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<document id="81AB4C5C28A0C9C77384E256C38C7530" ID-CLB-Dataset="45728" ID-DOI="10.5281/zenodo.206949" ID-GBIF-Dataset="759eec1b-c953-4b2d-8207-99324b9ceb6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="206949" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460422171139" checkinUser="plazi" docAuthor="Zrelli, Sonia, Sartori, Michel, Bejaoui, Mustapha &amp; Boumaiza, Moncef" docDate="2011" docId="038F87A4C61CFFD514AA417426ACD3DB" docLanguage="en" docName="zt03139p068.pdf" docOrigin="Zootaxa 3139" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Rhithrogena sartorii Zrelli &amp; Boumaiza, sp. nov." docType="treatment" docVersion="11" lastPageNumber="67" masterDocId="FFB6FFDCC61DFFD0143D473B2278D30F" masterDocTitle="Rhithrogena sartorii, a new mayfly species (Ephemeroptera: Heptageniidae) from North Africa" masterLastPageNumber="68" masterPageNumber="63" pageNumber="64" updateTime="1698330151770" updateUser="ExternalLinkService">
<mods:mods id="45D76B755F0504E517D1AD0E23A64AF9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8B721EE3DBD82952563D83C73E3E5C6C">
<mods:title id="7C15F8CD74A935C0AEBA1EA8B732210C">Rhithrogena sartorii, a new mayfly species (Ephemeroptera: Heptageniidae) from North Africa</mods:title>
</mods:titleInfo>
<mods:name id="A8941B6889D0A450A91B603AF68AB9B4" type="personal">
<mods:role id="DD64475A6349AE35AD50EE12B6D0BEB9">
<mods:roleTerm id="E70DD5E3F04D491E33E04B051F48BDE9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="545CC56681B4643C5BDB309D1ACA2A7D">Zrelli, Sonia</mods:namePart>
</mods:name>
<mods:name id="9D127F2732AD04FCA0BB29BD2B4BFF29" type="personal">
<mods:role id="55444547F65405F87FEFFCB1316C58D8">
<mods:roleTerm id="3EB2485CAC2C64935B0A47087E80DEDD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1ABC3C25308F3C50C8D3CA050EEAE385">Sartori, Michel</mods:namePart>
</mods:name>
<mods:name id="18CAE3441998C8E3D4A74A7BF9D0C667" type="personal">
<mods:role id="9E3B71FAB4150A7060F8935FFA7147DC">
<mods:roleTerm id="EE48D841DF81F197FECC1E7876AC2B74">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E95BFBDD9B264B29118B22D68D76BD58">Bejaoui, Mustapha</mods:namePart>
</mods:name>
<mods:name id="B4CAAB2071D3FDB5038A4D1687578CF1" type="personal">
<mods:role id="9D04998F74CD8D3BA51231937A31F53C">
<mods:roleTerm id="ED8C07A521ED43BC8CD89D0110D8D9A7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CB60CF4B640BCF7E7627441E2E515461">Boumaiza, Moncef</mods:namePart>
</mods:name>
<mods:typeOfResource id="57A66EE2D828A220262FF76D93ACF176">text</mods:typeOfResource>
<mods:relatedItem id="81AC54DAA4FBDDF3573C6C2EC6E17825" type="host">
<mods:titleInfo id="F8AB1E497DD1292685C0E0023820323B">
<mods:title id="9ED4B2ECDCE550B5407D58F50A66177E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B88C298A1C9A3F31CA3B07D0F36C4152">
<mods:date id="D3EB27B3DA476DD270C35EF95E289136">2011</mods:date>
<mods:detail id="0DDC0031F87AA065FA2BC4445A7782C4" type="volume">
<mods:number id="EB752BC8A9D66F05EE94606CEBB8B844">3139</mods:number>
</mods:detail>
<mods:extent id="7762D8072909DAB751E87329F756EF11" unit="page">
<mods:start id="B480304A256BEC559643BF0D6BA4AF4B">63</mods:start>
<mods:end id="BFFCB5377747275D2D1414A4367738A5">68</mods:end>
</mods:extent>
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<mods:classification id="2E976F4B61E96C982218C0844A63E441">journal article</mods:classification>
<mods:identifier id="49DF094084DFC2A398BDA438408E71ED" type="CLB-Dataset">45728</mods:identifier>
<mods:identifier id="D387E779A11A5CDAD9990E4C1C757D36" type="DOI">10.5281/zenodo.206949</mods:identifier>
<mods:identifier id="DE24219B81D72326DAFFF9CF5625A513" type="GBIF-Dataset">759eec1b-c953-4b2d-8207-99324b9ceb6e</mods:identifier>
<mods:identifier id="1BF8DC12AC78EFABD4D83A1B96B0C73E" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BDA7DF32F5B9B16D7D4B3BC181ED0E59" type="Zenodo-Dep">206949</mods:identifier>
</mods:mods>
<treatment id="038F87A4C61CFFD514AA417426ACD3DB" ID-DOI="http://doi.org/10.5281/zenodo.5688316" ID-GBIF-Taxon="119427626" ID-Zenodo-Dep="5688316" LSID="urn:lsid:plazi:treatment:038F87A4C61CFFD514AA417426ACD3DB" httpUri="http://treatment.plazi.org/id/038F87A4C61CFFD514AA417426ACD3DB" lastPageId="5" lastPageNumber="67" pageId="1" pageNumber="64">
<subSubSection id="C33C6539C61CFFD114AA41742358D585" pageId="1" pageNumber="64" type="nomenclature">
<paragraph id="8B9936B2C61CFFD114AA41742094D566" blockId="1.[151,748,1615,1674]" box="[151,748,1615,1641]" pageId="1" pageNumber="64">
<heading id="D0D181DEC61CFFD114AA41742094D566" bold="true" box="[151,748,1615,1641]" fontSize="11" level="1" pageId="1" pageNumber="64" reason="1">
<emphasis id="B952EAA0C61CFFD114AA41742094D566" bold="true" box="[151,748,1615,1641]" pageId="1" pageNumber="64">
