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76 KiB
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504 lines
76 KiB
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<document id="CB9A14C46CB91752CF8F9FEE7F2E7089" ID-CLB-Dataset="2551" ID-DOI="10.1206/3731.2" ID-GBIF-Dataset="d0a0671c-03b7-4e26-b5c5-4a9087cb4991" ID-ISSN="0003-0082" ID-Zenodo-Dep="5359990" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630229250042" checkinUser="carolina" docAuthor="Ojanguren-Affilastro, Andrés A., Mattoni, Camilo I., Ochoa, José A. & Prendini, Lorenzo" docDate="2012" docId="039887FFFFCCFFF2050AFC9CA1C794FF" docLanguage="en" docName="N3731.pdf" docOrigin="American Museum Novitates 2012 (3731)" docSource="http://www.bioone.org/doi/abs/10.1206/3731.2" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Rumikiru Ojanguren-Affilastro & Mattoni & Ochoa & Prendini 2012, n. gen." docType="treatment" docVersion="6" lastPageNumber="11" masterDocId="FFA1FF87FFC8FFF80731FFABA20D9039" masterDocTitle="Rumikiru, n. gen. (Scorpiones: Bothriuridae), a New Scorpion Genus from the Atacama Desert" masterLastPageNumber="44" masterPageNumber="1" pageNumber="5" updateTime="1699216958989" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0371EAA0FD8B7BC1C745DE3E9F564AAA">Rumikiru, n. gen. (Scorpiones: Bothriuridae), a New Scorpion Genus from the Atacama Desert</mods:title>
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captionTargetPageId-17="21" captionTargetPageId-18="22" captionTargetPageId-19="24" captionTargetPageId-2="6" captionTargetPageId-20="24" captionTargetPageId-21="25" captionTargetPageId-22="26" captionTargetPageId-23="27" captionTargetPageId-24="28" captionTargetPageId-25="29" captionTargetPageId-26="31" captionTargetPageId-27="32" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="11" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="FIGURE 1. Most parsimonious tree (length = 167; CI: 0.479; RI: 0.813), obtained by cladistic analysis of 65 morphological characters (appendix 1) scored for 30 species of Bothriuridae Simon, 1880, with the implied weighting regime that maximized average support (Mattoni et al., in press), showing the position of Rumikiru, n. gen., in the family. Synapomorphies optimized with accelerated transformation indicated with bars. Black bars indicate uniquely derived apomorphic states, white bars indicate parallel derivations of apomorphic states. Numbers above bars indicate characters, numbers below indicate states." captionText-1="FIGURE 2. Map of central Chile, plotting known locality records of Rumikiru atacama, n. sp. (circles), and Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb. (squares)." captionText-10="FIGURE 11. Rumikiru atacama, n. sp., sinistral chelicera, dorsal aspect. A. ♂ (MACN-Ar). B. ♀ (MACN-Ar). Scale bars = 0.3 mm." captionText-11="FIGURE 12. Rumikiru, n. gen., carapace. A, C. Rumikiru atacama, n. sp., ♂ (MACN-Ar). B, D. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb., ♂ (MACN-Ar). A, B. Dorsal aspect. C, D. Lateral aspect. Scale bars = 1 mm." captionText-12="FIGURE 13. Rumikiru atacama, n. sp., sternum, genital opercula and pectines. A, B. Sternum and genital opercula, ventral aspect. C, D. Sinistral pecten, ventral aspect. A, C, E, G. ♂ (MACN-Ar). B, D, F, H. ♀ (MACN-Ar). E, F. Pectinal teeth, close-up. G, H. Pectinal peg sensilla, close-up. Scale bars = 1 mm (A–D), 0.2 mm (E, F), 0.02 mm (G), 0.01 mm (H)." captionText-13="FIGURE 14. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb. A, B. Sternum and genital opercula, ventral aspect. C, D. Sinistral pecten, ventral aspect. A, C, E, G. ♂ (MACN-Ar). B, D, F, H. ♀ (MACN-Ar). E, F. Pectinal teeth, close-up. G, H. Pectinal peg sensilla, close-up. Scale bars = 1 mm (A–D), 0.2 mm (E, F), 0.02 mm (G), 0.01 mm (H)." captionText-14="FIGURE 15. Rumikiru atacama, n. sp., ♂ (MACN- Ar), sinistral telotarsi, prolateral aspect. A–D. Legs I–IV. Scale bar = 0.3 mm." captionText-15="FIGURE 16. Rumikiru atacama, n. sp., ♂ (MACN-Ar), dextral pedipalp femur. A. Dorsal aspect. B. External aspect. C. Ventral aspect. D. Internal aspect. Scale bar = 1 mm." captionText-16="FIGURE 17. Rumikiru atacama, n. sp., ♂ (MACN-Ar), dextral pedipalp patella. A. Dorsal aspect. B. External aspect. C. Ven- tral aspect. D. Internal aspect. Scale bar = 1 mm." captionText-17="FIGURE 18. Rumikiru atacama, n. sp., ♂ (MACN-Ar), dextral pedipalp chela. A. Dorsal aspect. B. External aspect. C. Ventral aspect. D. Internal aspect. Scale bar = 1 mm." captionText-18="FIGURE 19. Rumikiru atacama, n. sp., ♀ (MACN-Ar), dextral pedipalp chela. A. Dorsal aspect. B. External aspect. C. Ventral aspect. D. Internal aspect. Scale bar = 1 mm." captionText-19="FIGURE 20. Rumikiru lourencoi (Ojanguren- Affilastro, 2003), n. comb., ♂ (MACN-Ar), dextral pedipalp femur. A. Dorsal aspect. B. External aspect. C. Ventral aspect. D. Internal aspect. Scale bar = 1 mm." captionText-2="FIGURE 3. Map of central-western South America, plotting known locality records of Orobothriurus Maury, 1976 (circles), Pachakutej Ochoa, 2004 (triangles), and Rumikiru, n. gen. (crosses)." captionText-20="FIGURE 21. Rumikiru lourencoi (Ojanguren- Affilastro, 2003), n. comb., ♂ (MACN-Ar), dextral pedipalp patella. A. Dorsal aspect. B. External aspect. C. Ventral aspect. D. Internal aspect. Scale bar = 1 mm." captionText-21="FIGURE 22. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb., ♂ (MACN-Ar), dextral pedipalp chela. A. Dorsal aspect. B. External aspect. C. Ventral aspect. D. Internal aspect. Scale bar = 1 mm." captionText-22="FIGURE 23. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb., ♀ (MACN-Ar), dextral pedipalp chela. A. Dorsal aspect. B. External aspect. C. Ventral aspect. D. Internal aspect. Scale bar = 1 mm." captionText-23="FIGURE 24. Rumikiru atacama, n. sp., dextral pedipalp chela. A, B. ♂ (MACN-Ar). C, D. ♀ (MACN-Ar). A, C. Internal aspect. B, D. Ventral aspect. Scale bar = 1 mm." captionText-24="FIGURE 25. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb., dextral pedipalp chela. A, B. ♂ (MACN-Ar). C, D. ♀ (MACN-Ar). A, C. Internal aspect. B, D. Ventral aspect. Scale bar = 1 mm." captionText-25="FIGURE 26. Rumikiru, n. gen., sternite VII and metasomal segments. A. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb., ♀ (MACN-Ar), sternite VII and metasomal segments I and II, ventral aspect. B–D. Rumikiru atacama, n. sp., ♂ (MACN-Ar). B. Metasomal segments I and II, lateral aspect. C. Metasomal segments III and IV, lateral aspect. D. Metasomal segment V, lateral aspect. Annotations: DL: dorsolateral carina; LSM: lateral supramedian carina; LM: lateral median carina; LIM: lateral inframedian carina. Scale bar = 1 mm." captionText-26="FIGURE 27. Rumikiru, n. gen., metasomal segment V, ventral aspect. A, C. Rumikiru atacama, n. sp. B, D. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb. A, B. ♂ (MACN-Ar). C, D. ♀ (MACN-Ar). Scale bars = 1 mm." captionText-27="FIGURE 28. Rumikiru, n. gen., telson, lateral aspect (A–D), with close-up of posterior median ventral granule (E, F) and posterior median dorsal concavity (G, H). A, C, E, G. Rumikiru atacama, n. sp. B, D, F, H. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb. A, B. ♀ (MACN-Ar). C–H. ♂ (MACN-Ar). Scale bars = 1 mm (A–D), 0.5 mm (E), 0.2 mm (F, H), 0.25 mm (G)." captionText-3="FIGURE 4. Rumikiru atacama, n. sp., habitat and habitus. A. Type locality, Llanos de Challe National Park, Chile. B. ♂ (LBRE), displaying characteristic defense posture." captionText-4="FIGURE 5. Rumikiru, n. gen., habitus in life. A. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb., paratype ♂ (AMNH), from Pan de Azucar National Park, Chile. B. Rumikiru atacama, n. sp., gravid ♀ paratype (AMNH), from Llanos de Challe National Park, Chile." captionText-5="FIGURE 6. Rumikiru, n. gen., chela, dorsoexternal aspect, showing pigmentation pattern. A. Rumikiru atacama, n. sp., ♂ (MACN-Ar). B. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb., ♂ (MACN-Ar). Scale bars = 1 mm." captionText-6="FIGURE 7. Rumikiru, n. gen., telson, lateral aspect, showing pigmentation pattern. A. Rumikiru atacama, n. sp., ♂ (MACN-Ar). B. Rumikiru lourencoi (Ojan- guren-Affilastro, 2003), n. comb., ♂ (MACN-Ar). Scale bars = 1 mm." captionText-7="FIGURE 8. Rumikiru atacama, n. sp., habitus. A, B. Holotype ♂ (MNHNS). C, D. Paratype ♀ (MACN-Ar). A, C. Dorsal aspect. B, D. Ventral aspect. Scale bars = 10 mm." captionText-8="FIGURE 9. Rumikiru lourencoi (Ojanguren-Affilastro, 2003), n. comb., habitus. A, B. Holotype ♂ (MACN- Ar). C, D. Paratype ♀ (MACN-Ar). A, C. Dorsal aspect. B, D. Ventral aspect. Scale bars = 10 mm." captionText-9="FIGURE 10. Rumikiru, n. gen., sinistral hemispermatophore. A, C, D, E. Rumikiru lourencoi (Ojanguren- Affilastro, 2003), n. comb., ♂ (MACN-Ar). B, F, G, H, I. Rumikiru atacama, n. sp., ♂ (MACN-Ar). A, B. Distal lamina, ectal aspect, under UV light. C, H. Lobe region. E, G. Sinistral hemispermatophore ectal aspect. F. Sinistral hemispermatophore ental aspect. D, I. Basal lobe, internal fold, close-up. Scale bars = 1 mm (A, B, E–G), 0.3 mm (C, H), 0.2 mm (D, I)." figureDoi-0="http://doi.org/10.5281/zenodo.5359992" figureDoi-1="http://doi.org/10.5281/zenodo.5359994" figureDoi-10="http://doi.org/10.5281/zenodo.5360010" figureDoi-11="http://doi.org/10.5281/zenodo.5360012" figureDoi-12="http://doi.org/10.5281/zenodo.5360014" figureDoi-13="http://doi.org/10.5281/zenodo.5360016" figureDoi-14="http://doi.org/10.5281/zenodo.5866194" figureDoi-15="http://doi.org/10.5281/zenodo.5866196" figureDoi-16="http://doi.org/10.5281/zenodo.5866198" figureDoi-17="http://doi.org/10.5281/zenodo.5360020" figureDoi-18="http://doi.org/10.5281/zenodo.5360022" figureDoi-19="http://doi.org/10.5281/zenodo.5866200" figureDoi-2="http://doi.org/10.5281/zenodo.5359996" figureDoi-20="http://doi.org/10.5281/zenodo.5866202" figureDoi-21="http://doi.org/10.5281/zenodo.5360026" figureDoi-22="http://doi.org/10.5281/zenodo.5360032" figureDoi-23="http://doi.org/10.5281/zenodo.5360034" figureDoi-24="http://doi.org/10.5281/zenodo.5360036" figureDoi-25="http://doi.org/10.5281/zenodo.5360038" figureDoi-26="http://doi.org/10.5281/zenodo.5866204" figureDoi-27="http://doi.org/10.5281/zenodo.5360040" figureDoi-3="http://doi.org/10.5281/zenodo.5359998" figureDoi-4="http://doi.org/10.5281/zenodo.5360000" figureDoi-5="http://doi.org/10.5281/zenodo.5360002" figureDoi-6="http://doi.org/10.5281/zenodo.5866306" figureDoi-7="http://doi.org/10.5281/zenodo.5360004" figureDoi-8="http://doi.org/10.5281/zenodo.5360006" figureDoi-9="http://doi.org/10.5281/zenodo.5360008" httpUri-0="https://zenodo.org/record/5359992/files/figure.png" httpUri-1="https://zenodo.org/record/5359994/files/figure.png" httpUri-10="https://zenodo.org/record/5360010/files/figure.png" httpUri-11="https://zenodo.org/record/5360012/files/figure.png" httpUri-12="https://zenodo.org/record/5360014/files/figure.png" httpUri-13="https://zenodo.org/record/5360016/files/figure.png" httpUri-14="https://zenodo.org/record/5866194/files/figure.png" httpUri-15="https://zenodo.org/record/5866196/files/figure.png" httpUri-16="https://zenodo.org/record/5866198/files/figure.png" httpUri-17="https://zenodo.org/record/5360020/files/figure.png" httpUri-18="https://zenodo.org/record/5360022/files/figure.png" httpUri-19="https://zenodo.org/record/5866200/files/figure.png" httpUri-2="https://zenodo.org/record/5359996/files/figure.png" httpUri-20="https://zenodo.org/record/5866202/files/figure.png" httpUri-21="https://zenodo.org/record/5360026/files/figure.png" httpUri-22="https://zenodo.org/record/5360032/files/figure.png" httpUri-23="https://zenodo.org/record/5360034/files/figure.png" httpUri-24="https://zenodo.org/record/5360036/files/figure.png" httpUri-25="https://zenodo.org/record/5360038/files/figure.png" httpUri-26="https://zenodo.org/record/5866204/files/figure.png" httpUri-27="https://zenodo.org/record/5360040/files/figure.png" httpUri-3="https://zenodo.org/record/5359998/files/figure.png" httpUri-4="https://zenodo.org/record/5360000/files/figure.png" httpUri-5="https://zenodo.org/record/5360002/files/figure.png" httpUri-6="https://zenodo.org/record/5866306/files/figure.png" httpUri-7="https://zenodo.org/record/5360004/files/figure.png" httpUri-8="https://zenodo.org/record/5360006/files/figure.png" httpUri-9="https://zenodo.org/record/5360008/files/figure.png" pageId="4" pageNumber="5">Figures 1–28</figureCitation>
|
||
; table 1
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6562FFCCFFFC074BFC28A1CB93D9" pageId="4" pageNumber="5" type="reference_group">
|
||
<paragraph id="8B8E36E9FFCCFFFC074BFC28A1CB93D9" blockId="4.[122,1213,748,992]" pageId="4" pageNumber="5">
|
||
<taxonomicName id="4C314D6AFFCCFFFC074BFC28A0A693A2" ID-CoL="69YV" authority="Ojanguren-Affilastro, 2003 a: 117 - 122" authorityName="Ojanguren-Affilastro" authorityPageNumber="117 - 122" authorityYear="2003" box="[122,683,899,923]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFCCFFFC074BFC28A31C93A2" box="[122,273,899,923]" italics="true" pageId="4" pageNumber="5">Orobothriurus</emphasis>
|
||
<bibRefCitation id="EFA04B18FFCCFFFC0629FC28A0A693A2" author="Ojanguren-Affilastro, A. A." box="[280,683,899,923]" pageId="4" pageNumber="5" pagination="117 - 122" refId="ref14584" refString="Ojanguren-Affilastro, A. A. 2003 a. Un nuevo Orobothriurus (Scorpiones, Bothriuridae) de la region de Atacama, Chile. Revista Iberica de Aracnologia 7: 117 - 122." type="journal article" year="2003">Ojanguren-Affilastro, 2003a: 117–122</bibRefCitation>
|
||
</taxonomicName>
|
||
(part);
|
||
<bibRefCitation id="EFA04B18FFCCFFFC05CFFC28A64993A2" author="Ojanguren-Affilastro, A. A." box="[766,1092,899,923]" pageId="4" pageNumber="5" pagination="69 - 74" refId="ref14655" refString="Ojanguren-Affilastro, A. A. 2004. Un nuevo Brachistosternus Pocock 1893 del norte de Chile (Scorpiones, Bothriuridae). Revista Iberica de Aracnologia 10: 69 - 74." type="journal article" year="2004">Ojanguren-Affilastro, 2004: 72</bibRefCitation>
|
||
(part);
|
||
<bibRefCitation id="EFA04B18FFCCFFFC079BFC0EA3559384" author="Ochoa, J. A." box="[170,344,933,957]" pageId="4" pageNumber="5" pagination="43 - 73" refId="ref14266" refString="Ochoa, J. A. 2004. Filogenia del genero Orobothriurus y descripcion de un nuevo genero de Bothriuridae (Scorpiones). Revista Iberica de Aracnologia 9: 43 - 73." type="journal article" year="2004">Ochoa, 2004: 44</bibRefCitation>
|
||
(part);
|
||
<bibRefCitation id="EFA04B18FFCCFFFC069AFC0DA0D79384" author="Agusto, P. & C. I. Mattoni & J. Pizarro-Araya & J. Cepeda-Pizarro & F. Lopez-Cortes" box="[427,730,933,958]" pageId="4" pageNumber="5" pagination="407 - 421" refId="ref13878" refString="Agusto, P., C. I. Mattoni, J. Pizarro-Araya, J. Cepeda-Pizarro, and F. Lopez-Cortes. 2006. Comunidades de escorpiones (Arachnida: Scorpiones) del desierto costero transicional de Chile. Revista Chilena de Historia Natural 79: 407 - 421." type="journal article" year="2006">Agusto et al., 2006: 415–419</bibRefCitation>
|
||
(part);
|
||
<bibRefCitation id="EFA04B18FFCCFFFC041CFC0EA1B49384" author="Rein, J. O." box="[813,953,933,957]" pageId="4" pageNumber="5" pagination="1 - 12" refId="ref15049" refString="Rein, J. O. 2007. Taxonomic updates in scorpions (Arachnida: Scorpiones) since the publication of the Catalog of the scorpions of the world (1758 - 1998) (Fet, Sissom, Lowe, and Braunwalder, 2000). Part 1: Bothriuridae. The scorpion files occasional papers 1: 1 - 12. (http: // www. ub. ntnu. no / scorpion-files / sf _ occationalpapers. php)." type="journal article" year="2007">Rein, 2007: 5</bibRefCitation>
|
||
(part);
|
||
<bibRefCitation id="EFA04B18FFCCFFFC033DFC0EA33493D9" author="Pizarro-Araya, J. & J. Cepeda-Pizarro & G. E. Flores" pageId="4" pageNumber="5" pagination="257 - 274" refId="ref14874" refString="Pizarro-Araya, J., J. Cepeda-Pizarro, and G. E. Flores. 2008. Diversidad taxonomica de los artropodos epigeos de la region de Atacama (Chile): estado del conocimiento. In F. A. Squeo, G. Arancio and J. R. Gutierrez (editors). Libro rojo de la flora nativa y de los sitios prioritarios para su conservacion: region de Atacama: 257 - 274. Ediciones Universidad de La Serena, La Serena. Chile." type="book chapter" year="2008">Pizarro-Araya et al., 2008: 270</bibRefCitation>
|
||
, 271 (part);
|
||
<bibRefCitation id="EFA04B18FFCCFFFC068CFC6CA0A093E6" author="Vrech, D. E. & A. V. Peretti & C. I. Mattoni" box="[445,685,967,992]" pageId="4" pageNumber="5" pagination="463 - 483" refId="ref15193" refString="Vrech, D. E., A. V. Peretti, and C. I. Mattoni. 2011. Sperm package morphology in scorpions and its relation to phylogeny. Zoological Journal of the Linnean Society 161: 463 - 483." type="journal article" year="2011">Vrech et al., 2011: 465</bibRefCitation>
|
||
, 467, 470, 475, 482 (part).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6562FFCCFFFC079BFC55A05D9404" pageId="4" pageNumber="5" type="type_taxon">
|
||
<paragraph id="8B8E36E9FFCCFFFC079BFC55A05D9404" blockId="4.[122,1225,1022,1758]" pageId="4" pageNumber="5">
|
||
<typeStatus id="548A884BFFCCFFFC079BFC55A2E89421" box="[170,229,1022,1048]" pageId="4" pageNumber="5">TYPE</typeStatus>
|
||
SPECIES:
|
||
<taxonomicName id="4C314D6AFFCCFFFC066BFC55A1B49421" authority="Ojanguren-Affilastro, 2003" authorityName="Ojanguren-Affilastro" authorityYear="2003" box="[346,953,1022,1048]" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lourencoi">
|
||
<emphasis id="B945EAFBFFCCFFFC066BFC55A07E9421" box="[346,627,1022,1048]" italics="true" pageId="4" pageNumber="5">Orobothriurus lourencoi</emphasis>
|
||
Ojanguren-Affilastro, 2003
|
||
</taxonomicName>
|
||
[=
|
||
<taxonomicName id="4C314D6AFFCCFFFC04D6FC55A3C19404" ID-CoL="4TQ5W" authority="(Ojanguren-Affilastro, 2003)" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" baseAuthorityName="Ojanguren-Affilastro" baseAuthorityYear="2003" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lourencoi" status="comb. nov.">
|
||
<emphasis id="B945EAFBFFCCFFFC04D6FC55A6C59421" box="[999,1224,1022,1048]" italics="true" pageId="4" pageNumber="5">Rumikiru lourencoi</emphasis>
|
||
(Ojanguren-Affilastro, 2003)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFCCFFFC06E8FB8FA0489404" box="[473,581,1058,1086]" pageId="4" pageNumber="5" rank="species">n. comb.</taxonomicNameLabel>
|
||
].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6562FFCCFFFC079BFBE3A09E9524" pageId="4" pageNumber="5" type="etymology">
|
||
<paragraph id="8B8E36E9FFCCFFFC079BFBE3A09E9524" blockId="4.[122,1225,1022,1758]" pageId="4" pageNumber="5">
|
||
ETYMOLOGY: The indigenous peoples of the Chilean
|
||
<collectingRegion id="49F5F80BFFCCFFFC0413FBE2A184945A" box="[802,905,1097,1123]" country="Chile" name="Atacama" pageId="4" pageNumber="5">Atacama</collectingRegion>
|
||
originally spoke Kakan or Kunza, depending on the region and, after the Inca invasion, added Quechua and Aymara to their languages.
