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<document id="927CF5D6B64AB59DFB8ABB35D09F9332" ID-DOI="10.1080/00222930802124057" ID-ISSN="1464-5262" ID-Zenodo-Dep="4593937" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615397885932" checkinUser="carolina" docAuthor="Chatzimanolis, Stylianos" docDate="2008" docId="5C09879C776F1C187671FD715A086F78" docLanguage="en" docName="JNATHIST.42.25-26.1765-1792.pdf" docOrigin="Journal of Natural History 42 (25 - 26)" docSource="http://dx.doi.org/10.1080/00222930802124057" docStyle="DocumentStyle:FCA295E821436EF9307F50C7AC261B53.5:JNatHist.2008-2012.journal_article.1cover" docStyleId="FCA295E821436EF9307F50C7AC261B53" docStyleName="JNatHist.2008-2012.journal_article.1cover" docStyleVersion="5" docTitle="Isanopus hinojosai Chatzimanolis 2008, sp. n." docType="treatment" docVersion="3" lastPageNumber="1781" masterDocId="A030FFE477601C09744FFFB0586B6958" masterDocTitle="A revision of the neotropical beetle genus Isanopus Sharp, 1876 (Coleoptera: Staphylinidae: Staphylinini)" masterLastPageNumber="1792" masterPageNumber="1765" pageNumber="1779" updateTime="1698882181256" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="9B6713555C07A0C009D5DBDBA8A7FE11" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A3187B5DD7F93A73DDA5EC8BEF6463C9">
<mods:title id="0657674E9533DF224E032E8D34657BAF">A revision of the neotropical beetle genus Isanopus Sharp, 1876 (Coleoptera: Staphylinidae: Staphylinini)</mods:title>
</mods:titleInfo>
<mods:name id="315655D3DD2A78FAF32729DA67DBFB3B" type="personal">
<mods:role id="8110580DB766FC7ABD8AADE5DC89CB76">
<mods:roleTerm id="D91BE79EC56BB9D858A74096E2BE9ECC">Author</mods:roleTerm>
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<mods:namePart id="51126CD01DD53173A2F114E1D70C2484">Chatzimanolis, Stylianos</mods:namePart>
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<mods:typeOfResource id="F3C28260B30909865004AC975CBFCB3C">text</mods:typeOfResource>
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<mods:title id="0B4B0A9B9EA8CE1A5794422A8BE7C339">Journal of Natural History</mods:title>
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<mods:part id="1D93234D4A57F57F8C8A5EEFEA343532">
<mods:date id="A765BCBDC0AF787E6EAD9B6D3E935F8B">2008</mods:date>
<mods:detail id="3F9F68AAF01CACC3C6E71EA339A7A1B4" type="pubDate">
<mods:number id="009642CC0A5D61B4070C0F59C1271771">2008-07-31</mods:number>
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<mods:detail id="7E4D45FC6A9F9212046DB0912D6915CA" type="volume">
<mods:number id="794FF9CD9E90EB161B145F33169ECB5D">42</mods:number>
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<mods:detail id="490BBCA6535621C65719B2F9666377E1" type="issue">
<mods:number id="C7ABAC8A687B979FC4AF6718B73D7B2C">25 - 26</mods:number>
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<mods:start id="02ED27AE5527CF3887FA5F9FF6C9E3F0">1765</mods:start>
<mods:end id="FD9F6031290E9F04C29131C1269E5F9B">1792</mods:end>
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<mods:url id="C73FE57A586FA0BD4D996588AD9D6A35">http://dx.doi.org/10.1080/00222930802124057</mods:url>
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<mods:classification id="CFA17758389959AF8DE564B64D0188F7">journal article</mods:classification>
<mods:identifier id="3185F5F25E46492832AAD1BE2F4D7AA3" type="DOI">10.1080/00222930802124057</mods:identifier>
<mods:identifier id="7A62BE7BE35F129CFD3C526FD9B000B3" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="55CE0DAC4803DDED3D431CFE8676AB19" type="Zenodo-Dep">4593937</mods:identifier>
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<treatment id="5C09879C776F1C187671FD715A086F78" LSID="urn:lsid:plazi:treatment:5C09879C776F1C187671FD715A086F78" httpUri="http://treatment.plazi.org/id/5C09879C776F1C187671FD715A086F78" lastPageId="17" lastPageNumber="1781" pageId="15" pageNumber="1779">
<subSubSection id="9CBA6501776F1C067671FD715B396B81" box="[574,850,705,729]" pageId="15" pageNumber="1779" type="nomenclature">
<paragraph id="D41F368A776F1C067671FD715B396B81" blockId="15.[529,895,705,761]" box="[574,850,705,729]" pageId="15" pageNumber="1779">
<heading id="8F5781E6776F1C067671FD715B396B81" box="[574,850,705,729]" centered="true" fontSize="10" level="2" pageId="15" pageNumber="1779" reason="3">
<taxonomicName id="13A04D09776F1C067671FD715B616B81" ID-CoL="3Q34R" authority="Chatzimanolis, 2008" authorityName="Chatzimanolis" authorityYear="2008" box="[574,778,705,729]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1779" phylum="Arthropoda" rank="species" species="hinojosai" status="sp. nov.">
<emphasis id="E6D4EA98776F1C067671FD715B616B81" bold="true" box="[574,778,705,729]" italics="true" pageId="15" pageNumber="1779">Isanopus hinojosai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDE757E3776F1C06775CFD715B396B81" box="[787,850,705,729]" pageId="15" pageNumber="1779" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9CBA6501776F1C18765EFD515A086F78" lastPageId="17" lastPageNumber="1781" pageId="15" pageNumber="1779" type="description">
<paragraph id="D41F368A776F1C06765EFD515B146BA1" blockId="15.[529,895,705,761]" box="[529,895,737,761]" pageId="15" pageNumber="1779">
(
<figureCitation id="4C9B2A0F776F1C067657FD515AFD6BA1" box="[536,662,737,761]" captionStart="Figures 2" captionStartId="6.[146,222,1349,1371]" captionTargetBox="[149,1102,175,1302]" captionTargetId="figure-63@6.[146,1104,165,1312]" captionTargetPageId="6" captionText="Figures 2. Isanopus species, dorsum of head and pronotum. (A) I. ashei. (B) I. sahlbergi. (C) I. hinojosai. (D) I. eptaskouros." figureDoi="http://doi.org/10.5281/zenodo.4593941" httpUri="https://zenodo.org/record/4593941/files/figure.png" pageId="15" pageNumber="1779">Figures 2C</figureCitation>
,
<figureCitation id="4C9B2A0F776F1C0676EBFD515A9C6BA1" box="[676,759,737,761]" captionStart="Figure 3" captionStartId="8.[146,213,1346,1368]" captionTargetBox="[146,1104,165,1309]" captionTargetId="figure-86@8.[146,1104,165,1309]" captionTargetPageId="8" captionText="Figure 3. Isanopus hinojosai. (A) Left mandible, ventral aspect. (B) Right mandible, dorsal aspect, scale for 7–850.25 mm. (C) Abdominal sterna VII–VIII, scale50.81 mm. (D) Labium and hypopharynx, scale50.32 mm. (E) Left maxillae, ventral aspect, scale50.36 mm." figureDoi="http://doi.org/10.5281/zenodo.4593943" httpUri="https://zenodo.org/record/4593943/files/figure.png" pageId="15" pageNumber="1779">3A, 3E</figureCitation>
, 4E, 7A–C)
</paragraph>
<paragraph id="D41F368A776F1C06749AFC8259296A12" blockId="15.[213,322,818,842]" box="[213,322,818,842]" pageId="15" pageNumber="1779">
<emphasis id="E6D4EA98776F1C06749AFC8259296A12" box="[213,322,818,842]" italics="true" pageId="15" pageNumber="1779">Diagnosis</emphasis>
</paragraph>
<paragraph id="D41F368A776F1C06749AFCEA5C646A8A" blockId="15.[213,1169,858,978]" pageId="15" pageNumber="1779">
This species along with
<taxonomicName id="13A04D09776F1C0675A8FCEB5A526A2B" authorityName="Chatzimanolis" authorityYear="2008" box="[487,569,859,883]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1779" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="E6D4EA98776F1C0675A8FCEB5A526A2B" box="[487,569,859,883]" italics="true" pageId="15" pageNumber="1779">I. ashei</emphasis>
</taxonomicName>
and
<taxonomicName id="13A04D09776F1C06763BFCEB5ABF6A2B" authorityName="Sharp" authorityYear="1884" box="[628,724,859,883]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1779" phylum="Arthropoda" rank="species" species="sallaei">
<emphasis id="E6D4EA98776F1C06763BFCEB5ABF6A2B" box="[628,724,859,883]" italics="true" pageId="15" pageNumber="1779">I. sallaei</emphasis>
</taxonomicName>
has fewer and less dense punctures on the head and pronotum than other species of
<taxonomicName id="13A04D09776F1C067690FCCB5B2B6ACB" authorityName="Sharp" authorityYear="1876" box="[735,832,891,915]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1779" phylum="Arthropoda" rank="genus">
<emphasis id="E6D4EA98776F1C067690FCCB5B2B6ACB" box="[735,832,891,915]" italics="true" pageId="15" pageNumber="1779">Isanopus</emphasis>
</taxonomicName>
. It can be distinguished from all other species in the genus by the gradient colouration of the abdomen (see colouration section below) and the distinctive aedeagus (
<figureCitation id="4C9B2A0F776F1C06772DFC0A5BA16A8A" box="[866,970,954,978]" captionStart="Figure 7" captionStartId="16.[146,213,747,769]" captionTargetBox="[146,1099,165,690]" captionTargetId="figure-243@16.[146,1104,165,711]" captionTargetPageId="16" captionText="Figure 7. Aedeagus of I. hinojosai. (A) Dorsal view. (B) Lateral view. (C) Detail of paramere, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4593949" httpUri="https://zenodo.org/record/4593949/files/figure.png" pageId="15" pageNumber="1779">Figures 7</figureCitation>
A–C).
</paragraph>
<paragraph id="D41F368A776F1C06749AFBBB593F6D7B" blockId="15.[213,340,1035,1059]" box="[213,340,1035,1059]" pageId="15" pageNumber="1779">
<emphasis id="E6D4EA98776F1C06749AFBBB593F6D7B" box="[213,340,1035,1059]" italics="true" pageId="15" pageNumber="1779">Description</emphasis>
</paragraph>
<paragraph id="D41F368A776F1C06749AFB8359D66D13" blockId="15.[213,445,1075,1099]" box="[213,445,1075,1099]" pageId="15" pageNumber="1779">
Length
<quantity id="13589B6F776F1C067561FB8359D26D13" box="[302,441,1075,1099]" metricMagnitude="-2" metricUnit="m" metricValue="1.27" metricValueMax="1.34" metricValueMin="1.2" pageId="15" pageNumber="1779" unit="mm" value="12.7" valueMax="13.4" valueMin="12.0">12–13.4 mm</quantity>
.
</paragraph>
<paragraph id="D41F368A776F1C06749AFB34594D6C63" blockId="15.[213,1169,1155,1339]" pageId="15" pageNumber="1779">
<emphasis id="E6D4EA98776F1C06749AFB3459326DC4" box="[213,345,1156,1180]" italics="true" pageId="15" pageNumber="1779">Colouration</emphasis>
. Colour of head and pronotum brown to black with greenish overtones. Antenna yellow, except colour of antennomeres 1–3, dark yellow to brown. Scutellum dark brown to black. Elytra dark metallic blue, or purplish blue. Abdomen with gradient dark brown to yellow, usually segments II–IV dark brown, V–VI brown to dark yellow, VII–IX yellow. Legs dark brown except tarsi light brown.
</paragraph>
<paragraph id="D41F368A776F1C19749AFAC359146D7E" blockId="15.[213,1169,1395,1706]" lastBlockId="16.[146,1103,878,1062]" lastPageId="16" lastPageNumber="1780" pageId="15" pageNumber="1779">
<emphasis id="E6D4EA98776F1C06749AFAC3597A6CD3" box="[213,273,1395,1419]" italics="true" pageId="15" pageNumber="1779">Head</emphasis>
. Head with relatively small brown microsetae and longer black macrosetae along borders of head; head slightly transverse, width: length ratio51.28. Surface of epicranium glossy, with very few micropunctures distributed at centre. Epicranium flat, with sparse medium sized umbilicate setose punctures laterally, no punctures medially, setose punctures irregularly arranged in 3–4 rows anterior-posteriorly, punctures smaller around eyes, distance between punctures various, 0.5–1 times width of punctures. Eyes medium, length of eyes/length of head50.49, distance between eyes as wide as 1.8 times length of eye. Postgena with large, longitudinally elongate punctures (about four punctures/
<quantity id="13589B6F776F1C0676B2F9C25B386FD2" box="[765,851,1650,1674]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="1779" unit="mm" value="0.5">0.5 mm</quantity>
), punctures not uniformly distributed. Antennomeres with long macrosetae, antennomere 1 gradually club-like thickened, antennomeres 1–11 longer than wide, antennomere 1 about twice as long as antennomere 2, antennomere 3 1.5 times longer than antennomere 2, antennomere 4 slightly shorter than antennomere 3, antennomeres 5–10 almost identical, 3.5 times longer than wide, antennomere 11 subequal to previous antennomeres. Neck dorsally with no punctures or micropunctures, laterally with 3–4 small punctures.
</paragraph>
<caption id="80DF660277701C1974DDFD5B59646A47" ID-DOI="http://doi.org/10.5281/zenodo.4593949" ID-Zenodo-Dep="4593949" httpUri="https://zenodo.org/record/4593949/files/figure.png" pageId="16" pageNumber="1780" startId="16.[146,213,747,769]" targetBox="[146,1099,165,690]" targetPageId="16">
<paragraph id="D41F368A77701C1974DDFD5B59646A47" blockId="16.[146,1103,747,799]" pageId="16" pageNumber="1780">
Figure 7. Aedeagus of
<taxonomicName id="13A04D0977701C1975CFFD5C59856A59" authorityName="Chatzimanolis" authorityYear="2008" box="[384,494,748,769]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="1780" phylum="Arthropoda" rank="species" species="hinojosai">
<emphasis id="E6D4EA9877701C1975CFFD5C59856A59" box="[384,494,748,769]" italics="true" pageId="16" pageNumber="1780">I. hinojosai</emphasis>
</taxonomicName>
. (A) Dorsal view. (B) Lateral view. (C) Detail of paramere, dorsal view.
</paragraph>
</caption>
<paragraph id="D41F368A77701C1974DDFBEF59346F41" blockId="16.[146,1103,1119,1561]" pageId="16" pageNumber="1780">
<emphasis id="E6D4EA9877701C1974DDFBEF588F6D2F" box="[146,228,1119,1143]" italics="true" pageId="16" pageNumber="1780">Thorax</emphasis>
. Pronotum longer than wide, width: length ratio50.89; antero-lateral angles obtusely rounded, curved. Lateral margins of pronotum concave in dorsal aspect, pronotum broadest in apical 1/4 and narrower at basal angles. Surface of pronotum flat, glossy, with no micropunctures. Setose punctures absent on disc of pronotum, laterally present in 3–4 rows, distance between rows smaller towards lateral ends; punctures of medium size (about 4 punctures/
<quantity id="13589B6F77701C19770AFAB05BF66C40" box="[837,925,1280,1304]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="1780" unit="mm" value="0.5">0.5 mm</quantity>
); least distance between punctures for each row various, 1–2 times width of puncture for most. Pronotum with brown microsetae directed postero-medially in lateral and anterior areas and posteriorly in basal half, and few longer black macrosetae along borders. Scutellum prominent, with brown microsetae and sparse punctures, least distance between punctures 1–1.5 times width of puncture. Elytra subequal in length to pronotum, with medium setose uniform punctures (about six punctures/
<quantity id="13589B6F77701C1977A4FA715C296C81" box="[1003,1090,1473,1497]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="1780" unit="mm" value="0.5">0.5 mm</quantity>
), covered with yellowish brown microsetae and longer black macrosetae along borders of elytra.
</paragraph>
<paragraph id="D41F368A77701C1974DDF9E2594F6FF2" blockId="16.[146,1103,1618,1706]" pageId="16" pageNumber="1780">
<emphasis id="E6D4EA9877701C1974DDF9E258AC6F32" box="[146,199,1618,1642]" italics="true" pageId="16" pageNumber="1780">Legs</emphasis>
. Protarsus enlarged; meso and metatarsi elongate, enlarged, tarsomeres 1–4 asymmetrical, dilated distally. Legs almost completely covered with long yellow and brown setae.
</paragraph>
<paragraph id="D41F368A77711C18749AFF195C4568FB" blockId="17.[213,1169,168,419]" pageId="17" pageNumber="1781">
<emphasis id="E6D4EA9877711C18749AFF1959566999" box="[213,317,169,193]" italics="true" pageId="17" pageNumber="1781">Abdomen</emphasis>
. Abdominal terga III–V with tergal basal and no sub-basal (arch-like) carina. Terga III–VII with uniform punctuation patterns, densely covered with punctures and yellowish brown setae, punctures decreasing in size gradually from III–VII. Abdominal sterna III–IV with moderate transverse impressions, V with faint transverse impression, shining. Integument surface appearing slightly iridescent in strong direct light at moderate to high magnification. Sterna III–VII densely punctuated, punctures decreasing in size gradually from III–VII, with yellowish brown setae. Lateral tergal sclerites of the abdominal segment IX long and straight, covered with long black macrosetae.
</paragraph>
<paragraph id="D41F368A77711C18749AFE6C59F06B0D" blockId="17.[213,1168,476,597]" pageId="17" pageNumber="1781">
<emphasis id="E6D4EA9877711C18749AFE6C5A4B68AC" box="[213,544,476,500]" italics="true" pageId="17" pageNumber="1781">Secondary sexual structures</emphasis>
. Male: distal margin of sternum VII with slight emargination medially; sternum VIII with deep V-shaped emargination medially (
<figureCitation id="4C9B2A0F77711C187493FDAD59246B6D" box="[220,335,541,565]" captionStart="Figure 3" captionStartId="8.[146,213,1346,1368]" captionTargetBox="[146,1104,165,1309]" captionTargetId="figure-86@8.[146,1104,165,1309]" captionTargetPageId="8" captionText="Figure 3. Isanopus hinojosai. (A) Left mandible, ventral aspect. (B) Right mandible, dorsal aspect, scale for 7–850.25 mm. (C) Abdominal sterna VII–VIII, scale50.81 mm. (D) Labium and hypopharynx, scale50.32 mm. (E) Left maxillae, ventral aspect, scale50.36 mm." figureDoi="http://doi.org/10.5281/zenodo.4593943" httpUri="https://zenodo.org/record/4593943/files/figure.png" pageId="17" pageNumber="1781">Figure 3C</figureCitation>
); sternum IX with small emargination medially. Females: with no obvious sexual structures.
</paragraph>
<paragraph id="D41F368A77711C18749AFD3E5C256DB5" blockId="17.[213,1170,654,1262]" pageId="17" pageNumber="1781">
<emphasis id="E6D4EA9877711C18749AFD3E59546BFE" box="[213,319,654,678]" italics="true" pageId="17" pageNumber="1781">Aedeagus</emphasis>
. As in
<figureCitation id="4C9B2A0F77711C1875D7FD3E59976BFE" box="[408,508,654,678]" captionStart="Figure 7" captionStartId="16.[146,213,747,769]" captionTargetBox="[146,1099,165,690]" captionTargetId="figure-243@16.[146,1104,165,711]" captionTargetPageId="16" captionText="Figure 7. Aedeagus of I. hinojosai. (A) Dorsal view. (B) Lateral view. (C) Detail of paramere, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4593949" httpUri="https://zenodo.org/record/4593949/files/figure.png" pageId="17" pageNumber="1781">Figures 7</figureCitation>
A–C; paramere extremely reduced; paramere: median lobe ratio50.34. Paramere in dorsal view fairly wide but expanding to two apical processes, with a wide U-shaped emargination between them; in lateral outline almost straight; with no apparent sensory spinules. Median lobe in dorsal view wide, converging to apex; in lateral view becoming narrower and curved from middle to apex.
