treatments-xml/data/03/98/87/039887D2FF8EAF712A8A3E1CFB96F9B7.xml

<document id="0E9ECD5AE8CB46AD7D4CA0D9DAD4545F" ID-DOI="10.11646/zootaxa.4819.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="4396744" ID-ZooBank="80258C53-F94D-46CB-9E86-AF0C3226DF9E" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1609199984737" checkinUser="plazi" docAuthor="Polizei, Thiago T. S., Barclay, Maxwell V. L. &amp; Bispo, Pitágoras C." docDate="2020" docId="039887D2FF8EAF712A8A3E1CFB96F9B7" docLanguage="en" docName="zootaxa.4819.1.2.pdf" docOrigin="Zootaxa 4819 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hexacylloepus phalluspilosus Polizei &amp; Barclay &amp; Bispo 2020, sp. nov." docType="treatment" docUuid="49C4A1D5-2824-4FA2-81E5-AF4A3AB39EDC" docUuidSource="ZooBank" docVersion="1" lastPageNumber="33" masterDocId="FFA1FFAAFF97AF6A2A1D3A09FFC5FF9A" masterDocTitle="Hexacylloepus Hinton, 1940 rises again; 18 new species of a neglected genus (Coleoptera: Byrrhoidea: Elmidae)" masterLastPageNumber="48" masterPageNumber="6" pageNumber="31" updateTime="1730286293773" updateUser="diego" zenodo-license-document="CLOSED">
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<mods:title id="6E06E75C535C48AFD4657C9F2DA323E1">Hexacylloepus Hinton, 1940 rises again; 18 new species of a neglected genus (Coleoptera: Byrrhoidea: Elmidae)</mods:title>
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<mods:namePart id="D0ABDBD563841398472EB455B5EEDCF9">Polizei, Thiago T. S.</mods:namePart>
<mods:nameIdentifier id="35DA5D7E2BBD0A74760AA95FE896EF87" type="ORCID">0000-0003-3746-8865</mods:nameIdentifier>
<mods:affiliation id="C61E4330A469FE1E99AB0A9A1357ADB6">Laboratório de Coleoptera (LaC) - Museu de Zoologia da Universidade de São Paulo (MZSP). São Paulo, São Paulo State, Brazil. thiagopolizei @ hotmail. com; https: // orcid. org / 0000 - 0003 - 3746 - 8865 &amp; Programa de Pós-Graduação em Entomologia-Departamento de Biologia - Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto (FFCL-RP) - Universidade de São Paulo (USP). Ribeirão Preto, São Paulo State, Brazil.</mods:affiliation>
<mods:nameIdentifier id="C797F8AFAC6AE3BF073717313F14FBAD" type="email">thiagopolizei@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="409A3616BF45FEC305DA79824DC8682F">Barclay, Maxwell V. L.</mods:namePart>
<mods:affiliation id="34C656FB36A0F1055291CC398FD53AF4">Department of Life Sciences, Natural History Museum. London SW 7 5 BD, United Kingdom.</mods:affiliation>
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<mods:namePart id="B0FE67281900D6507AE2C72715F91266">Bispo, Pitágoras C.</mods:namePart>
<mods:affiliation id="DD540D91481BA259A30E8EB672B4DC93">Programa de Pós-Graduação em Entomologia-Departamento de Biologia - Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto (FFCL-RP) - Universidade de São Paulo (USP). Ribeirão Preto, São Paulo State, Brazil. &amp; Laboratório de Biologia Aquática (LABIA). Departamento de Ciências Biológicas - Faculdade de Ciências e Letras de Assis (FCL) - Universidade Estadual Paulista “ Júlio de Mesquita Filho ” (Unesp). Assis, São Paulo State, Brazil.</mods:affiliation>
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<subSubSection id="C32B654FFF8EAF732A8A3E1CFD9FFBB5" box="[151,602,1045,1071]" pageId="25" pageNumber="31" type="nomenclature">
<paragraph id="8B8E36C4FF8EAF732A8A3E1CFD9FFBB5" blockId="25.[151,602,1045,1107]" box="[151,602,1045,1071]" pageId="25" pageNumber="31">
<taxonomicName id="4C314D47FF8EAF732A8A3E1CFE30FBB5" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[151,501,1045,1071]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="31" phylum="Arthropoda" rank="species" species="phalluspilosus" status="sp. nov.">
<emphasis id="B945EAD6FF8EAF732A8A3E1CFE30FBB5" box="[151,501,1045,1071]" italics="true" pageId="25" pageNumber="31">Hexacylloepus phalluspilosus</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FF8EAF732BE13E1CFD9FFBB5" bold="true" box="[508,602,1045,1071]" pageId="25" pageNumber="31">
<taxonomicNameLabel id="A27657ADFF8EAF732BE13E1CFD9FFBB5" box="[508,602,1045,1071]" pageId="25" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C32B654FFF8EAF732A8A3E30FEF1FBC9" box="[151,308,1081,1107]" pageId="25" pageNumber="31" type="description">
<paragraph id="8B8E36C4FF8EAF732A8A3E30FEF1FBC9" blockId="25.[151,602,1045,1107]" box="[151,308,1081,1107]" pageId="25" pageNumber="31">
(
<figureCitation id="130A2A41FF8EAF732ABD3E30FF35FBC9" box="[160,240,1081,1107]" captionStart="FIGURE 12" captionStartId="26.[151,250,1978,2002]" captionTargetBox="[187,1369,484,1919]" captionTargetId="figure-125@26.[179,1408,453,1954]" captionTargetPageId="26" captionText="FIGURE 12. Hexacylloepus phalluspilosus sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="25" pageNumber="31">Fig. 12</figureCitation>
A–C)
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFF8EAF732A8A3E75FD6DFB0C" box="[151,680,1148,1174]" pageId="25" pageNumber="31" type="materials_examined">
<paragraph id="8B8E36C4FF8EAF732A8A3E75FD6DFB0C" blockId="25.[151,1437,1148,2039]" box="[151,680,1148,1174]" pageId="25" pageNumber="31">
<emphasis id="B945EAD6FF8EAF732A8A3E75FEFCFB0C" bold="true" box="[151,313,1148,1174]" pageId="25" pageNumber="31">
<typeStatus id="548A8866FF8EAF732A8A3E75FF17FB0C" box="[151,210,1148,1174]" pageId="25" pageNumber="31">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F3267654FF8EAF732B5D3E75FE42FB0C" box="[320,391,1148,1174]" name="Brazil" pageId="25" pageNumber="31">Brazil</collectingCountry>
, 
<collectingRegion id="49F5F826FF8EAF732B8E3E75FDD3FB0C" box="[403,534,1148,1174]" country="Brazil" name="Goias" pageId="25" pageNumber="31">Goiás State</collectingRegion>
, 
<location id="8EEE601FFF8EAF73283C3E75FD61FB0C" LSID="urn:lsid:plazi:treatment:039887D2FF8EAF712A8A3E1CFB96F9B7:8EEE601FFF8EAF73283C3E75FD61FB0C" box="[545,676,1148,1174]" country="Brazil" name="Pirenopolis" pageId="25" pageNumber="31" stateProvince="Goias">Pirenópolis</location>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFF8EAF732ADA3EA9FA58FA08" pageId="25" pageNumber="31" type="diagnosis">
<paragraph id="8B8E36C4FF8EAF732ADA3EA9FA58FA08" blockId="25.[151,1437,1148,2039]" pageId="25" pageNumber="31">
<emphasis id="B945EAD6FF8EAF732ADA3EA9FE81FB20" bold="true" box="[199,324,1184,1210]" pageId="25" pageNumber="31">Diagnosis:</emphasis>
Antennae with few thin setae on the apex of each antennomere. Frontoclypeal suture inconspicuous. Pronotum with a short, narrow, shallow longitudinal impression on disc, reaching the anterior and posterior margins. Hypomera with a complete belt of tomentum covering almost the entire anterior half portion. Hind wings macropterous. Epipleura with tomentum. Tibiae each with two rows of spines and setae. Abdomen covered with tomentum; first ventrite with a deep depression with a pair of carinae extending towards and reaching the metaventrite. Male genitalia spatulate; phallobasis twice as long as wide; parameres reaching 4/5 of the penis length, twice as wide at the base as at the apex; penis with the lateral margin slightly sinuous on anterior half and straight on posterior half.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFF8EAF712ADA3F95FD11FD57" lastPageId="27" lastPageNumber="33" pageId="25" pageNumber="31" type="description">
<paragraph id="8B8E36C4FF8EAF732ADA3F95FCD4FA41" blockId="25.[151,1437,1148,2039]" pageId="25" pageNumber="31">
<emphasis id="B945EAD6FF8EAF732ADA3F95FE9EFA2C" bold="true" box="[199,347,1436,1462]" pageId="25" pageNumber="31">Description:</emphasis>
Male. Body elongated, subparallel. Colour black, with antenna, mouthparts and legs reddishbrown. Total length: 
<quantity id="4CC99B21FF8EAF732B9C3FC9FE2DFA41" box="[385,488,1472,1499]" metricMagnitude="-3" metricUnit="m" metricValue="2.24" pageId="25" pageNumber="31" unit="mm" value="2.24">2.24 mm</quantity>
. Greatest width: 
<quantity id="4CC99B21FF8EAF7328BB3FC9FCC8FA41" box="[678,781,1472,1499]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="25" pageNumber="31" unit="mm" value="1.02">1.02 mm</quantity>
.
</paragraph>
<paragraph id="8B8E36C4FF8EAF732ADA3FEDFC16F929" blockId="25.[151,1437,1148,2039]" pageId="25" pageNumber="31">
<emphasis id="B945EAD6FF8EAF732ADA3FEDFEC8FA64" box="[199,269,1508,1534]" italics="true" pageId="25" pageNumber="31">Head:</emphasis>
surface with micropunctures separated from each other by their diameter. Antenna inserted at the anterior margin of the eyes, filiform, with 11 antennomeres each one with long thin setae at apex. Frontoclypeal suture inconspicuous. Clypeus 4x as wide as long; anterior margin straight and posterior margin sinuous; surface similar to that of head. Labrum twice as wide as long; antero-lateral angles rounded; and surface with long and thin setae. Maxillary palpus 4-segmented; terminal palpomere truncated apically. Labial palpus 3-segmented. Mentum and submentum rectangular. Gula rectangular. Genae covered with tomentum.
</paragraph>
<paragraph id="8B8E36C4FF8EAF732ADA3CB5FE20F8D8" blockId="25.[151,1437,1148,2039]" pageId="25" pageNumber="31">
<emphasis id="B945EAD6FF8EAF732ADA3CB5FEE4F94C" box="[199,289,1724,1750]" italics="true" pageId="25" pageNumber="31">Thorax:</emphasis>
Pronotum slightly as wide as long, arcuate anteriorly and laterally; posterior margin smooth with two prescutellar foveae (
<figureCitation id="130A2A41FF8EAF732B613CE9FE20F960" box="[380,485,1760,1786]" captionStart="FIGURE 12" captionStartId="26.[151,250,1978,2002]" captionTargetBox="[187,1369,484,1919]" captionTargetId="figure-125@26.[179,1408,453,1954]" captionTargetPageId="26" captionText="FIGURE 12. Hexacylloepus phalluspilosus sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="25" pageNumber="31">Fig. 12A</figureCitation>
). Surface with micropunctures and with posterior half alutaceous. Sublateral carinae complete, sinuous and raised. Longitudinal impression short, narrow and shallow on disc (
<figureCitation id="130A2A41FF8EAF732E973D0DFB36F884" box="[1162,1267,1796,1822]" captionStart="FIGURE 12" captionStartId="26.[151,250,1978,2002]" captionTargetBox="[187,1369,484,1919]" captionTargetId="figure-125@26.[179,1408,453,1954]" captionTargetPageId="26" captionText="FIGURE 12. Hexacylloepus phalluspilosus sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="25" pageNumber="31">Fig. 12A</figureCitation>
). A pair of gibbosities behind the scutellum.
</paragraph>
<paragraph id="8B8E36C4FF8EAF732ADA3D45FC63F86C" blockId="25.[151,1437,1148,2039]" pageId="25" pageNumber="31">
Elytra twice as long as wide; anterior margin arcuate; lateral margin moderately arcuate and serrate; rounded apex (
<figureCitation id="130A2A41FF8EAF732AC33D79FE8DF810" box="[222,328,1904,1930]" captionStart="FIGURE 12" captionStartId="26.[151,250,1978,2002]" captionTargetBox="[187,1369,484,1919]" captionTargetId="figure-125@26.[179,1408,453,1954]" captionTargetPageId="26" captionText="FIGURE 12. Hexacylloepus phalluspilosus sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="25" pageNumber="31">Fig. 12A</figureCitation>
). Humeri rounded, slightly prominent. Surface with puncture rows separated from each other by their diameter. Elytral suture moderately elevated. Carina short on interval IV, extending from base to 1/4 of the elytral length; sublateral carinae on intervals VI and VIII, extending from base to 2/3 of the elytral length. Epipleura with tomentum (
<figureCitation id="130A2A41FF8EAF732B4C3DD5FE72F86C" box="[337,439,2012,2038]" captionStart="FIGURE 12" captionStartId="26.[151,250,1978,2002]" captionTargetBox="[187,1369,484,1919]" captionTargetId="figure-125@26.[179,1408,453,1954]" captionTargetPageId="26" captionText="FIGURE 12. Hexacylloepus phalluspilosus sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="25" pageNumber="31">Fig. 12B</figureCitation>
). Hind wings micropterous. Scutellum oval.
</paragraph>
<paragraph id="8B8E36C4FF8DAF702ADA3A9EFC4CFEDB" blockId="26.[151,1436,151,429]" pageId="26" pageNumber="32">
Prosternum broad, with a pair of carinae reaching the anterior margin; surface with micropunctures; anterior margin straight and lateral margin with tomentum. Disc depressed. Prosternal process long and narrow, beyond the procoxae; lateral margin strongly prominent; rounded apex; disc strongly excavated. Notosternal suture slightly sinuous. Hypomera wider posteriorly than anteriorly, surface with micropunctures and with a complete belt of tomentum covering almost the entire anterior half portion (
<figureCitation id="130A2A41FF8DAF7029083B2EFCB9FEDB" box="[789,892,295,321]" captionStart="FIGURE 12" captionStartId="26.[151,250,1978,2002]" captionTargetBox="[187,1369,484,1919]" captionTargetId="figure-125@26.[179,1408,453,1954]" captionTargetPageId="26" captionText="FIGURE 12. Hexacylloepus phalluspilosus sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="26" pageNumber="32">Fig. 12B</figureCitation>
).
</paragraph>
<paragraph id="8B8E36C4FF8DAF702ADA3B42FD50FE37" blockId="26.[151,1436,151,429]" pageId="26" pageNumber="32">Mesoventrite with a slight groove to receive the prosternal process. Mesepimeron and mesepisternum with tomentum. Metaventrite broad with a strongly longitudinal medial depression; surface alutaceous with micropunctures; and with sides covered with tomentum.</paragraph>
<caption id="DF4E664CFF8DAF702A8A3DB3FC41F86C" ID-DOI="http://doi.org/10.5281/zenodo.4396770" ID-Zenodo-Dep="4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="26" pageNumber="32" startId="26.[151,250,1978,2002]" targetBox="[187,1369,484,1919]" targetPageId="26">
<paragraph id="8B8E36C4FF8DAF702A8A3DB3FC41F86C" blockId="26.[151,1436,1978,2038]" pageId="26" pageNumber="32">
<emphasis id="B945EAD6FF8DAF702A8A3DB3FEE4F848" bold="true" box="[151,289,1978,2002]" pageId="26" pageNumber="32">FIGURE 12.</emphasis>
<taxonomicName id="4C314D47FF8DAF702B353DB3FD92F848" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[296,599,1978,2002]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="32" phylum="Arthropoda" rank="species" species="phalluspilosus" status="sp. nov.">
<emphasis id="B945EAD6FF8DAF702B353DB3FD92F848" box="[296,599,1978,2002]" italics="true" pageId="26" pageNumber="32">Hexacylloepus phalluspilosus</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FF8DAF7028423DB3FD75F848" bold="true" box="[607,688,1978,2002]" pageId="26" pageNumber="32">
<taxonomicNameLabel id="A27657ADFF8DAF7028423DB3FD75F848" box="[607,688,1978,2002]" pageId="26" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm.
</paragraph>
</caption>
<paragraph id="8B8E36C4FF8CAF712ADA3A9EFAB8FE87" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="33">
Legs (except tarsi) covered with punctures and tomentum (
<figureCitation id="130A2A41FF8CAF7129753A9EFC14FF2B" box="[872,977,151,177]" captionStart="FIGURE 12" captionStartId="26.[151,250,1978,2002]" captionTargetBox="[187,1369,484,1919]" captionTargetId="figure-125@26.[179,1408,453,1954]" captionTargetPageId="26" captionText="FIGURE 12. Hexacylloepus phalluspilosus sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="27" pageNumber="33">Fig. 12B</figureCitation>
). Coxae rounded; trochanters elongated; femora 3x as long as wide, 1/3 of the length of the elytra; tibiae long and thin, 6x as long as wide, 1/3 of the length of the elytra. Profemora each with a dense row of long setae in distal 2/3. Tibiae each with two complete, dense rows of spines. Tarsi elongate, with setae; apical tarsomere longer than the basal four combined. Claws long and thin.
</paragraph>
<paragraph id="8B8E36C4FF8CAF712ADA3B2EFCB7FE4B" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="33">
<emphasis id="B945EAD6FF8CAF712ADA3B2EFEF4FEDB" box="[199,305,295,321]" italics="true" pageId="27" pageNumber="33">Abdomen</emphasis>
(
<figureCitation id="130A2A41FF8CAF712B5F3B2EFE6DFEDB" box="[322,424,295,321]" captionStart="FIGURE 12" captionStartId="26.[151,250,1978,2002]" captionTargetBox="[187,1369,484,1919]" captionTargetId="figure-125@26.[179,1408,453,1954]" captionTargetPageId="26" captionText="FIGURE 12. Hexacylloepus phalluspilosus sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="27" pageNumber="33">Fig. 12B</figureCitation>
): five ventrites; as long as wide. Anterior and posterior margins straight on the first and second ventrites and arcuate on the subsequent ventrites; lateral margins rounded. Surface densely covered with tomentum (excepted for disc of first ventrite). Disc of the first ventrite with a deep depression, and a pair of carinae extending towards and reaching the metaventrite. Second ventrite with the anterior half without tomentum. Fifth ventrite with a groove on disc, apex rounded and with long setae.
</paragraph>
<paragraph id="8B8E36C4FF8CAF712ADA3BD2FA89FD33" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="33">
<emphasis id="B945EAD6FF8CAF712ADA3BD2FEAEFE6F" box="[199,363,475,501]" italics="true" pageId="27" pageNumber="33">Male genitalia</emphasis>
(
<figureCitation id="130A2A41FF8CAF712B673BD2FE24FE6F" box="[378,481,475,501]" captionStart="FIGURE 12" captionStartId="26.[151,250,1978,2002]" captionTargetBox="[187,1369,484,1919]" captionTargetId="figure-125@26.[179,1408,453,1954]" captionTargetPageId="26" captionText="FIGURE 12. Hexacylloepus phalluspilosus sp. nov.. Male. Holotype. (A–B) Habitus. (A) Dorsal view. (B) Ventral view. (C) Ventral view of male genitalia. Scale bars: (A–B) = 0.5mm. (C) = 0.1mm." figureDoi="http://doi.org/10.5281/zenodo.4396770" httpUri="https://zenodo.org/record/4396770/files/figure.png" pageId="27" pageNumber="33">Fig. 12C</figureCitation>
): (Total length: 
<quantity id="4CC99B21FF8CAF71288F3BD2FD3CFE6F" box="[658,761,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="27" pageNumber="33" unit="mm" value="0.67">0.67 mm</quantity>
. Greatest width: 
<quantity id="4CC99B21FF8CAF7129A83BD2FBD8FE6F" box="[949,1053,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="27" pageNumber="33" unit="mm" value="0.17">0.17 mm</quantity>
), symmetrical and spatulate. Phallobase aproximately 3x as long as wide, slightly longer than penis length. Parameres fused and articulated with the phallobase, wide, reaching 4/5 of the penis length; twice as wide at the base as at the apex; apex slightly acute, curved to internal margin, strongly covered by setae in dorsal and ventral view. Penis spatulate with the lateral margin slightly sinuous on anterior half and straight on posterior half; apex rounded; fibula elongated, wide and approximately the same length as the penis; corona present, as an arrangement of tiny rod-like sclerotizations.
</paragraph>
<paragraph id="8B8E36C4FF8CAF712ADA38BAFD11FD57" blockId="27.[151,1437,151,1581]" box="[199,724,691,717]" pageId="27" pageNumber="33">
<emphasis id="B945EAD6FF8CAF712ADA38BAFEEDFD57" bold="true" box="[199,296,691,717]" pageId="27" pageNumber="33">Female:</emphasis>
External morphology similar to male.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFF8CAF712ADA38DFFD9CFD6B" box="[199,601,726,753]" pageId="27" pageNumber="33" type="materials_examined">
<paragraph id="8B8E36C4FF8CAF712ADA38DFFD9CFD6B" blockId="27.[151,1437,151,1581]" box="[199,601,726,753]" pageId="27" pageNumber="33">
<emphasis id="B945EAD6FF8CAF712ADA38DFFE76FD6A" bold="true" box="[199,435,726,752]" pageId="27" pageNumber="33">Material examined:</emphasis>
<specimenCount id="9D37FD4DFF8CAF712BA738DEFD90FD6B" box="[442,597,727,753]" pageId="27" pageNumber="33" type="generic">88 specimens</specimenCount>
.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFF8CAF712ADA38F3FC73FC8F" box="[199,950,762,789]" pageId="27" pageNumber="33" type="distribution">
<paragraph id="8B8E36C4FF8CAF712ADA38F3FC73FC8F" blockId="27.[151,1437,151,1581]" box="[199,950,762,789]" pageId="27" pageNumber="33">
<emphasis id="B945EAD6FF8CAF712ADA38F3FEA6FC8E" bold="true" box="[199,355,762,788]" pageId="27" pageNumber="33">Distribution:</emphasis>
<collectingCountry id="F3267654FF8CAF712B7738F2FE76FC8F" box="[362,435,763,789]" name="Brazil" pageId="27" pageNumber="33">Brazil</collectingCountry>
: 
<collectingRegion id="49F5F826FF8CAF712BA038F2FDC4FC8F" box="[445,513,763,789]" country="Brazil" name="Goias" pageId="27" pageNumber="33">Goiás</collectingRegion>
, Distrito Federal and 
<collectingRegion id="49F5F826FF8CAF7128EE38F2FCA1FC8F" box="[755,868,763,789]" country="Brazil" name="Sao Paulo" pageId="27" pageNumber="33">São Paulo</collectingRegion>
States.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFF8CAF712ADA3916FD4FFBE7" pageId="27" pageNumber="33" type="materials_examined">
<paragraph id="8B8E36C4FF8CAF712ADA3916FF36FCC7" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="33">
<materialsCitation id="3B593C99FF8CAF712ADA3916FF36FCC7" collectingDate="2010-07-09" collectingDateMax="2010-07-10" collectingDateMin="2010-07-09" collectionCode="MZSP" collectorName="G. Paciencia" country="Brazil" municipality="Pirenópolis" pageId="27" pageNumber="33" specimenCode="MZSP 35806" specimenCount="1" specimenCount-male="1" stateProvince="Goias" typeStatus="holotype">
<emphasis id="B945EAD6FF8CAF712ADA3916FEBBFCA3" bold="true" box="[199,382,799,825]" pageId="27" pageNumber="33">
<specimenCount id="9D37FD4DFF8CAF712ADA3916FEC1FCA3" box="[199,260,799,825]" pageId="27" pageNumber="33" type="male">Male</specimenCount>
<typeStatus id="548A8866FF8CAF712B133916FEBCFCA3" box="[270,377,799,825]" pageId="27" pageNumber="33" type="holotype">holotype</typeStatus>
:
</emphasis>
♂ | Holo- / type | BR-GO-Pirenópolis / 
<date id="FF8F1004FF8CAF7129463916FBCCFCA3" box="[859,1033,799,825]" pageId="27" pageNumber="33" value="2010-07-09" valueMax="2010-07-10" valueMin="2010-07-09">
<collectingDate id="EFCBE9ECFF8CAF7129463916FBCCFCA3" box="[859,1033,799,825]" pageId="27" pageNumber="33" value="2010-07-09" valueMax="2010-07-10" valueMin="2010-07-09">09-10.VII.2010</collectingDate>
</date>
/ 
<collectorName id="26C45312FF8CAF712E3B3916FB73FCA3" box="[1062,1206,799,825]" pageId="27" pageNumber="33">G. Paciencia</collectorName>
<emphasis id="B945EAD6FF8CAF712EDC3916FB2FFCA3" box="[1217,1258,799,825]" italics="true" pageId="27" pageNumber="33">leg.</emphasis>
| 
<specimenCode id="DB979EBFFF8CAF712F193916FA59FCA3" box="[1284,1436,798,825]" pageId="27" pageNumber="33">MZSP 35806</specimenCode>
(
<collectionCode id="ED20AE01FF8CAF712A82394AFF2EFCC7" box="[159,235,835,861]" pageId="27" pageNumber="33">MZSP</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph id="8B8E36C4FF8CAF712ADA396EFD4FFBE7" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="33">
<materialsCitation id="3B593C99FF8CAF712ADA396EFEDEFC3F" collectingDate="2010-07-09" collectingDateMax="2010-07-10" collectingDateMin="2010-07-09" collectionCode="MZSP" collectorName="G. Paciencia" country="Brazil" latitude="-15.889" longLatPrecision="1" longitude="-47.842667" municipality="Pirenópolis" pageId="27" pageNumber="33" specimenCode="MZSP 35806" specimenCount="87" specimenCount-male="1" stateProvince="Goias" typeStatus="paratype">
<emphasis id="B945EAD6FF8CAF712ADA396EFEFAFC1B" bold="true" box="[199,319,871,897]" pageId="27" pageNumber="33">
<typeStatus id="548A8866FF8CAF712ADA396EFEFAFC1B" box="[199,319,871,897]" pageId="27" pageNumber="33" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="9D37FD4DFF8CAF712B52396EFE29FC1B" box="[335,492,871,897]" pageId="27" pageNumber="33" type="generic">87 specimens</specimenCount>
: BR-GO-Pirenópolis / 
<date id="FF8F1004FF8CAF7128ED396EFC5BFC1B" box="[752,926,871,897]" pageId="27" pageNumber="33" value="2010-07-09" valueMax="2010-07-10" valueMin="2010-07-09">
<collectingDate id="EFCBE9ECFF8CAF7128ED396EFC5BFC1B" box="[752,926,871,897]" pageId="27" pageNumber="33" value="2010-07-09" valueMax="2010-07-10" valueMin="2010-07-09">09-10.VII.2010</collectingDate>
</date>
/ 
<collectorName id="26C45312FF8CAF7129AA396EFB80FC1B" box="[951,1093,871,897]" pageId="27" pageNumber="33">G. Paciencia</collectorName>
<emphasis id="B945EAD6FF8CAF712E53396EFBB2FC1B" box="[1102,1143,871,897]" italics="true" pageId="27" pageNumber="33">leg.</emphasis>
(38 MZSP; 12 NHMUK; 12 USNM).
</materialsCitation>
<materialsCitation id="3B593C99FF8CAF712B3C3982FD22FC53" collectingDate="2006-10-09" collectorName="P. C. Bispo" country="Brazil" location="Paraitinga-P. E. S. M. / nucleo Santa Virginia / Rib. Estrada Catucaba" municipality="São Luís do Paraitinga" pageId="27" pageNumber="33" specimenCount="1" stateProvince="Sao Paulo">
BR-SP-São Luís do / 
<location id="8EEE601FFF8CAF71280C3982FAD4FC3F" LSID="urn:lsid:plazi:treatment:039887D2FF8EAF712A8A3E1CFB96F9B7:8EEE601FFF8CAF71280C3982FAD4FC3F" box="[529,1297,907,933]" country="Brazil" municipality="São Luís do Paraitinga" name="Paraitinga-P. E. S. M. / nucleo Santa Virginia / Rib. Estrada Catucaba" pageId="27" pageNumber="33" stateProvince="Sao Paulo">Paraitinga—P.E.S.M. / núcleo Santa Virgínia / Rib. Estrada Catuçaba</location>
/ 
<date id="FF8F1004FF8CAF712F3B3982FA59FC3F" box="[1318,1436,907,933]" pageId="27" pageNumber="33" value="2006-10-09">
<collectingDate id="EFCBE9ECFF8CAF712F3B3982FA59FC3F" box="[1318,1436,907,933]" pageId="27" pageNumber="33" value="2006-10-09">09.X.2006</collectingDate>
</date>
/ 
<collectorName id="26C45312FF8CAF712AB739A6FEDCFC53" box="[170,281,943,969]" pageId="27" pageNumber="33">P.C.Bispo</collectorName>
<emphasis id="B945EAD6FF8CAF712B3939A6FE88FC53" box="[292,333,943,969]" italics="true" pageId="27" pageNumber="33">leg.</emphasis>
(4 MZSP; 3 NHMUK; 3 USNM).
</materialsCitation>
<materialsCitation id="3B593C99FF8CAF7128EF39A6FE3EFC77" collectingDate="2015-10-22" collectionCode="MZSP" collectorName="G. Paulon &amp; R. Moretto" country="Brazil" municipality="Cassia dos Coqueiros" pageId="27" pageNumber="33" specimenCount="1" stateProvince="Sao Paulo">
BR-SP-Cassia dos / Coqueiros / 
<date id="FF8F1004FF8CAF712E6E39A7FB2CFC53" box="[1139,1257,942,969]" pageId="27" pageNumber="33" value="2015-10-22">
<collectingDate id="EFCBE9ECFF8CAF712E6E39A7FB2CFC53" box="[1139,1257,942,969]" pageId="27" pageNumber="33" value="2015-10-22">22.X.2015</collectingDate>
</date>
/ 
<collectorName id="26C45312FF8CAF712F1A39A6FAB9FC53" box="[1287,1404,943,969]" pageId="27" pageNumber="33">G. Paulon</collectorName>
&amp; 
<collectorName id="26C45312FF8CAF712A8A39DAFED1FC77" box="[151,276,979,1005]" pageId="27" pageNumber="33">R. Moretto</collectorName>
/ 
<emphasis id="B945EAD6FF8CAF712B3239DAFE9CFC77" box="[303,345,979,1005]" italics="true" pageId="27" pageNumber="33">leg.</emphasis>
| (2 
<collectionCode id="ED20AE01FF8CAF712B8F39DAFE25FC77" box="[402,480,979,1005]" pageId="27" pageNumber="33">MZSP</collectionCode>
). |
</materialsCitation>
<materialsCitation id="3B593C99FF8CAF71281939DAFE1CFB8B" collectingDate="2012-08-19" collectionCode="MZSP" collectorName="T. T. S. Polizei &amp; L. S. M. Costa" country="Brazil" location="P. E. Morro do Diabo" municipality="BR-SP-Teodoro Sampaio" pageId="27" pageNumber="33" specimenCount="1" stateProvince="Sao Paulo">
<collectingMunicipality id="6BEAACBEFF8CAF71281939DAFCE1FC77" box="[516,804,979,1005]" pageId="27" pageNumber="33">BR-SP-Teodoro Sampaio</collectingMunicipality>
/ 
<location id="8EEE601FFF8CAF71292239DAFBE0FC77" LSID="urn:lsid:plazi:treatment:039887D2FF8EAF712A8A3E1CFB96F9B7:8EEE601FFF8CAF71292239DAFBE0FC77" box="[831,1061,979,1005]" country="Brazil" municipality="BR-SP-Teodoro Sampaio" name="P. E. Morro do Diabo" pageId="27" pageNumber="33" stateProvince="Sao Paulo">P.E.Morro do Diabo</location>
/ 
<date id="FF8F1004FF8CAF712E5D39DAFB17FC77" box="[1088,1234,979,1005]" pageId="27" pageNumber="33" value="2012-08-19">
<collectingDate id="EFCBE9ECFF8CAF712E5D39DAFB17FC77" box="[1088,1234,979,1005]" pageId="27" pageNumber="33" value="2012-08-19">19.VIII.2012</collectingDate>
</date>
/ 
<collectorName id="26C45312FF8CAF712EF139DAFAB8FC77" box="[1260,1405,979,1005]" pageId="27" pageNumber="33">T.T.S.Polizei</collectorName>
&amp; 
<collectorName id="26C45312FF8CAF712A8A39FEFEE3FB8B" box="[151,294,1015,1041]" pageId="27" pageNumber="33">L.S.M.Costa</collectorName>
<emphasis id="B945EAD6FF8CAF712B3339FEFE92FB8B" box="[302,343,1015,1041]" italics="true" pageId="27" pageNumber="33">leg.</emphasis>
(2 
<collectionCode id="ED20AE01FF8CAF712B6239FEFE08FB8B" box="[383,461,1015,1041]" pageId="27" pageNumber="33">MZSP</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B593C99FF8CAF712BFD39FEFBAEFBAF" collectingDate="2019-01-16" collectionCode="MZSP" collectorName="Renan Casimiro" country="Brazil" latitude="-15.889" location="APA GAMA-Jardim / Botanico. Corrego Cabeca de / Veado" longLatPrecision="1" longitude="-47.842667" municipality="Brasilia" pageId="27" pageNumber="33" specimenCount="1" stateProvince="Distrito Federal">
<collectingCountry id="F3267654FF8CAF712BFD39FEFD8DFB8B" box="[480,584,1015,1041]" name="Brazil" pageId="27" pageNumber="33">BRASIL</collectingCountry>
: 
<collectingRegion id="49F5F826FF8CAF71284939FEFCCCFB8B" box="[596,777,1015,1041]" pageId="27" pageNumber="33">Distrito Federal</collectingRegion>
: / 
<collectingMunicipality id="6BEAACBEFF8CAF71293839FEFC45FB8B" box="[805,896,1015,1041]" pageId="27" pageNumber="33">Brasília</collectingMunicipality>
. 
<location id="8EEE601FFF8CAF71299739FEFE8DFBAF" LSID="urn:lsid:plazi:treatment:039887D2FF8EAF712A8A3E1CFB96F9B7:8EEE601FFF8CAF71299739FEFE8DFBAF" country="Brazil" latitude="-15.889" longLatPrecision="1" longitude="-47.842667" municipality="Brasilia" name="APA GAMA-Jardim / Botanico. Corrego Cabeca de / Veado" pageId="27" pageNumber="33" stateProvince="Distrito Federal">APA GAMA—Jardim / Botânico. Córrego Cabeça de / Veado</location>
. 
<geoCoordinate id="EE055003FF8CAF712B4C3E13FE1EFBAF" box="[337,475,1050,1077]" degrees="15" direction="south" minutes="53" orientation="latitude" pageId="27" pageNumber="33" precision="1" seconds="20,4" value="-15.889">15º53’20,4S</geoCoordinate>
<geoCoordinate id="EE055003FF8CAF712BFF3E13FDB2FBAF" box="[482,631,1050,1077]" degrees="47" direction="west" minutes="50" orientation="longitude" pageId="27" pageNumber="33" precision="1" seconds="33,6" value="-47.842667">47º50’33,6W</geoCoordinate>
/ 
<date id="FF8F1004FF8CAF7128963E12FD33FBAF" box="[651,758,1051,1077]" pageId="27" pageNumber="33" value="2019-01-16">
<collectingDate id="EFCBE9ECFF8CAF7128963E12FD33FBAF" box="[651,758,1051,1077]" pageId="27" pageNumber="33" value="2019-01-16">16.I.2019</collectingDate>
</date>
<collectorName id="26C45312FF8CAF7128E13E12FC74FBAF" box="[764,945,1051,1077]" pageId="27" pageNumber="33">Renan Casimiro</collectorName>
<emphasis id="B945EAD6FF8CAF7129AB3E12FC25FBAF" box="[950,992,1051,1077]" italics="true" pageId="27" pageNumber="33">leg.</emphasis>
(10 
<collectionCode id="ED20AE01FF8CAF712E0C3E12FB9AFBAF" box="[1041,1119,1051,1077]" pageId="27" pageNumber="33">MZSP</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B593C99FF8CAF712E6C3E12FD4FFBE7" collectingDate="2008-05-18" collectionCode="MZSP" collectorName="Pitagoras Bispo" country="Brazil" location="Parque Estadual Campos / do Jordao. Riacho Sensitive" municipality="Campos do / Jordao" pageId="27" pageNumber="33" specimenCount="1" stateProvince="Sao Paulo">
<collectingCountry id="F3267654FF8CAF712E6C3E12FB1DFBAF" box="[1137,1240,1051,1077]" name="Brazil" pageId="27" pageNumber="33">BRASIL</collectingCountry>
. 
<collectingRegion id="49F5F826FF8CAF712EFC3E12FA91FBAF" box="[1249,1364,1051,1077]" country="Brazil" name="Sao Paulo" pageId="27" pageNumber="33">São Paulo</collectingRegion>
. 
<collectingMunicipality id="6BEAACBEFF8CAF712F433E12FE82FBC3" pageId="27" pageNumber="33">Campos do / Jordão</collectingMunicipality>
. 
<location id="8EEE601FFF8CAF712B4E3E36FC07FBC3" LSID="urn:lsid:plazi:treatment:039887D2FF8EAF712A8A3E1CFB96F9B7:8EEE601FFF8CAF712B4E3E36FC07FBC3" box="[339,962,1087,1113]" country="Brazil" municipality="Campos do / Jordao" name="Parque Estadual Campos / do Jordao. Riacho Sensitive" pageId="27" pageNumber="33" stateProvince="Sao Paulo">Parque Estadual Campos / do Jordão. Riacho Sensitive</location>
. 
<date id="FF8F1004FF8CAF7129D33E36FB81FBC3" box="[974,1092,1087,1113]" pageId="27" pageNumber="33" value="2008-05-18">
<collectingDate id="EFCBE9ECFF8CAF7129D33E36FB81FBC3" box="[974,1092,1087,1113]" pageId="27" pageNumber="33" value="2008-05-18">18.V.2008</collectingDate>
</date>
. 
<collectorName id="26C45312FF8CAF712E523E36FAC5FBC3" box="[1103,1280,1087,1113]" pageId="27" pageNumber="33">Pitágoras Bispo</collectorName>
leg. (
<collectionCode id="ED20AE01FF8CAF712F5C3E36FA55FBC3" box="[1345,1424,1087,1113]" pageId="27" pageNumber="33">MZSP</collectionCode>
). (MZSP 31170-31179, 31181, 35807-35852).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFF8CAF712ADA3E8EFE1AFB5F" pageId="27" pageNumber="33" type="etymology">
<paragraph id="8B8E36C4FF8CAF712ADA3E8EFE1AFB5F" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="33">
<emphasis id="B945EAD6FF8CAF712ADA3E8EFE97FB3B" bold="true" box="[199,338,1159,1185]" pageId="27" pageNumber="33">Etymology:</emphasis>
from Latin 
<emphasis id="B945EAD6FF8CAF712BC53E8EFD45FB3B" box="[472,640,1159,1185]" italics="true" pageId="27" pageNumber="33">
<taxonomicName id="4C314D47FF8CAF712BC53E8EFDB8FB3B" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[472,637,1159,1185]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="phalluspilosus">phalluspilosus</taxonomicName>
,
</emphasis>
(
<emphasis id="B945EAD6FF8CAF71288C3E8EFD26FB3B" box="[657,739,1159,1185]" italics="true" pageId="27" pageNumber="33">phallus</emphasis>
: penis plus 
<emphasis id="B945EAD6FF8CAF7129743E8EFC7DFB3B" box="[873,952,1159,1185]" italics="true" pageId="27" pageNumber="33">pilosus</emphasis>
: covered by hair), referring to male genitalia strongly covered by setae.
</paragraph>
</subSubSection>
<subSubSection id="C32B654FFF8CAF712ADA3EC7FB96F9B7" pageId="27" pageNumber="33" type="discussion">
<paragraph id="8B8E36C4FF8CAF712ADA3EC7FB96F9B7" blockId="27.[151,1437,151,1581]" pageId="27" pageNumber="33">
<emphasis id="B945EAD6FF8CAF712ADA3EC7FE7DFB73" bold="true" box="[199,440,1230,1257]" pageId="27" pageNumber="33">Comparative notes:</emphasis>
<taxonomicName id="4C314D47FF8CAF712BD93EC6FCD3FB73" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[452,790,1231,1257]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="phalluspilosus" status="sp. nov.">
<emphasis id="B945EAD6FF8CAF712BD93EC6FCD3FB73" box="[452,790,1231,1257]" italics="true" pageId="27" pageNumber="33">Hexacylloepus phalluspilosus</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FF8CAF71293F3EC6FC45FB73" bold="true" box="[802,896,1231,1257]" pageId="27" pageNumber="33">
<taxonomicNameLabel id="A27657ADFF8CAF71293F3EC6FC45FB73" box="[802,896,1231,1257]" pageId="27" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles 
<taxonomicName id="4C314D47FF8CAF712E153EC6FF1CFA97" authority="(Grouvelle, 1889)" baseAuthorityName="Grouvelle" baseAuthorityYear="1889" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="B945EAD6FF8CAF712E153EC6FAD5FB73" box="[1032,1296,1231,1257]" italics="true" pageId="27" pageNumber="33">Hexacylloepus flavipes</emphasis>
(
<bibRefCitation id="EFA04B35FF8CAF712F393EC6FF15FA97" author="Grouvelle, A." pageId="27" pageNumber="33" pagination="393 - 410" refId="ref25811" refString="Grouvelle, A. (1889) Nouvelles especes d'Helmides. Annales de la Societe Entomologique de France, 8 (6), 393 - 410." type="journal article" year="1889">Grouvelle, 1889</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C314D47FF8CAF712B0C3EFAFD3DFA97" authority="(Grouvelle, 1889)" baseAuthorityName="Grouvelle" baseAuthorityYear="1889" box="[273,760,1267,1293]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B945EAD6FF8CAF712B0C3EFAFDE1FA96" box="[273,548,1267,1293]" italics="true" pageId="27" pageNumber="33">Hexacylloepus granosus</emphasis>
(
<bibRefCitation id="EFA04B35FF8CAF7128293EFAFD35FA97" author="Grouvelle, A." box="[564,752,1267,1293]" pageId="27" pageNumber="33" pagination="393 - 410" refId="ref25811" refString="Grouvelle, A. (1889) Nouvelles especes d'Helmides. Annales de la Societe Entomologique de France, 8 (6), 393 - 410." type="journal article" year="1889">Grouvelle, 1889</bibRefCitation>
)
</taxonomicName>
, by having the pronotum as long as wide, Hypomera with a complete belt of tomentum covering almost the entire anterior half portion, and frontoclypeal suture inconspicuous. However, the general morphology of the male genitalia of the three species is completely different, and, in addition, the hind wings are micropterous in 
<emphasis id="B945EAD6FF8CAF7128433F56FCECFAE3" box="[606,809,1375,1401]" italics="true" pageId="27" pageNumber="33">
<taxonomicName id="4C314D47FF8CAF7128433F56FCE0FAE3" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[606,805,1375,1401]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="phalluspilosus">H. phalluspilosus</taxonomicName>
,
</emphasis>
while 
<taxonomicName id="4C314D47FF8CAF7129683F56FC2BFAE3" baseAuthorityName="Grouvelle" baseAuthorityYear="1889" box="[885,1006,1375,1401]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="B945EAD6FF8CAF7129683F56FC2BFAE3" box="[885,1006,1375,1401]" italics="true" pageId="27" pageNumber="33">H. flavipes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C314D47FF8CAF712E383F56FB6BFAE2" baseAuthorityName="Grouvelle" baseAuthorityYear="1889" box="[1061,1198,1375,1400]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B945EAD6FF8CAF712E383F56FB6BFAE2" box="[1061,1198,1375,1400]" italics="true" pageId="27" pageNumber="33">H. granosus</emphasis>
</taxonomicName>
are macropterous. In 
<taxonomicName id="4C314D47FF8CAF712A8A3F8AFED4FA07" baseAuthorityName="Grouvelle" baseAuthorityYear="1889" box="[151,273,1411,1437]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="flavipes">
<emphasis id="B945EAD6FF8CAF712A8A3F8AFED4FA07" box="[151,273,1411,1437]" italics="true" pageId="27" pageNumber="33">H. flavipes</emphasis>
</taxonomicName>
the tibiae each have a row of short spines at the distal 2/3 and, in males, a proleg with the internal claw shorter than the external claw, while in 
<taxonomicName id="4C314D47FF8CAF7128523FAEFCD7FA5B" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[591,786,1447,1473]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="phalluspilosus" status="sp. nov.">
<emphasis id="B945EAD6FF8CAF7128523FAEFCD7FA5B" box="[591,786,1447,1473]" italics="true" pageId="27" pageNumber="33">H. phalluspilosus</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FF8CAF7129043FAEFCB7FA5B" bold="true" box="[793,882,1447,1473]" pageId="27" pageNumber="33">
<taxonomicNameLabel id="A27657ADFF8CAF7129043FAEFCB7FA5B" box="[793,882,1447,1473]" pageId="27" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the tibiae each have two complete, dense rows of spines, and the claws on the proleg are of similar size. 
<taxonomicName id="4C314D47FF8CAF7128EE3FC2FCB9FA7E" baseAuthorityName="Grouvelle" baseAuthorityYear="1889" box="[755,892,1483,1508]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="granosus">
<emphasis id="B945EAD6FF8CAF7128EE3FC2FCB9FA7E" box="[755,892,1483,1508]" italics="true" pageId="27" pageNumber="33">H. granosus</emphasis>
</taxonomicName>
has the longitudinal impression on the pronotum long and very narrow and the metacoxae each with a short spine, while in 
<taxonomicName id="4C314D47FF8CAF7129D23FE6FB56F993" authority="Polizei &amp; Barclay &amp; Bispo, 2020" authorityName="Polizei &amp; Barclay &amp; Bispo" authorityYear="2020" box="[975,1171,1519,1545]" class="Insecta" family="Elmidae" genus="Hexacylloepus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="33" phylum="Arthropoda" rank="species" species="phalluspilosus" status="sp. nov.">
<emphasis id="B945EAD6FF8CAF7129D23FE6FB56F993" box="[975,1171,1519,1545]" italics="true" pageId="27" pageNumber="33">H. phalluspilosus</emphasis>
</taxonomicName>
<emphasis id="B945EAD6FF8CAF712E843FE6FB37F993" bold="true" box="[1177,1266,1519,1545]" pageId="27" pageNumber="33">
<taxonomicNameLabel id="A27657ADFF8CAF712E843FE6FB37F993" box="[1177,1266,1519,1545]" pageId="27" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the impression on the pronotum is short, narrow and shallow on disc, and the metacoxae lack spines.
</paragraph>
</subSubSection>
</treatment>
</document>