<taxonomicName id="4C264D31C61CFFD114AA417420FAD566" ID-CoL="7976L" authority="Zrelli &amp; Boumaiza" authorityName="Zrelli &amp; Boumaiza" box="[151,642,1615,1641]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B952EAA0C61CFFD114AA417423F7D566" bold="true" box="[151,399,1615,1641]" italics="true" pageId="1" pageNumber="64">Rhithrogena sartorii</emphasis>
Zrelli &amp; Boumaiza
</taxonomicName>
,
<taxonomicNameLabel id="A26157DBC61CFFD116B241742094D566" box="[655,748,1615,1641]" pageId="1" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9936B2C61CFFD114AA41492358D585" blockId="1.[151,748,1615,1674]" box="[151,288,1650,1674]" pageId="1" pageNumber="64">
(
<figureCitation id="131D2A37C61CFFD114A241492294D585" box="[159,236,1650,1674]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="1" pageNumber="64">Figs. 2</figureCitation>
19)
</paragraph>
</subSubSection>
<subSubSection id="C33C6539C61CFFD114AA418120ABD489" pageId="1" pageNumber="64" type="materials_examined">
<paragraph id="8B9936B2C61CFFD114AA41812357D415" blockId="1.[151,1436,1722,2034]" pageId="1" pageNumber="64">
<emphasis id="B952EAA0C61CFFD114AA418123FCD5DC" bold="true" box="[151,388,1722,1747]" pageId="1" pageNumber="64">Material examined.</emphasis>
<typeStatus id="549D8810C61CFFD115B241812383D5DD" box="[399,507,1722,1746]" pageId="1" pageNumber="64" type="holotype">Holotype</typeStatus>
: 1 nymph (preserved in ethanol), Ennour,
<geoCoordinate id="EE125075C61CFFD117D241812615D5DD" box="[1007,1133,1722,1746]" direction="north" orientation="latitude" pageId="1" pageNumber="64" precision="1" value="36.80072">36.80072N</geoCoordinate>
<geoCoordinate id="EE125075C61CFFD110454181269BD5DD" box="[1144,1251,1722,1746]" direction="east" orientation="longitude" pageId="1" pageNumber="64" precision="1" value="8.65871">8.65871E</geoCoordinate>
<specimenCount id="9D20FD3BC61CFFD110D24181273BD5DD" box="[1263,1347,1722,1746]" pageId="1" pageNumber="64" type="generic">
<quantity id="4CDE9B57C61CFFD110D241812747D5DD" box="[1263,1343,1722,1746]" metricMagnitude="2" metricUnit="m" metricValue="4.18" pageId="1" pageNumber="64" unit="m" value="418.0">418 m</quantity>
.
</specimenCount>
,
<date id="FF981072C61CFFD1116E418122AAD5F9" pageId="1" pageNumber="64" value="2010-04-28">28 IV 2010</date>
, S. Zrelli (
<collectionCode id="ED37AE77C61CFFD1157641E523F4D5F9" box="[331,396,1758,1782]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="1" pageNumber="64">MZL</collectionCode>
).
<typeStatus id="549D8810C61CFFD1159D41E42075D5F8" box="[416,525,1759,1783]" pageId="1" pageNumber="64" type="paratype">Paratypes</typeStatus>
: same data as the
<typeStatus id="549D8810C61CFFD116DC41E5213FD5F9" box="[737,839,1758,1782]" pageId="1" pageNumber="64" type="holotype">holotype</typeStatus>
: 30 nymphs,
<specimenCount id="9D20FD3BC61CFFD117E341E52652D5F9" box="[990,1066,1758,1782]" pageId="1" pageNumber="64" type="male">1 male</specimenCount>
subimago,
<specimenCount id="9D20FD3BC61CFFD1109241E52764D5F9" box="[1199,1308,1758,1782]" pageId="1" pageNumber="64" type="female">2 females</specimenCount>
subimagos (
<collectionCode id="ED37AE77C61CFFD114A2403922A6D415" box="[159,222,1794,1818]" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="1" pageNumber="64">MZL</collectionCode>
,
<collectionCode id="ED37AE77C61CFFD114D44039235BD415" box="[233,291,1794,1818]" pageId="1" pageNumber="64">LBE</collectionCode>
).
</paragraph>
<paragraph id="8B9936B2C61CFFD114F8401D20ABD489" blockId="1.[151,1436,1722,2034]" pageId="1" pageNumber="64">
<emphasis id="B952EAA0C61CFFD114F8401D2078D430" bold="true" box="[197,512,1830,1855]" pageId="1" pageNumber="64">Other material examined.</emphasis>
Same locality
<date id="FF981072C61CFFD1168C401D2153D431" box="[689,811,1830,1854]" pageId="1" pageNumber="64" value="2005-05-24">24 V 2005</date>
, 1 nymph,
<date id="FF981072C61CFFD1178F401D264DD431" box="[946,1077,1830,1854]" pageId="1" pageNumber="64" value="2006-04-30">30 IV 2006</date>
, 1 nymph; Berbeg,
<geoCoordinate id="EE125075C61CFFD11123401D27E4D431" box="[1310,1436,1830,1854]" direction="north" orientation="latitude" pageId="1" pageNumber="64" precision="1" value="36.74968">36.74968N</geoCoordinate>
<geoCoordinate id="EE125075C61CFFD114AA4071237DD46D" box="[151,261,1866,1890]" direction="east" orientation="longitude" pageId="1" pageNumber="64" precision="1" value="8.69693">8.69693E</geoCoordinate>
,
<specimenCount id="9D20FD3BC61CFFD1152C4071231AD46D" box="[273,354,1866,1890]" pageId="1" pageNumber="64" type="generic">
<quantity id="4CDE9B57C61CFFD1152C40712326D46D" box="[273,350,1866,1890]" metricMagnitude="2" metricUnit="m" metricValue="5.58" pageId="1" pageNumber="64" unit="m" value="558.0">558 m</quantity>
.
</specimenCount>
,
<date id="FF981072C61CFFD1155340712391D46D" box="[366,489,1866,1890]" pageId="1" pageNumber="64" value="2005-05-21">21 V 2005</date>
, 1 nymph; Rennagha,
<geoCoordinate id="EE125075C61CFFD116D84071211BD46D" box="[741,867,1866,1890]" direction="north" orientation="latitude" pageId="1" pageNumber="64" precision="1" value="36.85997">36.85997N</geoCoordinate>
<geoCoordinate id="EE125075C61CFFD11757407121A1D46D" box="[874,985,1866,1890]" direction="east" orientation="longitude" pageId="1" pageNumber="64" precision="1" value="8.72096">8.72096E</geoCoordinate>
,
<specimenCount id="9D20FD3BC61CFFD117D84071265DD46D" box="[997,1061,1866,1890]" pageId="1" pageNumber="64" type="generic">
<quantity id="4CDE9B57C61CFFD117D840712659D46D" box="[997,1057,1866,1890]" metricMagnitude="1" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="64" unit="m" value="58.0">58 m</quantity>
.
</specimenCount>
<date id="FF981072C61CFFD11011407126C3D46D" box="[1068,1211,1866,1890]" pageId="1" pageNumber="64" value="2005-07-29">29 VII 2005</date>
, 1 nymph; Ghezala,
<geoCoordinate id="EE125075C61CFFD114AA4055236CD489" box="[151,276,1902,1926]" direction="north" orientation="latitude" pageId="1" pageNumber="64" precision="1" value="36.64313">36.64313N</geoCoordinate>
<geoCoordinate id="EE125075C61CFFD11526405523F2D489" box="[283,394,1902,1926]" direction="east" orientation="longitude" pageId="1" pageNumber="64" precision="1" value="8.69852">8.69852E</geoCoordinate>
,
<quantity id="4CDE9B57C61CFFD115A8405523AFD489" box="[405,471,1902,1926]" metricMagnitude="2" metricUnit="m" metricValue="2.29" pageId="1" pageNumber="64" unit="m" value="229.0">229m</quantity>
,
<date id="FF981072C61CFFD115DE40552026D489" box="[483,606,1902,1926]" pageId="1" pageNumber="64" value="2005-05-24">24 V 2005</date>
, 1 nymph.
</paragraph>
</subSubSection>
<subSubSection id="C33C6539C61CFFD414FA40A92032D2C0" lastPageId="4" lastPageNumber="67" pageId="1" pageNumber="64" type="description">
<paragraph id="8B9936B2C61CFFD114FA40A92367D4FD" blockId="1.[151,1436,1722,2034]" pageId="1" pageNumber="64">
<emphasis id="B952EAA0C61CFFD114FA40A923C4D4A4" bold="true" box="[199,444,1938,1963]" pageId="1" pageNumber="64">
Description.
<emphasis id="B952EAA0C61CFFD1155C40A923CDD4A4" bold="true" box="[353,437,1938,1963]" italics="true" pageId="1" pageNumber="64">Nymph</emphasis>
.
</emphasis>
Body length of final instar, excluding caudal filaments, up to
<quantity id="4CDE9B57C61CFFD110B440A92699D4A5" box="[1161,1249,1938,1962]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="1" pageNumber="64" unit="mm" value="4.5">4.5 mm</quantity>
and
<quantity id="4CDE9B57C61CFFD1112640A9270BD4A5" box="[1307,1395,1938,1962]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="64" unit="mm" value="7.5">7.5 mm</quantity>
for male and female nymphs respectively. Cerci, 5.0
<quantity id="4CDE9B57C61CFFD116E9408D2148D4C1" box="[724,816,1974,1998]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="1" pageNumber="64" unit="mm" value="7.3">7.3 mm</quantity>
. Color: body brownish with visible tracheation on the whole body.
</paragraph>
<paragraph id="8B9936B2C61FFFD214F847AC20F8D3F8" blockId="2.[151,1436,151,247]" pageId="2" pageNumber="65">
Head dark brown, subtrapezoidal with convex sides and rounded hind corners (
<figureCitation id="131D2A37C61FFFD2107247AC26EED3A0" box="[1103,1174,151,175]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="2" pageNumber="65">Fig. 2</figureCitation>
). Labrum narrow, with distinct anterior teeth (
<figureCitation id="131D2A37C61FFFD215A8478723A0D3DB" box="[405,472,188,212]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="2" pageNumber="65">Fig. 3</figureCitation>
s), 9 11 comb-shaped bristles on fore margin of maxilla with 6 7 teeth (
<figureCitation id="131D2A37C61FFFD2113E47872731D3DB" box="[1283,1353,188,212]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="2" pageNumber="65">Fig. 4</figureCitation>
). Glossae subovoidal with rounded apex (
<figureCitation id="131D2A37C61FFFD2161E47E4201FD3F8" box="[547,615,223,247]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="2" pageNumber="65">Fig. 5</figureCitation>
s).
</paragraph>
<caption id="DF59663AC61FFFD214AA400B2357D4AD" httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="2" pageNumber="65" targetBox="[167,1383,288,1833]" targetPageId="2">
<paragraph id="8B9936B2C61FFFD214AA400B2357D4AD" blockId="2.[151,1436,1840,1954]" pageId="2" pageNumber="65">
<emphasis id="B952EAA0C61FFFD214AA400B2331D44A" bold="true" box="[151,329,1840,1863]" pageId="2" pageNumber="65">FIGURES 2–10.</emphasis>
<taxonomicName id="4C264D31C61FFFD2156F400B205CD449" box="[338,548,1840,1862]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B952EAA0C61FFFD2156F400B205CD449" box="[338,548,1840,1862]" italics="true" pageId="2" pageNumber="65">Rhithrogena sartorii</emphasis>
</taxonomicName>
<emphasis id="B952EAA0C61FFFD21610400A20F8D449" bold="true" box="[557,640,1841,1862]" pageId="2" pageNumber="65">
<taxonomicNameLabel id="A26157DBC61FFFD21610400A20F8D449" box="[557,640,1841,1862]" pageId="2" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, larva: 2. Head, dorsal. 3s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5s. Labium (dorsal view). 6. Hind femur. 7s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10a. Gill I. 10b. Gill II. 10c. Gill VII.
<taxonomicName id="4C264D31C61FFFD21066405526B7D48C" box="[1115,1231,1901,1923]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B952EAA0C61FFFD21066405526B7D48C" box="[1115,1231,1901,1923]" italics="true" pageId="2" pageNumber="65">R. insularis</emphasis>
</taxonomicName>
, 3i. 5i. 7i.
<taxonomicName id="4C264D31C61FFFD21106405522CED4AD" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="nuragica">
<emphasis id="B952EAA0C61FFFD21106405522CED4AD" italics="true" pageId="2" pageNumber="65">R. nuragica</emphasis>
</taxonomicName>
, 3n. 5n. 7n.
</paragraph>
</caption>
<caption id="DF59663AC61EFFD314AA45112300D150" httpUri="https://zenodo.org/record/206952/files/figure.png" pageId="3" pageNumber="66" targetBox="[198,1417,203,512]" targetPageId="3">
<paragraph id="8B9936B2C61EFFD314AA45112300D150" blockId="3.[151,1436,554,607]" pageId="3" pageNumber="66">
<emphasis id="B952EAA0C61EFFD314AA45112329D14F" bold="true" box="[151,337,554,577]" pageId="3" pageNumber="66">FIGURES 11–13.</emphasis>
<taxonomicName id="4C264D31C61EFFD3156A4511205FD14F" box="[343,551,554,576]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B952EAA0C61EFFD3156A4511205FD14F" box="[343,551,554,576]" italics="true" pageId="3" pageNumber="66">Rhithrogena sartorii</emphasis>
</taxonomicName>
<emphasis id="B952EAA0C61EFFD3161145102004D14F" bold="true" box="[556,636,555,576]" pageId="3" pageNumber="66">
<taxonomicNameLabel id="A26157DBC61EFFD3161145102004D14F" box="[556,636,555,576]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male subimago: 11. Styliger plate and gonopods in ventral view. 12. Penis. 13. Titillator, lateral view.
</paragraph>
</caption>
<caption id="DF59663AC61EFFD314AA4166275AD57C" box="[151,1314,1629,1652]" httpUri="https://zenodo.org/record/206953/files/figure.png" pageId="3" pageNumber="66" targetBox="[151,1436,657,1601]" targetPageId="3">
<paragraph id="8B9936B2C61EFFD314AA4166275AD57C" blockId="3.[151,1314,1629,1652]" box="[151,1314,1629,1652]" pageId="3" pageNumber="66">
<emphasis id="B952EAA0C61EFFD314AA4166232BD57D" bold="true" box="[151,339,1629,1652]" pageId="3" pageNumber="66">FIGURES 14–16.</emphasis>
<taxonomicName id="4C264D31C61EFFD3156741662052D57C" box="[346,554,1629,1651]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B952EAA0C61EFFD3156741662052D57C" box="[346,554,1629,1651]" italics="true" pageId="3" pageNumber="66">Rhithrogena sartorii</emphasis>
</taxonomicName>
<emphasis id="B952EAA0C61EFFD3160D416520F8D57C" bold="true" box="[560,640,1630,1651]" pageId="3" pageNumber="66">
<taxonomicNameLabel id="A26157DBC61EFFD3160D416520F8D57C" box="[560,640,1630,1651]" pageId="3" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, egg: 14. General view. 15. Detail of the micropyle. 16. Polar cap.
</paragraph>
</caption>
<paragraph id="8B9936B2C61EFFD314F841A12007D468" blockId="3.[151,1436,1690,2002]" pageId="3" pageNumber="66">
Dorsal surface of femora of all legs with few spines and an ovoid dark spot well visible on the center of the median pale area (
<figureCitation id="131D2A37C61EFFD31554418423D6D5D8" box="[361,430,1727,1751]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="3" pageNumber="66">Fig. 6</figureCitation>
). Bristles on hind femora very short with diverging margins (
<figureCitation id="131D2A37C61EFFD31050418426CBD5D8" box="[1133,1203,1727,1751]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="3" pageNumber="66">Fig. 7</figureCitation>
s). Tarsal claws with two or three denticles. Lateral sclerites of the first abdominal sternite rectangular with their anterior margin perpendicular to the body axis (
<figureCitation id="131D2A37C61EFFD3158C403C238ED410" box="[433,502,1799,1823]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="3" pageNumber="66">Fig. 8</figureCitation>
). Posterior margin of abdominal tergites regularly dentate in central part, with few supplementary microdenticles (
<figureCitation id="131D2A37C61EFFD315C540112045D44D" box="[504,573,1834,1858]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="3" pageNumber="66">Fig. 9</figureCitation>
). All gill lamellae crenulated (10a, 10b and 10c). Plica on the first gill triangular with slightly concave apex (
<figureCitation id="131D2A37C61EFFD3162F4074201CD468" box="[530,612,1871,1895]" captionStart="FIGURES 2 – 10" captionStartId="2.[151,264,1840,1863]" captionTargetBox="[167,1383,288,1833]" captionTargetId="figure@2.[151,1426,259,1834]" captionTargetPageId="2" captionText="FIGURES 2 – 10. Rhithrogena sartorii sp. nov., larva: 2. Head, dorsal. 3 s. Labrum, dorsal face 4. Comb-shaped setae from maxillary lacinia. 5 s. Labium (dorsal view). 6. Hind femur. 7 s. Bristles from the margin and the central part of hind femur. 8. First abdominal sternite. 9. Posterior margin of tergum V. 10 a. Gill I. 10 b. Gill II. 10 c. Gill VII. R. insularis, 3 i. 5 i. 7 i. R. nuragica, 3 n. 5 n. 7 n." httpUri="https://zenodo.org/record/206951/files/figure.png" pageId="3" pageNumber="66">Fig. 10</figureCitation>
a).
</paragraph>
<paragraph id="8B9936B2C61EFFD414FA40492760D3F8" blockId="3.[151,1436,1690,2002]" lastBlockId="4.[151,1437,151,1149]" lastPageId="4" lastPageNumber="67" pageId="3" pageNumber="66">
<emphasis id="B952EAA0C61EFFD314FA404923F9D484" bold="true" box="[199,385,1906,1931]" italics="true" pageId="3" pageNumber="66">Male subimago.</emphasis>
Body length 9.0 mm, forewing 8.0 mm. Cerci 9.0 mm. General color brown except the thorax, lighter. Eyes darkish brown. Sterna lighter than terga. Fore legs brown, femora with an elongated dark spot. Mid- and hind legs lighter, yellowish brown, and femora with a dark spot more elongated. Styliger plate and forceps brown, posterior margin deeply incised in the middle (
<figureCitation id="131D2A37C619FFD4170D47AC21FBD3A0" box="[816,899,151,175]" captionStart="FIGURES 11 – 13" captionStartId="3.[151,264,554,577]" captionTargetBox="[198,1417,203,512]" captionTargetId="figure@3.[151,1436,193,533]" captionTargetPageId="3" captionText="FIGURES 11 – 13. Rhithrogena sartorii sp. nov., male subimago: 11. Styliger plate and gonopods in ventral view. 12. Penis. 13. Titillator, lateral view." httpUri="https://zenodo.org/record/206952/files/figure.png" pageId="4" pageNumber="67">Fig. 11</figureCitation>
). Forceps uniformly brown, 3 segmented with a well defined basal segment, the 1st segment the longest, the 2nd and the 3rd equal in length. Genitalia as in
<figureCitation id="131D2A37C619FFD4117B478727E0D3DB" box="[1350,1432,188,212]" captionStart="FIGURES 11 – 13" captionStartId="3.[151,264,554,577]" captionTargetBox="[198,1417,203,512]" captionTargetId="figure@3.[151,1436,193,533]" captionTargetPageId="3" captionText="FIGURES 11 – 13. Rhithrogena sartorii sp. nov., male subimago: 11. Styliger plate and gonopods in ventral view. 12. Penis. 13. Titillator, lateral view." httpUri="https://zenodo.org/record/206952/files/figure.png" pageId="4" pageNumber="67">Fig. 12</figureCitation>
. Titillators with apical margin bearing 3 4 teeth with 3 subapical teeth visible on their surface (
<figureCitation id="131D2A37C619FFD4108547E42773D3F8" box="[1208,1291,223,247]" captionStart="FIGURES 11 – 13" captionStartId="3.[151,264,554,577]" captionTargetBox="[198,1417,203,512]" captionTargetId="figure@3.[151,1436,193,533]" captionTargetPageId="3" captionText="FIGURES 11 – 13. Rhithrogena sartorii sp. nov., male subimago: 11. Styliger plate and gonopods in ventral view. 12. Penis. 13. Titillator, lateral view." httpUri="https://zenodo.org/record/206952/files/figure.png" pageId="4" pageNumber="67">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8B9936B2C619FFD414FA46382713D230" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="67">
<emphasis id="B952EAA0C619FFD414FA463823E5D213" bold="true" box="[199,413,259,284]" italics="true" pageId="4" pageNumber="67">Female subimago.</emphasis>
Body length 7.3 8.0 mm, forewings 5.7 7.0 mm. General color yellowish brown. Thorax brown, abdomen with color pattern similar to male. All legs with elongated spots on the dorsal side of femora.
</paragraph>
<paragraph id="8B9936B2C619FFD414FA46772116D2A3" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="67">
<emphasis id="B952EAA0C619FFD414FA46772370D26C" bold="true" box="[199,264,332,355]" italics="true" pageId="4" pageNumber="67">Eggs.</emphasis>
Length: 180 200 µm, width: 150 170 µm. General shape ovoid (
<figureCitation id="131D2A37C619FFD4103C4677262ED26B" box="[1025,1110,332,356]" captionStart="FIGURES 14 – 16" captionStartId="3.[151,264,1629,1652]" captionTargetBox="[151,1436,657,1601]" captionTargetId="figure@3.[151,1436,657,1608]" captionTargetPageId="3" captionText="FIGURES 14 – 16. Rhithrogena sartorii sp. nov., egg: 14. General view. 15. Detail of the micropyle. 16. Polar cap." httpUri="https://zenodo.org/record/206953/files/figure.png" pageId="4" pageNumber="67">Fig. 14</figureCitation>
). Polar cap at one pole with numerous adhesive elements (KCTs). Micropyle with well developed margin (
<figureCitation id="131D2A37C619FFD4103C4654262CD288" box="[1025,1108,367,391]" captionStart="FIGURES 14 – 16" captionStartId="3.[151,264,1629,1652]" captionTargetBox="[151,1436,657,1601]" captionTargetId="figure@3.[151,1436,657,1608]" captionTargetPageId="3" captionText="FIGURES 14 – 16. Rhithrogena sartorii sp. nov., egg: 14. General view. 15. Detail of the micropyle. 16. Polar cap." httpUri="https://zenodo.org/record/206953/files/figure.png" pageId="4" pageNumber="67">Fig. 15</figureCitation>
). Chorionic surface with several KCTs and with microgranules irregularly arranged (
<figureCitation id="131D2A37C619FFD4173046AF2119D2A3" box="[781,865,404,428]" captionStart="FIGURES 14 – 16" captionStartId="3.[151,264,1629,1652]" captionTargetBox="[151,1436,657,1601]" captionTargetId="figure@3.[151,1436,657,1608]" captionTargetPageId="3" captionText="FIGURES 14 – 16. Rhithrogena sartorii sp. nov., egg: 14. General view. 15. Detail of the micropyle. 16. Polar cap." httpUri="https://zenodo.org/record/206953/files/figure.png" pageId="4" pageNumber="67">Fig. 16</figureCitation>
).
</paragraph>
<paragraph id="8B9936B2C619FFD414FA468C2032D2C0" blockId="4.[151,1437,151,1149]" box="[199,586,439,463]" pageId="4" pageNumber="67">Male and female imagos unknown</paragraph>
</subSubSection>
<subSubSection id="C33C6539C619FFD514FA46E726ACD3DB" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67" type="etymology">
<paragraph id="8B9936B2C619FFD414FA46E72164D118" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="67">
<emphasis id="B952EAA0C619FFD414FA46E72336D2FB" bold="true" box="[199,334,476,500]" pageId="4" pageNumber="67">Etymology.</emphasis>
The species is named to honour Dr. Michel Sartori (Museum of zoology, Lausanne,
<collectingCountry id="F3317622C619FFD4113B46E727E8D2FB" box="[1286,1424,476,500]" name="Switzerland" pageId="4" pageNumber="67">Switzerland</collectingCountry>
), who is a long-time investigator of the genus
<taxonomicName id="4C264D31C619FFD416B7453B216DD117" box="[650,789,512,536]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="B952EAA0C619FFD416B7453B216DD117" box="[650,789,512,536]" italics="true" pageId="4" pageNumber="67">Rhithrogena</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9936B2C619FFD414FA4518216CD053" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="67">
<emphasis id="B952EAA0C619FFD414FA45182345D133" bold="true" box="[199,317,547,572]" pageId="4" pageNumber="67">Affinities.</emphasis>
<taxonomicName id="4C264D31C619FFD4157E451F2050D133" box="[323,552,548,572]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B952EAA0C619FFD4157E451F2050D133" box="[323,552,548,572]" italics="true" pageId="4" pageNumber="67">Rhithrogena sartorii</emphasis>
</taxonomicName>
<emphasis id="B952EAA0C619FFD41612451F20F0D133" bold="true" box="[559,648,548,572]" pageId="4" pageNumber="67">
<taxonomicNameLabel id="A26157DBC619FFD41612451F20F0D133" box="[559,648,548,572]" pageId="4" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
seems to belong to the
<taxonomicName id="4C264D31C619FFD417B2451F2189D133" box="[911,1009,548,572]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B952EAA0C619FFD417B2451F2189D133" box="[911,1009,548,572]" italics="true" pageId="4" pageNumber="67">insularis</emphasis>
</taxonomicName>
-group which included up to now only
<taxonomicName id="4C264D31C619FFD414AA45722075D150" authority="Esben-Petersen 1913" authorityName="Esben-Petersen" authorityYear="1913" box="[151,525,583,608]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B952EAA0C619FFD414AA45722361D16F" box="[151,281,584,608]" italics="true" pageId="4" pageNumber="67">R. insularis</emphasis>
Esben-Petersen 1913
</taxonomicName>
and
<taxonomicName id="4C264D31C619FFD4167845722109D150" authority="Belfiore 1987" authorityName="Belfiore" authorityYear="1987" box="[581,881,583,608]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="nuragica">
<emphasis id="B952EAA0C619FFD416784572202ED16F" box="[581,598,585,608]" italics="true" pageId="4" pageNumber="67">R</emphasis>
.
<emphasis id="B952EAA0C619FFD41658457220B2D16F" box="[613,714,585,608]" italics="true" pageId="4" pageNumber="67">nuragica</emphasis>
<bibRefCitation id="EFB74B43C619FFD416EE457C2109D150" author="Belfiore" box="[723,881,583,607]" pageId="4" pageNumber="67" refString="Belfiore, C. (1987) Heptageniidae from Corsica and Sardinia. Rhithrogena nuragica n. sp., R. eatoni Esben Petersen 1912, and R. insularis Esben Petersen 1913 (Ephemeroptera). Annales de Limnologie, 23 (2), 87 94." type="journal article" year="1987">Belfiore 1987</bibRefCitation>
</taxonomicName>
. According to
<bibRefCitation id="EFB74B43C619FFD41024457C26BFD150" author="Belfiore" box="[1049,1223,583,607]" pageId="4" pageNumber="67" refString="Belfiore, C. (1987) Heptageniidae from Corsica and Sardinia. Rhithrogena nuragica n. sp., R. eatoni Esben Petersen 1912, and R. insularis Esben Petersen 1913 (Ephemeroptera). Annales de Limnologie, 23 (2), 87 94." type="journal article" year="1987">Belfiore (1987)</bibRefCitation>
and
<bibRefCitation id="EFB74B43C619FFD410C2457C2299D18B" pageId="4" pageNumber="67" refString="Belfiore, C., Picariello, O., Scillitani, G. &amp; Cretella, M. (1992) Genetic divergence between two insular species of the genus Rhithrogena (Ephemeroptera, Heptageniidae). Fragmenta Entomologica, 23, 235 242." type="journal article">
Belfiore
<emphasis id="B952EAA0C619FFD4115E457227E4D16F" box="[1379,1436,584,608]" italics="true" pageId="4" pageNumber="67">et al.</emphasis>
(1992)
</bibRefCitation>
the nymphs of both species shows a unique combination of characters: 1) lateral sclerites of the first abdominal sternite with anterior margin perpendicular to the body axis; 2) all gills crenulated; 3) a dark spot on the dorsal side of femora; 4) plica on the first gill triangular; and 5) the presence of well-developed tubercules on egg chorion. First 3 characters are also shared by species of the
<emphasis id="B952EAA0C619FFD416E345E321E5D1FF" box="[734,925,728,752]" italics="true" pageId="4" pageNumber="67">hybrida/hercynia</emphasis>
species group, but the last two clearly differ from it. In the present state of our knowledge,
<taxonomicName id="4C264D31C619FFD4169D45C6216BD01B" box="[672,787,765,788]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="sartorii">
<emphasis id="B952EAA0C619FFD4169D45C6216BD01B" box="[672,787,765,788]" italics="true" pageId="4" pageNumber="67">R. sartorii</emphasis>
</taxonomicName>
is morphologically close to the species of the
<taxonomicName id="4C264D31C619FFD4112645C627E4D01B" box="[1307,1436,764,788]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B952EAA0C619FFD4112645C627E4D01B" box="[1307,1436,764,788]" italics="true" pageId="4" pageNumber="67">R. insularis</emphasis>
</taxonomicName>
group, but genetic data based on both mitochondrial and nuclear markers are congruent and place
<taxonomicName id="4C264D31C619FFD410C9441A2711D037" box="[1268,1385,801,824]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="sartorii">
<emphasis id="B952EAA0C619FFD410C9441A2711D037" box="[1268,1385,801,824]" italics="true" pageId="4" pageNumber="67">R. sartorii</emphasis>
</taxonomicName>
in a cluster quite distant from them (L. Vuataz, comm. pers.).
</paragraph>
<paragraph id="8B9936B2C619FFD414F8445C2712D0C8" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="67">
When compared to the species found in Morroco,
<taxonomicName id="4C264D31C619FFD41731445221F9D08F" box="[780,897,873,896]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="sartorii">
<emphasis id="B952EAA0C619FFD41731445221F9D08F" box="[780,897,873,896]" italics="true" pageId="4" pageNumber="67">R. sartorii</emphasis>
</taxonomicName>
clearly differs from
<taxonomicName id="4C264D31C619FFD4104E4452269BD08F" box="[1139,1251,873,896]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="mariae">
<emphasis id="B952EAA0C619FFD4104E4452269BD08F" box="[1139,1251,873,896]" italics="true" pageId="4" pageNumber="67">R. mariae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C264D31C619FFD41122445227E4D08F" box="[1311,1436,872,896]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="ryszardi">
<emphasis id="B952EAA0C619FFD41122445227E4D08F" box="[1311,1436,872,896]" italics="true" pageId="4" pageNumber="67">R. ryszardi</emphasis>
</taxonomicName>
which belong to the
<emphasis id="B952EAA0C619FFD4154244B72069D0AB" box="[383,529,908,932]" italics="true" pageId="4" pageNumber="67">semicolorata</emphasis>
and the
<emphasis id="B952EAA0C619FFD4164944B62094D0AB" box="[628,748,909,932]" italics="true" pageId="4" pageNumber="67">germanica</emphasis>
species group respectively, which gills II VII are not crenulated. It also differs from
<emphasis id="B952EAA0C619FFD41589448A20F0D0C7" box="[436,648,944,968]" italics="true" pageId="4" pageNumber="67">
<taxonomicName id="4C264D31C619FFD41589448A2058D0C7" box="[436,544,944,968]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="ourika">R. ourika</taxonomicName>
,
<taxonomicName id="4C264D31C619FFD41616448A20F0D0C7" box="[555,648,944,968]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="ayadi">R. ayadi</taxonomicName>
</emphasis>
and
<taxonomicName id="4C264D31C619FFD41683448A21FAD0C7" box="[702,898,944,968]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="giudicelliorum">
<emphasis id="B952EAA0C619FFD41683448A21FAD0C7" box="[702,898,944,968]" italics="true" pageId="4" pageNumber="67">R. giudicelliorum</emphasis>
</taxonomicName>
, mainly by the shape of the male titillators.
</paragraph>
<paragraph id="8B9936B2C619FFD414F844EF26C6D773" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="67">
The main diagnostic nymphal characters are described and compared with the two
<taxonomicName id="4C264D31C619FFD4104A44EF26A1D0E3" box="[1143,1241,980,1004]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B952EAA0C619FFD4104A44EF26A1D0E3" box="[1143,1241,980,1004]" italics="true" pageId="4" pageNumber="67">insularis</emphasis>
</taxonomicName>
-group species in
<tableCitation id="C6A40309C619FFD414AA44CC2293D700" box="[151,235,1015,1039]" captionStart="TABLE 1" captionStartId="4.[151,235,1191,1212]" captionTargetBox="[151,1436,1384,1889]" captionTargetPageId="4" captionText="TABLE 1. Some descriptive characters of Rhithrogena sartorii sp. nov. compared with two related species: R. nuragica and R." httpUri="http://table.plazi.org/id/DF59663AC619FFD414AA439C27E4D7B2" pageId="4" pageNumber="67" tableUuid="DF59663AC619FFD414AA439C27E4D7B2">Table 1</tableCitation>
. The nymph of the new species can be distinguished especially by the number of comb-shaped bristles and the number of teeth on the comb-shaped bristles at the tip of lacinia. The shape of the chorionic tubercles is somewhat intermediate between those of
<taxonomicName id="4C264D31C619FFD4160F437A20CCD757" box="[562,692,1088,1112]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B952EAA0C619FFD4160F437A20CCD757" box="[562,692,1088,1112]" italics="true" pageId="4" pageNumber="67">R. insularis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C264D31C619FFD416D1437A210AD757" box="[748,882,1089,1112]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="nuragica">
<emphasis id="B952EAA0C619FFD416D1437A210AD757" box="[748,882,1089,1112]" italics="true" pageId="4" pageNumber="67">R. nuragica</emphasis>
</taxonomicName>
. KCTs lack the finger-like processes found in
<taxonomicName id="4C264D31C619FFD411B9437A2281D773" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B952EAA0C619FFD411B9437A2281D773" italics="true" pageId="4" pageNumber="67">R. insularis</emphasis>
</taxonomicName>
, and the arrangement of the KCTs is less regular than in
<taxonomicName id="4C264D31C619FFD4174A435E26C1D773" authority="Belfiore, 1987" authorityName="Belfiore" authorityYear="1987" box="[887,1209,1124,1148]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="nuragica">
<emphasis id="B952EAA0C619FFD4174A435E2183D773" box="[887,1019,1125,1148]" italics="true" pageId="4" pageNumber="67">R. nuragica</emphasis>
(
<bibRefCitation id="EFB74B43C619FFD41037435F26C9D773" author="Belfiore" box="[1034,1201,1124,1148]" pageId="4" pageNumber="67" refString="Belfiore, C. (1987) Heptageniidae from Corsica and Sardinia. Rhithrogena nuragica n. sp., R. eatoni Esben Petersen 1912, and R. insularis Esben Petersen 1913 (Ephemeroptera). Annales de Limnologie, 23 (2), 87 94." type="journal article" year="1987">Belfiore, 1987</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="DF59663AC619FFD414AA439C27E4D7B2" ID-Table-UUID="DF59663AC619FFD414AA439C27E4D7B2" box="[151,1436,1190,1213]" httpUri="http://table.plazi.org/id/DF59663AC619FFD414AA439C27E4D7B2" pageId="4" pageNumber="67" targetBox="[151,1436,1384,1889]" targetIsTable="true" targetPageId="4">
<paragraph id="8B9936B2C619FFD414AA439C27E4D7B2" blockId="4.[151,1436,1190,1365]" box="[151,1436,1190,1213]" pageId="4" pageNumber="67">
<emphasis id="B952EAA0C619FFD414AA439C237CD7B3" bold="true" box="[151,260,1191,1212]" pageId="4" pageNumber="67">TABLE 1.</emphasis>
Some descriptive characters of
<taxonomicName id="4C264D31C619FFD4167A439C216FD7B2" box="[583,791,1191,1213]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="sartorii" status="sp. nov.">
<emphasis id="B952EAA0C619FFD4167A439C216FD7B2" box="[583,791,1191,1213]" italics="true" pageId="4" pageNumber="67">Rhithrogena sartorii</emphasis>
</taxonomicName>
<emphasis id="B952EAA0C619FFD41720439C2115D7B3" bold="true" box="[797,877,1191,1212]" pageId="4" pageNumber="67">
<taxonomicNameLabel id="A26157DBC619FFD41720439C2115D7B3" box="[797,877,1191,1212]" pageId="4" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
compared with two related species:
<taxonomicName id="4C264D31C619FFD410E34393272DD7B2" box="[1246,1365,1192,1213]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="nuragica">
<emphasis id="B952EAA0C619FFD410E34393272DD7B2" box="[1246,1365,1192,1213]" italics="true" pageId="4" pageNumber="67">R. nuragica</emphasis>
</taxonomicName>
and R.
</paragraph>
</caption>
<paragraph id="8B9936B2C619FFD414AA43FE228DD7D4" blockId="4.[151,1436,1190,1365]" box="[151,245,1221,1243]" pageId="4" pageNumber="67">
<taxonomicName id="4C264D31C619FFD414AA43FE2288D7D4" box="[151,240,1221,1243]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="B952EAA0C619FFD414AA43FE2288D7D4" box="[151,240,1221,1243]" italics="true" pageId="4" pageNumber="67">insularis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9936B2C619FFD414A243CD2716D624" blockId="4.[151,1436,1190,1365]" pageId="4" pageNumber="67">
Character
<emphasis id="B952EAA0C619FFD4165E43CD270CD602" box="[611,1396,1270,1293]" italics="true" pageId="4" pageNumber="67">
<taxonomicName id="4C264D31C619FFD4165E43CD2150D604" box="[611,808,1270,1291]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="sartorii">Rhithrogena sartorii</taxonomicName>
<taxonomicName id="4C264D31C619FFD4175043CC2635D602" box="[877,1101,1271,1293]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="nuragica">Rhithrogena nuragica</taxonomicName>
<taxonomicName id="4C264D31C619FFD410AA43CC270CD602" box="[1175,1396,1271,1293]" class="Insecta" family="Heptageniidae" genus="Rhithrogena" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="insularis" status="sp. nov.">Rhithrogena insularis</taxonomicName>
</emphasis>
<emphasis id="B952EAA0C619FFD4165E422F20CCD626" bold="true" box="[611,692,1300,1321]" pageId="4" pageNumber="67">
<taxonomicNameLabel id="A26157DBC619FFD4165E422F20CCD626" box="[611,692,1300,1321]" pageId="4" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<bibRefCitation id="EFB74B43C619FFD41750422E2181D624" author="Belfiore" box="[877,1017,1301,1323]" pageId="4" pageNumber="67" refString="Belfiore, C. (1987) Heptageniidae from Corsica and Sardinia. Rhithrogena nuragica n. sp., R. eatoni Esben Petersen 1912, and R. insularis Esben Petersen 1913 (Ephemeroptera). Annales de Limnologie, 23 (2), 87 94." type="journal article" year="1987">Belfiore 1987</bibRefCitation>
Esben-Peterson 1913
</paragraph>
<paragraph id="8B9936B2C619FFD514A2420426ACD3DB" blockId="4.[151,1436,1190,1365]" lastBlockId="5.[151,1435,151,212]" lastPageId="5" lastPageNumber="68" pageId="4" pageNumber="67">
Mean body length (mm) 6 7,5 6,8
<emphasis id="B952EAA0C619FFD414FA409C2353D4B0" bold="true" box="[199,299,1959,1983]" pageId="4" pageNumber="67">Ecology.</emphasis>
Nymphs of the new species were collected in four different sites in strong current of streams and riffles of rivers with cobble substrates. Streams ranged in width from
<quantity id="4CDE9B57C619FFD417BF40F121D4D4ED" box="[898,940,1994,2018]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="67" unit="m" value="4.0">4 m</quantity>
(Berbeg) to
<quantity id="4CDE9B57C619FFD4100440F12609D4ED" box="[1081,1137,1994,2018]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="67" unit="m" value="10.0">10 m</quantity>
(Ennour), and water depth not exceeding
<quantity id="4CDE9B57C618FFD5157D47AC23F1D3A0" box="[320,393,151,175]" metricMagnitude="-1" metricUnit="m" metricValue="6.5" pageId="5" pageNumber="68" unit="cm" value="65.0">65 cm</quantity>
at all collecting sites. The very low mineralization of water (&lt;0.15 PSU) can explain the absence of the new species in the other streams of Northern
<collectingCountry id="F3317622C618FFD51704478721F4D3DB" box="[825,908,188,212]" name="Tunisia" pageId="5" pageNumber="68">Tunisia</collectingCountry>
which are more mineralized.
</paragraph>
</subSubSection>
</treatment>
</document>
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