358 lines
35 KiB
XML
358 lines
35 KiB
XML
<document id="F7473DFC74D2AED63867340B099F3D3E" ENCODING="UTF8" ID-GBIF-Dataset="9fcd647a-09f8-4ce9-ba68-778f173a6c1c" ModsDocID="22783" checkinTime="1249509757754" checkinUser="christiana" docAuthor="Hosoishi, S. & Ogata, K." docDate="2009" docId="658551BC1E82ECF36451764ADE4DB693" docLanguage="en" docName="22783.xml" docOrigin="Zootaxa 2062" docSource="http://hol.osu.edu/reference-full.html?id=22783" docTitle="Crematogaster (Physocrema) Forel" docType="treatment" docVersion="1" lastPageNumber="19" masterDocId="A92802EA8A918BF15D7133FD48DB5498" masterDocTitle="A taxonomic revision of the Asian endemic subgenus Physocrema of the genus Crematogaster (Hymenoptera: Formicidae)." masterLastPageNumber="36" masterPageNumber="15" pageNumber="17" updateTime="1694988214208" updateUser="admin">
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<mods:title id="2F6EFBE9C9EEEB8168ECDC0F4170D0F2">A taxonomic revision of the Asian endemic subgenus Physocrema of the genus Crematogaster (Hymenoptera: Formicidae).</mods:title>
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<mods:namePart id="08B4E050750957CF803424834CB56607">Hosoishi, S.</mods:namePart>
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<mods:namePart id="273DE0DBBA1AB2CC101E6B9A3646C9DA">Ogata, K.</mods:namePart>
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<mods:date id="FADA929F9B2D509D06E3BCCB3E936345">2009</mods:date>
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<mods:classification id="83BDDE0F5A3205618D97AC0EBEA6627B">journal article</mods:classification>
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<mods:identifier id="C9D2077175041BB883AE4104D0C6C638" type="HNS-PUB">22783</mods:identifier>
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<treatment id="658551BC1E82ECF36451764ADE4DB693" ID-DOI="http://doi.org/10.5281/zenodo.6214284" ID-GBIF-Taxon="100122499" ID-Zenodo-Dep="6214284" LSID="urn:lsid:plazi:treatment:658551BC1E82ECF36451764ADE4DB693" httpUri="http://treatment.plazi.org/id/658551BC1E82ECF36451764ADE4DB693" lastPageNumber="19" pageNumber="17">
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<subSubSection id="3C1E4354F2DE48B424198340ABEBBB38" type="nomenclature">
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<paragraph id="F3F2091156D0397B5AFE04AB5D6C49B8" pageNumber="17">
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The subgenus
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<taxonomicName id="D78307C77F82E2733F41C1AC62CB0B86" authority="Forel" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema Forel</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C55FB4F3FC534B825EDB6B8449F94A71" type="reference_group">
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<paragraph id="2392D8E4B36AE9C6D148CBDE5E040D73" pageNumber="17">
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<taxonomicName id="8EE1802BCF3BA6551536DBC92EF15D7C" authority="Forel, 1912" authorityName="Forel" authorityYear="1912" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema Forel, 1912</taxonomicName>
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: 220 [as subgenus of
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<taxonomicName id="69921DB992ADF60CE9AFA8DF331B8D04" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">Crematogaster</taxonomicName>
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]. Type-species:
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<taxonomicName id="1E40223C4DEFAC63EEB3E597CB6C907A" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="inflata">Crematogaster inflata</taxonomicName>
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, by subsequent designation of Wheeler 1913: 82.
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</paragraph>
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<paragraph id="742EEFFF625DC1057D3BA3B854E57B7E" pageNumber="17">
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<taxonomicName id="99C14270B39914D83A1ED62C75555A2E" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema</taxonomicName>
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as genus: Soulie, 1964: 398.
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</paragraph>
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<paragraph id="00194F0B54C8062460971543FDD96A63" pageNumber="17">
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<taxonomicName id="AB99D5DD0CCB4329A9A52B546BA25F4C" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema</taxonomicName>
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as junior synonym of
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<taxonomicName id="2CCF4B1EFAA9C750D5DAFFBE69A00608" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">Crematogaster</taxonomicName>
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: Brown, 1973: 183 [provisional]; Smith, D. R. 1979: 1376; Holldobler & Wilson, 1990: 13.
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</paragraph>
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<paragraph id="2DB82B5607EEED01121B3137524A566A" pageNumber="17">
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<taxonomicName id="5039DE49D0D408B9072C7843C43099B2" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema</taxonomicName>
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as subgenus of
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<taxonomicName id="4F36811BD7FF1AF0A2035F33ECBAAE65" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">Crematogaster</taxonomicName>
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: Forel, 1912: 220; Forel, 1917: 242; Emery, 1922: 139; Wheeler,W. M. 1922: 662; subsequent authors except those above; Bolton, 1995: 42.
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</paragraph>
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</subSubSection>
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<subSubSection id="765CD7AD0182E5B214A436F4D253096D" type="diagnosis">
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<paragraph id="5244C9FE4DA049659018AC3E64A56EC2" pageNumber="17">Diagnosis of worker. Head subquadrate, slightly broader than long, with weakly concave posterior margin, angular posterior corners and subparallel sides. Occipital carinae developed. Mandible striate, with four teeth in small workers and five in larger workers, apical and subapical teeth large. Anterior margin of clypeus convex with slightly impressed median portion (Figs 1, 2); anterolateral margins of clypeus protruded anteriorly, resulting in an appearance of the anterolateral and anteromedian margins being at about the same level (Figs 1, 2); posterior margin of clypeus rounded between frontal lobes. Frontal carinae almost parallel. Antennae 11 segmented; antennal club 4-jointed. Compound eyes large but not distinctly projecting beyond lateral margins of head in full face view.</paragraph>
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<paragraph id="B5F4543043C805C695E11510C37CD46B" pageNumber="17">Pronotum and mesonotum usually without defined suture, rarely promesonotal suture present in large workers. Mesothoracic spiracle reduced to form small pit dorsoventrally. Variable extent of posterior mesosoma enlarged and swollen (Figs 3, 4). Metapleural gland opening circular; propodeal spiracle facing posterolaterally (Figs 3, 4).</paragraph>
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<paragraph id="98E546438D2132A79C912B39D4BD01C7" pageNumber="17">Petiole flattened without node nor subpetiolar process; spiracle situated anteriorly midway between dorsal and ventral margins of petiole in lateral view, directed laterally; in dorsal view, elliptical with convex sides, longer than broad (Fig. 5). Postpetiole without distinctly longitudinal median sulcus, weakly bilobed behind in large workers (Fig. 5); spiracle situated distinctly anteriorly on the lateral surface.</paragraph>
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</subSubSection>
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<subSubSection id="D5D685C809B38386E2C8345449BDFCCA" type="discussion">
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<paragraph id="F2615C56B2DBE3390E6968F0E9FF5002" pageNumber="17">
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Comments. Workers of the subgenus
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<taxonomicName id="1EC6C9B1F7ED681CC47673BB0E9F826E" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema</taxonomicName>
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can be distinguished from those of all other subgenera of
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<taxonomicName id="25B1B5BA952950414B7F9D81DE8F0608" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">Crematogaster</taxonomicName>
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by the features of (1) anteromedian margin of clypeus convex with slightly impressed median portion; (2) anterolateral margins of clypeus produced anteriorly; (3) 4-jointed antennal club (but, 3- jointed in
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<taxonomicName id="75F21CF65B8D890821BB3D2C8CD4DCB8" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="mucronata">mucronata</taxonomicName>
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); (4) propodeum swollen; (5) metapleural gland orifice circular; (6) petiole elliptical; (7) petiole without node-like process posteriorly. The subgenus is unique among Asian
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<taxonomicName id="6DC9471058AC35D8D559AB7192D2BC1F" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">Crematogaster</taxonomicName>
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ants in having features (4), (5), (6) and (7). The workers of the subgenus
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<taxonomicName id="993B3F06D9C78BBE6AEF7954B5AD2B7D" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema</taxonomicName>
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share features (1), (2), (3) and (7) with the subgenus
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<taxonomicName id="FA40A1C24ABF948E9160211A0AC55F51" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Paracrema">Paracrema</taxonomicName>
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. In Asian
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<taxonomicName id="84749B7BBD553638344C064FEE0BDE5B" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="genus">Crematogaster</taxonomicName>
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specimens examined, the workers of the subgenera
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<taxonomicName id="D3EEBCFD5DCEA98A9D5EF85A16536CE5" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Crematogaster">Crematogaster</taxonomicName>
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,
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<taxonomicName id="22FBE54490E5A31037D1F5593B834366" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Decacrema">Decacrema</taxonomicName>
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,
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<taxonomicName id="F25B9365E04E78C2DF205C4F5303DA66" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Mesocrema">Mesocrema</taxonomicName>
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,
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<taxonomicName id="A878B13B3B249432F6C8B61EEEBFE22F" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Oxygyne">Oxygyne</taxonomicName>
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,
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<taxonomicName id="61EE971C941C66181F19FB3F64801D94" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Paracrema">Paracrema</taxonomicName>
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,
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<taxonomicName id="0F8299AA03CC3C44C39D4885887BB8F2" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema</taxonomicName>
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and
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<taxonomicName id="5CA0B7DF6158C02B4AD297BAA2C073FF" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Xiphocrema">Xiphocrema</taxonomicName>
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share features (2) and (7).
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</paragraph>
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</subSubSection>
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<subSubSection id="36C4EAC809C1DE8EC55B01C4521E248A" type="distribution">
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<paragraph id="FAF43AA903E4F7482C7F4EE959130607" pageNumber="18">
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<pageBreakToken id="DAD1A824B8B00951A682A25ACE17B91B" pageNumber="18">Distribution</pageBreakToken>
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. Southeast Asia: Vietnam (southern parts), Laos, Cambodia, Thailand, Myanmar (southeastern parts), Malaysia (Peninsula and Borneo), Singapore, Indonesia (Sumatra, Mentawai, Java, Bali, Lombok, Kalimantan, Sulawesi, Timor), Philippines (Fig. 21).
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</paragraph>
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</subSubSection>
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<subSubSection id="4D6EDA5F0550E4188B05E54E5F60C7B4" type="multiple">
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<caption id="E1EFDD1252918B5CEABFE5491424ACFE">
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<paragraph id="C0A02B0A39F33FE78A4AE95E1BED1A78" pageNumber="18">
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FIGURES 1-2. Clypeus in dorsal view: 1-
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<taxonomicName id="F0C1551239E120AD477FBE4FE3A02539" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="inflata" subGenus="Physocrema">C. (P.) inflata</taxonomicName>
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; 2-
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<taxonomicName id="A4F59C4CE89A5FD7DE0B7843FD34A75C" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aurita" subGenus="Physocrema">C. (P.) aurita</taxonomicName>
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. White arrow indicates anterolateral margin of clypeus.
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</paragraph>
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</caption>
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<caption id="BDC377DA42E17EF908446C75E87E81B4">
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<paragraph id="12AA40C771DD20AAF3E59E4904C7B53C" pageNumber="18">
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FIGURES 3-4. Mesosoma in lateral view: 3-
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<taxonomicName id="34368034539AD2B866662981415C5537" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ampullaris" subGenus="Physocrema">C. (P.) ampullaris</taxonomicName>
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; 4-
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<taxonomicName id="3380B4E11082D73DAC72C5241B8D9E0D" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="physothorax" subGenus="Physocrema">C. (P.) physothorax</taxonomicName>
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.
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</paragraph>
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</caption>
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<caption id="A8CBB0BC945D6372EE641F8A39930974">
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<paragraph id="4338ED4E754FD6E027808A6B4FC289A3" pageNumber="18">
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FIGURE 5. Petiole and postpetiole in dorsal view:
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<taxonomicName id="76511E32F03BF530D9CE74550AE16FAB" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="physothorax" subGenus="Physocrema">C. (P.) physothorax</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="3452379F3CDB32EDF59CD343FEF2AED1" type="synonymic_list">
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<paragraph id="D4B80C1758ED4ED38828EF399FEBE324" pageNumber="18">
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Synonymic list of the subgenus
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<taxonomicName id="AF6ADFB75B8FDFE68E2CB816892D890E" authority="Forel" authorityName="Forel" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema Forel</taxonomicName>
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</paragraph>
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<paragraph id="36C05B224259D938DAEE97AFCF76A303" pageNumber="18">
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<taxonomicName id="B03B6671E1CE80E784BBF64D5666DD38" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28069" authority="F. Smith, 1861" authorityName="F. Smith" authorityYear="1861" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster ampullaris Smith" order="Hymenoptera" pageNumber="18" phylum="Arthropoda" rank="species" species="ampullaris">ampullaris F. Smith, 1861</taxonomicName>
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.
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</paragraph>
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<paragraph id="8DA0D656F5F150624A6655E20571DB10" pageNumber="18">
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<taxonomicName id="A841D1E9E24B475BE22D1F0672DAAA7F" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28089" authority="Karavajew, 1935" authorityName="Karavajew" authorityYear="1935" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster aurita Karavaiev" order="Hymenoptera" pageNumber="18" phylum="Arthropoda" rank="species" species="aurita">aurita Karavajew, 1935</taxonomicName>
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.
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</paragraph>
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<paragraph id="122BD263CB2422D9FB74D297A01CAAA9" pageNumber="19">
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<taxonomicName id="B110CD3CC7699D7A8EFD4018B8CB1410" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28092" authority="Menozzi, 1925" authorityName="Menozzi" authorityYear="1925" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster bakeri Menozzi" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="bakeri">
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<pageBreakToken id="B56C3299DA16E79274E13852F2C7EFD8" pageNumber="19">bakeri</pageBreakToken>
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Menozzi, 1925
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="156483C7E0AF8241552C6A4460713108" pageNumber="19">
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<taxonomicName id="FD5E43354C6B8E19F6C05775F0B8AACD" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:137379" authority="F. Smith, 1857" authorityName="F. Smith" authorityYear="1857" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster difformis Smith" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="difformis">difformis F. Smith, 1857</taxonomicName>
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.
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</paragraph>
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<paragraph id="183DE73E83F9012879B9265444EC994F" pageNumber="19">
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=
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<taxonomicName id="6AB9EBE740F1FD158F426F7A0F8FAF49" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:137442" authority="Mayr, 1867" authorityName="Mayr" authorityYear="1867" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster edentata Mayr" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="edentata">edentata Mayr, 1867</taxonomicName>
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.
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</paragraph>
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<paragraph id="DD753253D08A8407BAB3AC6A40DC90DB" pageNumber="19">
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<taxonomicName id="CC59F7AFB94C062111AF2BF1AC0874C7" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28247" authority="F. Smith, 1857" authorityName="F. Smith" authorityYear="1857" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster inflata Smith" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="inflata">inflata F. Smith, 1857</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="068B0366D59334D237D6A806EA03E53D" pageNumber="19">
|
|
=
|
|
<taxonomicName id="BE4BF0A96CB219BE2C7B641BCF6017AF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28321" authority="Donisthorpe, 1941" authorityName="Donisthorpe" authorityYear="1941" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster moorei Donisthorpe" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="moorei">moorei Donisthorpe, 1941</taxonomicName>
|
|
.
|
|
<taxonomicNameLabel id="B82D4A2BD4DD4A39FFA317B80735C3A0">syn. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
<paragraph id="07259C112A54BDE4CA356F2F009C8EE5" pageNumber="19">
|
|
<taxonomicName id="8182748AA00F584BE6250375D5649F7B" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28325" authority="Emery, 1900" authorityName="Emery" authorityYear="1900" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster mucronata Emery" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="mucronata">mucronata Emery, 1900</taxonomicName>
|
|
. stat. rev.
|
|
</paragraph>
|
|
<paragraph id="16AE2B5729A15EEB35D97EC0AFE2B99B" pageNumber="19">
|
|
=
|
|
<taxonomicName id="C4054FDBFE69B66EE1BD2620C29DB187" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:137415" authority="Forel, 1922" authorityName="Forel" authorityYear="1922" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster fulmeki Forel" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="fulmeki">fulmeki Forel, 1922</taxonomicName>
|
|
.
|
|
<taxonomicNameLabel id="13857D06B1FF10A18AE277A47A00924F">syn. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
<paragraph id="06FFA4529A98C16A040791B1B792B68F" pageNumber="19">
|
|
<taxonomicName id="E8BCD9DF226AFCFF201DAC68C39597C8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:28345" authority="Stitz, 1925" authorityName="Stitz" authorityYear="1925" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster onusta Stitz" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="onusta">onusta Stitz, 1925</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="BAA1D73F9A1263FD501873F6FC424407" pageNumber="19">
|
|
<taxonomicName id="491A5B0EF3CB078242C7B1D577C3BFEF" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:149381" authority="Emery, 1889" authorityName="Emery" authorityYear="1889" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster difformis physothorax Emery" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="physothorax">physothorax Emery, 1889</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="952DC607CB31EAE513E60A6E5ACEB8AF" pageNumber="19">
|
|
<taxonomicName id="BD59F53FA92CF9158B17E17701EBC7E4" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:149437" authority="Forel, 1901" authorityName="Forel" authorityYear="1901" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster difformis sewardi Forel" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="sewardi">sewardi Forel, 1901</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="C24AD6F2CD1A7ED7E0B454D9089D6948" pageNumber="19">
|
|
<taxonomicName id="C3B7005AD41038625096CB96FA08DCB6" authority="Hosoishi & Ogata" authorityName="Hosoishi & Ogata" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tanakai" status="sp. nov.">tanakai Hosoishi & Ogata</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel id="9B6695246F7B1B265DE12451F2E366D3" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
<paragraph id="3B65F8A5184927332D4AC29ED51CCEE4" pageNumber="19">
|
|
<taxonomicName id="5364506FF843623277F7EF287CCA6295" authority="Forel, 1911" authorityName="Forel" authorityYear="1911" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="vacca">vacca Forel, 1911</taxonomicName>
|
|
.
|
|
<taxonomicNameLabel id="6384408A09A28B3409A77A504AF5701A">stat. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
<paragraph id="14A3F3EBE9358CC11D551EAC921E030A" pageNumber="19">
|
|
=
|
|
<taxonomicName id="9230DD771BB3AF7B835D3DFDE6454F63" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:149171" authority="Wheeler, W. M., 1919" authorityName="Wheeler, W. M." authorityYear="1919" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster stethogompha stethogompha Wheeler" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="stethogompha" subSpecies="stethogompha">stethogompha stethogompha Wheeler, W. M., 1919</taxonomicName>
|
|
.
|
|
<taxonomicNameLabel id="029B712A1F4A0B86CA46F9C5B04E43F9">syn. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
<paragraph id="DF8DC2BA2A619E4650E0491067A534A0" pageNumber="19">
|
|
=
|
|
<taxonomicName id="5DA204845E1FCBA1D49E328ABFC9A25A" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:149172" authority="Wheeler, W. M., 1919" authorityName="Wheeler, W. M." authorityYear="1919" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster stethogompha detritinodis Wheeler" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="subSpecies" species="stethogompha" subSpecies="detritinodis">stethogompha detritinodis Wheeler, W. M., 1919</taxonomicName>
|
|
.
|
|
<taxonomicNameLabel id="5CF406A224B7E29608227E4887B5076F">syn. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
<paragraph id="A6AFC8724492A02854438DFD74B2C97C" pageNumber="19">
|
|
<taxonomicName id="7445E3BEF4EE925E147182EE52DF535D" authority="Hosoishi & Ogata" authorityName="Hosoishi & Ogata" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="yamanei" status="sp. nov.">yamanei Hosoishi & Ogata</taxonomicName>
|
|
,
|
|
<taxonomicNameLabel id="EE79AD971ABEA7641796CC63999409CA" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="F6C2A28B20A16C79126FA88B4508501A" type="multiple">
|
|
<paragraph id="07590ECA4D2A526D23E481F4218A2A33" pageNumber="19">
|
|
Key to species based on the worker caste (except for
|
|
<taxonomicName id="4CE0B67DEB3393365E3EE7312C13E38C" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="bakeri">C. bakeri</taxonomicName>
|
|
, which is incertae sedis.)
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="FFA9FF169E0EB4E2BEC21EB11CDF56B2" type="key">
|
|
<keyStep id="FE3710636B0CE017132FEB364D0A1387">
|
|
<paragraph id="0D2D8CD78E98466C1451CC7CCA802411" pageNumber="19">
|
|
<keyLead id="8BCA4683C0BC29B57C76ECDD75095750">1 Propodeal spines developed and with pointed apices................................................................................................... 2</keyLead>
|
|
</paragraph>
|
|
<paragraph id="5B183087716C9F7A702CCAE63446E92B" pageNumber="19">
|
|
<keyLead id="739A8AAFCF53C4912FE62A0F673CE7D8">- Propodeal spines undeveloped and forming obtuse lobes ............................................................................................ 4</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="D7A4E7DE64536A71F0EEDE8F34B57597">
|
|
<paragraph id="82AE01B461D0B7F107319C3D27830E2C" pageNumber="19">
|
|
<keyLead id="CDCA0E27A303663A7A3067FDE6498A57">
|
|
2 Antennal club 3-jointed. Pronotum raised steeply toward mesonotum in lateral view................................
|
|
<taxonomicName id="DC43A5BA0530D5EFE664D2E347D264EE" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="mucronata">mucronata</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="C99E96F466754D4808E2A906FA769666" pageNumber="19">
|
|
<keyLead id="973CC5622339429C6C3CB36343EEF1A2">- Antennal club 4-jointed. Pronotum raised moderately toward mesonotum in lateral view.........................................3</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="F696B8BC968FBE92EA9536FD5BC9C6CA">
|
|
<paragraph id="CED021DA73DCD8E4D13E9DF97FBCC259" pageNumber="19">
|
|
<keyLead id="9351C24B287EC82BDDABD4F5926827CF">
|
|
3 In lateral view, pronotum and mesonotum forming a single convexity, propodeum distinctly raised relative to promesonotum, metanotal groove deep.........................................................................................................................
|
|
<taxonomicName id="9A6B297A5FC0A429248D67F5F6DFFE9C" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="vacca">vacca</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="EC56E22A84DA89B8834E135D6C06A103" pageNumber="19">
|
|
<keyLead id="CB6902C82A1A057C26D33CE5E60F8779">
|
|
- In lateral view, pronotum and mesonotum forming a single convexity, propodeum slightly raised relative to promesonotum, metanotal groove less deep..................................................................................................
|
|
<taxonomicName id="A954D36D6C731E01A9BE61C13FE36E06" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="yamanei" status="sp. nov.">yamanei</taxonomicName>
|
|
<taxonomicNameLabel id="31643EB4AF7E93D6005748F33F9120A1" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="CE599EF23E4AE2B1F185832AB719BC5E">
|
|
<paragraph id="7321FBE78FFD5C4C2E894A3857790FED" pageNumber="19">
|
|
<keyLead id="5ED7F15A27C8339FB352AE7DD7C635AE">
|
|
4 Dorsal median portion of head swollen................................................................................................
|
|
<taxonomicName id="A8612065D06E921581642899ECA2D9BF" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="tanakai" status="sp. nov.">tanakai</taxonomicName>
|
|
<taxonomicNameLabel id="6658FF5DAD97FF9967496FFB571E838A" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="F9B43F23279270129CC471084643E2FF" pageNumber="19">
|
|
<keyLead id="22E05C818E49363F5A6EED52DE0E3207">- Dorsal median portion of head not swollen..................................................................................................................5</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="FC11AE968C892B05A0B3FDE9B2436F9F">
|
|
<paragraph id="477E0295B6F0BD1CBE0A6205D0744155" pageNumber="19">
|
|
<keyLead id="7C4E353B9D24C6F67770151F1D8D9083">5 Propodeal spiracle situated near the metapleural gland orifice, the distance between them about the same as the propodeal spiracle diameter (Fig. 4) ............................................................................................................................. 6</keyLead>
|
|
</paragraph>
|
|
<paragraph id="1D59A4D9D4F5FBEBD1E45F6C320390B0" pageNumber="19">
|
|
<keyLead id="2D3C352188767AE7068A3965ABAB58D1">- Propodeal spiracle situated away from the metapleural gland orifice, the distance between them much greater than the propodeal spiracle diameter (Fig. 3) ....................................................................................................................... 9</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="0B11765949F3EA54C79E986B8B6F000A">
|
|
<paragraph id="4B6881BCA02F426544F7C41EF30C60B1" pageNumber="19">
|
|
<keyLead id="45AEFBAE7FA3F3B0DC3DD75D39D409F7">
|
|
6 Clypeus smooth and shining without longitudinal rugulae (Fig. 1).....................................................................
|
|
<taxonomicName id="11FA918D7F15CDC440E4A52AD5C998B8" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="inflata">inflata</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="138C6940A08267AAA47A9A0BCDFE4490" pageNumber="19">
|
|
<keyLead id="1527C877DABC5B2BDC6ABED5ABC77EC2">- Clypeus striated with rugulae (Fig. 2)..........................................................................................................................7</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="310ACD943483803500FFECCE5AE7E532">
|
|
<paragraph id="78838EBDD083B77B30863D7B1731301C" pageNumber="19">
|
|
<keyLead id="4415B39D5EF28556BB91683611D01528">
|
|
7 Dorsal face of head strongly striated with abundant longitudinal rugulae. Scape not reaching posterior corner of head in large workers. Workers with pronounced size polymorphism..........................................................................
|
|
<taxonomicName id="C8EA414975892E33D5A8C0E052817E5B" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="aurita">aurita</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="41FFB21E127AD8A052DF5CCA800DC1CC" pageNumber="19">
|
|
<keyLead id="88423D7984C7F91983FA0B0594D1D3DB">- Dorsal face of head not strongly striated with longitudinal rugulae. Scape reaching posterior corner of head. Workers monomorphic in size..................................................................................................................................................... 8</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="91356BD0F0F95E19E1B3B8DF4C0F2E67">
|
|
<paragraph id="4BF1B83752492BD023DA2FBCEE6E3649" pageNumber="19">
|
|
<keyLead id="2D845F1A61C2E7747D9041ADE46CB2AE">
|
|
8 The posterior portion of pronotum with transverse rugulae in dorsal view (Fig. 6)..................................
|
|
<taxonomicName id="7773EAD77B8585FCB69656FD7BFC965C" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="physothorax">physothorax</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="715B9D00FAD79B256DF1D366B492ABB5" pageNumber="19">
|
|
<keyLead id="317333439FAB5E3D568FA0B874CCF9A4">
|
|
- The posterior portion of pronotum without distinct rugulae in dorsal view (Fig. 7)..........................................
|
|
<taxonomicName id="545BBC8DA1FFC12CF03334A1A58393C4" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="onusta">onusta</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="F000B51DED546BA00ACD023115777177">
|
|
<paragraph id="AEC82D64356109330905F9C8238ED91D" pageNumber="19">
|
|
<keyLead id="216BDF1608E672DB83DD9A2AFFC5F942">
|
|
9 Dorsal setae on posterior portion of fourth abdominal tergite directed medially (Fig. 8)................................
|
|
<taxonomicName id="26E68F5183F551F557B3B0B10BC09CEE" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="difformis">difformis</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="D445D40467F1A7B5E2C4583E7A8D301D" pageNumber="19">
|
|
<keyLead id="F8987FFBE45BBD403BF6E1723C48AB2A">- Dorsal setae on fourth abdominal tergite directed posteriorly (Fig. 9).......................................................................10</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
<keyStep id="32E74BA8EE1CF9DABFD43DDD9E232518">
|
|
<paragraph id="B019B94CAD9BBEC8A38E5A1BA60F47CB" pageNumber="19">
|
|
<keyLead id="0073DAFEBF9CF979073874D54A47BB75">
|
|
10 In lateral view, dorsal profile of promesonotum convex, propodeum distinctly raised relative to promesonotum, metanotal groove very deep................................................................................................................................
|
|
<taxonomicName id="20E5E7B1A1C1D7C026FDD96A186E7C16" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" pageNumber="19" phylum="Arthropoda" rank="species" species="sewardi">sewardi</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
<paragraph id="1A29443B46E1BDCA70F621DDF6D1A62F" pageNumber="19">
|
|
<keyLead id="BCC7CD7F4BE41F29BC4E885551BB4496">
|
|
- In lateral view, dorsal profile of promesonotum convex, propodeum slightly raised relative to promesonotum, metanotal groove not so deep in dorsal median portion................................................................................
|
|
<taxonomicName id="476365199B9E943B54B28A76809C543C" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ampullaris">ampullaris</taxonomicName>
|
|
</keyLead>
|
|
</paragraph>
|
|
</keyStep>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |