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<document id="199B31F9C46E37A505CE166B00AC222A" ID-CLB-Dataset="24248" ID-DOI="10.5252/zoosystema2020v42a1" ID-GBIF-Dataset="f3328b3e-6059-40b6-b7f5-1c19aa90f676" ID-ISSN="1638-9387" ID-Zenodo-Dep="3627057" ID-ZooBank="urn:lsid:zoobank.org:pub:D14C124B-4CB6-4097-95F9-CACECBE4C844" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1579878451081" checkinUser="plazi" docAuthor="Barrientos, José Antonio, Brañas, Neus &amp; Mederos, Jorge" docDate="2020" docId="03C5C9567078FFCFFC34FF750718F833" docLanguage="en" docName="Zoosystema.42.1.1-29.pdf" docOrigin="Zoosystema 42 (1)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Lepthyphantes fadriquei Barrientos 2020, n. sp." docType="treatment" docVersion="9" lastPageNumber="11" masterDocId="FFFCB12E7072FFC3FFBBFFAD067AFFDE" masterDocTitle="The high complexity of Micronetinae Hull, 1920 (Araneae, Linyphiidae) evidenced through ten new cave-dweller species from the Morocco" masterLastPageNumber="29" masterPageNumber="1" pageNumber="9" updateTime="1698779390546" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FC47B49C5B5F0210BC93A5C5EB845B46">The high complexity of Micronetinae Hull, 1920 (Araneae, Linyphiidae) evidenced through ten new cave-dweller species from the Morocco</mods:title>
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<mods:namePart id="0E114A9707D978191430D15A3F2E41C2">Barrientos, José Antonio</mods:namePart>
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<mods:namePart id="80CF2B446E9403C4D27D048A133C9067">Brañas, Neus</mods:namePart>
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<mods:namePart id="C5EA2BDB2D4A74F735651AE24FA9B8A8">Mederos, Jorge</mods:namePart>
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<mods:typeOfResource id="7AB6E90B5C98F75092E61D9978C25D77">text</mods:typeOfResource>
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<mods:title id="73220690962CD4B2B75AFB189342C7BC">Zoosystema</mods:title>
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<mods:part id="84B3C592ACE4BE27192C6D23A168DFB2">
<mods:date id="3939C8CF1D4366801FC22B01764ECE9F">2020</mods:date>
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<mods:number id="0C622861FE5004375CD42FF9BEAB95DC">2020-01-23</mods:number>
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<mods:number id="62C2B08607EA7CDF15D6CE9F8C4AF490">42</mods:number>
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<mods:identifier id="A8FC7E9F3FC0BB1F070D1F93D8F8E69F" type="CLB-Dataset">24248</mods:identifier>
<mods:identifier id="509B65C960B3DDC07F982DA2EC5021C4" type="DOI">10.5252/zoosystema2020v42a1</mods:identifier>
<mods:identifier id="6FB9A8C751D30775C99B4EEF15309E7D" type="GBIF-Dataset">f3328b3e-6059-40b6-b7f5-1c19aa90f676</mods:identifier>
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<treatment id="03C5C9567078FFCFFC34FF750718F833" ID-DOI="http://doi.org/10.5281/zenodo.4327717" ID-GBIF-Taxon="161838936" ID-Zenodo-Dep="4327717" LSID="urn:lsid:plazi:treatment:03C5C9567078FFCFFC34FF750718F833" httpUri="http://treatment.plazi.org/id/03C5C9567078FFCFFC34FF750718F833" lastPageId="12" lastPageNumber="11" pageId="10" pageNumber="9">
<subSubSection id="C3762BCB7078FFC9FC34FF750337FF2A" box="[911,1357,216,244]" pageId="10" pageNumber="9" type="nomenclature">
<paragraph id="8BD378407078FFC9FC34FF750337FF2A" blockId="10.[911,1357,216,276]" box="[911,1357,216,244]" pageId="10" pageNumber="9">
<heading id="D09BCF2C7078FFC9FC34FF750337FF2A" box="[911,1357,216,244]" centered="true" fontSize="11" level="4" pageId="10" pageNumber="9" reason="4">
<taxonomicName id="4C6C03C37078FFC9FC34FF750371FF2D" ID-CoL="98ZX8" authority="Barrientos" authorityName="Barrientos" authorityYear="2020" box="[911,1291,216,243]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="fadriquei" status="sp. nov.">
<emphasis id="B918A4527078FFC9FC34FF7502EFFF2D" bold="true" box="[911,1173,216,243]" italics="true" pageId="10" pageNumber="9">Lepthyphantes fadriquei</emphasis>
Barrientos
</taxonomicName>
<taxonomicNameLabel id="A22B19297078FFC9FAA9FF770337FF2A" box="[1298,1357,218,244]" pageId="10" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3762BCB7078FFC9FBA2FF5402B9FECA" box="[1049,1219,249,276]" pageId="10" pageNumber="9" type="description">
<paragraph id="8BD378407078FFC9FBA2FF5402B9FECA" blockId="10.[911,1357,216,276]" box="[1049,1219,249,276]" pageId="10" pageNumber="9">
<heading id="D09BCF2C7078FFC9FBA2FF5402B9FECA" box="[1049,1219,249,276]" centered="true" fontSize="11" level="4" pageId="10" pageNumber="9" reason="1">
(
<figureCitation id="135764C57078FFC9FB9BFF540218FECA" box="[1056,1122,249,276]" captionStart="FIG" captionStartId="11.[133,144,1454,1472]" captionTargetBox="[264,1311,225,1398]" captionTargetId="figure@11.[246,1342,215,1406]" captionTargetPageId="11" captionText="FIG. 5. — Lepthyphantes fadriquei Barrientos n. sp., genital organs; A, B, male copulatory bulb, retrolateral view (A), ventral view (B); C, schema of the embolic division; D, E, epigyne, ventral view (D); idem, lateral view (E). Abbreviations: see Material and methods. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3627067" httpUri="https://zenodo.org/record/3627067/files/figure.png" pageId="10" pageNumber="9">Figs 5</figureCitation>
;
<figureCitation id="135764C57078FFC9FBCBFF5402F3FECD" box="[1136,1161,249,275]" captionStart="FIG" captionStartId="23.[133,144,1789,1807]" captionTargetBox="[135,1455,215,1732]" captionTargetId="figure@23.[135,1458,210,1732]" captionTargetPageId="23" captionText="FIG. 13. — A, B, Epigyne of Lepthyphantes almoravidus Barrientos n. sp. in ventral (A) and lateral (B) views; C, D, epigyne of Lepthyphantes biospeleologorum Barrientos n. sp. in dorsal (C) and lateral (D) views; E, F, epigyne of Lepthyphantes fadriquei Barrientos n. sp. in lateral (E) and ventral (F) views; G, male copulatory bulb of L. fadriquei Barrientos n. sp. retrolateral views; H, I, epigyne of Lepthyphantes ensiferus Barrientos n. sp. in lateral (H) and dorsal (I) views; J, male copulatory bulb of Lepthyphantes ensiferus Barrientos n. sp. in retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.3627083" httpUri="https://zenodo.org/record/3627083/files/figure.png" pageId="10" pageNumber="9">13</figureCitation>
E-G)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3762BCB7078FFC9FCFFFE9B03E2FE96" box="[836,1432,310,328]" pageId="10" pageNumber="9" type="nomenclature">
<paragraph id="8BD378407078FFC9FCFFFE9B03E2FE96" blockId="10.[836,1432,310,328]" box="[836,1432,310,328]" pageId="10" pageNumber="9">
<uri id="FFFD74427078FFC9FCFFFE9B03E2FE96" box="[836,1432,310,328]" pageId="10" pageNumber="9">
urn:lsid:zoobank.org:act:
<uuid id="FFCA42957078FFC9FBB0FE9B03E2FE96" box="[1035,1432,310,328]" pageId="10" pageNumber="9">64F9CBAA-7AE3-415D-89BE-5F842D0B23BD</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3762BCB7078FFC9FC96FEC5025DFDEA" pageId="10" pageNumber="9" type="materials_examined">
<paragraph id="8BD378407078FFC9FC96FEC50338FE45" blockId="10.[813,1456,360,490]" pageId="10" pageNumber="9">
<materialsCitation id="3B04721D7078FFC9FC96FEC50338FE45" ID-GBIF-Occurrence="2558291726" collectingDate="1970-08-28" collectionCode="TYPE" collectorName="O. Escola" country="Morocco" location="Morocco" pageId="10" pageNumber="9" specimenCode="MZB 73-2905" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54D7C6E27078FFC9FC96FEC50527FEA3" box="[813,861,360,384]" pageId="10" pageNumber="9">TYPE</typeStatus>
MATERIAL. —
<emphasis id="B918A4527078FFC9FC52FEC50232FE5E" bold="true" box="[1001,1096,360,384]" pageId="10" pageNumber="9">
<typeStatus id="54D7C6E27078FFC9FC52FEC50232FE5E" box="[1001,1096,360,384]" pageId="10" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
.
<emphasis id="B918A4527078FFC9FBE9FEC502D1FE5E" bold="true" box="[1106,1195,360,384]" pageId="10" pageNumber="9">
<collectingCountry id="F37B38D07078FFC9FBE9FEC502D1FE5E" box="[1106,1195,360,384]" name="Morocco" pageId="10" pageNumber="9">Morocco</collectingCountry>
</emphasis>
.
<specimenCount id="9D6AB3C97078FFC9FB0CFEC502A2FE5C" box="[1207,1240,360,386]" pageId="10" pageNumber="9" type="male">1♂</specimenCount>
; Kef Aziza, Boudenib, Meknès;
<date id="FFD25E807078FFC9FC3DFE2E0276FE45" box="[902,1036,387,411]" pageId="10" pageNumber="9" value="1970-08-28">
<collectingDate id="EF96A7687078FFC9FC3DFE2E0276FE45" box="[902,1036,387,411]" pageId="10" pageNumber="9" value="1970-08-28">28.VIII.1970</collectingDate>
</date>
;
<collectorName id="26991D967078FFC9FBADFE2E020EFE45" box="[1046,1140,387,411]" pageId="10" pageNumber="9">O. Escolá</collectorName>
leg.;
<specimenCode id="DBCAD03B7078FFC9FB12FE2E0347FE45" box="[1193,1341,387,411]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="9" type="Museum">MZB 73-2905</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD378407078FFC9FC96FE300240FE34" blockId="10.[813,1456,360,490]" pageId="10" pageNumber="9">
<materialsCitation id="3B04721D7078FFC9FC96FE3005C0FE11" ID-GBIF-Occurrence="2558291745" collectionCode="MZB" country="Morocco" location="Morocco" pageId="10" pageNumber="9" specimenCode="MZB 72-5577" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<emphasis id="B918A4527078FFC9FC96FE300589FE6B" bold="true" box="[813,1011,413,437]" pageId="10" pageNumber="9">
<typeStatus id="54D7C6E27078FFC9FC96FE3005F7FE6B" box="[813,909,413,437]" pageId="10" pageNumber="9" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F37B38D07078FFC9FC21FE300589FE6B" box="[922,1011,413,437]" name="Morocco" pageId="10" pageNumber="9">Morocco</collectingCountry>
</emphasis>
.
<specimenCount id="9D6AB3C97078FFC9FBB9FE300265FE69" box="[1026,1055,413,439]" pageId="10" pageNumber="9" type="female">3♀</specimenCount>
; same locality and date as the
<typeStatus id="54D7C6E27078FFC9FAE8FE3003D1FE6B" box="[1363,1451,413,437]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
;
<specimenCode id="DBCAD03B7078FFC9FC96FE1505C0FE11" box="[813,954,439,463]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="9" type="Museum">MZB 72-5577</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B04721D7078FFC9FC76FE1502D2FE0E" ID-GBIF-Occurrence="2558291795" box="[973,1192,440,465]" collectingDate="2. VI" collectionCode="VI" country="Morocco" pageId="10" pageNumber="9" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D6AB3C97078FFC9FC76FE150591FE0F" box="[973,1003,440,465]" pageId="10" pageNumber="9" type="female">1♀</specimenCount>
; same locality;
<date id="FFD25E807078FFC9FBC0FE1502D2FE0E" box="[1147,1192,440,464]" pageId="10" pageNumber="9" value="2001-06-02">
<collectingDate id="EF96A7687078FFC9FBC0FE1502D2FE0E" box="[1147,1192,440,464]" pageId="10" pageNumber="9">2.VI</collectingDate>
</date>
</materialsCitation>
<materialsCitation id="3B04721D7078FFC9FB13FE15024CFE34" ID-GBIF-Occurrence="2558291748" collectionCode="MZB" collectorName="F. Fadrique &amp; O. Escola" country="Morocco" pageId="10" pageNumber="9" specimenCode="MZB 2001-0414" specimenCount="1" typeStatus="paratype">
.2001;
<collectorName id="26991D967078FFC9FB51FE15032EFE0E" box="[1258,1364,440,464]" pageId="10" pageNumber="9">F. Fadrique</collectorName>
&amp;
<collectorName id="26991D967078FFC9FACAFE15052EFE34" pageId="10" pageNumber="9">O. Escolá</collectorName>
leg.;
<specimenCode id="DBCAD03B7078FFC9FC33FE7E024CFE34" box="[904,1078,466,490]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="9" type="Museum">MZB 2001-0414</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD378407078FFC9FC96FDAC025DFDEA" blockId="10.[813,1455,513,564]" pageId="10" pageNumber="9">
<materialsCitation id="3B04721D7078FFC9FC96FDAC03D5FDC4" ID-GBIF-Occurrence="2558291781" box="[813,1455,513,538]" collectionCode="OTHER" country="Morocco" location="Morocco" pageId="10" pageNumber="9" specimenCount="3" specimenCount-juvenile="3">
OTHER MATERIAL. —
<emphasis id="B918A4527078FFC9FC45FDAC022DFDC7" bold="true" box="[1022,1111,513,537]" pageId="10" pageNumber="9">
<collectingCountry id="F37B38D07078FFC9FC45FDAC022DFDC7" box="[1022,1111,513,537]" name="Morocco" pageId="10" pageNumber="9">Morocco</collectingCountry>
</emphasis>
.
<specimenCount id="9D6AB3C97078FFC9FBD8FDAF02E0FDC4" box="[1123,1178,514,538]" pageId="10" pageNumber="9" type="juvenile">3 juv.</specimenCount>
; same locality and date as the
</materialsCitation>
<materialsCitation id="3B04721D7078FFC9FC96FDB1025EFDEA" ID-GBIF-Occurrence="2558291723" box="[813,1060,540,564]" collectionCode="MZB" country="Morocco" pageId="10" pageNumber="9" specimenCode="MZB 72-5578" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D7C6E27078FFC9FC96FDB105FFFDEA" box="[813,901,540,564]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
;
<specimenCode id="DBCAD03B7078FFC9FC34FDB0025EFDEA" box="[911,1060,540,564]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="10" pageNumber="9" type="Museum">MZB 72-5578</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3762BCB7078FFC9FC96FDE60370FC2F" pageId="10" pageNumber="9" type="diagnosis">
<paragraph id="8BD378407078FFC9FC96FDE60370FC2F" blockId="10.[813,1456,587,1009]" pageId="10" pageNumber="9">
DIAGNOSIS. — In its posterior middle part, the paracymbium presents a very developed elongation, acute and curved towards the external side; its apical branch is narrow, straight and lanceolate. In the radix a sharp, spiny tip develops at its posterior end. The lamella characteristica consists of a long and narrow projection, which gradually sharpens towards the tip, maintaining a progressive curvature. The terminal apophysis, equally long and narrow, describes a curved path, parallel to the lamella characteristica, and somewhat longer. The epigyne is wide at its base and perpendicular to the surface; it narrows progressively distally where a proscape with an elliptical contour, flanked by the lateral walls, stands out. The set of the scape describes a structure in “S” (easily seen in side view). The morphology of the epigyne resembles that of
<taxonomicName id="4C6C03C37078FFC9FAAEFC27050AFC62" authority="Barrientos" authorityName="Barrientos" authorityYear="2020" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="ensiferus" status="sp. nov.">
<emphasis id="B918A4527078FFC9FAAEFC270305FC7F" box="[1301,1407,905,929]" italics="true" pageId="10" pageNumber="9">L. ensiferus</emphasis>
Barrientos
</taxonomicName>
<taxonomicNameLabel id="A22B19297078FFC9FCCEFC0905D1FC65" box="[885,939,932,955]" pageId="10" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
; however, the structure of the male genitalia separates the two species, which places
<taxonomicName id="4C6C03C37078FFC9FBEDFC12034CFC08" authority="Barrientos" authorityName="Barrientos" authorityYear="2020" box="[1110,1334,958,982]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="fadriquei" status="sp. nov.">
<emphasis id="B918A4527078FFC9FBEDFC1202B3FC08" box="[1110,1225,958,982]" italics="true" pageId="10" pageNumber="9">L. fadriquei</emphasis>
Barrientos
</taxonomicName>
<taxonomicNameLabel id="A22B19297078FFC9FA86FC12030EFC08" box="[1341,1396,959,982]" pageId="10" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
in an uncertain position in the group of
<taxonomicName id="4C6C03C37078FFC9FBC4FC74037CFC2F" authorityName="Hull" authorityYear="1920" box="[1151,1286,985,1009]" class="Arachnida" family="Linyphiidae" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="9" phylum="Arthropoda" rank="subFamily" subFamily="Micronetinae">Micronetinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3762BCB7078FFC9FC96FBA502B2FB54" pageId="10" pageNumber="9" type="etymology">
<paragraph id="8BD378407078FFC9FC96FBA502B2FB54" blockId="10.[813,1456,1032,1162]" pageId="10" pageNumber="9">
ETYMOLOGY. — We dedicate this species to Floren Fadrique, the main promoter of the latest biospeleological surveys in
<collectingCountry id="F37B38D07078FFC9FAEAFB8E03D6FBE4" box="[1361,1452,1059,1082]" name="Morocco" pageId="10" pageNumber="9">Morocco</collectingCountry>
; without his generosity, initiative and dedication on the ground (before, during and after each expedition) we would not have many of the interesting samples of this small study.
</paragraph>
</subSubSection>
<subSubSection id="C3762BCB7078FFC9FC96FB0C05B4FB31" pageId="10" pageNumber="9" type="distribution">
<paragraph id="8BD378407078FFC9FC96FB0C05B4FB31" blockId="10.[813,1455,1185,1263]" pageId="10" pageNumber="9">
DISTRIBUTION. —
<taxonomicName id="4C6C03C37078FFC9FC59FB0F02C3FB64" authority="Barrientos" authorityName="Barrientos" authorityYear="2020" box="[994,1209,1186,1210]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="fadriquei" status="sp. nov.">
<emphasis id="B918A4527078FFC9FC59FB0F022AFB64" box="[994,1104,1186,1210]" italics="true" pageId="10" pageNumber="9">L. fadriquei</emphasis>
Barrientos
</taxonomicName>
<taxonomicNameLabel id="A22B19297078FFC9FB06FB0F028BFB67" box="[1213,1265,1186,1209]" pageId="10" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
is known only from the
<typeStatus id="54D7C6E27078FFC9FCE8FB100507FB0B" box="[851,893,1213,1237]" pageId="10" pageNumber="9">type</typeStatus>
locality, a cavity of the province of Meknès, in the high Moroccan Atlas.
</paragraph>
</subSubSection>
<subSubSection id="C3762BCB7078FFCFFC96FAB80718F833" lastPageId="12" lastPageNumber="11" pageId="10" pageNumber="9" type="description">
<paragraph id="8BD378407078FFC9FC96FAB805C7FAF3" blockId="10.[813,1456,1301,1678]" box="[813,957,1301,1327]" pageId="10" pageNumber="9">DESCRIPTION</paragraph>
<paragraph id="8BD378407078FFC9FC96FA9805B4FA91" blockId="10.[813,1456,1301,1678]" box="[813,974,1333,1359]" pageId="10" pageNumber="9">
<emphasis id="B918A4527078FFC9FC96FA9805B4FA91" box="[813,974,1333,1359]" italics="true" pageId="10" pageNumber="9">
Male (
<typeStatus id="54D7C6E27078FFC9FCCBFA9805BDFA91" box="[880,967,1333,1359]" pageId="10" pageNumber="9" type="holotype">holotype</typeStatus>
)
</emphasis>
</paragraph>
<paragraph id="8BD378407078FFC9FC96FAF80264F950" blockId="10.[813,1456,1301,1678]" pageId="10" pageNumber="9">
<emphasis id="B918A4527078FFC9FC96FAF80517FAB1" bold="true" box="[813,877,1365,1391]" pageId="10" pageNumber="9">Body.</emphasis>
Carapace: 0.81 long, by 0.68 wide; coloration pale yellow uniform and without additional pigmentary pattern. Some hairs scattered in the cephalic area. Without eyes. Chelicerae: 0.39 (length of stem), with three teeth, short and separated on the promargin; unarmed retromargin. Legs of the same color as the carapace; relatively long and thin; measurements: leg I (1.32 + 0.25 + 1.42 + 1.32 + 0.78), leg II (1.22 + 0.22 + 1.25 + 1.13 + 0.64), leg III (1.13 + 0.20 + 0.81 + 0.91 + 0.59), and leg IV (1.27 + 0.25 + 1.08 + 1.18 + 0.64). Opisthosoma missing.
</paragraph>
<paragraph id="8BD378407078FFC9FC96F91E03D5F833" blockId="10.[813,1456,1715,2030]" pageId="10" pageNumber="9">
<emphasis id="B918A4527078FFC9FC96F91E0232F913" bold="true" box="[813,1096,1715,1741]" pageId="10" pageNumber="9">Palp and copulatory bulb</emphasis>
(
<figureCitation id="135764C57078FFC9FBE3F91902E3F911" box="[1112,1177,1716,1743]" captionStart="FIG" captionStartId="11.[133,144,1454,1472]" captionTargetBox="[264,1311,225,1398]" captionTargetId="figure@11.[246,1342,215,1406]" captionTargetPageId="11" captionText="FIG. 5. — Lepthyphantes fadriquei Barrientos n. sp., genital organs; A, B, male copulatory bulb, retrolateral view (A), ventral view (B); C, schema of the embolic division; D, E, epigyne, ventral view (D); idem, lateral view (E). Abbreviations: see Material and methods. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3627067" httpUri="https://zenodo.org/record/3627067/files/figure.png" pageId="10" pageNumber="9">Figs 5</figureCitation>
A-C; 13G). In the paracymbium, posterior middle part particularly well developed; prolonged by a long sharp projection and curved outwards (
<figureCitation id="135764C57078FFC9FC8DF8B905D9F8F0" box="[822,931,1811,1838]" captionStart="FIG" captionStartId="11.[133,144,1454,1472]" captionTargetBox="[264,1311,225,1398]" captionTargetId="figure@11.[246,1342,215,1406]" captionTargetPageId="11" captionText="FIG. 5. — Lepthyphantes fadriquei Barrientos n. sp., genital organs; A, B, male copulatory bulb, retrolateral view (A), ventral view (B); C, schema of the embolic division; D, E, epigyne, ventral view (D); idem, lateral view (E). Abbreviations: see Material and methods. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3627067" httpUri="https://zenodo.org/record/3627067/files/figure.png" pageId="10" pageNumber="9">Fig. 5A, B</figureCitation>
); proximal part wide and trapezoidal, with some hairs in its basal area; middle part marked by two protrusions or dilatations: one just mentioned in the posterior part and one in the anterior part, thick and blunt, delimiting the anterior pocket; from the anterior pocket, the apical branch narrow, straight, somewhat spatulate, and relatively short. Ventrally bulb showing on its prolateral side the basal structure of the
</paragraph>
<caption id="DF1328C87079FFC8FF3EFA0302FDFA08" ID-DOI="http://doi.org/10.5281/zenodo.3627067" ID-Zenodo-Dep="3627067" httpUri="https://zenodo.org/record/3627067/files/figure.png" pageId="11" pageNumber="10" startId="11.[133,144,1454,1472]" targetBox="[264,1311,225,1398]" targetPageId="11">
<paragraph id="8BD378407079FFC8FF3EFA0302FDFA08" blockId="11.[133,1455,1454,1495]" pageId="11" pageNumber="10">
FIG. 5. —
<taxonomicName id="4C6C03C37079FFC8FF6EFA030780FA1E" authority="Barrientos" authorityName="Barrientos" authorityYear="2020" box="[213,506,1454,1472]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="fadriquei" status="sp. nov.">
<emphasis id="B918A4527079FFC8FF6EFA03072AFA1E" box="[213,336,1454,1472]" italics="true" pageId="11" pageNumber="10">Lepthyphantes</emphasis>
<emphasis id="B918A4527079FFC8FEEDFA0307E4FA1E" box="[342,414,1454,1472]" italics="true" pageId="11" pageNumber="10">fadriquei</emphasis>
Barrientos
</taxonomicName>
<taxonomicNameLabel id="A22B19297079FFC8FE44FA03044BFA1E" box="[511,561,1454,1472]" pageId="11" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
, genital organs;
<emphasis id="B918A4527079FFC8FD02FA0304BCFA1E" bold="true" box="[697,710,1454,1472]" pageId="11" pageNumber="10">A</emphasis>
,
<emphasis id="B918A4527079FFC8FD6BFA0304A7FA1E" bold="true" box="[720,733,1454,1472]" pageId="11" pageNumber="10">B</emphasis>
, male copulatory bulb, retrolateral view (
<emphasis id="B918A4527079FFC8FB8AFA030244FA1E" bold="true" box="[1073,1086,1454,1472]" pageId="11" pageNumber="10">A</emphasis>
), ventral view (
<emphasis id="B918A4527079FFC8FB00FA0302B2FA1E" bold="true" box="[1211,1224,1454,1472]" pageId="11" pageNumber="10">B</emphasis>
);
<emphasis id="B918A4527079FFC8FB63FA03029CFA1E" bold="true" box="[1240,1254,1454,1472]" pageId="11" pageNumber="10">C</emphasis>
, schema of the embolic division;
<emphasis id="B918A4527079FFC8FF75FA6806A6FA09" bold="true" box="[206,220,1477,1495]" pageId="11" pageNumber="10">D</emphasis>
,
<emphasis id="B918A4527079FFC8FF5DFA680688FA09" bold="true" box="[230,242,1477,1495]" pageId="11" pageNumber="10">E</emphasis>
, epigyne, ventral view (
<emphasis id="B918A4527079FFC8FE0DFA6807BEFA09" bold="true" box="[438,452,1477,1495]" pageId="11" pageNumber="10">D</emphasis>
); idem, lateral view (
<emphasis id="B918A4527079FFC8FDD5FA680400FA09" bold="true" box="[622,634,1477,1495]" pageId="11" pageNumber="10">E</emphasis>
). Abbreviations: see Material and methods. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BD378407079FFC8FF3FF9990746F830" blockId="11.[132,776,1588,2030]" pageId="11" pageNumber="10">
embolic division (
<figureCitation id="135764C57079FFC8FEFFF99907E9F991" box="[324,403,1588,1615]" captionStart="FIG" captionStartId="11.[133,144,1454,1472]" captionTargetBox="[264,1311,225,1398]" captionTargetId="figure@11.[246,1342,215,1406]" captionTargetPageId="11" captionText="FIG. 5. — Lepthyphantes fadriquei Barrientos n. sp., genital organs; A, B, male copulatory bulb, retrolateral view (A), ventral view (B); C, schema of the embolic division; D, E, epigyne, ventral view (D); idem, lateral view (E). Abbreviations: see Material and methods. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3627067" httpUri="https://zenodo.org/record/3627067/files/figure.png" pageId="11" pageNumber="10">Fig. 5C</figureCitation>
), the radix. Radix with a sharp and spiny process in its posterior part, and immediately afterwards, on its internal side, the lamella characteristica. Lamella consisting of a long and narrow projection, progressively sharpened toward the tip, maintaining a progressive curvature. Terminal apophysis parallel to the lamella characteristica, equally long and narrow, and describing a curved path of somewhat greater length; these two similar structures running parallel. Embolus complex, arising from the anterior part of the radix; with a laminar expansion, thin and transparent, as protective element of the embolus proper. Suprategular apophysis as a sharp process in front and below the terminal apophysis and lamelle characteristica; sharp, sclerotized, thicker and curved (
<figureCitation id="135764C57079FFC8FF66F87E0756F830" box="[221,300,2003,2030]" captionStart="FIG" captionStartId="11.[133,144,1454,1472]" captionTargetBox="[264,1311,225,1398]" captionTargetId="figure@11.[246,1342,215,1406]" captionTargetPageId="11" captionText="FIG. 5. — Lepthyphantes fadriquei Barrientos n. sp., genital organs; A, B, male copulatory bulb, retrolateral view (A), ventral view (B); C, schema of the embolic division; D, E, epigyne, ventral view (D); idem, lateral view (E). Abbreviations: see Material and methods. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3627067" httpUri="https://zenodo.org/record/3627067/files/figure.png" pageId="11" pageNumber="10">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BD378407079FFC8FC96F99902C2F990" blockId="11.[813,1456,1588,2029]" box="[813,1208,1588,1615]" pageId="11" pageNumber="10">
<heading id="D09BCF2C7079FFC8FC96F99902C2F990" box="[813,1208,1588,1615]" fontSize="11" level="3" pageId="11" pageNumber="10" reason="1">
<emphasis id="B918A4527079FFC8FC96F99902C2F990" box="[813,1208,1588,1615]" italics="true" pageId="11" pageNumber="10">
Female (
<typeStatus id="54D7C6E27079FFC8FC3FF99805A7F991" box="[900,989,1589,1615]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
[MZB 2001-0414])
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD378407079FFC8FC96F9F90583F833" blockId="11.[813,1456,1588,2029]" pageId="11" pageNumber="10">
<emphasis id="B918A4527079FFC8FC96F9F90516F9B0" bold="true" box="[813,876,1620,1646]" pageId="11" pageNumber="10">Body.</emphasis>
Total body length: 2.15. Carapace: 0.88 long by 0.64 wide, yellow, very light, without pigmentary pattern; some scattered hairs in the cephalic part. Completely blind; without eyes or areolated remains. Chelicerae: 0.37 of length and a coloration more intense than carapace; with three tiny, equidistant teeth on the promargin; no teeth on the retromargin. Legs yellowish, very pale, long and thin; measurements: leg I (1.42 + 0.29 + 1.37 + 1.27 + 0.86), leg II (1.27 + 0.27 + 1.32 + 1.27 + 0.83), leg III (1.22 + 0.25 + 0.93 + 1.03 + 0.64), and leg IV (1.320 + 0.27 + 1.24 + 1.22 + 0.78). Opisthosoma 1.22 long by 0.78 wide, yellowish white, uniform and without pigmentary pattern, covered with fine, long, scattered and colorless hairs.
</paragraph>
<caption id="DF1328C8707EFFCFFF3FFB4A0475FAD1" ID-DOI="http://doi.org/10.5281/zenodo.3627069" ID-Zenodo-Dep="3627069" httpUri="https://zenodo.org/record/3627069/files/figure.png" pageId="12" pageNumber="11" startId="12.[132,143,1255,1273]" targetBox="[270,1314,229,1197]" targetPageId="12">
<paragraph id="8BD37840707EFFCFFF3FFB4A0475FAD1" blockId="12.[132,1454,1255,1295]" pageId="12" pageNumber="11">
FIG. 6. —
<taxonomicName id="4C6C03C3707EFFCFFF6FFB4A0472FB27" authority="Barrientos" authorityName="Barrientos" authorityYear="2020" box="[212,520,1255,1273]" class="Arachnida" family="Linyphiidae" genus="Lepthyphantes" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="imazigheni" status="sp. nov.">
<emphasis id="B918A452707EFFCFFF6FFB4A0735FB27" box="[212,335,1255,1273]" italics="true" pageId="12" pageNumber="11">Lepthyphantes</emphasis>
<taxonomicName id="4C6C03C3707EFFCFFEEEFB4A07D7FB27" box="[341,429,1255,1273]" pageId="12" pageNumber="11" rank="species" species="imazigheni">
<emphasis id="B918A452707EFFCFFEEEFB4A07D7FB27" box="[341,429,1255,1273]" italics="true" pageId="12" pageNumber="11">imazigheni</emphasis>
</taxonomicName>
Barrientos
</taxonomicName>
<taxonomicNameLabel id="A22B1929707EFFCFFDB7FB4A0445FB27" box="[524,575,1255,1273]" pageId="12" pageNumber="11" rank="species">n. sp.</taxonomicNameLabel>
, carapace and genital organs:
<emphasis id="B918A452707EFFCFFC87FB4A0533FB27" bold="true" box="[828,841,1255,1273]" pageId="12" pageNumber="11">A</emphasis>
, carapace;
<emphasis id="B918A452707EFFCFFC10FB4A05C2FB27" bold="true" box="[939,952,1255,1273]" pageId="12" pageNumber="11">B</emphasis>
,
<emphasis id="B918A452707EFFCFFC78FB4A05ABFB27" bold="true" box="[963,977,1255,1273]" pageId="12" pageNumber="11">C</emphasis>
, epigyne, ventral view (
<emphasis id="B918A452707EFFCFFB2FFB4A02DBFB27" bold="true" box="[1172,1185,1255,1273]" pageId="12" pageNumber="11">B</emphasis>
), lateral view (
<emphasis id="B918A452707EFFCFFAACFB4A035FFB27" bold="true" box="[1303,1317,1255,1273]" pageId="12" pageNumber="11">C</emphasis>
). Abbreviations: see Material and methods. Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BD37840707EFFCFFF3FFA790718F833" blockId="12.[132,775,1492,2030]" pageId="12" pageNumber="11">
<emphasis id="B918A452707EFFCFFF3FFA790730FA30" bold="true" box="[132,330,1492,1518]" pageId="12" pageNumber="11">Female genitalia.</emphasis>
Epigyne (
<figureCitation id="135764C5707EFFCFFE7BFA78044DFA31" box="[448,567,1493,1520]" captionStart="FIG" captionStartId="11.[133,144,1454,1472]" captionTargetBox="[264,1311,225,1398]" captionTargetId="figure@11.[246,1342,215,1406]" captionTargetPageId="11" captionText="FIG. 5. — Lepthyphantes fadriquei Barrientos n. sp., genital organs; A, B, male copulatory bulb, retrolateral view (A), ventral view (B); C, schema of the embolic division; D, E, epigyne, ventral view (D); idem, lateral view (E). Abbreviations: see Material and methods. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3627067" httpUri="https://zenodo.org/record/3627067/files/figure.png" pageId="12" pageNumber="11">Figs 5D, E</figureCitation>
;
<figureCitation id="135764C5707EFFCFFDFDFA7804EBFA31" box="[582,657,1493,1519]" captionStart="FIG" captionStartId="23.[133,144,1789,1807]" captionTargetBox="[135,1455,215,1732]" captionTargetId="figure@23.[135,1458,210,1732]" captionTargetPageId="23" captionText="FIG. 13. — A, B, Epigyne of Lepthyphantes almoravidus Barrientos n. sp. in ventral (A) and lateral (B) views; C, D, epigyne of Lepthyphantes biospeleologorum Barrientos n. sp. in dorsal (C) and lateral (D) views; E, F, epigyne of Lepthyphantes fadriquei Barrientos n. sp. in lateral (E) and ventral (F) views; G, male copulatory bulb of L. fadriquei Barrientos n. sp. retrolateral views; H, I, epigyne of Lepthyphantes ensiferus Barrientos n. sp. in lateral (H) and dorsal (I) views; J, male copulatory bulb of Lepthyphantes ensiferus Barrientos n. sp. in retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.3627083" httpUri="https://zenodo.org/record/3627083/files/figure.png" pageId="12" pageNumber="11">13E, F</figureCitation>
). Epigyne very developed and outstanding, in relation to the size of the opisthosoma; almost perpendicular to epigastrum surface. Its basal part narrowing progressively and formed by the anterior wall and the lateral walls, covered with long and scattered hairs. Distally, scape voluminous; proscape drawing a smooth surface with an elliptical contour; proscape framed between the extensions of the lateral walls. In the margins of the proscape, the fertilization ducts well visible. Scape: middle part located practically below the proscape; distal part oriented outward again, forming a narrow point (stretcher) protruding below. Entire set describing in a small space a double directional change in the shape of an “S”, which can be seen in a lateral view (
<figureCitation id="135764C5707EFFCFFF35F83E06A0F870" box="[142,218,1939,1966]" captionStart="FIG" captionStartId="11.[133,144,1454,1472]" captionTargetBox="[264,1311,225,1398]" captionTargetId="figure@11.[246,1342,215,1406]" captionTargetPageId="11" captionText="FIG. 5. — Lepthyphantes fadriquei Barrientos n. sp., genital organs; A, B, male copulatory bulb, retrolateral view (A), ventral view (B); C, schema of the embolic division; D, E, epigyne, ventral view (D); idem, lateral view (E). Abbreviations: see Material and methods. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3627067" httpUri="https://zenodo.org/record/3627067/files/figure.png" pageId="12" pageNumber="11">Fig. 5E</figureCitation>
). By transparency through the anterior wall of the epigyne, the receptacula observed in the pedunculated area, near the scape base.
</paragraph>
</subSubSection>
</treatment>
</document>
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