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<document id="D09A004B2E6C13F3136C57AC1E61BF4F" ID-CLB-Dataset="57159" ID-DOI="10.31610/zsr/2019.28.1.24" ID-GBIF-Dataset="2b690c17-f7fc-4c69-83d1-1d9047c876c9" ID-ISSN="2410-0226" ID-Zenodo-Dep="10124208" ID-ZooBank="urn:lsid:zoobank.org:pub:D82459C6-A090-4203-8D28-8BE062383D2B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1688479652846" checkinUser="felipe" docAuthor="Shamshev, I. V." docDate="2019" docId="039887FDFFD0FFA2783B196475BAF80D" docLanguage="en" docName="ZoosystRoss.28.1.24-41.pdf" docOrigin="Zoosystematica Rossica (Zoosyst. Rossica) 28 (1)" docSource="http://dx.doi.org/10.31610/zsr/2019.28.1.24" docStyle="DocumentStyle:8B7E50B51175314E898B46063364B751.1:ZoosystRoss.2019-.journal_article" docStyleId="8B7E50B51175314E898B46063364B751" docStyleName="ZoosystRoss.2019-.journal_article" docStyleVersion="1" docTitle="Empis (Polyblepharis) yakovlevi Shamshev 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="35" masterDocId="FFA1FF85FFD8FFA97B121C0A7019FF88" masterDocTitle="Six new species of the genus Empis (Diptera: Empididae) from the Altai Mountains of Russia" masterLastPageNumber="41" masterPageNumber="24" pageNumber="32" updateTime="1700049859112" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="39A9859A89648D44B36FDEECCB822885">Six new species of the genus Empis (Diptera: Empididae) from the Altai Mountains of Russia</mods:title>
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<mods:title id="9B542889A8675F72A9E1C3895C2BB8E4">Zoosystematica Rossica</mods:title>
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<mods:date id="F74E003D701022416A1F6BF26E12C90A">2019</mods:date>
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<treatment id="039887FDFFD0FFA2783B196475BAF80D" ID-DOI="http://doi.org/10.5281/zenodo.8114719" ID-GBIF-Taxon="210810427" ID-Zenodo-Dep="8114719" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D" httpUri="http://treatment.plazi.org/id/039887FDFFD0FFA2783B196475BAF80D" lastPageId="11" lastPageNumber="35" pageId="8" pageNumber="32">
<subSubSection id="C32B6560FFD0FFA1783B19647551FA01" box="[809,1352,1390,1417]" pageId="8" pageNumber="32" type="nomenclature">
<paragraph id="8B8E36EBFFD0FFA1783B19647551FA01" blockId="8.[809,1352,1390,1417]" box="[809,1352,1390,1417]" pageId="8" pageNumber="32">
<heading id="D0C68187FFD0FFA1783B19647551FA01" box="[809,1352,1390,1417]" fontSize="11" level="2" pageId="8" pageNumber="32" reason="2">
<emphasis id="B945EAF9FFD0FFA1783B19647551FA01" bold="true" box="[809,1352,1390,1417]" pageId="8" pageNumber="32">
<taxonomicName id="4C314D68FFD0FFA1783B196474C5FA01" ID-CoL="8BZQL" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[809,1244,1390,1417]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="yakovlevi" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B945EAF9FFD0FFA1783B19647362FA00" bold="true" box="[809,891,1390,1416]" italics="true" pageId="8" pageNumber="32">Empis</emphasis>
(
<emphasis id="B945EAF9FFD0FFA1789C1964745EFA01" bold="true" box="[910,1095,1390,1417]" italics="true" pageId="8" pageNumber="32">Polyblepharis</emphasis>
)
<emphasis id="B945EAF9FFD0FFA17F49196474C5FA01" bold="true" box="[1115,1244,1390,1417]" italics="true" pageId="8" pageNumber="32">yakovlevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765782FFD0FFA17FF619657551FA01" box="[1252,1352,1391,1417]" pageId="8" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C32B6560FFD0FFA2783B19987212F898" lastPageId="11" lastPageNumber="35" pageId="8" pageNumber="32" type="description">
<paragraph id="8B8E36EBFFD0FFA1783B199873D5FA24" blockId="8.[809,972,1426,1452]" box="[809,972,1426,1452]" pageId="8" pageNumber="32">
(
<figureCitation id="130A2A6EFFD0FFA17820199873DDFA24" box="[818,964,1426,1452]" captionStart="Figs 1012" captionStartId="9.[159,211,1772,1796]" captionTargetBox="[202,1398,201,1725]" captionTargetId="figure-22@9.[159,1444,181,1731]" captionTargetPageId="9" captionText="Figs 1012. Empis (Polyblepharis) yakovlevi sp. nov., male. 10, habitus, lateral view (holotype); 11, terminalia, lateral view; 12, phallus, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135539" httpUri="https://zenodo.org/record/10135539/files/figure.png" pageId="8" pageNumber="32">Figs 1012</figureCitation>
)
</paragraph>
<paragraph id="8B8E36EBFFD0FFA1785D19C07473F9EC" blockId="8.[809,1429,1482,1926]" pageId="8" pageNumber="32">
<materialsCitation id="3B593CB6FFD0FFA1785D19C07473F9EC" ID-GBIF-Occurrence="4144857321" collectionCode="V, ZIN, INS" country="Russia" latitude="49.716667" location="Republic" longLatPrecision="1101" longitude="89.183334" municipality="Male" pageId="8" pageNumber="32" specimenCount="1" stateProvince="Altay" typeStatus="holotype">
<emphasis id="B945EAF9FFD0FFA1785D19C073ACFA6B" box="[847,949,1482,1507]" italics="true" pageId="8" pageNumber="32">
<typeStatus id="548A8849FFD0FFA1785D19C073ACFA6B" box="[847,949,1482,1507]" pageId="8" pageNumber="32" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingMunicipality id="6BEAAC91FFD0FFA178D019C073E6FA6B" box="[962,1023,1482,1507]" pageId="8" pageNumber="32">Male</collectingMunicipality>
,
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<collectingCountry id="F326767BFFD0FFA17F1819C17445FA6B" box="[1034,1116,1483,1507]" name="Russia" pageId="8" pageNumber="32">Russia</collectingCountry>
</emphasis>
,
<emphasis id="B945EAF9FFD0FFA17F7B19C07533FA6B" box="[1129,1322,1482,1507]" italics="true" pageId="8" pageNumber="32">
<collectingRegion id="49F5F809FFD0FFA17F7B19C07533FA6B" box="[1129,1322,1482,1507]" country="Russia" name="Altay" pageId="8" pageNumber="32">
<location id="8EEE6030FFD0FFA17F7B19C074D4FA6B" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D:8EEE6030FFD0FFA17F7B19C074D4FA6B" box="[1129,1229,1482,1507]" country="Russia" latitude="49.716667" longLatPrecision="1101" longitude="89.183334" municipality="Male" name="Republic" pageId="8" pageNumber="32" stateProvince="Altay">Republic</location>
of Altai
</collectingRegion>
</emphasis>
, “[in Cyrillic, Russian] Tashanta
<quantity id="4CC99B0EFFD0FFA17F5B19E1749BF98C" box="[1097,1154,1515,1540]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="8" pageNumber="32" unit="km" value="4.0">4 km</quantity>
E [~
<geoCoordinate id="EE05502CFFD0FFA17FAA19E17501F98B" box="[1208,1304,1515,1539]" degrees="49" direction="north" minutes="43" orientation="latitude" pageId="8" pageNumber="32" precision="925" value="49.716667">49°43'N</geoCoordinate>
<geoCoordinate id="EE05502CFFD0FFA17E0C19E17560F98B" box="[1310,1401,1515,1539]" degrees="89" direction="east" minutes="11" orientation="longitude" pageId="8" pageNumber="32" precision="925" value="89.183334">89°11'E</geoCoordinate>
] / south-eastern Altai /
<collectionCode id="ED20AE2EFFD0FFA17F351A067425F9AB" box="[1063,1084,1548,1571]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="32" type="Museum">V</collectionCode>
. Richter 6 vii [1]977”, “
<taxonomicName id="4C314D68FFD0FFA17E421A0674DEF9CC" authority="Shamshev" authorityName="Shamshev" authorityYear="2019" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="32" phylum="Arthropoda" rank="species" species="yakovlevi" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B945EAF9FFD0FFA17E421A06758CF9AB" box="[1360,1429,1548,1571]" italics="true" pageId="8" pageNumber="32">Empis</emphasis>
(
<emphasis id="B945EAF9FFD0FFA178211A2173D5F9CC" box="[819,972,1579,1604]" italics="true" pageId="8" pageNumber="32">Polyblepharis</emphasis>
)
<emphasis id="B945EAF9FFD0FFA178CD1A217452F9CC" box="[991,1099,1579,1604]" italics="true" pageId="8" pageNumber="32">yakovlevi</emphasis>
Shamshev
</taxonomicName>
,
<taxonomicNameLabel id="A2765782FFD0FFA17FC01A267514F9CB" box="[1234,1293,1580,1603]" pageId="8" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
[red label]” (
<collectionCode id="ED20AE2EFFD0FFA178231A46737FF9EC" box="[817,870,1612,1636]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="32" type="Museum">ZIN</collectionCode>
,
<collectionCode id="ED20AE2EFFD0FFA178631A4173B8F9EC" box="[881,929,1611,1636]" pageId="8" pageNumber="32">INS</collectionCode>
_DIP_0000725).
