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<document ID-DOI="http://dx.doi.org/10.3897/fr.25.83034" ID-Pensoft-Pub="2193-0074-1-187" ID-Pensoft-UUID="86764B0ACD9453CD965B288014B441CD" ID-ZooBank="6402F8F152294153823FCAEA106F90A1" ModsDocID="2193-0074-25-1-187" checkinTime="1667366246929" checkinUser="pensoft" docAuthor="Spasojevic, Tamara, Broad, Gavin R. &amp; Klopfstein, Seraina" docDate="2022" docId="9BB900BF4F9E52F590BC81EF01D8A6AD" docLanguage="en" docName="FossilRecord 25(1): 187-212" docOrigin="Fossil Record 25 (1)" docPubDate="2022-06-07" docSource="http://dx.doi.org/10.3897/fr.25.83034" docTitle="Orthocentrus mortuaria Brues 1910, comb. nov." docType="treatment" docVersion="1" id="86764B0ACD9453CD965B288014B441CD" lastPageNumber="187" masterDocId="86764B0ACD9453CD965B288014B441CD" masterDocTitle="Revision of 18 ichneumonid fossil species (Hymenoptera, Ichneumonidae) highlights the need for open nomenclature in palaeontology" masterLastPageNumber="212" masterPageNumber="187" pageNumber="187" updateTime="1667366246929" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>Revision of 18 ichneumonid fossil species (Hymenoptera, Ichneumonidae) highlights the need for open nomenclature in palaeontology</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Spasojevic, Tamara</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5301-5722</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, Basel, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, Bern, Switzerland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Broad, Gavin R.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7223-5333</mods:nameIdentifier>
<mods:affiliation>Natural History Museum, Cromwell Road, London SW 7 6 BD, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Klopfstein, Seraina</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4025-975X</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum Basel, Augustinergasse 2, Basel, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, Baltzerstrasse 6, Bern, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">seraina.klopfstein@bs.ch</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Fossil Record</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-07</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>25</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>187</mods:start>
<mods:end>212</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/fr.25.83034</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/fr.25.83034</mods:identifier>
<mods:identifier type="Pensoft-Pub">2193-0074-1-187</mods:identifier>
<mods:identifier type="ZooBank">6402F8F152294153823FCAEA106F90A1</mods:identifier>
<mods:identifier type="Pensoft-UUID">86764B0ACD9453CD965B288014B441CD</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:9BB900BF4F9E52F590BC81EF01D8A6AD" httpUri="http://treatment.plazi.org/id/9BB900BF4F9E52F590BC81EF01D8A6AD" lastPageNumber="187" pageId="0" pageNumber="187">
<subSubSection pageId="0" pageNumber="187" type="nomenclature">
<paragraph pageId="0" pageNumber="187">
<taxonomicName LSID="9BB900BF-4F9E-52F5-90BC-81EF01D8A6AD" authority="Brues, 1910" authorityName="Brues" authorityYear="1910" baseAuthorityName="Spasojevic &amp; Broad &amp; Klopfstein" baseAuthorityYear="1910" class="Insecta" family="Ichneumonidae" genus="Orthocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthocentrus mortuaria" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="mortuaria" status="comb. nov.">Orthocentrus? mortuaria Brues, 1910</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="187">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="187" type="description">
<paragraph pageId="0" pageNumber="187">
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Orthocentrus? mortuaria (Brues, 1910) comb. nov. A. Photograph of the holotype obtained from the MCZ; B. Our interpretative drawing of the fossil. Thicker lines indicate outlines of body structures, thinner lines show characters inside these outlines, and dotted lines represent uncertain interpretations." figureDoi="10.3897/fr.25.83034.figure8" httpUri="https://binary.pensoft.net/fig/693067" pageId="0" pageNumber="187">Fig. 8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="187" type="reference_group">
<paragraph pageId="0" pageNumber="187">
<taxonomicName authorityName="Spasojevic &amp; Broad &amp; Klopfstein" authorityYear="2022" class="Insecta" family="Ichneumonidae" genus="Polysphincta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polysphincta mortuaria" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="mortuaria" status="comb. nov.">* Polysphincta mortuaria</taxonomicName>
Brues, 1910
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="187" type="materials_examined">
<paragraph pageId="0" pageNumber="187">Material.</paragraph>
<paragraph pageId="0" pageNumber="187">
Photographs of the
<typeStatus>holotype</typeStatus>
(part, #PALE-2134) obtained from the MCZ.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="187" type="stratum">
<paragraph pageId="0" pageNumber="187">Stratum.</paragraph>
<paragraph pageId="0" pageNumber="187">Teller County, Florissant shales, Colorado, USA. Late Eocene (Chadronian), 37.2-33.9 Ma.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="187" type="description">
<paragraph pageId="0" pageNumber="187">Description.</paragraph>
<paragraph pageId="0" pageNumber="187">
<emphasis bold="true" pageId="0" pageNumber="187">
Female.
<emphasis italics="true" pageId="0" pageNumber="187">Holotype</emphasis>
</emphasis>
in lateral view, with head and a few segments of antennae, mesosoma, fore wings, and metasoma with ovipositor, all rather poorly preserved; legs and hind wings missing. Body length 6.3 mm.
</paragraph>
<paragraph pageId="0" pageNumber="187">Head light brown, mesosoma black with light brown portions, metasoma mostly light brown on T1-T3, remainder dark brown (the light brown colouration could actually be a preservation artefact: it shows different sculpture and its texture is repeated on the rock outside the body).</paragraph>
<paragraph pageId="0" pageNumber="187">
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Head</emphasis>
with face bulging below antennal sockets, with eye very small, with very long malar space.
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Antennae</emphasis>
more than 3.9 mm, incompletely preserved.
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Mesosoma</emphasis>
rather short, poorly preserved, unclear if with epicnemial carina. Propodeum with pleural carina and lateral longitudinal carina, maybe also with posterior transverse carina.
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Fore wing</emphasis>
4.1 mm, rather short and stout, with areolet open and vein 2Rs longer than 3M; vein 1cu-a strongly inclivous meeting M + Cu clearly distally of 1M; cell 2R1 only 2.9
<normalizedToken originalValue="×">x</normalizedToken>
longer than wide.
<emphasis bold="true" italics="true" pageId="0" pageNumber="187">Metasoma</emphasis>
compressed from about fourth segment; T1 short and tapering towards base, with latero-median carinae converging on basal half, parallel on apical half; T2 and following tergites transverse. Ovipositor 0.5 mm.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="187" type="interpretation">
<paragraph pageId="0" pageNumber="187">Interpretation.</paragraph>
<paragraph pageId="0" pageNumber="187">
The bulging face with small eyes and long malar space are only found in
<taxonomicName lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Metopiinae">Metopiinae</taxonomicName>
and
<taxonomicName lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Orthocentrinae">Orthocentrinae</taxonomicName>
, and wing venation and ovipositor shape point to the latter. Given the size, 1cu-a meeting M + Cu clearly distally of 1M, and possible presence of the lateral portion of the epicnemial carina,
<taxonomicName class="Insecta" family="Ichneumonidae" genus="Orthocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthocentrus" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="187">Orthocentrus</emphasis>
</taxonomicName>
is the best guess, but the poor preservation of the fossil precludes a certain placement.
</paragraph>
<caption doi="10.3897/fr.25.83034.figure8" httpUri="https://binary.pensoft.net/fig/693067" pageId="0" pageNumber="187" start="Figure 8" startId="F8">
<paragraph pageId="0" pageNumber="187">
<emphasis bold="true" pageId="0" pageNumber="187">Figure 8.</emphasis>
<taxonomicName authorityName="Brues" authorityYear="1910" baseAuthorityName="Spasojevic &amp; Broad &amp; Klopfstein" baseAuthorityYear="1910" class="Insecta" family="Ichneumonidae" genus="Orthocentrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthocentrus mortuaria" order="Hymenoptera" pageId="0" pageNumber="187" phylum="Arthropoda" rank="species" species="mortuaria">Orthocentrus? mortuaria</taxonomicName>
(Brues, 1910) comb. nov.
<emphasis bold="true" pageId="0" pageNumber="187">A.</emphasis>
Photograph of the holotype obtained from the MCZ;
<emphasis bold="true" pageId="0" pageNumber="187">B.</emphasis>
Our interpretative drawing of the fossil. Thicker lines indicate outlines of body structures, thinner lines show characters inside these outlines, and dotted lines represent uncertain interpretations.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="187">
<emphasis bold="true" pageId="0" pageNumber="187">
<taxonomicName baseAuthorityName="Forster" baseAuthorityYear="1869" lsidName="" pageId="0" pageNumber="187" rank="subFamily" subFamily="Phygadeuontinae">Phygadeuontinae</taxonomicName>
<normalizedToken originalValue="Förster">Foerster</normalizedToken>
, 1869 (
<emphasis italics="true" pageId="0" pageNumber="187">sensu</emphasis>
Santos (2017)
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>