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<document id="77116CE46C6742C97A7FC394C74D3916" ID-CLB-Dataset="22138" ID-DOI="10.5852/ejt.2017.316" ID-GBIF-Dataset="42070ad7-c2d0-418a-8742-bafd806a0b35" ID-ISSN="2118-9773" ID-Zenodo-Dep="3827391" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589469189246" checkinUser="carolina" docAuthor="Olesen, Jørgen, Meland, Kenneth, Glenner, Henrik, Van Hengstum, Peter J. &amp; Iliffe, Thomas M." docDate="2017" docId="03984D24FFCB6449FDE5896EFCF4AA4A" docLanguage="en" docName="ejt-2017-316.pdf.imf" docOrigin="European Journal of Taxonomy 316" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Xibalbanus cozumelensis Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe 2017, sp. nov." docType="treatment" docUuid="B1758E15-23A9-4432-A9D5-DBD10EF11233" docUuidSource="ZooBank" docVersion="9" lastPageNumber="20" masterDocId="FFA1355CFFC3645AFFB58A3DFF9FA828" masterDocTitle="Xibalbanus cozumelensis, a new species of Remipedia (Crustacea) from Cozumel, Mexico, and a molecular phylogeny of Xibalbanus on the Yucatán Peninsula" masterLastPageNumber="27" masterPageNumber="1" pageNumber="9" updateTime="1698839709709" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B2F2A16C17FE6AC668966F6C97E51F34">Xibalbanus cozumelensis, a new species of Remipedia (Crustacea) from Cozumel, Mexico, and a molecular phylogeny of Xibalbanus on the Yucatán Peninsula</mods:title>
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<mods:namePart id="50F79E22FCACC1F10AA4896228F6F655">Olesen, Jørgen</mods:namePart>
<mods:affiliation id="E22A6BB7E515242447A7D7BD17445AA9">Natural History Museum of Denmark, University of Copenhagen, Denmark. &amp; Corresponding author: jolesen @ snm. ku. dk</mods:affiliation>
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<mods:namePart id="D3708E7740B6E11074A4E4A995DEA02E">Meland, Kenneth</mods:namePart>
<mods:affiliation id="3461C3ADF27ED6A4AFF71C0FA927D9AE">Department of Biology, University of Bergen, Norway. &amp; Email: Kenneth. Meland @ uib. no</mods:affiliation>
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<mods:affiliation id="BFECA8DC1AC95D5EDD0FAE311C1D8BC9">Department of Biology, University of Bergen, Norway. &amp; Email: Henrik. Glenner @ uib. no</mods:affiliation>
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<mods:namePart id="F7983AB96192450C447C268A3707B95F">Van Hengstum, Peter J.</mods:namePart>
<mods:affiliation id="70BE54879A3897D34F0A1450B5533FB8">Department of Marine Sciences, Texas A &amp; M University at Galveston, USA. &amp; Email: vanhenp @ tamug. edu</mods:affiliation>
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<emphasis id="B9452020FFCB6452FDE5896EFC1DAB46" bold="true" box="[592,898,851,878]" italics="true" pageId="8" pageNumber="9">Xibalbanus cozumelensis</emphasis>
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<figureCitation id="130AE0B7FFCB6452FD6A89A7FCCBAB9C" box="[735,852,922,948]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStartId-0="9.[189,232,1929,1955]" captionStartId-1="11.[189,232,1893,1919]" captionStartId-2="12.[189,232,1790,1816]" captionStartId-3="13.[189,232,1789,1815]" captionStartId-4="15.[189,232,1860,1886]" captionStartId-5="16.[189,232,1792,1818]" captionStartId-6="17.[189,232,1929,1955]" captionStartId-7="18.[189,232,1824,1850]" captionTargetBox-0="[189,1398,901,1904]" captionTargetBox-1="[231,1357,263,1858]" captionTargetBox-2="[229,1358,264,1753]" captionTargetBox-3="[276,1311,262,1755]" captionTargetBox-4="[236,1353,268,1818]" captionTargetBox-5="[276,1313,267,1763]" captionTargetBox-6="[189,1398,678,1893]" captionTargetBox-7="[190,1398,436,1789]" captionTargetId-0="figure@9.[189,1398,901,1904]" captionTargetId-1="figure@11.[231,1357,263,1858]" captionTargetId-2="figure@12.[229,1358,263,1753]" captionTargetId-3="figure@13.[276,1311,261,1755]" captionTargetId-4="figure@15.[234,1353,268,1818]" captionTargetId-5="figure@16.[274,1313,265,1763]" captionTargetId-6="figure@17.[189,1398,678,1893]" captionTargetId-7="figure@18.[189,1398,435,1790]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionTargetPageId-2="12" captionTargetPageId-3="13" captionTargetPageId-4="15" captionTargetPageId-5="16" captionTargetPageId-6="17" captionTargetPageId-7="18" captionText-0="Fig. 3. Xibalbanus cozumelensis sp. nov., light microscopy. A. Holotype, adult specimen, lateral view (ZMUC-CRU-4791). B. Holotype, cephalon, lateral view. C. Paratype, cephalon, dorsal view (ZMUC- CRU-4793)." captionText-1="Fig. 4. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), light microscopy. A. Antenna 1, left side from anterior. B. Antenna 2, left side from anterior. C. Maxilla 1, left side from anterior. D. Maxilla 2, left side from anterior. E. Maxilliped, left side from anterior. F. Caudal rami and posterior part of trunk. G. Caudal ramus, left side seen from median." captionText-2="Fig. 5. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792). Cephalon and cephalic structures. Scanning electron microscopy. A. Cephalon with cephalic shield and appendages. B. Antenna 1, right side, proximal parts of dorsal and ventral branches. C–D. Unidentified suctorians attached at the proximo-ventral parts of antenna 1. E. Labrum, frontal filaments, and parts of antennae 1 and 2, right side. F. Frontal filaments. G. Antenna 2, left side from anterior. H. Mandible, left side from anterior. I. Mandible, left side from posterior. J. Mandible, left side, apical view on gnathal edge. K. Mandible, left side, apical view on molar process." captionText-3="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." captionText-4="Fig. 7. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. A. Complete mx2 from anterior. B. Close-up of terminal claw-complex from anterior (segment 7). C. Close-up of terminal claw-complex from lateral (segment 7). D. Median side of segments 1 and 2 with endites (e1–3) from anterior. E. Endites of segment 1 (e1–3) from median. F. Complete mx2 from median." captionText-5="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." captionText-6="Fig. 9. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), trunk limbs, left side from anterior, light microscopy. A–J. Selected trunk segments (number specified on figure). All to the same scale." captionText-7="Fig. 10. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), cephalon, body segments, and trunk limbs, scanning electron microscopy. A. Cephalon, cephalic shield, and head appendages from lateral. B. Trunk segments 5–10. C. Trunk limb 2, right side. D. Close-up of serrated setae in latero-distal corner of exopod segment 1. E. Close-up of serrated setae in latero-distal corner of exopod segment 2. F–G. Two unidentified suctorians attached to exopod of trunk limb 3. H–I. Posterior body segments, trunk limbs, and caudal rami from ventral and lateral." figureDoi-0="http://doi.org/10.5281/zenodo.3827397" figureDoi-1="http://doi.org/10.5281/zenodo.3827399" figureDoi-2="http://doi.org/10.5281/zenodo.3827401" figureDoi-3="http://doi.org/10.5281/zenodo.3827403" figureDoi-4="http://doi.org/10.5281/zenodo.3827405" figureDoi-5="http://doi.org/10.5281/zenodo.3827407" figureDoi-6="http://doi.org/10.5281/zenodo.3827409" figureDoi-7="http://doi.org/10.5281/zenodo.3827411" httpUri-0="https://zenodo.org/record/3827397/files/figure.png" httpUri-1="https://zenodo.org/record/3827399/files/figure.png" httpUri-2="https://zenodo.org/record/3827401/files/figure.png" httpUri-3="https://zenodo.org/record/3827403/files/figure.png" httpUri-4="https://zenodo.org/record/3827405/files/figure.png" httpUri-5="https://zenodo.org/record/3827407/files/figure.png" httpUri-6="https://zenodo.org/record/3827409/files/figure.png" httpUri-7="https://zenodo.org/record/3827411/files/figure.png" pageId="8" pageNumber="9">Figs 3–10</figureCitation>
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<taxonomicName id="4C3187B1FFCB6452FF0889E6FECAABDD" ID-CoL="84TQT" authorityName="Yager" authorityYear="1981" box="[189,341,987,1013]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B9452020FFCB6452FF0889E6FECAABDD" box="[189,341,987,1013]" italics="true" pageId="8" pageNumber="9">Speleonectes</emphasis>
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sp. – 
<treatmentCitation id="0A90DA23FFCB6452FE2C89E6FC92ABDE" author="Yanez-Mendoza G. &amp; Zarza-Gonzalez E. &amp; Mejia-Ortiz L. M." box="[409,781,987,1014]" page="54" pageId="8" pageNumber="9" year="2007">
<bibRefCitation id="EFA081C3FFCB6452FE2C89E6FC92ABDE" author="Yanez-Mendoza G. &amp; Zarza-Gonzalez E. &amp; Mejia-Ortiz L. M." box="[409,781,987,1014]" pageId="8" pageNumber="9" pagination="49 - 70" refId="ref13029" refString="Yanez-Mendoza G., Zarza-Gonzalez E. &amp; Mejia-Ortiz L. M. 2007. Sistemas anquihalinos. In: Mejia- Ortiz L. M. (ed.) Biodiversidad acuatica de la Isla de Cozumel: 49 - 70. Universidad de Quintana Roo- CONABIO, Plaza y Valdes, Mexico." type="book chapter" year="2007">
Yañez-Mendoza 
<emphasis id="B9452020FFCB6452FDD589E1FD0CABDD" box="[608,659,987,1013]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. 2007: 54
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.
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<emphasis id="B9452020FFCB6452FF088E21FEA9AC1E" bold="true" box="[189,310,1052,1078]" pageId="8" pageNumber="9">Diagnosis</emphasis>
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<taxonomicName id="4C3187B1FFCB6452FF088E76FE78AC4D" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[189,487,1099,1125]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFCB6452FF088E76FE78AC4D" box="[189,487,1099,1125]" italics="true" pageId="8" pageNumber="9">Xibalbanus cozumelensis</emphasis>
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is a slender and comparatively large species (
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), up to 
<quantity id="4CC951D7FFCB6452FA978E76FAE8AC4E" box="[1314,1399,1099,1126]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="9" unit="mm" value="35.0">35 mm</quantity>
in length; largest specimens with 39 (38 limb-bearing) trunk segments. The species is very similar to 
<taxonomicName id="4C3187B1FFCB6452FF088EAFFDE6AC84" authority="(Yager, 1987)" baseAuthorityName="Yager" baseAuthorityYear="1987" box="[189,633,1170,1196]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="tulumensis">
<emphasis id="B9452020FFCB6452FF088EAFFE56AC84" box="[189,457,1170,1196]" italics="true" pageId="8" pageNumber="9">Xibalbanus tulumensis</emphasis>
(
<bibRefCitation id="EFA081C3FFCB6452FE698EAFFDEFAC84" author="Yager J." box="[476,624,1170,1196]" pageId="8" pageNumber="9" pagination="160 - 166" refId="ref12897" refString="Yager J. 1987. Speleonectes tulumensis n. sp. (Crustacea: Remipedia) from two anchialine cenotes of the Yucatan Peninsula, Mexico. Stygologia 3: 160 - 166." type="journal article" year="1987">Yager, 1987</bibRefCitation>
)
</taxonomicName>
, but is genetically different from this species and is so far only known from Cozumel.
</paragraph>
</subSubSection>
<subSubSection id="C32BAFB9FFCB6452FF088ECAFCF7AD17" box="[189,872,1271,1344]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8B8EFC32FFCB6452FF088ECAFEDBAD39" blockId="8.[189,324,1271,1297]" box="[189,324,1271,1297]" pageId="8" pageNumber="9">
<heading id="D0C64B5EFFCB6452FF088ECAFEDBAD39" bold="true" box="[189,324,1271,1297]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B9452020FFCB6452FF088ECAFEDBAD39" bold="true" box="[189,324,1271,1297]" pageId="8" pageNumber="9">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8EFC32FFCB6452FF088F18FCF7AD17" blockId="8.[189,872,1317,1344]" box="[189,872,1317,1344]" pageId="8" pageNumber="9">
The specific epithet refers to the 
<typeStatus id="548A4290FFCB6452FD888F1BFDEFAD68" box="[573,624,1318,1344]" pageId="8" pageNumber="9">type</typeStatus>
locality of Cozumel.
</paragraph>
</subSubSection>
<subSubSection id="C32BAFB9FFCB6452FF088F5BFAE8AEEC" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8B8EFC32FFCB6452FF088F5BFE35ADA8" blockId="8.[189,426,1382,1408]" box="[189,426,1382,1408]" pageId="8" pageNumber="9">
<heading id="D0C64B5EFFCB6452FF088F5BFE35ADA8" bold="true" box="[189,426,1382,1408]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B9452020FFCB6452FF088F5BFE35ADA8" bold="true" box="[189,426,1382,1408]" pageId="8" pageNumber="9">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="3B59F66FFFCB6452FF5F8FA8FE65ADDE" ID-GBIF-Occurrence="2618819301" collectingDate="2014-07-07" collectionCode="T" collectorName="Iliffe &amp; P. Van Hengstum" country="Mexico" location="MEXICO" pageId="8" pageNumber="9" specimenCode="ZMUC-CRU-4791" specimenCount="1" typeStatus="holotype">
<paragraph id="8B8EFC32FFCB6452FF5F8FA8FEC3AD87" blockId="8.[189,1399,1429,1526]" box="[234,348,1429,1455]" pageId="8" pageNumber="9">
<emphasis id="B9452020FFCB6452FF5F8FA8FEC3AD87" bold="true" box="[234,348,1429,1455]" pageId="8" pageNumber="9">
<typeStatus id="548A4290FFCB6452FF5F8FA8FEC3AD87" box="[234,348,1429,1455]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8EFC32FFCB6452FF088F85FE65ADDE" blockId="8.[189,1399,1429,1526]" pageId="8" pageNumber="9">
<collectingCountry id="F326BCA2FFCB6452FF088F85FEAAADFA" box="[189,309,1464,1490]" name="Mexico" pageId="8" pageNumber="9">MEXICO</collectingCountry>
: 
<quantity id="4CC951D7FFCB6452FE8A8F85FE0BADFB" box="[319,404,1464,1491]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="8" pageNumber="9" unit="mm" value="33.0">33 mm</quantity>
, 39 trunk segments (38 bearing trunk limbs), T. 
<collectorName id="26C499E4FFCB6452FC758F85FC65ADFA" box="[960,1018,1464,1490]" pageId="8" pageNumber="9">Iliffe</collectorName>
and 
<collectorName id="26C499E4FFCB6452FB848F84FB66ADFB" box="[1073,1273,1465,1491]" pageId="8" pageNumber="9">P. Van Hengstum</collectorName>
leg., 
<date id="FF8FDAF2FFCB6452FA8D8F84FF67ADDE" pageId="8" pageNumber="9" value="2014-07-07">
<collectingDate id="EFCB231AFFCB6452FA8D8F84FF67ADDE" pageId="8" pageNumber="9" value="2014-07-07">7 Jul. 2014</collectingDate>
</date>
(
<specimenCode id="DB975449FFCB6452FEB28FE1FE72ADDE" box="[263,493,1500,1526]" pageId="8" pageNumber="9">ZMUC-CRU-4791</specimenCode>
).
</paragraph>
</materialsCitation>
<paragraph id="8B8EFC32FFCB6452FF5F8C20FEF8AE1F" blockId="8.[189,1399,1565,1733]" box="[234,359,1565,1591]" pageId="8" pageNumber="9">
<emphasis id="B9452020FFCB6452FF5F8C20FEF8AE1F" bold="true" box="[234,359,1565,1591]" pageId="8" pageNumber="9">
<typeStatus id="548A4290FFCB6452FF5F8C20FEF8AE1F" box="[234,359,1565,1591]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B8EFC32FFCB6452FF088C7DFAE8AEEC" blockId="8.[189,1399,1565,1733]" pageId="8" pageNumber="9">
<collectingCountry id="F326BCA2FFCB6452FF088C7DFEAEAE72" box="[189,305,1600,1626]" name="Mexico" pageId="8" pageNumber="9">MEXICO</collectingCountry>
: 
<specimenCount id="9D3737BBFFCB6452FEF58C7DFE4FAE72" box="[320,464,1600,1626]" pageId="8" pageNumber="9" type="generic">2 specimens</specimenCount>
(bearing 37 and 38 trunk limbs respectively), same locality as 
<typeStatus id="548A4290FFCB6452FB018C7DFA81AE72" box="[1204,1310,1600,1626]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, part of one individual removed for molecular analysis, other parts dissected for SEM and mounted separately on stubs, 
<collectionCode id="ED2064F7FFCB6452FE9C8CBAFEA3AE89" box="[297,316,1671,1697]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="9">T</collectionCode>
. Iliffe and P. Van Hengstum leg., 
<date id="FF8FDAF2FFCB6452FD0E8CBAFCA5AE89" box="[699,826,1671,1697]" pageId="8" pageNumber="9" value="2014-07-07">7 Jul. 2014</date>
(ZMUC-CRU-4792); 
<specimenCount id="9D3737BBFFCB6452FBF58CBAFB4DAE89" box="[1088,1234,1671,1697]" pageId="8" pageNumber="9" type="generic">2 specimens</specimenCount>
, same locality as 
<typeStatus id="548A4290FFCB6452FF6E8C97FEDAAEEC" box="[219,325,1706,1732]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED2064F7FFCB6452FEFA8C96FEFDAEED" box="[335,354,1707,1733]" name="Tavera, Department of Geology and Geophysics" pageId="8" pageNumber="9">T</collectionCode>
. Iliffe and P. Van Hengstum leg., 
<date id="FF8FDAF2FFCB6452FD548C97FCFEAEEC" box="[737,865,1706,1732]" pageId="8" pageNumber="9" value="2014-07-03">3 Jul. 2014</date>
(ZMUC-CRU-4793 and ZMUC-CRU-4794).
</paragraph>
</subSubSection>
<subSubSection id="C32BAFB9FFCB6452FF088CD1FC56AF7F" pageId="8" pageNumber="9" type="distribution">
<paragraph id="8B8EFC32FFCB6452FF088CD1FEC2AF2D" blockId="8.[189,349,1771,1798]" box="[189,349,1771,1798]" pageId="8" pageNumber="9">
<heading id="D0C64B5EFFCB6452FF088CD1FEC2AF2D" bold="true" box="[189,349,1771,1798]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B9452020FFCB6452FF088CD1FEC2AF2D" bold="true" box="[189,349,1771,1798]" pageId="8" pageNumber="9">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B8EFC32FFCB6452FF088D27FC56AF7F" blockId="8.[189,1398,1818,1880]" pageId="8" pageNumber="9">
<collectingCountry id="F326BCA2FFCB6452FF088D27FEAAAF1C" box="[189,309,1818,1844]" name="Mexico" pageId="8" pageNumber="9">MEXICO</collectingCountry>
: Cueva Quebrada, a flooded cave system in Chankannaab National Park on the west coast of Cozumel in 
<collectingRegion id="49F532D0FFCB6452FEF98D00FE6CAF70" box="[332,499,1853,1880]" country="Mexico" name="Quintana Roo" pageId="8" pageNumber="9">Quintana Roo</collectingRegion>
, 
<collectingRegion id="49F532D0FFCB6452FE4A8D00FDC6AF7F" box="[511,601,1853,1879]" country="Mexico" name="Mexico" pageId="8" pageNumber="9">Mexico</collectingRegion>
at 
<geoCoordinate id="EE059AF5FFCB6452FDC98D01FD77AF70" box="[636,744,1852,1880]" degrees="20" direction="north" minutes="26" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="20.433332">20°26′ N</geoCoordinate>
, 
<geoCoordinate id="EE059AF5FFCB6452FD418D01FCFCAF70" box="[756,867,1852,1880]" degrees="86" direction="west" minutes="59" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="-86.98333">86°59′ W</geoCoordinate>
(
<figureCitation id="130AE0B7FFCB6452FCC68D00FC24AF7F" box="[883,955,1853,1879]" captionStart="Fig" captionStartId="3.[189,232,1613,1639]" captionTargetBox="[193,1392,264,1601]" captionTargetId="figure@3.[193,1392,264,1601]" captionTargetPageId="3" captionText="Fig. 2. Maps of the Yucatán Peninsula showing the known distribution of four species of Xibalbanus (Remipedia) and water depths around Cozumel. A. Yucatán Peninsula showing Xibalbanus localities (1–10) in the northeastern region of the Yucatán Peninsula and one locality in Belize further south (11). B. Close-up of the northeastern region of the Yucatán Peninsula with the positions of 10 Xibalbanus localities indicated. C. Close-up of eastern Cozumel with the branches of the anchialine cave “Cueva Quebrada” indicated as an overlay, derived from Yañez-Mendoza et al. (2007). The collecting site of Xibalbanus cozumelensis sp. nov. is indicated as “10”. Numbers refer to the following cenotes/caves: 1 = Cenote Crustacea; 2 = Cenote Chac Mool; 3 = Cenote Ponderosa (= Cenote Jardín del Edén); 4 = Cenote Tajma Ha; 5 = Cenote Carwash (=Cenote Aktun Ha); 6 = Cenote Vaca Ha; 7 = Cenote Temple of Doom (Cenote Calavera, Cenote Esqueleto); 8 = Cenote Naharon (Cenote Cristal); 9 = Cenote Maya Blue (Cenote Escondido); 10 = Cueva Quebrada; 11 = Caye Chapel Cave. 1–9 are in the northeastern Yucatán Peninsula, 10 is on the island of Cozumel, 11 is in Belize." figureDoi="http://doi.org/10.5281/zenodo.3827395" httpUri="https://zenodo.org/record/3827395/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32BAFB9FFCB6454FF088D42FEF4ADE3" lastPageId="14" lastPageNumber="15" pageId="8" pageNumber="9" type="description">
<paragraph id="8B8EFC32FFCB6452FF088D42FED1AFB1" blockId="8.[189,334,1919,1945]" box="[189,334,1919,1945]" pageId="8" pageNumber="9">
<heading id="D0C64B5EFFCB6452FF088D42FED1AFB1" bold="true" box="[189,334,1919,1945]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B9452020FFCB6452FF088D42FED1AFB1" bold="true" box="[189,334,1919,1945]" pageId="8" pageNumber="9">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8EFC32FFCB6453FF088D93FCC8A9D1" blockId="8.[189,1399,1965,2027]" lastBlockId="9.[189,1399,268,506]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
BODY. Elongate, without eyes or pigment (
<figureCitation id="130AE0B7FFCB6452FD1D8D90FD71AFE0" box="[680,750,1965,1992]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[189,1398,901,1904]" captionTargetId="figure@9.[189,1398,901,1904]" captionTargetPageId="9" captionText="Fig. 3. Xibalbanus cozumelensis sp. nov., light microscopy. A. Holotype, adult specimen, lateral view (ZMUC-CRU-4791). B. Holotype, cephalon, lateral view. C. Paratype, cephalon, dorsal view (ZMUC- CRU-4793)." figureDoi="http://doi.org/10.5281/zenodo.3827397" httpUri="https://zenodo.org/record/3827397/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
). Maximum length of specimens examined 
<quantity id="4CC951D7FFCB6452FB5E8D90FAA0AFE0" box="[1259,1343,1965,1992]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="9" unit="mm" value="35.0">35 mm</quantity>
. The shortest specimen examined is 
<quantity id="4CC951D7FFCB6452FD948DECFDEBAFC3" box="[545,628,2001,2027]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="9" unit="mm" value="30.0">30 mm</quantity>
. Cephalic shield small, tapering anteriorly with two lateral notches, about 1/12 of the total body length; cephalic shield partly covering first trunk segment and proximal parts of cephalic appendages; with distinct transverse suture in the region of maxillae 1, and another transverse but weaker suture in the region of antennae 1 (
<figureCitation id="130AE0B7FFCA6453FCCF8B6EFC42A945" box="[890,989,339,365]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[189,1398,901,1904]" captionTargetId="figure@9.[189,1398,901,1904]" captionTargetPageId="9" captionText="Fig. 3. Xibalbanus cozumelensis sp. nov., light microscopy. A. Holotype, adult specimen, lateral view (ZMUC-CRU-4791). B. Holotype, cephalon, lateral view. C. Paratype, cephalon, dorsal view (ZMUC- CRU-4793)." figureDoi="http://doi.org/10.5281/zenodo.3827397" httpUri="https://zenodo.org/record/3827397/files/figure.png" pageId="9" pageNumber="10">Figs 3C</figureCitation>
, 
<figureCitation id="130AE0B7FFCA6453FC598B6EFBBCA945" box="[1004,1059,339,365]" captionStart="Fig" captionStartId="18.[189,232,1824,1850]" captionTargetBox="[190,1398,436,1789]" captionTargetId="figure@18.[189,1398,435,1790]" captionTargetPageId="18" captionText="Fig. 10. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), cephalon, body segments, and trunk limbs, scanning electron microscopy. A. Cephalon, cephalic shield, and head appendages from lateral. B. Trunk segments 5–10. C. Trunk limb 2, right side. D. Close-up of serrated setae in latero-distal corner of exopod segment 1. E. Close-up of serrated setae in latero-distal corner of exopod segment 2. F–G. Two unidentified suctorians attached to exopod of trunk limb 3. H–I. Posterior body segments, trunk limbs, and caudal rami from ventral and lateral." figureDoi="http://doi.org/10.5281/zenodo.3827411" httpUri="https://zenodo.org/record/3827411/files/figure.png" pageId="9" pageNumber="10">10A</figureCitation>
). Trunk elongate, maximum number of trunk segments of specimens examined 39 (38 limb bearing). Most trunk segments with similar subrectangular tergites with rounded lateral margins (
<figureCitation id="130AE0B7FFCA6453FC358BA4FC75A99B" box="[896,1002,409,435]" captionStart="Fig" captionStartId="18.[189,232,1824,1850]" captionTargetBox="[190,1398,436,1789]" captionTargetId="figure@18.[189,1398,435,1790]" captionTargetPageId="18" captionText="Fig. 10. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), cephalon, body segments, and trunk limbs, scanning electron microscopy. A. Cephalon, cephalic shield, and head appendages from lateral. B. Trunk segments 5–10. C. Trunk limb 2, right side. D. Close-up of serrated setae in latero-distal corner of exopod segment 1. E. Close-up of serrated setae in latero-distal corner of exopod segment 2. F–G. Two unidentified suctorians attached to exopod of trunk limb 3. H–I. Posterior body segments, trunk limbs, and caudal rami from ventral and lateral." figureDoi="http://doi.org/10.5281/zenodo.3827411" httpUri="https://zenodo.org/record/3827411/files/figure.png" pageId="9" pageNumber="10">Fig. 10B</figureCitation>
); posteriormost segments reduced with posteriorly pointing lateral margins of tergites (
<figureCitation id="130AE0B7FFCA6453FCAA8B81FC1FA9FE" box="[799,896,444,470]" captionStart="Fig" captionStartId="18.[189,232,1824,1850]" captionTargetBox="[190,1398,436,1789]" captionTargetId="figure@18.[189,1398,435,1790]" captionTargetPageId="18" captionText="Fig. 10. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), cephalon, body segments, and trunk limbs, scanning electron microscopy. A. Cephalon, cephalic shield, and head appendages from lateral. B. Trunk segments 5–10. C. Trunk limb 2, right side. D. Close-up of serrated setae in latero-distal corner of exopod segment 1. E. Close-up of serrated setae in latero-distal corner of exopod segment 2. F–G. Two unidentified suctorians attached to exopod of trunk limb 3. H–I. Posterior body segments, trunk limbs, and caudal rami from ventral and lateral." figureDoi="http://doi.org/10.5281/zenodo.3827411" httpUri="https://zenodo.org/record/3827411/files/figure.png" pageId="9" pageNumber="10">Fig. 10I</figureCitation>
). Appendages are lacking on posteriormost segment (
<figureCitation id="130AE0B7FFCA6453FE9A8BE2FE16A9D2" box="[303,393,479,506]" captionStart="Fig" captionStartId="11.[189,232,1893,1919]" captionTargetBox="[231,1357,263,1858]" captionTargetId="figure@11.[231,1357,263,1858]" captionTargetPageId="11" captionText="Fig. 4. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), light microscopy. A. Antenna 1, left side from anterior. B. Antenna 2, left side from anterior. C. Maxilla 1, left side from anterior. D. Maxilla 2, left side from anterior. E. Maxilliped, left side from anterior. F. Caudal rami and posterior part of trunk. G. Caudal ramus, left side seen from median." figureDoi="http://doi.org/10.5281/zenodo.3827399" httpUri="https://zenodo.org/record/3827399/files/figure.png" pageId="9" pageNumber="10">Fig. 4F</figureCitation>
). Sternal bars very narrow, isomorphic.
</paragraph>
<paragraph id="8B8EFC32FFCA6453FF08881CFED1AA77" blockId="9.[189,1399,545,607]" pageId="9" pageNumber="10">
FRONTAL FILAMENTS. Long, distally bifurcated into a large anterior branch and a smaller posterior branch (
<figureCitation id="130AE0B7FFCA6453FF738879FE94AA77" box="[198,267,580,607]" captionStart="Fig" captionStartId="12.[189,232,1790,1816]" captionTargetBox="[229,1358,264,1753]" captionTargetId="figure@12.[229,1358,263,1753]" captionTargetPageId="12" captionText="Fig. 5. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792). Cephalon and cephalic structures. Scanning electron microscopy. A. Cephalon with cephalic shield and appendages. B. Antenna 1, right side, proximal parts of dorsal and ventral branches. C–D. Unidentified suctorians attached at the proximo-ventral parts of antenna 1. E. Labrum, frontal filaments, and parts of antennae 1 and 2, right side. F. Frontal filaments. G. Antenna 2, left side from anterior. H. Mandible, left side from anterior. I. Mandible, left side from posterior. J. Mandible, left side, apical view on gnathal edge. K. Mandible, left side, apical view on molar process." figureDoi="http://doi.org/10.5281/zenodo.3827401" httpUri="https://zenodo.org/record/3827401/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
E–F).
</paragraph>
<paragraph id="8B8EFC32FFCA6453FF0888BBFC03AB5B" blockId="9.[189,1399,645,883]" pageId="9" pageNumber="10">
ANTENNA 1 (
<figureCitation id="130AE0B7FFCA6453FEFE88B8FE37AA88" box="[331,424,645,672]" captionStart="Fig" captionStartId="9.[189,232,1929,1955]" captionTargetBox="[189,1398,901,1904]" captionTargetId="figure@9.[189,1398,901,1904]" captionTargetPageId="9" captionText="Fig. 3. Xibalbanus cozumelensis sp. nov., light microscopy. A. Holotype, adult specimen, lateral view (ZMUC-CRU-4791). B. Holotype, cephalon, lateral view. C. Paratype, cephalon, dorsal view (ZMUC- CRU-4793)." figureDoi="http://doi.org/10.5281/zenodo.3827397" httpUri="https://zenodo.org/record/3827397/files/figure.png" pageId="9" pageNumber="10">Figs 3B</figureCitation>
, 
<figureCitation id="130AE0B7FFCA6453FE0688BBFE48AA88" box="[435,471,646,672]" captionStart="Fig" captionStartId="11.[189,232,1893,1919]" captionTargetBox="[231,1357,263,1858]" captionTargetId="figure@11.[231,1357,263,1858]" captionTargetPageId="11" captionText="Fig. 4. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), light microscopy. A. Antenna 1, left side from anterior. B. Antenna 2, left side from anterior. C. Maxilla 1, left side from anterior. D. Maxilla 2, left side from anterior. E. Maxilliped, left side from anterior. F. Caudal rami and posterior part of trunk. G. Caudal ramus, left side seen from median." figureDoi="http://doi.org/10.5281/zenodo.3827399" httpUri="https://zenodo.org/record/3827399/files/figure.png" pageId="9" pageNumber="10">4A</figureCitation>
, 
<figureCitation id="130AE0B7FFCA6453FE5088B8FD95AA88" box="[485,522,645,672]" captionStart="Fig" captionStartId="12.[189,232,1790,1816]" captionTargetBox="[229,1358,264,1753]" captionTargetId="figure@12.[229,1358,263,1753]" captionTargetPageId="12" captionText="Fig. 5. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792). Cephalon and cephalic structures. Scanning electron microscopy. A. Cephalon with cephalic shield and appendages. B. Antenna 1, right side, proximal parts of dorsal and ventral branches. C–D. Unidentified suctorians attached at the proximo-ventral parts of antenna 1. E. Labrum, frontal filaments, and parts of antennae 1 and 2, right side. F. Frontal filaments. G. Antenna 2, left side from anterior. H. Mandible, left side from anterior. I. Mandible, left side from posterior. J. Mandible, left side, apical view on gnathal edge. K. Mandible, left side, apical view on molar process." figureDoi="http://doi.org/10.5281/zenodo.3827401" httpUri="https://zenodo.org/record/3827401/files/figure.png" pageId="9" pageNumber="10">5B</figureCitation>
). Two-branched, slender, relatively short, about 1/6 of body length. Protopod 2-segmented; proximal segment is largest with rows of dense, long ribbon-like aesthetascs along median margin; distal segment continues into a long dorsal/lateral branch and a shorter ventral/median branch. Dorsal branch with 11 slender segments most of which have clusters of aesthetascs along medial margin, and small scattered setae, terminal segment with fine terminal setae. Ventral ramus short, less than ½ the length of dorsal ramus, 9 segments, basal segments weakly divided, clusters of distomedial aesthetascs and short setae on margins; distal segment with terminal setae.
</paragraph>
<caption id="DF4EACBAFFCA6453FF088DB4FED0AFC2" ID-DOI="http://doi.org/10.5281/zenodo.3827397" ID-Zenodo-Dep="3827397" httpUri="https://zenodo.org/record/3827397/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1929,1955]" targetBox="[189,1398,901,1904]" targetPageId="9">
<paragraph id="8B8EFC32FFCA6453FF088DB4FED0AFC2" blockId="9.[189,1398,1929,2026]" pageId="9" pageNumber="10">
<emphasis id="B9452020FFCA6453FF088DB4FE93AF8B" bold="true" box="[189,268,1929,1955]" pageId="9" pageNumber="10">Fig. 3.</emphasis>
<taxonomicName id="4C3187B1FFCA6453FEA18DB4FDA1AF8B" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[276,574,1929,1955]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFCA6453FEA18DB4FDA1AF8B" box="[276,574,1929,1955]" italics="true" pageId="9" pageNumber="10">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFCA6453FDF38DB7FD3BAF8C" box="[582,676,1930,1956]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
, light microscopy. 
<emphasis id="B9452020FFCA6453FC328DB4FC03AF8B" bold="true" box="[903,924,1929,1955]" pageId="9" pageNumber="10">A</emphasis>
. Holotype, adult specimen, lateral view (ZMUC-CRU-4791). 
<emphasis id="B9452020FFCA6453FE0A8D90FE4CAFEF" bold="true" box="[447,467,1965,1991]" pageId="9" pageNumber="10">B</emphasis>
. Holotype, cephalon, lateral view. 
<emphasis id="B9452020FFCA6453FCD98D91FC1EAFEE" bold="true" box="[876,897,1964,1990]" pageId="9" pageNumber="10">C</emphasis>
. Paratype, cephalon, dorsal view (ZMUC- CRU-4793).
</paragraph>
</caption>
<paragraph id="8B8EFC32FFC96450FF088B31FE13A9F0" blockId="10.[189,1399,267,472]" pageId="10" pageNumber="11">
ANTENNA 2 (
<figureCitation id="130AE0B7FFC96450FEE68B36FE2DA90D" box="[339,434,267,294]" captionStart="Fig" captionStartId="11.[189,232,1893,1919]" captionTargetBox="[231,1357,263,1858]" captionTargetId="figure@11.[231,1357,263,1858]" captionTargetPageId="11" captionText="Fig. 4. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), light microscopy. A. Antenna 1, left side from anterior. B. Antenna 2, left side from anterior. C. Maxilla 1, left side from anterior. D. Maxilla 2, left side from anterior. E. Maxilliped, left side from anterior. F. Caudal rami and posterior part of trunk. G. Caudal ramus, left side seen from median." figureDoi="http://doi.org/10.5281/zenodo.3827399" httpUri="https://zenodo.org/record/3827399/files/figure.png" pageId="10" pageNumber="11">Fig. 4B</figureCitation>
). A paddle-like biramous appendage. Protopod divided in two segments, a coxa (proximally) and a basis; coxa with 4 medial setae, basis with 9 medial setae. Endopod 3-segmented and curves laterally; segment 1 with 10–11 setae on lateral (outer) margin; segment 2 with 7; segment 3 ovoid, bearing about 27–28 setae along entire margin, including a terminal double row of about 10 setae. Exopod an unsegmented, large oval plate with about 29–35 setae along margin. All setae plumose with long setules.
</paragraph>
<paragraph id="8B8EFC32FFC96450FF08883CFAC0AA17" blockId="10.[189,1398,512,575]" pageId="10" pageNumber="11">
LABRUM (
<figureCitation id="130AE0B7FFC96450FE9F883DFE1CAA33" box="[298,387,512,539]" captionStart="Fig" captionStartId="12.[189,232,1790,1816]" captionTargetBox="[229,1358,264,1753]" captionTargetId="figure@12.[229,1358,263,1753]" captionTargetPageId="12" captionText="Fig. 5. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792). Cephalon and cephalic structures. Scanning electron microscopy. A. Cephalon with cephalic shield and appendages. B. Antenna 1, right side, proximal parts of dorsal and ventral branches. C–D. Unidentified suctorians attached at the proximo-ventral parts of antenna 1. E. Labrum, frontal filaments, and parts of antennae 1 and 2, right side. F. Frontal filaments. G. Antenna 2, left side from anterior. H. Mandible, left side from anterior. I. Mandible, left side from posterior. J. Mandible, left side, apical view on gnathal edge. K. Mandible, left side, apical view on molar process." figureDoi="http://doi.org/10.5281/zenodo.3827401" httpUri="https://zenodo.org/record/3827401/files/figure.png" pageId="10" pageNumber="11">Fig. 5E</figureCitation>
). Prominent, bulbous, subdivided by a transverse suture in an anterior trapezoid shaped part and a posterior (distal) subrectangular part rounded laterally; distally is a dense tuft of fine setae.
</paragraph>
<paragraph id="8B8EFC32FFC96450FF08885AFA8EAB54" blockId="10.[189,1399,614,892]" pageId="10" pageNumber="11">
MANDIBLES (
<figureCitation id="130AE0B7FFC96450FEF8885BFE10AAA9" box="[333,399,614,641]" captionStart="Fig" captionStartId="12.[189,232,1790,1816]" captionTargetBox="[229,1358,264,1753]" captionTargetId="figure@12.[229,1358,263,1753]" captionTargetPageId="12" captionText="Fig. 5. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792). Cephalon and cephalic structures. Scanning electron microscopy. A. Cephalon with cephalic shield and appendages. B. Antenna 1, right side, proximal parts of dorsal and ventral branches. C–D. Unidentified suctorians attached at the proximo-ventral parts of antenna 1. E. Labrum, frontal filaments, and parts of antennae 1 and 2, right side. F. Frontal filaments. G. Antenna 2, left side from anterior. H. Mandible, left side from anterior. I. Mandible, left side from posterior. J. Mandible, left side, apical view on gnathal edge. K. Mandible, left side, apical view on molar process." figureDoi="http://doi.org/10.5281/zenodo.3827401" httpUri="https://zenodo.org/record/3827401/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
H–K). Well developed with a gnathal edge consisting of three parts, a molar, a lacinia mobilis, and an incisor; only mandible of left side has been examined in detail. Left molar process occupies approximately ⅔ of the gnathal edge; in apical view the molar is crescent-shaped, composed of a dense row of multi-tipped setules bending towards each other forming a large cavity in the center of the molar; on the functionally anterior side of the molar are clusters of short setules and one cluster of longer. Left incisor broad, flattened, distal margin subdivided into four cusps of subequal morphology (three are pointed, one spade-shaped). The lacinia mobilis is positioned between the incisor and the molar as an extension of the latter, subequal in length to incisor but more narrow terminating in three cusps.
</paragraph>
<paragraph id="8B8EFC32FFC96450FF088999FD18AE6B" blockId="10.[189,1399,931,1603]" pageId="10" pageNumber="11">
FIRST MAXILLA (
<figureCitation id="130AE0B7FFC96450FEDA899EFE52AB95" box="[367,461,931,957]" captionStart="Fig" captionStartId="11.[189,232,1893,1919]" captionTargetBox="[231,1357,263,1858]" captionTargetId="figure@11.[231,1357,263,1858]" captionTargetPageId="11" captionText="Fig. 4. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), light microscopy. A. Antenna 1, left side from anterior. B. Antenna 2, left side from anterior. C. Maxilla 1, left side from anterior. D. Maxilla 2, left side from anterior. E. Maxilliped, left side from anterior. F. Caudal rami and posterior part of trunk. G. Caudal ramus, left side seen from median." figureDoi="http://doi.org/10.5281/zenodo.3827399" httpUri="https://zenodo.org/record/3827399/files/figure.png" pageId="10" pageNumber="11">Figs 4C</figureCitation>
, 
<figureCitation id="130AE0B7FFC96450FE6D899EFE76AB96" box="[472,489,931,958]" captionStart="Fig" captionStartId="13.[189,232,1789,1815]" captionTargetBox="[276,1311,262,1755]" captionTargetId="figure@13.[276,1311,261,1755]" captionTargetPageId="13" captionText="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." figureDoi="http://doi.org/10.5281/zenodo.3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="10" pageNumber="11">6</figureCitation>
). 7-segmented, uniramous, robust. Point of flexion between segments 4 and 5. Segment 1 (
<figureCitation id="130AE0B7FFC96450FEE789FAFE04ABCA" box="[338,411,967,994]" captionStart="Fig" captionStartId="13.[189,232,1789,1815]" captionTargetBox="[276,1311,262,1755]" captionTargetId="figure@13.[276,1311,261,1755]" captionTargetPageId="13" captionText="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." figureDoi="http://doi.org/10.5281/zenodo.3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
A–B) relatively short and drawn out medially in a flattened endite with 7 robust setae, posteriormost seta largest, anteriormost serrated; subterminally with row of 4 setules. Segment 2 (
<figureCitation id="130AE0B7FFC96450FF738E32FED8AC01" box="[198,327,1039,1065]" captionStart="Fig" captionStartId="13.[189,232,1789,1815]" captionTargetBox="[276,1311,262,1755]" captionTargetId="figure@13.[276,1311,261,1755]" captionTargetPageId="13" captionText="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." figureDoi="http://doi.org/10.5281/zenodo.3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="10" pageNumber="11">Fig. 6A, D</figureCitation>
) narrow, of subtriangular shape, with elongate, plate-like endite articulated to the segment and directed obliquely medially/anteriorly; terminal margin with one row of 7 robust spines and another more disorganized row of about 17 moderately long or smaller setae; at the posterior margin is a cluster of 4 long setae; at the anterior margin is a row of 5 long setae. Segment 3 (
<figureCitation id="130AE0B7FFC96450FBBB8E47FBCEACBD" box="[1038,1105,1146,1173]" captionStart="Fig" captionStartId="13.[189,232,1789,1815]" captionTargetBox="[276,1311,262,1755]" captionTargetId="figure@13.[276,1311,261,1755]" captionTargetPageId="13" captionText="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." figureDoi="http://doi.org/10.5281/zenodo.3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
A–B) is trapezoid shaped with a well-developed conical medial endite bearing 2 robust, broad-based cone-shaped setae, each with tiny papilla-like projections on distomedial surface (
<figureCitation id="130AE0B7FFC96450FC868EFFFC14ACF4" box="[819,907,1218,1244]" captionStart="Fig" captionStartId="13.[189,232,1789,1815]" captionTargetBox="[276,1311,262,1755]" captionTargetId="figure@13.[276,1311,261,1755]" captionTargetPageId="13" captionText="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." figureDoi="http://doi.org/10.5281/zenodo.3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="10" pageNumber="11">Fig. 6F</figureCitation>
, G–I); anteriorly on endite is a row of 5 long, slender setae, posteriorly is 5 setae of different lengths. Segment 4 (
<figureCitation id="130AE0B7FFC96450FB948EDBFBE2AD29" box="[1057,1149,1254,1281]" captionStart="Fig" captionStartId="13.[189,232,1789,1815]" captionTargetBox="[276,1311,262,1755]" captionTargetId="figure@13.[276,1311,261,1755]" captionTargetPageId="13" captionText="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." figureDoi="http://doi.org/10.5281/zenodo.3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
) as large as the three preceding segments, bearing a well-developed conical median endite with 2 large cone-shaped setae as on segment 3, also with tiny papilla-like projections on distomedial surface; anteriorly on endite is a double-row of 9 setae, posteriorly on endite is a cluster of 9 setae, 4 relatively long and 5 relatively short. Segment 5 (
<figureCitation id="130AE0B7FFC96450FEFE8F48FE37ADB8" box="[331,424,1397,1424]" captionStart="Fig" captionStartId="13.[189,232,1789,1815]" captionTargetBox="[276,1311,262,1755]" captionTargetId="figure@13.[276,1311,261,1755]" captionTargetPageId="13" captionText="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." figureDoi="http://doi.org/10.5281/zenodo.3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="10" pageNumber="11">Fig. 6A</figureCitation>
) about the same length as segment 4 but more slender; cluster of about 12 slender setae in medio-distal corner of segment. Segment 6 (
<figureCitation id="130AE0B7FFC96450FC9A8FA4FC2FAD9B" box="[815,944,1433,1459]" captionStart="Fig" captionStartId="13.[189,232,1789,1815]" captionTargetBox="[276,1311,262,1755]" captionTargetId="figure@13.[276,1311,261,1755]" captionTargetPageId="13" captionText="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." figureDoi="http://doi.org/10.5281/zenodo.3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="10" pageNumber="11">Fig. 6A, C</figureCitation>
) very short not clearly separated from segment 5 dorsally, with two rows of setae anteriorly and posteriorly, about 8 setae in each. Segment 7 (
<figureCitation id="130AE0B7FFC96450FF698FDCFEBDADD4" box="[220,290,1505,1532]" captionStart="Fig" captionStartId="13.[189,232,1789,1815]" captionTargetBox="[276,1311,262,1755]" captionTargetId="figure@13.[276,1311,261,1755]" captionTargetPageId="13" captionText="Fig. 6. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. A. Complete mx1 from anterior. B. Median view. C. Close-up of median structures (e.g., endites) and distal claw from anterior. D. Distal setation of plate-like endite of segment 2 seen from ‘below’. E. Distal setation of endite of segment 1 seen from ‘below’. F. Close-up of two robust, broad-based cone-shaped setae of segment 3. G. Close-up of papillae of cone-shaped seta. H. Close-up of median structures (e.g., endites) and distal claw from posterior. I. Fang-like claw with visible subterminal duct opening." figureDoi="http://doi.org/10.5281/zenodo.3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="10" pageNumber="11">Fig. 6</figureCitation>
A–C) short, not clearly separated from segment 6 dorsally, terminating in stout fang-like claw with visible subterminal duct opening, more than 3 times the length of segment 6; a cluster of about 6 long simple setae basally at segment 7.
</paragraph>
<paragraph id="8B8EFC32FFC96454FF088C57FE2CA9FE" blockId="10.[189,1399,1642,2027]" lastBlockId="14.[189,1399,267,470]" lastPageId="14" lastPageNumber="15" pageId="10" pageNumber="11">
SECOND MAXILLA (
<figureCitation id="130AE0B7FFC96450FE3C8C57FE79AEAD" box="[393,486,1642,1669]" captionStart="Fig" captionStartId="11.[189,232,1893,1919]" captionTargetBox="[231,1357,263,1858]" captionTargetId="figure@11.[231,1357,263,1858]" captionTargetPageId="11" captionText="Fig. 4. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), light microscopy. A. Antenna 1, left side from anterior. B. Antenna 2, left side from anterior. C. Maxilla 1, left side from anterior. D. Maxilla 2, left side from anterior. E. Maxilliped, left side from anterior. F. Caudal rami and posterior part of trunk. G. Caudal ramus, left side seen from median." figureDoi="http://doi.org/10.5281/zenodo.3827399" httpUri="https://zenodo.org/record/3827399/files/figure.png" pageId="10" pageNumber="11">Figs 4D</figureCitation>
, 
<figureCitation id="130AE0B7FFC96450FE418C56FD9AAEAD" box="[500,517,1643,1669]" captionStart="Fig" captionStartId="15.[189,232,1860,1886]" captionTargetBox="[236,1353,268,1818]" captionTargetId="figure@15.[234,1353,268,1818]" captionTargetPageId="15" captionText="Fig. 7. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. A. Complete mx2 from anterior. B. Close-up of terminal claw-complex from anterior (segment 7). C. Close-up of terminal claw-complex from lateral (segment 7). D. Median side of segments 1 and 2 with endites (e1–3) from anterior. E. Endites of segment 1 (e1–3) from median. F. Complete mx2 from median." figureDoi="http://doi.org/10.5281/zenodo.3827405" httpUri="https://zenodo.org/record/3827405/files/figure.png" pageId="10" pageNumber="11">7</figureCitation>
). 7-segmented, uniramous, longer and more slender than maxilla 1; point of flexion between segments 3 and 4; segments 4–7 decreasing in size distally. Segment 1 with 3 digitiform endites (
<figureCitation id="130AE0B7FFC96450FE958C8FFEE1AEE4" box="[288,382,1714,1740]" captionStart="Fig" captionStartId="15.[189,232,1860,1886]" captionTargetBox="[236,1353,268,1818]" captionTargetId="figure@15.[234,1353,268,1818]" captionTargetPageId="15" captionText="Fig. 7. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. A. Complete mx2 from anterior. B. Close-up of terminal claw-complex from anterior (segment 7). C. Close-up of terminal claw-complex from lateral (segment 7). D. Median side of segments 1 and 2 with endites (e1–3) from anterior. E. Endites of segment 1 (e1–3) from median. F. Complete mx2 from median." figureDoi="http://doi.org/10.5281/zenodo.3827405" httpUri="https://zenodo.org/record/3827405/files/figure.png" pageId="10" pageNumber="11">Fig. 7A</figureCitation>
, D–E) increasing in size anteriorly; all endites with 1 long apical spine-like seta, a row of short, spine-like setae medially (least on endite 1, most in endite 3) and a row of long setae laterally (least on endite 1, most in endite 3). Segment 2 (
<figureCitation id="130AE0B7FFC96450FD5B8CC7FCF8AF3C" box="[750,871,1786,1812]" captionStart="Fig" captionStartId="15.[189,232,1860,1886]" captionTargetBox="[236,1353,268,1818]" captionTargetId="figure@15.[234,1353,268,1818]" captionTargetPageId="15" captionText="Fig. 7. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. A. Complete mx2 from anterior. B. Close-up of terminal claw-complex from anterior (segment 7). C. Close-up of terminal claw-complex from lateral (segment 7). D. Median side of segments 1 and 2 with endites (e1–3) from anterior. E. Endites of segment 1 (e1–3) from median. F. Complete mx2 from median." figureDoi="http://doi.org/10.5281/zenodo.3827405" httpUri="https://zenodo.org/record/3827405/files/figure.png" pageId="10" pageNumber="11">Fig. 7A, F</figureCitation>
) short, wedge-shaped, ‘squeezed in’ between segments 1 and 3 with the broad end of the ‘wedge’ facing medio-anteriorly, medially bearing 1 short, stout, curved medial spine, with a short row of about 4 median setae and short row or cluster of about 5 short to moderately long slender setae at medio-distal corner. Segment 3 (
<figureCitation id="130AE0B7FFC96450FBAF8D58FB0AAFA8" box="[1050,1173,1893,1920]" captionStart="Fig" captionStartId="15.[189,232,1860,1886]" captionTargetBox="[236,1353,268,1818]" captionTargetId="figure@15.[234,1353,268,1818]" captionTargetPageId="15" captionText="Fig. 7. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. A. Complete mx2 from anterior. B. Close-up of terminal claw-complex from anterior (segment 7). C. Close-up of terminal claw-complex from lateral (segment 7). D. Median side of segments 1 and 2 with endites (e1–3) from anterior. E. Endites of segment 1 (e1–3) from median. F. Complete mx2 from median." figureDoi="http://doi.org/10.5281/zenodo.3827405" httpUri="https://zenodo.org/record/3827405/files/figure.png" pageId="10" pageNumber="11">Fig. 7A, F</figureCitation>
) elongate, rounded medially with row of about 11 long setae and 8 shorter setae. Segment 4 (
<figureCitation id="130AE0B7FFC96450FBAA8DB4FBE4AF8B" box="[1055,1147,1929,1955]" captionStart="Fig" captionStartId="15.[189,232,1860,1886]" captionTargetBox="[236,1353,268,1818]" captionTargetId="figure@15.[234,1353,268,1818]" captionTargetPageId="15" captionText="Fig. 7. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. A. Complete mx2 from anterior. B. Close-up of terminal claw-complex from anterior (segment 7). C. Close-up of terminal claw-complex from lateral (segment 7). D. Median side of segments 1 and 2 with endites (e1–3) from anterior. E. Endites of segment 1 (e1–3) from median. F. Complete mx2 from median." figureDoi="http://doi.org/10.5281/zenodo.3827405" httpUri="https://zenodo.org/record/3827405/files/figure.png" pageId="10" pageNumber="11">Fig. 7A</figureCitation>
) slightly shorter than segment 3, with small distomedial lobe terminating in a cluster of about 10 short to long, thin setae; distolateral corner with 4 slender setae of varying lengths. Segment 5 (
<figureCitation id="130AE0B7FFC96450FC4B8DECFBC4AFC3" box="[1022,1115,2001,2027]" captionStart="Fig" captionStartId="15.[189,232,1860,1886]" captionTargetBox="[236,1353,268,1818]" captionTargetId="figure@15.[234,1353,268,1818]" captionTargetPageId="15" captionText="Fig. 7. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. A. Complete mx2 from anterior. B. Close-up of terminal claw-complex from anterior (segment 7). C. Close-up of terminal claw-complex from lateral (segment 7). D. Median side of segments 1 and 2 with endites (e1–3) from anterior. E. Endites of segment 1 (e1–3) from median. F. Complete mx2 from median." figureDoi="http://doi.org/10.5281/zenodo.3827405" httpUri="https://zenodo.org/record/3827405/files/figure.png" pageId="10" pageNumber="11">Fig. 7A</figureCitation>
) shorter than segment 4 with approximate same configuration of setae. Segment 6 (
<figureCitation id="130AE0B7FFCD6454FCCD8B36FC6AA90E" box="[888,1013,267,294]" captionStart="Fig" captionStartId="15.[189,232,1860,1886]" captionTargetBox="[236,1353,268,1818]" captionTargetId="figure@15.[234,1353,268,1818]" captionTargetPageId="15" captionText="Fig. 7. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. A. Complete mx2 from anterior. B. Close-up of terminal claw-complex from anterior (segment 7). C. Close-up of terminal claw-complex from lateral (segment 7). D. Median side of segments 1 and 2 with endites (e1–3) from anterior. E. Endites of segment 1 (e1–3) from median. F. Complete mx2 from median." figureDoi="http://doi.org/10.5281/zenodo.3827405" httpUri="https://zenodo.org/record/3827405/files/figure.png" pageId="14" pageNumber="15">Fig. 7A, F</figureCitation>
) bearing a large cluster of about 15 setae at distomedial corner and a pair of clusters at the disto-lateral corners (about 6 setae in each). Segment 7 (
<figureCitation id="130AE0B7FFCD6454FEE58B6FFE4BA944" box="[336,468,338,364]" captionStart="Fig" captionStartId="15.[189,232,1860,1886]" captionTargetBox="[236,1353,268,1818]" captionTargetId="figure@15.[234,1353,268,1818]" captionTargetPageId="15" captionText="Fig. 7. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. A. Complete mx2 from anterior. B. Close-up of terminal claw-complex from anterior (segment 7). C. Close-up of terminal claw-complex from lateral (segment 7). D. Median side of segments 1 and 2 with endites (e1–3) from anterior. E. Endites of segment 1 (e1–3) from median. F. Complete mx2 from median." figureDoi="http://doi.org/10.5281/zenodo.3827405" httpUri="https://zenodo.org/record/3827405/files/figure.png" pageId="14" pageNumber="15">Fig. 7A, C</figureCitation>
) very short, about half the length of segment 6, with terminal claw complex consisting of fused arch of at least 17 denticles and 1 long, stout anterior spine; 1 short, stout anterior spine subterminal to fused arch; opposable thumb-like pad bearing many (at least 40) setae which fan under claw complex.
</paragraph>
<caption id="DF4EACBAFFC86451FF088D58FCEBAFC1" ID-DOI="http://doi.org/10.5281/zenodo.3827399" ID-Zenodo-Dep="3827399" httpUri="https://zenodo.org/record/3827399/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,232,1893,1919]" targetBox="[231,1357,263,1858]" targetPageId="11">
<paragraph id="8B8EFC32FFC86451FF088D58FCEBAFC1" blockId="11.[189,1399,1893,2026]" pageId="11" pageNumber="12">
<emphasis id="B9452020FFC86451FF088D58FE97AF57" bold="true" box="[189,264,1893,1919]" pageId="11" pageNumber="12">Fig. 4.</emphasis>
<taxonomicName id="4C3187B1FFC86451FEBB8D5BFDABAFA8" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[270,564,1894,1920]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFC86451FEBB8D5BFDABAFA8" box="[270,564,1894,1920]" italics="true" pageId="11" pageNumber="12">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFC86451FD8C8D5BFD0BAFA8" box="[569,660,1894,1920]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (ZMUC-CRU-4791), light microscopy. 
<emphasis id="B9452020FFC86451FB6D8D58FB72AF57" bold="true" box="[1240,1261,1893,1919]" pageId="11" pageNumber="12">A</emphasis>
. Antenna 1, left side from anterior. 
<emphasis id="B9452020FFC86451FE698DB4FE6FAF8B" bold="true" box="[476,496,1929,1955]" pageId="11" pageNumber="12">B</emphasis>
. Antenna 2, left side from anterior. 
<emphasis id="B9452020FFC86451FC078DB4FC58AF8B" bold="true" box="[946,967,1929,1955]" pageId="11" pageNumber="12">C</emphasis>
. Maxilla 1, left side from anterior. 
<emphasis id="B9452020FFC86451FF088D91FF4DAFEE" bold="true" box="[189,210,1964,1990]" pageId="11" pageNumber="12">D</emphasis>
. Maxilla 2, left side from anterior. 
<emphasis id="B9452020FFC86451FDDE8D91FDE0AFEE" bold="true" box="[619,639,1964,1990]" pageId="11" pageNumber="12">E</emphasis>
. Maxilliped, left side from anterior. 
<emphasis id="B9452020FFC86451FB928D91FBA6AFEE" bold="true" box="[1063,1081,1964,1990]" pageId="11" pageNumber="12">F</emphasis>
. Caudal rami and posterior part of trunk. 
<emphasis id="B9452020FFC86451FEEB8DF2FEEAAFC1" bold="true" box="[350,373,1999,2025]" pageId="11" pageNumber="12">G</emphasis>
. Caudal ramus, left side seen from median.
</paragraph>
</caption>
<caption id="DF4EACBAFFCF6456FF088CC3FCB3AFC4" ID-DOI="http://doi.org/10.5281/zenodo.3827401" ID-Zenodo-Dep="3827401" httpUri="https://zenodo.org/record/3827401/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,232,1790,1816]" targetBox="[229,1358,264,1753]" targetPageId="12">
<paragraph id="8B8EFC32FFCF6456FF088CC3FCB3AFC4" blockId="12.[189,1399,1790,2028]" pageId="12" pageNumber="13">
<emphasis id="B9452020FFCF6456FF088CC3FE8CAF31" bold="true" box="[189,275,1790,1817]" pageId="12" pageNumber="13">Fig. 5.</emphasis>
<taxonomicName id="4C3187B1FFCF6456FE978CC3FDCCAF30" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[290,595,1790,1816]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFCF6456FE978CC3FDCCAF30" box="[290,595,1790,1816]" italics="true" pageId="12" pageNumber="13">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFCF6456FDD78CC2FD57AF31" box="[610,712,1791,1817]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ZMUC-CRU-4792). Cephalon and cephalic structures. Scanning electron microscopy. 
<emphasis id="B9452020FFCF6456FD6B8D1CFD6CAF13" bold="true" box="[734,755,1825,1851]" pageId="12" pageNumber="13">A</emphasis>
. Cephalon with cephalic shield and appendages. 
<emphasis id="B9452020FFCF6456FF088D78FF4EAF77" bold="true" box="[189,209,1861,1887]" pageId="12" pageNumber="13">B</emphasis>
. Antenna 1, right side, proximal parts of dorsal and ventral branches. 
<emphasis id="B9452020FFCF6456FBAF8D78FBCCAF77" bold="true" box="[1050,1107,1861,1887]" pageId="12" pageNumber="13">C–D</emphasis>
. Unidentified suctorians attached at the proximo-ventral parts of antenna 1. 
<emphasis id="B9452020FFCF6456FCAF8D55FCB1AFAA" bold="true" box="[794,814,1896,1922]" pageId="12" pageNumber="13">E</emphasis>
. Labrum, frontal filaments, and parts of antennae 1 and 2, right side. 
<emphasis id="B9452020FFCF6456FE1C8DB1FE24AF8E" bold="true" box="[425,443,1932,1958]" pageId="12" pageNumber="13">F</emphasis>
. Frontal filaments. 
<emphasis id="B9452020FFCF6456FD138DB6FD22AF8D" bold="true" box="[678,701,1931,1957]" pageId="12" pageNumber="13">G</emphasis>
. Antenna 2, left side from anterior. 
<emphasis id="B9452020FFCF6456FBDB8DB1FB1AAF8E" bold="true" box="[1134,1157,1932,1958]" pageId="12" pageNumber="13">H</emphasis>
. Mandible, left side from anterior. 
<emphasis id="B9452020FFCF6456FED38D92FEEEAFE1" bold="true" box="[358,369,1967,1993]" pageId="12" pageNumber="13">I</emphasis>
. Mandible, left side from posterior. 
<emphasis id="B9452020FFCF6456FCA98D92FCB4AFE1" bold="true" box="[796,811,1967,1993]" pageId="12" pageNumber="13">J</emphasis>
. Mandible, left side, apical view on gnathal edge. 
<emphasis id="B9452020FFCF6456FF088DEFFF4BAFC4" bold="true" box="[189,212,2002,2028]" pageId="12" pageNumber="13">K</emphasis>
. Mandible, left side, apical view on molar process.
</paragraph>
</caption>
<caption id="DF4EACBAFFCE6457FF088CC0FDDAAFC2" ID-DOI="http://doi.org/10.5281/zenodo.3827403" ID-Zenodo-Dep="3827403" httpUri="https://zenodo.org/record/3827403/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,232,1789,1815]" targetBox="[276,1311,262,1755]" targetPageId="13">
<paragraph id="8B8EFC32FFCE6457FF088CC0FDDAAFC2" blockId="13.[189,1399,1789,2026]" pageId="13" pageNumber="14">
<emphasis id="B9452020FFCE6457FF088CC0FE91AF3F" bold="true" box="[189,270,1789,1815]" pageId="13" pageNumber="14">Fig. 6.</emphasis>
<taxonomicName id="4C3187B1FFCE6457FEAC8CC0FDDAAF3F" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[281,581,1789,1815]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFCE6457FEAC8CC0FDDAAF3F" box="[281,581,1789,1815]" italics="true" pageId="13" pageNumber="14">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFCE6457FDE58CC3FD2EAF30" box="[592,689,1790,1816]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ZMUC-CRU-4792), maxilla 1 (mx1), left side, scanning electron microscopy. 
<emphasis id="B9452020FFCE6457FD898D1DFDCEAF12" bold="true" box="[572,593,1824,1850]" pageId="13" pageNumber="14">A</emphasis>
. Complete mx1 from anterior. 
<emphasis id="B9452020FFCE6457FC6A8D1DFC6CAF12" bold="true" box="[991,1011,1824,1850]" pageId="13" pageNumber="14">B</emphasis>
. Median view. 
<emphasis id="B9452020FFCE6457FB098D1DFB4EAF12" bold="true" box="[1212,1233,1824,1850]" pageId="13" pageNumber="14">C</emphasis>
. Close-up of median structures (e.g., endites) and distal claw from anterior. 
<emphasis id="B9452020FFCE6457FC148D79FC29AF76" bold="true" box="[929,950,1860,1886]" pageId="13" pageNumber="14">D</emphasis>
. Distal setation of plate-like endite of segment 2 seen from ‘below’. 
<emphasis id="B9452020FFCE6457FD968D5AFDA8AFA9" bold="true" box="[547,567,1895,1921]" pageId="13" pageNumber="14">E</emphasis>
. Distal setation of endite of segment 1 seen from ‘below’. 
<emphasis id="B9452020FFCE6457FB598D5AFB61AFA9" bold="true" box="[1260,1278,1895,1921]" pageId="13" pageNumber="14">F</emphasis>
. Close-up of two robust, broad-based cone-shaped setae of segment 3. 
<emphasis id="B9452020FFCE6457FCC08DB7FC13AF8C" bold="true" box="[885,908,1930,1956]" pageId="13" pageNumber="14">G</emphasis>
. Close-up of papillae of cone-shaped seta. 
<emphasis id="B9452020FFCE6457FF088D90FF4BAFEF" bold="true" box="[189,212,1965,1991]" pageId="13" pageNumber="14">H</emphasis>
. Close-up of median structures (e.g., endites) and distal claw from posterior. 
<emphasis id="B9452020FFCE6457FBDB8D90FBE6AFEF" bold="true" box="[1134,1145,1965,1991]" pageId="13" pageNumber="14">I</emphasis>
. Fang-like claw with visible subterminal duct opening.
</paragraph>
</caption>
<paragraph id="8B8EFC32FFCD6454FF088BC2FD74AC67" blockId="14.[189,1399,510,1103]" pageId="14" pageNumber="15">
MAXILLIPED (
<figureCitation id="130AE0B7FFCD6454FEE28BC3FE29AA31" box="[343,438,510,537]" captionStart="Fig" captionStartId="11.[189,232,1893,1919]" captionTargetBox="[231,1357,263,1858]" captionTargetId="figure@11.[231,1357,263,1858]" captionTargetPageId="11" captionText="Fig. 4. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), light microscopy. A. Antenna 1, left side from anterior. B. Antenna 2, left side from anterior. C. Maxilla 1, left side from anterior. D. Maxilla 2, left side from anterior. E. Maxilliped, left side from anterior. F. Caudal rami and posterior part of trunk. G. Caudal ramus, left side seen from median." figureDoi="http://doi.org/10.5281/zenodo.3827399" httpUri="https://zenodo.org/record/3827399/files/figure.png" pageId="14" pageNumber="15">Figs 4E</figureCitation>
, 
<figureCitation id="130AE0B7FFCD6454FE7D8BC3FE47AA30" box="[456,472,510,536]" captionStart="Fig" captionStartId="16.[189,232,1792,1818]" captionTargetBox="[276,1313,267,1763]" captionTargetId="figure@16.[274,1313,265,1763]" captionTargetPageId="16" captionText="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." figureDoi="http://doi.org/10.5281/zenodo.3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="14" pageNumber="15">8</figureCitation>
). 8-segmented, generally similar in shape but longer and more slender than maxilla 2, with one more segment beyond point of flexure. Segment 1 (
<figureCitation id="130AE0B7FFCD6454FBB1881FFB1CAA14" box="[1028,1155,546,572]" captionStart="Fig" captionStartId="16.[189,232,1792,1818]" captionTargetBox="[276,1313,267,1763]" captionTargetId="figure@16.[274,1313,265,1763]" captionTargetPageId="16" captionText="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." figureDoi="http://doi.org/10.5281/zenodo.3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="14" pageNumber="15">Fig. 8A, C</figureCitation>
) is short with 5 long and 2 short setae along median margin and about 3–4 small setae in a row more proximally. Segment 2 is large and undivided in anterior view (
<figureCitation id="130AE0B7FFCD6454FDD98855FD57AAAB" box="[620,712,616,643]" captionStart="Fig" captionStartId="16.[189,232,1792,1818]" captionTargetBox="[276,1313,267,1763]" captionTargetId="figure@16.[274,1313,265,1763]" captionTargetPageId="16" captionText="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." figureDoi="http://doi.org/10.5281/zenodo.3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="14" pageNumber="15">Fig. 8A</figureCitation>
), but in posterior view it is subdivided by an oblique furrow giving the appearance of two separate triangular segments (
<figureCitation id="130AE0B7FFCD6454FCC488B1FC52AA8E" box="[881,973,652,678]" captionStart="Fig" captionStartId="16.[189,232,1792,1818]" captionTargetBox="[276,1313,267,1763]" captionTargetId="figure@16.[274,1313,265,1763]" captionTargetPageId="16" captionText="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." figureDoi="http://doi.org/10.5281/zenodo.3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="14" pageNumber="15">Fig. 8C</figureCitation>
); at the medio-distal margin is a row of 12 setae of varying length. Segment 3 as long as the combined length of segments 1–2; with round medial margin with about 15 slender setae of varying length in a dense row in the middle of the segment. Segment 4 (
<figureCitation id="130AE0B7FFCD6454FEF888CBFE50AB38" box="[333,463,758,784]" captionStart="Fig" captionStartId="16.[189,232,1792,1818]" captionTargetBox="[276,1313,267,1763]" captionTargetId="figure@16.[274,1313,265,1763]" captionTargetPageId="16" captionText="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." figureDoi="http://doi.org/10.5281/zenodo.3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="14" pageNumber="15">Fig. 8A, C</figureCitation>
) the same length as segment 3, subtriangular in shape, with a cluster of about 15 setae of varying size at disto-medial corner, and a small cluster of about 4 short setae in latero-distal corner. Segment 5 (
<figureCitation id="130AE0B7FFCD6454FE118900FDBAAB7F" box="[420,549,829,855]" captionStart="Fig" captionStartId="16.[189,232,1792,1818]" captionTargetBox="[276,1313,267,1763]" captionTargetId="figure@16.[274,1313,265,1763]" captionTargetPageId="16" captionText="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." figureDoi="http://doi.org/10.5281/zenodo.3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="14" pageNumber="15">Fig. 8A, C</figureCitation>
) approximately the same size as segment 4 but more rectangular; about 12 setae in a cluster at disto-medial corner, 
<quantity id="4CC951D7FFCD6454FD00895DFD7DAB53" box="[693,738,864,891]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="14" pageNumber="15" unit="in" value="5.0">5 in</quantity>
latero-distal corner. Segment 6 (
<figureCitation id="130AE0B7FFCD6454FBEB895CFB44AB53" box="[1118,1243,865,891]" captionStart="Fig" captionStartId="16.[189,232,1792,1818]" captionTargetBox="[276,1313,267,1763]" captionTargetId="figure@16.[274,1313,265,1763]" captionTargetPageId="16" captionText="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." figureDoi="http://doi.org/10.5281/zenodo.3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="14" pageNumber="15">Fig. 8A, C</figureCitation>
) very similar to segment 5 with respect to setation, but only slightly more than half as long. Segment 7 (
<figureCitation id="130AE0B7FFCD6454FB5B89B9FAF1ABB6" box="[1262,1390,900,926]" captionStart="Fig" captionStartId="16.[189,232,1792,1818]" captionTargetBox="[276,1313,267,1763]" captionTargetId="figure@16.[274,1313,265,1763]" captionTargetPageId="16" captionText="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." figureDoi="http://doi.org/10.5281/zenodo.3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="14" pageNumber="15">Fig. 8A, C</figureCitation>
) about the same length as segment 6 with rows of short setae along distal margin of segment: two rows at the disto-lateral corners of the anterior and posterior side of segment, each with 4–6 setae, and two rows at the disto-medial corners with about 10 (anterior side) or 5 (posterior side) setae. Segment 8 (
<figureCitation id="130AE0B7FFCD6454FF738E2FFEDCAC04" box="[198,323,1042,1068]" captionStart="Fig" captionStartId="16.[189,232,1792,1818]" captionTargetBox="[276,1313,267,1763]" captionTargetId="figure@16.[274,1313,265,1763]" captionTargetPageId="16" captionText="Fig. 8. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. A. Complete mxp from anterior. B. Close-up of terminal claw-complex from anterior (segment 8). C. Complete mxp from posterior. D. Close-up of terminal claw-complex from posterior (segment 8). E. Close-up of terminal claw-complex from lateral (segment 8). F. Close-up of small cluster of setae along distal margin of segment 7. G. Close-up of pore at close to latero-distal margin of segment 7. H. Close-up of cluster of setae at latero-distal corner of segment 4. I. Close-up of cluster of setae at latero-distal corner of segment 5." figureDoi="http://doi.org/10.5281/zenodo.3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="14" pageNumber="15">Fig. 8B, D</figureCitation>
) similar to claw complex on maxilla 2, with stout anterior spine subterminal to arched claw complex and with opposable setae-bearing pad.
</paragraph>
<paragraph id="8B8EFC32FFCD6454FF088E45FC98AD4E" blockId="14.[189,1399,1143,1382]" pageId="14" pageNumber="15">
TRUNK. Limbs biramous (
<figureCitation id="130AE0B7FFCD6454FDB18E4AFD18ACB9" box="[516,647,1143,1169]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="17.[189,232,1929,1955]" captionStartId-1="18.[189,232,1824,1850]" captionTargetBox-0="[189,1398,678,1893]" captionTargetBox-1="[190,1398,436,1789]" captionTargetId-0="figure@17.[189,1398,678,1893]" captionTargetId-1="figure@18.[189,1398,435,1790]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Fig. 9. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), trunk limbs, left side from anterior, light microscopy. A–J. Selected trunk segments (number specified on figure). All to the same scale." captionText-1="Fig. 10. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792), cephalon, body segments, and trunk limbs, scanning electron microscopy. A. Cephalon, cephalic shield, and head appendages from lateral. B. Trunk segments 5–10. C. Trunk limb 2, right side. D. Close-up of serrated setae in latero-distal corner of exopod segment 1. E. Close-up of serrated setae in latero-distal corner of exopod segment 2. F–G. Two unidentified suctorians attached to exopod of trunk limb 3. H–I. Posterior body segments, trunk limbs, and caudal rami from ventral and lateral." figureDoi-0="http://doi.org/10.5281/zenodo.3827409" figureDoi-1="http://doi.org/10.5281/zenodo.3827411" httpUri-0="https://zenodo.org/record/3827409/files/figure.png" httpUri-1="https://zenodo.org/record/3827411/files/figure.png" pageId="14" pageNumber="15">Figs 9–10</figureCitation>
), natatory, with large protopod, 4-segmented endopod and 3-segmented exopod; long plumose setae along margins; size of limbs varies along body, increasing in size from trunk limb 1 to approximately trunk limb 7, decreasing in size approximately from trunk limb 29 onwards; trunk limbs 37 and 38 are significantly reduced in size (
<figureCitation id="130AE0B7FFCD6454FC6B8EDCFBA7ACD3" box="[990,1080,1249,1275]" captionStart="Fig" captionStartId="11.[189,232,1893,1919]" captionTargetBox="[231,1357,263,1858]" captionTargetId="figure@11.[231,1357,263,1858]" captionTargetPageId="11" captionText="Fig. 4. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), light microscopy. A. Antenna 1, left side from anterior. B. Antenna 2, left side from anterior. C. Maxilla 1, left side from anterior. D. Maxilla 2, left side from anterior. E. Maxilliped, left side from anterior. F. Caudal rami and posterior part of trunk. G. Caudal ramus, left side seen from median." figureDoi="http://doi.org/10.5281/zenodo.3827399" httpUri="https://zenodo.org/record/3827399/files/figure.png" pageId="14" pageNumber="15">Fig. 4F</figureCitation>
); in the most anterior trunk limbs the exopod is longer than the endopod, in the middle range of trunk limbs the rami are sub-equal in size, around trunk limb 30 onwards the endopod is clearly larger, while in the last two reduced trunk limbs (37 and 38) the exopods are largest (
<figureCitation id="130AE0B7FFCD6454FD048F71FD66AD4E" box="[689,761,1356,1382]" captionStart="Fig" captionStartId="17.[189,232,1929,1955]" captionTargetBox="[189,1398,678,1893]" captionTargetId="figure@17.[189,1398,678,1893]" captionTargetPageId="17" captionText="Fig. 9. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), trunk limbs, left side from anterior, light microscopy. A–J. Selected trunk segments (number specified on figure). All to the same scale." figureDoi="http://doi.org/10.5281/zenodo.3827409" httpUri="https://zenodo.org/record/3827409/files/figure.png" pageId="14" pageNumber="15">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B8EFC32FFCD6454FF088FB0FEF4ADE3" blockId="14.[189,1398,1421,1483]" pageId="14" pageNumber="15">
CAUDAL RAMI. Very slender, about 4–5 times length of anal segment with small terminal setae (
<figureCitation id="130AE0B7FFCD6454FAF78FB0FF54ADE3" captionStart="Fig" captionStartId="11.[189,232,1893,1919]" captionTargetBox="[231,1357,263,1858]" captionTargetId="figure@11.[231,1357,263,1858]" captionTargetPageId="11" captionText="Fig. 4. Xibalbanus cozumelensis sp. nov., holotype (ZMUC-CRU-4791), light microscopy. A. Antenna 1, left side from anterior. B. Antenna 2, left side from anterior. C. Maxilla 1, left side from anterior. D. Maxilla 2, left side from anterior. E. Maxilliped, left side from anterior. F. Caudal rami and posterior part of trunk. G. Caudal ramus, left side seen from median." figureDoi="http://doi.org/10.5281/zenodo.3827399" httpUri="https://zenodo.org/record/3827399/files/figure.png" pageId="14" pageNumber="15">Figs 4</figureCitation>
F–G, 10H–I).
</paragraph>
</subSubSection>
<subSubSection id="C32BAFB9FFCD6449FF088FCEFCF4AA4A" lastPageId="19" lastPageNumber="20" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B8EFC32FFCD6454FF088FCEFEB0AE25" blockId="14.[189,303,1523,1549]" box="[189,303,1523,1549]" pageId="14" pageNumber="15">
<heading id="D0C64B5EFFCD6454FF088FCEFEB0AE25" bold="true" box="[189,303,1523,1549]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B9452020FFCD6454FF088FCEFEB0AE25" bold="true" box="[189,303,1523,1549]" pageId="14" pageNumber="15">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8EFC32FFCD6454FF088C1CFD20AE77" blockId="14.[189,1399,1569,1631]" pageId="14" pageNumber="15">
Unidentified suctorians of varying shapes were found attached at the proximo-ventral parts of antennae 1 and at the trunk limbs (
<figureCitation id="130AE0B7FFCD6454FE798C78FD8AAE77" box="[460,533,1604,1631]" captionStart="Fig" captionStartId="12.[189,232,1790,1816]" captionTargetBox="[229,1358,264,1753]" captionTargetId="figure@12.[229,1358,263,1753]" captionTargetPageId="12" captionText="Fig. 5. Xibalbanus cozumelensis sp. nov., paratype (ZMUC-CRU-4792). Cephalon and cephalic structures. Scanning electron microscopy. A. Cephalon with cephalic shield and appendages. B. Antenna 1, right side, proximal parts of dorsal and ventral branches. C–D. Unidentified suctorians attached at the proximo-ventral parts of antenna 1. E. Labrum, frontal filaments, and parts of antennae 1 and 2, right side. F. Frontal filaments. G. Antenna 2, left side from anterior. H. Mandible, left side from anterior. I. Mandible, left side from posterior. J. Mandible, left side, apical view on gnathal edge. K. Mandible, left side, apical view on molar process." figureDoi="http://doi.org/10.5281/zenodo.3827401" httpUri="https://zenodo.org/record/3827401/files/figure.png" pageId="14" pageNumber="15">Figs 5</figureCitation>
C–D, 10F–G).
</paragraph>
<paragraph id="8B8EFC32FFCD6454FF088CBBFCA4AE88" blockId="14.[189,827,1670,1697]" box="[189,827,1670,1697]" pageId="14" pageNumber="15">
<heading id="D0C64B5EFFCD6454FF088CBBFCA4AE88" bold="true" box="[189,827,1670,1697]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B9452020FFCD6454FF088CBBFCA4AE88" bold="true" box="[189,827,1670,1697]" pageId="14" pageNumber="15">
<taxonomicName id="4C3187B1FFCD6454FF088CBBFEDBAE88" authorityName="Yager" authorityYear="1981" box="[189,324,1670,1696]" class="Remipedia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Arthropoda" rank="class">Remipedia</taxonomicName>
CO1 and 16S mtDNA distance matrices
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8EFC32FFCD6448FF088C88FDF9A9A7" blockId="14.[189,1399,1717,2027]" lastBlockId="18.[189,1399,267,399]" lastPageId="18" lastPageNumber="19" pageId="14" pageNumber="15">
<tableCitation id="C6B3C989FFCD6454FF088C88FE8CAEE7" box="[189,275,1717,1743]" captionStart="Table 2" captionStartId="6.[188,257,539,567]" captionTargetPageId="6" captionText="Table 2 (continued on next page). Genetic distance matrices for CO1 and 16S including three different specimens of Xibalbanus cozumelensis sp. nov. and comparable sequences of other remipedes." httpUri="http://table.plazi.org/id/DF4EACBAFFC5645CFF098826FD7FAA72" pageId="14" pageNumber="15" tableUuid="DF4EACBAFFC5645CFF098826FD7FAA72">Table 2</tableCitation>
shows uncorrected genetic distances of CO1 and 16S mtDNA among populations of 
<taxonomicName id="4C3187B1FFCD6454FB448C88FEC7AEDB" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFCD6454FB448C88FEC7AEDB" italics="true" pageId="14" pageNumber="15">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFCD6454FED08CE4FE59AEDB" box="[357,454,1753,1779]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
obtained in this study and other 
<taxonomicName id="4C3187B1FFCD6454FCC48CE5FC6FAEDA" authorityName="Yager" authorityYear="1981" box="[881,1008,1752,1778]" class="Remipedia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Arthropoda" rank="class">Remipedia</taxonomicName>
of which data are available in GenBank (see also 
<bibRefCitation id="EFA081C3FFCD6454FE198CC1FD10AF3E" author="Neiber M. T. &amp; Hansen F. C. &amp; Iliffe T. M. &amp; Gonzalez B. C. &amp; Koenemann S." box="[428,655,1788,1814]" pageId="14" pageNumber="15" pagination="31 - 46" refId="ref12178" refString="Neiber M. T., Hansen F. C., Iliffe T. M., Gonzalez B. C. &amp; Koenemann S. 2012. Molecular taxonomy of Speleonectes fuchscockburni, a new pseudocryptic species of Remipedia (Crustacea) from an anchialine cave system on the Yucatan Peninsula, Quintana Roo, Mexico. Zootaxa 3190: 31 - 46. https: // doi. org / 10.15468 / ztlpt 9" type="journal article" year="2012">
Neiber 
<emphasis id="B9452020FFCD6454FDB28CC0FDD9AF3E" box="[519,582,1788,1814]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2012
</bibRefCitation>
; 
<bibRefCitation id="EFA081C3FFCD6454FD2A8CC0FC5EAF3E" author="Hoenemann M. &amp; Neiber M. T. &amp; Koenemann S. &amp; Humphreys W. F. &amp; Iliffe T. M. &amp; Li D. &amp; Schram F. R. &amp; Koenemann S." box="[671,961,1788,1815]" pageId="14" pageNumber="15" pagination="603 - 619" refId="ref11721" refString="Hoenemann M., Neiber M. T., Koenemann S., Humphreys W. F., Iliffe T. M., Li D., Schram F. R. &amp; Koenemann S. 2013. Phylogenetic analysis and systematic revision of Remipedia (Nectiopoda) from Bayesian analysis of molecular data. Journal of Crustacean Biology 33: 603 - 619. https: // doi. org / 10.1163 / 1937240 X- 00002179" type="journal article" year="2013">
Hoenemann 
<emphasis id="B9452020FFCD6454FC8D8CC0FCE9AF3E" box="[824,886,1788,1814]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2013
</bibRefCitation>
). The CO1 gene was sequenced for three specimens of 
<taxonomicName id="4C3187B1FFCD6454FE1F8D1DFDF6AF11" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[426,617,1823,1849]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFCD6454FE1F8D1DFDF6AF11" box="[426,617,1823,1849]" italics="true" pageId="14" pageNumber="15">X. cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFCD6454FDC18D1DFD4DAF12" box="[628,722,1824,1850]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
yielding a 0–3% pairwise genetic distance between the individuals. The 
<taxonomicName id="4C3187B1FFCD6454FECA8D79FDA7AF75" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[383,568,1859,1885]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cozumelensis">
<emphasis id="B9452020FFCD6454FECA8D79FDA7AF75" box="[383,568,1859,1885]" italics="true" pageId="14" pageNumber="15">X. cozumelensis</emphasis>
</taxonomicName>
specimens are genetically (CO1) closest to populations of 
<taxonomicName id="4C3187B1FFCD6454FB6F8D79FAE9AF75" baseAuthorityName="Yager" baseAuthorityYear="1987" box="[1242,1398,1859,1885]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="tulumensis">
<emphasis id="B9452020FFCD6454FB6F8D79FAE9AF75" box="[1242,1398,1859,1885]" italics="true" pageId="14" pageNumber="15">X. tulumensis</emphasis>
</taxonomicName>
from the ‘mainland’ 
<collectingRegion id="49F532D0FFCD6454FE018D5BFD8DAFA8" box="[436,530,1894,1920]" country="Mexico" name="Yucatan" pageId="14" pageNumber="15">Yucatán</collectingRegion>
Peninsula with a 9–11% genetic distance. The genetic distance between 
<taxonomicName id="4C3187B1FFCD6454FF088DB6FEE7AF8C" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[189,376,1930,1956]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cozumelensis">
<emphasis id="B9452020FFCD6454FF088DB6FEE7AF8C" box="[189,376,1930,1956]" italics="true" pageId="14" pageNumber="15">X. cozumelensis</emphasis>
</taxonomicName>
and other remipedes varies between 13% and 25%. The genetic distance (CO1) among species of 
<taxonomicName id="4C3187B1FFCD6454FE8C8D90FE27AFEF" authorityName="Yager" authorityYear="1981" box="[313,440,1965,1991]" class="Remipedia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="14" pageNumber="15" phylum="Arthropoda" rank="class">Remipedia</taxonomicName>
in general varies between 12% and 31%, but only with few values in the low end of the spectrum, e.g., the distance between two species of 
<taxonomicName id="4C3187B1FFCD6454FCCD8DECFBA5AFC3" authorityName="Yager" authorityYear="1987" box="[888,1082,2001,2027]" class="Remipedia" family="Speleonectidae" genus="Cryptocorynetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B9452020FFCD6454FCCD8DECFBA5AFC3" box="[888,1082,2001,2027]" italics="true" pageId="14" pageNumber="15">Cryptocorynetes</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3187B1FFCD644BFBF88DECFE64A90D" authority="Wollermann, Koenemann &amp; Iliffe, 2007" authorityName="Wollermann, Koenemann &amp; Iliffe" authorityYear="2007" class="Remipedia" family="Speleonectidae" genus="Cryptocorynetes" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="17" lastPageNumber="18" order="Nectiopoda" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="longulus">
<emphasis id="B9452020FFCD6454FBF88DECFB49AFC3" box="[1101,1238,2001,2027]" italics="true" pageId="14" pageNumber="15">C. longulus</emphasis>
Wollermann, Koenemann &amp; Iliffe, 2007
</taxonomicName>
and 
<taxonomicName id="4C3187B1FFD2644BFD8D8B31FC1DA90D" authority="Yager, 1987" authorityName="Yager" authorityYear="1987" box="[568,898,267,294]" class="Remipedia" family="Speleonectidae" genus="Cryptocorynetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="haptodiscus">
<emphasis id="B9452020FFD2644BFD8D8B31FD76A90D" box="[568,745,267,294]" italics="true" pageId="17" pageNumber="18">C. haptodiscus</emphasis>
<bibRefCitation id="EFA081C3FFD2644BFD448B31FC1DA90D" author="Yager J." box="[753,898,267,294]" pageId="17" pageNumber="18" pagination="160 - 166" refId="ref12897" refString="Yager J. 1987. Speleonectes tulumensis n. sp. (Crustacea: Remipedia) from two anchialine cenotes of the Yucatan Peninsula, Mexico. Stygologia 3: 160 - 166." type="journal article" year="1987">Yager, 1987</bibRefCitation>
</taxonomicName>
) and two species of 
<taxonomicName id="4C3187B1FFD2644BFBCB8B36FB68A90D" authorityName="Schram, Yager &amp; Emerson" authorityYear="1986" box="[1150,1271,267,293]" class="Remipedia" family="Godzilliidae" genus="Godzillius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9452020FFD2644BFBCB8B36FB68A90D" box="[1150,1271,267,293]" italics="true" pageId="17" pageNumber="18">Godzillius</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3187B1FFD2644BFABF8B31FD16A961" authority="Gonzalez, Singpiel &amp; Schlagner, 2013" authorityName="Gonzalez, Singpiel &amp; Schlagner" authorityYear="2013" class="Remipedia" family="Godzilliidae" genus="Godzillius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="fuchsi">
<emphasis id="B9452020FFD2644BFABF8B31FAE9A90D" box="[1290,1398,267,294]" italics="true" pageId="17" pageNumber="18">G. fuchsi</emphasis>
Gonzalez, Singpiel &amp; Schlagner, 2013
</taxonomicName>
and 
<taxonomicName id="4C3187B1FFD2644BFD738B12FB75A961" authority="Schram, Yager &amp; Emerson, 1986" authorityName="Schram, Yager &amp; Emerson" authorityYear="1986" box="[710,1258,302,329]" class="Remipedia" family="Godzilliidae" genus="Godzillius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B9452020FFD2644BFD738B12FCCFA961" box="[710,848,303,329]" italics="true" pageId="17" pageNumber="18">G. robustus</emphasis>
Schram, Yager &amp; Emerson, 1986
</taxonomicName>
), which are 13%, and the distances between two species of 
<taxonomicName id="4C3187B1FFD2644BFD4B8B6FFCE7A944" authorityName="Garcia-Valdecasas" authorityYear="1984" box="[766,888,338,364]" class="Remipedia" family="Morlockiidae" genus="Morlockia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B9452020FFD2644BFD4B8B6FFCE7A944" box="[766,888,338,364]" italics="true" pageId="17" pageNumber="18">Morlockia</emphasis>
</taxonomicName>
(
<taxonomicName id="4C3187B1FFD2644BFC398B6EFADCA944" authority="Garcia-Valdecasas, 1984" authorityName="Garcia-Valdecasas" authorityYear="1984" box="[908,1347,338,364]" class="Remipedia" family="Morlockiidae" genus="Morlockia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ondinae">
<emphasis id="B9452020FFD2644BFC398B6EFB8BA944" box="[908,1044,338,364]" italics="true" pageId="17" pageNumber="18">M. ondinae</emphasis>
García-Valdecasas, 1984
</taxonomicName>
and 
<taxonomicName id="4C3187B1FFD2644BFF088B4BFD0CA9A7" authority="(Koenemann et al., 2009))" baseAuthorityName="Koenemann" baseAuthorityYear="2009" box="[189,659,373,399]" class="Remipedia" family="Morlockiidae" genus="Morlockia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="atlantida">
<emphasis id="B9452020FFD2644BFF088B4BFED0A9A7" box="[189,335,373,399]" italics="true" pageId="17" pageNumber="18">M. atlantida</emphasis>
(
<bibRefCitation id="EFA081C3FFD2644BFEEA8B48FD1FA9A7" author="Koenemann S. &amp; Bloechl A. &amp; Martinez A. &amp; Iliffe T. M. &amp; Hoenemann M. &amp; Oromi P." box="[351,640,373,399]" pageId="17" pageNumber="18" pagination="215 - 225" refId="ref11883" refString="Koenemann S., Bloechl A., Martinez A., Iliffe T. M., Hoenemann M. &amp; Oromi P. 2009. A new, disjunct species of Speleonectes (Remipedia, Crustacea) from the Canary Islands. Marine Biodiversity 39: 215 - 225. https: // doi. org / 10.1007 / s 12526 - 009 - 0021 - 8" type="journal article" year="2009">
Koenemann 
<emphasis id="B9452020FFD2644BFE428B4BFDB4A9A7" box="[503,555,373,399]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
., 2009
</bibRefCitation>
))
</taxonomicName>
, which is 12%. The rest of the CO1 genetic distances between species of 
<taxonomicName id="4C3187B1FFD2644BFE8C8BA5FE27A99A" authorityName="Yager" authorityYear="1981" box="[313,440,408,434]" class="Remipedia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Arthropoda" rank="class">Remipedia</taxonomicName>
based on sequences available in GenBank are included for completeness but will not be described further here (
<tableCitation id="C6B3C989FFD2644BFDAD8B86FDEDA9FD" box="[536,626,443,469]" captionStart="Table 2" captionStartId="6.[188,257,539,567]" captionTargetPageId="6" captionText="Table 2 (continued on next page). Genetic distance matrices for CO1 and 16S including three different specimens of Xibalbanus cozumelensis sp. nov. and comparable sequences of other remipedes." httpUri="http://table.plazi.org/id/DF4EACBAFFC5645CFF098826FD7FAA72" pageId="17" pageNumber="18" tableUuid="DF4EACBAFFC5645CFF098826FD7FAA72">Table 2</tableCitation>
). 16S was sequenced for one specimen of 
<taxonomicName id="4C3187B1FFD2644BFBE88B81FA88A9FD" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[1117,1303,443,469]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFD2644BFBE88B81FA88A9FD" box="[1117,1303,443,469]" italics="true" pageId="17" pageNumber="18">X. cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFD2644BFAA88B81FAE8A9FE" box="[1309,1399,444,470]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
The specimen is genetically (16S) closest to 
<taxonomicName id="4C3187B1FFD2644BFD648BE2FCEFA9D1" baseAuthorityName="Yager" baseAuthorityYear="1987" box="[721,880,479,505]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tulumensis">
<emphasis id="B9452020FFD2644BFD648BE2FCEFA9D1" box="[721,880,479,505]" italics="true" pageId="17" pageNumber="18">X. tulumensis</emphasis>
</taxonomicName>
showing genetic distances of 14–19%. The genetic distances in the 16S gene between species of 
<taxonomicName id="4C3187B1FFD2644BFCF0883FFC5BAA34" authorityName="Yager" authorityYear="1981" box="[837,964,514,540]" class="Remipedia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="17" pageNumber="18" phylum="Arthropoda" rank="class">Remipedia</taxonomicName>
generally are between 20–30%, but with the distance between a few species being lower and at the same level as between 
<taxonomicName id="4C3187B1FFD2644BFB0F881BFAE8AA17" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[1210,1399,549,575]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFD2644BFB0F881BFAE8AA17" box="[1210,1399,549,575]" italics="true" pageId="17" pageNumber="18">X. cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFD2644BFF088874FE85AA4B" box="[189,282,585,611]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C3187B1FFD2644BFEE28874FE68AA4A" baseAuthorityName="Yager" baseAuthorityYear="1987" box="[343,503,584,610]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tulumensis">
<emphasis id="B9452020FFD2644BFEE28874FE68AA4A" box="[343,503,584,610]" italics="true" pageId="17" pageNumber="18">X. tulumensis</emphasis>
</taxonomicName>
(e.g., 
<taxonomicName id="4C3187B1FFD2644BFDF38874FC02AA4A" authorityName="Yager" authorityYear="1987" box="[582,925,584,611]" class="Remipedia" family="Speleonectidae" genus="Cryptocorynetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="haptodiscus">
<emphasis id="B9452020FFD2644BFDF38874FC02AA4A" box="[582,925,584,611]" italics="true" pageId="17" pageNumber="18">Cryptocorynetes haptodiscus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3187B1FFD26448FC6E8874FEDDA90D" authority="Hazerli, Koenemann &amp; Iliffe, 2009" authorityName="Hazerli, Koenemann &amp; Iliffe" authorityYear="2009" class="Remipedia" family="Speleonectidae" genus="Cryptocorynetes" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="18" lastPageNumber="19" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="elmorei">
<emphasis id="B9452020FFD2644BFC6E8874FBC7AA4A" box="[987,1112,584,611]" italics="true" pageId="17" pageNumber="18">C. elmorei</emphasis>
Hazerli, 
<bibRefCitation id="EFA081C3FFD26448FB7D8874FEDDA90D" author="Koenemann S. &amp; Bloechl A. &amp; Martinez A. &amp; Iliffe T. M. &amp; Hoenemann M. &amp; Oromi P." lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" pagination="215 - 225" refId="ref11883" refString="Koenemann S., Bloechl A., Martinez A., Iliffe T. M., Hoenemann M. &amp; Oromi P. 2009. A new, disjunct species of Speleonectes (Remipedia, Crustacea) from the Canary Islands. Marine Biodiversity 39: 215 - 225. https: // doi. org / 10.1007 / s 12526 - 009 - 0021 - 8" type="journal article" year="2009">Koenemann &amp; Iliffe, 2009</bibRefCitation>
</taxonomicName>
at 16%, and 
<taxonomicName id="4C3187B1FFD16448FE568B36FC0AA90D" authority="Yager, 1981" authorityName="Yager" authorityYear="1981" box="[483,917,267,294]" class="Remipedia" family="Speleonectidae" genus="Speleonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lucayensis">
<emphasis id="B9452020FFD16448FE568B36FD62A90D" box="[483,765,267,293]" italics="true" pageId="18" pageNumber="19">Speleonectes lucayensis</emphasis>
<bibRefCitation id="EFA081C3FFD16448FCB28B31FC0AA90D" author="Yager J." box="[775,917,267,294]" pageId="18" pageNumber="19" pagination="328 - 333" refId="ref12854" refString="Yager J. 1981. Remipedia, a new class of Crustacea from a marine cave in the Bahamas. Journal of Crustacean Biology 1 (3): 328 - 333. https: // doi. org / 10.2307 / 1547965" type="journal article" year="1981">Yager, 1981</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C3187B1FFD16448FC678B31FE09A961" authority="Daenekas, Iliffe, Yager &amp; Koenemann, 2009" authorityName="Daenekas, Iliffe, Yager &amp; Koenemann" authorityYear="2009" class="Remipedia" family="Speleonectidae" genus="Speleonectes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kakuki">
<emphasis id="B9452020FFD16448FC678B31FBA1A90D" box="[978,1086,267,294]" italics="true" pageId="18" pageNumber="19">S. kakuki</emphasis>
<bibRefCitation id="EFA081C3FFD16448FBF38B36FE09A961" author="Daenekas J. &amp; Iliffe T. &amp; Yager J. &amp; Koenemann S." pageId="18" pageNumber="19" pagination="51 - 66" refId="ref11481" refString="Daenekas J., Iliffe T. M, Yager J. &amp; Koenemann S. 2009. Speleonectes kakuki, a new species of Remipedia (Crustacea) from anchialine and sub-seafloor caves on Andros and Cat Island, Bahamas. Zootaxa 2016: 51 - 66. https: // doi. org / 10.15468 / c 8 ooey" type="journal article" year="2009">Daenekas, Iliffe, Yager &amp; Koenemann, 2009</bibRefCitation>
</taxonomicName>
at 17%). CytB was compared to CytB extracted from the complete mitochondrion of 
<taxonomicName id="4C3187B1FFD16448FF6A8B6EFEE0A944" baseAuthorityName="Yager" baseAuthorityYear="1987" box="[223,383,338,364]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tulumensis">
<emphasis id="B9452020FFD16448FF6A8B6EFEE0A944" box="[223,383,338,364]" italics="true" pageId="18" pageNumber="19">X. tulumensis</emphasis>
</taxonomicName>
(
<accessionNumber id="946261D1FFD16448FE248B6FFD87A945" box="[401,536,338,365]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY456190" pageId="18" pageNumber="19">AY456190</accessionNumber>
). The 
<taxonomicName id="4C3187B1FFD16448FDD18B6EFCBEA944" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[612,801,338,364]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFD16448FDD18B6EFCBEA944" box="[612,801,338,364]" italics="true" pageId="18" pageNumber="19">X. cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFD16448FC9E8B6FFC17A944" box="[811,904,338,364]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
sequences were identical and had a 15% genetic distance from 
<taxonomicName id="4C3187B1FFD16448FE748B4BFDC1A9A7" baseAuthorityName="Yager" baseAuthorityYear="1987" box="[449,606,373,399]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="tulumensis">
<emphasis id="B9452020FFD16448FE748B4BFDC1A9A7" box="[449,606,373,399]" italics="true" pageId="18" pageNumber="19">X. tulumensis</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF4EACBAFFCC6455FF088D79FDAEAFC3" ID-DOI="http://doi.org/10.5281/zenodo.3827405" ID-Zenodo-Dep="3827405" httpUri="https://zenodo.org/record/3827405/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1860,1886]" targetBox="[236,1353,268,1818]" targetPageId="15">
<paragraph id="8B8EFC32FFCC6455FF088D79FDAEAFC3" blockId="15.[189,1399,1860,2028]" pageId="15" pageNumber="16">
<emphasis id="B9452020FFCC6455FF088D79FE91AF76" bold="true" box="[189,270,1860,1886]" pageId="15" pageNumber="16">Fig. 7.</emphasis>
<taxonomicName id="4C3187B1FFCC6455FEAC8D79FDDAAF76" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[281,581,1860,1886]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFCC6455FEAC8D79FDDAAF76" box="[281,581,1860,1886]" italics="true" pageId="15" pageNumber="16">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFCC6455FDE58D78FD2EAF77" box="[592,689,1861,1887]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ZMUC-CRU-4792), maxilla 2 (mx2), left side, scanning electron microscopy. 
<emphasis id="B9452020FFCC6455FD9E8D5AFDDFAFA9" bold="true" box="[555,576,1895,1921]" pageId="15" pageNumber="16">A</emphasis>
. Complete mx2 from anterior. 
<emphasis id="B9452020FFCC6455FC058D55FC5BAFAA" bold="true" box="[944,964,1896,1922]" pageId="15" pageNumber="16">B</emphasis>
. Close-up of terminal claw-complex from anterior (segment 7). 
<emphasis id="B9452020FFCC6455FE498DB7FD8EAF8C" bold="true" box="[508,529,1930,1956]" pageId="15" pageNumber="16">C</emphasis>
. Close-up of terminal claw-complex from lateral (segment 7). 
<emphasis id="B9452020FFCC6455FB4D8DB6FA92AF8D" bold="true" box="[1272,1293,1931,1957]" pageId="15" pageNumber="16">D</emphasis>
. Median side of segments 1 and 2 with endites (e1–3) from anterior. 
<emphasis id="B9452020FFCC6455FCD98D93FC1FAFE0" bold="true" box="[876,896,1966,1992]" pageId="15" pageNumber="16">E</emphasis>
. Endites of segment 1 (e1–3) from median. 
<emphasis id="B9452020FFCC6455FF088DECFF50AFC3" bold="true" box="[189,207,2001,2027]" pageId="15" pageNumber="16">F</emphasis>
. Complete mx2 from median.
</paragraph>
</caption>
<caption id="DF4EACBAFFD3644AFF088D3DFC89AFC6" ID-DOI="http://doi.org/10.5281/zenodo.3827407" ID-Zenodo-Dep="3827407" httpUri="https://zenodo.org/record/3827407/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1792,1818]" targetBox="[276,1313,267,1763]" targetPageId="16">
<paragraph id="8B8EFC32FFD3644AFF088D3DFC89AFC6" blockId="16.[189,1399,1792,2030]" pageId="16" pageNumber="17">
<emphasis id="B9452020FFD3644AFF088D3DFE92AF32" bold="true" box="[189,269,1792,1818]" pageId="16" pageNumber="17">Fig. 8.</emphasis>
<taxonomicName id="4C3187B1FFD3644AFEA28D3DFDDDAF32" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[279,578,1792,1818]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFD3644AFEA28D3DFDDDAF32" box="[279,578,1792,1818]" italics="true" pageId="16" pageNumber="17">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFD3644AFDF98D3CFD33AF33" box="[588,684,1793,1819]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ZMUC-CRU-4792), maxilliped (mxp), left side, scanning electron microscopy. 
<emphasis id="B9452020FFD3644AFD9E8D1EFDDFAF15" bold="true" box="[555,576,1827,1853]" pageId="16" pageNumber="17">A</emphasis>
. Complete mxp from anterior. 
<emphasis id="B9452020FFD3644AFC058D19FC5BAF16" bold="true" box="[944,964,1828,1854]" pageId="16" pageNumber="17">B</emphasis>
. Close-up of terminal claw-complex from anterior (segment 8). 
<emphasis id="B9452020FFD3644AFDB28D7AFD83AF49" bold="true" box="[519,540,1863,1889]" pageId="16" pageNumber="17">C</emphasis>
. Complete mxp from posterior. 
<emphasis id="B9452020FFD3644AFC108D7AFC25AF49" bold="true" box="[933,954,1863,1889]" pageId="16" pageNumber="17">D</emphasis>
. Close-up of terminal claw-complex from posterior (segment 8). 
<emphasis id="B9452020FFD3644AFDB78D57FD89AFAC" bold="true" box="[514,534,1898,1924]" pageId="16" pageNumber="17">E</emphasis>
. Close-up of terminal claw-complex from lateral (segment 8). 
<emphasis id="B9452020FFD3644AFB588D57FB60AFAC" bold="true" box="[1261,1279,1898,1924]" pageId="16" pageNumber="17">F</emphasis>
. Close-up of small cluster of setae along distal margin of segment 7. 
<emphasis id="B9452020FFD3644AFCC18DB0FC14AF8F" bold="true" box="[884,907,1933,1959]" pageId="16" pageNumber="17">G</emphasis>
. Close-up of pore at close to latero-distal margin of segment 7. 
<emphasis id="B9452020FFD3644AFE088D8CFE4BAFE3" bold="true" box="[445,468,1969,1995]" pageId="16" pageNumber="17">H</emphasis>
. Close-up of cluster of setae at latero-distal corner of segment 4. 
<emphasis id="B9452020FFD3644AFB668D8CFB41AFE3" bold="true" box="[1235,1246,1969,1995]" pageId="16" pageNumber="17">I</emphasis>
. Close-up of cluster of setae at latero-distal corner of segment 5.
</paragraph>
</caption>
<caption id="DF4EACBAFFD2644BFF088DB4FF60AFC1" ID-DOI="http://doi.org/10.5281/zenodo.3827409" ID-Zenodo-Dep="3827409" httpUri="https://zenodo.org/record/3827409/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1929,1955]" targetBox="[189,1398,678,1893]" targetPageId="17">
<paragraph id="8B8EFC32FFD2644BFF088DB4FF60AFC1" blockId="17.[189,1398,1929,2025]" pageId="17" pageNumber="18">
<emphasis id="B9452020FFD2644BFF088DB4FE91AF8B" bold="true" box="[189,270,1929,1955]" pageId="17" pageNumber="18">Fig. 9.</emphasis>
<taxonomicName id="4C3187B1FFD2644BFEAC8DB4FDDBAF8B" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[281,580,1929,1955]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFD2644BFEAC8DB4FDDBAF8B" box="[281,580,1929,1955]" italics="true" pageId="17" pageNumber="18">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFD2644BFDFA8DB7FD30AF8C" box="[591,687,1930,1956]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (ZMUC-CRU-4791), trunk limbs, left side from anterior, light microscopy. 
<emphasis id="B9452020FFD2644BFE4E8D91FDB1AFEE" bold="true" box="[507,558,1964,1990]" pageId="17" pageNumber="18">A–J</emphasis>
. Selected trunk segments (number specified on figure). All to the same scale.
</paragraph>
</caption>
<caption id="DF4EACBAFFD16448FF088D1DFC32AFC2" ID-DOI="http://doi.org/10.5281/zenodo.3827411" ID-Zenodo-Dep="3827411" httpUri="https://zenodo.org/record/3827411/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,232,1824,1850]" targetBox="[190,1398,436,1789]" targetPageId="18">
<paragraph id="8B8EFC32FFD16448FF088D1DFC32AFC2" blockId="18.[189,1399,1824,2027]" pageId="18" pageNumber="19">
<emphasis id="B9452020FFD16448FF088D1DFE82AF12" bold="true" box="[189,285,1824,1850]" pageId="18" pageNumber="19">Fig. 10.</emphasis>
<taxonomicName id="4C3187B1FFD16448FE928D1CFDCDAF13" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[295,594,1825,1851]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFD16448FE928D1CFDCDAF13" box="[295,594,1825,1851]" italics="true" pageId="18" pageNumber="19">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFD16448FDE88D1CFD22AF13" box="[605,701,1825,1851]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, paratype (ZMUC-CRU-4792), cephalon, body segments, and trunk limbs, scanning electron microscopy. 
<emphasis id="B9452020FFD16448FCB38D79FC84AF76" bold="true" box="[774,795,1860,1886]" pageId="18" pageNumber="19">A</emphasis>
. Cephalon, cephalic shield, and head appendages from lateral. 
<emphasis id="B9452020FFD16448FEE98D5AFEEFAFA9" bold="true" box="[348,368,1895,1921]" pageId="18" pageNumber="19">B</emphasis>
. Trunk segments 5–10. 
<emphasis id="B9452020FFD16448FD228D5AFD33AFA9" bold="true" box="[663,684,1895,1921]" pageId="18" pageNumber="19">C</emphasis>
. Trunk limb 2, right side. 
<emphasis id="B9452020FFD16448FC418D5AFB96AFA9" bold="true" box="[1012,1033,1895,1921]" pageId="18" pageNumber="19">D</emphasis>
. Close-up of serrated setae in latero-distal corner of exopod segment 1. 
<emphasis id="B9452020FFD16448FD108DB7FD26AF8C" bold="true" box="[677,697,1930,1956]" pageId="18" pageNumber="19">E</emphasis>
. Close-up of serrated setae in latero-distal corner of exopod segment 2. 
<emphasis id="B9452020FFD16448FEF38D90FEE2AFEF" bold="true" box="[326,381,1965,1991]" pageId="18" pageNumber="19">F–G</emphasis>
. Two unidentified suctorians attached to exopod of trunk limb 3. 
<emphasis id="B9452020FFD16448FB3C8D93FB25AFE0" bold="true" box="[1161,1210,1966,1992]" pageId="18" pageNumber="19">H–I</emphasis>
. Posterior body segments, trunk limbs, and caudal rami from ventral and lateral.
</paragraph>
</caption>
<paragraph id="8B8EFC32FFD06449FF088B36FB59A90D" blockId="19.[189,1222,267,293]" box="[189,1222,267,293]" pageId="19" pageNumber="20">
<heading id="D0C64B5EFFD06449FF088B36FB59A90D" bold="true" box="[189,1222,267,293]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B9452020FFD06449FF088B36FB59A90D" bold="true" box="[189,1222,267,293]" pageId="19" pageNumber="20">
Phylogeny of 
<taxonomicName id="4C3187B1FFD06449FED38B36FE72A90D" authorityName="Yager" authorityYear="1981" box="[358,493,267,293]" class="Remipedia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="20" phylum="Arthropoda" rank="class">Remipedia</taxonomicName>
based on Bayesian analysis of CO1 and 16S mtDNA genes
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8EFC32FFD06449FF088B70FCF4AA4A" blockId="19.[189,1399,333,610]" pageId="19" pageNumber="20">
The phylogenetic tree of 
<taxonomicName id="4C3187B1FFD06449FE458B70FDF0A94F" authorityName="Yager" authorityYear="1981" box="[496,623,333,359]" class="Remipedia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="20" phylum="Arthropoda" rank="class">Remipedia</taxonomicName>
in 
<figureCitation id="130AE0B7FFD06449FD2F8B70FD70A94F" box="[666,751,333,359]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[231,1350,265,1910]" captionTargetId="figure@20.[231,1350,265,1910]" captionTargetPageId="20" captionText="Fig. 11. Phylogenetic position of Xibalbanus cozumelensis sp. nov. based on a Bayesian analysis of partial CO1 and 16S rDNA sequences of X. cozumelensis sp. nov. and other remipedes available from GenBank." figureDoi="http://doi.org/10.5281/zenodo.3827413" httpUri="https://zenodo.org/record/3827413/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
is based on a Bayesian analysis of the CO1 and 16S mtDNA genes. New sequences of both CO1 and 16S have been obtained for 
<taxonomicName id="4C3187B1FFD06449FBFB8B4CFAE8A9A3" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[1102,1399,369,395]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFD06449FBFB8B4CFAE8A9A3" box="[1102,1399,369,395]" italics="true" pageId="19" pageNumber="20">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFD06449FF088BA8FE85A987" box="[189,282,405,431]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
Sequences from other species of 
<taxonomicName id="4C3187B1FFD06449FD078BA9FCAEA986" authorityName="Yager" authorityYear="1981" box="[690,817,404,430]" class="Remipedia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="19" pageNumber="20" phylum="Arthropoda" rank="class">Remipedia</taxonomicName>
are from GenBank and were previously used in 
<bibRefCitation id="EFA081C3FFD06449FF088B85FE34A9FA" author="Neiber M. T. &amp; Hansen F. C. &amp; Iliffe T. M. &amp; Gonzalez B. C. &amp; Koenemann S." box="[189,427,440,466]" pageId="19" pageNumber="20" pagination="31 - 46" refId="ref12178" refString="Neiber M. T., Hansen F. C., Iliffe T. M., Gonzalez B. C. &amp; Koenemann S. 2012. Molecular taxonomy of Speleonectes fuchscockburni, a new pseudocryptic species of Remipedia (Crustacea) from an anchialine cave system on the Yucatan Peninsula, Quintana Roo, Mexico. Zootaxa 3190: 31 - 46. https: // doi. org / 10.15468 / ztlpt 9" type="journal article" year="2012">
Neiber 
<emphasis id="B9452020FFD06449FEA38B84FECCA9FA" box="[278,339,440,466]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
(2012)
</bibRefCitation>
and 
<bibRefCitation id="EFA081C3FFD06449FE5D8B84FC87A9FA" author="Hoenemann M. &amp; Neiber M. T. &amp; Koenemann S. &amp; Humphreys W. F. &amp; Iliffe T. M. &amp; Li D. &amp; Schram F. R. &amp; Koenemann S." box="[488,792,440,467]" pageId="19" pageNumber="20" pagination="603 - 619" refId="ref11721" refString="Hoenemann M., Neiber M. T., Koenemann S., Humphreys W. F., Iliffe T. M., Li D., Schram F. R. &amp; Koenemann S. 2013. Phylogenetic analysis and systematic revision of Remipedia (Nectiopoda) from Bayesian analysis of molecular data. Journal of Crustacean Biology 33: 603 - 619. https: // doi. org / 10.1163 / 1937240 X- 00002179" type="journal article" year="2013">
Hoenemann 
<emphasis id="B9452020FFD06449FDCA8B84FD23A9FA" box="[639,700,440,466]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
(2013)
</bibRefCitation>
. In the phylogeny (
<figureCitation id="130AE0B7FFD06449FBB78B85FBC7A9FA" box="[1026,1112,440,466]" captionStart="Fig" captionStartId="20.[189,232,1930,1956]" captionTargetBox="[231,1350,265,1910]" captionTargetId="figure@20.[231,1350,265,1910]" captionTargetPageId="20" captionText="Fig. 11. Phylogenetic position of Xibalbanus cozumelensis sp. nov. based on a Bayesian analysis of partial CO1 and 16S rDNA sequences of X. cozumelensis sp. nov. and other remipedes available from GenBank." figureDoi="http://doi.org/10.5281/zenodo.3827413" httpUri="https://zenodo.org/record/3827413/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
) all included species of 
<taxonomicName id="4C3187B1FFD06449FF088BE1FEDCA9DE" authorityName="Hoenemann, Neiber, Schram &amp; Koenemann" authorityYear="2013" box="[189,323,476,502]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9452020FFD06449FF088BE1FEDCA9DE" box="[189,323,476,502]" italics="true" pageId="19" pageNumber="20">Xibalbanus</emphasis>
</taxonomicName>
were in the same clade (monophyletic). 
<taxonomicName id="4C3187B1FFD06449FCAE8BE1FBDDA9DE" authority="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe, 2017" authorityName="Olesen &amp; Meland &amp; Glenner &amp; Van Hengstum &amp; Iliffe" authorityYear="2017" box="[795,1090,476,502]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="cozumelensis" status="sp. nov.">
<emphasis id="B9452020FFD06449FCAE8BE1FBDDA9DE" box="[795,1090,476,502]" italics="true" pageId="19" pageNumber="20">Xibalbanus cozumelensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2769D5BFFD06449FBFC8BE0FB3CA9DF" box="[1097,1187,477,503]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
appeared as sister species to 
<taxonomicName id="4C3187B1FFD06449FE82883CFE4BAA32" baseAuthorityName="Yager" baseAuthorityYear="1987" box="[311,468,512,538]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tulumensis">
<emphasis id="B9452020FFD06449FE82883CFE4BAA32" box="[311,468,512,538]" italics="true" pageId="19" pageNumber="20">X. tulumensis</emphasis>
</taxonomicName>
, and these two together appeared as sister of 
<emphasis id="B9452020FFD06449FC5D883CFB5BAA32" box="[1000,1220,512,538]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="4C3187B1FFD06449FC5D883CFB5FAA32" box="[1000,1216,512,538]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="fuchscockburni">X. fuchscockburni</taxonomicName>
.
</emphasis>
The remaining part of the phylogeny was largely similar to the topology obtained by 
<bibRefCitation id="EFA081C3FFD06449FC4A8819FAB8AA16" author="Hoenemann M. &amp; Neiber M. T. &amp; Koenemann S. &amp; Humphreys W. F. &amp; Iliffe T. M. &amp; Li D. &amp; Schram F. R. &amp; Koenemann S." box="[1023,1319,548,574]" pageId="19" pageNumber="20" pagination="603 - 619" refId="ref11721" refString="Hoenemann M., Neiber M. T., Koenemann S., Humphreys W. F., Iliffe T. M., Li D., Schram F. R. &amp; Koenemann S. 2013. Phylogenetic analysis and systematic revision of Remipedia (Nectiopoda) from Bayesian analysis of molecular data. Journal of Crustacean Biology 33: 603 - 619. https: // doi. org / 10.1163 / 1937240 X- 00002179" type="journal article" year="2013">
Hoenemann 
<emphasis id="B9452020FFD06449FB208819FB4EAA16" box="[1173,1233,548,574]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
(2013)
</bibRefCitation>
and is not further described here as the focus was on 
<taxonomicName id="4C3187B1FFD06449FD6B887AFCFBAA49" authorityName="Hoenemann, Neiber, Schram &amp; Koenemann" authorityYear="2013" box="[734,868,583,609]" class="Remipedia" family="Speleonectidae" genus="Xibalbanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nectiopoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B9452020FFD06449FD6B887AFCFBAA49" box="[734,868,583,609]" italics="true" pageId="19" pageNumber="20">Xibalbanus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>