chocó">
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<collectingRegion country="Colombia" name="Choco">Choco</collectingRegion>
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</normalizedToken>
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</emphasis>
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: Mun. Nuqui, Corregimiento Termales, Quebrada Piedra Piedra,
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<normalizedToken originalValue="Acevedo-Rodríguez">Acevedo-Rodriguez</normalizedToken>
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, Callejas and Churchill 6800 (NY);
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<collectingCounty>
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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<normalizedToken originalValue="Mutatá">Mutata</normalizedToken>
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</collectingCounty>
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tributary of
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<collectingMunicipality>
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<normalizedToken originalValue="Río">Rio</normalizedToken>
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El Valle
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</collectingMunicipality>
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||
between base of
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<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:FE1B91BB86A1C98C1C9D56CD9159181F" country="US" county="Rio Mutata" municipality="Rio El Valle" name="Alto de Buey" stateProvince="Choco">
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Alto
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<collectorName>de Buey</collectorName>
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</location>
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and mouth of river, Gentry and Fallen 17474 (MO, NY).
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<emphasis bold="true" pageId="124" pageNumber="125">
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<collectorName>El Valle</collectorName>
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</emphasis>
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:
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<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:A294E4B82382BFEFDAA1CF5F9C80A304" country="US" county="Rio Mutata" municipality="Rio El Valle" name="Costa del Pacifico" stateProvince="Choco">
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Costa del
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<normalizedToken originalValue="Pacífico">Pacifico</normalizedToken>
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</location>
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,
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<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:E52C2A236949393AD82B4EB4A12E8BC0" country="US" county="Rio Mutata" municipality="Rio El Valle" name="rio Cajambre" stateProvince="Choco">
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<normalizedToken originalValue="río">rio</normalizedToken>
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Cajambre
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</location>
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, Barco, Cuatrecasas 17143 (NY,
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<collectingCountry name="United States of America">US</collectingCountry>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Silva, Cuatrecasas" country="United States of America" county="Costa del Pacifico" location="San Isidro" municipality="rio Cajambre" specimenCount="1" stateProvince="Choco">
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||
<collectingCounty>
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||
Costa del
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||
<normalizedToken originalValue="Pacífico">Pacifico</normalizedToken>
|
||
</collectingCounty>
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||
,
|
||
<collectingMunicipality>
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||
<normalizedToken originalValue="río">rio</normalizedToken>
|
||
Cajambre
|
||
</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:801A9053DB64611DE21482B6F265978E" country="United States of America" county="Costa del Pacifico" municipality="rio Cajambre" name="San Isidro" stateProvince="Choco">San Isidro</location>
|
||
,
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||
<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:503355A196174C82CBEAE14A83896E92" country="United States of America" county="Costa del Pacifico" municipality="rio Cajambre" name="Cuatrecasas" stateProvince="Choco">Cuatrecasas</location>
|
||
17280 (NY);
|
||
<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:E5E91FB4F8633B3C362D57001B1F7D22" country="United States of America" county="Costa del Pacifico" municipality="rio Cajambre" name="Costa del Pacifico" stateProvince="Choco">
|
||
Costa del
|
||
<normalizedToken originalValue="Pacífico">Pacifico</normalizedToken>
|
||
</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:9A3E908CAA5EE9B4B8B648F59B3C4503" country="United States of America" county="Costa del Pacifico" municipality="rio Cajambre" name="rio Cajambre" stateProvince="Choco">
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||
<normalizedToken originalValue="río">rio</normalizedToken>
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||
Cajambre
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||
</location>
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||
,
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||
<collectorName>Silva</collectorName>
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||
,
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||
<collectorName>Cuatrecasas</collectorName>
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||
17668 (NY)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" collectorName="Barbacoas, Ewan" country="United States of America" county="Buenaventura Bay" location="Narino" municipality="Killip" specimenCount="1" stateProvince="Colorado">
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||
<collectingRegion country="United States of America" name="Colorado">Colorado</collectingRegion>
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||
, north shore of
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||
<collectingCounty>Buenaventura Bay</collectingCounty>
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||
,
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||
<collectingMunicipality>Killip</collectingMunicipality>
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||
38772 (NY).
|
||
<emphasis bold="true" pageId="124" pageNumber="125">
|
||
<normalizedToken originalValue="Nariño">
|
||
<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:D7BF8CF24E0674E3DFC63BE87BF2BEE4" country="United States of America" county="Buenaventura Bay" municipality="Killip" name="Narino" stateProvince="Colorado">Narino</location>
|
||
</normalizedToken>
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||
</emphasis>
|
||
: Quebrada Mongon at
|
||
<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:D5CCDA27E92787012FA10C8D81D35A43" country="United States of America" county="Buenaventura Bay" municipality="Killip" name="Camp Mongon" stateProvince="Colorado">Camp Mongon</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:9BE05C4DBB54346593EA873B5E66ABAD:13038709B71F9118920136E5686F7D4E" country="United States of America" county="Buenaventura Bay" municipality="Killip" name="Rio Telembi" stateProvince="Colorado">
|
||
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
||
Telembi
|
||
</location>
|
||
above
|
||
<collectorName>Barbacoas</collectorName>
|
||
,
|
||
<collectorName>Ewan</collectorName>
|
||
16877 (NY)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="124" pageNumber="125">
|
||
<emphasis bold="true" pageId="124" pageNumber="125">
|
||
<collectingCountry name="Ecuador">ECUADOR</collectingCountry>
|
||
.
|
||
<collectingRegion country="Ecuador" name="Esmeraldas">Esmeraldas</collectingRegion>
|
||
</emphasis>
|
||
: Eloy Alfaro, Reserva
|
||
<normalizedToken originalValue="Ecológica">Ecologica</normalizedToken>
|
||
Cotacachi-Cayapas,
|
||
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
||
Santiago, Tirado 571 (MO, NY).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |