treatments-xml/data/76/47/F4/7647F4658334DB5121F2BB764377FC8E.xml

<document id="D1B066BB20A08AE1DF23D06DEAA02691" ID-DOI="10.11646/zootaxa.4758.1.9" ID-GBIF-Dataset="ef209b80-8915-4fe9-a46b-c0ef681bdba0" ID-ISSN="1175-5326" ID-Zenodo-Dep="3730773" ID-ZooBank="A25C4257-7242-45F2-AE27-3CAB1436AB3A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585304271945" checkinUser="plazi" docAuthor="Mielke, Carlos G. C., Grehan, John R. &amp; Cock, Matthew J. W." docDate="2020" docId="7647F4658334DB5121F2BB764377FC8E" docLanguage="en" docName="zootaxa.4758.1.9.pdf.imf" docOrigin="Zootaxa 4758 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Wallacella Mielke &amp; Grehan &amp; Cock 2020, gen. nov." docType="treatment" docVersion="7" lastPageNumber="184" masterDocId="8A7E8C1D8335DB522165BE4B4106FF99" masterDocTitle="Ghost-moths of Trinidad and Tobago with description of a new genus and a new species (Lepidoptera: Hepialidae)" masterLastPageNumber="190" masterPageNumber="181" pageNumber="182" updateTime="1698732493479" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D11D53426CFCDD9815D2AF6BF2D630A4">Ghost-moths of Trinidad and Tobago with description of a new genus and a new species (Lepidoptera: Hepialidae)</mods:title>
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<mods:namePart id="CD6321C9D289198190FD99DB379DAC13">Mielke, Carlos G. C.</mods:namePart>
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<mods:namePart id="5F4BDAAE9ED6914ADFE386D9BD2CD87F">Grehan, John R.</mods:namePart>
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<mods:namePart id="6B412CD17D6FD6EDC94B7921A1718C3E">Cock, Matthew J. W.</mods:namePart>
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<mods:date id="B32E7CB09E3072ADB2DF7F26D1143D8E">2020</mods:date>
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<treatment id="7647F4658334DB5121F2BB764377FC8E" ID-DOI="http://doi.org/10.5281/zenodo.3812899" ID-GBIF-Taxon="163300081" ID-Zenodo-Dep="3812899" LSID="urn:lsid:plazi:treatment:7647F4658334DB5121F2BB764377FC8E" httpUri="http://treatment.plazi.org/id/7647F4658334DB5121F2BB764377FC8E" lastPageId="3" lastPageNumber="184" pageId="1" pageNumber="182">
<subSubSection id="B6F416F88334DB5321F2BB764094FACE" box="[151,402,1341,1367]" pageId="1" pageNumber="182" type="nomenclature">
<paragraph id="FE5145738334DB5321F2BB764094FACE" blockId="1.[151,808,1341,1403]" box="[151,402,1341,1367]" pageId="1" pageNumber="182">
<heading id="A519F21F8334DB5321F2BB764094FACE" bold="true" box="[151,402,1341,1367]" fontSize="11" level="1" pageId="1" pageNumber="182" reason="1">
<emphasis id="CC9A99618334DB5321F2BB764094FACE" bold="true" box="[151,402,1341,1367]" pageId="1" pageNumber="182">
<taxonomicName id="39EE3EF08334DB5321F2BB764010FACE" ID-CoL="9FSH8" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[151,278,1341,1367]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="182" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618334DB5321F2BB764010FACE" bold="true" box="[151,278,1341,1367]" italics="true" pageId="1" pageNumber="182">Wallacella</emphasis>
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, 
<taxonomicNameLabel id="D7A9241A8334DB532041BB764094FACE" box="[292,402,1341,1367]" pageId="1" pageNumber="182" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="B6F416F88334DB5321F2BB2A450EFA27" pageId="1" pageNumber="182" type="description">
<paragraph id="FE5145738334DB5321F2BB2A422EFAE2" blockId="1.[151,808,1341,1403]" box="[151,808,1376,1403]" pageId="1" pageNumber="182">
(
<figureCitation id="66D559F68334DB5321FABB2A40ACFAE2" box="[159,426,1376,1403]" captionStart="FIGURES 1−6" captionStartId="2.[151,264,1458,1483]" captionTargetBox="[154,1433,489,1433]" captionTargetId="figure@2.[154,1433,489,1433]" captionTargetPageId="2" captionText="FIGURES 1−6. Wallacella guianensis, comb. nov. 1−4, Male: 1a, dorsal view; 1b, ventral view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.217; MJWC); 2, dorsal view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.356; MJWC; lines: i—proximal antemedial, ii—distal antemedial, iii—proximal postmedial, iv—premarginal); 3, dorsal view (French Guiana; CGCM 33.107); 4, dorsal view, holotype of Phassus guianensis (USNM). 5, Female: dorsal view (MHNH). 6, Male wing venation (CGCM 30.158). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3730775" httpUri="https://zenodo.org/record/3730775/files/figure.png" pageId="1" pageNumber="182">Figs 1a, 1b, 2, 3, 4, 5, 6</figureCitation>
, 
<figureCitation id="66D559F68334DB5320D3BB2A4219FAE2" box="[438,799,1376,1403]" captionStart="FIGURES 10−15" captionStartId="5.[151,264,1886,1911]" captionTargetBox="[161,1424,640,1859]" captionTargetId="figure@5.[151,1436,633,1861]" captionTargetPageId="5" captionText="FIGURES 10−15. Wallacella guianensis, comb. nov. 10, Legs (CGCM 30.158). 11−12, Male abdomen (CGCM 35.217): 11, upper dorsal, lower ventral view; 12, tergum and sternum VII and VIII. 13, Male genitalia (CGCM 35.217): 13a, lateral view; 13b, anterior view; 13c, ventral view (pseudotegumen upwards). 14, Male abdomen (holotype), tergite and sternite VII and VIII. 15, Male genitalia (holotype), ventral view (unscaled). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3730779" httpUri="https://zenodo.org/record/3730779/files/figure.png" pageId="1" pageNumber="182">10, 11, 12, 13a, 13b, 13c, 14, 15</figureCitation>
)
</paragraph>
<paragraph id="FE5145738334DB5321F2BBED450EFA27" blockId="1.[151,1032,1445,1470]" box="[151,1032,1445,1470]" pageId="1" pageNumber="182">
<typeStatus id="2155FBD18334DB5321F2BBED41CFFA27" box="[151,201,1446,1470]" pageId="1" pageNumber="182">Type</typeStatus>
species: 
<taxonomicName id="39EE3EF08334DB532043BBEE437AFA24" authority="Schaus, 1940" authorityName="Schaus" authorityYear="1940" box="[294,636,1445,1470]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="182" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis id="CC9A99618334DB532043BBEE40EAFA24" box="[294,492,1445,1469]" italics="true" pageId="1" pageNumber="182">Phassus guianensis</emphasis>
<bibRefCitation id="9A7F38828334DB532097BBEE437AFA24" author="Schaus, W." box="[498,636,1445,1470]" pageId="1" pageNumber="182" pagination="83 - 88" refId="ref6293" refString="Schaus, W. (1940) New species of British Guiana Heterocera. Zoologica, 25, 83 - 88." type="journal article" year="1940">Schaus, 1940</bibRefCitation>
</taxonomicName>
, by present designation and monotypy.
</paragraph>
</subSubSection>
<subSubSection id="B6F416F88334DB5321F2BBA041DAF90F" pageId="1" pageNumber="182" type="diagnosis">
<paragraph id="FE5145738334DB5321F2BBA041DAF90F" blockId="1.[151,1436,1515,2010]" pageId="1" pageNumber="182">
<emphasis id="CC9A99618334DB5321F2BBA0400CF99F" bold="true" box="[151,266,1515,1542]" pageId="1" pageNumber="182">Diagnosis</emphasis>
. Recognized by the distinctive and unique shape of the posterior male abdomen (asymmetrical tergite VIII) and genitalia (tergal lobes asymmetrical and pocket-like ventral projection of the saccus). In addition by the following combination of characters: i) male labial palpus two-segmented, ii) antennal rami long and flattened, ii) ‘hepialine’ venation (
<bibRefCitation id="9A7F38828334DB5320ECB8134348F9EB" author="Dumbleton, L. J." box="[393,590,1623,1650]" pageId="1" pageNumber="182" pagination="920 - 981" refId="ref5499" refString="Dumbleton, L. J. (1966) Genitalia, classification and zoogeography of the New Zealand Hepialidae (Lepidoptera). New Zealand Journal of Science, 9 (4), 920 - 981." type="journal article" year="1966">Dumbleton 1966</bibRefCitation>
), and iv) the pseudotegumen dorso-mesally unfused while ventro-mesally fused.
</paragraph>
</subSubSection>
<subSubSection id="B6F416F88334DB5021A2B8D442FCFE4A" lastPageId="2" lastPageNumber="183" pageId="1" pageNumber="182" type="description">
<paragraph id="FE5145738334DB5321A2B8D443B9F8F7" blockId="1.[151,1436,1515,2010]" pageId="1" pageNumber="182">
<emphasis id="CC9A99618334DB5321A2B8D44057F923" bold="true" box="[199,337,1695,1722]" pageId="1" pageNumber="182">Description</emphasis>
. 
<emphasis id="CC9A99618334DB532006B8EB409BF923" box="[355,413,1696,1722]" italics="true" pageId="1" pageNumber="182">Male</emphasis>
(
<figureCitation id="66D559F68334DB5320CBB8EB43B7F920" box="[430,689,1695,1722]" captionStart="FIGURES 1−6" captionStartId="2.[151,264,1458,1483]" captionTargetBox="[154,1433,489,1433]" captionTargetId="figure@2.[154,1433,489,1433]" captionTargetPageId="2" captionText="FIGURES 1−6. Wallacella guianensis, comb. nov. 1−4, Male: 1a, dorsal view; 1b, ventral view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.217; MJWC); 2, dorsal view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.356; MJWC; lines: i—proximal antemedial, ii—distal antemedial, iii—proximal postmedial, iv—premarginal); 3, dorsal view (French Guiana; CGCM 33.107); 4, dorsal view, holotype of Phassus guianensis (USNM). 5, Female: dorsal view (MHNH). 6, Male wing venation (CGCM 30.158). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3730775" httpUri="https://zenodo.org/record/3730775/files/figure.png" pageId="1" pageNumber="182">Figs 1a, 1b, 2, 3, 4, 6</figureCitation>
, 
<figureCitation id="66D559F68334DB5323A5B8EB4263F923" box="[704,869,1696,1722]" captionStart="FIGURES 10−15" captionStartId="5.[151,264,1886,1911]" captionTargetBox="[161,1424,640,1859]" captionTargetId="figure@5.[151,1436,633,1861]" captionTargetPageId="5" captionText="FIGURES 10−15. Wallacella guianensis, comb. nov. 10, Legs (CGCM 30.158). 11−12, Male abdomen (CGCM 35.217): 11, upper dorsal, lower ventral view; 12, tergum and sternum VII and VIII. 13, Male genitalia (CGCM 35.217): 13a, lateral view; 13b, anterior view; 13c, ventral view (pseudotegumen upwards). 14, Male abdomen (holotype), tergite and sternite VII and VIII. 15, Male genitalia (holotype), ventral view (unscaled). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3730779" httpUri="https://zenodo.org/record/3730779/files/figure.png" pageId="1" pageNumber="182">10, 11, 12, 14</figureCitation>
). 
<emphasis id="CC9A99618334DB532218B8EB42BCF923" box="[893,954,1696,1722]" italics="true" pageId="1" pageNumber="182">Head</emphasis>
. Clypeus glabrous anteriorly and mesally projected, and differentiated from the frons. Frons with long scales at the distal edge of the scape, piliform, and porrect. Vertex scales as the frons. Eyes large, occupying 4/5 of the head in anterior view. Labial palpus stout with two palpomeres; distal palpomere embedded dorsally to the basal palpomere; length of palpomeres as long as the subtriangular prelabium. Antenna bipectinate, dark brown; dorsal rami shorter than ventral rami; rami flattened; sensilla caetica present, sensilla trichodea absent.
</paragraph>
<paragraph id="FE5145738334DB5321A2B933428EF843" blockId="1.[151,1436,1515,2010]" pageId="1" pageNumber="182">
<emphasis id="CC9A99618334DB5321A2B933405DF808" box="[199,347,1912,1938]" italics="true" pageId="1" pageNumber="182">Thorax. Legs</emphasis>
(
<figureCitation id="66D559F68334DB53200EB93340B8F80B" box="[363,446,1912,1938]" captionStart="FIGURES 10−15" captionStartId="5.[151,264,1886,1911]" captionTargetBox="[161,1424,640,1859]" captionTargetId="figure@5.[151,1436,633,1861]" captionTargetPageId="5" captionText="FIGURES 10−15. Wallacella guianensis, comb. nov. 10, Legs (CGCM 30.158). 11−12, Male abdomen (CGCM 35.217): 11, upper dorsal, lower ventral view; 12, tergum and sternum VII and VIII. 13, Male genitalia (CGCM 35.217): 13a, lateral view; 13b, anterior view; 13c, ventral view (pseudotegumen upwards). 14, Male abdomen (holotype), tergite and sternite VII and VIII. 15, Male genitalia (holotype), ventral view (unscaled). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3730779" httpUri="https://zenodo.org/record/3730779/files/figure.png" pageId="1" pageNumber="182">Fig. 10</figureCitation>
). Male legs lacking specialized androconia; metaleg reduced; arolium absent. 
<emphasis id="CC9A99618334DB532449B933400CF82F" italics="true" pageId="1" pageNumber="182">Fore- and hindwings</emphasis>
(
<figureCitation id="66D559F68334DB53207FB9D74059F82F" box="[282,351,1947,1974]" captionStart="FIGURES 1−6" captionStartId="2.[151,264,1458,1483]" captionTargetBox="[154,1433,489,1433]" captionTargetId="figure@2.[154,1433,489,1433]" captionTargetPageId="2" captionText="FIGURES 1−6. Wallacella guianensis, comb. nov. 1−4, Male: 1a, dorsal view; 1b, ventral view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.217; MJWC); 2, dorsal view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.356; MJWC; lines: i—proximal antemedial, ii—distal antemedial, iii—proximal postmedial, iv—premarginal); 3, dorsal view (French Guiana; CGCM 33.107); 4, dorsal view, holotype of Phassus guianensis (USNM). 5, Female: dorsal view (MHNH). 6, Male wing venation (CGCM 30.158). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3730775" httpUri="https://zenodo.org/record/3730775/files/figure.png" pageId="1" pageNumber="182">Fig. 6</figureCitation>
). Hepialine venation as in 
<taxonomicName id="39EE3EF08334DB5323E9B9D74237F82F" authority="Walker" authorityName="Walker" authorityYear="1856" box="[652,817,1948,1974]" class="Insecta" family="Hepialidae" genus="Cibyra" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="182" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618334DB5323E9B9D743DCF82F" box="[652,730,1948,1974]" italics="true" pageId="1" pageNumber="182">Cibyra</emphasis>
Walker
</taxonomicName>
(
<bibRefCitation id="9A7F38828334DB532225B9D7457AF82F" author="Mielke, C. G. C. &amp; Casagrande, M. M." box="[832,1148,1948,1974]" pageId="1" pageNumber="182" pagination="73 - 86" refId="ref5865" refString="Mielke, C. G. C. &amp; Casagrande, M. M. (2013) A new Cibyra Walker, 1856 from Southern Brazil (Lepidoptera, Hepialidae). Nachrichten entomologische Vereins Apollo, Neue Golge, 34 (1 / 2), 73 - 86." type="journal article" year="2013">Mielke &amp; Casagrande 2013</bibRefCitation>
); Sc1 present on the forewing and hindwing with CuP complete and a single A vein present.
</paragraph>
<paragraph id="FE5145738337DB5021A2BED245CDFF4E" blockId="2.[151,1437,153,467]" pageId="2" pageNumber="183">
<emphasis id="CC9A99618337DB5021A2BED24037FF2A" box="[199,305,153,179]" italics="true" pageId="2" pageNumber="183">Abdomen</emphasis>
(
<figureCitation id="66D559F68337DB502027BED240F4FF2A" box="[322,498,153,179]" captionStart="FIGURES 10−15" captionStartId="5.[151,264,1886,1911]" captionTargetBox="[161,1424,640,1859]" captionTargetId="figure@5.[151,1436,633,1861]" captionTargetPageId="5" captionText="FIGURES 10−15. Wallacella guianensis, comb. nov. 10, Legs (CGCM 30.158). 11−12, Male abdomen (CGCM 35.217): 11, upper dorsal, lower ventral view; 12, tergum and sternum VII and VIII. 13, Male genitalia (CGCM 35.217): 13a, lateral view; 13b, anterior view; 13c, ventral view (pseudotegumen upwards). 14, Male abdomen (holotype), tergite and sternite VII and VIII. 15, Male genitalia (holotype), ventral view (unscaled). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3730779" httpUri="https://zenodo.org/record/3730779/files/figure.png" pageId="2" pageNumber="183">Figs 11, 12, 14</figureCitation>
). Tergite VIII bilobed, asymmetrical, with right lobe greatly projected posteriorly; sternite VIII in two portions with a main central sclerite and two detached postero-lateral plates.
</paragraph>
<paragraph id="FE5145738337DB5021A2BEAA408FFE36" blockId="2.[151,1437,153,467]" pageId="2" pageNumber="183">
<emphasis id="CC9A99618337DB5021A2BEAA406DFF62" box="[199,363,225,251]" italics="true" pageId="2" pageNumber="183">Male genitalia</emphasis>
(
<figureCitation id="66D559F68337DB50201DBEAA4376FF62" box="[376,624,224,251]" captionStart="FIGURES 10−15" captionStartId="5.[151,264,1886,1911]" captionTargetBox="[161,1424,640,1859]" captionTargetId="figure@5.[151,1436,633,1861]" captionTargetPageId="5" captionText="FIGURES 10−15. Wallacella guianensis, comb. nov. 10, Legs (CGCM 30.158). 11−12, Male abdomen (CGCM 35.217): 11, upper dorsal, lower ventral view; 12, tergum and sternum VII and VIII. 13, Male genitalia (CGCM 35.217): 13a, lateral view; 13b, anterior view; 13c, ventral view (pseudotegumen upwards). 14, Male abdomen (holotype), tergite and sternite VII and VIII. 15, Male genitalia (holotype), ventral view (unscaled). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3730779" httpUri="https://zenodo.org/record/3730779/files/figure.png" pageId="2" pageNumber="183">Figs 13a, 13b, 13c, 15</figureCitation>
). Tegumen (= intermediate plate) as a narrow bar, fused to the pseudotegumen, but distinguished by sclerotization and ventrally articulated with saccus. Saccus U-shaped with a conspicuous and ventral projection forming a pocket with its posterior edge heavily sclerotized. Tergal lobes asymmetrical as an extension of dorsal processes of the pseudotegumen. Pseudotegumen dorso-mesally unfused and ventro-mesally fused. Fultura inferior (= juxta) and fultura superior (= trulleum) both form well sclerotized plates. Valva digitiform. Phallus membranous.
</paragraph>
<paragraph id="FE5145738337DB5021A2BFF242FCFE4A" blockId="2.[151,1437,153,467]" box="[199,1018,440,467]" pageId="2" pageNumber="183">
<emphasis id="CC9A99618337DB5021A2BFF2401DFE4A" box="[199,283,441,467]" italics="true" pageId="2" pageNumber="183">Female</emphasis>
(
<figureCitation id="66D559F68337DB50204FBFF24069FE4A" box="[298,367,440,467]" captionStart="FIGURES 1−6" captionStartId="2.[151,264,1458,1483]" captionTargetBox="[154,1433,489,1433]" captionTargetId="figure@2.[154,1433,489,1433]" captionTargetPageId="2" captionText="FIGURES 1−6. Wallacella guianensis, comb. nov. 1−4, Male: 1a, dorsal view; 1b, ventral view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.217; MJWC); 2, dorsal view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.356; MJWC; lines: i—proximal antemedial, ii—distal antemedial, iii—proximal postmedial, iv—premarginal); 3, dorsal view (French Guiana; CGCM 33.107); 4, dorsal view, holotype of Phassus guianensis (USNM). 5, Female: dorsal view (MHNH). 6, Male wing venation (CGCM 30.158). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3730775" httpUri="https://zenodo.org/record/3730775/files/figure.png" pageId="2" pageNumber="183">Fig. 5</figureCitation>
). Examined only as a photograph. Hindwing includes 2A.
</paragraph>
</subSubSection>
<caption id="AA9115FB8337DB5021F2BBF9432DF9C2" ID-DOI="http://doi.org/10.5281/zenodo.3730775" ID-Zenodo-Dep="3730775" httpUri="https://zenodo.org/record/3730775/files/figure.png" pageId="2" pageNumber="183" startId="2.[151,264,1458,1483]" targetBox="[154,1433,489,1433]" targetPageId="2">
<paragraph id="FE5145738337DB5021F2BBF9432DF9C2" blockId="2.[151,1437,1458,1627]" pageId="2" pageNumber="183">
<emphasis id="CC9A99618337DB5021F2BBF94031FA52" bold="true" box="[151,311,1458,1483]" pageId="2" pageNumber="183">FIGURES 1−6</emphasis>
. 
<taxonomicName id="39EE3EF08337DB502021BBF84325FA52" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" baseAuthorityName="Schaus" baseAuthorityYear="1940" box="[324,547,1459,1483]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="guianensis" status="comb. nov.">
<emphasis id="CC9A99618337DB502021BBF84325FA52" box="[324,547,1459,1483]" italics="true" pageId="2" pageNumber="183">Wallacella guianensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8337DB502354BBF843A1FA52" box="[561,679,1459,1483]" pageId="2" pageNumber="183" rank="species">
<emphasis id="CC9A99618337DB502354BBF8436AFA52" bold="true" box="[561,620,1459,1483]" pageId="2" pageNumber="183">comb</emphasis>
. 
<emphasis id="CC9A99618337DB50231FBBF843A7FA52" bold="true" box="[634,673,1459,1483]" pageId="2" pageNumber="183">nov</emphasis>
.
</taxonomicNameLabel>
1−4, Male: 1a, dorsal view; 1b, ventral view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.217; MJWC); 2, dorsal view (Trinidad, Off Saunders Rd, Inniss Field; CGCM 35.356; MJWC; lines: i—proximal antemedial, ii—distal antemedial, iii—proximal postmedial, iv—premarginal); 3, dorsal view (French Guiana; CGCM 33.107); 4, dorsal view, holotype of 
<taxonomicName id="39EE3EF08337DB502336B854421EF9AE" authorityName="Schaus" authorityYear="1940" box="[595,792,1567,1591]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis id="CC9A99618337DB502336B854421EF9AE" box="[595,792,1567,1591]" italics="true" pageId="2" pageNumber="183">Phassus guianensis</emphasis>
</taxonomicName>
(USNM). 5, Female: dorsal view (MHNH). 6, Male wing venation (CGCM 30.158). Scale bar = 1 cm.
</paragraph>
</caption>
<subSubSection id="B6F416F88337DB5021A2B8C142EAF8C1" pageId="2" pageNumber="183" type="etymology">
<paragraph id="FE5145738337DB5021A2B8C145DBF896" blockId="2.[151,1436,1673,2024]" pageId="2" pageNumber="183">
<emphasis id="CC9A99618337DB5021A2B8C1404EF93D" bold="true" box="[199,328,1674,1700]" pageId="2" pageNumber="183">Etymology</emphasis>
. 
<taxonomicName id="39EE3EF08337DB50203FB8C140D7F93D" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[346,465,1674,1700]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618337DB50203FB8C140D7F93D" box="[346,465,1674,1700]" italics="true" pageId="2" pageNumber="183">Wallacella</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8337DB502087B8C14348F93D" box="[482,590,1674,1700]" pageId="2" pageNumber="183" rank="genus">
<emphasis id="CC9A99618337DB502087B8C1430AF93D" bold="true" box="[482,524,1674,1700]" pageId="2" pageNumber="183">gen</emphasis>
. 
<emphasis id="CC9A99618337DB502378B8C1434FF93D" bold="true" box="[541,585,1674,1700]" pageId="2" pageNumber="183">nov</emphasis>
.
</taxonomicNameLabel>
, is named after Alfred R. Wallace for his important contributions on the 
<taxonomicName id="39EE3EF08337DB5021F2B8E64018F95E" authorityName="Linnaeus" authorityYear="1758" box="[151,286,1709,1735]" class="Insecta" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
in the 19 
<superScript id="099BE83B8337DB5020EFB8E74091F923" attach="left" box="[394,407,1708,1722]" fontSize="6" pageId="2" pageNumber="183">th</superScript>
century. The name follows the tradition of 
<taxonomicName id="39EE3EF08337DB5022E6B8E5455DF951" authority="(Viette)" baseAuthorityName="Viette" box="[899,1115,1709,1736]" class="Insecta" family="Hepialidae" genus="Druceiella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618337DB5022E6B8E542FDF951" box="[899,1019,1710,1736]" italics="true" pageId="2" pageNumber="183">Druceiella</emphasis>
(Viette)
</taxonomicName>
, 
<taxonomicName id="39EE3EF08337DB502502B8E541F5F975" authority="(Grehan &amp; C. Mielke)" baseAuthorityName="Grehan &amp; C. Mielke" class="Insecta" family="Hepialidae" genus="Dugdaleiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618337DB502502B8E545FFF951" box="[1127,1273,1710,1736]" italics="true" pageId="2" pageNumber="183">Dugdaleiella</emphasis>
(Grehan &amp; C. Mielke)
</taxonomicName>
, 
<taxonomicName id="39EE3EF08337DB50219BB89940FCF975" authority="(Viette)" baseAuthorityName="Viette" box="[254,506,1745,1772]" class="Insecta" family="Limacodidae" genus="Hampsoniella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618337DB50219BB899409DF975" box="[254,411,1746,1772]" italics="true" pageId="2" pageNumber="183">Hampsoniella</emphasis>
(Viette)
</taxonomicName>
, 
<emphasis id="CC9A99618337DB502360B899439FF975" box="[517,665,1746,1772]" italics="true" pageId="2" pageNumber="183">Huebneriella</emphasis>
(C. Mielke &amp; Grehan), 
<emphasis id="CC9A99618337DB5022C3B8994520F975" box="[934,1062,1746,1772]" italics="true" pageId="2" pageNumber="183">Kozloviella</emphasis>
(Grehan &amp; C. Mielke), 
<taxonomicName id="39EE3EF08337DB502457B8994027F889" authority="(Viette)" baseAuthorityName="Viette" class="Insecta" family="Hepialidae" genus="Pfitzneriella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618337DB502457B89941C7F889" italics="true" pageId="2" pageNumber="183">Pfitzneriella</emphasis>
(Viette)
</taxonomicName>
, and 
<taxonomicName id="39EE3EF08337DB50203EB8BD423BF889" authority="(C. Mielke, Grehan &amp; Grados)" baseAuthorityName="C. Mielke, Grehan &amp; Grados" box="[347,829,1781,1808]" class="Insecta" family="Hepialidae" genus="Walkeriella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618337DB50203EB8BD40DDF889" box="[347,475,1782,1808]" italics="true" pageId="2" pageNumber="183">Walkeriella</emphasis>
(C. Mielke, Grehan &amp; Grados)
</taxonomicName>
. The gender of the name is feminine.
</paragraph>
<paragraph id="FE5145738337DB5021A2B95242EAF8C1" blockId="2.[151,1436,1673,2024]" pageId="2" pageNumber="183">
<emphasis id="CC9A99618337DB5021A2B9524307F8AD" bold="true" box="[199,513,1817,1844]" pageId="2" pageNumber="183">Geographical distribution</emphasis>
. 
<taxonomicName id="39EE3EF08337DB502377B951438FF8AD" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[530,649,1818,1844]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618337DB502377B951438FF8AD" box="[530,649,1818,1844]" italics="true" pageId="2" pageNumber="183">Wallacella</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8337DB5023FFB9514201F8AD" box="[666,775,1818,1844]" pageId="2" pageNumber="183" rank="genus">
<emphasis id="CC9A99618337DB5023FFB95143C2F8AD" bold="true" box="[666,708,1818,1844]" pageId="2" pageNumber="183">gen</emphasis>
. 
<emphasis id="CC9A99618337DB5023B0B9514207F8AD" bold="true" box="[725,769,1818,1844]" pageId="2" pageNumber="183">nov</emphasis>
.
</taxonomicNameLabel>
, is known to occur in the Northern South America from 
<collectingCountry id="86F905E38337DB5021F2B976400CF8CE" box="[151,266,1853,1879]" name="Venezuela" pageId="2" pageNumber="183">Venezuela</collectingCountry>
to Northern (
<collectingRegion id="3C2A8B918337DB5020C7B97540D1F8C1" box="[418,471,1854,1880]" country="Brazil" name="Para" pageId="2" pageNumber="183">Pará</collectingRegion>
) and Northeastern (
<collectingRegion id="3C2A8B918337DB5023D1B976422FF8C1" box="[692,809,1853,1880]" country="Brazil" name="Maranhao" pageId="2" pageNumber="183">Maranhão</collectingRegion>
) 
<collectingCountry id="86F905E38337DB50225DB976427AF8CE" box="[824,892,1853,1879]" name="Brazil" pageId="2" pageNumber="183">Brazil</collectingCountry>
(
<figureCitation id="66D559F68337DB5022EEB97642E6F8C1" box="[907,992,1853,1880]" captionStart="FIGURE 20" captionStartId="7.[151,250,1980,2005]" captionTargetBox="[198,1390,957,1955]" captionTargetId="figure@7.[198,1390,957,1955]" captionTargetPageId="7" captionText="FIGURE 20. Geographical distribution of Wallacella guianensis, comb. nov., and Gymelloxes juliusboosi, sp. nov., in Northern South America. Trinidad and Tobago inset." figureDoi="http://doi.org/10.5281/zenodo.3730783" httpUri="https://zenodo.org/record/3730783/files/figure.png" pageId="2" pageNumber="183">Fig. 20</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B6F416F88337DB5121A2B9294377FC8E" lastPageId="3" lastPageNumber="184" pageId="2" pageNumber="183" type="discussion">
<paragraph id="FE5145738337DB5121A2B9294240FD56" blockId="2.[151,1436,1673,2024]" lastBlockId="3.[151,1437,153,791]" lastPageId="3" lastPageNumber="184" pageId="2" pageNumber="183">
<emphasis id="CC9A99618337DB5021A2B9294032F8E5" bold="true" box="[199,308,1890,1916]" pageId="2" pageNumber="183">Remarks</emphasis>
. The presence of a posterior lateral knob (not strongly developed) on the tergosternal connection, a broad intermediate connection between the tergosternal bar and the lateral ridge, and a broken anterior margin at the tergosternal connection, along with a very close parallel position between the hindwing Sc and Rs veins, support inclusion of 
<taxonomicName id="39EE3EF08337DB502032B98540C8F871" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[343,462,1998,2024]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618337DB502032B98540C8F871" box="[343,462,1998,2024]" italics="true" pageId="2" pageNumber="183">Wallacella</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8337DB5020BEB9854343F871" box="[475,581,1998,2024]" pageId="2" pageNumber="183" rank="genus">
<emphasis id="CC9A99618337DB5020BEB9854303F871" bold="true" box="[475,517,1998,2024]" pageId="2" pageNumber="183">gen</emphasis>
. 
<emphasis id="CC9A99618337DB502376B9854339F871" bold="true" box="[531,575,1998,2024]" pageId="2" pageNumber="183">nov</emphasis>
.
</taxonomicNameLabel>
, within the ‘cibyrine’ cluster of genera (
<bibRefCitation id="9A7F38828337DB502562B9864599F871" author="Grehan, J. R." box="[1031,1183,1997,2024]" pageId="2" pageNumber="183" pagination="1 - 30" refId="ref5577" refString="Grehan, J. R. (2012) Morphological evidence for phylogenetic relationships within the Hepialidae (Lepidoptera: Exoporia). Bulletin of the Buffalo Society of Natural Sciences, 42, 1 - 30." type="journal article" year="2012">Grehan 2012</bibRefCitation>
). Phylogenetic affinity within the cibyrine cluster is currently unresolved due to the lack of identified apomorphies shared between genera. The female genitalia are not known for many taxa and male genitalia often show distinct features shared by only one or a few species and this has contributed to the designation of several monotypic genera. A revision of 
<taxonomicName id="39EE3EF08336DB512441BEAA41EBFEB9" authority="(Viette)" baseAuthorityName="Viette" class="Insecta" family="Hepialidae" genus="Druceiella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618336DB512441BEAA449AFF62" box="[1316,1436,225,251]" italics="true" pageId="3" pageNumber="184">Druceiella</emphasis>
(Viette)
</taxonomicName>
by 
<bibRefCitation id="9A7F38828336DB51207DBF4E4339FE86" author="Grehan, J. R. &amp; Rawlins, J. E." box="[280,575,261,287]" pageId="3" pageNumber="184" pagination="165 - 192" refId="ref5730" refString="Grehan, J. R. &amp; Rawlins, J. E. (2018) Taxonomic revision and vicariance biogeography of the central and South American ghost moth genus Druceiella (Lepidoptera: Hepialidae). Annals of the Carnegie Museum, 85, 165 - 192. https: // doi. org / 10.2992 / 007.085.0203" type="journal article" year="2018">Grehan &amp; Rawlins (2018)</bibRefCitation>
noted that this genus shared a pattern of three darker forewing patches with 
<taxonomicName id="39EE3EF08336DB5121F2BF624079FEDD" authority="(Viette)" baseAuthorityName="Viette" box="[151,383,297,324]" class="Insecta" family="Hepialidae" genus="Pfitzneriana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618336DB5121F2BF624027FEDA" box="[151,289,297,323]" italics="true" pageId="3" pageNumber="184">Pfitzneriana</emphasis>
(Viette)
</taxonomicName>
and 
<taxonomicName id="39EE3EF08336DB5120D2BF624328FEDA" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[439,558,297,323]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618336DB5120D2BF624328FEDA" box="[439,558,297,323]" italics="true" pageId="3" pageNumber="184">Wallacella</emphasis>
</taxonomicName>
[as ‘ 
<taxonomicName id="39EE3EF08336DB512302BF6243C5FEDA" box="[615,707,297,323]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618336DB512302BF6243C5FEDA" box="[615,707,297,323]" italics="true" pageId="3" pageNumber="184">Phassus</emphasis>
</taxonomicName>
’] 
<taxonomicName id="39EE3EF08336DB5123BBBF614250FEDA" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" baseAuthorityName="Mielke &amp; Grehan &amp; Cock" baseAuthorityYear="2020" box="[734,854,298,323]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="guianensis" status="comb. nov.">
<emphasis id="CC9A99618336DB5123BBBF614250FEDA" box="[734,854,298,323]" italics="true" pageId="3" pageNumber="184">guianensis</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8336DB512201BF6242CCFEDA" box="[868,970,297,323]" pageId="3" pageNumber="184" rank="species">
<emphasis id="CC9A99618336DB512201BF6242A3FEDA" bold="true" box="[868,933,297,323]" pageId="3" pageNumber="184">comb</emphasis>
. 
<emphasis id="CC9A99618336DB5122D0BF6242C3FEDA" bold="true" box="[949,965,297,323]" pageId="3" pageNumber="184">n</emphasis>
.
</taxonomicNameLabel>
, although this is less distinct in the latter species. The three genera also shared an apically fused apex of the pseudotegumen that is also strongly sclerotized and lined with transverse grooves and a sclerotized expansion of the posterior saccus in the form of a central rectangular protuberance in 
<taxonomicName id="39EE3EF08336DB5120F5BFDE431CFE36" box="[400,538,405,431]" class="Insecta" family="Hepialidae" genus="Pfitzneriana" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618336DB5120F5BFDE431CFE36" box="[400,538,405,431]" italics="true" pageId="3" pageNumber="184">Pfitzneriana</emphasis>
</taxonomicName>
and an enlarged rim with laterally projecting tubercles in 
<taxonomicName id="39EE3EF08336DB5125CEBFDE4424FE36" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[1195,1314,405,431]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618336DB5125CEBFDE4424FE36" box="[1195,1314,405,431]" italics="true" pageId="3" pageNumber="184">Wallacella</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8336DB512454BFDE449AFE36" box="[1329,1436,405,431]" pageId="3" pageNumber="184" rank="genus">
<emphasis id="CC9A99618336DB512454BFDE445DFE36" bold="true" box="[1329,1371,405,431]" pageId="3" pageNumber="184">gen</emphasis>
. 
<emphasis id="CC9A99618336DB51240FBFDE4490FE36" bold="true" box="[1386,1430,405,431]" pageId="3" pageNumber="184">nov</emphasis>
.
</taxonomicNameLabel>
<taxonomicName id="39EE3EF08336DB5121F2BFF24009FE4A" box="[151,271,441,467]" class="Insecta" family="Hepialidae" genus="Druceiella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618336DB5121F2BFF24009FE4A" box="[151,271,441,467]" italics="true" pageId="3" pageNumber="184">Druceiella</emphasis>
</taxonomicName>
and 
<taxonomicName id="39EE3EF08336DB512026BFF240BCFE4A" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[323,442,441,467]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618336DB512026BFF240BCFE4A" box="[323,442,441,467]" italics="true" pageId="3" pageNumber="184">Wallacella</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8336DB5120A2BFF24336FE4A" box="[455,560,441,467]" pageId="3" pageNumber="184" rank="genus">
<emphasis id="CC9A99618336DB5120A2BFF240F7FE4A" bold="true" box="[455,497,441,467]" pageId="3" pageNumber="184">gen</emphasis>
. 
<emphasis id="CC9A99618336DB51209BBFF2432CFE4A" bold="true" box="[510,554,441,467]" pageId="3" pageNumber="184">nov</emphasis>
.
</taxonomicNameLabel>
, have an asymmetrical posterior abdomen margin with a posteriorly projecting lobe which is on the left in 
<taxonomicName id="39EE3EF08336DB5120A9BF964345FE6E" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[460,579,477,503]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618336DB5120A9BF964345FE6E" box="[460,579,477,503]" italics="true" pageId="3" pageNumber="184">Wallacella</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8336DB512337BF9643BAFE6E" box="[594,700,477,503]" pageId="3" pageNumber="184" rank="genus">
<emphasis id="CC9A99618336DB512337BF96437AFE6E" bold="true" box="[594,636,477,503]" pageId="3" pageNumber="184">gen</emphasis>
. 
<emphasis id="CC9A99618336DB5123EFBF9643B0FE6E" bold="true" box="[650,694,477,503]" pageId="3" pageNumber="184">nov</emphasis>
.
</taxonomicNameLabel>
, and on the right in 
<taxonomicName id="39EE3EF08336DB5122C7BF96451CFE6E" box="[930,1050,477,503]" class="Insecta" family="Hepialidae" genus="Druceiella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618336DB5122C7BF96451CFE6E" box="[930,1050,477,503]" italics="true" pageId="3" pageNumber="184">Druceiella</emphasis>
</taxonomicName>
. Since this projection represents a uniquely derived feature within the 
<taxonomicName id="39EE3EF08336DB512344BC4A439CFD82" box="[545,666,513,539]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
it is tempting to see it as a potential synapomorphy for these two genera since they both occur in Northern South America and the feature has not been recorded in any other 
<taxonomicName id="39EE3EF08336DB512479BC6E449FFDA6" box="[1308,1433,549,575]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
. An additional feature indicative of potential close relationship is the shape and size of the sclerotized tergal lobes which are asymmetrically expanded on the right side in 
<taxonomicName id="39EE3EF08336DB512262BC264279FD1E" box="[775,895,621,647]" class="Insecta" family="Hepialidae" genus="Druceiella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618336DB512262BC264279FD1E" box="[775,895,621,647]" italics="true" pageId="3" pageNumber="184">Druceiella</emphasis>
</taxonomicName>
and on the left side in 
<taxonomicName id="39EE3EF08336DB512519BC2645F5FD1E" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[1148,1267,621,647]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618336DB512519BC2645F5FD1E" box="[1148,1267,621,647]" italics="true" pageId="3" pageNumber="184">Wallacella</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8336DB512464BC26446CFD1E" box="[1281,1386,621,647]" pageId="3" pageNumber="184" rank="genus">
<emphasis id="CC9A99618336DB512464BC26442DFD1E" bold="true" box="[1281,1323,621,647]" pageId="3" pageNumber="184">gen</emphasis>
. 
<emphasis id="CC9A99618336DB51245DBC264462FD1E" bold="true" box="[1336,1380,621,647]" pageId="3" pageNumber="184">nov</emphasis>
.
</taxonomicNameLabel>
, but the presence of this feature also needs to be further assessed in other genera. The male antennae of 
<taxonomicName id="39EE3EF08336DB512590BCDA446BFD32" box="[1269,1389,657,683]" class="Insecta" family="Hepialidae" genus="Druceiella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus">
<emphasis id="CC9A99618336DB512590BCDA446BFD32" box="[1269,1389,657,683]" italics="true" pageId="3" pageNumber="184">Druceiella</emphasis>
</taxonomicName>
and 
<taxonomicName id="39EE3EF08336DB5121F2BCFE4008FD56" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[151,270,693,719]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618336DB5121F2BCFE4008FD56" box="[151,270,693,719]" italics="true" pageId="3" pageNumber="184">Wallacella</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8336DB512079BCFE4080FD56" box="[284,390,693,719]" pageId="3" pageNumber="184" rank="genus">
<emphasis id="CC9A99618336DB512079BCFE4040FD56" bold="true" box="[284,326,693,719]" pageId="3" pageNumber="184">gen</emphasis>
. 
<emphasis id="CC9A99618336DB512031BCFE4086FD56" bold="true" box="[340,384,693,719]" pageId="3" pageNumber="184">nov</emphasis>
.
</taxonomicNameLabel>
share similarly long and flattened rami.
</paragraph>
<paragraph id="FE5145738336DB5121A2BC924377FC8E" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="184">
The Brazilian specimens examined of 
<taxonomicName id="39EE3EF08336DB512312BC9243E8FD6A" authority="Mielke &amp; Grehan &amp; Cock, 2020" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" box="[631,750,729,755]" class="Insecta" family="Hepialidae" genus="Wallacella" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CC9A99618336DB512312BC9243E8FD6A" box="[631,750,729,755]" italics="true" pageId="3" pageNumber="184">Wallacella</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="D7A9241A8336DB512399BC924260FD6A" box="[764,870,729,755]" pageId="3" pageNumber="184" rank="genus">
<emphasis id="CC9A99618336DB512399BC924220FD6A" bold="true" box="[764,806,729,755]" pageId="3" pageNumber="184">gen</emphasis>
. 
<emphasis id="CC9A99618336DB512251BC924266FD6A" bold="true" box="[820,864,729,755]" pageId="3" pageNumber="184">nov</emphasis>
.
</taxonomicNameLabel>
, have not been assigned to 
<taxonomicName id="39EE3EF08336DB5125F8BC91443DFD6A" authorityName="Mielke &amp; Grehan &amp; Cock" authorityYear="2020" baseAuthorityName="Mielke &amp; Grehan &amp; Cock" baseAuthorityYear="2020" box="[1181,1339,729,755]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="guianensis">
<emphasis id="CC9A99618336DB5125F8BC9145B2FD6A" box="[1181,1204,730,755]" italics="true" pageId="3" pageNumber="184">W</emphasis>
. 
<emphasis id="CC9A99618336DB5125A6BC91443DFD6A" box="[1219,1339,730,755]" italics="true" pageId="3" pageNumber="184">guianensis</emphasis>
</taxonomicName>
pending future investigation of their species status.
</paragraph>
</subSubSection>
</treatment>
</document>