<typeStatus id="0B1B882877711C1877FDFCA05C766A70" box="[946,1053,784,808]" pageId="17" pageNumber="1781" type="holotype">Holotype</typeStatus>
, male with labels as follows: ‘‘
<collectingCountry id="ACB7761A77711C1875EFFC805A256A10" box="[416,590,816,840]" name="Costa Rica" pageId="17" pageNumber="1781">COSTA RICA</collectingCountry>
:
<collectingRegion id="1664F86877711C187616FC805B716A10" box="[601,794,816,840]" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="1781">Puntarenas Prov.</collectingRegion>
Altamira Biol. Sta.,
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,
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09
<emphasis id="E6D4EA9877711C1874A1FCE058936A30" bold="true" box="[238,248,848,872]" italics="true" pageId="17" pageNumber="1781">°</emphasis>
01.769N
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,
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83
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00.499W
</geoCoordinate>
, vi.7.2004, J.S. Ashe, Z. Falin, I. Hinojosa, ex: treefall litter’’/ [barcode label] ‘‘SM0607654’’/‘‘
<typeStatus id="0B1B882877711C187671FCC15ACC6AD1" box="[574,679,881,905]" pageId="17" pageNumber="1781" type="holotype">Holotype</typeStatus>
<taxonomicName id="13A04D0977711C1876FAFCC15C596AD1" authority="Chatzimanolis" authorityName="Chatzimanolis" authorityYear="2008" box="[693,1074,881,905]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1781" phylum="Arthropoda" rank="species" species="hinojosai">
<emphasis id="E6D4EA9877711C1876FAFCC15BEA6AD1" box="[693,897,881,905]" italics="true" pageId="17" pageNumber="1781">Isanopus hinojosai</emphasis>
Chatzimanolis
</taxonomicName>
, design. Chatzimanolis 2007’’. In the collection of INBio, Costa Rica. Note that the
<typeStatus id="0B1B882877711C187057FC215C126AF1" box="[1048,1145,913,937]" pageId="17" pageNumber="1781" type="holotype">holotype</typeStatus>
is currently housed in Snow Entomological Collection, Division of Entomology, Natural History Museum/BRC, University of Kansas, Lawrence but based on a collecting agreement between the two institution the
<typeStatus id="0B1B882877711C18768FFC425B4A6D52" box="[704,801,1010,1034]" pageId="17" pageNumber="1781" type="holotype">holotype</typeStatus>
would be transferred to INBio.
<typeStatus id="0B1B882877711C18749AFBA3592E6D73" box="[213,325,1043,1067]" pageId="17" pageNumber="1781" type="paratype">Paratypes</typeStatus>
: 13, same locality label as above, barcode labels SM0607656, SM0607655, SM0607657, SM0607557, SM0607653 (
<quantity id="13589B6F77711C1876DDFB835AD56D13" box="[658,702,1075,1099]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="17" pageNumber="1781" unit="in" value="5.0">5 in</quantity>
SEMC), SM0607658 (
<quantity id="13589B6F77711C187788FB835B9A6D13" box="[967,1009,1075,1099]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="17" pageNumber="1781" unit="in" value="1.0">1 in</quantity>
UTCI); same data except vi.6.2004, SM0607670 (
<quantity id="13589B6F77711C1876C3FBE45AD46D34" box="[652,703,1108,1132]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="17" pageNumber="1781" unit="in" value="1.0">1 in</quantity>
SEMC); same data except vi.5.2004, SM0607584, SM0607587, SM0607586, SM0607588, SM0607585 (
<quantity id="13589B6F77711C1877A5FBC45C4A6DD4" box="[1002,1057,1140,1164]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="17" pageNumber="1781" unit="in" value="5.0">5 in</quantity>
SEMC);
<collectingCountry id="ACB7761A77711C18749AFB24593F6DF4" box="[213,340,1172,1196]" name="Panama" pageId="17" pageNumber="1781">PANAMA</collectingCountry>
:
<collectingRegion id="1664F86877711C18752BFB2459A86DF4" box="[356,451,1172,1196]" country="Panama" name="Chiriqui" pageId="17" pageNumber="1781">Chiriqui</collectingRegion>
, Finca La Suisa,
<quantity id="13589B6F77711C1876D6FB245AB06DF4" box="[665,731,1172,1196]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="17" pageNumber="1781" unit="km" value="24.0">24 km</quantity>
S Gualaca, 8
<emphasis id="E6D4EA9877711C1877CFFB255BE16DF5" bold="true" box="[896,906,1173,1197]" italics="true" pageId="17" pageNumber="1781">°</emphasis>
349N 82
<emphasis id="E6D4EA9877711C1877A6FB255B986DF5" bold="true" box="[1001,1011,1173,1197]" italics="true" pageId="17" pageNumber="1781">°</emphasis>
129W,
<quantity id="13589B6F77711C18700CFB245CFA6DF4" box="[1091,1169,1172,1196]" metricMagnitude="3" metricUnit="m" metricValue="1.285" pageId="17" pageNumber="1781" unit="m" value="1285.0">1285 m</quantity>
v.31.2000, HandA Howden, barcode label SM0235827 (
<quantity id="13589B6F77711C187775FB055B0A6D95" box="[826,865,1205,1229]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="17" pageNumber="1781" unit="in" value="1.0">1 in</quantity>
SEMC). All
<typeStatus id="0B1B882877711C1877A0FB055C316D95" box="[1007,1114,1205,1229]" pageId="17" pageNumber="1781" type="paratype">paratypes</typeStatus>
with label: ‘‘
<typeStatus id="0B1B882877711C18756EFB6559EC6DB5" box="[289,391,1237,1261]" pageId="17" pageNumber="1781" type="paratype">Paratype</typeStatus>
<taxonomicName id="13A04D0977711C1875C0FB665A956DB5" authority="Chatzimanolis" authorityName="Chatzimanolis" authorityYear="2008" box="[399,766,1237,1262]" class="Insecta" family="Staphylinidae" genus="Isanopus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="1781" phylum="Arthropoda" rank="species" species="hinojosai">
<emphasis id="E6D4EA9877711C1875C0FB665A3E6DB6" box="[399,597,1238,1262]" italics="true" pageId="17" pageNumber="1781">Isanopus hinojosai</emphasis>
Chatzimanolis
</taxonomicName>
, design. Chatzimanolis 2007’’.
</paragraph>
<paragraph id="D41F368A77711C18749AFA965AC46C07" blockId="17.[213,1169,1318,1375]" pageId="17" pageNumber="1781">
<emphasis id="E6D4EA9877711C18749AFA9659256C66" box="[213,334,1318,1342]" italics="true" pageId="17" pageNumber="1781">Etymology</emphasis>
. This species is named in honor of Ismael Hinojosa-Diaz, who collected many of the specimens of the
<typeStatus id="0B1B882877711C18767FFAF75A0B6C07" box="[560,608,1351,1375]" pageId="17" pageNumber="1781">type</typeStatus>
series.
</paragraph>
<paragraph id="D41F368A77711C18749AFA28593C6C88" blockId="17.[213,1168,1431,1488]" pageId="17" pageNumber="1781">
<emphasis id="E6D4EA9877711C18749AFA2859306CE8" box="[213,347,1432,1456]" italics="true" pageId="17" pageNumber="1781">Distribution</emphasis>
. Known from elevations of
<quantity id="13589B6F77711C1876FEFA275B286CF7" box="[689,835,1431,1455]" metricMagnitude="3" metricUnit="m" metricValue="1.4425" metricValueMax="1.6" metricValueMin="1.285" pageId="17" pageNumber="1781" unit="m" value="1442.5" valueMax="1600.0" valueMin="1285.0">1285–1600 m</quantity>
in
<collectingCountry id="ACB7761A77711C18773FFA275B986CF7" box="[880,1011,1431,1455]" name="Costa Rica" pageId="17" pageNumber="1781">Costa Rica</collectingCountry>
and
<collectingCountry id="ACB7761A77711C18707BFA275CFB6CF7" box="[1076,1168,1431,1455]" name="Panama" pageId="17" pageNumber="1781">Panama</collectingCountry>
(
<figureCitation id="4C9B2A0F77711C187493FA0859206C88" box="[220,331,1464,1488]" captionStart="Figure 10" captionStartId="23.[213,280,856,878]" captionTargetBox="[213,1171,165,820]" captionTargetId="figure-204@23.[213,1171,165,820]" captionTargetPageId="23" captionText="Figure 10. Distribution map of I. ashei, I. hinojosai and I. sallaei." figureDoi="http://doi.org/10.5281/zenodo.4593955" httpUri="https://zenodo.org/record/4593955/files/figure.png" pageId="17" pageNumber="1781">Figure 10</figureCitation>
).
</paragraph>
<paragraph id="D41F368A77711C18749AF9B95A086F78" blockId="17.[213,611,1544,1569]" box="[213,611,1544,1569]" pageId="17" pageNumber="1781">
<emphasis id="E6D4EA9877711C18749AF9B959476F79" box="[213,300,1545,1569]" italics="true" pageId="17" pageNumber="1781">Habitat</emphasis>
. Collected in treefall litter.
</paragraph>
</subSubSection>
</treatment>
</document>
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