415 lines
69 KiB
XML
415 lines
69 KiB
XML
<document id="7B2A66C0E129DAE62C53C69120AAE98D" ID-DOI="10.5281/zenodo.186465" ID-GBIF-Dataset="7ee3040e-29c4-4368-8915-ba2f7d6b14fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="186465" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461052118242" checkinUser="plazi" docAuthor="Vargovitsh, Robert S." docDate="2009" docId="BA7487DEEF60FF9AFF28418FFE4509E5" docLanguage="en" docName="zt02047p047.pdf" docOrigin="Zootaxa 2047" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pygmarrhopalites pseudoprincipalis Vargovitsh, 2009, sp. nov." docType="treatment" docUuid="2EFB7950-0501-45D2-9E87-85EB5BCEFB0F" docUuidSource="ZooBank" docVersion="12" lastPageNumber="36" masterDocId="464DFFA6EF7EFFB9FFBF4746FFA00F77" masterDocTitle="New Cave Arrhopalitidae (Collembola: Symphypleona) from the Crimea (Ukraine)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="31" updateTime="1698579300348" updateUser="plazi">
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<mods:mods id="488468D4A9112B66C834E106FC288D06" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="79F08DA309CE949ED9FADCFDCF9EEF99">
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<mods:title id="F95F0A46736DC58DDAFE8F37B1C6B7BD">New Cave Arrhopalitidae (Collembola: Symphypleona) from the Crimea (Ukraine)</mods:title>
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</mods:titleInfo>
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<mods:name id="30683DC33770479CD866F9AB06AAD7D1" type="personal">
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<mods:role id="40B94DC7036B76C91B1572A9E6347FB2">
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<mods:roleTerm id="2E36C6723130038B5E408DFAF78BFF9A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F6642089C95615A260D93F6E6F9DB4A3">Vargovitsh, Robert S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="9C245BAD32001B4B1D705CA1925F0F6E">text</mods:typeOfResource>
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<mods:titleInfo id="C6FF990D97556E1A1AA5DCEC160EC8AA">
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<mods:title id="63533F139ADB2D8CD5B604CCDDF7E6DE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="60EBEC25ED7480B6AE841790C3801F9D">
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<mods:date id="2EFD9F2BA5855380E9C798EC7B080536">2009</mods:date>
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<mods:detail id="FE2B873CE6536F8C85A1AE2E10AA3ADD" type="volume">
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<mods:number id="BC9214DF853B7771D28E18ADEAAC8795">2047</mods:number>
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<mods:start id="5313A27845C22582D5EBAF3670E59004">1</mods:start>
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<mods:classification id="D3FF309560D678EEF918ECF30F70B9C3">journal article</mods:classification>
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<mods:identifier id="A54A3E50C6FF82D854B54B20ED2563E1" type="DOI">10.5281/zenodo.186465</mods:identifier>
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<mods:identifier id="AB8CAE20AC7208AC756DD22C009A8BE9" type="GBIF-Dataset">7ee3040e-29c4-4368-8915-ba2f7d6b14fc</mods:identifier>
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<mods:identifier id="79702808854A613C1D3F0247EF00433D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8ADD6531A6266D812254F8CACC6719C8" type="Zenodo-Dep">186465</mods:identifier>
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<mods:identifier id="BCF514EF6091EE366AFB80512EE9012F" type="ZooBank">9232BEFF-077C-4C10-8E17-85CBA2AD1428</mods:identifier>
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</mods:mods>
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<treatment id="BA7487DEEF60FF9AFF28418FFE4509E5" ID-DOI="http://doi.org/10.5281/zenodo.5618666" ID-GBIF-Taxon="119612339" ID-Zenodo-Dep="5618666" LSID="urn:lsid:zoobank.org:act:2EFB7950-0501-45D2-9E87-85EB5BCEFB0F" httpUri="http://treatment.plazi.org/id/BA7487DEEF60FF9AFF28418FFE4509E5" lastPageId="35" lastPageNumber="36" pageId="30" pageNumber="31">
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<subSubSection id="7AC76543EF60FFA7FF28418FFE3C0871" pageId="30" pageNumber="31" type="nomenclature">
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<paragraph id="326236C8EF60FFA7FF28418FFD0F0995" blockId="30.[151,687,1736,1798]" box="[151,687,1736,1763]" pageId="30" pageNumber="31">
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<heading id="692A81A4EF60FFA7FF28418FFD0F0995" bold="true" box="[151,687,1736,1763]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
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<emphasis id="00A9EADAEF60FFA7FF28418FFD0F0995" bold="true" box="[151,687,1736,1763]" pageId="30" pageNumber="31">
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<taxonomicName id="F5DD4D4BEF60FFA7FF28418FFDEA0994" box="[151,586,1737,1763]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="pseudoprincipalis" status="sp. nov.">
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<emphasis id="00A9EADAEF60FFA7FF28418FFDEA0994" bold="true" box="[151,586,1737,1763]" italics="true" pageId="30" pageNumber="31">Pygmarrhopalites pseudoprincipalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1B9A57A1EF60FFA7FDED418EFD0F0995" box="[594,687,1736,1762]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="326236C8EF60FFA7FF2841AAFE3C0871" blockId="30.[151,687,1736,1798]" box="[151,412,1772,1798]" pageId="30" pageNumber="31">
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<figureCitation id="AAE62A4DEF60FFA7FF2841AAFE990871" box="[151,313,1772,1798]" captionStart-0="FIGURES 118 – 131" captionStart-1="FIGURES 132 – 136" captionStart-2="FIGURES 137 – 143" captionStartId-0="32.[151,272,1873,1897]" captionStartId-1="33.[151,269,1721,1745]" captionStartId-2="34.[151,270,1942,1966]" captionTargetBox-0="[154,1433,190,1872]" captionTargetBox-1="[164,1405,190,1678]" captionTargetBox-2="[191,1388,205,1941]" captionTargetId-0="figure@32.[151,1436,145,1907]" captionTargetId-1="figure@33.[151,1436,158,1747]" captionTargetId-2="figure@34.[191,1398,157,1949]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-2="34" captionText-0="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." captionText-1="FIGURES 132 – 136. Pygmarrhopalites pseudoprincipalis sp. nov.: 132 – 134, chaetotaxy of antennal segments: Ant I – II, anterior view (132), Ant III, posterior view (133), Ant IV, anterior view (134); 135 – 136, chaetotaxy of Abd VI: female (135), male (136)." captionText-2="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri-0="https://zenodo.org/record/186476/files/figure.png" httpUri-1="https://zenodo.org/record/186477/files/figure.png" httpUri-2="https://zenodo.org/record/186478/files/figure.png" pageId="30" pageNumber="31">Figs 118–143</figureCitation>
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,
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<tableCitation id="7F5F0373EF60FFA7FEFA41AAFE3C0871" box="[325,412,1772,1798]" captionStart="TABLE 3" captionStartId="24.[151,239,577,601]" captionText="TABLE 3. Dens chaetotaxy of Pygmarrhopalites tauricus sp. nov., P. kaprusi sp. nov., P. pseudoprincipalis sp. nov. and P. principalis skelicus ssp. nov.; + = simple seta; S = spine-like; s = weakly spine-like; v. p. = varying position; in brackets = variability." pageId="30" pageNumber="31">Table 3</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="7AC76543EF60FFA6FF28407EFD660E51" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="diagnosis">
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<paragraph id="326236C8EF60FFA6FF28407EFD660E51" blockId="30.[151,1437,1848,2033]" lastBlockId="31.[151,1437,152,2034]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
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<emphasis id="00A9EADAEF60FFA7FF28407EFEB90825" bold="true" box="[151,281,1848,1874]" pageId="30" pageNumber="31">Diagnosis.</emphasis>
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Body with brownish spotted pigmentation, eyes pigmented; trichobothrium B somewhat out of AC (ABC=~160o) and AB≤BC; mesothorax with 2 + 2 thickened or spiny setae; posterior dorsal setae of great abdomen distinctly shorter than hind claw; antenna in females 1.6 times as long as head; Ant IV with 5–6 subsegments; head with 4 + 4 weakly spine-like setae; all claws with inner tooth, mid and hind claw with tunica; empodia with apical filament overtopping tip of claw, fore and mid empodia with, and hind–without corner tooth; trochanter II with 5 setae; tenaculum with 2 setulae; manubrium with 7 + 7 setae; dens with 5 spines or spiny setae, Ie very strong and looks somewhat articulated, ventrally with 3, 2, 1, 1 setae; tip of mucro narrow and almost pointed; small abdomen with distinctly broadened circumanal setae and seta ms1 not forked; appendices anales strongly palmate.
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</paragraph>
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</subSubSection>
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<subSubSection id="7AC76543EF61FFA6FF7A4674FB0B0D43" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="326236C8EF61FFA6FF7A4674FB7B0EED" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
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<emphasis id="00A9EADAEF61FFA6FF7A4674FE600E3B" bold="true" box="[197,448,306,332]" pageId="31" pageNumber="32">Material examined.</emphasis>
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<typeStatus id="ED66886AEF61FFA6FE764674FD990E3B" box="[457,569,306,332]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
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on slide (C-266-46): female,
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<collectingCountry id="4ACA7658EF61FFA6FC1A4674FBA90E3B" box="[933,1033,306,332]" name="Ukraine" pageId="31" pageNumber="32">Ukraine</collectingCountry>
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, Crimea, Dolgorukovsky Massif, Kizil-Coba Cave (= Krasnaya Cave),
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<date id="46631008EF61FFA6FDE3461FFD460E04" box="[604,742,345,371]" pageId="31" pageNumber="32" value="1998-12-08">12.08.1998</date>
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.
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<typeStatus id="ED66886AEF61FFA6FD4D461FFCC60E04" box="[754,870,345,371]" pageId="31" pageNumber="32" type="paratype">Paratypes</typeStatus>
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on slides:
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<specimenCount id="24DBFD41EF61FFA6FC56461FFBE90E04" box="[1001,1097,345,371]" pageId="31" pageNumber="32" type="male">5 males</specimenCount>
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,
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<specimenCount id="24DBFD41EF61FFA6FBE9461FFB7B0E04" box="[1110,1243,345,371]" pageId="31" pageNumber="32" type="female">11 females</specimenCount>
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,
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<specimenCount id="24DBFD41EF61FFA6FB58461FFA8E0E04" box="[1255,1326,345,371]" pageId="31" pageNumber="32" type="generic">6 juv.</specimenCount>
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, together with
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<typeStatus id="ED66886AEF61FFA6FF6D46C6FE9C0EED" box="[210,316,384,410]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
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.
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<typeStatus id="ED66886AEF61FFA6FEF846C6FE140EED" box="[327,436,384,410]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
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and 20
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<typeStatus id="ED66886AEF61FFA6FDAD46C6FD220EED" box="[530,642,384,410]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
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are preserved in SIZNAS, 2
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<typeStatus id="ED66886AEF61FFA6FC6846C6FBE70EED" box="[983,1095,384,410]" pageId="31" pageNumber="32" type="paratype">paratypes</typeStatus>
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in
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<collectionCode id="54CCAE0DEF61FFA6FBD346C6FB770EED" box="[1132,1239,384,410]" pageId="31" pageNumber="32">SNHML</collectionCode>
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.
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</paragraph>
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<paragraph id="326236C8EF61FFA6FF7A46E0FB0B0D43" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
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<emphasis id="00A9EADAEF61FFA6FF7A46E0FE230EB7" bold="true" box="[197,387,422,448]" pageId="31" pageNumber="32">Other material</emphasis>
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(on slides).
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<emphasis id="00A9EADAEF61FFA6FDA746E0FD780EB7" bold="true" box="[536,728,422,448]" pageId="31" pageNumber="32">Karabi Massif:</emphasis>
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<specimenCount id="24DBFD41EF61FFA6FD5F46E0FCF60EB7" box="[736,854,422,448]" pageId="31" pageNumber="32" type="female">2 females</specimenCount>
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, Egiz-Tinakh-3 Cave (=Viola Cave),
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<date id="46631008EF61FFA6FAAC46E0FA370EB7" box="[1299,1431,422,448]" pageId="31" pageNumber="32" value="1993-08-07">7.viii.1993</date>
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; female, juv., Gvozdetskogo Cave (-
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<quantity id="F5259B2DEF61FFA6FD82468BFDCE0E90" box="[573,622,461,487]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="m" value="5.0">5 m</quantity>
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; -
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<quantity id="F5259B2DEF61FFA6FD3B468BFD750E90" box="[644,725,461,487]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="m" value="150.0">150 m</quantity>
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),
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<date id="46631008EF61FFA6FD54468BFCCF0E90" box="[747,879,461,487]" pageId="31" pageNumber="32" value="2003-08-09">9.viii.2003</date>
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; female, juv., Molodezhnaya Cave (-
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<quantity id="F5259B2DEF61FFA6FA9A468BFAC70E90" box="[1317,1383,461,487]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="32" unit="m" value="20.0">20 m</quantity>
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; -
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<quantity id="F5259B2DEF61FFA6FAC3468BFF110D79" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="m" value="50.0">50 m</quantity>
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),
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<date id="46631008EF61FFA6FF7846B2FEF70D79" box="[199,343,500,526]" pageId="31" pageNumber="32" value="2003-08-11">11.viii.2003</date>
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;
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<specimenCount id="24DBFD41EF61FFA6FEDB46B2FE620D79" box="[356,450,500,526]" pageId="31" pageNumber="32" type="male">4 males</specimenCount>
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,
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<specimenCount id="24DBFD41EF61FFA6FE7146B2FD980D79" box="[462,568,500,526]" pageId="31" pageNumber="32" type="female">1 female</specimenCount>
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,
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<specimenCount id="24DBFD41EF61FFA6FDFB46B2FD290D79" box="[580,649,500,526]" pageId="31" pageNumber="32" type="generic">3 juv.</specimenCount>
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, Monastyr’ Chokrak Cave (-
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<quantity id="F5259B2DEF61FFA6FC6446B2FB8C0D79" box="[987,1068,500,526]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="m" value="100.0">100 m</quantity>
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),
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<date id="46631008EF61FFA6FBFD46B2FB650D79" box="[1090,1221,500,526]" pageId="31" pageNumber="32" value="2003-08-06">6.viii.2003</date>
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||
;
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||
<specimenCount id="24DBFD41EF61FFA6FB6D46B2FAE60D79" box="[1234,1350,500,526]" pageId="31" pageNumber="32" type="female">6 females</specimenCount>
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||
,
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||
<specimenCount id="24DBFD41EF61FFA6FAEC46B2FA380D79" box="[1363,1432,500,526]" pageId="31" pageNumber="32" type="generic">2 juv.</specimenCount>
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, Skazka Cave (-
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||
<quantity id="F5259B2DEF61FFA6FEF8455CFE280D43" box="[327,392,538,564]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="31" pageNumber="32" unit="m" value="20.0">20 m</quantity>
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||
),
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||
<date id="46631008EF61FFA6FE21455CFD810D43" box="[414,545,538,564]" pageId="31" pageNumber="32" value="2003-08-04">4.viii.2003</date>
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. Besides slides, part of material is preserved in alcohol.
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</paragraph>
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</subSubSection>
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<subSubSection id="7AC76543EF61FF9AFF7A4506FB310CFB" lastPageId="35" lastPageNumber="36" pageId="31" pageNumber="32" type="description">
|
||
<paragraph id="326236C8EF61FFA6FF7A4506FE6D0DF5" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
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||
<emphasis id="00A9EADAEF61FFA6FF7A4506FE6C0D2D" bold="true" box="[197,460,576,602]" pageId="31" pageNumber="32">Description. Female:</emphasis>
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body (
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<figureCitation id="AAE62A4DEF61FFA6FD9E4507FD260D2C" box="[545,646,577,603]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 118</figureCitation>
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)
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<quantity id="F5259B2DEF61FFA6FD294507FC850D2C" box="[662,805,577,603]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="31" pageNumber="32" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
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length, with brownish pigmentation more developed on head and body dorsum.
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||
</paragraph>
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<paragraph id="326236C8EF61FFA6FF7A45C8FC7D0C6B" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
|
||
Head (
|
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<figureCitation id="AAE62A4DEF61FFA6FEA945C8FE220DDF" box="[278,386,654,680]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 119</figureCitation>
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): eyes 1 + 1, black in alcohol and on slides. Pigmented spots often larger than cornea. Clypeal area with axial seta in row a. Interantennal area with axial seta in row β. Frontal area with 3 axial setae in rows A, B and C. 8 setae of frontal area are spine-like (in row C: 2 + 2; D: 2 + 2), spines thin (as in
|
||
<figureCitation id="AAE62A4DEF61FFA6FF284444FF5C0C6B" box="[151,252,770,796]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 127</figureCitation>
|
||
); axial seta of row C and setae of row B are slightly thickened.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF61FFA6FF7A446FFD010B25" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
|
||
Antennae 1.54–1.7 times as long as head. Ant I: II: III: IV = 1: 2.1–2.5: 3.1–3.6: 6.8–8.3. Ant I with 7 setae; Ant II with 15 setae (
|
||
<figureCitation id="AAE62A4DEF61FFA6FE5B4416FDEE0C1D" box="[484,590,848,874]" captionStart="FIGURES 132 – 136" captionStartId="33.[151,269,1721,1745]" captionTargetBox="[164,1405,190,1678]" captionTargetId="figure@33.[151,1436,158,1747]" captionTargetPageId="33" captionText="FIGURES 132 – 136. Pygmarrhopalites pseudoprincipalis sp. nov.: 132 – 134, chaetotaxy of antennal segments: Ant I – II, anterior view (132), Ant III, posterior view (133), Ant IV, anterior view (134); 135 – 136, chaetotaxy of Abd VI: female (135), male (136)." httpUri="https://zenodo.org/record/186477/files/figure.png" pageId="31" pageNumber="32">Fig. 132</figureCitation>
|
||
). Ant III (
|
||
<figureCitation id="AAE62A4DEF61FFA6FD784416FC910C1D" box="[711,817,848,874]" captionStart="FIGURES 132 – 136" captionStartId="33.[151,269,1721,1745]" captionTargetBox="[164,1405,190,1678]" captionTargetId="figure@33.[151,1436,158,1747]" captionTargetPageId="33" captionText="FIGURES 132 – 136. Pygmarrhopalites pseudoprincipalis sp. nov.: 132 – 134, chaetotaxy of antennal segments: Ant I – II, anterior view (132), Ant III, posterior view (133), Ant IV, anterior view (134); 135 – 136, chaetotaxy of Abd VI: female (135), male (136)." httpUri="https://zenodo.org/record/186477/files/figure.png" pageId="31" pageNumber="32">Fig. 133</figureCitation>
|
||
): subbasally slightly thickened, with 18 setae and 2 sense rods; seta Aai very small and curved; setae Api and Ape shorter and thinner than others. Ant IV (
|
||
<figureCitation id="AAE62A4DEF61FFA6FAD34430FF650CC0" captionStart="FIGURES 132 – 136" captionStartId="33.[151,269,1721,1745]" captionTargetBox="[164,1405,190,1678]" captionTargetId="figure@33.[151,1436,158,1747]" captionTargetPageId="33" captionText="FIGURES 132 – 136. Pygmarrhopalites pseudoprincipalis sp. nov.: 132 – 134, chaetotaxy of antennal segments: Ant I – II, anterior view (132), Ant III, posterior view (133), Ant IV, anterior view (134); 135 – 136, chaetotaxy of Abd VI: female (135), male (136)." httpUri="https://zenodo.org/record/186477/files/figure.png" pageId="31" pageNumber="32">Fig. 134</figureCitation>
|
||
) distinctly subdivided into 5, rarely 6 subsegments; subsegmental formula: 1 + 3 + 1 = (A + M1–M2) + (M3–M5) + (B). 6 subsegments are present if whorl BA separated by suture from whorl BM1. If 5 subsegments present, basal one is 1.3–1.5 times as long as apical one and 0.8–0.9 times as long as Ant
|
||
<date id="46631008EF61FFA6FAED44ACFA3C0B73" box="[1362,1436,1002,1028]" pageId="31" pageNumber="32" value="1913-03">III. 13</date>
|
||
whorls of setae on Ant IV present; whorl BM2 of basal subsegment with 1–3 setae, rarely without setae and then total number of whorls on Ant IV is 12.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF61FFA6FF7A4318FAC50A64" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
|
||
Foreleg: precoxae and coxa with 1, 0, 1 setae (
|
||
<figureCitation id="AAE62A4DEF61FFA6FD564318FCF00B0F" box="[745,848,1118,1144]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 120</figureCitation>
|
||
). Trochanter with 3 anterior and 1 posterior setae; femur with 13 setae, a4 turned perpendicularly to the longitudinal axis of the segment (
|
||
<figureCitation id="AAE62A4DEF61FFA6FB2343C3FAA40BE8" box="[1180,1284,1157,1183]" captionStart="FIGURES 137 – 143" captionStartId="34.[151,270,1942,1966]" captionTargetBox="[191,1388,205,1941]" captionTargetId="figure@34.[191,1398,157,1949]" captionTargetPageId="34" captionText="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="31" pageNumber="32">Fig. 137</figureCitation>
|
||
). Tibiotarsus (
|
||
<figureCitation id="AAE62A4DEF61FFA6FF2043EAFEA40BB1" box="[159,260,1196,1222]" captionStart="FIGURES 137 – 143" captionStartId="34.[151,270,1942,1966]" captionTargetBox="[191,1388,205,1941]" captionTargetId="figure@34.[191,1398,157,1949]" captionTargetPageId="34" captionText="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="31" pageNumber="32">Fig. 138</figureCitation>
|
||
) with 3 setae FP and seta FSa; whorl I with 9 setae among which Ja curved and thickened; each of whorls II–V with 8 setae. Claw without tunica, with inner tooth and 2 pairs of lateral teeth; empodium narrow, with corner tooth, apical filament exceeding claw (
|
||
<figureCitation id="AAE62A4DEF61FFA6FD5443BFFCF10A64" box="[747,849,1273,1299]" captionStart="FIGURES 137 – 143" captionStartId="34.[151,270,1942,1966]" captionTargetBox="[191,1388,205,1941]" captionTargetId="figure@34.[191,1398,157,1949]" captionTargetPageId="34" captionText="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="31" pageNumber="32">Fig. 138</figureCitation>
|
||
). Claw 4.2–4.6 times shorter than tibiotarsus.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF61FFA6FF7A4266FB600AA2" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
|
||
Mid leg: precoxae and coxa with 1, 1, 3 setae (
|
||
<figureCitation id="AAE62A4DEF61FFA6FD464266FCC20A4D" box="[761,866,1312,1338]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 120</figureCitation>
|
||
). Trochanter with anterior trochanteral organ, 3 anterior and 1 posterior setae; femur with 14–15 setae (
|
||
<figureCitation id="AAE62A4DEF61FFA6FC974200FC300A17" box="[808,912,1350,1376]" captionStart="FIGURES 137 – 143" captionStartId="34.[151,270,1942,1966]" captionTargetBox="[191,1388,205,1941]" captionTargetId="figure@34.[191,1398,157,1949]" captionTargetPageId="34" captionText="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="31" pageNumber="32">Fig. 140</figureCitation>
|
||
). Tibiotarsus (
|
||
<figureCitation id="AAE62A4DEF61FFA6FB854200FB030A17" box="[1082,1187,1350,1376]" captionStart="FIGURES 137 – 143" captionStartId="34.[151,270,1942,1966]" captionTargetBox="[191,1388,205,1941]" captionTargetId="figure@34.[191,1398,157,1949]" captionTargetPageId="34" captionText="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="31" pageNumber="32">Fig. 139</figureCitation>
|
||
): 3 setae FP and seta FSa present; whorl I with 9 setae, each of whorls II–IV with 8 setae, whorl V with 7 setae. Claw with tunica, broader than in foreleg, with inner tooth and 2 pairs of lateral teeth; empodium broadened in basal half, with corner tooth and apical filament exceeding claw. Claw 4.6–5 times shorter than tibiotarsus.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF61FFA6FF7A42A7FA3D0918" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
|
||
<collectingCountry id="4ACA7658EF61FFA6FF7A42A7FEA10A8C" box="[197,257,1505,1531]" name="India" pageId="31" pageNumber="32">Hind</collectingCountry>
|
||
leg: precoxae and coxa with 1, 1, 3 setae (
|
||
<figureCitation id="AAE62A4DEF61FFA6FCB542A7FCD40A8C" box="[778,884,1505,1531]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 120</figureCitation>
|
||
). Trochanter as in mid leg; femur with 13–14 setae: 2 posterior ones are very small (
|
||
<figureCitation id="AAE62A4DEF61FFA6FDE7414EFD1F0955" box="[600,703,1544,1570]" captionStart="FIGURES 137 – 143" captionStartId="34.[151,270,1942,1966]" captionTargetBox="[191,1388,205,1941]" captionTargetId="figure@34.[191,1398,157,1949]" captionTargetPageId="34" captionText="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="31" pageNumber="32">Fig. 142</figureCitation>
|
||
). Chaetotaxy of tibiotarsus as in mid leg (
|
||
<figureCitation id="AAE62A4DEF61FFA6FB17414EFAAE0955" box="[1192,1294,1544,1570]" captionStart="FIGURES 137 – 143" captionStartId="34.[151,270,1942,1966]" captionTargetBox="[191,1388,205,1941]" captionTargetId="figure@34.[191,1398,157,1949]" captionTargetPageId="34" captionText="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="31" pageNumber="32">Fig. 141</figureCitation>
|
||
). Claw with tunica, a little broader than mid claw, with inner tooth and 2 pairs of lateral teeth; empodium broader than in mid leg, without tooth, apical filament exceeding claw (
|
||
<figureCitation id="AAE62A4DEF61FFA6FC9B4113FC2A0918" box="[804,906,1621,1647]" captionStart="FIGURES 137 – 143" captionStartId="34.[151,270,1942,1966]" captionTargetBox="[191,1388,205,1941]" captionTargetId="figure@34.[191,1398,157,1949]" captionTargetPageId="34" captionText="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="31" pageNumber="32">Fig. 141</figureCitation>
|
||
). Claw 5.5–6.2 times shorter than tibiotarsus.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF61FFA6FF7A413AFCBD09E1" blockId="31.[151,1437,152,2034]" box="[197,797,1660,1686]" pageId="31" pageNumber="32">Length ratio of tibiotarsi I: II: III = 1: 1–1.1: 1.3.</paragraph>
|
||
<paragraph id="326236C8EF61FFA6FF7A41E5FE57087D" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
|
||
Ventral tube with 2 small subapical setulae on corpus and 2 long and smooth eversible sacks. Tenaculum (
|
||
<figureCitation id="AAE62A4DEF61FFA6FF1F418FFEAA0994" box="[160,266,1737,1763]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 131</figureCitation>
|
||
): ramus 3-dentate, with basal appendage; anterior lobe with 2 apical setulae; tip of posterior lobe exceeding tip of anterior lobe.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF61FFA6FF7A4050FCB208D2" blockId="31.[151,1437,152,2034]" pageId="31" pageNumber="32">
|
||
Furca (
|
||
<figureCitation id="AAE62A4DEF61FFA6FEA74050FEDF0847" box="[280,383,1814,1840]" captionStart="FIGURES 137 – 143" captionStartId="34.[151,270,1942,1966]" captionTargetBox="[191,1388,205,1941]" captionTargetId="figure@34.[191,1398,157,1949]" captionTargetPageId="34" captionText="FIGURES 137 – 143. Pygmarrhopalites pseudoprincipalis sp. nov.: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138 – 139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="31" pageNumber="32">Fig. 143</figureCitation>
|
||
): manubrium with 7 + 7 posterior setae. Dens with Ie, Ii, IIpe, IIIpi and IVpi as spines, Ie very strong and looks somewhat articulated (
|
||
<figureCitation id="AAE62A4DEF61FFA6FD18407BFCAE0820" box="[679,782,1853,1879]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 128</figureCitation>
|
||
); setae Ipe, IIIpe–Vpe thickened in basal half; 3, 2, 1, 1 setae on anterior side. Mucro with serrated edges; tip narrow and almost pointed, somewhat curved in lateral view (
|
||
<figureCitation id="AAE62A4DEF61FFA6FF5F40CDFEE608D2" box="[224,326,1931,1957]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 130</figureCitation>
|
||
). Dens 1.3–1.5 times as long as mucro.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF61FF9AFF7A40F7FCBD0F8B" blockId="31.[151,1437,152,2034]" lastBlockId="35.[151,1437,152,1682]" lastPageId="35" lastPageNumber="36" pageId="31" pageNumber="32">
|
||
Great abdomen (
|
||
<figureCitation id="AAE62A4DEF61FFA6FE3840F7FE4F08BC" box="[391,495,1969,1995]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 120</figureCitation>
|
||
): mesothorax with 2 + 2 spine-like or thickened dorsal setae (
|
||
<figureCitation id="AAE62A4DEF61FFA6FB0640F7FABC08BC" box="[1209,1308,1969,1995]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 126</figureCitation>
|
||
) in row m, which are longer than other anterior setae (
|
||
<figureCitation id="AAE62A4DEF61FFA6FD32409EFD540885" box="[653,756,2008,2034]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="31" pageNumber="32">Fig. 125</figureCitation>
|
||
); posterior lateral complex with 6 setae; furca base complex with 9 setae; ventral complex with 3 setae; posterior dorsal complex with 3 rows of relatively short setae, most of them 1.5 times shorter than hind claw (
|
||
<figureCitation id="AAE62A4DEF5FFF98FCB3402CFCD208F3" box="[780,882,1898,1924]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="33" pageNumber="34">Fig. 124</figureCitation>
|
||
), except two the most posterior setae of rows d I and d II which are as long as hindclaw but distinctly shorter than axial seta ms1 of Abd VI (
|
||
<figureCitation id="AAE62A4DEF5FFF98FA9D40D7FA2E08DC" box="[1314,1422,1937,1963]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="33" pageNumber="34">Fig. 123</figureCitation>
|
||
). Trichobothria ABC equidistant (AB≤BC) and form an angle about 160o. Single seta of p-row of Abd I is located below the level of trichobothrium B anteriorly to the trichobothrial complex (marked with arrow); seta b1 lies approximately in the middle of the line between trichobothria B and C; seta c1 of thichobothrial complex lies on the level of trichobothrium C or lower.
|
||
</paragraph>
|
||
<caption id="66A26640EF5EFF99FF284017FC8E0884" httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="32" pageNumber="33" targetBox="[154,1433,190,1872]" targetPageId="32">
|
||
<paragraph id="326236C8EF5EFF99FF284017FC8E0884" blockId="32.[151,1436,1873,2035]" pageId="32" pageNumber="33">
|
||
<emphasis id="00A9EADAEF5EFF99FF284017FEDD081E" bold="true" box="[151,381,1873,1897]" pageId="32" pageNumber="33">FIGURES 118–131.</emphasis>
|
||
<taxonomicName id="F5DD4D4BEF5EFF99FE3B4014FCB6081E" box="[388,790,1874,1897]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="pseudoprincipalis" status="sp. nov.">
|
||
<emphasis id="00A9EADAEF5EFF99FE3B4014FCB6081E" box="[388,790,1874,1897]" italics="true" pageId="32" pageNumber="33">Pygmarrhopalites pseudoprincipalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1B9A57A1EF5EFF99FCA14017FCD6081E" box="[798,886,1873,1897]" pageId="32" pageNumber="33" rank="species">
|
||
<emphasis id="00A9EADAEF5EFF99FCA14017FCD6081E" bold="true" box="[798,886,1873,1897]" pageId="32" pageNumber="33">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121–129, shape of setae: (ms1) of Abd VI (121), (mps2) of Abd VI (122), (dI-1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="66A26640EF5FFF98FF2841FFFEF90862" httpUri="https://zenodo.org/record/186477/files/figure.png" pageId="33" pageNumber="34" targetBox="[164,1405,190,1678]" targetPageId="33">
|
||
<paragraph id="326236C8EF5FFF98FF2841FFFEF90862" blockId="33.[151,1436,1721,1813]" pageId="33" pageNumber="34">
|
||
<emphasis id="00A9EADAEF5FFF98FF2841FFFED809A6" bold="true" box="[151,376,1721,1745]" pageId="33" pageNumber="34">FIGURES 132–136.</emphasis>
|
||
<taxonomicName id="F5DD4D4BEF5FFF98FEC141FFFCA309A7" box="[382,771,1721,1744]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pseudoprincipalis" status="sp. nov.">
|
||
<emphasis id="00A9EADAEF5FFF98FEC141FFFCA309A7" box="[382,771,1721,1744]" italics="true" pageId="33" pageNumber="34">Pygmarrhopalites pseudoprincipalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1B9A57A1EF5FFF98FCB541FFFCFF09A6" box="[778,863,1721,1745]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="00A9EADAEF5FFF98FCB541FFFCFF09A6" bold="true" box="[778,863,1721,1745]" pageId="33" pageNumber="34">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 132–134, chaetotaxy of antennal segments: Ant I–II, anterior view (132), Ant III, posterior view (133), Ant IV, anterior view (134); 135–136, chaetotaxy of Abd VI: female (135), male (136).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="66A26640EF5CFF9BFF2840D0FAD30884" httpUri="https://zenodo.org/record/186478/files/figure.png" pageId="34" pageNumber="35" targetBox="[191,1388,205,1941]" targetPageId="34">
|
||
<paragraph id="326236C8EF5CFF9BFF2840D0FAD30884" blockId="34.[151,1437,1942,2035]" pageId="34" pageNumber="35">
|
||
<emphasis id="00A9EADAEF5CFF9BFF2840D0FEDA08D9" bold="true" box="[151,378,1942,1966]" pageId="34" pageNumber="35">FIGURES 137–143.</emphasis>
|
||
<taxonomicName id="F5DD4D4BEF5CFF9BFE3E40D1FCAA08D9" box="[385,778,1943,1966]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pseudoprincipalis" status="sp. nov.">
|
||
<emphasis id="00A9EADAEF5CFF9BFE3E40D1FCAA08D9" box="[385,778,1943,1966]" italics="true" pageId="34" pageNumber="35">Pygmarrhopalites pseudoprincipalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1B9A57A1EF5CFF9BFCAD40D0FCC808D9" box="[786,872,1942,1966]" pageId="34" pageNumber="35" rank="species">
|
||
<emphasis id="00A9EADAEF5CFF9BFCAD40D0FCC808D9" bold="true" box="[786,872,1942,1966]" pageId="34" pageNumber="35">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: 137, 140, 142, chaetotaxy of trochanter and femur, anterior view: foreleg (137), mid leg (140), hind leg (142); 138–139, 141, chaetotaxy of tibiotarsus and foot complex: fore leg, anterior view (138), mid- (139) and hind leg (141), posterior view; 143, chaetotaxy of furca, posterior view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="326236C8EF5DFF9AFF7A464EFBA60E55" blockId="35.[151,1437,152,1682]" box="[197,1030,264,290]" pageId="35" pageNumber="36">
|
||
Fifth abdominal segment with trichobothrium D and 4 setae (
|
||
<figureCitation id="AAE62A4DEF5DFF9AFC2D464EFC590E55" box="[914,1017,264,290]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="35" pageNumber="36">Fig. 120</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="326236C8EF5DFF9AFF7A4668FC9B0EE1" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
Sixth abdominal segment (
|
||
<figureCitation id="AAE62A4DEF5DFF9AFDB84668FDD10E3F" box="[519,625,302,328]" captionStart="FIGURES 132 – 136" captionStartId="33.[151,269,1721,1745]" captionTargetBox="[164,1405,190,1678]" captionTargetId="figure@33.[151,1436,158,1747]" captionTargetPageId="33" captionText="FIGURES 132 – 136. Pygmarrhopalites pseudoprincipalis sp. nov.: 132 – 134, chaetotaxy of antennal segments: Ant I – II, anterior view (132), Ant III, posterior view (133), Ant IV, anterior view (134); 135 – 136, chaetotaxy of Abd VI: female (135), male (136)." httpUri="https://zenodo.org/record/186477/files/figure.png" pageId="35" pageNumber="36">Fig. 135</figureCitation>
|
||
): circumanal setae distinctly thickened (
|
||
<figureCitation id="AAE62A4DEF5DFF9AFBE94668FAA10E3F" box="[1110,1281,302,328]" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="35" pageNumber="36">Figs 121, 122</figureCitation>
|
||
); appendices anales inserted in papilla, turned towards genital opening, deeply serrated and palmated in distal part (
|
||
<figureCitation id="AAE62A4DEF5DFF9AFAD34613FF670EE1" captionStart="FIGURES 118 – 131" captionStartId="32.[151,272,1873,1897]" captionTargetBox="[154,1433,190,1872]" captionTargetId="figure@32.[151,1436,145,1907]" captionTargetPageId="32" captionText="FIGURES 118 – 131. Pygmarrhopalites pseudoprincipalis sp. nov.: 118, outline of habitus; 119, chaetotaxy of head, frontal view; 120, chaetotaxy of great abdomen, lateral view; 121 – 129, shape of setae: (ms 1) of Abd VI (121), (mps 2) of Abd VI (122), (dI- 1) of great abdomen (123), posterior dorsal seta of great abdomen (124), anterior dorsal seta of abdomen (125), dorsal spine-like seta of mesothorax (126), spine-like seta of head vertex (127), (Ie) of dens (128), appendices anales (129); 130, apex of mucro; 131, tenaculum." httpUri="https://zenodo.org/record/186476/files/figure.png" pageId="35" pageNumber="36">Fig. 129</figureCitation>
|
||
); hind claw 1–1.4 times as long as appendices anales.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF5DFF9AFF7A46E4FDEB0D47" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
<emphasis id="00A9EADAEF5DFF9AFF7A46E4FEAE0ECB" bold="true" box="[197,270,418,444]" pageId="35" pageNumber="36">Male:</emphasis>
|
||
body length
|
||
<quantity id="F5259B2DEF5DFF9AFE1446E4FDFB0ECB" box="[427,603,418,444]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="35" pageNumber="36" unit="mm" value="0.8" valueMax="0.85" valueMin="0.75">0.75–0.85 mm</quantity>
|
||
. Antenna: head = 1.8–1.9. Ant I: II: III: IV = 1: 2.1–2.3: 3.2–3.3: 7.8–8.3. Ant IV subdivided into 7–8 subsegments, basal one or two are separated by annulations with several rings; number of setal whorls on Ant IV as in female. Anal valves without thickened setae and with lesser than in female number of setae (
|
||
<figureCitation id="AAE62A4DEF5DFF9AFE684550FD9E0D47" box="[471,574,534,560]" captionStart="FIGURES 132 – 136" captionStartId="33.[151,269,1721,1745]" captionTargetBox="[164,1405,190,1678]" captionTargetId="figure@33.[151,1436,158,1747]" captionTargetPageId="33" captionText="FIGURES 132 – 136. Pygmarrhopalites pseudoprincipalis sp. nov.: 132 – 134, chaetotaxy of antennal segments: Ant I – II, anterior view (132), Ant III, posterior view (133), Ant IV, anterior view (134); 135 – 136, chaetotaxy of Abd VI: female (135), male (136)." httpUri="https://zenodo.org/record/186477/files/figure.png" pageId="35" pageNumber="36">Fig. 136</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="326236C8EF5DFF9AFF7A457BFE940D09" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
<emphasis id="00A9EADAEF5DFF9AFF7A457BFEF50D20" bold="true" box="[197,341,573,599]" pageId="35" pageNumber="36">Variability.</emphasis>
|
||
In single female seta ms1 of Abd VI is apically bifurcated. Ant IV rarely not divided into subsegments.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF5DFF9AFF7A45CCFB310CFB" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
<emphasis id="00A9EADAEF5DFF9AFF7A45CCFDAD0DD3" bold="true" box="[197,525,650,676]" pageId="35" pageNumber="36">Bionomy and distribution.</emphasis>
|
||
Most of the specimens were collected from water surface, and some from wet walls. In Kizil-Coba Cave this species inhabits dark zone but penetrates not far from entrance (about
|
||
<quantity id="F5259B2DEF5DFF9AFA8A45F7FF0E0D85" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="35" pageNumber="36" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100–200 m</quantity>
|
||
distance, down to -
|
||
<quantity id="F5259B2DEF5DFF9AFE21459EFE7E0D85" box="[414,478,728,754]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="36" unit="m" value="30.0">30 m</quantity>
|
||
depth) and has not been found mixed together with other species of the genus which inhabit more distant and deep parts of the cave. Troglophile.
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFC1345B9FB3A0C6F" box="[940,1178,767,792]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="pseudoprincipalis" status="sp. nov.">
|
||
<emphasis id="00A9EADAEF5DFF9AFC1345B9FB3A0C6F" box="[940,1178,767,792]" italics="true" pageId="35" pageNumber="36">P. pseudoprincipalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1B9A57A1EF5DFF9AFB1E45B8FB5E0C6F" box="[1185,1278,766,792]" pageId="35" pageNumber="36" rank="species">
|
||
<emphasis id="00A9EADAEF5DFF9AFB1E45B8FB5E0C6F" bold="true" box="[1185,1278,766,792]" pageId="35" pageNumber="36">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
was found in the caves of only two adjacent massifs of Eastern biospeleological region of Crimea (Karabi and Dolgorukovsky massifs) (
|
||
<tableCitation id="7F5F0373EF5DFF9AFE73440AFD860C11" box="[460,550,844,870]" captionStart="TABLE 1" captionStartId="8.[151,239,151,175]" captionTargetBox="[158,1429,242,593]" captionTargetPageId="8" captionText="TABLE 1. Records of Arrhopalitidae species in the caves of Crimea; (ske) = Pygmarrhopalites principalis skelicus ssp. nov.; (? ssp) =? independent subspecies." httpUri="http://table.plazi.org/id/66A26640EF76FFB1FF2847D1FD9E0FA6" pageId="35" pageNumber="36" tableUuid="66A26640EF76FFB1FF2847D1FD9E0FA6">Table 1</tableCitation>
|
||
). It shows allopatric distribution with close related
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFB29440AFA3C0C12" box="[1174,1436,844,869]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="principalis" status="subsp. nov." subSpecies="skelicus">
|
||
<emphasis id="00A9EADAEF5DFF9AFB29440AFA3C0C12" box="[1174,1436,844,869]" italics="true" pageId="35" pageNumber="36">P. principalis skelicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1B9A57A1EF5DFF9AFF284434FEA00CFB" box="[151,256,882,908]" pageId="35" pageNumber="36" rank="subSpecies">
|
||
<emphasis id="00A9EADAEF5DFF9AFF284434FEA00CFB" bold="true" box="[151,256,882,908]" pageId="35" pageNumber="36">ssp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(Aj-Petri Massif) and another variance of
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFD4C4434FB2D0CFB" authority="Chatyr-Dag Massif" authorityName="Chatyr-Dag Massif" box="[755,1165,882,908]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="principalis">
|
||
<emphasis id="00A9EADAEF5DFF9AFD4C4434FC2E0CFC" box="[755,910,882,907]" italics="true" pageId="35" pageNumber="36">P. principalis</emphasis>
|
||
(Chatyr-Dag Massif)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7AC76543EF5DFF9AFF7A44DEFF430CAD" pageId="35" pageNumber="36" type="etymology">
|
||
<paragraph id="326236C8EF5DFF9AFF7A44DEFF430CAD" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
<emphasis id="00A9EADAEF5DFF9AFF7A44DEFEF40CC5" bold="true" box="[197,340,920,946]" pageId="35" pageNumber="36">Etymology.</emphasis>
|
||
The name of the new species reflects its similarity to
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFC5F44DFFF7F0CAD" authority="Stach, 1945" authorityName="Stach" authorityYear="1945" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="principalis">
|
||
<emphasis id="00A9EADAEF5DFF9AFC5F44DFFAE00CC5" box="[992,1344,921,946]" italics="true" pageId="35" pageNumber="36">Pygmarrhopalites principalis</emphasis>
|
||
(
|
||
<bibRefCitation id="564C4B39EF5DFF9AFAEE44DFFF760CAD" author="Stach" pageId="35" pageNumber="46" refString="Stach, J. (1945) The species of the genus Arrhopalites occurring in European caves. Acta Musei Historiae Naturalis, Academia Polona Litterarum et Scientiarum, 1, 1 - 47." type="journal article" year="1945">Stach, 1945</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7AC76543EF5DFF9AFF7A44A0FCBF0B39" pageId="35" pageNumber="36" type="discussion">
|
||
<paragraph id="326236C8EF5DFF9AFF7A44A0FCBF0B39" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
<emphasis id="00A9EADAEF5DFF9AFF7A44A0FE980B77" bold="true" box="[197,312,998,1024]" pageId="35" pageNumber="36">Remarks</emphasis>
|
||
. The new species belongs to the
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFD7B44A1FCE40B77" box="[708,836,999,1024]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="principalis">
|
||
<emphasis id="00A9EADAEF5DFF9AFD7B44A1FCE40B77" box="[708,836,999,1024]" italics="true" pageId="35" pageNumber="36">principalis</emphasis>
|
||
</taxonomicName>
|
||
-group with 13 Palaearctic species (
|
||
<bibRefCitation id="564C4B39EF5DFF9AFB5244A0FA370B77" author="Bretfeld" box="[1261,1431,998,1024]" pageId="35" pageNumber="45" refString="Bretfeld, G. (1999) Symphypleona. In: Dunger W. (Ed.) Synopses on Palaearctic Collembola, vol. 2. Abhandlungen und Berichte des Naturkundemuseum Gorlitz, 71, 1 - 318." type="book chapter" year="1999">Bretfeld 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="564C4B39EF5DFF9AFF28434BFE8F0B50" author="Palissa" box="[151,303,1037,1063]" pageId="35" pageNumber="46" refString="Palissa, A. (2000) Beitrage zur Collembolenfauna der Hohlen deutscher Mittelgebirge. Teil II (mit Anhang uber einige Dipluren). Beitrage zur Entomologie 50: 199 - 236." type="journal article" year="2000">Palissa 2000</bibRefCitation>
|
||
). Accordingly to the palmate shape of appendices anales and presence of spine-like setae on head dorsum, the closest to the new species are five of them.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7AC76543EF5DFF9AFF7A431DFB990AF4" pageId="35" pageNumber="36" type="reference_group">
|
||
<paragraph id="326236C8EF5DFF9AFF7A431DFB300A78" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
From
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFEB4431DFDE90B02" authority="Stach, 1945" authorityName="Stach" authorityYear="1945" box="[267,585,1115,1141]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="principalis">
|
||
<emphasis id="00A9EADAEF5DFF9AFEB4431DFE060B03" box="[267,422,1115,1140]" italics="true" pageId="35" pageNumber="36">P. principalis</emphasis>
|
||
(
|
||
<bibRefCitation id="564C4B39EF5DFF9AFE0A431DFDE10B02" author="Stach" box="[437,577,1115,1141]" pageId="35" pageNumber="46" refString="Stach, J. (1945) The species of the genus Arrhopalites occurring in European caves. Acta Musei Historiae Naturalis, Academia Polona Litterarum et Scientiarum, 1, 1 - 47." type="journal article" year="1945">Stach, 1945</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
the new species differs mainly by acuminate tip of mucro (shovel-like in
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFF2843C7FEBB0BED" box="[151,283,1153,1178]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="principalis">
|
||
<emphasis id="00A9EADAEF5DFF9AFF2843C7FEBB0BED" box="[151,283,1153,1178]" italics="true" pageId="35" pageNumber="36">principalis</emphasis>
|
||
</taxonomicName>
|
||
) and thickened setae Ipe, IIIpe–Vpe of dens. Besides original Stach’s description, some other characteristics of
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFED343EEFDAC0BB6" box="[364,524,1192,1217]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="principalis">
|
||
<emphasis id="00A9EADAEF5DFF9AFED343EEFDAC0BB6" box="[364,524,1192,1217]" italics="true" pageId="35" pageNumber="36">P. principalis</emphasis>
|
||
</taxonomicName>
|
||
which differ it from the new species were figured: forked axial seta ms1 of small abdomen and heavy spines of head dorsum (
|
||
<bibRefCitation id="564C4B39EF5DFF9AFD534388FCD50B9F" author="Gisin" box="[748,885,1230,1256]" pageId="35" pageNumber="45" refString="Gisin, H. (1947) Sur les insectes apterygotes du Parc National Suisse. Especes et groupements euedaphiques. Resultats des recheerches scientifiques enterprises au Parc Natational Suisse, 2, 77 - 91." type="journal article" year="1947">Gisin 1947</bibRefCitation>
|
||
:
|
||
<figureCitation id="AAE62A4DEF5DFF9AFC3F4388FC680B9F" box="[896,968,1230,1256]" captionStart="FIGURES 2 – 18" captionStartId="10.[151,275,1867,1891]" captionTargetBox="[186,1386,191,1866]" captionTargetId="figure@10.[184,1399,125,1866]" captionTargetPageId="10" captionText="FIGURES 2 – 18. Arrhopalites karabiensis sp. nov.: 2, outline of habitus; 3, chaetotaxy of head, frontal view; 4, chaetotaxy of great abdomen, lateral view; 5 – 16, shape of setae: (mpi 1) of Abd VI (5), (mps 2) of Abd VI (6), (ms 1) of Abd VI (7), posterior dorsal seta of great abdomen (8), (dI- 1) of great abdomen (9), anterior dorsal seta of abdomen (10), dorsal seta of mesothorax (11), spine-like (Ia) seta of dens (12), spine-like (Ipe) seta of dens (13), spine-like seta of head vertex (14), appendices anales, lateral view (15) and dorsal view (16); 17, tenaculum; 18, apex of mucro." httpUri="https://zenodo.org/record/186466/files/figure.png" pageId="35" pageNumber="36">Fig. 2</figureCitation>
|
||
;
|
||
<bibRefCitation id="564C4B39EF5DFF9AFC694388FAE40B9F" author="Christiansen" box="[982,1348,1230,1256]" pageId="35" pageNumber="45" refString="Christiansen, K. & Bellinger, P. (1998) The Collembola of North America North of the Rio Grande. 2 nd edition, Grinnell College, Grinnell, 1518 pp." type="book" year="1998">Christiansen & Bellinger 1998</bibRefCitation>
|
||
: 1234). However, belonging of these descriptions to the same species seems rather ambiguous.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF5DFF9AFF7A425AFB990AF4" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
From Palaearctic
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFE25425AFD530A41" authority="Gisin, 1958" authorityName="Gisin" authorityYear="1958" box="[410,755,1308,1334]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="secundarius">
|
||
<emphasis id="00A9EADAEF5DFF9AFE25425AFDEA0A42" box="[410,586,1308,1333]" italics="true" pageId="35" pageNumber="36">P. secundarius</emphasis>
|
||
(
|
||
<bibRefCitation id="564C4B39EF5DFF9AFDE5425AFD4A0A41" author="Gisin" box="[602,746,1308,1334]" pageId="35" pageNumber="45" refString="Gisin, H. (1958) Sur la faune europeenne des Collemboles II. Revue Suisse de Zoologie, 65 (36), 773 - 778." type="journal article" year="1958">Gisin, 1958</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
described species differs by: much larger body (
|
||
<quantity id="F5259B2DEF5DFF9AFAFF425AFF650A2A" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="35" pageNumber="36" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
|
||
in female of
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFED64205FDB80A2B" box="[361,536,1347,1372]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="secundarius">
|
||
<emphasis id="00A9EADAEF5DFF9AFED64205FDB80A2B" box="[361,536,1347,1372]" italics="true" pageId="35" pageNumber="36">P. secundarius</emphasis>
|
||
</taxonomicName>
|
||
); thin spines of head dorsum (stout in
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFC504205FB3E0A2B" box="[1007,1182,1347,1372]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="secundarius">
|
||
<emphasis id="00A9EADAEF5DFF9AFC504205FB3E0A2B" box="[1007,1182,1347,1372]" italics="true" pageId="35" pageNumber="36">P. secundarius</emphasis>
|
||
</taxonomicName>
|
||
); elongated shape of outer Ie spine of dens (oval and short in
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFDD0422FFCBA0AF5" box="[623,794,1385,1410]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="secundarius">
|
||
<emphasis id="00A9EADAEF5DFF9AFDD0422FFCBA0AF5" box="[623,794,1385,1410]" italics="true" pageId="35" pageNumber="36">P. secundarius</emphasis>
|
||
</taxonomicName>
|
||
); shape of mucronal tip.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7AC76543EF5DFF9AFF7A42D6FE4509E5" pageId="35" pageNumber="36" type="description">
|
||
<paragraph id="326236C8EF5DFF9AFF7A42D6FCAB0AA7" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
From Hungarian
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFE2642D6FC130ADD" authority="Loksa & Rubio, 1966" authorityName="Loksa & Rubio" authorityYear="1966" box="[409,947,1424,1450]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="infrasecundarius">
|
||
<emphasis id="00A9EADAEF5DFF9AFE2642D6FD290ADE" box="[409,649,1424,1449]" italics="true" pageId="35" pageNumber="36">P. infrasecundarius</emphasis>
|
||
(
|
||
<bibRefCitation id="564C4B39EF5DFF9AFD2542D6FC0A0ADD" author="Loksa" box="[666,938,1424,1450]" pageId="35" pageNumber="46" refString="Loksa, I. & Rubio, I. (1966) Angaben zu den Kenntnissen uber die Collembolenfauna des Bakony-Gebirges. Opuscula Zoologica Budapest, 6 (1), 139 - 156." type="journal article" year="1966">Loksa & Rubio, 1966</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
the new species differs by: presence of corner tooth on fore and mid claw; shape of empodia.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF5DFF9AFF7A429BFBD30969" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
From East Russian
|
||
<emphasis id="00A9EADAEF5DFF9AFE13429BFDE60A81" box="[428,582,1501,1526]" italics="true" pageId="35" pageNumber="36">P. k o l y m e n s i s</emphasis>
|
||
(
|
||
<bibRefCitation id="564C4B39EF5DFF9AFDE5429BFC8E0A80" author="Tshelnokov" box="[602,814,1501,1527]" pageId="35" pageNumber="46" refString="Tshelnokov, V. G. (1990) A new genus, new and little known species of springtails (Collembola) in the fauna of North- East Asia. Entomologicheskoe Obozrenie, 69 (2), 342 - 352. [in Russian]." type="journal article" year="1990">Tshelnokov, 1990</bibRefCitation>
|
||
) the new species differs by: larger body, presence of Ip–IVp setae on posterior dens; presence of tooth on claw and empodium of foreleg.
|
||
</paragraph>
|
||
<paragraph id="326236C8EF5DFF9AFF7A416DFE4509E5" blockId="35.[151,1437,152,1682]" pageId="35" pageNumber="36">
|
||
From Japanese
|
||
<taxonomicName id="F5DD4D4BEF5DFF9AFE3F416DFD3F0932" authority="Yosii, 1954" authorityName="Yosii" authorityYear="1954" box="[384,671,1579,1605]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="alticola">
|
||
<emphasis id="00A9EADAEF5DFF9AFE3F416DFE5A0933" box="[384,506,1579,1604]" italics="true" pageId="35" pageNumber="36">P. alticola</emphasis>
|
||
(
|
||
<bibRefCitation id="564C4B39EF5DFF9AFDB5416DFD370932" author="Yosii" box="[522,663,1579,1605]" pageId="35" pageNumber="47" refString="Yosii, R. (1954) Hohlencollembolen Japans I. Kontyu, 20 (3 - 4), 62 - 70." type="journal article" year="1954">Yosii, 1954</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
according to redescription of
|
||
<bibRefCitation id="564C4B39EF5DFF9AFBB3416DFB760932" author="Zeppelini" box="[1036,1238,1579,1605]" pageId="35" pageNumber="47" refString="Zeppelini, D. (2004) The genus Arrhopalites (Collembola, Arrhopalitidae) in Asia, with the description of two new Japanese species of Yosiis collection. Zootaxa, 430, 1 - 26." type="journal article" year="2004">Zeppelini (2004)</bibRefCitation>
|
||
the new species differs by: presence of spiny cephalic setae, presence of corner tooth on fore claw and longer than claw apical filament on hind empodium.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |