treatments-xml/data/D9/53/DC/D953DC9C1F975E74B5F6D8C40446EEAC.xml
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<document ID-DOI="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" ID-Pensoft-Pub="2511-6428-1-13" ID-Pensoft-UUID="F4C1C6A2DFD95D3BAF5AF1CE09226837" ID-ZooBank="589077DFF8BB45ABB30D7E0DE1C307FE" ModsDocID="2511-6428-74-1-13" checkinTime="1706792588976" checkinUser="pensoft" docAuthor="Irmler, Ulrich" docDate="2024" docId="D953DC9C1F975E74B5F6D8C40446EEAC" docLanguage="en" docName="ContribEntomol 74(1): 13-34" docOrigin="Contributions to Entomology 74 (1)" docPubDate="2024-01-31" docSource="http://dx.doi.org/10.3897/contrib.entomol.74.e114543" docTitle="Lithocharodes andersoni Irmler 2024, sp. nov." docType="treatment" docUuid="8583C739-19A7-4D4A-82D0-B530D880FD4A" docUuidSource="ZooBank" docVersion="1" id="F4C1C6A2DFD95D3BAF5AF1CE09226837" lastPageNumber="13" masterDocId="F4C1C6A2DFD95D3BAF5AF1CE09226837" masterDocTitle="Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)" masterLastPageNumber="34" masterPageNumber="13" pageNumber="13" updateTime="1706792588976" updateUser="pensoft">
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<mods:title id="D35B3727E2F7E844EE4C7C7FE4C9B5B5">Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae)</mods:title>
</mods:titleInfo>
<mods:name id="EA27A5B62C93FD5D2B7FD470608D208A" type="personal">
<mods:role id="9D7A86ADB276D0A1BCF202427BF15A53">
<mods:roleTerm id="11F8C12277C9A71DF56CDBE912A78E2D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D7A7B7B9E0A276E8AC14843A0AA73B68">Irmler, Ulrich</mods:namePart>
<mods:affiliation id="53CAE95349C545A55331A195F46E4012">Institute for Ecosystem Research, University of Kiel, Olshausenstrasse 40, 24098 Kiel, Germany</mods:affiliation>
<mods:nameIdentifier id="44DB59478DCA673811EFA9F6E6EEAED5" type="email">uirmler@ecology.uni-kiel.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="3F704AAD0FB9EF319B13DC794F5A3039">text</mods:typeOfResource>
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<mods:titleInfo id="D7018633E7470525CA4B9AAAF2A66DA6">
<mods:title id="1453F0E9193E2A170325C87AAA380EA7">Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="68DF92002330EE1CB29D2FEBFBBB159D">
<mods:date id="0F67965A070CD9112ED037089F4FCEA2">2024</mods:date>
<mods:detail id="2F7AE6329565B38D05FD43FCD64555B2" type="pubDate">
<mods:number id="ACED261BEF3F8973CB50E6C50A848388">2024-01-31</mods:number>
</mods:detail>
<mods:detail id="AFD9FB48BF7B9002C92D559FA4E753B2" type="volume">
<mods:number id="6E231BD3182EA1216136658A2B169734">74</mods:number>
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<mods:number id="A08ED4408235176C9A3118FC952C8D6D">1</mods:number>
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<mods:start id="5ED2A11E8C7B699AF099ED2EBA8D253D">13</mods:start>
<mods:end id="FFC1B50BF818E7B4EB5FBFA60927433C">34</mods:end>
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<mods:location id="AFC7D7F9ABAA05859D85397F3F6EBD98">
<mods:url id="C96146ACA45295133F03DC3996AD95B6">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:url>
</mods:location>
<mods:classification id="DCDB80F87385D74F6E5CD0A66C6E7A5F">journal article</mods:classification>
<mods:identifier id="697C7C24385C7E5CB9F9320D09BBC07E" type="DOI">http://dx.doi.org/10.3897/contrib.entomol.74.e114543</mods:identifier>
<mods:identifier id="58AA8A987465E7216AF370F34EAFE6B1" type="Pensoft-Pub">2511-6428-1-13</mods:identifier>
<mods:identifier id="2EF1756DF124B57FB03466A86794CCC9" type="ZooBank">589077DFF8BB45ABB30D7E0DE1C307FE</mods:identifier>
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</mods:mods>
<treatment id="D953DC9C1F975E74B5F6D8C40446EEAC" LSID="urn:lsid:zoobank.org:act:8583C739-19A7-4D4A-82D0-B530D880FD4A" httpUri="http://treatment.plazi.org/id/D953DC9C1F975E74B5F6D8C40446EEAC" lastPageNumber="13" pageId="0" pageNumber="13">
<subSubSection id="BB62BB3C3135CDE82F584E2BD9C2E61A" pageId="0" pageNumber="13" type="nomenclature">
<paragraph id="98420E1619805975FF54697695DC9D7B" pageId="0" pageNumber="13">
<taxonomicName id="86FB39FB36B08D40B1AAD93FF85FCC79" LSID="https://zoobank.org/8583C739-19A7-4D4A-82D0-B530D880FD4A" authority="Irmler, 2024" authorityName="Irmler" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Lithocharodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lithocharodes andersoni" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="species" species="andersoni" status="sp. nov.">Lithocharodes andersoni</taxonomicName>
<taxonomicNameLabel id="46284EC46108A9F183D4B48F3CCA175E" pageId="0" pageNumber="13">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="3C5B6CDF88CD489F646AA04D4F7F15FC" pageId="0" pageNumber="13" type="description">
<paragraph id="BB2898A8A279371EB036E661C006176C" pageId="0" pageNumber="13">
<figureCitation id="F7D69B4AB367EFB2ED1B5D467139D1A9" captionStart="Figures 14" captionStartId="F1" captionText="Figures 1 - 4. 1. Lithocharodes andersoni; 2. L. dubia; 3. L. esmeraldae; 4. L. falini. a. Aedeagus; b. Paramere; c. Sternite VII; d. Tergite VII. Scale bars: 0.5 mm (a, c, d); 0.1 mm (b)." figureDoi="10.3897/contrib.entomol.74.e114543.figures1-4" httpUri="https://binary.pensoft.net/fig/975715" pageId="0" pageNumber="13">Figs 1a-d</figureCitation>
<figureCitation id="99613F2D3D05DB9CFFE5DF841A807493" captionStart="Figure 19" captionStartId="F6" captionText="Figure 19. A. Fore-body of Lithocharodes andersoni; B. L. dubia; C. L. esmeraldae; D. L. falini; E. L. hansoni; F. L. hibbsi. Scale bars: 0.5 mm." figureDoi="10.3897/contrib.entomol.74.e114543.figure19" httpUri="https://binary.pensoft.net/fig/975720" pageId="0" pageNumber="13">, 19A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="BF17F76828439113C80FE8B16AF311FB" pageId="0" pageNumber="13" type="materials_examined">
<paragraph id="CFFAE36307EBA9C0A169ECE2E35E25E0" pageId="0" pageNumber="13">Material examined.</paragraph>
<paragraph id="AB01EF4F9462F64A944D281A35DDDB5B" pageId="0" pageNumber="13">
<materialsCitation id="ADDF0163D14A574F99B57D4E1E3A33F5" collectingDate="1995-05-02" collectorName="Berlese, R. Anderson" country="COSTA RICA" elevation="600" latitude="10.966666" location="Pitilla Biological Station" longLatPrecision="21" longitude="-85.416664" specimenCount="1" stateProvince="Guanacaste" typeStatus="Holotype">
<emphasis id="56A9CBA666AA8010916CCDF63116A08D" bold="true" italics="true" pageId="0" pageNumber="13">
<typeStatus id="40E9BCF2A5EB7917816E156BF8F9AF24" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="B4B7DB8208BF90AB1BA50121ADE8053F" name="Costa Rica">COSTA RICA</collectingCountry>
male;
<collectingRegion id="A4CF6B0CD201E8F20FFCBAED8063B875" country="Costa Rica" name="Guanacaste">Guanacaste</collectingRegion>
;
<location id="F1AA435E0A7D55671628946EE6AA0E56" LSID="urn:lsid:plazi:treatment:D953DC9C1F975E74B5F6D8C40446EEAC:F1AA435E0A7D55671628946EE6AA0E56" country="COSTA RICA" latitude="10.966666" longLatPrecision="21" longitude="-85.416664" name="Pitilla Biological Station" stateProvince="Guanacaste">Pitilla Biological Station</location>
;
<quantity id="7EC6BF695A7D659388D994CE6AA627D5" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<elevation id="4C15C50881162B89EACD50A2D37D5AEA" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
</quantity>
elevation;
<geoCoordinate id="3D26BCC977C2165CC5D0BDCE3E98DFFB" degrees="10" direction="north" minutes="58" orientation="latitude" precision="15" seconds="0" value="10.966666">10°58'0&quot;N</geoCoordinate>
,
<geoCoordinate id="F6275619CAA591E12C0846028963AE1D" degrees="85" direction="west" minutes="25" orientation="longitude" precision="15" seconds="0" value="-85.416664">85°25'0&quot;W</geoCoordinate>
;
<collectorName id="9EE9A123C379CE69F6933A6B684219F5">Berlese</collectorName>
; leaf litter,
<collectingDate id="4F1C7474540D739A7E0A4F35B2D74094" value="1995-05-02">2 May 1995</collectingDate>
;
<collectorName id="E9E64F2C4B5A18F05343685A6746BBAC">R. Anderson</collectorName>
leg.; KNHM CR1A95 4B.
</materialsCitation>
</paragraph>
<paragraph id="B8F6EC2A99EE1404F40B8BEB1F09CF69" pageId="0" pageNumber="13">
<materialsCitation id="D00B6B6679A991AAC36DAA2AF19FAADC" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="COSTA RICA" location="COSTA RICA" specimenCount="1" specimenCount-male="1" typeStatus="Paratypes">
<emphasis id="B4FD5BA6CEEF865169DF4F1A21C75380" bold="true" italics="true" pageId="0" pageNumber="13">
<typeStatus id="2B6558E65E1A86F519478609143351AD" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<collectingCountry id="2635617ACCDB5CE4E6EBBAD4D9627D2A" name="Costa Rica">COSTA RICA</collectingCountry>
<specimenCount id="D4A3ABFFA126D368304C71E92B2366FD" count="1" type="male">1 male</specimenCount>
</materialsCitation>
;
<specimenCount id="ECB3E1C04DFCB7ECC56B98906B7572B3" count="3" type="female">3 females</specimenCount>
;
<materialsCitation id="7072BBF401063F81A5E16BE892170DE0" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="Berlese &amp; R. Anderson" country="Costa Rica" elevation="600" latitude="10.966666" location="Pitilla Biological Station" longLatPrecision="21" longitude="-85.416664" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">same data as holotype KNHM CR1A95 4B and 4D</materialsCitation>
;
<specimenCount id="36DB24CBC1440F2F7BDB9CC96A81496B" count="1" type="male">1 male</specimenCount>
;
<materialsCitation id="0E15120AD0A4C52613E05F9BFA6CFF0F" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" collectorName="Berlese &amp; R. Anderson" country="Costa Rica" elevation="600" latitude="10.966666" location="Pitilla Biological Station" longLatPrecision="21" longitude="-85.416664" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">same data; UIC CR1A944B</materialsCitation>
;
<materialsCitation id="5B95233FC4C82A5BC9A31AEF1E32446A" collectingDate="2024-01-01" collectingDateMax="2024-12-31" collectingDateMin="2024-01-01" country="PANAMA" location="PANAMA" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="28692987BDFDA97EDF83C971EEE47674" name="Panama">PANAMA</collectingCountry>
<specimenCount id="FAAC8E76123C26F5088E75623C665145" count="1" type="male">1 male</specimenCount>
</materialsCitation>
;
<materialsCitation id="3F4B41CEA2E92D51BD2CF7C703DF2409" collectingDate="1995-06-05" collectingMethod="flight intercept trap" collectorName="J. Ashe, R. Brooks" country="Panama" elevation="790" latitude="8.733334" location="Capira" longLatPrecision="1300" longitude="-79.95" municipality="Cerro Campana" specimenCount="1" typeStatus="paratype">
<collectingMunicipality id="E76E27D7B9BC2DE194A6649E9DFE6EDD">Cerro Campana</collectingMunicipality>
(
<location id="C11D9E97295C4F5501DE4CD2E8D58519" LSID="urn:lsid:plazi:treatment:D953DC9C1F975E74B5F6D8C40446EEAC:C11D9E97295C4F5501DE4CD2E8D58519" country="Panama" latitude="8.733334" longLatPrecision="1300" longitude="-79.95" municipality="Cerro Campana" name="Capira">Capira</location>
);
<quantity id="0C918595AF2F987C4A6E5D182D9056EB" metricMagnitude="2" metricUnit="m" metricValue="7.9" unit="m" value="790.0">
<elevation id="10E80288327009DEC028EDB567F4622C" metricMagnitude="2" metricUnit="m" metricValue="7.9" unit="m" value="790.0">790 m</elevation>
</quantity>
elevation,
<geoCoordinate id="BFF5854837DCCE16AB0B118AA6CD8783" degrees="8" direction="north" minutes="44" orientation="latitude" precision="925" value="8.733334">8°44'N</geoCoordinate>
,
<geoCoordinate id="AE829A9A0DE6A3B091B2E7C9EA56700E" degrees="79" direction="west" minutes="57" orientation="longitude" precision="925" value="-79.95">79°57'W</geoCoordinate>
;
<collectingMethod id="E3DE9F8176698B61BFBDDF59400DAEE9">flight intercept trap</collectingMethod>
;
<collectingDate id="938A51BDD9D128EADEB280AA2F1CCD54" value="1995-06-05">5 June 1995</collectingDate>
;
<collectorName id="18687E35DC975A173F3A2336015F9D30">J. Ashe</collectorName>
,
<collectorName id="3791AF46EDE6DCCD70C3F288EDBB74BB">R. Brooks</collectorName>
leg.; KNHM #129
</materialsCitation>
;
<specimenCount id="BBADB09ED5CCD4CC440F8C4AA3114F22" count="1" type="male">1 male</specimenCount>
;
<materialsCitation id="6E9302B42CE01EA5B53FE7BB80A9A078" collectingDate="1996-06-06" collectorName="R. Anderson" country="Panama" elevation="1300" latitude="7.7555556" location="Cerro Pirre" longLatPrecision="21" longitude="-77.685" municipality="Estacion Ambiental Cana" specimenCount="1" stateProvince="Darien" typeStatus="paratype">
<normalizedToken id="04F4803A0B23D1F1DC96CD7EAB2A8CA2" originalValue="Darién">
<collectingRegion id="D4CBEC54A9AD9CEBE9C31A60BDEEB013" country="Panama" name="Darien">Darien</collectingRegion>
</normalizedToken>
,
<collectingMunicipality id="1BFE89A186288927B2D80C1666F86DCC">Estacion Ambiental Cana</collectingMunicipality>
;
<location id="7564E2AAC905A69D94D6FDAB58443BCD" LSID="urn:lsid:plazi:treatment:D953DC9C1F975E74B5F6D8C40446EEAC:7564E2AAC905A69D94D6FDAB58443BCD" country="Panama" latitude="7.7555556" longLatPrecision="21" longitude="-77.685" municipality="Estacion Ambiental Cana" name="Cerro Pirre" stateProvince="Darien">Cerro Pirre</location>
;