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</paragraph>
<paragraph id="8B8E36EBFFD0FFA3785D1A6772B8FE7A" blockId="8.[809,1429,1482,1926]" lastBlockId="10.[144,764,184,1927]" lastPageId="10" lastPageNumber="34" pageId="8" pageNumber="32">
<materialsCitation id="3B593CB6FFD0FFA1785D1A677327F8ED" ID-GBIF-Occurrence="4144857314" collectingDate="1964-07-24" collectingDateMax="2008-07-12" collectingDateMin="1964-07-24" collectionCode="ZIN, V" collectorName="E. Nartshuk &amp; A. Barkalov" country="Russia" county="Same" elevation="2450" latitude="49.3" location="Chikhachev" longLatPrecision="5567" longitude="87.65" municipality="Republic" pageId="8" pageNumber="32" specimenCount="47" specimenCount-female="16" specimenCount-male="31" stateProvince="Altay" typeStatus="paratype">
<emphasis id="B945EAF9FFD0FFA1785D1A6773D8F90C" box="[847,961,1645,1668]" italics="true" pageId="8" pageNumber="32">
<typeStatus id="548A8849FFD0FFA1785D1A6773D8F90C" box="[847,961,1645,1668]" pageId="8" pageNumber="32" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCounty id="62EF4E67FFD0FFA178C11A667408F90C" box="[979,1041,1644,1668]" pageId="8" pageNumber="32">Same</collectingCounty>
data as for holotype:
<specimenCount id="9D37FD62FFD0FFA17E371A617589F90C" box="[1317,1424,1643,1668]" pageId="8" pageNumber="32" type="male">15 males</specimenCount>
,
<specimenCount id="9D37FD62FFD0FFA1783B1A867382F92D" box="[809,923,1676,1701]" pageId="8" pageNumber="32" type="female">2 females</specimenCount>
,
<date id="FF8F102BFFD0FFA178BE1A867431F92D" box="[940,1064,1676,1701]" pageId="8" pageNumber="32" value="1977-07-06">
<collectingDate id="EFCBE9C3FFD0FFA178BE1A867431F92D" box="[940,1064,1676,1701]" pageId="8" pageNumber="32" value="1977-07-06">6.VII.1977</collectingDate>
</date>
;
<specimenCount id="9D37FD62FFD0FFA17F2B1A867495F92D" box="[1081,1164,1676,1701]" pageId="8" pageNumber="32" type="male">1 male</specimenCount>
,
<specimenCount id="9D37FD62FFD0FFA17F8C1A86751FF92D" box="[1182,1286,1676,1701]" pageId="8" pageNumber="32" type="female">1 female</specimenCount>
,
<date id="FF8F102BFFD0FFA17E041A867589F92C" box="[1302,1424,1676,1700]" pageId="8" pageNumber="32" value="1977-07-07">
<collectingDate id="EFCBE9C3FFD0FFA17E041A867589F92C" box="[1302,1424,1676,1700]" pageId="8" pageNumber="32" value="1977-07-07">7.VII.1977</collectingDate>
</date>
;
<specimenCount id="9D37FD62FFD0FFA1783B1AA77388F94D" box="[809,913,1708,1733]" pageId="8" pageNumber="32" type="female">1 female</specimenCount>
,
<date id="FF8F102BFFD0FFA178B11AA67435F94D" box="[931,1068,1708,1733]" pageId="8" pageNumber="32" value="1977-07-15">
<collectingDate id="EFCBE9C3FFD0FFA178B11AA67435F94D" box="[931,1068,1708,1733]" pageId="8" pageNumber="32" value="1977-07-15">15.VII.1977</collectingDate>
</date>
.
<emphasis id="B945EAF9FFD0FFA17F2C1AA67489F94C" bold="true" box="[1086,1168,1708,1732]" pageId="8" pageNumber="32">
<collectingCountry id="F326767BFFD0FFA17F2C1AA67489F94C" box="[1086,1168,1708,1732]" name="Russia" pageId="8" pageNumber="32">Russia</collectingCountry>
</emphasis>
,
<emphasis id="B945EAF9FFD0FFA17FB61AA6756BF94D" box="[1188,1394,1708,1733]" italics="true" pageId="8" pageNumber="32">
<collectingMunicipality id="6BEAAC91FFD0FFA17FB61AA67511F94D" box="[1188,1288,1708,1733]" pageId="8" pageNumber="32">Republic</collectingMunicipality>
of
<collectingRegion id="49F5F809FFD0FFA17E281AA6756BF94D" box="[1338,1394,1708,1733]" country="Russia" name="Altay" pageId="8" pageNumber="32">Altai</collectingRegion>
</emphasis>
:
<specimenCount id="9D37FD62FFD0FFA17E951AA77375F96D" pageId="8" pageNumber="32" type="male">2 males</specimenCount>
,
<specimenCount id="9D37FD62FFD0FFA178651AC773FBF96D" box="[887,994,1740,1765]" pageId="8" pageNumber="32" type="female">4 females</specimenCount>
,
<location id="8EEE6030FFD0FFA178FF1AC67461F96D" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D:8EEE6030FFD0FFA178FF1AC67461F96D" box="[1005,1144,1740,1765]" country="Russia" county="Same" latitude="49.3" longLatPrecision="5567" longitude="87.65" municipality="Republic" name="Chikhachev" pageId="8" pageNumber="32" stateProvince="Altay">Chikhachev</location>
ridge, pass,
<date id="FF8F102BFFD0FFA17E111AC67589F96D" box="[1283,1424,1740,1765]" pageId="8" pageNumber="32" value="1964-07-24">
<collectingDate id="EFCBE9C3FFD0FFA17E111AC67589F96D" box="[1283,1424,1740,1765]" pageId="8" pageNumber="32" value="1964-07-24">24.VII.1964</collectingDate>
</date>
,
<collectorName id="26C4533DFFD0FFA1783B1AE473AFF88D" box="[809,950,1772,1797]" pageId="8" pageNumber="32">E. Nartshuk</collectorName>
leg. (all
<collectionCode id="ED20AE2EFFD0FFA17F081AE77449F88D" box="[1050,1104,1773,1797]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="8" pageNumber="32" type="Museum">ZIN</collectionCode>
);
<specimenCount id="9D37FD62FFD0FFA17F771AE774D2F88D" box="[1125,1227,1772,1797]" pageId="8" pageNumber="32" type="male">13 males</specimenCount>
,
<specimenCount id="9D37FD62FFD0FFA17FC51AE7755DF88D" box="[1239,1348,1772,1797]" pageId="8" pageNumber="32" type="female">8 females</specimenCount>
,
<location id="8EEE6030FFD0FFA17E5D1AE673A4F8AE" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D:8EEE6030FFD0FFA17E5D1AE673A4F8AE" country="Russia" county="Same" latitude="49.3" longLatPrecision="5567" longitude="87.65" municipality="Republic" name="Agach distr." pageId="8" pageNumber="32" stateProvince="Altay">Kosh-Agach distr.</location>
, plateau
<location id="8EEE6030FFD0FFA17F221B07746AF8AE" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D:8EEE6030FFD0FFA17F221B07746AF8AE" box="[1072,1139,1805,1830]" country="Russia" county="Same" latitude="49.3" longLatPrecision="5567" longitude="87.65" municipality="Republic" name="Ukok" pageId="8" pageNumber="32" stateProvince="Altay">Ukok</location>
,
<quantity id="4CC99B0EFFD0FFA17F971B0774FCF8AD" box="[1157,1253,1805,1830]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="8" pageNumber="32" unit="m" value="2450.0">
<elevation id="001CD1D8FFD0FFA17F971B0774FCF8AD" box="[1157,1253,1805,1830]" metricMagnitude="3" metricUnit="m" metricValue="2.45" pageId="8" pageNumber="32" unit="m" value="2450.0">2450 m</elevation>
</quantity>
,
<location id="8EEE6030FFD0FFA17FE41B07737DF8CE" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D:8EEE6030FFD0FFA17FE41B07737DF8CE" country="Russia" county="Same" latitude="49.3" longLatPrecision="5567" longitude="87.65" municipality="Republic" name="Muzdy-Bulak Lake" pageId="8" pageNumber="32" stateProvince="Altay">Muzdy-Bulak Lake</location>
,
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<geoCoordinate id="EE05502CFFD0FFA178DA1B27743DF8CE" box="[968,1060,1837,1862]" degrees="87.65" direction="east" orientation="longitude" pageId="8" pageNumber="32" precision="555" value="87.65">87.65°E</geoCoordinate>
,
<date id="FF8F102BFFD0FFA17F231B2474C6F8CE" box="[1073,1247,1838,1862]" pageId="8" pageNumber="32" value="2008-07-08" valueMax="2008-07-12" valueMin="2008-07-08">
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</date>
,
<collectorName id="26C4533DFFD0FFA17FFE1B24756EF8CE" box="[1260,1399,1837,1862]" pageId="8" pageNumber="32">A. Barkalov</collectorName>
&amp;
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</materialsCitation>
.
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<taxonomicName id="4C314D68FFD0FFA178551B4773B6F8EE" authority="T. Novgorodova" authorityName="P.A.Saccardo" authorityYear="1892" box="[839,943,1869,1894]" class="Pezizomycetes" family="Rhizinaceae" genus="Sorokina" kingdom="Fungi" order="Pezizales" pageId="8" pageNumber="34" phylum="Ascomycota" rank="genus">Sorokina</taxonomicName>
leg. (
<collectionCode id="ED20AE2EFFD0FFA178FF1B44745CF8EE" box="[1005,1093,1870,1894]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="8" pageNumber="32">SZMN</collectionCode>
);
<specimenCount id="9D37FD62FFD0FFA17F4B1B4474A7F8EE" box="[1113,1214,1869,1894]" pageId="8" pageNumber="32" type="male">12 males</specimenCount>
,
<specimenCount id="9D37FD62FFD0FFA17FD81B47752CF8EE" box="[1226,1333,1869,1894]" pageId="8" pageNumber="32" type="female">5 females</specimenCount>
, plateau
<collectingMunicipality id="6BEAAC91FFD0FFA1783B1B677375F80E" box="[809,876,1901,1926]" pageId="8" pageNumber="32">Ukok</collectingMunicipality>
, Kaljin-Kul-Bas and
<location id="8EEE6030FFD0FFA17F7A1B6774FDF80E" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D:8EEE6030FFD0FFA17F7A1B6774FDF80E" box="[1128,1252,1901,1926]" country="Russia" latitude="49.316666" longLatPrecision="1104" longitude="87.433334" municipality="Ukok" name="Kaljin-Kul" pageId="8" pageNumber="32" stateProvince="Altay">Kaljin-Kul</location>
lakes,
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<geoCoordinate id="EE05502CFFD2FFA37B821CB270F7FF58" box="[144,238,184,209]" degrees="87" direction="east" minutes="26" orientation="longitude" pageId="10" pageNumber="34" precision="925" value="87.433334">87°26'E</geoCoordinate>
,
<quantity id="4CC99B0EFFD2FFA37A111CB271A8FF58" box="[259,433,184,209]" metricMagnitude="3" metricUnit="m" metricValue="2.425" metricValueMax="2.45" metricValueMin="2.4" pageId="10" pageNumber="34" unit="m" value="2425.0" valueMax="2450.0" valueMin="2400.0">
<elevation id="001CD1D8FFD2FFA37A111CB271A8FF58" box="[259,433,184,209]" metricMagnitude="3" metricUnit="m" metricValue="2.425" metricValueMax="2.45" metricValueMin="2.4" pageId="10" pageNumber="34" unit="m" value="2425.0" valueMax="2450.0" valueMin="2400.0">24002450 m</elevation>
</quantity>
,
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</date>
,
<collectionCode id="ED20AE2EFFD2FFA379861CB372B0FF58" box="[660,681,185,208]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="34" type="Museum">V</collectionCode>
</materialsCitation>
.
<taxonomicName id="4C314D68FFD2FFA379AE1CB270DAFF79" authority="T. Novgorodova" authorityName="P.A.Saccardo" authorityYear="1892" class="Pezizomycetes" family="Rhizinaceae" genus="Sorokina" kingdom="Fungi" order="Pezizales" pageId="10" pageNumber="34" phylum="Ascomycota" rank="genus">Sorokina</taxonomicName>
&amp; T.
<materialsCitation id="3B593CB6FFD2FFA37A001CD271FDFEB8" ID-GBIF-Occurrence="4144857309" collectingDate="2006-07-08" collectionCode="SZMN, V" collectorName="Novgorodova" country="Russia" elevation="2450" latitude="49.566666" location="Ukok" longLatPrecision="1103" longitude="87.71667" pageId="10" pageNumber="34" specimenCount="2" specimenCount-male="2" stateProvince="Altay" typeStatus="paratype">
<collectorName id="26C4533DFFD2FFA37A001CD271B2FF79" box="[274,427,216,241]" pageId="10" pageNumber="34">Novgorodova</collectorName>
leg. (
<collectionCode id="ED20AE2EFFD2FFA37AE01CD37253FF79" box="[498,586,217,241]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="10" pageNumber="34">SZMN</collectionCode>
);
<specimenCount id="9D37FD62FFD2FFA379711CD372D9FF79" box="[611,704,216,241]" pageId="10" pageNumber="34" type="male">2 males</specimenCount>
, plateau
<location id="8EEE6030FFD2FFA37BDE1CF27116FE99" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D:8EEE6030FFD2FFA37BDE1CF27116FE99" box="[204,271,248,273]" country="Russia" latitude="49.566666" longLatPrecision="1103" longitude="87.71667" name="Ukok" pageId="10" pageNumber="34" stateProvince="Altay">Ukok</location>
,
<locationDeviation id="9A3C2FC9FFD2FFA37A0C1CF3723BFE99" box="[286,546,248,273]" location="Maytobe" pageId="10" pageNumber="34">
<quantity id="4CC99B0EFFD2FFA37A0C1CF37142FE99" box="[286,347,248,273]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="34" unit="km" value="8.0">8 km</quantity>
NE of Maytobe
</locationDeviation>
,
<geoCoordinate id="EE05502CFFD2FFA379231CF37288FE98" box="[561,657,249,273]" degrees="49" direction="north" minutes="34" orientation="latitude" pageId="10" pageNumber="34" precision="925" value="49.566666">49°34'N</geoCoordinate>
<geoCoordinate id="EE05502CFFD2FFA379881CF372EFFE98" box="[666,758,249,273]" degrees="87" direction="east" minutes="43" orientation="longitude" pageId="10" pageNumber="34" precision="925" value="87.71667">87°43'E</geoCoordinate>
,
<quantity id="4CC99B0EFFD2FFA37B821D12712CFEB8" box="[144,309,280,305]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="10" pageNumber="34" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">
<elevation id="001CD1D8FFD2FFA37B821D12712CFEB8" box="[144,309,280,305]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="10" pageNumber="34" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">24002500 m</elevation>
</quantity>
,
<date id="FF8F102BFFD2FFA37A531D1271DAFEB9" box="[321,451,280,305]" pageId="10" pageNumber="34" value="2006-07-08">
<collectingDate id="EFCBE9C3FFD2FFA37A531D1271DAFEB9" box="[321,451,280,305]" pageId="10" pageNumber="34" value="2006-07-08">8.VII.2006</collectingDate>
</date>
,
<collectionCode id="ED20AE2EFFD2FFA37ADD1D1371FDFEB8" box="[463,484,281,304]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="10" pageNumber="34" type="Museum">V</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B593CB6FFD2FFA37AFD1D1270BAFE19" ID-GBIF-Occurrence="4144857305" collectingDate="2006-07-10" collectionCode="SZMN, T" country="Russia" elevation="2500" latitude="49.566666" location="Ildegem River" longLatPrecision="1103" longitude="87.71667" pageId="10" pageNumber="34" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Altay" typeStatus="paratype">
<taxonomicName id="4C314D68FFD2FFA37AFD1D12724EFEB9" authority="T. Novgorodova" authorityName="P.A.Saccardo" authorityYear="1892" box="[495,599,280,305]" class="Pezizomycetes" family="Rhizinaceae" genus="Sorokina" kingdom="Fungi" order="Pezizales" pageId="10" pageNumber="34" phylum="Ascomycota" rank="genus">Sorokina</taxonomicName>
leg. (
<collectionCode id="ED20AE2EFFD2FFA379851D1372F6FEB9" box="[663,751,281,305]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="10" pageNumber="34">SZMN</collectionCode>
);
<specimenCount id="9D37FD62FFD2FFA37B821D3370C6FED9" box="[144,223,312,337]" pageId="10" pageNumber="34" type="male">1 male</specimenCount>
,
<specimenCount id="9D37FD62FFD2FFA37BFF1D337142FED9" box="[237,347,312,337]" pageId="10" pageNumber="34" type="female">2 females</specimenCount>
,
<locationDeviation id="9A3C2FC9FFD2FFA37A7A1D33727FFED9" box="[360,614,312,337]" location="Maytobe" pageId="10" pageNumber="34">
<quantity id="4CC99B0EFFD2FFA37A7A1D3371BDFED9" box="[360,420,312,337]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="34" unit="km" value="8.0">8 km</quantity>
NE of Maytobe
</locationDeviation>
, tributary of
<location id="8EEE6030FFD2FFA37B821D52712BFEF9" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D:8EEE6030FFD2FFA37B821D52712BFEF9" box="[144,306,344,369]" country="Russia" latitude="49.566666" longLatPrecision="1103" longitude="87.71667" name="Ildegem River" pageId="10" pageNumber="34" stateProvince="Altay">Ildegem River</location>
,
<geoCoordinate id="EE05502CFFD2FFA37A2E1D537185FEF8" box="[316,412,345,369]" degrees="49" direction="north" minutes="34" orientation="latitude" pageId="10" pageNumber="34" precision="925" value="49.566666">49°34'N</geoCoordinate>
<geoCoordinate id="EE05502CFFD2FFA37AB21D5371E5FEF8" box="[416,508,345,369]" degrees="87" direction="east" minutes="43" orientation="longitude" pageId="10" pageNumber="34" precision="925" value="87.71667">87°43'E</geoCoordinate>
,
<quantity id="4CC99B0EFFD2FFA379151D527246FEF8" box="[519,607,344,369]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="34" unit="m" value="2500.0">
<elevation id="001CD1D8FFD2FFA379151D527246FEF8" box="[519,607,344,369]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="34" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="FF8F102BFFD2FFA3797B1D5272EEFEF9" box="[617,759,344,369]" pageId="10" pageNumber="34" value="2006-07-10">
<collectingDate id="EFCBE9C3FFD2FFA3797B1D5272EEFEF9" box="[617,759,344,369]" pageId="10" pageNumber="34" value="2006-07-10">10.VII.2006</collectingDate>
</date>
,
<collectionCode id="ED20AE2EFFD2FFA37B821D7070BAFE19" box="[144,163,378,401]" name="Tavera, Department of Geology and Geophysics" pageId="10" pageNumber="34">T</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B593CB6FFD2FFA37BA41D7270CBFE7A" ID-GBIF-Occurrence="4144857303" collectionCode="SZMN, VI" collectorName="Novgorodova" country="Russia" county="Ust' - Koksa distr." elevation="2332" location="Kamdyt" municipality="Katunskiy" pageId="10" pageNumber="34" specimenCount="1" specimenCount-male="1" stateProvince="Altay" typeStatus="paratype">
<collectorName id="26C4533DFFD2FFA37BA41D727156FE19" box="[182,335,376,401]" pageId="10" pageNumber="34">Novgorodova</collectorName>
leg. (
<collectionCode id="ED20AE2EFFD2FFA37A8F1D7371ECFE19" box="[413,501,377,401]" country="Russia" httpUri="http://grbio.org/cool/wy0h-yads" name="Siberian Zoological Museum" pageId="10" pageNumber="34">SZMN</collectionCode>
);
<specimenCount id="9D37FD62FFD2FFA379001D73727EFE19" box="[530,615,376,401]" pageId="10" pageNumber="34" type="male">1 male</specimenCount>
,
<collectingCounty id="62EF4E67FFD2FFA379691D7270D4FE3A" pageId="10" pageNumber="34">Ust-Koksa distr.</collectingCounty>
,
<collectingMunicipality id="6BEAAC91FFD2FFA37BF61D937144FE3A" box="[228,349,409,434]" pageId="10" pageNumber="34">Katunskiy</collectingMunicipality>
ridge,
<location id="8EEE6030FFD2FFA37AD11D93723DFE3A" LSID="urn:lsid:plazi:treatment:039887FDFFD0FFA2783B196475BAF80D:8EEE6030FFD2FFA37AD11D93723DFE3A" box="[451,548,409,434]" country="Russia" county="Ust' - Koksa distr." municipality="Katunskiy" name="Kamdyt" pageId="10" pageNumber="34" stateProvince="Altay">Kamdyt</location>
, 49°58.749.14'N 86°3334.9'E,
<quantity id="4CC99B0EFFD2FFA37A531DB371F8FE59" box="[321,481,441,466]" metricMagnitude="3" metricUnit="m" metricValue="2.332" metricValueMax="2.514" metricValueMin="2.15" pageId="10" pageNumber="34" unit="m" value="2332.0" valueMax="2514.0" valueMin="2150.0">
<elevation id="001CD1D8FFD2FFA37A531DB371F8FE59" box="[321,481,441,466]" metricMagnitude="3" metricUnit="m" metricValue="2.332" metricValueMax="2.514" metricValueMin="2.15" pageId="10" pageNumber="34" unit="m" value="2332.0" valueMax="2514.0" valueMin="2150.0">21502514 m</elevation>
</quantity>
, subalpine-alpine [zone],
<date id="FF8F102BFFD2FFA37B821DD070CBFE7A" box="[144,210,474,498]" pageId="10" pageNumber="34" value="2007-06-28" valueMax="2007-07-04" valueMin="2007-06-28">
28.
<collectionCode id="ED20AE2EFFD2FFA37BA31DD070CBFE7A" box="[177,210,474,498]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="10" pageNumber="34">VI</collectionCode>
</date>
</materialsCitation>
4.VII.2007,
<collectorName id="26C4533DFFD2FFA37A7E1DD071EDFE7A" box="[364,500,473,498]" pageId="10" pageNumber="34">O. Kosterin</collectorName>
leg. (ZMUM).
</paragraph>
<caption id="DF4E6663FFD1FFA07B8D1AE672CCF8CB" ID-DOI="http://doi.org/10.5281/zenodo.10135539" ID-Zenodo-Dep="10135539" httpUri="https://zenodo.org/record/10135539/files/figure.png" pageId="9" pageNumber="33" startId="9.[159,211,1772,1796]" targetBox="[202,1398,201,1725]" targetPageId="9" targetType="figure">
<paragraph id="8B8E36EBFFD1FFA07B8D1AE672CCF8CB" blockId="9.[159,1444,1771,1860]" pageId="9" pageNumber="33">
<emphasis id="B945EAF9FFD1FFA07B8D1AE67132F88C" bold="true" box="[159,299,1772,1796]" pageId="9" pageNumber="33">Figs 1012.</emphasis>
<taxonomicName id="4C314D68FFD1FFA07A211AE772BAF88C" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[307,675,1771,1796]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="33" phylum="Arthropoda" rank="species" species="yakovlevi" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B945EAF9FFD1FFA07A211AE77161F88C" box="[307,376,1773,1796]" italics="true" pageId="9" pageNumber="33">Empis</emphasis>
(
<emphasis id="B945EAF9FFD1FFA07A981AE1723AF88C" box="[394,547,1771,1796]" italics="true" pageId="9" pageNumber="33">Polyblepharis</emphasis>
)
<emphasis id="B945EAF9FFD1FFA079251AE172BAF88C" box="[567,675,1771,1796]" italics="true" pageId="9" pageNumber="33">yakovlevi</emphasis>
</taxonomicName>
<emphasis id="B945EAF9FFD1FFA079B81AE6731EF88C" bold="true" box="[682,775,1772,1796]" pageId="9" pageNumber="33">
<taxonomicNameLabel id="A2765782FFD1FFA079B81AE6731EF88C" box="[682,775,1772,1796]" pageId="9" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B945EAF9FFD1FFA0784B1AE6736DF88C" bold="true" box="[857,884,1772,1796]" pageId="9" pageNumber="33">10</emphasis>
, habitus, lateral view (holotype);
<emphasis id="B945EAF9FFD1FFA07FEE1AE6750EF88C" bold="true" box="[1276,1303,1772,1796]" pageId="9" pageNumber="33">11</emphasis>
, terminalia, lateral view;
<emphasis id="B945EAF9FFD1FFA07A221B067155F8AC" bold="true" box="[304,332,1804,1828]" pageId="9" pageNumber="33">12</emphasis>
, phallus, lateral view. Abbreviations:
<emphasis id="B945EAF9FFD1FFA079E31B077307F8AC" box="[753,798,1805,1828]" italics="true" pageId="9" pageNumber="33">cerc</emphasis>
, cercus;
<emphasis id="B945EAF9FFD1FFA0786C1B0773D7F8AC" box="[894,974,1803,1828]" italics="true" pageId="9" pageNumber="33">ej apod</emphasis>
, ejaculatory apodeme;
<emphasis id="B945EAF9FFD1FFA07FC21B01750FF8AC" box="[1232,1302,1803,1828]" italics="true" pageId="9" pageNumber="33">epand</emphasis>
, epandrium;
<emphasis id="B945EAF9FFD1FFA07B8D1B2170CEF8CC" box="[159,215,1835,1860]" italics="true" pageId="9" pageNumber="33">hypd</emphasis>
, hypandrium;
<emphasis id="B945EAF9FFD1FFA07A6F1B217180F8CC" box="[381,409,1835,1860]" italics="true" pageId="9" pageNumber="33">ph</emphasis>
, phallus. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B8E36EBFFD2FFA37BA71DF172CBFC80" blockId="10.[144,764,184,1927]" pageId="10" pageNumber="34">
<emphasis id="B945EAF9FFD2FFA37BA71DF17128FD9C" box="[181,305,507,532]" italics="true" pageId="10" pageNumber="34">Diagnosis</emphasis>
. Large species (body length about
<quantity id="4CC99B0EFFD2FFA37B821E1470C0FDB0" box="[144,217,542,568]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="34" unit="mm" value="8.0">8 mm</quantity>
) with extensively yellow legs almost entirely covered with long dense hair-like setae, face bearing marginal setae, palpus black, scutum with four vittae, laterotergite with black setae, abdominal tergites extensively shiny. Male: eyes almost touching on frons, tarsomeres of fore and mid tarsi somewhat thickened. Female: legs not pinnate.
</paragraph>
<paragraph id="8B8E36EBFFD2FFA37BA71F1871FFFCF9" blockId="10.[144,764,184,1927]" pageId="10" pageNumber="34">
<emphasis id="B945EAF9FFD2FFA37BA71F18715CFCA3" box="[181,325,786,811]" italics="true" pageId="10" pageNumber="34">Description</emphasis>
.
<emphasis id="B945EAF9FFD2FFA37A701F1B71B8FCA3" box="[354,417,785,811]" italics="true" pageId="10" pageNumber="34">Male</emphasis>
(
<figureCitation id="130A2A6EFFD2FFA37AD31F1B7235FCA3" box="[449,556,785,811]" captionStart="Figs 1012" captionStartId="9.[159,211,1772,1796]" captionTargetBox="[202,1398,201,1725]" captionTargetId="figure-22@9.[159,1444,181,1731]" captionTargetPageId="9" captionText="Figs 1012. Empis (Polyblepharis) yakovlevi sp. nov., male. 10, habitus, lateral view (holotype); 11, terminalia, lateral view; 12, phallus, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135539" httpUri="https://zenodo.org/record/10135539/files/figure.png" pageId="10" pageNumber="34">Fig. 10</figureCitation>
). Body length
<quantity id="4CC99B0EFFD2FFA37B821F39713CFCC7" box="[144,293,819,847]" metricMagnitude="-3" metricUnit="m" metricValue="7.85" metricValueMax="8.1" metricValueMin="7.6" pageId="10" pageNumber="34" unit="mm" value="7.85" valueMax="8.1" valueMin="7.6">7.68.1 mm</quantity>
(
<typeStatus id="548A8849FFD2FFA37A271F3E71B0FCC6" box="[309,425,820,846]" pageId="10" pageNumber="34" type="holotype">holotype</typeStatus>
,
<quantity id="4CC99B0EFFD2FFA37AA71F3E720DFCC7" box="[437,532,820,847]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="10" pageNumber="34" unit="mm" value="7.9">7.9 mm</quantity>
); wing length
<quantity id="4CC99B0EFFD2FFA379D51F3E70F2FCF9" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.4" metricValueMin="7.1" pageId="10" pageNumber="34" unit="mm" value="7.25" valueMax="7.4" valueMin="7.1">7.1 7.4 mm</quantity>
(
<typeStatus id="548A8849FFD2FFA37BE81F5D7177FCF9" box="[250,366,855,881]" pageId="10" pageNumber="34" type="holotype">holotype</typeStatus>
,
<quantity id="4CC99B0EFFD2FFA37A6B1F5D71C0FCF9" box="[377,473,855,881]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="10" pageNumber="34" unit="mm" value="7.3">7.3 mm</quantity>
).
</paragraph>
<paragraph id="8B8E36EBFFD2FFA37BA71F7072B4FA6C" blockId="10.[144,764,184,1927]" pageId="10" pageNumber="34">Head black, with black setation. Eyes dichoptic, ommatidia of equal size. Frons very narrow in upper part, at most as broad as anterior ocellus, widened above antennae, densely brownish pruinose, bare. Face wide, almost parallel-sided, largely brownish grey pruinose, narrowly shiny along lower margin, with 34 black marginal setae in upper part; clypeus shiny. Occiput densely greyish pruinose, covered with dense, long, thin setae. Ocellar tubercle concolorous with occiput, with numerous similar setae. Antenna black; scape about 1.5 times as long as pedicel, with black setulae longer than in pedicel; postpedicel moderately long, nearly 3 times as long as wide, with straight margins; stylus about half the postpedicel length. Proboscis moderately long; labrum brownish, 1.31.5 times as long as head height; palpus long, black, with numerous very long black setae.</paragraph>
<paragraph id="8B8E36EBFFD2FFA37BA719E67509FD2B" blockId="10.[144,764,184,1927]" lastBlockId="10.[809,1429,185,1926]" pageId="10" pageNumber="34">Thorax black, densely brownish grey pruinose (except as noted); scutum with four indistinct, broad, brownish vittae (in dorsal view); setation black. Prosternum bare. Proepisternum and antepronotum with numerous uniform long hair-like setae. Postpronotal lobe shiny on outer side, with numerous moderately long to long subequally thin setae. Mesonotal setation: presutural and postsutural surfaces with numerous long setae of different robustness (23 setae usually stronger but highly variable), 46 long notopleurals (number and position highly variable, often even on right and left sides; additional long thin setae present on anterior part of notopleuron), 12 very long and 12 short postalars, numerous scutellars of different robustness (number and position highly variable; at least 7 setae present on each side); acrostichals long, thin, numerous, arranged in 23 irregular rows, lacking on prescutellar depression; dorsocentrals arranged in 35 irregular dense rows before suture and in 23-serial sparser rows toward scutellum, somewhat longer than acrostichals, of subequal length and robustness throughout (some prescutellars somewhat longer and stronger). Laterotergite with numerous setae. Anterior and posterior spiracles yellow.</paragraph>
<paragraph id="8B8E36EBFFD2FFA3785D1EA47540FA51" blockId="10.[809,1429,185,1926]" pageId="10" pageNumber="34">Legs with coxae and trochanters black, faintly greyish pruinose; femora and tibiae extensively yellowish, very narrowly brownish at apices (in paler specimens, femora entirely yellowish); tarsi almost uniformly brownish, basitarsi somewhat paler near base. All parts of legs with black setation; coxae and trochanters with numerous long setae. Fore and mid tibiae and tarsi somewhat thickened; hind femur slender, only slightly broader than fore and mid femora. All femora entirely covered with long thin setae; also, hind femur with some stronger anteroventral setae in subapical part. Fore and mid tibiae with similar but denser and longer setae (especially dorsally) spreading on to corresponding tarsomeres 14; both without strong setae (except circlets of subapicals). Hind tibia with shorter thin setae than on preceding tibiae, also bearing 56 anterodorsal and 56 posterodorsal short stronger setae (besides circlet of subapicals); posteroapical comb with one moderately long, strong seta. Hind basitarsus with sparser and somewhat shorter setation dorsally; tarsomeres 25 with simple setulae.</paragraph>
<paragraph id="8B8E36EBFFD2FFA3785D19E9754EF938" blockId="10.[809,1429,185,1926]" pageId="10" pageNumber="34">
Wing membrane brownish infuscate. Pterostigma brownish yellow, narrow. Basal costal seta moderately long. Veins
<emphasis id="B945EAF9FFD2FFA37F801A2174A9F9C3" box="[1170,1200,1579,1611]" italics="true" pageId="10" pageNumber="34">
R
<subScript id="17B534AEFFD2FFA37FB51A3774A9F9C3" attach="left" box="[1191,1200,1597,1611]" fontSize="6" pageId="10" pageNumber="34">5</subScript>
</emphasis>
and
<emphasis id="B945EAF9FFD2FFA37FE01A21750DF9C3" box="[1266,1300,1579,1611]" italics="true" pageId="10" pageNumber="34">
M
<subScript id="17B534AEFFD2FFA37E191A37750DF9C3" attach="left" box="[1291,1300,1597,1611]" fontSize="6" pageId="10" pageNumber="34">1</subScript>
</emphasis>
divergent in subapical parts; radial fork acute. Anal angle acute, subsequently anal lobe well-developed. Calypter brownish, dark fringed. Halter yellow.
</paragraph>
<paragraph id="8B8E36EBFFD2FFA2785D1AB37257FC8F" blockId="10.[809,1429,185,1926]" lastBlockId="11.[159,779,185,1693]" lastPageId="11" lastPageNumber="35" pageId="10" pageNumber="34">
Abdomen entirely black, with tergites (except hidden tergite 8) almost entirely shiny; tergites 1 and 2 narrowly brownish grey pruinose anteriorly, tergites 6 and 7 with brownish iridescent spots; sternites 12 entirely brownish grey pruinose, remaining sternites narrowly pruinose anteriorly. Abdomen covered with uniform, long, dense, black setae, without stronger posteromarginal setae. Sclerites of segment 7 unmodified; tergite 8 separated from sternite 8, divided medially, represented by two subtriangular sclerites bearing scattered setulae; sternite 8 large, scoop-shaped, with three tubercles near base laterally on either side. Terminalia (
<figureCitation id="130A2A6EFFD3FFA27A6E1DBA71C1FE42" box="[380,472,432,458]" captionStart="Figs 1012" captionStartId="9.[159,211,1772,1796]" captionTargetBox="[202,1398,201,1725]" captionTargetId="figure-22@9.[159,1444,181,1731]" captionTargetPageId="9" captionText="Figs 1012. Empis (Polyblepharis) yakovlevi sp. nov., male. 10, habitus, lateral view (holotype); 11, terminalia, lateral view; 12, phallus, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135539" httpUri="https://zenodo.org/record/10135539/files/figure.png" pageId="11" pageNumber="35">Fig. 11</figureCitation>
) moderately large, dark brown, only phallus brownish yellow. Cercus elongate oval (in lateral view), with smoothed inner margins (in dorsal view), covered with black setulae. Epandrial lamella rather subrectangular (in lateral view), covered with black setulae being somewhat longer along lower margin. Hypandrium subtriangular in ventral view, bare. Phallus (
<figureCitation id="130A2A6EFFD3FFA27BBA1EC0711DFD6C" box="[168,260,714,740]" captionStart="Figs 1012" captionStartId="9.[159,211,1772,1796]" captionTargetBox="[202,1398,201,1725]" captionTargetId="figure-22@9.[159,1444,181,1731]" captionTargetPageId="9" captionText="Figs 1012. Empis (Polyblepharis) yakovlevi sp. nov., male. 10, habitus, lateral view (holotype); 11, terminalia, lateral view; 12, phallus, lateral view. Abbreviations: cerc, cercus; ej apod, ejaculatory apodeme; epand, epandrium; hypd, hypandrium; ph, phallus. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10135539" httpUri="https://zenodo.org/record/10135539/files/figure.png" pageId="11" pageNumber="35">Fig. 12</figureCitation>
) divided; apical part of distiphallus hooklike, very slender, moderately long.
</paragraph>
<paragraph id="8B8E36EBFFD3FFA27BD71F1A72A1FBCD" blockId="11.[159,779,185,1693]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27BD71F1A7105FCA2" box="[197,284,784,810]" italics="true" pageId="11" pageNumber="35">Female</emphasis>
. Body length
<quantity id="4CC99B0EFFD3FFA27ADF1F1A7271FCA3" box="[461,616,784,812]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" metricValueMax="6.8" metricValueMin="6.4" pageId="11" pageNumber="35" unit="mm" value="6.6" valueMax="6.8" valueMin="6.4">6.46.8 mm</quantity>
; wing length
<quantity id="4CC99B0EFFD3FFA27B8D1F39712FFCC6" box="[159,310,819,847]" metricMagnitude="-3" metricUnit="m" metricValue="7.15" metricValueMax="7.4" metricValueMin="6.9" pageId="11" pageNumber="35" unit="mm" value="7.15" valueMax="7.4" valueMin="6.9">6.97.4 mm</quantity>
. Similar to male, except the following characters. Frons broad, parallel-sided, with marginal setulae; occiput with sparse setae; thorax and legs with similar but shorter setation; fore and mid tarsomeres slender; abdomen with scattered short setae, posterior part of tergite 2 and entire tergites 38 shiny, sternites 68 subshiny. Cercus slender, moderately long, covered with dark setulae.
</paragraph>
<paragraph id="8B8E36EBFFD3FFA27BD7184571A8FAB5" blockId="11.[159,779,185,1693]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27BD718457142FBE1" box="[197,347,1103,1129]" italics="true" pageId="11" pageNumber="35">Comparison</emphasis>
. The new species is most similar to
<taxonomicName id="4C314D68FFD3FFA27BD6187871F7FB05" authority="Becker, 1915" authorityName="Becker" authorityYear="1915" box="[196,494,1137,1165]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="35" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B945EAF9FFD3FFA27BD6187870CCFB03" box="[196,213,1138,1163]" italics="true" pageId="11" pageNumber="35">E</emphasis>
.
<emphasis id="B945EAF9FFD3FFA27BFA187B7159FB03" box="[232,320,1137,1163]" italics="true" pageId="11" pageNumber="35">hirsuta</emphasis>
Becker, 1915
</taxonomicName>
known only from the North Urals (
<bibRefCitation id="EFA04B1AFFD3FFA27A42189F723FFB38" author="Shamshev I. V." box="[336,550,1172,1200]" pageId="11" pageNumber="35" pagination="1 - 184" refId="ref10818" refString="Shamshev I. V. 2016. An annotated checklist of empidoid flies (Diptera: Empidoidea, except Dolichopodidae) of Russia. Proceedings of the Russian entomological Society, 87: 1 - 184." type="journal article" year="2016">Shamshev, 2016</bibRefCitation>
).
<taxonomicName id="4C314D68FFD3FFA2792C189F7312FB27" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[574,779,1173,1199]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="35" phylum="Arthropoda" rank="species" species="yakovlevi" status="sp. nov.">
<emphasis id="B945EAF9FFD3FFA2792C189F7312FB27" box="[574,779,1173,1199]" italics="true" pageId="11" pageNumber="35">Empis yakovlevi</emphasis>
</taxonomicName>
<emphasis id="B945EAF9FFD3FFA27B8D18B27111FB5A" bold="true" box="[159,264,1208,1234]" pageId="11" pageNumber="35">
<taxonomicNameLabel id="A2765782FFD3FFA27B8D18B27111FB5A" box="[159,264,1208,1234]" pageId="11" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C314D68FFD3FFA27AA718B37228FB5A" authorityName="Becker" authorityYear="1915" box="[437,561,1208,1235]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="35" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B945EAF9FFD3FFA27AA718B371DFFB5A" box="[437,454,1209,1234]" italics="true" pageId="11" pageNumber="35">E</emphasis>
.
<emphasis id="B945EAF9FFD3FFA27ACB18B27228FB5A" box="[473,561,1208,1234]" italics="true" pageId="11" pageNumber="35">hirsuta</emphasis>
</taxonomicName>
primarily in the black palpus (versus yellow) and the longer setae on the femora; also, the male has evenly thickened fore tibia (vs. clavate).
</paragraph>
<paragraph id="8B8E36EBFFD3FFA27BD7194F7217FA41" blockId="11.[159,779,185,1693]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27BD7194F715EFAD7" box="[197,327,1349,1375]" italics="true" pageId="11" pageNumber="35">Etymology</emphasis>
. The new species is named in honour of Russian entomologist Roman V. Yakovlev (Barnaul) in recognition of his contribution to the knowledge of insects of
<collectingRegion id="49F5F809FFD3FFA27AD519A57213FA41" box="[455,522,1455,1481]" country="Russia" name="Altay" pageId="11" pageNumber="35">Altai</collectingRegion>
.
</paragraph>
<paragraph id="8B8E36EBFFD3FFA27BD719D870E8F998" blockId="11.[159,779,185,1693]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27BD719D87147FA64" box="[197,350,1490,1516]" italics="true" pageId="11" pageNumber="35">Distribution</emphasis>
. Palaearctic:
<collectingCountry id="F326767BFFD3FFA2791C19D87278FA64" box="[526,609,1490,1516]" name="Russia" pageId="11" pageNumber="35">Russia</collectingCountry>
(Republic of
<collectingRegion id="49F5F809FFD3FFA27B8D19FF70FDF998" box="[159,228,1525,1552]" country="Russia" name="Altay" pageId="11" pageNumber="35">Altai</collectingRegion>
).
</paragraph>
<paragraph id="8B8E36EBFFD3FFA27BD71A1371DFF915" blockId="11.[159,779,185,1693]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27BD71A137241F9BB" box="[197,600,1561,1587]" italics="true" pageId="11" pageNumber="35">Habitat and seasonal occurrence</emphasis>
. According to the label data, the species occurs in subalpine and alpine zones (
<quantity id="4CC99B0EFFD3FFA27A501A5571ECF9F2" box="[322,501,1631,1659]" metricMagnitude="3" metricUnit="m" metricValue="2.3" metricValueMax="2.5" metricValueMin="2.1" pageId="11" pageNumber="35" unit="m" value="2300.0" valueMax="2500.0" valueMin="2100.0">21002500 m</quantity>
). It was collected from late June until late July.
</paragraph>
<paragraph id="8B8E36EBFFD3FFA27B8D1AC77212F898" blockId="11.[159,671,1740,1808]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27B8D1AC77212F898" bold="true" pageId="11" pageNumber="35">
A key to species of
<taxonomicName id="4C314D68FFD3FFA27AA31AC67262F961" authorityName="Bezzi" authorityYear="1909" box="[433,635,1740,1769]" pageId="11" pageNumber="41" rank="subGenus" subGenus="Polyblepharis">
<emphasis id="B945EAF9FFD3FFA27AA31AC67262F961" bold="true" box="[433,635,1740,1769]" italics="true" pageId="11" pageNumber="35">Polyblepharis</emphasis>
</taxonomicName>
of the
<collectingRegion id="49F5F809FFD3FFA27BC61AF97166F898" box="[212,383,1779,1808]" country="Russia" name="Altay" pageId="11" pageNumber="35">Altai region</collectingRegion>
of
<collectingCountry id="F326767BFFD3FFA27AB81AF97212F898" box="[426,523,1779,1808]" name="Russia" pageId="11" pageNumber="35">Russia</collectingCountry>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C32B6560FFD3FFA27B8D1B2475BAF80D" pageId="11" pageNumber="35" type="key">
<key id="FFA061DAFFD3FFA27B8D1B2475BAF80D" pageId="11" pageNumber="35">
<keyStep id="30C52E4EFFD3FFA27B8D1B2475BAFEBA" pageId="11" pageNumber="35">
<paragraph id="8B8E36EBFFD3FFA27B8D1B247310F80E" blockId="11.[159,778,1837,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA27B8D1B247310F80E" pageId="11" pageNumber="35">1. Scutum (in dorsal view) with three indistinct vittae running beneath the rows of acrostichal and dorsocentral setae or without vittae............ 2</keyLead>
</paragraph>
<paragraph id="8B8E36EBFFD3FFA2782A1CB375BAFEBA" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1CB375BAFEBA" pageId="11" pageNumber="35"> Scutum (in dorsal view) with four distinct brownish, dark or shiny vittae (one between the rows of acrostichals and dorsocentrals and one between the dorsocentrals and supra-alars on either side).... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E4EFFD3FFA2782A1D3075BBFDBC" pageId="11" pageNumber="35">
<paragraph id="8B8E36EBFFD3FFA2782A1D3075BAFE3B" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1D3075BAFE3B" pageId="11" pageNumber="35">
2. Acrostichal and dorsocentral setae multiserial, long. Male: fore basitarsus about half the fore tibia length. Female: palpus largely yellow, darkened at apex........................
<emphasis id="B945EAF9FFD3FFA27FD11D9175BAFE3B" bold="true" box="[1219,1443,411,435]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27FD11D9174CAFE3A" bold="true" box="[1219,1235,411,434]" italics="true" pageId="11" pageNumber="35">E</emphasis>
. (
<emphasis id="B945EAF9FFD3FFA27FFD1D917519FE3A" bold="true" box="[1263,1280,411,434]" italics="true" pageId="11" pageNumber="35">P</emphasis>
.)
<emphasis id="B945EAF9FFD3FFA27E0B1D9175BAFE3B" bold="true" box="[1305,1443,411,435]" italics="true" pageId="11" pageNumber="35">sedelnikovi</emphasis>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8E36EBFFD3FFA2782A1DB675BBFDBC" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1DB675BBFDBC" pageId="11" pageNumber="35">
Acrostichal setae biserial, dorsocentrals arranged in one to two irregular rows, very short. Male: fore basitarsus at least two-thirds the fore tibia length. Female: palpus black.........
<emphasis id="B945EAF9FFD3FFA27FDE1E1775BBFDBC" bold="true" box="[1228,1442,540,565]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27FDE1E1774C5FDBC" bold="true" box="[1228,1244,541,564]" italics="true" pageId="11" pageNumber="35">E</emphasis>
. (
<emphasis id="B945EAF9FFD3FFA27FE51E177511FDBC" bold="true" box="[1271,1288,541,564]" italics="true" pageId="11" pageNumber="35">P</emphasis>
.)
<emphasis id="B945EAF9FFD3FFA27E331E1675BBFDBC" bold="true" box="[1313,1442,540,564]" italics="true" pageId="11" pageNumber="35">longimana</emphasis>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E4EFFD3FFA2782A1E3775BAFDFE" pageId="11" pageNumber="35">
<paragraph id="8B8E36EBFFD3FFA2782A1E3775BAFDDD" blockId="11.[824,1444,184,1926]" box="[824,1443,572,597]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1E3775BAFDDD" box="[824,1443,572,597]" pageId="11" pageNumber="35">3. Palpus black................................. 4</keyLead>
</paragraph>
<paragraph id="8B8E36EBFFD3FFA2782A1E5475BAFDFE" blockId="11.[824,1444,184,1926]" box="[824,1443,605,630]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1E5475BAFDFE" box="[824,1443,605,630]" pageId="11" pageNumber="35"> Palpus yellowish............................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E4EFFD3FFA2782A1E7475BBFC90" pageId="11" pageNumber="35">
<paragraph id="8B8E36EBFFD3FFA2782A1E7475BBFD5F" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1E7475BBFD5F" pageId="11" pageNumber="35">
4. Smaller, body length about
<quantity id="4CC99B0EFFD3FFA27FA11E7474E5FD1D" box="[1203,1276,638,662]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="35" unit="mm" value="4.0">4 mm</quantity>
; scutum with shiny blackish vittae; legs entirely black.................................
<emphasis id="B945EAF9FFD3FFA27F871EB575BBFD5F" bold="true" box="[1173,1442,702,727]" pageId="11" pageNumber="35">
<taxonomicName id="4C314D68FFD3FFA27F871EB57558FD5E" ID-CoL="8BZMN" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[1173,1345,702,726]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="35" phylum="Arthropoda" rank="species" species="altaica" status="sp. nov." subGenus="Polyblepharis">
<emphasis id="B945EAF9FFD3FFA27F871EB574B6FD5E" bold="true" box="[1173,1199,703,726]" italics="true" pageId="11" pageNumber="35">E.</emphasis>
(
<emphasis id="B945EAF9FFD3FFA27FD21EB574CEFD5E" bold="true" box="[1216,1239,703,726]" italics="true" pageId="11" pageNumber="35">P.</emphasis>
)
<emphasis id="B945EAF9FFD3FFA27FF91EB47558FD5E" bold="true" box="[1259,1345,702,726]" italics="true" pageId="11" pageNumber="35">altaica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765782FFD3FFA27E551EB575BBFD5F" box="[1351,1442,703,727]" pageId="11" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8E36EBFFD3FFA2782A1EEA75BBFC90" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1EEA75BBFC90" pageId="11" pageNumber="35">
Larger, body length
<quantity id="4CC99B0EFFD3FFA27F4B1ED574DAFD7F" box="[1113,1219,735,759]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="11" pageNumber="35" unit="mm" value="8.5" valueMax="9.0" valueMin="8.0">89 mm</quantity>
; scutum with dull brownish vittae; legs extensively yellow........ 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E4EFFD3FFA2782A1F1575BBFBD4" pageId="11" pageNumber="35">
<paragraph id="8B8E36EBFFD3FFA2782A1F1575BBFC32" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1F1575BBFC32" pageId="11" pageNumber="35">
5. Face bare; frons broad, only slightly narrower than the distance between outer margins of posterior ocelli; fore and mid tarsomeres slender; legs covered with short setae; tibiae black. Female unknown...................
<emphasis id="B945EAF9FFD3FFA27F511FA875BBFC32" bold="true" box="[1091,1442,929,954]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27F511FA8744AFC31" bold="true" box="[1091,1107,930,953]" italics="true" pageId="11" pageNumber="35">E</emphasis>
.
<taxonomicName id="4C314D68FFD3FFA27F711FAB7558FC31" ID-CoL="8BZPW" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[1123,1345,929,953]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="35" phylum="Arthropoda" rank="species" species="seminskiensis" status="sp. nov." subGenus="Polyblepharis">
(
<emphasis id="B945EAF9FFD3FFA27F7C1FA87466FC31" bold="true" box="[1134,1151,930,953]" italics="true" pageId="11" pageNumber="35">P</emphasis>
.)
<emphasis id="B945EAF9FFD3FFA27F8A1FAB7558FC31" bold="true" box="[1176,1345,929,953]" italics="true" pageId="11" pageNumber="35">seminskiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765782FFD3FFA27E551FA875BBFC32" box="[1351,1442,930,954]" pageId="11" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8E36EBFFD3FFA2782A1FC875BBFBD4" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1FC875BBFBD4" pageId="11" pageNumber="35">
Face with marginal setae; frons narrow, eyes almost touching; fore and mid tarsomeres somewhat thickened; legs almost entirely covered with long dense hair-like setae; tibiae extensively yellowish...........................
<emphasis id="B945EAF9FFD3FFA27F67184E75BBFBD4" bold="true" box="[1141,1442,1091,1116]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27F67184E749CFBD3" bold="true" box="[1141,1157,1092,1115]" italics="true" pageId="11" pageNumber="35">E</emphasis>
.
<taxonomicName id="4C314D68FFD3FFA27F8718497559FBD3" ID-CoL="8BZQL" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[1173,1344,1091,1115]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="35" phylum="Arthropoda" rank="species" species="yakovlevi" status="sp. nov." subGenus="Polyblepharis">
(
<emphasis id="B945EAF9FFD3FFA27FB2184E74A8FBD3" bold="true" box="[1184,1201,1092,1115]" italics="true" pageId="11" pageNumber="35">P</emphasis>
.)
<emphasis id="B945EAF9FFD3FFA27FD818497559FBD3" bold="true" box="[1226,1344,1091,1115]" italics="true" pageId="11" pageNumber="35">yakovlevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765782FFD3FFA27E55184E75BBFBD4" box="[1351,1442,1092,1116]" pageId="11" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E4EFFD3FFA2782A186975BBFAD7" pageId="11" pageNumber="35">
<paragraph id="8B8E36EBFFD3FFA2782A186975BBFB55" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A186975BBFB55" pageId="11" pageNumber="35">6. Acrostichal setae biserial, dorsocentrals usually biserial anteriorly and uniserial toward scutellum; scutellum with 46 setae; laterotergite mostly with black setae................................... 7</keyLead>
</paragraph>
<paragraph id="8B8E36EBFFD3FFA2782A18EC75BBFAD7" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A18EC75BBFAD7" pageId="11" pageNumber="35"> Acrostichal setae arranged at least in three to four irregular rows, dorsocentrals multiserial anteriorly; scutellum at least with 10 setae; laterotergite with pale to yellowish setae........................ 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E4EFFD3FFA2782A196275BBF9A9" pageId="11" pageNumber="35">
<paragraph id="8B8E36EBFFD3FFA2782A196275BBFA37" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A196275BBFA37" pageId="11" pageNumber="35">
7. Postpronotal lobe shiny on outer side. Male: eyes holoptic. Female: mid and hind femora with short posteroventral pennate setae..
<emphasis id="B945EAF9FFD3FFA27FC119A275BBFA37" bold="true" box="[1235,1442,1447,1472]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27FC119A274FAFA37" bold="true" box="[1235,1251,1448,1471]" italics="true" pageId="11" pageNumber="35">E</emphasis>
. (
<emphasis id="B945EAF9FFD3FFA27FEC19A27516FA37" bold="true" box="[1278,1295,1448,1471]" italics="true" pageId="11" pageNumber="35">P</emphasis>
.)
<emphasis id="B945EAF9FFD3FFA27E3A19AD75BBFA37" bold="true" box="[1320,1442,1447,1471]" italics="true" pageId="11" pageNumber="35">subciliata</emphasis>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8E36EBFFD3FFA2782A19C375BBF9A9" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A19C375BBF9A9" pageId="11" pageNumber="35">
Postpronotal lobe entirely pruinose. Male: eyes dichoptic, separated by broad frons. Female unknown..................
<emphasis id="B945EAF9FFD3FFA27F801A0375BBF9A9" bold="true" box="[1170,1442,1545,1569]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27F801A0374BBF9A8" bold="true" box="[1170,1186,1545,1568]" italics="true" pageId="11" pageNumber="35">E</emphasis>
.
<taxonomicName id="4C314D68FFD3FFA27FA31A037558F9A9" ID-CoL="8BZQC" authority="Shamshev, 2019" authorityName="Shamshev" authorityYear="2019" box="[1201,1345,1545,1569]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="35" phylum="Arthropoda" rank="species" species="turalae" status="sp. nov." subGenus="Polyblepharis">
(
<emphasis id="B945EAF9FFD3FFA27FAF1A0374D7F9A8" bold="true" box="[1213,1230,1545,1568]" italics="true" pageId="11" pageNumber="35">P</emphasis>
.)
<emphasis id="B945EAF9FFD3FFA27FF51A037558F9A9" bold="true" box="[1255,1345,1545,1569]" italics="true" pageId="11" pageNumber="35">turalae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2765782FFD3FFA27E551A0375BBF9A9" box="[1351,1442,1545,1569]" pageId="11" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C52E4EFFD3FFA2782A1A2075BAF80D" pageId="11" pageNumber="35">
<paragraph id="8B8E36EBFFD3FFA2782A1A2075BBF94A" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1A2075BBF94A" pageId="11" pageNumber="35">
8. Hind femur thickened, with numerous spine-like setae throughout ventrally; hind tibia slightly thickened toward apex; hind femur about 1.5 times as wide as hind tibia in middle part; legs usually uniformly brownish.............
<emphasis id="B945EAF9FFD3FFA27FEF1AA175BBF94A" bold="true" box="[1277,1442,1707,1731]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27FEF1AA17514F94A" bold="true" box="[1277,1293,1707,1730]" italics="true" pageId="11" pageNumber="35">E</emphasis>
. (
<emphasis id="B945EAF9FFD3FFA27E3A1AA17520F94A" bold="true" box="[1320,1337,1707,1730]" italics="true" pageId="11" pageNumber="35">P</emphasis>
.)
<emphasis id="B945EAF9FFD3FFA27E401AA175BBF94A" bold="true" box="[1362,1442,1707,1730]" italics="true" pageId="11" pageNumber="35">crassa</emphasis>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8B8E36EBFFD3FFA2782A1AC675BAF80D" blockId="11.[824,1444,184,1926]" pageId="11" pageNumber="35">
<keyLead id="30C095DEFFD3FFA2782A1AC675BAF80D" pageId="11" pageNumber="35">
Hind femur slender, covered only with simple setulae ventrally, at most with 34 short anteroventral setae in subapical part; hind tibia almost evenly thickened, in middle part nearly as wide as hind femur; mid and hind femora usually yellowish (except extreme apex).................
<emphasis id="B945EAF9FFD3FFA27FF11B6475BAF80D" bold="true" box="[1251,1443,1901,1926]" pageId="11" pageNumber="35">
<emphasis id="B945EAF9FFD3FFA27FF11B6474EAF80D" bold="true" box="[1251,1267,1902,1925]" italics="true" pageId="11" pageNumber="35">E</emphasis>
. (
<emphasis id="B945EAF9FFD3FFA27E1D1B647539F80D" bold="true" box="[1295,1312,1902,1925]" italics="true" pageId="11" pageNumber="35">P</emphasis>
.)
<emphasis id="B945EAF9FFD3FFA27E2B1B6775BAF80D" bold="true" box="[1337,1443,1901,1925]" italics="true" pageId="11" pageNumber="35">gravipes</emphasis>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>