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<mods:title id="39AF7C4E406E838EC2410E33FD133C5D">Euconnus Thomson of Japan: redescriptions of species established by Reitter, Sharp and Franz, new synonyms, and summary of current state of knowledge (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<treatment id="03C03B6BFF87FFEAFF7B2AA8FE24FB28" ID-DOI="http://doi.org/10.5281/zenodo.6028756" ID-GBIF-Taxon="192597249" ID-Zenodo-Dep="6028756" LSID="urn:lsid:plazi:treatment:03C03B6BFF87FFEAFF7B2AA8FE24FB28" httpUri="http://treatment.plazi.org/id/03C03B6BFF87FFEAFF7B2AA8FE24FB28" lastPageNumber="29" pageId="28" pageNumber="29">
<subSubSection id="C373D9F6FF87FFEAFF7B2AA8FD83FD8B" box="[151,613,510,537]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BD68A7DFF87FFEAFF7B2AA8FD83FD8B" blockId="28.[151,613,510,537]" box="[151,613,510,537]" pageId="28" pageNumber="29">
<heading id="D09E3D11FF87FFEAFF7B2AA8FD83FD8B" bold="true" box="[151,613,510,537]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
<taxonomicName id="4C69F1FEFF87FFEAFF7B2AA8FD83FD8B" ID-CoL="9G95S" authority="Hoshina, 2019" authorityName="Hoshina" authorityYear="2019" box="[151,613,510,537]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="touko" subGenus="Euconnus">
<emphasis id="B91D566FFF87FFEAFF7B2AA8FD83FD8B" bold="true" box="[151,613,510,537]" pageId="28" pageNumber="29">
<emphasis id="B91D566FFF87FFEAFF7B2AA8FEF5FD8A" bold="true" box="[151,275,511,537]" italics="true" pageId="28" pageNumber="29">Euconnus</emphasis>
(
<emphasis id="B91D566FFF87FFEAFEC82AA8FE46FD8A" bold="true" box="[292,416,511,537]" italics="true" pageId="28" pageNumber="29">Euconnus</emphasis>
)
<emphasis id="B91D566FFF87FFEAFE5D2AA9FE13FD8A" bold="true" box="[433,501,510,537]" italics="true" pageId="28" pageNumber="29">touko</emphasis>
Hoshina
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C373D9F6FF87FFEAFF2B2913FC72FDE9" pageId="28" pageNumber="29" type="reference_group">
<paragraph id="8BD68A7DFF87FFEAFF2B2913FDDBFDC8" blockId="28.[199,916,579,634]" box="[199,573,579,603]" pageId="28" pageNumber="29">
<treatmentCitationGroup id="AB79AD53FF87FFEAFF2B2913FDDBFDC8" box="[199,573,579,603]" pageId="28" pageNumber="29">
<treatmentCitation id="0AC8AC6CFF87FFEAFF2B2913FDDEFDC8" author="Hoshina, H." box="[199,568,579,603]" page="56" pageId="28" pageNumber="29" year="2019">
<taxonomicName id="4C69F1FEFF87FFEAFF2B2913FDDEFDC8" ID-CoL="9G95S" authority="Hoshina, 2019 " authorityName="Hoshina" authorityPageNumber="56" authorityYear="2019" box="[199,568,579,603]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="touko">
<emphasis id="B91D566FFF87FFEAFF2B2913FE8FFDC8" box="[199,361,579,603]" italics="true" pageId="28" pageNumber="29">Euconnus touko</emphasis>
<bibRefCitation id="EFF8F78CFF87FFEAFE9C2914FDDEFDC8" author="Hoshina, H." box="[368,568,579,603]" pageId="28" pageNumber="29" pagination="49 - 65" refId="ref24877" refString="Hoshina, H. (2019 a) Seven new species of the subfamily Scydmaeninae (Coleoptera: Staphylinidae) from Japan. The Memoirs of the Research and Education Center for Regional Environment, Fukui University, 25, 49 - 65. [2018]" type="journal article" year="2019">Hoshina, 2019a: 56</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8BD68A7DFF87FFEAFF2B2934FC72FDE9" blockId="28.[199,916,579,634]" box="[199,916,610,634]" pageId="28" pageNumber="29">
<taxonomicName id="4C69F1FEFF87FFEAFF2B2934FDA5FDE9" ID-CoL="9G95S" authority="Hoshina, 2019" authorityName="Hoshina" authorityYear="2019" box="[199,579,610,634]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="touko" subGenus="Euconnus">
<emphasis id="B91D566FFF87FFEAFF2B2934FECDFDE9" box="[199,299,611,634]" italics="true" pageId="28" pageNumber="29">Euconnus</emphasis>
(
<emphasis id="B91D566FFF87FFEAFED62934FE78FDE9" box="[314,414,611,634]" italics="true" pageId="28" pageNumber="29">Euconnus</emphasis>
)
<emphasis id="B91D566FFF87FFEAFE412935FE03FDE9" box="[429,485,610,634]" italics="true" pageId="28" pageNumber="29">touko</emphasis>
Hoshina
</taxonomicName>
;
<emphasis id="B91D566FFF87FFEAFDA12935FC72FDE9" bold="true" box="[589,916,610,634]" pageId="28" pageNumber="29">
here placed in
<taxonomicName id="4C69F1FEFF87FFEAFD072935FCB0FDEA" ID-CoL="8VRNT" box="[747,854,610,633]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91D566FFF87FFEAFD072935FCB0FDEA" bold="true" box="[747,854,610,633]" italics="true" pageId="28" pageNumber="29">Euconnus</emphasis>
</taxonomicName>
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</emphasis>
</paragraph>
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<subSubSection id="C373D9F6FF87FFEAFF7B29FFFE24FB28" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BD68A7DFF87FFEAFF7B29FFFBC2FBE1" blockId="28.[151,1437,680,1211]" pageId="28" pageNumber="29">
<emphasis id="B91D566FFF87FFEAFF7B29FFFEEDFD51" bold="true" box="[151,267,680,706]" pageId="28" pageNumber="29">Remarks.</emphasis>
This species is known to occur on Shimokoshiki-jima, a small island off NW coast of Kyushu. It seems to be a member of the
<taxonomicName id="4C69F1FEFF87FFEAFE99299AFE2FFD75" authorityName="Vit" authorityYear="2011" box="[373,457,716,742]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="alesi">
<emphasis id="B91D566FFF87FFEAFE99299AFE2FFD75" box="[373,457,716,742]" italics="true" pageId="28" pageNumber="29">E. alesi</emphasis>
</taxonomicName>
species group. Three males are known, with the body length 1.471.53, which overlaps with variability of
<taxonomicName id="4C69F1FEFF87FFEAFE8629A6FDC7FC99" authorityName="Hoshina" authorityYear="2013" box="[362,545,752,778]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kumejimensis">
<emphasis id="B91D566FFF87FFEAFE8629A6FDC7FC99" box="[362,545,752,778]" italics="true" pageId="28" pageNumber="29">E. kumejimensis</emphasis>
</taxonomicName>
(
<quantity id="4C912798FF87FFEAFDDD29A7FD58FC98" box="[561,702,752,779]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="28" pageNumber="29" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.51.7 mm</quantity>
), a species similar to
<taxonomicName id="4C69F1FEFF87FFEAFC5D29A6FBE3FC99" authorityName="Vit" authorityYear="2011" box="[945,1029,752,778]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="alesi">
<emphasis id="B91D566FFF87FFEAFC5D29A6FBE3FC99" box="[945,1029,752,778]" italics="true" pageId="28" pageNumber="29">E. alesi</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFF8F78CFF87FFEAFBF829A7FB36FC99" author="Hoshina, H." box="[1044,1232,752,778]" pageId="28" pageNumber="29" pagination="49 - 65" refId="ref24877" refString="Hoshina, H. (2019 a) Seven new species of the subfamily Scydmaeninae (Coleoptera: Staphylinidae) from Japan. The Memoirs of the Research and Education Center for Regional Environment, Fukui University, 25, 49 - 65. [2018]" type="journal article" year="2019">Hoshina (2019a)</bibRefCitation>
concluded that
<taxonomicName id="4C69F1FEFF87FFEAFA6829A6FF33FCBD" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="touko">
<emphasis id="B91D566FFF87FFEAFA6829A6FF33FCBD" italics="true" pageId="28" pageNumber="29">E. touko</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C69F1FEFF87FFEAFE872842FDC7FCBD" authorityName="Hoshina" authorityYear="2013" box="[363,545,788,814]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kumejimensis">
<emphasis id="B91D566FFF87FFEAFE872842FDC7FCBD" box="[363,545,788,814]" italics="true" pageId="28" pageNumber="29">E. kumejimensis</emphasis>
</taxonomicName>
“in the median lobe of aedeagus sharply curved ventrally, but can be separated from it by having the median lobe of aedeagus with the apex at a ventro-apical corner, which is distinctly projected at middle and both corners of apical margin in ventral and dorsal views”, vs.
<taxonomicName id="4C69F1FEFF87FFEAFBED280AFB5EFCE5" authorityName="Hoshina" authorityYear="2013" box="[1025,1208,860,886]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kumejimensis">
<emphasis id="B91D566FFF87FFEAFBED280AFB5EFCE5" box="[1025,1208,860,886]" italics="true" pageId="28" pageNumber="29">E. kumejimensis</emphasis>
</taxonomicName>
having “the median lobe with the apex simply round in ventral and dorsal views”. The aedeagus illustrated for
<taxonomicName id="4C69F1FEFF87FFEAFB2F28D6FA9AFC09" authorityName="Hoshina" authorityYear="2013" box="[1219,1404,896,922]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kumejimensis">
<emphasis id="B91D566FFF87FFEAFB2F28D6FA9AFC09" box="[1219,1404,896,922]" italics="true" pageId="28" pageNumber="29">E. kumejimensis</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFF8F78CFF87FFEAFF7B28F3FEA0FC2D" author="Hoshina, H." box="[151,326,932,958]" pageId="28" pageNumber="29" pagination="285 - 288" refId="ref24786" refString="Hoshina, H. (2013) New record of the subfamily Scydmaeninae (Coleoptera, Staphylinidae) from Kumejima Island, the Ryukyus, Japan, with description of a new species. Japanese Journal of Systematic Entomology, 19 (2), 285 - 288." type="journal article" year="2013">Hoshina (2013)</bibRefCitation>
has a partly everted endophallus, and even the one later figured in
<bibRefCitation id="EFF8F78CFF87FFEAFBC128F3FB0EFC2D" author="Hoshina, H." box="[1069,1256,932,958]" pageId="28" pageNumber="29" pagination="49 - 65" refId="ref24877" refString="Hoshina, H. (2019 a) Seven new species of the subfamily Scydmaeninae (Coleoptera: Staphylinidae) from Japan. The Memoirs of the Research and Education Center for Regional Environment, Fukui University, 25, 49 - 65. [2018]" type="journal article" year="2019">Hoshina (2019a)</bibRefCitation>
is inconclusive, as it is also partly distorted and simplified. The aedeagus of
<taxonomicName id="4C69F1FEFF87FFEAFCA8289EFC44FC71" box="[836,930,968,994]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="touko">
<emphasis id="B91D566FFF87FFEAFCA8289EFC44FC71" box="[836,930,968,994]" italics="true" pageId="28" pageNumber="29">E. touko</emphasis>
</taxonomicName>
illustrated in
<bibRefCitation id="EFF8F78CFF87FFEAFBAC289FFB1BFC71" author="Hoshina, H." box="[1088,1277,968,994]" pageId="28" pageNumber="29" pagination="49 - 65" refId="ref24877" refString="Hoshina, H. (2019 a) Seven new species of the subfamily Scydmaeninae (Coleoptera: Staphylinidae) from Japan. The Memoirs of the Research and Education Center for Regional Environment, Fukui University, 25, 49 - 65. [2018]" type="journal article" year="2019">Hoshina (2019a)</bibRefCitation>
seems almost non-distorted, and it is not possible to compare shapes of the apical structures (entirely obscured and distorted by the erected endophallus in the
<typeStatus id="54D234DFFF87FFEAFE032F47FDB7FBB9" box="[495,593,1040,1066]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C69F1FEFF87FFEAFD942F46FCC9FBB9" authorityName="Hoshina" authorityYear="2013" box="[632,815,1040,1066]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kumejimensis">
<emphasis id="B91D566FFF87FFEAFD942F46FCC9FBB9" box="[632,815,1040,1066]" italics="true" pageId="28" pageNumber="29">E. kumejimensis</emphasis>
</taxonomicName>
). Comparisons will be possible only when non-erected aedeagi of both species are illustrated, and the cephalic modification in males properly studied (preferably by scanning electron microscopy). Therefore, the status of
<taxonomicName id="4C69F1FEFF87FFEAFCEB2F0EFC82FBE1" box="[775,868,1112,1138]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="touko">
<emphasis id="B91D566FFF87FFEAFCEB2F0EFC82FBE1" box="[775,868,1112,1138]" italics="true" pageId="28" pageNumber="29">E. touko</emphasis>
</taxonomicName>
remains unclear.
</paragraph>
<paragraph id="8BD68A7DFF87FFEAFF2B2F2BFE24FB28" blockId="28.[151,1437,680,1211]" pageId="28" pageNumber="29">
<bibRefCitation id="EFF8F78CFF87FFEAFF2B2F2BFE64FB05" author="Hoshina, H." box="[199,386,1148,1174]" pageId="28" pageNumber="29" pagination="49 - 65" refId="ref24877" refString="Hoshina, H. (2019 a) Seven new species of the subfamily Scydmaeninae (Coleoptera: Staphylinidae) from Japan. The Memoirs of the Research and Education Center for Regional Environment, Fukui University, 25, 49 - 65. [2018]" type="journal article" year="2019">Hoshina (2019a)</bibRefCitation>
did not place this species in any subgenus. As all other members of the
<taxonomicName id="4C69F1FEFF87FFEAFB752F2AFB0AFB05" authorityName="Vit" authorityYear="2011" box="[1177,1260,1148,1174]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="alesi">
<emphasis id="B91D566FFF87FFEAFB752F2AFB0AFB05" box="[1177,1260,1148,1174]" italics="true" pageId="28" pageNumber="29">E. alesi</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C69F1FEFF87FFEAFAAC2F2AFA7AFB05" box="[1344,1436,1148,1174]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="touko">
<emphasis id="B91D566FFF87FFEAFAAC2F2AFA7AFB05" box="[1344,1436,1148,1174]" italics="true" pageId="28" pageNumber="29">E. touko</emphasis>
</taxonomicName>
belongs in
<taxonomicName id="4C69F1FEFF87FFEAFEFE2FF6FE66FB29" box="[274,384,1185,1210]" class="Insecta" family="Staphylinidae" genus="Euconnus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B91D566FFF87FFEAFEFE2FF6FE66FB29" box="[274,384,1185,1210]" italics="true" pageId="28" pageNumber="29">Euconnus</emphasis>
</taxonomicName>
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