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<document id="DDA7B12D33DE48D0CB74007FECC13BB4" ID-DOI="10.11646/zootaxa.5089.1.1" ID-GBIF-Dataset="67fbf2b0-b944-485d-90b5-976914d338ad" ID-ISSN="1175-5326" ID-Zenodo-Dep="5836060" ID-ZooBank="E12F2B20-A84A-48E2-9C77-B281F1BFC62E" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1641892626941" checkinUser="plazi" docAuthor="Ortega-Salas, Héctor, González-Soriano, Enrique &amp; Jocque, Merlijn" docDate="2022" docId="039887EB3034FFFFFF73FAE6F61CFB27" docLanguage="en" docName="zootaxa.5089.1.1.pdf" docOrigin="Zootaxa 5089 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Paraphlebia ixchel Ortega-Salas &amp; Gonzalez-Soriano 2022, sp. nov." docType="treatment" docVersion="8" lastPageNumber="20" masterDocId="FFA1FF933026FFECFFE4FFC9F553FF86" masterDocTitle="Untangling the waterfall damsels: a review of the Mesoamerican genus Paraphlebia Selys in Hagen, 1861 (Odonata: Thaumatoneuridae) with descriptions of 11 new species" masterLastPageNumber="66" masterPageNumber="1" pageNumber="19" updateTime="1732192187527" updateUser="juliana">
<mods:mods id="27FEF1E64A0333EDCE7E68A179ECCB48" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F156B6CD320CD4D2C1C55A9B2EEB1211">
<mods:title id="5FAF48485702DB41DB30CE847A1FC459">Untangling the waterfall damsels: a review of the Mesoamerican genus Paraphlebia Selys in Hagen, 1861 (Odonata: Thaumatoneuridae) with descriptions of 11 new species</mods:title>
</mods:titleInfo>
<mods:name id="AA1CB05D6B80FD1654E2CBCE94750EC0" type="personal">
<mods:role id="54D6461F9E2B57C5BA14AA7A1CBD3929">
<mods:roleTerm id="851A0ACF4B00E51A07FE1A7A9714862F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B97E6C7E92868529C8B1B7B2878ED4A6">Ortega-Salas, Héctor</mods:namePart>
<mods:nameIdentifier id="57B0F1359E23C06A75EC943EE172FDDC" type="ORCID">0000-0002-5373-4839</mods:nameIdentifier>
<mods:affiliation id="ECCEE4F43F09EE7F6237EDE78EB4C898">Posgrado en Ciencias Biológicas, UNAM, México. &amp; Departamento de Zoología, Instituto de Biología, UNAM, Apartado Postal 70 - 153, México. &amp; Current address: Naturalis Biodiversity Center, Leiden, The Netherlands.</mods:affiliation>
<mods:nameIdentifier id="B1D164060395DF1D1EE136F1E58B8E0F" type="email">hector.ortegasalas@naturalis.nl</mods:nameIdentifier>
</mods:name>
<mods:name id="E1B60287D50B7C08D5B4B5B1FC4A5B49" type="personal">
<mods:role id="B357E110ADFD1C2E55654189EDD96EB0">
<mods:roleTerm id="552A35E3FFE0694DA01B0F454B716EC3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1D573120EC87CAA1A39F11F42A7A9C2A">González-Soriano, Enrique</mods:namePart>
<mods:nameIdentifier id="3CD5A9BB8E924098630130AF77BA8E9E" type="ORCID">0000-0002-4798-7274</mods:nameIdentifier>
<mods:affiliation id="A880305F2A0F7E8341D074F37B25C167">Departamento de Zoología, Instituto de Biología, UNAM, Apartado Postal 70 - 153, México.</mods:affiliation>
<mods:nameIdentifier id="975CFC5F44C4D8B85124607DC1FB5164" type="email">esoriano@ib.unam.mx</mods:nameIdentifier>
</mods:name>
<mods:name id="E978A66453F84EB67722CCD4F715D3DB" type="personal">
<mods:role id="071CCB88E3B1921D09B811DC8B801A89">
<mods:roleTerm id="4C4182F5D9A06C9FFDB3B43473CC0EA9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E0730753BC5950441B73872FA8A889F6">Jocque, Merlijn</mods:namePart>
<mods:nameIdentifier id="EFA11482B5782E92C2E101A98BBE39D5" type="ORCID">0000-0002-7196-7476</mods:nameIdentifier>
<mods:affiliation id="948541795BCCEC7CA8608013CBDD262B">Royal Belgian Institute of Natural Sciences, Brussels, Belgium. &amp; Operation Wallacea, Hope House, Old Bolingbroke, Lincolnshire, United Kingdom. &amp; Biodiversity inventory for Conservation (BINCO) NPO, Walmersumstraat 44, 3380 Glabbeek, Belgium.</mods:affiliation>
<mods:nameIdentifier id="698D0D4F30C4EA2B9CE65163E5E16A93" type="email">merlijn.jocque@binco.eu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A18E6DB022981F00E378B7DBE9F90DF2">text</mods:typeOfResource>
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<mods:titleInfo id="71B1C846D8601B5A60F93982B90DBEA4">
<mods:title id="0E649EDC2DDC959D72EA8FDDF6BA2CB6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="D14FD7F48C915D297288CE0BAA833C09">
<mods:date id="D3246FB876F63B43278661051E651011">2022</mods:date>
<mods:detail id="E5DEBFDA55406230080F0A5901B126C4" type="pubDate">
<mods:number id="9A08994D2019F98F923786314F8ABD0A">2022-01-11</mods:number>
</mods:detail>
<mods:detail id="CE4C56B9BD294EBFCE6AC17CE7C98E5C" type="volume">
<mods:number id="A0C90F41DF3644E497707664C6376AC5">5089</mods:number>
</mods:detail>
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<mods:number id="CC2954DAD6FC931A5C8D3D0A830B65F0">1</mods:number>
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<mods:start id="C777751BD10B10956E40B93F75000C9B">1</mods:start>
<mods:end id="07E84C42C369A07FA207FBF245069F25">66</mods:end>
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<mods:classification id="551E718DBADA0D32A44A8B18EC1CAB24">journal article</mods:classification>
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<treatment id="039887EB3034FFFFFF73FAE6F61CFB27" ID-DOI="http://doi.org/10.5281/zenodo.5836100" ID-GBIF-Taxon="209152404" ID-Zenodo-Dep="5836100" LSID="urn:lsid:plazi:treatment:039887EB3034FFFFFF73FAE6F61CFB27" httpUri="http://treatment.plazi.org/id/039887EB3034FFFFFF73FAE6F61CFB27" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="C32B65763034FFFEFF73FAE6F6C1FACC" box="[151,914,1327,1354]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8B8E36FD3034FFFEFF73FAE6F6C1FACC" blockId="18.[151,1096,1327,1392]" box="[151,914,1327,1354]" pageId="18" pageNumber="19">
<heading id="D0C681913034FFFEFF73FAE6F6C1FACC" bold="true" box="[151,914,1327,1354]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="4">
<emphasis id="B945EAEF3034FFFEFF73FAE6F6C1FACC" bold="true" box="[151,914,1327,1354]" pageId="18" pageNumber="19">
<taxonomicName id="4C314D7E3034FFFEFF73FAE6F67EFACC" authority="Ortega-Salas &amp; Gonzalez-Soriano" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[151,813,1327,1354]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ixchel" status="sp. nov.">
<emphasis id="B945EAEF3034FFFEFF73FAE6F42FFACC" bold="true" box="[151,380,1327,1354]" italics="true" pageId="18" pageNumber="19">Paraphlebia ixchel</emphasis>
Ortega-Salas &amp; González-Soriano
</taxonomicName>
<taxonomicNameLabel id="A27657943034FFFEFCD0FAF9F6C1FACC" box="[820,914,1328,1354]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B65763034FFFEFF73FA9DF45EF9C7" pageId="18" pageNumber="19" type="description">
<paragraph id="8B8E36FD3034FFFEFF73FA9DF11BFAE8" blockId="18.[151,1096,1327,1392]" box="[151,1096,1364,1392]" pageId="18" pageNumber="19">
<figureCitation id="130A2A783034FFFEFF73FA9DF5A2FAE9" box="[151,241,1364,1391]" captionStart="FIGURES 20–27" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[207,1373,193,1901]" captionTargetId="figure-17@25.[197,1390,181,1919]" captionTargetPageId="25" captionText="FIGURES 20–27. Paraphlebia spp. colour pattern of thorax." figureDoi="http://doi.org/10.5281/zenodo.5836070" httpUri="https://zenodo.org/record/5836070/files/figure.png" pageId="18" pageNumber="19">Figs. 25</figureCitation>
(thorax
<specimenCount id="9D37FD743034FFFEFEB4FA9CF436FAF6" box="[336,357,1365,1392]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
), 121 (posterior lobe of prothorax
<specimenCount id="9D37FD743034FFFEFD0FFA9CF653FAF6" box="[747,768,1365,1392]" pageId="18" pageNumber="19" type="female">♀</specimenCount>
), 130 (S8–
<specimenCount id="9D37FD743034FFFEFC99FA9DF6E6FAF6" box="[893,949,1364,1392]" pageId="18" pageNumber="19" type="female">10 ♀</specimenCount>
), 150 (map).
</paragraph>
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<uri id="FFA03AFF3034FFFEFF73FA51F626FA36" box="[151,885,1432,1456]" pageId="18" pageNumber="19">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8B8E36FD3034FFFEFF73FA16F45EF9C7" blockId="18.[151,1437,1503,2033]" pageId="18" pageNumber="19">
<materialsCitation id="3B593CA03034FFFEFF73FA16F45EF9C7" ID-GBIF-Occurrence="4023503590" collectingDate="1962-06-29" collectionCode="TWD, IBUNAM" collectorName="T. Donnelly" country="Guatemala" county="Santa Catarina Ixtahuacan" elevation="200" latitude="14.8" location="Rio Nahulate" longLatPrecision="76" longitude="-91.366" municipality="Rio Nahulate" pageId="18" pageNumber="19" specimenCode="CNIN: OD22715" specimenCount="1" specimenCount-female="1" stateProvince="Solola" typeStatus="holotype">
<typeStatus id="548A885F3034FFFEFF73FA16F450FA7F" box="[151,259,1503,1529]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F326766D3034FFFEFEF4FA16F4EEFA7F" box="[272,445,1503,1529]" name="Guatemala" pageId="18" pageNumber="19">GUATEMALA</collectingCountry>
•
<specimenCount id="9D37FD743034FFFEFE3EFA16F755FA7D" box="[474,518,1503,1531]" pageId="18" pageNumber="19" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49F5F81F3034FFFEFDFCFA16F731FA7F" box="[536,610,1503,1529]" country="Guatemala" name="Solola" pageId="18" pageNumber="19">Solola</collectingRegion>
,
<collectingCounty id="62EF4E713034FFFEFD94FA16F6CDFA7F" box="[624,926,1503,1529]" pageId="18" pageNumber="19">Santa Catarina Ixtahuacán</collectingCounty>
,
<location id="8EEE60263034FFFEFC48FA16F115FA7F" LSID="urn:lsid:plazi:treatment:039887EB3034FFFFFF73FAE6F61CFB27:8EEE60263034FFFEFC48FA16F115FA7F" box="[940,1094,1503,1529]" country="Guatemala" county="Santa Catarina Ixtahuacan" latitude="14.8" longLatPrecision="76" longitude="-91.366" municipality="Rio Nahulate" name="Rio Nahulate" pageId="18" pageNumber="19" stateProvince="Solola">Río Nahulate</location>
,
<locationDeviation id="9A3C2FDF3034FFFEFBB7FA16F190FA7F" box="[1107,1219,1503,1529]" pageId="18" pageNumber="19">3.6 km S.</locationDeviation>
of
<location id="8EEE60263034FFFEFB0BFA16F0CBFA7F" LSID="urn:lsid:plazi:treatment:039887EB3034FFFFFF73FAE6F61CFB27:8EEE60263034FFFEFB0BFA16F0CBFA7F" box="[1263,1432,1503,1529]" country="Guatemala" county="Santa Catarina Ixtahuacan" latitude="14.8" longLatPrecision="76" longitude="-91.366" municipality="Rio Nahulate" name="Santa Catarina" pageId="18" pageNumber="19" stateProvince="Solola">Santa Catarina</location>
, forest (200 m above river); [
<geoCoordinate id="EE05503A3034FFFEFE29F9CAF76CF99B" box="[461,575,1539,1565]" degrees="14.800" direction="north" orientation="latitude" pageId="18" pageNumber="19" precision="55" value="14.8">14.800°N</geoCoordinate>
,
<geoCoordinate id="EE05503A3034FFFEFDADF9CBF790F99B" box="[585,707,1538,1565]" degrees="91.366" direction="west" orientation="longitude" pageId="18" pageNumber="19" precision="55" value="-91.366">91.366°W</geoCoordinate>
];
<quantity id="4CC99B183034FFFEFD37F9CAF679F99B" box="[723,810,1539,1565]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="18" pageNumber="19" unit="m" value="1900.0">
<elevation id="001CD1CE3034FFFEFD37F9CAF679F99B" box="[723,810,1539,1565]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="18" pageNumber="19" unit="m" value="1900.0">1900 m</elevation>
</quantity>
;
<date id="FF8F103D3034FFFEFCD1F9CAF6EDF99B" box="[821,958,1539,1565]" pageId="18" pageNumber="19" value="1962-06-29">
<collectingDate id="EFCBE9D53034FFFEFCD1F9CAF6EDF99B" box="[821,958,1539,1565]" pageId="18" pageNumber="19" value="1962-06-29">29 Jun 1962</collectingDate>
</date>
;
<collectorName id="26C4532B3034FFFEFC2DF9CAF11EF99B" box="[969,1101,1539,1565]" pageId="18" pageNumber="19">T. Donnelly</collectorName>
leg.;
<collectionCode id="ED20AE383034FFFEFB6DF9CAF19FF99B" box="[1161,1228,1539,1565]" pageId="18" pageNumber="19">TWD</collectionCode>
;
<collectionCode id="ED20AE383034FFFEFB33F9CAF01DF99B" box="[1239,1358,1539,1565]" country="Mexico" httpUri="http://grbio.org/cool/9wfd-7x2v" name="Instituto de BiIología, Universidad Nacional Autónoma de México" pageId="18" pageNumber="19" type="University or college">IBUNAM</collectionCode>
:
<specimenCode id="DB979E863034FFFEFAB5F9CAF459F9C7" pageId="18" pageNumber="19">CNIN: OD22715</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B65763034FFFEFF23F982F00FF92B" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8B8E36FD3034FFFEFF23F982F00FF92B" blockId="18.[151,1437,1503,2033]" pageId="18" pageNumber="19">
<emphasis id="B945EAEF3034FFFEFF23F982F41BF9E3" bold="true" box="[199,328,1611,1637]" pageId="18" pageNumber="19">Etymology</emphasis>
. Named
<taxonomicName id="4C314D7E3034FFFEFE4DF982F4BFF9E3" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[425,492,1611,1637]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ixchel">
<emphasis id="B945EAEF3034FFFEFE4DF982F4BFF9E3" box="[425,492,1611,1637]" italics="true" pageId="18" pageNumber="19">ixchel</emphasis>
</taxonomicName>
/ɪksˈʃel/ (L. noun in apposition), after Ix Chel (also Chac Chel, Chak Chel or Sak Chel) the Mayan old goddess of creation, divination, and curing. Ix Chel is one of the few known feminine divinities in the Mayan pantheon, is therefore suitable to give its name to the only species described from a female specimen.
</paragraph>
</subSubSection>
<subSubSection id="C32B65763034FFFFFF23F97FF73CFE57" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="description">
<paragraph id="8B8E36FD3034FFFEFF23F97FF767F957" blockId="18.[151,1437,1503,2033]" box="[199,564,1718,1745]" pageId="18" pageNumber="19">
<emphasis id="B945EAEF3034FFFEFF23F97FF767F957" bold="true" box="[199,564,1718,1745]" pageId="18" pageNumber="19">
Description of female
<typeStatus id="548A885F3034FFFEFE29F97EF767F957" box="[461,564,1719,1745]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8E36FD3034FFFEFF23F912F06AF8BB" blockId="18.[151,1437,1503,2033]" pageId="18" pageNumber="19">
<emphasis id="B945EAEF3034FFFEFF23F912F457F973" box="[199,260,1755,1781]" italics="true" pageId="18" pageNumber="19">Head</emphasis>
. Chiefly black, labrum dark glossy blue, anteclypeus pale, postclypeus pale, frons pale cream with black lateral spots, vertex with two pale spots anterolateral to lateral ocelli, occiput with occipital bar restricted to spots on the lateral edges, eyes dark brown to black; antennal scape with distal margin pale, flagellum light brown.
</paragraph>
<paragraph id="8B8E36FD3034FFFFFF23F88EF070FF53" blockId="18.[151,1437,1503,2033]" lastBlockId="19.[151,1437,151,1185]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B945EAEF3034FFFEFF23F88EF44BF8E7" box="[199,280,1863,1889]" italics="true" pageId="18" pageNumber="19">Thorax</emphasis>
. Prothorax: chiefly black; anterior lobe mostly pale, with black line on anterior margin; middle lobe with pale markings along central pit and outer margins; posterior lobe brown to black towards middle, see
<figureCitation id="130A2A783034FFFEFA94F8A2F592F82F" captionStart="FIGURES 117–128" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[208,1385,195,1907]" captionTargetId="figure-17@37.[190,1398,181,1919]" captionTargetPageId="37" captionText="FIGURES 117–128. Paraphlebia spp. posterior lobe of prothorax dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5836120" httpUri="https://zenodo.org/record/5836120/files/figure.png" pageId="18" pageNumber="19">Fig. 121</figureCitation>
for morphology. Pterothorax: chiefly black with diffuse green metallic reflections over dark colouration on mesepisternum and mesepimeron; mediodorsal carina with mediodorsal pale line covering one-fourth width of mesepisternum; see
<figureCitation id="130A2A783034FFFEFE9AF81EF49CF877" box="[382,463,2006,2033]" captionStart="FIGURES 20–27" captionStartId="25.[151,264,1943,1967]" captionTargetBox="[207,1373,193,1901]" captionTargetId="figure-17@25.[197,1390,181,1919]" captionTargetPageId="25" captionText="FIGURES 20–27. Paraphlebia spp. colour pattern of thorax." figureDoi="http://doi.org/10.5281/zenodo.5836070" httpUri="https://zenodo.org/record/5836070/files/figure.png" pageId="18" pageNumber="19">Fig. 25</figureCitation>
for colour pattern; metepimeron pale, with a hint of a black curved line, may be lost in preservation;; sternum pale; depression mesad to mesostigmal lobe in mesostigmal plate angulated; legs mostly pale; femora with black extensor surfaces and distal joints; tibiae with black distal joints; claws reddish.
</paragraph>
<paragraph id="8B8E36FD3035FFFFFF23FF16F447FEC7" blockId="19.[151,1437,151,1185]" pageId="19" pageNumber="20">
<emphasis id="B945EAEF3035FFFFFF23FF16F458FF7E" box="[199,267,223,248]" italics="true" pageId="19" pageNumber="20">Wings</emphasis>
. Hyaline, with a slightly amber tint; FW with one post-quadrangular cell, the vein that descends from the subnodus at the first Vx; HW field between CuA and posterior margin lacking supplementary veins. Px: FW 40/38, HW 27/26.
</paragraph>
<paragraph id="8B8E36FD3035FFFFFF23FE82F609FE0F" blockId="19.[151,1437,151,1185]" pageId="19" pageNumber="20">
<emphasis id="B945EAEF3035FFFFFF23FE82F462FEE3" box="[199,305,331,357]" italics="true" pageId="19" pageNumber="20">Abdomen</emphasis>
. Black with pale areas as follow: S1 with anteroventral spots and posterior margin; S9 with dorsal spot widened posteriorly. Ovipositor short, not reaching tip of cerci.
</paragraph>
<paragraph id="8B8E36FD3035FFFFFF23FE5AF64CFE2B" blockId="19.[151,1437,151,1185]" box="[199,799,403,429]" pageId="19" pageNumber="20">Measurements. Abdomen: 39.0, FW: 37.0, HW: 34.6.</paragraph>
<paragraph id="8B8E36FD3035FFFFFF23FE7FF73CFE57" blockId="19.[151,1437,151,1185]" box="[199,623,438,465]" pageId="19" pageNumber="20">
<emphasis id="B945EAEF3035FFFFFF23FE7FF4E4FE57" bold="true" box="[199,439,438,465]" pageId="19" pageNumber="20">Description of male.</emphasis>
Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32B65763035FFFFFF23FE13F6F7FCBF" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8B8E36FD3035FFFFFF23FE13F68DFC93" blockId="19.[151,1437,151,1185]" pageId="19" pageNumber="20">
<emphasis id="B945EAEF3035FFFFFF23FE13F411FE72" bold="true" box="[199,322,474,500]" pageId="19" pageNumber="20">Diagnosis.</emphasis>
<taxonomicName id="4C314D7E3035FFFFFEA9FE12F774FE73" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[333,551,475,501]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ixchel">
<emphasis id="B945EAEF3035FFFFFEA9FE12F774FE73" box="[333,551,475,501]" italics="true" pageId="19" pageNumber="20">Paraphlebia ixchel</emphasis>
</taxonomicName>
,
<taxonomicName id="4C314D7E3035FFFFFDDEFE12F79FFE72" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[570,716,475,500]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="esperanza">
<emphasis id="B945EAEF3035FFFFFDDEFE12F79FFE72" box="[570,716,475,500]" italics="true" pageId="19" pageNumber="20">P. esperanza</emphasis>
</taxonomicName>
and
<taxonomicName id="4C314D7E3035FFFFFCEFFE12F6C3FE73" authorityName="Jocque &amp; Ortega-Salas" authorityYear="2022" box="[779,912,475,501]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kukulkan">
<emphasis id="B945EAEF3035FFFFFCEFFE12F6C3FE73" box="[779,912,475,501]" italics="true" pageId="19" pageNumber="20">P. kukulkan</emphasis>
</taxonomicName>
belong to the group of species with a short ovipositor that never surpasses the tip of the cercus (
<figureCitation id="130A2A783035FFFFFD3AFE36F6D7FD9F" box="[734,900,511,537]" captionStart="FIGURES 129–137" captionStartId="38.[151,264,1943,1967]" captionTargetBox="[218,1370,193,1907]" captionTargetId="figure-18@38.[202,1385,181,1919]" captionTargetPageId="38" captionText="FIGURES 129–137. Paraphlebia spp. S8–10 lateral view." figureDoi="http://doi.org/10.5281/zenodo.5836126" httpUri="https://zenodo.org/record/5836126/files/figure.png" pageId="19" pageNumber="20">Figs. 129–130</figureCitation>
). The posterior lobe of the prothorax in
<taxonomicName id="4C314D7E3035FFFFFADAFE36F0CFFD9F" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[1342,1436,511,537]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ixchel">
