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<document ID-CLB-Dataset="229647" ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-GBIF-Dataset="6ba6d2ca-5dad-4270-914f-1042327d503c" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="5349EAC2B36F57B1AD74C62925B85724" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana asymmetrica Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="9B4393E3-772A-4CF6-A99E-F3A3313E8582" docUuidSource="ZooBank" docVersion="5" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1732669275789" updateUser="ExternalLinkService">
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<mods:title id="2A2787028DD03300254368A321617FA7">Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:name id="FAC06B1267D3C8B31E85A3F261490D31" type="personal">
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<mods:namePart id="94B08994F885A2FC27A70F6EED3BD291">Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier id="375464259C695C234900A2A6A37B5AB1" type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation id="1D41096289A5D862F5CFB4B71430AC5A">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier id="F5D0EADAED1097C4146D526C024A585D" type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="D12233794BE0311AAFA690BDFE60D82E">Cavichioli, Rodney Ramiro</mods:namePart>
<mods:nameIdentifier id="365EA8159B0BDAEF5F4E222653C3FBE0" type="ORCID">https://orcid.org/0000-0001-5996-0629</mods:nameIdentifier>
<mods:affiliation id="63E852712E905799B556EA6A175F3045">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:title id="2E52DE6FC213C15EC1DCD541808FB1A1">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date id="687AB9E8932C6DB0CF3E903132128154">2023</mods:date>
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<treatment id="5349EAC2B36F57B1AD74C62925B85724" ID-GBIF-Taxon="209013706" LSID="urn:lsid:zoobank.org:act:9B4393E3-772A-4CF6-A99E-F3A3313E8582" httpUri="http://treatment.plazi.org/id/5349EAC2B36F57B1AD74C62925B85724" lastPageNumber="79" pageId="0" pageNumber="79" scope_family="Cicadellidae" scope_order="Hemiptera" scope_tribe="Gyponini">
<subSubSection id="3113AD18CC61FC0BF2E2A37254A5D472" pageId="0" pageNumber="79" type="nomenclature">
<paragraph id="561E4BF268FAEE7C28C05AF2ADFBDF47" pageId="0" pageNumber="79">
<taxonomicName id="982BAA6577AE2CE14727F7CB906AB788" LSID="https://zoobank.org/9B4393E3-772A-4CF6-A99E-F3A3313E8582" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana asymmetrica" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="asymmetrica" status="sp. nov.">Coarctana asymmetrica</taxonomicName>
<taxonomicNameLabel id="0BBAD75142E3EE1D50E9C794B9222EB0" pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="27381E2DAECA03AB2CE9E63EB89BB58E" pageId="0" pageNumber="79" type="description">
<paragraph id="58CD64851648D98F4C20DF0B6F6C56B2" pageId="0" pageNumber="79">
<figureCitation id="F931651B48CE999AC0BBD0A27D351CF1" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">Figs 18</figureCitation>
<figureCitation id="1D222F19E2F6E24098B780B1E55D0D0C" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">, 31C, D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="FD54270E9D94118DA83CA9198636E2DC" pageId="0" pageNumber="79" type="diagnosis">
<paragraph id="0CFCF190B7A6A75B7FA8530B9599AAB8" pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph id="9A115D4A8AF9C2358231A34BAC12E100" pageId="0" pageNumber="79">
Head (Fig.
<figureCitation id="CFBE48A2EF8603C93E82ACB9817DAABD" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
) with portion between ocelli and anterior margin dark-brown. Pygofer (Fig.
<figureCitation id="18570C6561F4CB56C74503D6AB46E73B" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18G</figureCitation>
) with apex excavated. Subgenital plate (Fig.
<figureCitation id="2C2B9CB0313CEDE0AAEDFBF98F71D2E6" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18H</figureCitation>
) with long filiform setae. Style (Fig.
<figureCitation id="C098DA53A7D39EFDF478142F33DDF2F0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18K</figureCitation>
) apex truncated, with one process directed dorsally and one ventrally, both with equivalent sizes and acute. Aedeagus (Fig.
<figureCitation id="F945C05D9CD6F06770F31F054D7EA88F" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18L, M</figureCitation>
) with ventral margin of apodemal process forming rounded lobe produced ventrally; shaft with pair of processes directed ventrally: left process short, about 1/4 length of shaft, right process longer, about 2/3 length of shaft and directed to left side.
</paragraph>
</subSubSection>
<subSubSection id="736932A11A2729C94FF51DB1F77419E3" pageId="0" pageNumber="79" type="description">
<paragraph id="01663350CC713D6A17689C54C63515BA" pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph id="F34E2F03C275C997CD489A507F4336D7" pageId="0" pageNumber="79">Total length: holotype male 7.6 mm.</paragraph>
</subSubSection>
<subSubSection id="16F5DFCF992E6A0D2C36AEF19A13D26A" pageId="0" pageNumber="79" type="description">
<paragraph id="70A7013992A297ADB20027D6D3252CDB" pageId="0" pageNumber="79">Description.</paragraph>
<paragraph id="0BC6B02B3D65F84A598B53E586730723" pageId="0" pageNumber="79">
Head, in frontal view (Fig.
<figureCitation id="28BBDB414332CB53DBAD0AB53AADD644" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18B</figureCitation>
), frons 1.2
<normalizedToken id="E5CE88EA1513DF47AF1FE103FC38E6BC" originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by approximately half maximum width of clypeus; clypeus 1.2
<normalizedToken id="E7FCF17448209EDE0C829E0DFF283392" originalValue="×">x</normalizedToken>
wider than long; lateral margins straight, slightly convergent apically; apex straight. Head, in lateral view (Fig.
<figureCitation id="7C1D9902AD1D13A864EEC5B3C2FDDDA7" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18C</figureCitation>
), crown-face transition thin, with 3 transverse carinae; clypeus not inflated. Forewing (Fig.
<figureCitation id="28F092E86C3B39E4C8444D73A134F469" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18D</figureCitation>
) with apex of anal veins bifid. Profemur AV row with 5 setae; PV row with 1 seta near apex. Protibia PD row with 3 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 22-24, 13 and 15 macrosetae respectively. Metatarsomere I 4
<normalizedToken id="B775F687FBFD0692CDA19C1F96D3ADB1" originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 5 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection id="9E54F5503DC825E3593312EEDE9FF4E5" pageId="0" pageNumber="79" type="coloration">
<paragraph id="87217DA412283952C1ECC9A465F4B505" pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph id="88255160F646288A5C58BCDA0503ED1B" pageId="0" pageNumber="79">
Head and thorax (Figs
<figureCitation id="A01D41CA1F3A6A19CF26A776A123A259" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A-D</figureCitation>
,
<figureCitation id="9A96017D2028D892F2448B27D58D8129" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31C, D</figureCitation>
) yellowish-brown. Head (Fig.
<figureCitation id="2D449A1182CF2DB0486320DB271212F5" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
) with pair of small circular black spots, near posterior margin, behind ocelli; portion between ocelli dark brown. Face (Fig.
<figureCitation id="058D218EF33830D5AD6901C034E0E220" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18B</figureCitation>
) yellowish; antennal pit dark brown. Pronotum (Fig.
<figureCitation id="424C4872EF6ECE6533F5953ED24F6E46" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
) with black punctures; small black maculae near anterior margin, behind eye and ocellus; proepimeron (Fig.
<figureCitation id="9407DA757447D0968E4F25B0C1DD49FF" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18C</figureCitation>
) black, except ventral third yellow. Mesonotum (Fig.
<figureCitation id="8429B55370919DB1C96D3ACEDDD1DAC4" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18A</figureCitation>
) reddish-brown. Scutellum yellowish. Forewing (Fig.
<figureCitation id="31EF0A48E03B32E292F77E0ED736FAA3" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18D</figureCitation>
) with brown veins, larger dark-brown maculae on apex of anal veins, outer discal cell, costal margin, cross veins of discal and apical cells; transverse brown stripe over subapical cells; apical cells smoky apically. Metatibia (Fig.
<figureCitation id="2A1BF809513D8E00BD51FA5F58202AE7" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection id="DCA0AC5E3C68DBFD37327E1342701DEF" pageId="0" pageNumber="79" type="male terminalia">
<paragraph id="99D945B30C560CEA364442DDF8841DC1" pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph id="EF029E34E524E1AEE6D1C52D19813C26" pageId="0" pageNumber="79">
Sternite VIII (Fig.
<figureCitation id="AFED03613FB822E58C86E155DF3B4856" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18E</figureCitation>
) 1.5
<normalizedToken id="E51518A201B5BD3DE981590DB85413E8" originalValue="×">x</normalizedToken>
wider than long; lateral margins weakly convergent posterad; posterior margin straight. Valve (Fig.
<figureCitation id="FA559E1B33A7738DA2BCE23A57D93D7D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18F</figureCitation>
) 1.8
<normalizedToken id="C39196408A3A304989F8C1C6BFD0BFD3" originalValue="×">x</normalizedToken>
wider than long; posterior margin rounded. Pygofer, in lateral view (Fig.
<figureCitation id="0524F896DB181D6734E12A8F09B03CA5" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18G</figureCitation>
), 1.6
<normalizedToken id="9E066CFA8F21780ACD704AD3B0F4BB33" originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process thin and short, extending ventrally for 1\5 height of lateral lobe; ventral margin rounded, posterodorsal and posteroventral margins straight and weakly convergent posterad; apex broad and excavated, forming two rounded tips: the dorsal wider than the ventral. Subgenital plate, in lateral view (Fig.
<figureCitation id="9C99A0C59428484059EB86C62C082BD4" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18G</figureCitation>
), not reaching pygofer apex; in ventral view (Fig.
<figureCitation id="C22F5F1703A8487E77F2034083B7C35E" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18H</figureCitation>
), elongated, 3.3
<normalizedToken id="CDB729B11A2015C7ED6630011C004813" originalValue="×">x</normalizedToken>
longer than wide, maximum width near median third; long filiform setae on ventral surface near outer margin and dorsal surface near base; lateral margins rounded; apex tapered, subacute. Connective (Fig.
<figureCitation id="059CDAA1A54714ACB74B576B1ED029B8" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18I</figureCitation>
) about as wide as long; anterior margin excavated; dorsal keel reduced; stem short. Style, in dorsal view (Fig.
<figureCitation id="D261CD1565787CDBD6DE93CD9EC83D5C" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18J</figureCitation>
), with outer lobe developed and rounded; in lateral view (Fig.
<figureCitation id="1BAC1A0D39AD7219657BC6397CCF5383" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18K</figureCitation>
), blade narrow, slightly wider on apical half; dorsal margin almost straight; ventral margin with basal half straight and apical half slightly rounded and serrated; apex truncated, with one process directed dorsally and one ventrally, both with equivalent sizes. Aedeagus (Fig.
<figureCitation id="869336A6EBA787513E66B4A0B84A81C0" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18L, M</figureCitation>
) preatrium weakly developed; dorsal apodeme with dorsal margin straight, lateral margins not produced laterally; apodemal processes narrow basally, curved in right angle dorsally in the basal third; ventral margin forming rounded lobe produced ventrally; distal portion straight, apex weakly broadened, curved over the apex of shaft; shaft approximately cylindrical, strongly curved dorsally near base, with small basolateral expansion that surrounds the previous portion of shaft; apex with pair of processes directed ventrally: left process short, about 1/4 length of shaft, right process longer, about 2/3 length of shaft and directed to left side. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection id="70EB4F595504D84E0975A43EABA17FC2" pageId="0" pageNumber="79" type="female">
<paragraph id="F9308396184DAEC7695D601A57D5CDD0" pageId="0" pageNumber="79">Female.</paragraph>
<paragraph id="7A505F9E7D1C909A75195717538B13C2" pageId="0" pageNumber="79">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C4597686752EDDA31388B70CAE809B80" pageId="0" pageNumber="79" type="materials_examined">
<paragraph id="B93C9F71F868C6DAD2A0DF9C86A46159" pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph id="DB3EE268D63433A2DDE164CFC03073B3" pageId="0" pageNumber="79">
<materialsCitation id="945F11CB19B317095423E8D956286248" ID-GBIF-Occurrence="4017211339" collectingDate="1984-11-20" collectorName="C. Elias" country="Brazil" location="Caceres" specimenCount="1" stateProvince="Mato Grosso" typeStatus="Holotype">
<emphasis id="C8C3925ED2B21D6DC9E81E9E4E8F42E1" bold="true" pageId="0" pageNumber="79">
<typeStatus id="5477E34AA5CC2B756529F2C1CDF5EF18">Holotype</typeStatus>
male:
<collectingCountry id="D2AE481F9D32C61BF3756EBF306AEC55" name="Brazil">Brazil</collectingCountry>
:
<collectingRegion id="A1E355FF34FA9411435044406015A65B" country="Brazil" name="Mato Grosso">Mato Grosso</collectingRegion>
</emphasis>
:
<normalizedToken id="DAA57575F1F1039AAF83C88EC5166CA1" originalValue="“Cáceres">
&quot;
<location id="13B56F5D68FA63AE16CE914F3682D798" LSID="urn:lsid:plazi:treatment:5349EAC2B36F57B1AD74C62925B85724:13B56F5D68FA63AE16CE914F3682D798" country="Brazil" name="Caceres" stateProvince="Mato Grosso">Caceres</location>
</normalizedToken>
, MT\
<collectingDate id="F7E18539D831BDB533688E00447D0171" value="1984-11-20">20.XI.1984</collectingDate>
\
<collectorName id="E9F97D4B9CD94D4A1CE39E21FC712194">C. Elias</collectorName>
leg.\ Polonoroeste&quot; (DZUP).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="978B6B4466A5052905376FEDBFEB112F" pageId="0" pageNumber="79" type="etymology">
<paragraph id="7F8ECC3EC119691AE0AE04B749A8EA63" pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph id="4C054F5CBA268D084C3AC457682F6947" pageId="0" pageNumber="79">The new species name refers to asymmetric apical processes of the aedeagus.</paragraph>
</subSubSection>
<subSubSection id="976611643E5479F296AE9A3240C418FD" pageId="0" pageNumber="79" type="remarks">
<paragraph id="ED44F59B5A7AD5AFB9E8043A58526F34" pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph id="8B661292782F161FFE9A0C03DE6A3376" pageId="0" pageNumber="79">
<taxonomicName id="9DC661D2E75F8BA9D5012AD6F097E909" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana asymmetrica" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="asymmetrica">
<emphasis id="A1913E5E8B0B9EDF0D5F6C62BFDAA72A" italics="true" pageId="0" pageNumber="79">Coarctana asymmetrica</emphasis>
</taxonomicName>
<emphasis id="4E3EF5126045ED85EF533926329A9B1F" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
was recovered as sister group to
<taxonomicName id="528B7E8930728259E677FA0A2C7B1FC9" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. nigromedia" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="nigromedia">
<emphasis id="0A960BAEFD7CD5975361A5A837ED26D6" italics="true" pageId="0" pageNumber="79">C. nigromedia</emphasis>
</taxonomicName>
<emphasis id="FCB8EBBF6E728EB77C5DA95705143A22" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
with moderate branch support (SR = 67) (Fig.
<figureCitation id="ADEA1E6E7D23B28235B1E2BFE4A8F9F1" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
), both of them have the pygofer apex excavated (Fig.
<figureCitation id="0DF24F9E07FCEB034B1B930212B0F473" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18G</figureCitation>
). The new species is easily differentiated by having the style (Fig.
<figureCitation id="5D92D1463A32F4FA7D46B0996D1EA9C8" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18K</figureCitation>
) with apex truncated, bearing one small process directed dorsally and one ventrally, both similar in size; and in having the aedeagal shaft (Fig.
<figureCitation id="A53D7A61ECF5355F56B438A4BCB59E9D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Coarctana asymmetrica sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure18" httpUri="https://binary.pensoft.net/fig/798171" pageId="0" pageNumber="79">18L, M</figureCitation>
) with pair of asymmetric processes.
</paragraph>
</subSubSection>
</treatment>
</document>
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