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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.505.9862" ID-GBIF-Dataset="2e13b190-dff9-41bd-b2db-d6f2b4c82731" ID-PMC="PMC4453233" ID-Pensoft-Pub="1313-2970-505-1" ID-PubMed="26052236" ID-ZBK="7F1C641D389940BD8E9B1F812D4509D1" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-505-1" ModsDocOrigin="ZooKeys 505" ModsDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910, in China, with descriptions of new cavernicolous species (Diplopoda, Polydesmida, Haplodesmidae)" checkinTime="1451244309637" checkinUser="pensoft" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauries, Jean-Paul &amp; VandenSpiegel, Didier" docDate="2015" docId="570EDBE91E7F15FEAC2BB7047CFC7B06" docLanguage="en" docName="ZooKeys 505: 1-34" docOrigin="ZooKeys 505" docSource="http://dx.doi.org/10.3897/zookeys.505.9862" docTitle="Eutrichodesmus troglobius Golovatch, Geoffroy, Mauries &amp; VandenSpiegel, sp. n." docType="treatment" docUuid="BA529FEC-19CB-46E2-867D-B3621667978E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="22" masterDocId="FFCBFF9AFFE6FFEDFFA5644EFF808470" masterDocTitle="Review of the millipede genus Eutrichodesmus Silvestri, 1910, in China, with descriptions of new cavernicolous species (Diplopoda, Polydesmida, Haplodesmidae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="20" updateTime="1668160367798" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Review of the millipede genus Eutrichodesmus Silvestri, 1910, in China, with descriptions of new cavernicolous species (Diplopoda, Polydesmida, Haplodesmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Geoffroy, Jean-Jacques</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mauries, Jean-Paul</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>VandenSpiegel, Didier</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>505</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>34</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.505.9862</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.505.9862</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-505-1</mods:identifier>
<mods:identifier type="ZBK">7F1C641D389940BD8E9B1F812D4509D1</mods:identifier>
<mods:identifier type="ZooBank">7F1C641D389940BD8E9B1F812D4509D1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152060117" LSID="urn:lsid:zoobank.org:act:BA529FEC-19CB-46E2-867D-B3621667978E" httpUri="http://treatment.plazi.org/id/570EDBE91E7F15FEAC2BB7047CFC7B06" lastPageId="22" lastPageNumber="22" pageId="19" pageNumber="20">
<subSubSection pageId="19" pageNumber="20" type="multiple">
<paragraph pageId="19" pageNumber="20">
<pageBreakToken pageId="19" pageNumber="20" start="start">Taxon</pageBreakToken>
classification Animalia Polydesmida Haplodesmidae
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="nomenclature">
<paragraph pageId="19" pageNumber="20">
<taxonomicName LSID="http://zoobank.org/BA529FEC-19CB-46E2-867D-B3621667978E" authority="Golovatch, Geoffroy, Mauries &amp; VandenSpiegel" class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus troglobius" order="Polydesmida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="troglobius">
Eutrichodesmus troglobius Golovatch, Geoffroy,
<normalizedToken originalValue="Mauriès">Mauries</normalizedToken>
&amp; VandenSpiegel
</taxonomicName>
<taxonomicNameLabel pageId="19" pageNumber="20">sp. n.</taxonomicNameLabel>
Figs 20, 21
</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="type material">
<paragraph pageId="19" pageNumber="20">Type material.</paragraph>
<paragraph pageId="19" pageNumber="20">
Holotype ♂ (MNHN JC 375), China, Guizhou Prov., Kaiyang, Cave Xianyan Dong, 19.II.2004, leg. S.
<normalizedToken originalValue="Prevorčnik">Prevorcnik</normalizedToken>
&amp; B. Sket.
</paragraph>
<paragraph pageId="19" pageNumber="20">Paratypes: 1 ♂, 2 ♀, 2 subadult ♀ (MNHN JC 375), 1 ♀ (SEM), same data, together with holotype.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="name">
<paragraph pageId="19" pageNumber="20">Name.</paragraph>
<paragraph pageId="19" pageNumber="20">To emphasize cavernicoly; adjective.</paragraph>
</subSubSection>
<subSubSection pageId="19" pageNumber="20" type="diagnosis">
<paragraph pageId="19" pageNumber="20">Diagnosis.</paragraph>
<paragraph pageId="19" pageNumber="20">Differs from congeners by the relatively broad and modestly declivous paraterga which are set low on the body at about 45° to the vertical axis and distinctly discontinue the more strongly declined outline of the sides above paraterga, coupled with low, but distinct, mostly longitudinally oblong, metatergal tuberculations arranged in three transverse irregular rows, and the rather simple gonopod acropodite which only shows a small distodorsal tooth (see also Key below).</paragraph>
</subSubSection>
<subSubSection lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="description">
<paragraph pageId="20" pageNumber="21">
<pageBreakToken pageId="20" pageNumber="21" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="20" pageNumber="21">Length of adults ca 8-9 mm, width 1.0 and 1.9-2.0 mm on midbody pro- and metazonae, respectively. Holotype ca 9 mm long, 1.0 and 2.0 mm wide on pro- and metazonae, respectiverly. Coloration entirely pallid.</paragraph>
<paragraph pageId="20" pageNumber="21">
All characters as in
<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus triangularis" order="Polydesmida" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="triangularis">Eutrichodesmus triangularis</taxonomicName>
sp. n., except as follows.
</paragraph>
<paragraph lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
Body with 20 segments (♂, ♀), conglobation pattern typical of
<normalizedToken originalValue="“doratodesmoids”">&quot;doratodesmoids&quot;</normalizedToken>
, volvation apparently being incomplete because of broad and modestly declivous paraterga. Antennae rather long and poorly clavate (Fig. 20H, L). Collum not covering the head from above, fore margin clearly lobulate and slightly elevated, with abundant distinct tuberculations arranged in regular rows, but lobulations observed only at anterior margin. Metaterga behind collum with three transverse, rather irregular, mixostictic
<pageBreakToken pageId="21" pageNumber="22" start="start">rows</pageBreakToken>
of similarly distinct, longitudinally oblong, setigerous tuberculations extending onto paraterga, usually about 8-10+8-10 per row (Fig. 20
<normalizedToken originalValue="AF">A-F</normalizedToken>
); mid-dorsal regions of metaterga not elevated; caudomarginal lobulations numerous, 2-3 more evident ones only on paraterga (Fig. 20
<normalizedToken originalValue="AF">A-F</normalizedToken>
); limbus microcrenulate (Fig. 20M). Paraterga with evident shoulders anteriorly, broad, modestly declivous, directed ventrolaterad at about 45° to even more strongly declined sides above paraterga (Fig. 20N), tips lying clearly below level of venter, usually distinctly trilobate laterally; anterolaterals evident only in segment 2 (Fig. 20A, D). Paraterga 2 strongly enlarged, directed ventrad (Fig. 20A, D, H), lateral margin broadly rounded, with few, but very distinct lobulations; a full row of similarly large caudolaterals located above schism, both schism and hyposchism being small (Fig. 20A). Tergal setae short, 2-segmented, apical part setoid (Fig. 20G). Pore formula apparently normal, ozopores indistinct. Epiproct strongly flattened dorsoventrally and tuberculate dorsally (Fig. 20C, F, J). Hypoproct subtrapeziform (Fig. 20J).
</paragraph>
<caption pageId="21" pageNumber="22">
<paragraph pageId="21" pageNumber="22">
Figure 20.
<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus troglobius" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="troglobius">Eutrichodesmus troglobius</taxonomicName>
sp. n., ♀ paratype; A, D, H anterior part of body, lateral, dorsal and ventral views, respectively B, E, I midbody segments, lateral, dorsal and ventral views, respectively C, F, J posterior part of body, lateral, dorsal and ventral views, respectively G tergal seta, subdorsal view K midbody paratergite, lateral view L head, ventral view M limbus and prozonite texture, dorsal view N cross-section of a midbody segment, caudal view. Scale bars: 0.5 mm (I, J), 0.2 mm (
<normalizedToken originalValue="AF">A-F</normalizedToken>
, H, N), 0.1 mm (L), 0.02 mm (M).
</paragraph>
</caption>
<paragraph pageId="21" pageNumber="22">Sterna usually with a rather deep, narrow depression between coxae (Fig. 20I, J). Legs long and slender, 1.1-1.2 times as long as body height (Fig. 20I, J, N), only coxae and most surface of prefemora finely micropapillate (Fig. 21A).</paragraph>
<caption pageId="21" pageNumber="22">
<paragraph pageId="21" pageNumber="22">
Figure 21.
<taxonomicName class="Diplopoda" family="Haplodesmidae" genus="Eutrichodesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrichodesmus troglobius" order="Polydesmida" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="troglobius">Eutrichodesmus troglobius</taxonomicName>
sp. n., ♂ paratype; A leg 9, lateral view B, C left gonopod, mesal and lateral views, respectively. Scale bar: 0.2 mm. Designations in text.
</paragraph>
</caption>
<paragraph lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
Gonopods (Fig. 21B, C) rather simple. Coxae subquadrate, large, micropapillate and setose mostly on lateral face, with a small, subtriangular, setigerous tooth caudola
<pageBreakToken pageId="22" pageNumber="23" start="start">terally</pageBreakToken>
. Telopodite considerably longer than coxite, moderately and regularly curved ventrad, setose nearly over its basal half until base of a prominent, curved, subspiniform, abundantly microtuberculate, distofemoral process (dp), the latter situated at about basal 1/3 of telopodite, acropodite twisted, with a small, subapical, ventral tooth (d); seminal groove terminating subapically; a hairpad wanting.
</paragraph>
</subSubSection>
</treatment>
</document>