<emphasis id="B945EAEF3035FFFFFADAFE36F0CFFD9F" box="[1342,1436,511,537]" italics="true" pageId="19" pageNumber="20">P. ixchel</emphasis>
</taxonomicName>
is similar to that of
<taxonomicName id="4C314D7E3035FFFFFE95FDEAF4ADFDBA" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[369,510,547,572]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="esperanza">
<emphasis id="B945EAEF3035FFFFFE95FDEAF4ADFDBA" box="[369,510,547,572]" italics="true" pageId="19" pageNumber="20">P. esperanza</emphasis>
</taxonomicName>
. It differs from that of
<taxonomicName id="4C314D7E3035FFFFFD1BFDEAF6D3FDBB" authorityName="Jocque &amp; Ortega-Salas" authorityYear="2022" box="[767,896,547,573]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kukulkan">
<emphasis id="B945EAEF3035FFFFFD1BFDEAF6D3FDBB" box="[767,896,547,573]" italics="true" pageId="19" pageNumber="20">P. kukulkan</emphasis>
</taxonomicName>
in presenting digitiform projections towards the lateral edges of the posterior margin while in the latter the posterior lobe is entire, without projections. In dorsal view, these projections are directed posterodorsally, making a V-shaped gap between them and the posterior margin of the medial lobe in
<taxonomicName id="4C314D7E3035FFFFFE60FD46F4B1FD2F" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[388,482,655,681]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ixchel">
<emphasis id="B945EAEF3035FFFFFE60FD46F4B1FD2F" box="[388,482,655,681]" italics="true" pageId="19" pageNumber="20">P. ixchel</emphasis>
</taxonomicName>
, while in
<taxonomicName id="4C314D7E3035FFFFFDB5FD46F78CFD2E" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[593,735,655,680]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="esperanza">
<emphasis id="B945EAEF3035FFFFFDB5FD46F78CFD2E" box="[593,735,655,680]" italics="true" pageId="19" pageNumber="20">P. esperanza</emphasis>
</taxonomicName>
they are directed posteroventrally, not forming such gap (
<figureCitation id="130A2A783035FFFFFA80FD46F5AEFD4B" captionStart="FIGURES 117–128" captionStartId="37.[151,264,1943,1967]" captionTargetBox="[208,1385,195,1907]" captionTargetId="figure-17@37.[190,1398,181,1919]" captionTargetPageId="37" captionText="FIGURES 117–128. Paraphlebia spp. posterior lobe of prothorax dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5836120" httpUri="https://zenodo.org/record/5836120/files/figure.png" pageId="19" pageNumber="20">Figs. 121–122</figureCitation>
). The female of
<taxonomicName id="4C314D7E3035FFFFFE55FD7AF76DFD4A" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[433,574,691,716]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="esperanza">
<emphasis id="B945EAEF3035FFFFFE55FD7AF76DFD4A" box="[433,574,691,716]" italics="true" pageId="19" pageNumber="20">P. esperanza</emphasis>
</taxonomicName>
can be readily separated from all other species by the position of the vein that descend from the subnodus, in the FW this vein is always closer to the second post-quadrangular Vx than to the first. In all other species there is always closer to the first post-quadrangular Vx.
</paragraph>
<paragraph id="8B8E36FD3035FFFFFF23FCD7F6F7FCBF" blockId="19.[151,1437,151,1185]" box="[199,932,798,825]" pageId="19" pageNumber="20">
<emphasis id="B945EAEF3035FFFFFF23FCD7F470FCBE" bold="true" box="[199,291,798,824]" pageId="19" pageNumber="20">Habitat</emphasis>
. This species was collected in forest,
<quantity id="4CC99B183035FFFFFD2EFCD6F642FCBF" box="[714,785,799,825]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="20" unit="m" value="200.0">200 m</quantity>
from a river.
</paragraph>
</subSubSection>
<subSubSection id="C32B65763035FFFFFF23FC8BF7FFFC07" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8B8E36FD3035FFFFFF23FC8BF7FFFC07" blockId="19.[151,1437,151,1185]" pageId="19" pageNumber="20">
<emphasis id="B945EAEF3035FFFFFF23FC8BF40AFCDA" bold="true" box="[199,345,834,860]" pageId="19" pageNumber="20">Distribution</emphasis>
. Southwest
<collectingCountry id="F326766D3035FFFFFE01FC8AF730FCDB" box="[485,611,835,861]" name="Guatemala" pageId="19" pageNumber="20">Guatemala</collectingCountry>
: it is only known from the
<typeStatus id="548A885F3035FFFFFC71FC8AF696FCDB" box="[917,965,835,861]" pageId="19" pageNumber="20">type</typeStatus>
locality near Santa Catarina in the
<collectingRegion id="49F5F81F3035FFFFFAB1FC8AF44AFC07" country="Guatemala" name="Solola" pageId="19" pageNumber="20">Solola department</collectingRegion>
. It was collected at ca.
<quantity id="4CC99B183035FFFFFDF8FCAEF72BFC07" box="[540,632,871,897]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="19" pageNumber="20" unit="m" value="1900.0">1900 m</quantity>
.a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="C32B65763035FFFFFF23FC42F61CFB27" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8B8E36FD3035FFFFFF23FC42F61CFB27" blockId="19.[151,1437,151,1185]" pageId="19" pageNumber="20">
<emphasis id="B945EAEF3035FFFFFF23FC42F467FC23" bold="true" box="[199,308,907,933]" pageId="19" pageNumber="20">Remarks</emphasis>
. Although not all the females of the species hereby described are known, it is clear that the morphology of the posterior lobe of prothorax presents a strong parallelism between males and females at the species level. Therefore
<taxonomicName id="4C314D7E3035FFFFFEE8FC1AF43AFC6B" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[268,361,979,1005]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="ixchel">
<emphasis id="B945EAEF3035FFFFFEE8FC1AF43AFC6B" box="[268,361,979,1005]" italics="true" pageId="19" pageNumber="20">P. ixchel</emphasis>
</taxonomicName>
is considered as a distinct species as, with the exception of
<taxonomicName id="4C314D7E3035FFFFFC1EFC1AF1D4FC6A" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" box="[1018,1159,979,1004]" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="esperanza">
<emphasis id="B945EAEF3035FFFFFC1EFC1AF1D4FC6A" box="[1018,1159,979,1004]" italics="true" pageId="19" pageNumber="20">P. esperanza</emphasis>
</taxonomicName>
, no other male described presents a similar structure. Based on the female morphology, the male probably will be similar to that of
<taxonomicName id="4C314D7E3035FFFFFA6CFC3EF45AFBB2" authorityName="Ortega-Salas &amp; Gonzalez-Soriano" authorityYear="2022" class="Insecta" family="Megapodagrionidae" genus="Paraphlebia" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="esperanza">
<emphasis id="B945EAEF3035FFFFFA6CFC3EF45AFBB2" italics="true" pageId="19" pageNumber="20">P. esperanza</emphasis>
</taxonomicName>
: it should have a convex posterior margin on the posterior lobe of prothorax with digitiform projections, and it is possible that such projections are projected posteriorly. The cerci should have the mediodorsal flange welldeveloped and the vein that descends from the nodus in the FW should be closer to the first Vx than to the second, the latter character allowing differentiation of the two species.
</paragraph>
</subSubSection>
</treatment>
</document>
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