|
||
<taxonomicName id="4C314D6AFFCCFFFC0608FB38A3A59494" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[313,424,1171,1197]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFCCFFFC0608FB38A3A59494" box="[313,424,1171,1197]" italics="true" pageId="4" pageNumber="5">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
is formed from two Quechua words,
|
||
<emphasis id="B945EAFBFFCCFFFC046EFB3FA1959494" box="[863,920,1172,1197]" italics="true" pageId="4" pageNumber="5">rumi</emphasis>
|
||
meaning “stone” and
|
||
<emphasis id="B945EAFBFFCCFFFC03A9FB38A6C59494" box="[1176,1224,1171,1197]" italics="true" pageId="4" pageNumber="5">kiru</emphasis>
|
||
meaning “tooth,” and refers to the large denticles on the movable fingers of the pedipalp chelae in this genus, unique in the family
|
||
<taxonomicName id="4C314D6AFFCCFFFC052BFB75A0BE94C1" authorityName="Simon" authorityYear="1880" box="[538,691,1246,1272]" class="Arachnida" family="Bothriuridae" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Bothriuridae</taxonomicName>
|
||
, and to the habitat of its two species, both of which appear to be restricted to rocky slopes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6562FFCCFFF1079BFA83A0F69624" lastPageId="9" lastPageNumber="10" pageId="4" pageNumber="5" type="diagnosis">
|
||
<paragraph id="8B8E36E9FFCCFFFD079BFA83A3B4966E" blockId="4.[122,1225,1022,1758]" lastBlockId="5.[120,1225,1409,1735]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
DIAGNOSIS: Species of
|
||
<taxonomicName id="4C314D6AFFCCFFFC0687FA83A029957B" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[438,548,1320,1346]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFCCFFFC0687FA83A029957B" box="[438,548,1320,1346]" italics="true" pageId="4" pageNumber="5">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFCCFFFC0505FA82A088957A" box="[564,645,1321,1347]" pageId="4" pageNumber="5" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, are easily recognized by the enlarged basal denticle of the median denticle row of the pedipalp chela movable finger (figs. 18A, 19A, 24C, 25A, C), which is approximately three times larger than and replaces the first five or six median denticles. Such a hypertrophied basal denticle is unique among bothriurids. An enlarged basal denticle on the movable finger is exhibited by some bothriurids, e.g.,
|
||
<taxonomicName id="4C314D6AFFCCFFFC048BFA15A37295C4" authority="(Gervais, 1841)" baseAuthorityName="Gervais" baseAuthorityYear="1841" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ehrenbergii">
|
||
<emphasis id="B945EAFBFFCCFFFC048BFA15A2B695C4" italics="true" pageId="4" pageNumber="5">Brachistosternus ehrenbergii</emphasis>
|
||
(Gervais, 1841)
|
||
</taxonomicName>
|
||
, but replaces no more than two or three denticles (
|
||
<bibRefCitation id="EFA04B18FFCCFFFC04C3FA48A38D961A" author="Ochoa J. A. & A. A. Ojanguren-Affilastro" pageId="4" pageNumber="5" pagination="61 - 69" refId="ref14371" refString="Ochoa J. A., and A. A. Ojanguren-Affilastro. 2007. Systematics and distribution of Brachistosternus (Brachistosternus) ehrenbergii (Gervais, 1841), with the first record of stridulation in the genus Brachistosternus. Studies on Neotropical Fauna and Environment 42 (1): 61 - 69." type="journal article" year="2007">Ochoa and Ojanguren-Affilastro, 2007</bibRefCitation>
|
||
). The submedial position of the apophysis on the internal surface of the pedipalp chela manus in the adult male of
|
||
<taxonomicName id="4C314D6AFFCCFFFC0544F985A0EE9671" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[629,739,1582,1608]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFCCFFFC0544F985A0EE9671" box="[629,739,1582,1608]" italics="true" pageId="4" pageNumber="5">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFCCFFFC05C3F985A14F9671" box="[754,834,1582,1608]" pageId="4" pageNumber="5" rank="genus">n. gen.</taxonomicNameLabel>
|
||
(figs. 18C, D, 22A, C, D, 24A, B, 25A, B), is also unique in the family. The apophysis is situated in the distal third of the surface in all other bothriurid taxa in which it occurs, and is absent from the pedipalp chela manus of basal bothriurids, e.g.,
|
||
<taxonomicName id="4C314D6AFFCCFFFC06B7F934A0CD9681" authority="Lawrence, 1928" authorityName="Lawrence" authorityYear="1928" box="[390,704,1694,1720]" class="Arachnida" family="Bothriuridae" genus="Lisposoma" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFCCFFFC06B7F934A3F29681" box="[390,511,1695,1720]" italics="true" pageId="4" pageNumber="5">Lisposoma</emphasis>
|
||
Lawrence, 1928
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C314D6AFFCCFFFC05CEF935A60E9681" authority="Simon, 1880" authorityName="Simon" authorityYear="1880" box="[767,1027,1694,1720]" class="Arachnida" family="Bothriuridae" genus="Thestylus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFCCFFFC05CEF935A1649681" box="[767,873,1694,1720]" italics="true" pageId="4" pageNumber="5">Thestylus</emphasis>
|
||
Simon, 1880
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFA04B18FFCCFFFC0323F935A6C99681" author="Prendini, L." box="[1042,1220,1694,1720]" pageId="4" pageNumber="5" pagination="1 - 78" refId="ref14965" refString="Prendini, L. 2000. Phylogeny and classification of the superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): an exemplar approach. Cladistics 16 (1): 1 - 78." type="journal article" year="2000">Prendini, 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA04B18FFCCFFFC074BF968A2BA96E4" author="Prendini, L." box="[122,183,1731,1757]" pageId="4" pageNumber="5" pagination="149 - 172" refId="ref15000" refString="Prendini, L. 2003. A new genus and species of bothriurid scorpion from the Brandberg Massif, Namibia, with a reanalysis of bothriurid phylogeny and a discussion of the phylogenetic position of Lisposoma Lawrence. Systematic Entomology 28 (2): 149 - 172." type="journal article" year="2003">2003</bibRefCitation>
|
||
). The shape of the apophysis and the position of trichobothium
|
||
<emphasis id="B945EAFBFFCCFFFC04F7F968A1D196E4" box="[966,988,1731,1757]" italics="true" pageId="4" pageNumber="5">ib</emphasis>
|
||
distal to it are also unique to
|
||
<taxonomicName id="4C314D6AFFCDFFFD07C3FA2AA36D95A2" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[242,352,1409,1435]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFCDFFFD07C3FA2AA36D95A2" box="[242,352,1409,1435]" italics="true" pageId="5" pageNumber="6">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFCDFFFD065EFA29A3B295A5" box="[367,447,1410,1436]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
|
||
The new genus may be further separated by a twist in the medial to distal third of the dentate margin of the pedipalp chela fingers, which abruptly alters the orientation of the median denticle row (more conspicuously on the movable finger), in the male (figs. 18A, 22A), that is absent in other bothriurid genera. Additionally, the carapace of
|
||
<taxonomicName id="4C314D6AFFCDFFFD0749F9BCA2EB9608" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[120,230,1559,1585]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFCDFFFD0749F9BCA2EB9608" box="[120,230,1559,1585]" italics="true" pageId="5" pageNumber="6">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFCDFFFD07C7F9BCA3459608" box="[246,328,1559,1585]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, is dorsoventrally compressed (fig. 12C, D), unlike many other bothriurids, in which it is more convex.
|
||
</paragraph>
|
||
<caption id="DF4E6661FFCDFFFD0749FA80A13E9566" ID-DOI="http://doi.org/10.5281/zenodo.5359994" ID-Zenodo-Dep="5359994" httpUri="https://zenodo.org/record/5359994/files/figure.png" pageId="5" pageNumber="6" startId="5.[120,206,1323,1344]" targetBox="[222,1118,230,1296]" targetPageId="5">
|
||
<paragraph id="8B8E36E9FFCDFFFD0749FA80A13E9566" blockId="5.[120,1224,1321,1375]" pageId="5" pageNumber="6">
|
||
FIGURE 2. Map of central Chile, plotting known locality records of
|
||
<emphasis id="B945EAFBFFCDFFFD040BFA82A1F09578" box="[826,1021,1321,1345]" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C314D6AFFCDFFFD040BFA82A1F49578" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[826,1017,1321,1345]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="atacama" status="sp. nov.">Rumikiru atacama</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A2765780FFCDFFFD0335FA81A6319578" box="[1028,1084,1322,1345]" pageId="5" pageNumber="6" rank="species">n. sp.</taxonomicNameLabel>
|
||
(circles), and
|
||
<taxonomicName id="4C314D6AFFCDFFFD0749FAECA0689566" authority="(Ojanguren-Affilastro, 2003)" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" baseAuthorityName="Ojanguren-Affilastro" baseAuthorityYear="2003" box="[120,613,1351,1375]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lourencoi" status="comb. nov.">
|
||
<emphasis id="B945EAFBFFCDFFFD0749FAECA3349566" box="[120,313,1351,1375]" italics="true" pageId="5" pageNumber="6">Rumikiru lourencoi</emphasis>
|
||
(Ojanguren-Affilastro, 2003)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFCDFFFD0541FAECA0C49566" box="[624,713,1351,1375]" pageId="5" pageNumber="6" rank="species">n. comb.</taxonomicNameLabel>
|
||
(squares).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E36E9FFCDFFF10799F9CAA0F69624" blockId="5.[120,1225,1409,1735]" lastBlockId="9.[120,1225,1165,1752]" lastPageId="9" lastPageNumber="10" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C314D6AFFCDFFFD0799F9CAA3199642" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[168,276,1633,1659]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFCDFFFD0799F9CAA3199642" box="[168,276,1633,1659]" italics="true" pageId="5" pageNumber="6">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFCDFFFD0615F9C9A3789645" box="[292,373,1634,1660]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, is most closely related to
|
||
<taxonomicName id="4C314D6AFFCDFFFD059EF9CAA1279642" authorityName="Ochoa" authorityYear="2004" box="[687,810,1633,1659]" class="Arachnida" family="Bothriuridae" genus="Pachakutej" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFCDFFFD059EF9CAA1279642" box="[687,810,1633,1659]" italics="true" pageId="5" pageNumber="6">Pachakutej</emphasis>
|
||
</taxonomicName>
|
||
, based mainly on characters of the hemispermatophore (fig. 10). The two genera share the presence of one sclerotized apophysis on the internal fold of the internal lobe, and a papillose fold in the basal lobe (figs. 10C, H). Species of
|
||
<taxonomicName id="4C314D6AFFC1FFF107A4FB26A302949E" authorityName="Ochoa" authorityYear="2004" box="[149,271,1165,1191]" class="Arachnida" family="Bothriuridae" genus="Pachakutej" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFC1FFF107A4FB26A302949E" box="[149,271,1165,1191]" italics="true" pageId="9" pageNumber="10">Pachakutej</emphasis>
|
||
</taxonomicName>
|
||
, however, possess a unique synapomorphy, a spatulate terminal process on the basal lobe of the hemispermatophore (
|
||
<bibRefCitation id="EFA04B18FFC1FFF106C0FB18A08B94F4" author="Ochoa, J. A." box="[497,646,1203,1229]" pageId="9" pageNumber="10" pagination="43 - 73" refId="ref14266" refString="Ochoa, J. A. 2004. Filogenia del genero Orobothriurus y descripcion de un nuevo genero de Bothriuridae (Scorpiones). Revista Iberica de Aracnologia 9: 43 - 73." type="journal article" year="2004">Ochoa, 2004</bibRefCitation>
|
||
) that is absent in
|
||
<taxonomicName id="4C314D6AFFC1FFF10460FB18A1B094F4" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[849,957,1203,1229]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF10460FB18A1B094F4" box="[849,957,1203,1229]" italics="true" pageId="9" pageNumber="10">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF104FBFB18A61794F4" box="[970,1050,1203,1229]" pageId="9" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, and are markedly more pigmented. The pedipalp carinae are more pronounced, especially on the femur and patella, in
|
||
<taxonomicName id="4C314D6AFFC1FFF107DFFB56A357952E" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[238,346,1277,1303]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF107DFFB56A357952E" box="[238,346,1277,1303]" italics="true" pageId="9" pageNumber="10">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF10659FB55A3B89521" box="[360,437,1278,1304]" pageId="9" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
|
||
(figs. 16, 17, 20, 21), than in
|
||
<taxonomicName id="4C314D6AFFC1FFF10430FB56A171952E" authorityName="Ochoa" authorityYear="2004" box="[769,892,1277,1303]" class="Arachnida" family="Bothriuridae" genus="Pachakutej" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFC1FFF10430FB56A171952E" box="[769,892,1277,1303]" italics="true" pageId="9" pageNumber="10">Pachakutej</emphasis>
|
||
</taxonomicName>
|
||
. The two genera may also be separated by the trichobothrial pattern of the pedipalp chela manus. Trichobothrium
|
||
<emphasis id="B945EAFBFFC1FFF10367FA8FA6649504" box="[1110,1129,1316,1341]" italics="true" pageId="9" pageNumber="10">V</emphasis>
|
||
<subScript id="17B534ACFFC1FFF10358FA9BA67F9578" attach="left" box="[1129,1138,1328,1345]" fontSize="7" pageId="9" pageNumber="10">2</subScript>
|
||
is situated in the same axis as
|
||
<emphasis id="B945EAFBFFC1FFF106BCFAE2A3AD955B" box="[397,416,1353,1378]" italics="true" pageId="9" pageNumber="10">V</emphasis>
|
||
<subScript id="17B534ACFFC1FFF10691FAFDA3A4955E" attach="left" box="[416,425,1366,1383]" fontSize="7" pageId="9" pageNumber="10">1</subScript>
|
||
and
|
||
<emphasis id="B945EAFBFFC1FFF106D1FAE2A3FE955B" box="[480,499,1353,1378]" italics="true" pageId="9" pageNumber="10">V</emphasis>
|
||
<quantity id="4CC99B0CFFC1FFF106C5FAFDA011955B" box="[500,540,1352,1383]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="10" unit="in" value="3.0">
|
||
<subScript id="17B534ACFFC1FFF106C5FAFDA3F0955E" attach="left" box="[500,509,1366,1383]" fontSize="7" pageId="9" pageNumber="10">3</subScript>
|
||
in
|
||
</quantity>
|
||
<taxonomicName id="4C314D6AFFC1FFF10513FAE3A083955B" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[546,654,1352,1378]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF10513FAE3A083955B" box="[546,654,1352,1378]" italics="true" pageId="9" pageNumber="10">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF105ADFAE2A0E7955A" box="[668,746,1353,1379]" pageId="9" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
|
||
(figs. 18C, 19C, 22C, 23C), but not in the same axis, forming an angle less than 180°, in
|
||
<taxonomicName id="4C314D6AFFC1FFF105BAFAC6A10B95BE" authorityName="Ochoa" authorityYear="2004" box="[651,774,1389,1415]" class="Arachnida" family="Bothriuridae" genus="Pachakutej" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFC1FFF105BAFAC6A10B95BE" box="[651,774,1389,1415]" italics="true" pageId="9" pageNumber="10">Pachakutej</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<emphasis id="B945EAFBFFC1FFF10476FAC6A16795BE" box="[839,874,1389,1415]" italics="true" pageId="9" pageNumber="10">Db</emphasis>
|
||
is equidistant between
|
||
<emphasis id="B945EAFBFFC1FFF10349FAC5A69895BE" box="[1144,1173,1390,1415]" italics="true" pageId="9" pageNumber="10">Dt</emphasis>
|
||
and
|
||
<emphasis id="B945EAFBFFC1FFF10749FA38A29B9594" box="[120,150,1427,1453]" italics="true" pageId="9" pageNumber="10">Eb</emphasis>
|
||
<quantity id="4CC99B0CFFC1FFF107A7FA0BA2B09594" box="[150,189,1427,1457]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="10" unit="in" value="3.0">
|
||
<subScript id="17B534ACFFC1FFF107A7FA0BA2929588" attach="left" box="[150,159,1440,1457]" fontSize="7" pageId="9" pageNumber="10">3</subScript>
|
||
in
|
||
</quantity>
|
||
<taxonomicName id="4C314D6AFFC1FFF107F2FA38A3229594" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[195,303,1427,1453]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF107F2FA38A3229594" box="[195,303,1427,1453]" italics="true" pageId="9" pageNumber="10">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF1060AFA38A3859594" box="[315,392,1427,1453]" pageId="9" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
|
||
(figs. 18B, 19B, 22B, 23B), but close to
|
||
<emphasis id="B945EAFBFFC1FFF10471FA38A1509595" box="[832,861,1427,1452]" italics="true" pageId="9" pageNumber="10">Dt</emphasis>
|
||
in
|
||
<taxonomicName id="4C314D6AFFC1FFF104B0FA38A1F19594" authorityName="Ochoa" authorityYear="2004" box="[897,1020,1427,1453]" class="Arachnida" family="Bothriuridae" genus="Pachakutej" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFC1FFF104B0FA38A1F19594" box="[897,1020,1427,1453]" italics="true" pageId="9" pageNumber="10">Pachakutej</emphasis>
|
||
</taxonomicName>
|
||
. Finally, the VSM carinae of metasomal segment V are subparallel and situated close to the VL carinae in
|
||
<taxonomicName id="4C314D6AFFC1FFF10367FA13A6CF95EB" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[1110,1218,1464,1490]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF10367FA13A6CF95EB" box="[1110,1218,1464,1490]" italics="true" pageId="9" pageNumber="10">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF10749FA75A2C895C1" box="[120,197,1502,1528]" pageId="9" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
|
||
(fig. 27), whereas the VSM carinae are slightly inclined toward the external margin in the posterior third of the segment, or absent, in
|
||
<taxonomicName id="4C314D6AFFC1FFF10548F9A8A0F99624" authorityName="Ochoa" authorityYear="2004" box="[633,756,1539,1565]" class="Arachnida" family="Bothriuridae" genus="Pachakutej" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFC1FFF10548F9A8A0F99624" box="[633,756,1539,1565]" italics="true" pageId="9" pageNumber="10">Pachakutej</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4E6661FFCEFFFE0749F9A7A1CE9679" ID-DOI="http://doi.org/10.5281/zenodo.5359996" ID-Zenodo-Dep="5359996" httpUri="https://zenodo.org/record/5359996/files/figure.png" pageId="6" pageNumber="7" startId="6.[120,206,1548,1569]" targetBox="[167,1181,227,1518]" targetPageId="6">
|
||
<paragraph id="8B8E36E9FFCEFFFE0749F9A7A1CE9679" blockId="6.[120,1224,1546,1600]" pageId="6" pageNumber="7">
|
||
FIGURE 3. Map of central-western South America, plotting known locality records of
|
||
<taxonomicName id="4C314D6AFFCEFFFE04DBF9A1A2A49606" authority="Maury, 1976" authorityName="Maury" authorityYear="1976" class="Arachnida" family="Bothriuridae" genus="Orobothriurus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFCEFFFE04DBF9A1A674961B" box="[1002,1145,1546,1570]" italics="true" pageId="6" pageNumber="7">Orobothriurus</emphasis>
|
||
<bibRefCitation id="EFA04B18FFCEFFFE03B1F9A0A2A49606" author="Maury, E. A." pageId="6" pageNumber="7" pagination="14 - 25" refId="ref14224" refString="Maury, E. A. 1976. Escorpiones y escorpionismo en el Peru. V. Orobothriurus, un nuevo genero de escorpiones altoandinos (Bothriuridae). Revista Peruana de Entomologia 18 (1): 14 - 25." type="journal article" year="1976">Maury, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
(circles),
|
||
<taxonomicName id="4C314D6AFFCEFFFE063CF98CA00F9679" authority="Ochoa, 2004" authorityName="Ochoa" authorityYear="2004" box="[269,514,1575,1600]" class="Arachnida" family="Bothriuridae" genus="Pachakutej" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFCEFFFE063CF98CA3779606" box="[269,378,1575,1599]" italics="true" pageId="6" pageNumber="7">Pachakutej</emphasis>
|
||
<bibRefCitation id="EFA04B18FFCEFFFE06B0F98CA00F9679" author="Ochoa, J. A." box="[385,514,1575,1600]" pageId="6" pageNumber="7" pagination="43 - 73" refId="ref14266" refString="Ochoa, J. A. 2004. Filogenia del genero Orobothriurus y descripcion de un nuevo genero de Bothriuridae (Scorpiones). Revista Iberica de Aracnologia 9: 43 - 73." type="journal article" year="2004">Ochoa, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
(triangles), and
|
||
<emphasis id="B945EAFBFFCEFFFE059AF98CA11C9606" box="[683,785,1575,1599]" italics="true" pageId="6" pageNumber="7">
|
||
<taxonomicName id="4C314D6AFFCEFFFE059AF98CA1009606" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[683,781,1575,1599]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">Rumikiru</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A2765780FFCEFFFE0426F983A1509679" box="[791,861,1576,1600]" pageId="6" pageNumber="7" rank="genus">n. gen.</taxonomicNameLabel>
|
||
(crosses).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E6661FFCFFFFF0749F9D3A0FD9692" ID-DOI="http://doi.org/10.5281/zenodo.5359998" ID-Zenodo-Dep="5359998" httpUri="https://zenodo.org/record/5359998/files/figure.png" pageId="7" pageNumber="8" startId="7.[120,206,1656,1677]" targetBox="[191,1163,224,1629]" targetPageId="7">
|
||
<paragraph id="8B8E36E9FFCFFFFF0749F9D3A0FD9692" blockId="7.[120,1222,1653,1707]" pageId="7" pageNumber="8">
|
||
FIGURE 4.
|
||
<taxonomicName id="4C314D6AFFCFFFFF07C1F9DDA3A196B4" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[240,428,1654,1678]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="atacama" status="sp. nov.">
|
||
<emphasis id="B945EAFBFFCFFFFF07C1F9DDA3A196B4" box="[240,428,1654,1678]" italics="true" pageId="7" pageNumber="8">Rumikiru atacama</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFCFFFFF068BF9DDA3FF96B4" box="[442,498,1654,1677]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
||
, habitat and habitus.
|
||
<emphasis id="B945EAFBFFCFFFFF05E1F9DEA0E596B4" bold="true" box="[720,744,1653,1677]" pageId="7" pageNumber="8">A.</emphasis>
|
||
Type locality, Llanos de Challe National Park, Chile.
|
||
<emphasis id="B945EAFBFFCFFFFF078AF938A2DF9692" bold="true" box="[187,210,1683,1707]" pageId="7" pageNumber="8">B.</emphasis>
|
||
♂ (LBRE), displaying characteristic defense posture.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E6661FFC0FFF0074BF927A0D796E4" ID-DOI="http://doi.org/10.5281/zenodo.5360000" ID-Zenodo-Dep="5360000" httpUri="https://zenodo.org/record/5360000/files/figure.png" pageId="8" pageNumber="9" startId="8.[122,208,1676,1697]" targetBox="[171,1179,221,1649]" targetPageId="8">
|
||
<paragraph id="8B8E36E9FFC0FFF0074BF927A0D796E4" blockId="8.[122,1226,1674,1757]" pageId="8" pageNumber="9">
|
||
FIGURE 5.
|
||
<taxonomicName id="4C314D6AFFC0FFF007C2F921A35F969B" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[243,338,1674,1698]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC0FFF007C2F921A35F969B" box="[243,338,1674,1698]" italics="true" pageId="8" pageNumber="9">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC0FFF00651F920A3A5969A" box="[352,424,1675,1699]" pageId="8" pageNumber="9" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, habitus in life.
|
||
<emphasis id="B945EAFBFFC0FFF0057FF921A06A969B" bold="true" box="[590,615,1674,1698]" pageId="8" pageNumber="9">A.</emphasis>
|
||
<taxonomicName id="4C314D6AFFC0FFF0055EF921A652969B" authority="(Ojanguren-Affilastro, 2003)" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" baseAuthorityName="Ojanguren-Affilastro" baseAuthorityYear="2003" box="[623,1119,1674,1698]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lourencoi" status="comb. nov.">
|
||
<emphasis id="B945EAFBFFC0FFF0055EF921A13C969B" box="[623,817,1674,1698]" italics="true" pageId="8" pageNumber="9">Rumikiru lourencoi</emphasis>
|
||
(Ojanguren-Affilastro, 2003)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC0FFF0035AF920A6CA969B" box="[1131,1223,1674,1698]" pageId="8" pageNumber="9" rank="species">n. comb.</taxonomicNameLabel>
|
||
, paratype ♂ (AMNH), from Pan de Azucar National Park, Chile.
|
||
<emphasis id="B945EAFBFFC0FFF0043CF903A12996F9" bold="true" box="[781,804,1704,1728]" pageId="8" pageNumber="9">B.</emphasis>
|
||
<taxonomicName id="4C314D6AFFC0FFF0041BF903A1E89686" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[810,997,1704,1728]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="atacama" status="sp. nov.">
|
||
<emphasis id="B945EAFBFFC0FFF0041BF903A1E89686" box="[810,997,1704,1728]" italics="true" pageId="8" pageNumber="9">Rumikiru atacama</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC0FFF004C0F903A6259686" box="[1009,1064,1704,1727]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
||
, gravid ♀ paratype (AMNH), from Llanos de Challe National Park, Chile.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E6661FFC1FFF10749FC55A34E9477" ID-DOI="http://doi.org/10.5281/zenodo.5360002" ID-Zenodo-Dep="5360002" httpUri="https://zenodo.org/record/5360002/files/figure.png" pageId="9" pageNumber="10" startId="9.[120,206,1022,1043]" targetBox="[254,1099,224,996]" targetPageId="9">
|
||
<paragraph id="8B8E36E9FFC1FFF10749FC55A34E9477" blockId="9.[120,1225,1020,1103]" pageId="9" pageNumber="10">
|
||
FIGURE 6.
|
||
<taxonomicName id="4C314D6AFFC1FFF107DEFC57A343942D" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[239,334,1020,1044]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF107DEFC57A343942D" box="[239,334,1020,1044]" italics="true" pageId="9" pageNumber="10">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF1066DFC56A3AE942C" box="[348,419,1021,1045]" pageId="9" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, chela, dorsoexternal aspect, showing pigmentation pattern.
|
||
<emphasis id="B945EAFBFFC1FFF10321FC57A624942D" bold="true" box="[1040,1065,1020,1044]" pageId="9" pageNumber="10">A.</emphasis>
|
||
<taxonomicName id="4C314D6AFFC1FFF10301FC57A2A99408" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atacama" status="sp. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF10301FC57A2A99408" italics="true" pageId="9" pageNumber="10">Rumikiru atacama</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF10783FBB1A2E69408" box="[178,235,1050,1073]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
, ♂ (MACN-Ar).
|
||
<emphasis id="B945EAFBFFC1FFF10692FBB1A3B4940B" bold="true" box="[419,441,1050,1074]" pageId="9" pageNumber="10">B.</emphasis>
|
||
<taxonomicName id="4C314D6AFFC1FFF106F0FBB2A1BC940B" authority="(Ojanguren-Affilastro, 2003)" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" baseAuthorityName="Ojanguren-Affilastro" baseAuthorityYear="2003" box="[449,945,1049,1074]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lourencoi" status="comb. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF106F0FBB2A08E9408" box="[449,643,1049,1073]" italics="true" pageId="9" pageNumber="10">Rumikiru lourencoi</emphasis>
|
||
(Ojanguren-Affilastro, 2003)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF1048CFBB1A6179408" box="[957,1050,1049,1073]" pageId="9" pageNumber="10" rank="species">n. comb.</taxonomicNameLabel>
|
||
, ♂ (MACN-Ar). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32B6562FFC1FFF10799F983A12B965E" pageId="9" pageNumber="10" type="reference_group">
|
||
<paragraph id="8B8E36E9FFC1FFF10799F983A12B965E" blockId="9.[120,1225,1165,1752]" pageId="9" pageNumber="10">
|
||
INCLUDED SPECIES: This genus contains two species:
|
||
<taxonomicName id="4C314D6AFFC1FFF1047AF983A623967B" ID-CoL="4TQ5V" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[843,1070,1576,1602]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atacama" status="sp. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF1047AF983A623967B" box="[843,1070,1576,1602]" italics="true" pageId="9" pageNumber="10">Rumikiru atacama</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF10373F982A684967A" box="[1090,1161,1577,1603]" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C314D6AFFC1FFF10749F9E6A0BE9651" ID-CoL="4TQ5W" authority="(Ojanguren-Affilastro, 2003)" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" baseAuthorityName="Ojanguren-Affilastro" baseAuthorityYear="2003" box="[120,691,1613,1640]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lourencoi" status="comb. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF10749F9E6A354965E" box="[120,345,1613,1639]" italics="true" pageId="9" pageNumber="10">Rumikiru lourencoi</emphasis>
|
||
(Ojanguren-Affilastro, 2003)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF1058EF9E5A12B965E" box="[703,806,1613,1640]" pageId="9" pageNumber="10" rank="species">n. comb.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6562FFC1FFF20799F9D8A194917F" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="distribution">
|
||
<paragraph id="8B8E36E9FFC1FFF20799F9D8A194917F" blockId="9.[120,1225,1165,1752]" lastBlockId="10.[689,1224,225,887]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
DISTRIBUTION:
|
||
<taxonomicName id="4C314D6AFFC1FFF10657F9D8A3D896B4" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[358,469,1651,1677]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC1FFF10657F9D8A3D896B4" box="[358,469,1651,1677]" italics="true" pageId="9" pageNumber="10">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC1FFF106D7F9D8A03796B4" box="[486,570,1651,1677]" pageId="9" pageNumber="10" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, is endemic to the
|
||
<taxonomicName id="4C314D6AFFC1FFF10416F9D8A1EA96B4" authority="Desert" authorityName="Desert" box="[807,999,1651,1677]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="atacama">Atacama Desert</taxonomicName>
|
||
of northern
|
||
<collectingCountry id="F3267679FFC1FFF103B9F9D8A6C596B4" box="[1160,1224,1651,1677]" name="Chile" pageId="9" pageNumber="10">Chile</collectingCountry>
|
||
(figs. 2, 3, 4A). All known records are located in the
|
||
<collectingRegion id="49F5F80BFFC1FFF105E2F933A1C6968B" box="[723,971,1688,1714]" country="Chile" name="Antofagasta" pageId="9" pageNumber="10">Antofagasta Province</collectingRegion>
|
||
of Region II (
|
||
<collectingRegion id="49F5F80BFFC1FFF1035BF933A2B896E1" country="Chile" name="Antofagasta" pageId="9" pageNumber="10">Antofagasta</collectingRegion>
|
||
) and the Chañaral and Huasco provinces of Region III (
|
||
<collectingRegion id="49F5F80BFFC1FFF10453F915A1DD96E1" box="[866,976,1726,1752]" country="Chile" name="Atacama" pageId="9" pageNumber="10">Atacama</collectingRegion>
|
||
). The distribution of this genus is allopatric with that of its sister genus,
|
||
<taxonomicName id="4C314D6AFFC2FFF20433FEACA1729118" authorityName="Ochoa" authorityYear="2004" box="[770,895,263,289]" class="Arachnida" family="Bothriuridae" genus="Pachakutej" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFC2FFF20433FEACA1729118" box="[770,895,263,289]" italics="true" pageId="10" pageNumber="11">Pachakutej</emphasis>
|
||
</taxonomicName>
|
||
, from the inter-Andean valleys of
|
||
<collectingCountry id="F3267679FFC2FFF20434FE86A131917E" box="[773,828,301,327]" name="Peru" pageId="10" pageNumber="11">Peru</collectingCountry>
|
||
(fig. 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B6562FFC2FFF205D0FEF9A1C794FF" pageId="10" pageNumber="11" type="biology_ecology">
|
||
<paragraph id="8B8E36E9FFC2FFF205D0FEF9A00F93F8" blockId="10.[689,1224,225,887]" lastBlockId="10.[120,1225,897,1223]" pageId="10" pageNumber="11">
|
||
ECOLOGY. In addition to its unusual morphology,
|
||
<taxonomicName id="4C314D6AFFC2FFF2041DFEDCA19291A8" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[812,927,375,401]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC2FFF2041DFEDCA19291A8" box="[812,927,375,401]" italics="true" pageId="10" pageNumber="11">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC2FFF20488FEDCA61A91A8" box="[953,1047,375,401]" pageId="10" pageNumber="11" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, occupies an unusual habitat among bothriurid scorpions. Most bothriurids are fossorial, requiring exposed soil to construct burrows. However, all personally collected specimens of
|
||
<taxonomicName id="4C314D6AFFC2FFF20580FD9AA1129272" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[689,799,561,587]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC2FFF20580FD9AA1129272" box="[689,799,561,587]" italics="true" pageId="10" pageNumber="11">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC2FFF2041EFD99A18F9275" box="[815,898,562,588]" pageId="10" pageNumber="11" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, were found in rocky habitats, with almost no exposed soil. At Pan de Azúcar National Park, the
|
||
<typeStatus id="548A884BFFC2FFF204C2FDD6A62B92AE" box="[1011,1062,637,663]" pageId="10" pageNumber="11">type</typeStatus>
|
||
locality of
|
||
<emphasis id="B945EAFBFFC2FFF2039EFDD6A1119282" italics="true" pageId="10" pageNumber="11">
|
||
<taxonomicName id="4C314D6AFFC2FFF2039EFDD6A6C592AF" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[1199,1224,637,662]" pageId="10" pageNumber="11" status="comb. nov.">R.</taxonomicName>
|
||
lourencoi
|
||
</emphasis>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC2FFF2041EFD09A1919282" box="[815,924,673,700]" pageId="10" pageNumber="11" rank="species">n. comb.</taxonomicNameLabel>
|
||
, several specimens were captured on vegetationless scree slopes com- prising piles of sharp, loose stones accumu- lated below steep cliff faces. At Llanos de Challe National Park, specimens of
|
||
<emphasis id="B945EAFBFFC2FFF2035BFC93A0E8934F" italics="true" pageId="10" pageNumber="11">R. atac- ama</emphasis>
|
||
, n. sp., were collected in a slightly more vegetated environment, but this species was common only on scree slopes, cliff faces, and exposed rocky outcrops (fig. 4A).
|
||
</paragraph>
|
||
<caption id="DF4E6661FFC2FFF20749FD79A3309365" ID-DOI="http://doi.org/10.5281/zenodo.5866306" ID-Zenodo-Dep="5866306" httpUri="https://zenodo.org/record/5866306/files/figure.png" pageId="10" pageNumber="11" startId="10.[120,207,722,743]" targetBox="[120,665,226,695]" targetPageId="10">
|
||
<paragraph id="8B8E36E9FFC2FFF20749FD79A3309365" blockId="10.[120,667,720,861]" pageId="10" pageNumber="11">
|
||
FIGURE 7.
|
||
<taxonomicName id="4C314D6AFFC2FFF207C4FD7BA35B92D1" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[245,342,720,744]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC2FFF207C4FD7BA35B92D1" box="[245,342,720,744]" italics="true" pageId="10" pageNumber="11">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC2FFF20657FD7BA3BD92D1" box="[358,432,720,744]" pageId="10" pageNumber="11" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, telson, lateral aspect, showing pigmentation pattern.
|
||
<emphasis id="B945EAFBFFC2FFF20688FD46A3DF933C" bold="true" box="[441,466,749,773]" pageId="10" pageNumber="11">A.</emphasis>
|
||
<taxonomicName id="4C314D6AFFC2FFF206E8FD46A099933C" box="[473,660,749,773]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="atacama">
|
||
<emphasis id="B945EAFBFFC2FFF206E8FD46A099933C" box="[473,660,749,773]" italics="true" pageId="10" pageNumber="11">Rumikiru atacama</emphasis>
|
||
</taxonomicName>
|
||
, n. sp.,
|
||
<specimenCount id="9D37FD60FFC2FFF2078FFCA0A2DE931A" box="[190,211,779,803]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
|
||
(MACN-Ar).
|
||
<emphasis id="B945EAFBFFC2FFF2065AFCA1A38C931B" bold="true" box="[363,385,778,802]" pageId="10" pageNumber="11">B.</emphasis>
|
||
<taxonomicName id="4C314D6AFFC2FFF206BBFCA1A040931B" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" baseAuthorityName="Ojanguren-Affilastro" baseAuthorityYear="2003" box="[394,589,778,802]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lourencoi">
|
||
<emphasis id="B945EAFBFFC2FFF206BBFCA1A040931B" box="[394,589,778,802]" italics="true" pageId="10" pageNumber="11">Rumikiru lourencoi</emphasis>
|
||
</taxonomicName>
|
||
(Ojan- guren-Affilastro, 2003), n. comb.,
|
||
<specimenCount id="9D37FD60FFC2FFF206DEFC83A0099379" box="[495,516,808,832]" pageId="10" pageNumber="11" type="male">♂</specimenCount>
|
||
(MACN-Ar). Scale bars =
|
||
<quantity id="4CC99B0CFFC2FFF207C8FCEEA3359365" box="[249,312,837,861]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8E36E9FFC2FFF20799FC67A6439408" blockId="10.[120,1225,897,1223]" pageId="10" pageNumber="11">
|
||
The environment inhabited by
|
||
<taxonomicName id="4C314D6AFFC2FFF2052CFC67A08693DF" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[541,651,972,998]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC2FFF2052CFC67A08693DF" box="[541,651,972,998]" italics="true" pageId="10" pageNumber="11">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC2FFF205ADFC66A0E293DE" box="[668,751,973,999]" pageId="10" pageNumber="11" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, differs markedly from that of its sister genus,
|
||
<taxonomicName id="4C314D6AFFC2FFF207F8FC5AA34A9432" authorityName="Ochoa" authorityYear="2004" box="[201,327,1009,1035]" class="Arachnida" family="Bothriuridae" genus="Pachakutej" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B945EAFBFFC2FFF207F8FC5AA34A9432" box="[201,327,1009,1035]" italics="true" pageId="10" pageNumber="11">Pachakutej</emphasis>
|
||
</taxonomicName>
|
||
, which occurs under stones in more humid habitats, in inter-Andean valleys and montane rainforests on the eastern slopes of the Andes in
|
||
<collectingCountry id="F3267679FFC2FFF2046FFBBCA1989408" box="[862,917,1047,1073]" name="Peru" pageId="10" pageNumber="11">Peru</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFA04B18FFC2FFF20494FBBCA64F9408" author="Ochoa, J. A." box="[933,1090,1047,1073]" pageId="10" pageNumber="11" pagination="43 - 73" refId="ref14266" refString="Ochoa, J. A. 2004. Filogenia del genero Orobothriurus y descripcion de un nuevo genero de Bothriuridae (Scorpiones). Revista Iberica de Aracnologia 9: 43 - 73." type="journal article" year="2004">Ochoa, 2004</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8E36E9FFC2FFF20799FB97A1C794FF" blockId="10.[120,1225,897,1223]" pageId="10" pageNumber="11">
|
||
<taxonomicName id="4C314D6AFFC2FFF20799FB97A31B946F" authority="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini, 2012" authorityName="Ojanguren-Affilastro & Mattoni & Ochoa & Prendini" authorityYear="2012" box="[168,278,1084,1110]" class="Arachnida" family="Bothriuridae" genus="Rumikiru" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B945EAFBFFC2FFF20799FB97A31B946F" box="[168,278,1084,1110]" italics="true" pageId="10" pageNumber="11">Rumikiru</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2765780FFC2FFF20615FB96A378946E" box="[292,373,1085,1111]" pageId="10" pageNumber="11" rank="genus">n. gen.</taxonomicNameLabel>
|
||
, displays an unusual defensive behavior, not reported among other scorpions, which typically curve the metasoma and strike out with the aculeus and, in some species, the pedipalp chelae, when threatened: males spread the pedipalps wide open, making short pulses or vibrations with them, without curving the metasoma (fig. 4B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |