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<document id="FB562DE1A41099AFDB5689859B7C9ECF" ID-DOI="10.5281/zenodo.186465" ID-GBIF-Dataset="7ee3040e-29c4-4368-8915-ba2f7d6b14fc" ID-ISSN="1175-5326" ID-Zenodo-Dep="186465" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461052118242" checkinUser="plazi" docAuthor="Vargovitsh, Robert S." docDate="2009" docId="BA7487DEEF5DFF91FF2841AAFF7C08D0" docLanguage="en" docName="zt02047p047.pdf" docOrigin="Zootaxa 2047" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pygmarrhopalites principalis subsp. skelicus Vargovitsh, 2009, ssp. nov." docType="treatment" docUuid="5362D932-004C-459A-825D-E84D8EA93773" docUuidSource="ZooBank" docVersion="12" lastPageNumber="41" masterDocId="464DFFA6EF7EFFB9FFBF4746FFA00F77" masterDocTitle="New Cave Arrhopalitidae (Collembola: Symphypleona) from the Crimea (Ukraine)" masterLastPageNumber="47" masterPageNumber="1" pageNumber="36" updateTime="1698579300348" updateUser="plazi">
<mods:mods id="2309093968BFAEF7A270E1BDD070B32D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B45E3932B9F2A49E6D999C33A4B9DFF4">
<mods:title id="B5BB74FE6A080E40C4BCAF5947F3DE05">New Cave Arrhopalitidae (Collembola: Symphypleona) from the Crimea (Ukraine)</mods:title>
</mods:titleInfo>
<mods:name id="7ADFE3650CD8539F499DC898821EA50F" type="personal">
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</mods:role>
<mods:namePart id="F38FCDD7CBE2F669495B6D12FAA85746">Vargovitsh, Robert S.</mods:namePart>
</mods:name>
<mods:typeOfResource id="29A2B4486EDDA48B18DC09FE3CDA4179">text</mods:typeOfResource>
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<mods:title id="3CD4B9665FFECAB8B14CF25DF4A7517B">Zootaxa</mods:title>
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<mods:part id="8E9A43726BCEAF99F23B9B392100C89B">
<mods:date id="47C222557533010133B7F69453C1E5F2">2009</mods:date>
<mods:detail id="F3FBAE6EA9021F26B59116EAF8D51EFF" type="volume">
<mods:number id="F74ED808951B3FA201E46A92B2E3A047">2047</mods:number>
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<mods:classification id="B5A9D150E6D9C0A7E5C80FBA2A097C57">journal article</mods:classification>
<mods:identifier id="9B0FA9246ED40BF147BC7B9B8DB7BCF4" type="DOI">10.5281/zenodo.186465</mods:identifier>
<mods:identifier id="43F923962DC4D18A2EED44F8E5ADB34F" type="GBIF-Dataset">7ee3040e-29c4-4368-8915-ba2f7d6b14fc</mods:identifier>
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<treatment id="BA7487DEEF5DFF91FF2841AAFF7C08D0" ID-DOI="http://doi.org/10.5281/zenodo.5618668" ID-GBIF-Taxon="119612340" ID-Zenodo-Dep="5618668" LSID="urn:lsid:zoobank.org:act:5362D932-004C-459A-825D-E84D8EA93773" httpUri="http://treatment.plazi.org/id/BA7487DEEF5DFF91FF2841AAFF7C08D0" lastPageId="40" lastPageNumber="41" pageId="35" pageNumber="36">
<subSubSection id="7AC76543EF5DFF9AFF2841AAFE3D085F" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="326236C8EF5DFF9AFF2841AAFD6D0872" blockId="35.[151,717,1771,1832]" box="[151,717,1771,1798]" pageId="35" pageNumber="36">
<heading id="692A81A4EF5DFF9AFF2841AAFD6D0872" bold="true" box="[151,717,1771,1798]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<emphasis id="00A9EADAEF5DFF9AFF2841AAFD6D0872" bold="true" box="[151,717,1771,1798]" pageId="35" pageNumber="36">
<taxonomicName id="F5DD4D4BEF5DFF9AFF2841AAFDFD0871" box="[151,605,1772,1798]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="35" pageNumber="36" phylum="Arthropoda" rank="subSpecies" species="principalis" status="subsp. nov." subSpecies="skelicus">
<emphasis id="00A9EADAEF5DFF9AFF2841AAFDFD0871" bold="true" box="[151,605,1772,1798]" italics="true" pageId="35" pageNumber="36">Pygmarrhopalites principalis skelicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1B9A57A1EF5DFF9AFDDB41ADFD6D0872" box="[612,717,1771,1797]" pageId="35" pageNumber="36" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="326236C8EF5DFF9AFF284048FE3D085F" blockId="35.[151,717,1771,1832]" box="[151,413,1806,1832]" pageId="35" pageNumber="36">
<figureCitation id="AAE62A4DEF5DFF9AFF284048FE9A085F" box="[151,314,1806,1832]" captionStart-0="FIGURES 144 – 157" captionStart-1="FIGURES 158 – 162" captionStart-2="FIGURES 163 – 171" captionStartId-0="37.[151,270,1874,1898]" captionStartId-1="38.[151,274,1970,1994]" captionStartId-2="39.[151,269,1910,1934]" captionTargetBox-0="[189,1394,186,1873]" captionTargetBox-1="[201,1356,184,1944]" captionTargetBox-2="[173,1414,186,1857]" captionTargetId-0="figure@37.[189,1401,129,1878]" captionTargetId-1="figure@38.[195,1391,154,1968]" captionTargetId-2="figure@39.[169,1423,162,1905]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionText-0="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." captionText-1="FIGURES 158 – 162. Pygmarrhopalites principalis skelicus ssp. nov.: 158 – 160, chaetotaxy of antennal segments, anterior view: Ant I – II (158), Ant III (159), Ant IV (160); 161, outline of antenna of male; 162, apex of mucro." captionText-2="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri-0="https://zenodo.org/record/186479/files/figure.png" httpUri-1="https://zenodo.org/record/186480/files/figure.png" httpUri-2="https://zenodo.org/record/186481/files/figure.png" pageId="35" pageNumber="36">Figs 144–171</figureCitation>
,
<tableCitation id="7F5F0373EF5DFF9AFEF94048FE3D085F" box="[326,413,1806,1832]" captionStart="TABLE 3" captionStartId="24.[151,239,577,601]" captionText="TABLE 3. Dens chaetotaxy of Pygmarrhopalites tauricus sp. nov., P. kaprusi sp. nov., P. pseudoprincipalis sp. nov. and P. principalis skelicus ssp. nov.; + = simple seta; S = spine-like; s = weakly spine-like; v. p. = varying position; in brackets = variability." pageId="35" pageNumber="36">Table 3</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="7AC76543EF5DFF9DFF28401DFD6B0E3B" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="326236C8EF5DFF9DFF28401DFD6B0E3B" blockId="35.[151,1436,1883,2028]" lastBlockId="36.[151,1437,152,2034]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="00A9EADAEF5DFF9AFF28401DFEB60802" bold="true" box="[151,278,1883,1909]" pageId="35" pageNumber="36">Diagnosis</emphasis>
. Body with weak reddish-brownish spotted pigmentation, eyes pigmented; trichobothrium B somewhat out of AC (ABC=~160o) and AB&lt;BC; mesothorax with thickened dorsal setae; posterior dorsal setae of great abdomen as long as hind claw and longer; antenna in females 2 times as long as head; Ant IV with 5–7 subsegments; head with 4 + 4 spine-like setae; all claws with inner tooth, mid and hind claw with tunica; empodia with apical filament overtopping tip of claw, fore and mid empodia with, and hind – without corner tooth; trochanter II with 5 setae; tenaculum with 2 setulae; manubrium with 7 + 7 setae; dens with 5 spines or spiny setae, Ie very strong and looks somewhat articulated, anteriorly with 3, 2, 1, 1 setae; tip of mucro broadened or rounded; small abdomen with rather strong but not broadened circumanal setae and axial seta ms1 not forked; appendices anales palmate.
</paragraph>
</subSubSection>
<subSubSection id="7AC76543EF5AFF9DFF7A461EFCB70DDF" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="326236C8EF5AFF9DFF7A461EFB8B0EB7" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
<emphasis id="00A9EADAEF5AFF9DFF7A461EFE1C0E05" bold="true" box="[197,444,344,370]" pageId="36" pageNumber="37">Material examined.</emphasis>
<typeStatus id="ED66886AEF5AFF9DFE7B461FFD920E04" box="[452,562,345,371]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
on slide (C-277-1): female,
<collectingCountry id="4ACA7658EF5AFF9DFC38461FFC490E04" box="[903,1001,345,371]" name="Ukraine" pageId="36" pageNumber="37">Ukraine</collectingCountry>
, Crimea, Aj-Petri Massif, Skelskaya Cave,
<date id="46631008EF5AFF9DFF5746C6FE240EED" box="[232,388,384,410]" pageId="36" pageNumber="37" value="1998-08-14">14.viii.1998</date>
.
<typeStatus id="ED66886AEF5AFF9DFE2E46C6FDAB0EED" box="[401,523,384,410]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
on slides:
<specimenCount id="24DBFD41EF5AFF9DFD2946C6FD5B0EED" box="[662,763,384,410]" pageId="36" pageNumber="37" type="male">8 males</specimenCount>
,
<specimenCount id="24DBFD41EF5AFF9DFCB646C6FC360EED" box="[777,918,384,410]" pageId="36" pageNumber="37" type="female">23 females</specimenCount>
,
<specimenCount id="24DBFD41EF5AFF9DFC1C46C6FC4E0EED" box="[931,1006,384,410]" pageId="36" pageNumber="37" type="generic">5 juv.</specimenCount>
, collected together with
<typeStatus id="ED66886AEF5AFF9DFA9846C6FA390EED" box="[1319,1433,384,410]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
.
<typeStatus id="ED66886AEF5AFF9DFF2846E0FEA40EB7" box="[151,260,422,448]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
and 34
<typeStatus id="ED66886AEF5AFF9DFEDE46E0FE710EB7" box="[353,465,422,448]" pageId="36" pageNumber="37" type="paratype">paratypes</typeStatus>
are preserved in SIZNAS, 2
<typeStatus id="ED66886AEF5AFF9DFC9846E0FC370EB7" box="[807,919,422,448]" pageId="36" pageNumber="37" type="paratype">paratypes</typeStatus>
in
<collectionCode id="54CCAE0DEF5AFF9DFC0346E0FB870EB7" box="[956,1063,422,448]" pageId="36" pageNumber="37">SNHML</collectionCode>
.
</paragraph>
<paragraph id="326236C8EF5AFF9DFF7A468BFCB70DDF" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
<emphasis id="00A9EADAEF5AFF9DFF7A468BFE220E90" bold="true" box="[197,386,461,487]" pageId="36" pageNumber="37">Other material</emphasis>
(on slides).
<emphasis id="00A9EADAEF5AFF9DFDAA468BFD400E90" bold="true" box="[533,736,461,487]" pageId="36" pageNumber="37">Aj-Petri Massif:</emphasis>
female, Kaskadnaya Cave (-
<quantity id="F5259B2DEF5AFF9DFB8A468BFB250E90" box="[1077,1157,461,487]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="100.0">100 m</quantity>
),
<date id="46631008EF5AFF9DFB24468BFA8D0E90" box="[1179,1325,461,487]" pageId="36" pageNumber="37" value="1998-08-17">17.viii.1998</date>
;
<specimenCount id="24DBFD41EF5AFF9DFA86468BFA370E90" box="[1337,1431,461,487]" pageId="36" pageNumber="37" type="male">2 males</specimenCount>
, female,
<specimenCount id="24DBFD41EF5AFF9DFF4346B2FEE60D79" box="[252,326,500,526]" pageId="36" pageNumber="37" type="generic">3 juv.</specimenCount>
, same cave (-
<quantity id="F5259B2DEF5AFF9DFE4F46B2FDE50D79" box="[496,581,500,526]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="36" pageNumber="37" unit="m" value="100.0">100 m</quantity>
, -
<quantity id="F5259B2DEF5AFF9DFDE446B2FD100D79" box="[603,688,500,526]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="36" pageNumber="37" unit="m" value="150.0">150 m</quantity>
),
<date id="46631008EF5AFF9DFD7846B2FCC30D79" box="[711,867,500,526]" pageId="36" pageNumber="37" value="1998-08-20">20.viii.1998</date>
; male, female,
<specimenCount id="24DBFD41EF5AFF9DFB9C46B2FBCC0D79" box="[1059,1132,500,526]" pageId="36" pageNumber="37" type="generic">8 juv.</specimenCount>
, Drugaya Cave (-
<quantity id="F5259B2DEF5AFF9DFAF746B2FA2E0D79" box="[1352,1422,500,526]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="36" pageNumber="37" unit="m" value="27.0">27 m</quantity>
),
<date id="46631008EF5AFF9DFF28455CFEBC0D43" box="[151,284,538,564]" pageId="36" pageNumber="37" value="2005-08-01">1.viii.2005</date>
; male, female, 200-4 Cave (-
<quantity id="F5259B2DEF5AFF9DFDC6455CFD1C0D43" box="[633,700,538,564]" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="36" pageNumber="37" unit="m" value="23.0">23 m</quantity>
),
<date id="46631008EF5AFF9DFD6D455CFCF70D43" box="[722,855,538,564]" pageId="36" pageNumber="37" value="2005-08-01">1.viii.2005</date>
;
<specimenCount id="24DBFD41EF5AFF9DFCDB455CFC7B0D43" box="[868,987,538,564]" pageId="36" pageNumber="37" type="female">2 females</specimenCount>
, juv., Kamnepadnaya Cave (-
<quantity id="F5259B2DEF5AFF9DFA83455CFA2E0D43" box="[1340,1422,538,564]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="36" pageNumber="37" unit="m" value="105.0">105 m</quantity>
),
<date id="46631008EF5AFF9DFF284507FE800D2C" box="[151,288,577,603]" pageId="36" pageNumber="37" value="2005-07-29">29.vii.2005</date>
;
<specimenCount id="24DBFD41EF5AFF9DFE944507FE3F0D2C" box="[299,415,577,603]" pageId="36" pageNumber="37" type="female">2 females</specimenCount>
, Oreshek Cave (-
<quantity id="F5259B2DEF5AFF9DFDD94507FD060D2C" box="[614,678,577,603]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="36" pageNumber="37" unit="m" value="25.0">25 m</quantity>
),
<date id="46631008EF5AFF9DFD044507FCE40D2C" box="[699,836,577,603]" pageId="36" pageNumber="37" value="2005-07-28">28.vii.2005</date>
; male, Paniya Cave (-
<quantity id="F5259B2DEF5AFF9DFBFD4507FB220D2C" box="[1090,1154,577,603]" metricMagnitude="1" metricUnit="m" metricValue="1.7" pageId="36" pageNumber="37" unit="m" value="17.0">17 m</quantity>
),
<date id="46631008EF5AFF9DFB294507FAB90D2C" box="[1174,1305,577,603]" pageId="36" pageNumber="37" value="2005-08-02">2.viii.2005</date>
;
<specimenCount id="24DBFD41EF5AFF9DFA9B4507FA370D2C" box="[1316,1431,577,603]" pageId="36" pageNumber="37" type="female">5 females</specimenCount>
, Bakhchisarai Distr., Vil'ya-Burun Mt., Vilyaburunskaya Cave, traps,
<date id="46631008EF5AFF9DFC02452EFBE30DF5" box="[957,1091,616,642]" pageId="36" pageNumber="37" value="2004-07-19">19.vii.2004</date>
–
<date id="46631008EF5AFF9DFBEB452EFB7A0DF5" box="[1108,1242,616,642]" pageId="36" pageNumber="37" value="2006-07-17">17.vii.2006</date>
. A.G. Koval leg. Besides slides, part of material is preserved in alcohol.
</paragraph>
</subSubSection>
<subSubSection id="7AC76543EF5AFF91FF7A45F3FEFA0C96" lastPageId="40" lastPageNumber="41" pageId="36" pageNumber="37" type="description">
<paragraph id="326236C8EF5AFF9DFF7A45F3FE1C0D81" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
<emphasis id="00A9EADAEF5AFF9DFF7A45F3FE6F0DB8" bold="true" box="[197,463,693,719]" pageId="36" pageNumber="37">Description. Female:</emphasis>
body (
<figureCitation id="AAE62A4DEF5AFF9DFD9B45F3FD2B0DB8" box="[548,651,693,719]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="36" pageNumber="37">Fig. 144</figureCitation>
) length
<quantity id="F5259B2DEF5AFF9DFD5145F3FCCB0DB8" box="[750,875,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="36" pageNumber="37" unit="mm" value="1.15" valueMax="1.3" valueMin="1.0">1–1.3 mm</quantity>
, dorsally with reddish-brownish pigmentation, sometimes unpigmented.
</paragraph>
<paragraph id="326236C8EF5AFF9DFF7A4444FA2A0C1D" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
Head (
<figureCitation id="AAE62A4DEF5AFF9DFEAD4444FED90C6B" box="[274,377,770,796]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="36" pageNumber="37">Fig. 145</figureCitation>
): Eyes 1 + 1, more or less pigmented (with pigmented spots in alcohol and clear on slides). Clypeal area with axial seta in row a. Interantennal area with axial seta in row β. Frontal area with 3 axial setae in rows A, B and C. 8 setae of frontal area are weakly spine-like (in row C: 2 + 2; D: 2 + 2) (
<figureCitation id="AAE62A4DEF5AFF9DFAA84416FADE0C1D" box="[1303,1406,848,874]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="36" pageNumber="37">Fig. 151</figureCitation>
).
</paragraph>
<paragraph id="326236C8EF5AFF9DFF7A4430FC600B25" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
Antennae 1.9–2.1 times as long as head. Ant I: II: III: IV = 1: 2.1–2.5: 3.3–4.2: 8.3–10.2. Ant I with 7 setae; Ant II with 15 setae (
<figureCitation id="AAE62A4DEF5AFF9DFE6444DBFDE20CC0" box="[475,578,925,951]" captionStart="FIGURES 158 – 162" captionStartId="38.[151,274,1970,1994]" captionTargetBox="[201,1356,184,1944]" captionTargetId="figure@38.[195,1391,154,1968]" captionTargetPageId="38" captionText="FIGURES 158 – 162. Pygmarrhopalites principalis skelicus ssp. nov.: 158 – 160, chaetotaxy of antennal segments, anterior view: Ant I – II (158), Ant III (159), Ant IV (160); 161, outline of antenna of male; 162, apex of mucro." httpUri="https://zenodo.org/record/186480/files/figure.png" pageId="36" pageNumber="37">Fig. 158</figureCitation>
). Ant III (
<figureCitation id="AAE62A4DEF5AFF9DFD0844DBFCBF0CC0" box="[695,799,925,951]" captionStart="FIGURES 158 – 162" captionStartId="38.[151,274,1970,1994]" captionTargetBox="[201,1356,184,1944]" captionTargetId="figure@38.[195,1391,154,1968]" captionTargetPageId="38" captionText="FIGURES 158 – 162. Pygmarrhopalites principalis skelicus ssp. nov.: 158 – 160, chaetotaxy of antennal segments, anterior view: Ant I – II (158), Ant III (159), Ant IV (160); 161, outline of antenna of male; 162, apex of mucro." httpUri="https://zenodo.org/record/186480/files/figure.png" pageId="36" pageNumber="37">Fig. 159</figureCitation>
): slightly thickened in subbasal – middle part, with 18 setae and 2 sense rods; seta Aai very small and curved; setae Api and Ape shorter and thinner than others. Ant IV (
<figureCitation id="AAE62A4DEF5AFF9DFF7844ACFE8E0B73" box="[199,302,1002,1028]" captionStart="FIGURES 158 – 162" captionStartId="38.[151,274,1970,1994]" captionTargetBox="[201,1356,184,1944]" captionTargetId="figure@38.[195,1391,154,1968]" captionTargetPageId="38" captionText="FIGURES 158 – 162. Pygmarrhopalites principalis skelicus ssp. nov.: 158 – 160, chaetotaxy of antennal segments, anterior view: Ant I – II (158), Ant III (159), Ant IV (160); 161, outline of antenna of male; 162, apex of mucro." httpUri="https://zenodo.org/record/186480/files/figure.png" pageId="36" pageNumber="37">Fig. 160</figureCitation>
) usually distinctly subdivided into 5 (rarely 6 or 7) subsegments with 13(12) setal whorls. If 5 subsegments present, basal one is 1.6–1.8 times as long as apical one and 1–1.1 times as long as Ant III. Subsegmental formula: 1 + 3 + 1 = (A + M1–M2) + (M3–M5) + (B).
</paragraph>
<paragraph id="326236C8EF5AFF9DFF7A4318FE520A4D" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
Foreleg: precoxae and coxa with 1, 0, 1 setae respectively (
<figureCitation id="AAE62A4DEF5AFF9DFC2B4318FC5E0B0F" box="[916,1022,1118,1144]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="36" pageNumber="37">Fig. 146</figureCitation>
). Trochanter with 3 anterior and 1 posterior setae; femur with 12–13 setae (
<figureCitation id="AAE62A4DEF5AFF9DFDC543C3FD400BE8" box="[634,736,1157,1183]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 163</figureCitation>
), seta a 4 turned perpendicularly to the longitudinal axis of the segment. Tibiotarsus (
<figureCitation id="AAE62A4DEF5AFF9DFE6D43EAFD9B0BB1" box="[466,571,1196,1222]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 164</figureCitation>
) with 3 setae FP and seta FSa; whorl I with 9 setae among which Ja is curved and spiny; each of whorls II–V with 8 setae. Claw without tunica, with inner tooth and 2 pairs of lateral teeth; empodium narrow, with corner tooth, apical filament exceeding claw (
<figureCitation id="AAE62A4DEF5AFF9DFBC143BFFB470A64" box="[1150,1255,1273,1299]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 164</figureCitation>
). Claw 4.0–4.6 times shorter than tibiotarsus.
</paragraph>
<paragraph id="326236C8EF5AFF9DFF7A4200FEB90955" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
Mid leg: precoxae 1, 2 and coxa with 1, 1, 3 setae respectively (
<figureCitation id="AAE62A4DEF5AFF9DFC574200FBF60A17" box="[1000,1110,1350,1376]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="36" pageNumber="37">Fig. 146</figureCitation>
). Trochanter with anterior trochanteral organ, 3 anterior and 1 posterior setae; femur with 14 setae (
<figureCitation id="AAE62A4DEF5AFF9DFC43422BFBC50AF0" box="[1020,1125,1389,1415]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 166</figureCitation>
). Tibiotarsus (
<figureCitation id="AAE62A4DEF5AFF9DFAB0422BFAD80AF0" box="[1295,1400,1389,1415]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 165</figureCitation>
): 3 setae FP and seta FSa present; whorl I with 9 setae, each of whorls II–IV with 8 setae, whorl V with 7 setae. Claw with tunica, broader than in foreleg, with inner tooth and 2 pairs of lateral teeth; empodium broadened in basal half, with corner tooth and apical filament exceeding claw (
<figureCitation id="AAE62A4DEF5AFF9DFC0F42A7FBBA0A8C" box="[944,1050,1505,1531]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 165</figureCitation>
). Claw 4.5–5 times shorter than tibiotarsus.
</paragraph>
<paragraph id="326236C8EF5AFF9DFF7A4168FA3D09CA" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
<collectingCountry id="4ACA7658EF5AFF9DFF7A4168FEA1093F" box="[197,257,1582,1608]" name="India" pageId="36" pageNumber="37">Hind</collectingCountry>
leg: precoxae and coxa with 1, 1, 3 setae respectively (
<figureCitation id="AAE62A4DEF5AFF9DFC214168FBA8093F" box="[926,1032,1582,1608]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="36" pageNumber="37">Fig. 146</figureCitation>
). Trochanter as in mid leg; femur with 14 setae: 2 posterior ones are very small (
<figureCitation id="AAE62A4DEF5AFF9DFD7A4113FC8E0918" box="[709,814,1621,1647]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 167</figureCitation>
). Chaetotaxy of tibiotarsus as in mid leg. Claw with tunica, a little broader than mid claw, with inner tooth and 2 pairs of lateral teeth; empodium broader than in mid leg, without tooth, apical filament exceeding claw (
<figureCitation id="AAE62A4DEF5AFF9DFC9B41E5FC2A09CA" box="[804,906,1699,1725]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 168</figureCitation>
). Claw 5.5–6.2 times shorter than tibiotarsus.
</paragraph>
<paragraph id="326236C8EF5AFF9DFF7A418FFCBD0994" blockId="36.[151,1437,152,2034]" box="[197,797,1737,1763]" pageId="36" pageNumber="37">Length ratio of tibiotarsi I: II: III = 1: 1–1.1: 1.3.</paragraph>
<paragraph id="326236C8EF5AFF9DFF7A41B6FE930820" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
Ventral tube with 2 small subapical setulae, smooth and long eversible sacks. Tenaculum (
<figureCitation id="AAE62A4DEF5AFF9DFB5641B6FAF1087D" box="[1257,1361,1776,1802]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="36" pageNumber="37">Fig. 157</figureCitation>
): rami 3-dentate, with basal appendage; anterior lobe with 2 apical setulae; tip of posterior lobe exceeding tip of anterior lobe.
</paragraph>
<paragraph id="326236C8EF5AFF9DFF7A4022FD320885" blockId="36.[151,1437,152,2034]" pageId="36" pageNumber="37">
Furca (
<figureCitation id="AAE62A4DEF5AFF9DFEA64022FE200809" box="[281,384,1892,1918]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 171</figureCitation>
): manubrium with 7 + 7 posterior setae. Dens: setae Ie, Ii, IIpe, IIIpi and IVpi are spinelike; outer spine Ie very strong, elongated and looks somewhat articulated (
<figureCitation id="AAE62A4DEF5AFF9DFB9940CDFB3008D2" box="[1062,1168,1931,1957]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="36" pageNumber="37">Fig. 156</figureCitation>
); setae Ipe, IIIpe–Vpe simple; 3, 2, 1, 1 setae on anterior side (
<figureCitation id="AAE62A4DEF5AFF9DFDC440F7FD4408BC" box="[635,740,1969,1995]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="36" pageNumber="37">Fig. 171</figureCitation>
;
<tableCitation id="7F5F0373EF5AFF9DFD4E40F7FCEC08BC" box="[753,844,1969,1995]" captionStart="TABLE 3" captionStartId="24.[151,239,577,601]" captionText="TABLE 3. Dens chaetotaxy of Pygmarrhopalites tauricus sp. nov., P. kaprusi sp. nov., P. pseudoprincipalis sp. nov. and P. principalis skelicus ssp. nov.; + = simple seta; S = spine-like; s = weakly spine-like; v. p. = varying position; in brackets = variability." pageId="36" pageNumber="37">Table 3</tableCitation>
). Tip of mucro rounded, slightly broadened (
<figureCitation id="AAE62A4DEF5AFF9DFAD340F7FF670885" captionStart="FIGURES 158 – 162" captionStartId="38.[151,274,1970,1994]" captionTargetBox="[201,1356,184,1944]" captionTargetId="figure@38.[195,1391,154,1968]" captionTargetPageId="38" captionText="FIGURES 158 – 162. Pygmarrhopalites principalis skelicus ssp. nov.: 158 – 160, chaetotaxy of antennal segments, anterior view: Ant I – II (158), Ant III (159), Ant IV (160); 161, outline of antenna of male; 162, apex of mucro." httpUri="https://zenodo.org/record/186480/files/figure.png" pageId="36" pageNumber="37">Fig. 162</figureCitation>
). Dens 1.5–1.7 times as long as mucro.
</paragraph>
<caption id="66A26640EF5BFF9CFF284014FCE40883" httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="37" pageNumber="38" targetBox="[189,1394,186,1873]" targetPageId="37">
<paragraph id="326236C8EF5BFF9CFF284014FCE40883" blockId="37.[151,1436,1874,2036]" pageId="37" pageNumber="38">
<emphasis id="00A9EADAEF5BFF9CFF284014FEDA081D" bold="true" box="[151,378,1874,1898]" pageId="37" pageNumber="38">FIGURES 144–157.</emphasis>
<taxonomicName id="F5DD4D4BEF5BFF9CFE3E4015FCBA081D" box="[385,794,1875,1898]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="37" pageNumber="38" phylum="Arthropoda" rank="subSpecies" species="principalis" status="subsp. nov." subSpecies="skelicus">
<emphasis id="00A9EADAEF5BFF9CFE3E4015FCBA081D" box="[385,794,1875,1898]" italics="true" pageId="37" pageNumber="38">Pygmarrhopalites principalis skelicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1B9A57A1EF5BFF9CFC9E4014FC22081D" box="[801,898,1874,1898]" pageId="37" pageNumber="38" rank="subSpecies">
<emphasis id="00A9EADAEF5BFF9CFC9E4014FC22081D" bold="true" box="[801,898,1874,1898]" pageId="37" pageNumber="38">ssp. nov.</emphasis>
</taxonomicNameLabel>
: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147–156, shape of setae: (dI-1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps2) of Abd VI (152), (ms1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum.
</paragraph>
</caption>
<caption id="66A26640EF58FF9FFF2840F4FA99089B" httpUri="https://zenodo.org/record/186480/files/figure.png" pageId="38" pageNumber="39" targetBox="[201,1356,184,1944]" targetPageId="38">
<paragraph id="326236C8EF58FF9FFF2840F4FA99089B" blockId="38.[151,1436,1970,2028]" pageId="38" pageNumber="39">
<emphasis id="00A9EADAEF58FF9FFF2840F4FE2208BD" bold="true" box="[151,386,1970,1994]" pageId="38" pageNumber="39">FIGURES 158–162.</emphasis>
<taxonomicName id="F5DD4D4BEF58FF9FFE3540F5FC9708BD" box="[394,823,1971,1994]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="38" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="principalis" status="subsp. nov." subSpecies="skelicus">
<emphasis id="00A9EADAEF58FF9FFE3540F5FC9708BD" box="[394,823,1971,1994]" italics="true" pageId="38" pageNumber="39">Pygmarrhopalites principalis skelicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1B9A57A1EF58FF9FFC8040F4FC0508BD" box="[831,933,1970,1994]" pageId="38" pageNumber="39" rank="subSpecies">
<emphasis id="00A9EADAEF58FF9FFC8040F4FC0508BD" bold="true" box="[831,933,1970,1994]" pageId="38" pageNumber="39">ssp. nov.</emphasis>
</taxonomicNameLabel>
: 158–160, chaetotaxy of antennal segments, anterior view: Ant I–II (158), Ant III (159), Ant IV (160); 161, outline of antenna of male; 162, apex of mucro.
</paragraph>
</caption>
<caption id="66A26640EF59FF9EFF284030FC670881" httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="39" pageNumber="40" targetBox="[173,1414,186,1857]" targetPageId="39">
<paragraph id="326236C8EF59FF9EFF284030FC670881" blockId="39.[151,1436,1910,2038]" pageId="39" pageNumber="40">
<emphasis id="00A9EADAEF59FF9EFF284030FED808F9" bold="true" box="[151,376,1910,1934]" pageId="39" pageNumber="40">FIGURES 163–171.</emphasis>
<taxonomicName id="F5DD4D4BEF59FF9EFEC24031FCB108F9" box="[381,785,1911,1934]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="39" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="principalis" status="subsp. nov." subSpecies="skelicus">
<emphasis id="00A9EADAEF59FF9EFEC24031FCB108F9" box="[381,785,1911,1934]" italics="true" pageId="39" pageNumber="40">Pygmarrhopalites principalis skelicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1B9A57A1EF59FF9EFCA84030FCD608F9" box="[791,886,1910,1934]" pageId="39" pageNumber="40" rank="subSpecies">
<emphasis id="00A9EADAEF59FF9EFCA84030FCD608F9" bold="true" box="[791,886,1910,1934]" pageId="39" pageNumber="40">ssp. nov.</emphasis>
</taxonomicNameLabel>
: 163, 166–167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164–165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169–170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view.
</paragraph>
</caption>
<paragraph id="326236C8EF56FF91FF7A47DEFCBD0EB4" blockId="40.[151,1437,152,1959]" pageId="40" pageNumber="41">
Great abdomen (
<figureCitation id="AAE62A4DEF56FF91FE3047DEFE5B0FC5" box="[399,507,152,178]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="40" pageNumber="41">Fig. 146</figureCitation>
): mesothorax with spiny dorsal setae (
<figureCitation id="AAE62A4DEF56FF91FC7247DEFB970FC5" box="[973,1079,152,178]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="40" pageNumber="41">Fig. 150</figureCitation>
), which are longer than other anterior setae (
<figureCitation id="AAE62A4DEF56FF91FEF247F8FE180FAF" box="[333,440,190,216]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="40" pageNumber="41">Fig. 149</figureCitation>
); posterior lateral complex with 6 setae; furca base complex with 9 setae; ventral complex with 3 setae; posterior dorsal complex with setae at least as long as hind claw (
<figureCitation id="AAE62A4DEF56FF91FB0847A3FABF0F88" box="[1207,1311,229,255]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="40" pageNumber="41">Fig. 148</figureCitation>
); the most posterior seta of dI-row (
<figureCitation id="AAE62A4DEF56FF91FE06464AFDBC0E51" box="[441,540,268,294]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="40" pageNumber="41">Fig. 147</figureCitation>
) is 1.5 times longer than hind claw and longer than axial seta ms1 of Abd VI (
<figureCitation id="AAE62A4DEF56FF91FF204673FEA60E38" box="[159,262,309,335]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="40" pageNumber="41">Fig. 153</figureCitation>
). Trichobothrial complex: AB a little shorter than BC; ABC form an angle about 160o. Single seta of p-row of Abd I is located below the level of trichobothrium B anteriorly to the trichobothrial complex (marked with arrow); seta b1 lies close to the line between trichobothria B and C; seta c1 of thichobothrial complex lies on the level of trichobothrium C or lower.
</paragraph>
<paragraph id="326236C8EF56FF91FF7A4696FBA60E9D" blockId="40.[151,1437,152,1959]" box="[197,1030,464,490]" pageId="40" pageNumber="41">
Fifth abdominal segment with trichobothrium D and 4 setae (
<figureCitation id="AAE62A4DEF56FF91FC2D4696FC590E9D" box="[914,1017,464,490]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="40" pageNumber="41">Fig. 146</figureCitation>
).
</paragraph>
<paragraph id="326236C8EF56FF91FF7A46B1FD7D0D29" blockId="40.[151,1437,152,1959]" pageId="40" pageNumber="41">
Sixth abdominal segment (
<figureCitation id="AAE62A4DEF56FF91FDB546B1FDD50D66" box="[522,629,503,529]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="40" pageNumber="41">Fig. 169</figureCitation>
): circumanal setae strong but not much thickened (
<figureCitation id="AAE62A4DEF56FF91FB5D46B1FA2E0D66" box="[1250,1422,503,529]" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="40" pageNumber="41">Figs 152, 153</figureCitation>
). Appendices anales inserted in papilla, turned towards genital opening and palmated in distal part (
<figureCitation id="AAE62A4DEF56FF91FADA455BFEA20D29" captionStart="FIGURES 144 – 157" captionStartId="37.[151,270,1874,1898]" captionTargetBox="[189,1394,186,1873]" captionTargetId="figure@37.[189,1401,129,1878]" captionTargetPageId="37" captionText="FIGURES 144 – 157. Pygmarrhopalites principalis skelicus ssp. nov.: 144, outline of habitus; 145, chaetotaxy of head, frontal view; 146, chaetotaxy of great abdomen, lateral view; 147 – 156, shape of setae: (dI- 1) of great abdomen (147), posterior dorsal seta of great abdomen (148), anterior dorsal seta of abdomen (149), dorsal spine-like seta of mesothorax (150), spine-like seta of head vertex (151), (mps 2) of Abd VI (152), (ms 1) of Abd VI (153), appendices anales, dorsal (154) and lateral view (155), (Ie) of dens (156); 157, tenaculum." httpUri="https://zenodo.org/record/186479/files/figure.png" pageId="40" pageNumber="41">Figs 154–155</figureCitation>
); about 1.3 times shorter than hind claw.
</paragraph>
<paragraph id="326236C8EF56FF91FF7A452CFD100DA5" blockId="40.[151,1437,152,1959]" pageId="40" pageNumber="41">
<emphasis id="00A9EADAEF56FF91FF7A452CFEAF0DF3" bold="true" box="[197,271,618,644]" pageId="40" pageNumber="41">Male:</emphasis>
body length
<quantity id="F5259B2DEF56FF91FE11452CFDF00DF3" box="[430,592,618,644]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="9.0" metricValueMin="6.5" pageId="40" pageNumber="41" unit="mm" value="0.775" valueMax="0.9" valueMin="0.65">0.65–0.9 mm</quantity>
. Antenna: head = 1.9–2.2. Ant I: II: III: IV = 1: 2.1–2.4: 3.7–3.9: 8.9–10.3. Ant IV subdivided into 7–8 subsegments; one or two basal subsegments separated by weak annulations (
<figureCitation id="AAE62A4DEF56FF91FE9145FEFE340DA5" box="[302,404,696,722]" captionStart="FIGURES 158 – 162" captionStartId="38.[151,274,1970,1994]" captionTargetBox="[201,1356,184,1944]" captionTargetId="figure@38.[195,1391,154,1968]" captionTargetPageId="38" captionText="FIGURES 158 – 162. Pygmarrhopalites principalis skelicus ssp. nov.: 158 – 160, chaetotaxy of antennal segments, anterior view: Ant I – II (158), Ant III (159), Ant IV (160); 161, outline of antenna of male; 162, apex of mucro." httpUri="https://zenodo.org/record/186480/files/figure.png" pageId="40" pageNumber="41">Fig. 161</figureCitation>
). Abd VI as in
<figureCitation id="AAE62A4DEF56FF91FDF845FEFD0C0DA5" box="[583,684,696,722]" captionStart="FIGURES 163 – 171" captionStartId="39.[151,269,1910,1934]" captionTargetBox="[173,1414,186,1857]" captionTargetId="figure@39.[169,1423,162,1905]" captionTargetPageId="39" captionText="FIGURES 163 – 171. Pygmarrhopalites principalis skelicus ssp. nov.: 163, 166 – 167, chaetotaxy of trochanter and femur, anterior view: foreleg (163), mid leg (166), hind leg (167); 164 – 165, chaetotaxy of tibiotarsus and foot complex: foreleg, posterior view (164), mid leg, anterior view (165); 168, foot complex of hind leg; 169 – 170, chaetotaxy of Abd VI, lateral view: female (169) and male (170); 171, chaetotaxy of furca, posterior view." httpUri="https://zenodo.org/record/186481/files/figure.png" pageId="40" pageNumber="41">Fig. 170</figureCitation>
.
</paragraph>
<paragraph id="326236C8EF56FF91FF7A4598FEFA0C96" blockId="40.[151,1437,152,1959]" pageId="40" pageNumber="41">
<emphasis id="00A9EADAEF56FF91FF7A4598FDAF0D8F" bold="true" box="[197,527,734,760]" pageId="40" pageNumber="41">Bionomy and distribution.</emphasis>
In most caves specimens were taken from water surface of small pools, from stalagmites and walls, and from the wooden substance. In Kaskadnaya Cave specimens were collected from litter penetrating into the cave from surface down to -
<quantity id="F5259B2DEF56FF91FCB7446AFCF40C31" box="[776,852,812,838]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="40" pageNumber="41" unit="m" value="150.0">150 m</quantity>
vertical depth. Troglophile or “recent” troglobite. Subspecies is distributed only in Western biospeleological region, sometimes shares caves with
<taxonomicName id="F5DD4D4BEF56FF91FB4A4415FAD20C1B" box="[1269,1394,851,876]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="tauricus" status="sp. nov.">
<emphasis id="00A9EADAEF56FF91FB4A4415FAD20C1B" box="[1269,1394,851,876]" italics="true" pageId="40" pageNumber="41">P. tauricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1B9A57A1EF56FF91FAC64414FF6B0CE4" pageId="40" pageNumber="41" rank="species">
<emphasis id="00A9EADAEF56FF91FAC64414FF6B0CE4" bold="true" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="F5DD4D4BEF56FF91FEB7443FFE030CE5" box="[264,419,889,914]" class="Entognatha" family="Arrhopalitidae" genus="Arrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="peculiaris" status="sp. nov.">
<emphasis id="00A9EADAEF56FF91FEB7443FFE030CE5" box="[264,419,889,914]" italics="true" pageId="40" pageNumber="41">A. peculiaris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1B9A57A1EF56FF91FE14443FFDAC0CE4" box="[427,524,889,915]" pageId="40" pageNumber="41" rank="species">
<emphasis id="00A9EADAEF56FF91FE14443FFDAC0CE4" bold="true" box="[427,524,889,915]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
and shows allopatric mode of distribution with close related variance of
<taxonomicName id="F5DD4D4BEF56FF91FA39443FFEB70CCE" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="00A9EADAEF56FF91FA39443FFEB70CCE" italics="true" pageId="40" pageNumber="41">P. principalis</emphasis>
</taxonomicName>
from Chatyr-Dag and
<taxonomicName id="F5DD4D4BEF56FF91FD9B44E6FCB50CCE" box="[548,789,928,953]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pseudoprincipalis" status="sp. nov.">
<emphasis id="00A9EADAEF56FF91FD9B44E6FCB50CCE" box="[548,789,928,953]" italics="true" pageId="40" pageNumber="41">P. pseudoprincipalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1B9A57A1EF56FF91FCA244E6FCDB0CCD" box="[797,891,928,954]" pageId="40" pageNumber="41" rank="species">
<emphasis id="00A9EADAEF56FF91FCA244E6FCDB0CCD" bold="true" box="[797,891,928,954]" pageId="40" pageNumber="41">sp. nov.</emphasis>
</taxonomicNameLabel>
from two massifs of Eastern biospeleological region (
<tableCitation id="7F5F0373EF56FF91FF4E4481FEE80C96" box="[241,328,967,993]" captionStart="TABLE 1" captionStartId="8.[151,239,151,175]" captionTargetBox="[158,1429,242,593]" captionTargetPageId="8" captionText="TABLE 1. Records of Arrhopalitidae species in the caves of Crimea; (ske) = Pygmarrhopalites principalis skelicus ssp. nov.; (? ssp) =? independent subspecies." httpUri="http://table.plazi.org/id/66A26640EF76FFB1FF2847D1FD9E0FA6" pageId="40" pageNumber="41" tableUuid="66A26640EF76FFB1FF2847D1FD9E0FA6">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="7AC76543EF56FF91FF7A44ABFBED0B23" pageId="40" pageNumber="41" type="etymology">
<paragraph id="326236C8EF56FF91FF7A44ABFBF80B70" blockId="40.[151,1437,152,1959]" box="[197,1112,1005,1031]" pageId="40" pageNumber="41">
<emphasis id="00A9EADAEF56FF91FF7A44ABFEEA0B70" bold="true" box="[197,330,1005,1031]" pageId="40" pageNumber="41">Etymology</emphasis>
. The new subspecies is named after
<typeStatus id="ED66886AEF56FF91FD4744ABFC8A0B70" box="[760,810,1005,1031]" pageId="40" pageNumber="41">type</typeStatus>
locality, Skelskaya Cave.
</paragraph>
<paragraph id="326236C8EF56FF91FF7A4355FBED0B23" blockId="40.[151,1437,152,1959]" pageId="40" pageNumber="41">
<emphasis id="00A9EADAEF56FF91FF7A4355FEF10B5A" bold="true" box="[197,337,1043,1069]" pageId="40" pageNumber="41">Variability.</emphasis>
Ant IV in females normaly with 5, rarely 6 or even 7 subsegments, in males 7–8 subsegments. Whorl BM2 of Ant IV with 0 to 4 setae. Seta pi2 of fore femur present or absent.
</paragraph>
</subSubSection>
<subSubSection id="7AC76543EF56FF91FF7A4327FF7C08D0" pageId="40" pageNumber="41" type="discussion">
<paragraph id="326236C8EF56FF91FF7A4327FE080A89" blockId="40.[151,1437,152,1959]" pageId="40" pageNumber="41">
<emphasis id="00A9EADAEF56FF91FF7A4327FE980B0C" bold="true" box="[197,312,1121,1147]" pageId="40" pageNumber="41">Remarks</emphasis>
. The new subspecies generally fits to Stach’s (1945) description of
<taxonomicName id="F5DD4D4BEF56FF91FBD84327FAA60B0D" box="[1127,1286,1121,1146]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="00A9EADAEF56FF91FBD84327FAA60B0D" box="[1127,1286,1121,1146]" italics="true" pageId="40" pageNumber="41">P. principalis</emphasis>
</taxonomicName>
(appendices anales, strong but not winged circumanal setae, foot complex, dens chaetotaxy and shape of mucro tip, antennae), although some important features (number and shape of cephalic spines, shape of axial seta ms1 on small abdomen) are not mentioned in original description. Subsequently these important features were described by
<bibRefCitation id="564C4B39EF56FF91FE8443BAFE770A61" author="Gisin" box="[315,471,1276,1302]" pageId="40" pageNumber="45" refString="Gisin, H. (1947) Sur les insectes apterygotes du Parc National Suisse. Especes et groupements euedaphiques. Resultats des recheerches scientifiques enterprises au Parc Natational Suisse, 2, 77 - 91." type="journal article" year="1947">Gisin (1947)</bibRefCitation>
but his figures of mucro and winged circumanal setae contradict with original description; besides, the size of animals from Stach’s and Gisin’s descriptions is quite different (adult female not less than
<quantity id="F5259B2DEF56FF91FE8F420FFED40A14" box="[304,372,1353,1379]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="40" pageNumber="41" unit="mm" value="1.0">1 mm</quantity>
and
<quantity id="F5259B2DEF56FF91FE13420FFDE80A14" box="[428,584,1353,1379]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="40" pageNumber="41" unit="mm" value="0.675" valueMax="0.75" valueMin="0.6">0.6–0.75 mm</quantity>
respectively). Until full redescription of
<taxonomicName id="F5DD4D4BEF56FF91FB98420FFA3C0A14" authority="Stach, 1945" authorityName="Stach" authorityYear="1945" box="[1063,1436,1353,1379]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="00A9EADAEF56FF91FB98420FFB630A15" box="[1063,1219,1353,1378]" italics="true" pageId="40" pageNumber="41">P. principalis</emphasis>
sensu
<bibRefCitation id="564C4B39EF56FF91FAAD420FFA3C0A14" author="Stach" box="[1298,1436,1353,1379]" pageId="40" pageNumber="46" refString="Stach, J. (1945) The species of the genus Arrhopalites occurring in European caves. Acta Musei Historiae Naturalis, Academia Polona Litterarum et Scientiarum, 1, 1 - 47." type="journal article" year="1945">Stach, 1945</bibRefCitation>
</taxonomicName>
(especially such critical features as bifurcated or not axial seta ms1 of small abdomen and number and shape of cephalic spines) we consider subspecies described above as belonging to this species but with different antenna/head ratio (antennae 2 times as long as head instead of 1.3–1.6 times) and female with normally 5 subsegments in Ant IV.
</paragraph>
<paragraph id="326236C8EF56FF91FF7A414DFC7209EE" blockId="40.[151,1437,152,1959]" pageId="40" pageNumber="41">
<taxonomicName id="F5DD4D4BEF56FF91FF7A414DFE640953" box="[197,452,1547,1572]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="principalis" status="subsp. nov." subSpecies="skelicus">
<emphasis id="00A9EADAEF56FF91FF7A414DFE640953" box="[197,452,1547,1572]" italics="true" pageId="40" pageNumber="41">P. principalis skelicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="1B9A57A1EF56FF91FE73414CFD940953" box="[460,564,1546,1572]" pageId="40" pageNumber="41" rank="subSpecies">
<emphasis id="00A9EADAEF56FF91FE73414CFD940953" bold="true" box="[460,564,1546,1572]" pageId="40" pageNumber="41">ssp. nov.</emphasis>
</taxonomicNameLabel>
is also very close to described above
<taxonomicName id="F5DD4D4BEF56FF91FC52414DFB7C0953" box="[1005,1244,1547,1572]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pseudoprincipalis">
<emphasis id="00A9EADAEF56FF91FC52414DFB7C0953" box="[1005,1244,1547,1572]" italics="true" pageId="40" pageNumber="41">P. pseudoprincipalis</emphasis>
</taxonomicName>
but differs from it in following features: 1) broadened tip of mucro; 2) longer antennae; 3) not thickened setae Ipe, IIIpe–Vpe of dens; 4) longer dorsal posterior setae of great abdomen and longer setae of a and m rows of Abd VI; 5) less palmated appendices anales; 6) somewhat greater ratio ‘dens: mucro’.
</paragraph>
<paragraph id="326236C8EF56FF91FF7A41E3FF7C08D0" blockId="40.[151,1437,152,1959]" pageId="40" pageNumber="41">
Specimens belonging to
<taxonomicName id="F5DD4D4BEF56FF91FE5B41E0FDDF09C8" box="[484,639,1702,1727]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="00A9EADAEF56FF91FE5B41E0FDDF09C8" box="[484,639,1702,1727]" italics="true" pageId="40" pageNumber="41">P. principalis</emphasis>
</taxonomicName>
were found also in caves of Chatyr-Dag Massif (
<tableCitation id="7F5F0373EF56FF91FB0541E3FAB209C8" box="[1210,1298,1701,1727]" captionStart="TABLE 1" captionStartId="8.[151,239,151,175]" captionTargetBox="[158,1429,242,593]" captionTargetPageId="8" captionText="TABLE 1. Records of Arrhopalitidae species in the caves of Crimea; (ske) = Pygmarrhopalites principalis skelicus ssp. nov.; (? ssp) =? independent subspecies." httpUri="http://table.plazi.org/id/66A26640EF76FFB1FF2847D1FD9E0FA6" pageId="40" pageNumber="41" tableUuid="66A26640EF76FFB1FF2847D1FD9E0FA6">Table 1</tableCitation>
). However, intraspecific taxonomical position of population from Chatyr-Dag remains unclear: Ipe, IIIpe–Vpe setae of dens thickened as in
<taxonomicName id="F5DD4D4BEF56FF91FE2741B5FD31087B" box="[408,657,1779,1804]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pseudoprincipalis">
<emphasis id="00A9EADAEF56FF91FE2741B5FD31087B" box="[408,657,1779,1804]" italics="true" pageId="40" pageNumber="41">P. pseudoprincipalis</emphasis>
</taxonomicName>
, but body proportions more close to
<taxonomicName id="F5DD4D4BEF56FF91FBE441B5FAC8087B" box="[1115,1384,1779,1804]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="principalis" subSpecies="skelicus">
<emphasis id="00A9EADAEF56FF91FBE441B5FAC8087B" box="[1115,1384,1779,1804]" italics="true" pageId="40" pageNumber="41">P. principalis skelicus</emphasis>
</taxonomicName>
and mucronal apex more swollen than in last subspecies. It is possible that specimens from Chatyr-Dag represent an independent subspecies. The question of geographical variability and intraspecific taxonomical borders of conspecific Crimean populations of
<taxonomicName id="F5DD4D4BEF56FF91FD814021FD7908F7" box="[574,729,1895,1920]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="principalis">
<emphasis id="00A9EADAEF56FF91FD814021FD7908F7" box="[574,729,1895,1920]" italics="true" pageId="40" pageNumber="41">P. principalis</emphasis>
</taxonomicName>
and their relation to
<taxonomicName id="F5DD4D4BEF56FF91FC734021FB1B08F7" box="[972,1211,1895,1920]" class="Entognatha" family="Arrhopalitidae" genus="Pygmarrhopalites" kingdom="Animalia" order="Collembola" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="pseudoprincipalis">
<emphasis id="00A9EADAEF56FF91FC734021FB1B08F7" box="[972,1211,1895,1920]" italics="true" pageId="40" pageNumber="41">P. pseudoprincipalis</emphasis>
</taxonomicName>
requires additional study.
</paragraph>
</subSubSection>
</treatment>
</document>
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