<geoCoordinate id="F09F58719901B487B2925E9E3434BD7F" degrees="7" direction="north" minutes="45" orientation="latitude" precision="15" seconds="20" value="7.7555556">7°45'20&quot;N</geoCoordinate>
,
<geoCoordinate id="FCBC4EB027C42FF1DA81B6E2A128CD47" degrees="77" direction="west" minutes="41" orientation="longitude" precision="15" seconds="6" value="-77.685">77°41'6&quot;W</geoCoordinate>
;
<quantity id="E26DAE2CA8C592F9DBF1DE4D94BEEFA1" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
<elevation id="2A9136329414195BD26B1E825BA68DCE" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</elevation>
</quantity>
elevation; cloud forest transition litter,
<collectingDate id="2AA06F60229D30A27EC7C8CEFDC85C03" value="1996-06-06">6 VI 1996</collectingDate>
,
<collectorName id="A56D336F60921FBE33E3984D715A4E22">R. Anderson</collectorName>
leg.; KNHM #96-113B
</materialsCitation>
;
<specimenCount id="DC5CE3DE18F110F0DC300816DA5DF0AA" count="1" type="male">1 male</specimenCount>
;
<materialsCitation id="ECAB68615117A01CD8910E5877F03022" collectingDate="1995-05-27" collectingMethod="intercept trap" collectorName="J. Jolly, C. Chaboo" country="Panama" latitude="9.116667" location="Pipeline Rd." longLatPrecision="1300" longitude="-79.75" municipality="Parque Nac. Soberania" specimenCount="1" stateProvince="Colon" typeStatus="paratype">
<collectingRegion id="DCA45E3FBE5A020643BF1C431D68E346" country="Panama" name="Colon">Colon</collectingRegion>
,
<collectingMunicipality id="FEC5178BF64818A868B01B4AFD3719AC">Parque Nac. Soberania</collectingMunicipality>
;
<location id="6F4318A8F1286F2C7224C39441005F8A" LSID="urn:lsid:plazi:treatment:D953DC9C1F975E74B5F6D8C40446EEAC:6F4318A8F1286F2C7224C39441005F8A" country="Panama" latitude="9.116667" longLatPrecision="1300" longitude="-79.75" municipality="Parque Nac. Soberania" name="Pipeline Rd." stateProvince="Colon">Pipeline Rd.</location>
;
<geoCoordinate id="D1FF9780B4025521C2180D093A88BD31" degrees="9" direction="north" minutes="07" orientation="latitude" precision="925" value="9.116667">9°07'N</geoCoordinate>
,
<geoCoordinate id="2DAA224D24D19CFD1875BE63D667FCEF" degrees="79" direction="west" minutes="45" orientation="longitude" precision="925" value="-79.75">79°45'W</geoCoordinate>
; flt.
<collectingMethod id="72A26245EDB58F67ACE815CA255B5FEC">intercept trap</collectingMethod>
;
<collectingDate id="E14AA8C8777B1C2BEDDE29DE811AE2CD" value="1995-05-27">27 May 1995</collectingDate>
;
<collectorName id="40B0DE770A70FACB48989DB0C9048BA4">J. Jolly</collectorName>
,
<collectorName id="0CCF623DEC53201BB899983C9A70C15B">C. Chaboo</collectorName>
leg.; KNHM
</materialsCitation>
;
<specimenCount id="3E625E864362759317A3E701063F8595" count="1" type="male">1 male</specimenCount>
;
<specimenCount id="FBF8CD2B7326F80345A617F4CEFCEA2E" count="1" type="female">1 female</specimenCount>
;
<materialsCitation id="40ED8D31546110928174EE6F59DEEDC5" collectingDate="1995-05-31" collectingDateMax="1995-06-02" collectingDateMin="1995-05-31" collectingMethod="flight intercept trap" collectorName="J. Ashe, R. Brooks" country="Panama" latitude="9.116667" location="Pipeline Road" longLatPrecision="1300" longitude="-79.75" municipality="Parque Nac. Soberania" specimenCount="1" stateProvince="Colon" typeStatus="paratype">
same location;
<location id="E8C5069F5EE3CE8EBCD7779AA6A9175F" LSID="urn:lsid:plazi:treatment:D953DC9C1F975E74B5F6D8C40446EEAC:E8C5069F5EE3CE8EBCD7779AA6A9175F" country="Panama" latitude="9.116667" longLatPrecision="1300" longitude="-79.75" name="Pipeline Road" stateProvince="Colon">Pipeline Road</location>
km 5.3;
<collectingMethod id="77D6F742A78B5A60B3444499ABAC35E7">flight intercept trap</collectingMethod>
;
<collectingDate id="71E7CCF12B31C8859306C67D462E336F" value="1995-05-31" valueMax="1995-06-02" valueMin="1995-05-31">31 May-2 June 1995</collectingDate>
;
<collectorName id="EC4CA1FD2B40EDA3B1A3D13C97ADC15E">J. Ashe</collectorName>
,
<collectorName id="CE0902B68355358254176B2EF5850785">R. Brooks</collectorName>
leg.; KNHM #098
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="6EADCEF07CEF12869EF8B6E68F60B070" pageId="0" pageNumber="13" type="diagnosis">
<paragraph id="6556D6BFC0369B9CBEB4C11634130642" pageId="0" pageNumber="13">Diagnosis.</paragraph>
<paragraph id="3BA92A2903430775F724CCD2F00D4844" pageId="0" pageNumber="13">
According to the structure of the aedeagus with absent sclerotised endophallus, the species is most similar to the
<taxonomicName id="E98075C88DE2D099D45FA6FEC6F8064C" lsidName="L. hanagarthi" pageId="0" pageNumber="13" rank="species" species="hanagarthi">
<emphasis id="9323819C4ECB6B30BDD41B09A6A187F0" italics="true" pageId="0" pageNumber="13">L. hanagarthi</emphasis>
</taxonomicName>
-group that is characterised by a short transparent endophallus. In contrast to the bicoloured other two species of the group,
<taxonomicName id="0BAED2803F425FE184BFDF2F3E11AB83" lsidName="L. andersoni" pageId="0" pageNumber="13" rank="species" species="andersoni">
<emphasis id="71B3B1C6D484C4A7569529763879B56E" italics="true" pageId="0" pageNumber="13">L. andersoni</emphasis>
</taxonomicName>
is unicoloured as
<taxonomicName id="89A2C9C1DBE2D280922242E3619543AC" lsidName="L. boliviana" pageId="0" pageNumber="13" rank="species" species="boliviana">
<emphasis id="BC4F7315284713F352858A51E132717A" italics="true" pageId="0" pageNumber="13">L. boliviana</emphasis>
</taxonomicName>
Irmler, 2021 and not bicoloured as
<taxonomicName id="98BA9D62BDF7BA146D193F7AF54A033E" lsidName="L. hanagarthi" pageId="0" pageNumber="13" rank="species" species="hanagarthi">
<emphasis id="C00E1C9B2E632EE06E178A0EE7A36B10" italics="true" pageId="0" pageNumber="13">L. hanagarthi</emphasis>
</taxonomicName>
<bibRefCitation id="94CBD667FD2BDC29A1273418A0FDC37D" DOI="https://doi.org/10.21248/contrib.entomol.71.1.029-085" author="Irmler, U" journalOrPublisher="Contributions to Entomology" pageId="0" pageNumber="13" pagination="29 - 86" refId="B3" refString="Irmler, U, 2021. The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini). Contributions to Entomology 71: 29 - 86, DOI: https://doi.org/10.21248/contrib.entomol.71.1.029-085" title="The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)." url="https://doi.org/10.21248/contrib.entomol.71.1.029-085" volume="71" year="2021">Irmler 2021</bibRefCitation>
. It differs from
<taxonomicName id="E78837C26875343B3E4026B1BA40CEFF" lsidName="L. boliviana" pageId="0" pageNumber="13" rank="species" species="boliviana">
<emphasis id="52D116BF33670A9FEE61023FAB48971D" italics="true" pageId="0" pageNumber="13">L. boliviana</emphasis>
</taxonomicName>
by the much sparser punctation of the head.
</paragraph>
</subSubSection>
<subSubSection id="02BBE617F31ABA6EA98733645C639921" pageId="0" pageNumber="13" type="description">
<paragraph id="81BBE4DECA71545D61F1E8755FCB30C6" pageId="0" pageNumber="13">Description.</paragraph>
<paragraph id="331B69336A9307F70E2E169939FE6E5B" pageId="0" pageNumber="13">Length: 5.3 mm; Colouration: blackish, legs light brown yellowish, antennae darker light brown.</paragraph>
<paragraph id="74F1097D17941675A19EC6EAA31FBE5E" pageId="0" pageNumber="13">Head: 0.82 mm long, 0.67 mm wide; eyes not prominent; PS:E ratio 3.73; postocular sides smoothly curved to rounded posterior angles; without distinct posterior angles; interantennal furrows reaching anterior edge of eyes; centrally between eyes with indistinct depression; setiferous punctation scarce and moderately deep; distance between punctures irregular; on average, two to three times as wide as diameter of punctures; without microsculpture; surface polished; antennae with first antennomere as long as half-length of head; second and third antennomere conical; combined half as long as half-length of first antennomere; following antennomeres transverse; on average, twice as wide as long; slightly increasing in width; all antennomeres pubescent. Pronotum: 1.04 mm long, 0.63 mm wide; only slightly wider in anterior third; widest part at anterior third; slightly narrowed to posterior angles; posterior angles obtuse; posterior edge smoothly rounded; setiferous punctation moderately deep and dense; along impunctate mid-line with irregular row of 10-15 punctures; on average, interstices between punctures irregular; at least as wide as diameter of punctures; surface without microsculpture; polished. Elytra: 0.95 mm long, 0.85 mm wide; humeral angles obtuse; sides posteriorly slightly divergent; posterior angles sub-rectangular; posterior margin slightly retreated to suture; setiferous punctation dense and moderately deep; on average, interstices between punctures as wide as diameter of punctures; surface without microsculpture; polished. Abdomen finely and moderately densely punctate; surface without microsculpture; polished; sternite VII of male with straight posterior margin; slightly emarginate at outer angles; tergite VII of male straight. Meso- and meta-tibia with four ctenidia. Aedeagus widest near anterior edge; anterior edge straight; slightly convergent posteriorly; without distinct endophallus; parameres thick; longer than half-length of central lobe; C:A ratio 0.5; slightly narrowed anteriorly; in anterior third abruptly narrowed; along inner edge and in anterior half with numerous sensillae.</paragraph>
</subSubSection>
<subSubSection id="D9CCB6E5A3529F083885B5FC61A5119E" pageId="0" pageNumber="13" type="etymology">
<paragraph id="C7CD9EDA2D9E994B6D4881A8BF82D638" pageId="0" pageNumber="13">Etymology.</paragraph>
<paragraph id="EE9D0B08FBF9EC959AAE81926E3967BC" pageId="0" pageNumber="13">
The species name honours the collector of the species Richard Anderson, who sampled
<taxonomicName id="4B23F7A88C9B909FF04A65E128CDFF98" class="Hexapoda" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="13" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
extensively in the Neotropical Region.
</paragraph>
</subSubSection>
<subSubSection id="629254092BE20918392E9DFE2769A2B2" pageId="0" pageNumber="13" type="geography">
<paragraph id="395E590EC82DF6AF8CC97BC039E41525" pageId="0" pageNumber="13">Geography.</paragraph>
<paragraph id="0167350F1C3C71CA66C20FF82893A8CA" pageId="0" pageNumber="13">Costa Rica and Panama.</paragraph>
</subSubSection>
<subSubSection id="63286013B7E7914BEEF005124D61141A" pageId="0" pageNumber="13" type="ecology">
<paragraph id="E8A7C48849D744258490C142E6A1AE10" pageId="0" pageNumber="13">Ecology.</paragraph>
<paragraph id="C31486ADE1FECFA64E851B2081672E33" pageId="0" pageNumber="13">Floor of montane forests and lower cloud forests in May and June.</paragraph>
</subSubSection>
</treatment>
</document>