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<document id="054402ABAB630B2A1A0CB900DF4C68EB" ID-DOI="10.11646/zootaxa.1448.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5077273" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625629497592" checkinUser="felipe" docAuthor="De Moraes, Gilberto J., Zannou, Ignace D., Ueckermann, Eddie A., Oliveira, Anibal R., Hanna, Rachid &amp; Yaninek, John S." docDate="2007" docId="832E87FA1516EE5DE7EDFD00CCBFA262" docLanguage="en" docName="zt01448p039.pdf" docOrigin="Zootaxa 1448 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Proprioseiopsis bremaensis Moraes, Zannou &amp; Oliveira 2007, n. sp." docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="7F17FF82151AEE50E77AFF9FCD7FA640" masterDocTitle="Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa" masterLastPageNumber="39" masterPageNumber="1" pageNumber="13" updateTime="1698993771546" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="EA4B1E789F864ECC5E25033E2ABB7446" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="66EA0B3338F600E23131AC56DC20D4EE">
<mods:title id="F59E1253FA32058118797B1EF1964B42">Species of the subtribes Arrenoseiina and Proprioseiopsina (Tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa</mods:title>
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</mods:role>
<mods:namePart id="56AC4D54B4BE260E00A304039F5FF609">De Moraes, Gilberto J.</mods:namePart>
<mods:affiliation id="AA54CAF1640C07029D3263E2F1D390B6">Depto. Entomologia, Fitopatologia e Zoologia Agrícola, ESALQ-Universidade de São Paulo- 13418 - 900 Piracicaba-SP, Brazil. E-mail: gjmoraes @ carpa. ciagri. usp. br Plant Protection Research Institute, Private Bag X 134, Queenwood, Pretoria; 0121 South Africa Biological Control Centre for Africa, International Institute of Tropical Agriculture, 08 B. P. 0932, Cotonou, Benin, West Africa Dept. Entomology, Smith Hall Room 100, 901 W State Street, Purdue University, West Lafayette, IN 47907 - 2089; USA</mods:affiliation>
</mods:name>
<mods:name id="2389F6108A937FD70FF29EEBBAAB536D" type="personal">
<mods:role id="B010CF160BE7D53F4325B58F2A04E92B">
<mods:roleTerm id="C6F1172989EA59F3596DDB7353855DEA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8465181039A55B36B347625C51A6CE98">Zannou, Ignace D.</mods:namePart>
<mods:affiliation id="7391FBC0ADF1B8C547D0F8FC211B79CE">Depto. Entomologia, Fitopatologia e Zoologia Agrícola, ESALQ-Universidade de São Paulo- 13418 - 900 Piracicaba-SP, Brazil. E-mail: gjmoraes @ carpa. ciagri. usp. br Plant Protection Research Institute, Private Bag X 134, Queenwood, Pretoria; 0121 South Africa Biological Control Centre for Africa, International Institute of Tropical Agriculture, 08 B. P. 0932, Cotonou, Benin, West Africa Dept. Entomology, Smith Hall Room 100, 901 W State Street, Purdue University, West Lafayette, IN 47907 - 2089; USA</mods:affiliation>
</mods:name>
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<mods:role id="166EA5897DCE8A45F050F051B03E3047">
<mods:roleTerm id="09AEE7BCC3F1EF29D6F9B95B0F0B9B49">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3B2A95BF135D2897A210D4F0E2C91E1">Ueckermann, Eddie A.</mods:namePart>
</mods:name>
<mods:name id="39062BA51E0A67E09A3A7F646E0D7D85" type="personal">
<mods:role id="F8369466DAC84B90B1EC776902727C91">
<mods:roleTerm id="DC06957ED524439CE73C0988FE0628F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A4B44B0FF553D154AE3F2A13C4E5AE39">Oliveira, Anibal R.</mods:namePart>
<mods:affiliation id="F8181580A0A875C9EFD21060B77F1BC7">Depto. Entomologia, Fitopatologia e Zoologia Agrícola, ESALQ-Universidade de São Paulo- 13418 - 900 Piracicaba-SP, Brazil. E-mail: gjmoraes @ carpa. ciagri. usp. br Plant Protection Research Institute, Private Bag X 134, Queenwood, Pretoria; 0121 South Africa Biological Control Centre for Africa, International Institute of Tropical Agriculture, 08 B. P. 0932, Cotonou, Benin, West Africa Dept. Entomology, Smith Hall Room 100, 901 W State Street, Purdue University, West Lafayette, IN 47907 - 2089; USA</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="862A05736AD3DF7F9F7D95B8C114D976">Hanna, Rachid</mods:namePart>
</mods:name>
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<mods:role id="3D5BC382166F6A5A1F33697DD2F3F17C">
<mods:roleTerm id="E896BF4FC8B4DE964A17320E6F034F73">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F54CFF0D3CB006C74977D8B82DA0B2A9">Yaninek, John S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="3213BE5C9382C504318D592B9477FE9A">text</mods:typeOfResource>
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<mods:title id="517B29652488B9716450FB5D86948456">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D32875DA7C6F8BA9C90308BFAC2F02B7">
<mods:date id="9FFA9A5F7602BDC257F5DEB7C533F674">2007</mods:date>
<mods:detail id="AC15944F65AF27DC2442D4B16B9E8A12" type="pubDate">
<mods:number id="12AD595FA8DB7A6E859F24BBADBB9489">2007-04-16</mods:number>
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<mods:number id="4DE828735A244E4756A98299328ED8E3">1448</mods:number>
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<mods:number id="5BD8E4EB0780BBE147A7065BB3F01B95">1</mods:number>
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<mods:start id="78ADD2F19D3B5B90672C49F29F45DBD1">1</mods:start>
<mods:end id="17E464077DA710A79FB6481065FD519F">39</mods:end>
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<mods:location id="86FF3F4709BF8A2E4C01BAA094DD4E42">
<mods:url id="0C0380E7EC4D7848C6F214710D2B1518">https://biotaxa.org/Zootaxa/article/view/zootaxa.1448.1.1</mods:url>
</mods:location>
<mods:classification id="A1654827D35C8D9D47D2D5C9E15C7968">journal article</mods:classification>
<mods:identifier id="637500D34B1008A2E09AFC76489E7B53" type="DOI">10.11646/zootaxa.1448.1.1</mods:identifier>
<mods:identifier id="7E3CDE44ED652813DB497C2957AF6903" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="85D72A0E48E1ABC7B6FB0381A8A16458" type="Zenodo-Dep">5077273</mods:identifier>
<mods:identifier id="9AA84EF64997D329B0A7BC6DD89B4172" type="ZooBank">10385A17-BFD2-4460-9CAC-2B11114DC576</mods:identifier>
</mods:mods>
<treatment id="832E87FA1516EE5DE7EDFD00CCBFA262" LSID="urn:lsid:plazi:treatment:832E87FA1516EE5DE7EDFD00CCBFA262" httpUri="http://treatment.plazi.org/id/832E87FA1516EE5DE7EDFD00CCBFA262" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">
<subSubSection id="439D65671516EE5CE7EDFD00CEF2A4F9" box="[151,909,671,697]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="0B3836EC1516EE5CE7EDFD00CEF2A4F9" blockId="12.[151,909,671,732]" box="[151,909,671,697]" pageId="12" pageNumber="13">
<heading id="507081801516EE5CE7EDFD00CEF2A4F9" bold="true" box="[151,909,671,697]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="39F3EAFE1516EE5CE7EDFD00CEF2A4F9" bold="true" box="[151,909,671,697]" pageId="12" pageNumber="13">
<taxonomicName id="CC874D6F1516EE5CE7EDFD00CE3CA4F9" ID-CoL="77XTX" authority="Moraes, Zannou &amp; Oliveira" authorityName="Moraes, Zannou &amp; Oliveira" authorityYear="2007" box="[151,835,671,697]" class="Arachnida" family="Phytoseiidae" genus="Proprioseiopsis" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bremaensis" status="sp. nov.">
<emphasis id="39F3EAFE1516EE5CE7EDFD00CC9EA4F9" bold="true" box="[151,481,671,697]" italics="true" pageId="12" pageNumber="13">Proprioseiopsis bremaensis</emphasis>
Moraes, Zannou &amp; Oliveira
</taxonomicName>
<taxonomicNameLabel id="22C057851516EE5CE431FD00CEF2A4F9" box="[843,909,671,697]" pageId="12" pageNumber="13" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="439D65671516EE5DE7EDFD5DCCBFA262" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13" type="description">
<paragraph id="0B3836EC1516EE5CE7EDFD5DCD90A49C" blockId="12.[151,909,671,732]" box="[151,239,706,732]" pageId="12" pageNumber="13">
(
<figureCitation id="93BC2A691516EE5CE7E5FD5DCD9AA49C" box="[159,229,706,732]" captionStart="FIGURE 8" captionStartId="12.[151,255,1978,2002]" captionTargetBox="[167,1420,1042,1922]" captionTargetId="figure-246@12.[151,1436,1029,1955]" captionTargetPageId="12" captionText="FIGURE 8. Proprioseiopsis bremaensis Moraes, Zannou &amp; Oliveira n. sp. (female): A. Dorsal shield; B. Ventral sur-" figureDoi="http://doi.org/10.5281/zenodo.5077291" httpUri="https://zenodo.org/record/5077291/files/figure.png" pageId="12" pageNumber="13">Fig. 8</figureCitation>
)
</paragraph>
<paragraph id="0B3836EC1516EE5CE7EDFC8DCE66A58C" blockId="12.[151,1436,786,972]" pageId="12" pageNumber="13">Diagnosis. Species characterized by having dorsal shield mostly smooth, with few anterolateral striae; seta Z5 slightly shorter than setae z4, Z4 and s4; setae z4, Z4, Z5, s4 and Jv5 serrate; z2 and z4 about as long as distances between their bases and the bases of the following setae; sternal shield with few lateral striae; ventrianal shield subtriangular, reticulate, about as long as wide at level of Zv2; fixed cheliceral digit with few teeth (apparently 3 teeth); calyx of spermatheca cup-shaped.</paragraph>
<caption id="5FF866641516EE5CE7EDF825C8E4A192" ID-DOI="http://doi.org/10.5281/zenodo.5077291" ID-Zenodo-Dep="5077291" box="[151,1435,1978,2002]" httpUri="https://zenodo.org/record/5077291/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,255,1978,2002]" targetBox="[167,1420,1042,1922]" targetPageId="12">
<paragraph id="0B3836EC1516EE5CE7EDF825C8E4A192" blockId="12.[151,1435,1978,2002]" box="[151,1435,1978,2002]" pageId="12" pageNumber="13">
<emphasis id="39F3EAFE1516EE5CE7EDF825CC65A192" bold="true" box="[151,282,1978,2002]" pageId="12" pageNumber="13">FIGURE 8.</emphasis>
<taxonomicName id="CC874D6F1516EE5CE659F824CEFDA192" authority="Moraes, Zannou &amp; Oliveira" authorityName="Moraes, Zannou &amp; Oliveira" authorityYear="2007" box="[291,898,1979,2002]" class="Arachnida" family="Phytoseiidae" genus="Proprioseiopsis" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="bremaensis" status="sp. nov.">
<emphasis id="39F3EAFE1516EE5CE659F824CF34A192" box="[291,587,1979,2002]" italics="true" pageId="12" pageNumber="13">Proprioseiopsis bremaensis</emphasis>
Moraes, Zannou &amp; Oliveira
</taxonomicName>
<taxonomicNameLabel id="22C057851516EE5CE4F0F825CEB8A192" box="[906,967,1978,2002]" pageId="12" pageNumber="13" rank="species">
<emphasis id="39F3EAFE1516EE5CE4F0F825CEBEA192" bold="true" box="[906,961,1978,2002]" pageId="12" pageNumber="13">n. sp</emphasis>
.
</taxonomicNameLabel>
(female): A. Dorsal shield; B. Ventral sur-
</paragraph>
</caption>
<paragraph id="0B3836EC1517EE5DE7EDFF06CC86A6F0" blockId="13.[151,1436,153,1058]" box="[151,505,153,176]" pageId="13" pageNumber="14">face; C. Spermatheca; D. Leg IV.</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFF5FCF3EA69A" blockId="13.[151,1436,153,1058]" box="[197,577,192,218]" pageId="13" pageNumber="14">
Female. (
<specimenCount id="1D81FD651517EE5DE64FFF5FCCC7A69A" box="[309,440,192,218]" pageId="13" pageNumber="14" type="generic">1 specimen</specimenCount>
measured).
</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFF77CFFEA712" blockId="13.[151,1436,153,1058]" pageId="13" pageNumber="14">
<materialsCitation id="BBEF3CB11517EE5DE7BFFF77CF02A712" collectionCode="R" latitude="-5.233333" pageId="13" pageNumber="14" specimenCode="R1" specimenCount="1">
Dorsum. Dorsal shield smooth, with few anterolateral striae, 341 long and 240 wide. Setae j1 24, j3 41, j4 5, j5 5, j6 5, J5 10, z2 29, z4 70, z5 5, Z1 17, Z4 74, Z5 62, s4 74,
<geoCoordinate id="6EB3502B1517EE5DE4E1FE8FCE9DA76A" box="[923,994,272,298]" degrees="2" direction="south" minutes="34" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="-2.5666666">S2 34</geoCoordinate>
,
<geoCoordinate id="6EB3502B1517EE5DE497FE8FC94BA76A" box="[1005,1076,272,298]" degrees="4" direction="south" minutes="24" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="-4.4">S4 24</geoCoordinate>
,
<geoCoordinate id="6EB3502B1517EE5DE345FE8FC9FAA76A" box="[1087,1157,272,298]" degrees="5" direction="south" minutes="14" orientation="latitude" pageId="13" pageNumber="14" precision="925" value="-5.233333">S5 14</geoCoordinate>
, r3 17,
<specimenCode id="5B219E971517EE5DE3A7FE8FC981A76A" box="[1245,1278,272,298]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="14">R1</specimenCode>
12. Setae z4, Z4, Z5 and s4 serrate, other setae smooth
</materialsCitation>
.
</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFEFFC905A7B2" blockId="13.[151,1436,153,1058]" pageId="13" pageNumber="14">Venter. Sternal shield mostly smooth, few anterior and lateral striae; posterior margin concave; distances between St1-St3 50, St2-St2 67. Genital shield smooth; distance between St5-St5 91. Ventrianal shield subtriangular, reticulate, anterior margin very slightly concave, 118 long, 127 wide at level of Zv2, elliptical preanal pores mesad and slightly posterior to Jv2; Jv5 serrate and other ventral setae smooth.</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFD9FCD8BA402" blockId="13.[151,1436,153,1058]" pageId="13" pageNumber="14">
Chelicera. Movable digit 30 long, with 1 tooth; fixed digit 27 long, apparently with 3 teeth and a
<emphasis id="39F3EAFE1517EE5DE219FD9FCD92A401" italics="true" pageId="13" pageNumber="14">pilus dentilis</emphasis>
.
</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFDCFC9A2A42A" blockId="13.[151,1436,153,1058]" box="[197,1245,592,618]" pageId="13" pageNumber="14">
Spermatheca. Calyx cup-shaped 7 long,
<quantity id="CC7F9B091517EE5DE5E7FDCFCFA6A42A" box="[669,729,592,618]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="13" pageNumber="14" unit="in" value="10.0">10 in</quantity>
diameter at its distal extreme; atrium small.
</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFDE7CCC9A4FA" blockId="13.[151,1436,153,1058]" pageId="13" pageNumber="14">Legs. Sharp-tipped macrosetae; Sge III 19, Sge IV 34, Sti IV 19, St IV 65. Chaetotaxy: genu II: 2–2/0,2/0– 1; genu III: 1–2/1,2/0–1.</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFD57CCF4A4A2" blockId="13.[151,1436,153,1058]" box="[197,395,712,738]" pageId="13" pageNumber="14">Male. Unknown.</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFD6FCFA7A572" blockId="13.[151,1436,153,1058]" pageId="13" pageNumber="14">
<materialsCitation id="BBEF3CB11517EE5DE7BFFD6FCFA7A572" collectingDate="1990-04-05" collectionCode="ESALQ-USP" collectorName="B. Haruna &amp; K. Antwi" country="Ghana" location="Locality" pageId="13" pageNumber="14" specimenCount="1" typeStatus="holotype">
<location id="0E5860371517EE5DE7BFFD6FCC58A54A" LSID="urn:lsid:plazi:treatment:832E87FA1516EE5DE7EDFD00CCBFA262:0E5860371517EE5DE7BFFD6FCC58A54A" box="[197,295,752,778]" country="Ghana" name="Locality" pageId="13" pageNumber="14">Locality</location>
and type material.
<typeStatus id="D43C884E1517EE5DE577FD6FCF05A54A" box="[525,634,752,778]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
female from
<taxonomicName id="CC874D6F1517EE5DE466FD6FCEC1A54A" box="[796,958,752,778]" class="Magnoliopsida" family="Solanaceae" genus="Capsicum" kingdom="Plantae" order="Solanales" pageId="13" pageNumber="14" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="39F3EAFE1517EE5DE466FD6FCEEEA549" box="[796,913,752,777]" italics="true" pageId="13" pageNumber="14">Capsicum</emphasis>
sp.
</taxonomicName>
,
<locationDeviation id="1A8A2FCE1517EE5DE4B6FD6FC9FFA54A" box="[972,1152,752,778]" location="Brema" pageId="13" pageNumber="14">
<quantity id="CC7F9B091517EE5DE4B6FD6FC977A54A" box="[972,1032,752,778]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="14" unit="km" value="3.0">3 km</quantity>
N Brema
</locationDeviation>
,
<collectingCountry id="7390767C1517EE5DE3F7FD6FC9A3A54A" box="[1165,1244,752,778]" name="Ghana" pageId="13" pageNumber="14">Ghana</collectingCountry>
,
<date id="7F39102C1517EE5DE393FD6FC80BA54A" box="[1257,1396,752,778]" pageId="13" pageNumber="14" value="1990-04-05">
<collectingDate id="6F7DE9C41517EE5DE393FD6FC80BA54A" box="[1257,1396,752,778]" pageId="13" pageNumber="14" value="1990-04-05">05-IV-1990</collectingDate>
</date>
,
<collectorName id="A672533A1517EE5DE2FBFD6FCD92A572" pageId="13" pageNumber="14">B. Haruna</collectorName>
and
<collectorName id="A672533A1517EE5DE65CFC87CCE9A572" box="[294,406,792,818]" pageId="13" pageNumber="14">K. Antwi</collectorName>
, deposited at
<collectionCode id="6D96AE291517EE5DE54FFC87CFABA572" box="[565,724,792,818]" pageId="13" pageNumber="14">ESALQ-USP</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFCDFC8E3A51A" blockId="13.[151,1436,153,1058]" box="[197,1436,832,858]" pageId="13" pageNumber="14">
Etymology. The epithet
<taxonomicName id="CC874D6F1517EE5DE6A4FCDFCF1DA519" authorityName="Moraes, Zannou &amp; Oliveira" authorityYear="2007" box="[478,610,832,857]" class="Arachnida" family="Phytoseiidae" genus="Proprioseiopsis" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bremaensis">
<emphasis id="39F3EAFE1517EE5DE6A4FCDFCF1DA519" box="[478,610,832,857]" italics="true" pageId="13" pageNumber="14">bremaensis</emphasis>
</taxonomicName>
refers to the location where the
<typeStatus id="D43C884E1517EE5DE4A3FCDFC973A51A" box="[985,1036,832,858]" pageId="13" pageNumber="14">type</typeStatus>
of this new species was collected.
</paragraph>
<paragraph id="0B3836EC1517EE5DE7BFFCF7CCBFA262" blockId="13.[151,1436,153,1058]" pageId="13" pageNumber="14">
Remarks.
<taxonomicName id="CC874D6F1517EE5DE647FCF7CF00A5C1" baseAuthorityName="Abbasova" baseAuthorityYear="1970" box="[317,639,872,897]" class="Arachnida" family="Phytoseiidae" genus="Proprioseiopsis" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bregetovae">
<emphasis id="39F3EAFE1517EE5DE647FCF7CF00A5C1" box="[317,639,872,897]" italics="true" pageId="13" pageNumber="14">Proprioseiopsis bregetovae</emphasis>
</taxonomicName>
differs from this new species by having the posterior region of the dorsal shield reticulate and because its z4 and Jv5 are smooth.
<taxonomicName id="CC874D6F1517EE5DE4FBFC0FC9ACA5E9" baseAuthorityName="Chant" baseAuthorityYear="1957" box="[897,1235,912,937]" class="Arachnida" family="Phytoseiidae" genus="Proprioseiopsis" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="okanagensis">
<emphasis id="39F3EAFE1517EE5DE4FBFC0FC9ACA5E9" box="[897,1235,912,937]" italics="true" pageId="13" pageNumber="14">Proprioseiopsis okanagensis</emphasis>
</taxonomicName>
differs from this new species by having Z5 longer than z4, Z4 and s4; in addition, it has all dorsal setae as well as Jv5 smooth, and fixed cheliceral digit multidentate. Both of those species differ from the new species by having calyx of spermatheca bell-shaped.
</paragraph>
</subSubSection>
</treatment>
</document>
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