261 lines
31 KiB
XML
261 lines
31 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3795974" ID-GBIF-Dataset="ad3e7788-7b8c-48c6-980c-76eac8bd2e0c" ID-GBIF-Taxon="163877326" ID-ISSN="1734-1833" ID-Zenodo-Dep="3795974" approvalRequired="6" approvalRequired_for_textStreams="6" checkinTime="1588190933256" checkinUser="felipe" docAuthor="Szadziewski, Ryszard, Dominiak, Patrycja & Withers, Phil" docDate="2020" docId="3D528C47FF9CBF2D37A5E569FCE7FBEA" docLanguage="en" docName="AnnalesZoologici.70.1.113-120.pdf.imf" docOrigin="Annales Zoologici 70 (1)" docStyle="DocumentStyle{}" docStyleName="annaleszoologici.2020.journal_article" docTitle="Monohelea mediterranea Szadziewski & Dominiak & Withers 2020, sp. nov." docType="treatment" docVersion="5" lastPageId="6" lastPageNumber="119" masterDocId="C16BF43FFF99BF2B375AE477FF91FF8F" masterDocTitle="Two New Species Of Biting Midges From France And Algeria (Diptera: Ceratopogonidae)" masterLastPageNumber="120" masterPageNumber="113" pageId="5" pageNumber="118" updateTime="1643440664791" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two New Species Of Biting Midges From France And Algeria (Diptera: Ceratopogonidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Szadziewski, Ryszard</mods:namePart>
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<mods:affiliation>Department of Invertebrate Zoology and Parasitology, Faculty of Biology, University of Gdańsk, Wita Stwosza 59, 80 - 308 Gdańsk, Poland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Dominiak, Patrycja</mods:namePart>
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<mods:affiliation>Norges arktiske universitetsmuseum og akademi for kunstfag, UiT Norges Arktiske Universitet, NO- 9037 TromsØ, Norway</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Withers, Phil</mods:namePart>
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<mods:affiliation>40 Montee du Cimetiere, Sainte Euphemie, 01600, France</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Annales Zoologici</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2020-03-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>70</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>113</mods:start>
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<mods:end>120</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3161/00034541ANZ2020.70.1.006</mods:identifier>
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<mods:identifier type="GBIF-Dataset">ad3e7788-7b8c-48c6-980c-76eac8bd2e0c</mods:identifier>
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<mods:identifier type="ISSN">1734-1833</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3776615</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3795974" ID-GBIF-Taxon="163877326" ID-Zenodo-Dep="3795974" LSID="urn:lsid:plazi:treatment:3D528C47FF9CBF2D37A5E569FCE7FBEA" httpUri="http://treatment.plazi.org/id/3D528C47FF9CBF2D37A5E569FCE7FBEA" lastPageId="6" lastPageNumber="119" pageId="5" pageNumber="118">
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<subSubSection box="[255,660,286,308]" pageId="5" pageNumber="118" type="nomenclature">
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<paragraph blockId="5.[255,660,286,340]" box="[255,660,286,308]" pageId="5" pageNumber="118">
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<heading box="[255,657,286,308]" level="2" pageId="5" pageNumber="118">
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<taxonomicName authority="Szadziewski & Dominiak & Withers, 2020" authorityName="Szadziewski & Dominiak & Withers" authorityYear="2020" box="[255,565,286,308]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="species" species="mediterranea" status="sp. nov.">
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<emphasis bold="true" box="[255,565,286,308]" italics="true" pageId="5" pageNumber="118">Monohelea mediterranea</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[573,660,287,308]" pageId="5" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[400,515,318,340]" pageId="5" pageNumber="118" type="description">
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<paragraph blockId="5.[255,660,286,340]" box="[400,515,318,340]" pageId="5" pageNumber="118">
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(
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<figureCitation box="[408,507,318,340]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStartId-0="4.[144,203,1856,1873]" captionStartId-1="5.[406,465,1881,1898]" captionStartId-2="6.[145,204,1876,1893]" captionTargetBox-0="[144,1443,196,1797]" captionTargetBox-1="[202,1385,706,1852]" captionTargetBox-2="[399,1225,1252,1827]" captionTargetId-0="figure@4.[144,1443,196,1798]" captionTargetId-1="figure@5.[202,1385,706,1853]" captionTargetId-2="figure@6.[334,1291,1228,1851]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Figure 4. Monohelea mediterranea sp. nov., male; a – head of paratype, dorsal aspect, b – palpi of paratype, c – total habitus of paratype, without head and genitalia, d – genitalia of holotype, ventral view." captionText-1="Figure 5. Monohelea mediterranea sp. nov., wing of male; a – holotype, b – paratype." captionText-2="Figure 6. Monohelea mediterranea sp. nov., male genitalia; a – ventral aspect of holotype, b – aedeagus of holotype, c – parameres of paratype." figureDoi-0="http://doi.org/10.5281/zenodo.3776623" figureDoi-1="http://doi.org/10.5281/zenodo.3776625" figureDoi-2="http://doi.org/10.5281/zenodo.3776627" httpUri-0="https://zenodo.org/record/3776623/files/figure.png" httpUri-1="https://zenodo.org/record/3776625/files/figure.png" httpUri-2="https://zenodo.org/record/3776627/files/figure.png" pageId="5" pageNumber="118">Figs 4–6</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="118" type="diagnosis">
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<paragraph blockId="5.[144,771,379,586]" pageId="5" pageNumber="118">
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<emphasis bold="true" box="[181,303,379,400]" italics="true" pageId="5" pageNumber="118">Diagnosis</emphasis>
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. The species is distinct in having second radial cell without dark spot, dark spot below first radial cell rectangular and uniformly dark, without pale spot or spots, legs with brown and pale rings and stripes, gonocoxite in male genitalia without ventral tubercle and apex of paramere T-shaped. Female unknown.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="119" pageId="5" pageNumber="118" type="description">
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<paragraph blockId="5.[144,771,379,586]" lastBlockId="5.[816,1444,180,586]" pageId="5" pageNumber="118">
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<emphasis bold="true" box="[181,325,564,585]" italics="true" pageId="5" pageNumber="118">Description</emphasis>
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. Male. Body yellowish brown (
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<figureCitation box="[677,758,564,586]" captionStart="Figure 4" captionStartId="4.[144,203,1856,1873]" captionTargetBox="[144,1443,196,1797]" captionTargetId="figure@4.[144,1443,196,1798]" captionTargetPageId="4" captionText="Figure 4. Monohelea mediterranea sp. nov., male; a – head of paratype, dorsal aspect, b – palpi of paratype, c – total habitus of paratype, without head and genitalia, d – genitalia of holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3776623" httpUri="https://zenodo.org/record/3776623/files/figure.png" pageId="5" pageNumber="118">Fig. 4c</figureCitation>
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). Eyes bare, broadly separated by vertex and frons (
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<figureCitation captionStart="Figure 4" captionStartId="4.[144,203,1856,1873]" captionTargetBox="[144,1443,196,1797]" captionTargetId="figure@4.[144,1443,196,1798]" captionTargetPageId="4" captionText="Figure 4. Monohelea mediterranea sp. nov., male; a – head of paratype, dorsal aspect, b – palpi of paratype, c – total habitus of paratype, without head and genitalia, d – genitalia of holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3776623" httpUri="https://zenodo.org/record/3776623/files/figure.png" pageId="5" pageNumber="118">Fig. 4a</figureCitation>
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). Flagellum with well developed plume (
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<figureCitation box="[1342,1430,212,234]" captionStart="Figure 4" captionStartId="4.[144,203,1856,1873]" captionTargetBox="[144,1443,196,1797]" captionTargetId="figure@4.[144,1443,196,1798]" captionTargetPageId="4" captionText="Figure 4. Monohelea mediterranea sp. nov., male; a – head of paratype, dorsal aspect, b – palpi of paratype, c – total habitus of paratype, without head and genitalia, d – genitalia of holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3776623" httpUri="https://zenodo.org/record/3776623/files/figure.png" pageId="5" pageNumber="118">Fig. 4a</figureCitation>
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), length
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<quantity box="[896,1055,244,265]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.299999999999999" metricValueMin="8.0" pageId="5" pageNumber="118" unit="mm" value="0.815" valueMax="0.83" valueMin="0.8">0.80–0.83 mm</quantity>
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, AR 0.93–0.95. Palpus 5-segment- ed (
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<figureCitation box="[862,947,276,298]" captionStart="Figure 4" captionStartId="4.[144,203,1856,1873]" captionTargetBox="[144,1443,196,1797]" captionTargetId="figure@4.[144,1443,196,1798]" captionTargetPageId="4" captionText="Figure 4. Monohelea mediterranea sp. nov., male; a – head of paratype, dorsal aspect, b – palpi of paratype, c – total habitus of paratype, without head and genitalia, d – genitalia of holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3776623" httpUri="https://zenodo.org/record/3776623/files/figure.png" pageId="5" pageNumber="118">Fig. 4b</figureCitation>
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); third palpal segment with small sensory pit, length
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<quantity box="[939,1051,308,330]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.3" metricValueMin="4.8" pageId="5" pageNumber="118" unit="mm" value="50.5" valueMax="53.0" valueMin="48.0">48–53 µm</quantity>
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, PR 2.11–2.30.
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</paragraph>
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<paragraph blockId="5.[816,1444,180,586]" pageId="5" pageNumber="118">
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Thorax brownish with indistinct ornamentation in slide mounted specimen, paratergite narrow, scutellum yellowish with 4 marginal bristles. Wing length
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<quantity box="[816,974,436,458]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" metricValueMax="1.3" metricValueMin="1.22" pageId="5" pageNumber="118" unit="mm" value="1.26" valueMax="1.3" valueMin="1.22">1.22–1.30 mm</quantity>
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, CR 0.70–0.71. Both first radial cells well developed, macrotrichia restricted to few at wing tip. Wing pattern as in
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<figureCitation box="[1052,1121,500,522]" captionStart="Figure 5" captionStartId="5.[406,465,1881,1898]" captionTargetBox="[202,1385,706,1852]" captionTargetId="figure@5.[202,1385,706,1853]" captionTargetPageId="5" captionText="Figure 5. Monohelea mediterranea sp. nov., wing of male; a – holotype, b – paratype." figureDoi="http://doi.org/10.5281/zenodo.3776625" httpUri="https://zenodo.org/record/3776625/files/figure.png" pageId="5" pageNumber="118">Fig. 5</figureCitation>
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; second radial cell without dark spot; dark spot below first radial cell rectangular, uniformly dark, without pale spot or spots.
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||
</paragraph>
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||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3776625" ID-Zenodo-Dep="3776625" box="[406,1182,1881,1898]" httpUri="https://zenodo.org/record/3776625/files/figure.png" pageId="5" pageNumber="118" startId="5.[406,465,1881,1898]" targetBox="[202,1385,706,1852]" targetPageId="5">
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<paragraph blockId="5.[406,1182,1881,1898]" box="[406,1182,1881,1898]" pageId="5" pageNumber="118">
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Figure 5.
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<taxonomicName authority="Szadziewski & Dominiak & Withers, 2020" authorityName="Szadziewski & Dominiak & Withers" authorityYear="2020" box="[492,737,1881,1898]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="species" species="mediterranea" status="sp. nov.">
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||
<emphasis box="[492,737,1881,1898]" italics="true" pageId="5" pageNumber="118">Monohelea mediterranea</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[742,813,1881,1897]" pageId="5" pageNumber="118" rank="species">sp. nov.</taxonomicNameLabel>
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, wing of male; a – holotype, b – paratype.
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</paragraph>
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</caption>
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<paragraph blockId="6.[144,771,180,1125]" pageId="6" pageNumber="119">Fore and hind legs somewhat thickened, mid legs slender. Legs brownish, femora without spines; coxae brown; trochanter of fore leg yellow, of mid and hind legs brown; femur of hind leg yellowish with distinctly brown proximal portion, narrow brown ring at middle and ventral subapical dark spot; fore and mid femora with similar colour patterns, however paler and indistinct; hind tibia with distinct brown ring at mid length and apex, fore and mid tibiae paler, with less developed brownish patterns as in hind tibia; tarsomere 1 of fore leg straight, with 1 subapical and 1 apical spine; tarsomere 1 of mid leg straight, with 2 subbasal and 2 apical spines; tarsomere 1 of hind leg strongly bent at base, with one row of palisade setae and strong subbasal spine; tibial spur of hind leg short, tibial comb composed of 9 spines; tarsomeres 4 cylindrical; claws small, equal; TR(1) 1.85–1.92, TR(2) 2.00–2.17, TR(3) 1.67–1.71.</paragraph>
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<paragraph blockId="6.[144,771,180,1125]" pageId="6" pageNumber="119">
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Genitalia brown (
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<figureCitation box="[385,470,734,756]" captionStart="Figure 4" captionStartId="4.[144,203,1856,1873]" captionTargetBox="[144,1443,196,1797]" captionTargetId="figure@4.[144,1443,196,1798]" captionTargetPageId="4" captionText="Figure 4. Monohelea mediterranea sp. nov., male; a – head of paratype, dorsal aspect, b – palpi of paratype, c – total habitus of paratype, without head and genitalia, d – genitalia of holotype, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3776623" httpUri="https://zenodo.org/record/3776623/files/figure.png" pageId="6" pageNumber="119">Fig. 4d</figureCitation>
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). Gonocoxite stout, mesal margin without lobe or tubercle (
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<figureCitation box="[536,621,765,787]" captionStart="Figure 6" captionStartId="6.[145,204,1876,1893]" captionTargetBox="[399,1225,1252,1827]" captionTargetId="figure@6.[334,1291,1228,1851]" captionTargetPageId="6" captionText="Figure 6. Monohelea mediterranea sp. nov., male genitalia; a – ventral aspect of holotype, b – aedeagus of holotype, c – parameres of paratype." figureDoi="http://doi.org/10.5281/zenodo.3776627" httpUri="https://zenodo.org/record/3776627/files/figure.png" pageId="6" pageNumber="119">Fig. 6a</figureCitation>
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). Gonostylus with slightly curved, pointed dark tip. Aedeagus with sclerotized basal loop, a pair of pointed, oblique lateral sclerites (
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<figureCitation box="[261,350,857,879]" captionStart="Figure 6" captionStartId="6.[145,204,1876,1893]" captionTargetBox="[399,1225,1252,1827]" captionTargetId="figure@6.[334,1291,1228,1851]" captionTargetPageId="6" captionText="Figure 6. Monohelea mediterranea sp. nov., male genitalia; a – ventral aspect of holotype, b – aedeagus of holotype, c – parameres of paratype." figureDoi="http://doi.org/10.5281/zenodo.3776627" httpUri="https://zenodo.org/record/3776627/files/figure.png" pageId="6" pageNumber="119">Fig. 6b</figureCitation>
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). Parameres separated, with winglike basal apodeme and T-shaped apex (
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<figureCitation box="[607,689,888,910]" captionStart="Figure 6" captionStartId="6.[145,204,1876,1893]" captionTargetBox="[399,1225,1252,1827]" captionTargetId="figure@6.[334,1291,1228,1851]" captionTargetPageId="6" captionText="Figure 6. Monohelea mediterranea sp. nov., male genitalia; a – ventral aspect of holotype, b – aedeagus of holotype, c – parameres of paratype." figureDoi="http://doi.org/10.5281/zenodo.3776627" httpUri="https://zenodo.org/record/3776627/files/figure.png" pageId="6" pageNumber="119">Fig. 6c</figureCitation>
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).
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</paragraph>
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<paragraph blockId="6.[144,771,180,1125]" box="[181,397,919,941]" pageId="6" pageNumber="119">Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="119" type="materials_examined">
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<paragraph blockId="6.[144,771,180,1125]" lastBlockId="6.[816,1445,180,1125]" pageId="6" pageNumber="119">
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<emphasis bold="true" box="[181,417,949,971]" italics="true" pageId="6" pageNumber="119">Material examined</emphasis>
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.
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<materialsCitation ID-GBIF-Occurrence="2609448602" collectingDate="1981-04-27" collectingDateMax="2009-07-15" collectingDateMin="1981-04-27" collectingMethod="collected by entomological net / Malaise trap" collectionCode="R" collectorName="Szadziewski & Pyrenees-Orientales & de la Massane & P. Withers. The" country="Algeria" county="Collection of Extant Invertebrates" location="University of Gdansk." municipality="Museum of Amber Inclusions" pageId="6" pageNumber="119" specimenCount="2" specimenCount-male="2" stateProvince="Biskra" typeStatus="holotype">
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<typeStatus box="[430,532,949,971]" pageId="6" pageNumber="119">Holotype</typeStatus>
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<specimenCount box="[539,597,949,971]" pageId="6" pageNumber="119" type="male">male</specimenCount>
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, North
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<collectingCountry box="[681,767,949,971]" name="Algeria" pageId="6" pageNumber="119">Algeria</collectingCountry>
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, Sahara,
|
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<quantity box="[240,310,980,1002]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="119" unit="km" value="30.0">
|
||
<locationDeviation box="[240,310,980,1002]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="119" unit="km" value="30.0">30 km</locationDeviation>
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</quantity>
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north of
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<collectingRegion box="[415,493,980,1002]" country="Algeria" name="Biskra" pageId="6" pageNumber="119">Biskra</collectingRegion>
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||
,
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<date box="[505,658,980,1002]" pageId="6" pageNumber="119" value="1981-04-27">
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||
<collectingDate box="[505,658,980,1002]" pageId="6" pageNumber="119" value="1981-04-27">27 April 1981</collectingDate>
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</date>
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||
,
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<collectingMethod pageId="6" pageNumber="119">collected by entomological net</collectingMethod>
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,
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<collectionCode box="[403,424,1011,1032]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6" pageNumber="119">R</collectionCode>
|
||
.
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||
<collectorName box="[437,582,1011,1033]" pageId="6" pageNumber="119">Szadziewski</collectorName>
|
||
;
|
||
<typeStatus box="[597,700,1011,1032]" pageId="6" pageNumber="119">paratype</typeStatus>
|
||
<specimenCount box="[709,767,1011,1033]" pageId="6" pageNumber="119" type="male">male</specimenCount>
|
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,
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<collectingCountry box="[144,228,1042,1063]" name="France" pageId="6" pageNumber="119">France</collectingCountry>
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,
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<collectorName box="[238,473,1042,1064]" pageId="6" pageNumber="119">Pyrenees-Orientales</collectorName>
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, RNN foret
|
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<collectorName box="[603,767,1042,1064]" pageId="6" pageNumber="119">de la Massane</collectorName>
|
||
,
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||
<collectingMethod box="[144,290,1072,1094]" pageId="6" pageNumber="119">Malaise trap</collectingMethod>
|
||
,
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||
<date box="[303,424,1072,1094]" pageId="6" pageNumber="119" value="2009-07-15">
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||
<collectingDate box="[303,424,1072,1094]" pageId="6" pageNumber="119" value="2009-07-15">15.07.2009</collectingDate>
|
||
</date>
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||
,
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||
<collectorName box="[436,615,1072,1094]" pageId="6" pageNumber="119">P. Withers. The</collectorName>
|
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<typeStatus box="[623,721,1072,1094]" pageId="6" pageNumber="119">holotype</typeStatus>
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and
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<typeStatus box="[144,247,1104,1125]" pageId="6" pageNumber="119">paratype</typeStatus>
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are deposited in the
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<collectingCounty box="[528,971,180,1125]" pageId="6" pageNumber="119">Collection of Extant Invertebrates</collectingCounty>
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the
|
||
<collectingMunicipality box="[1030,1379,180,202]" pageId="6" pageNumber="119">Museum of Amber Inclusions</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:3D528C47FF9CBF2D37A5E569FCE7FBEA:B0246B8AFF9FBF2D3229E4C3FB90FF66" country="Algeria" county="Collection of Extant Invertebrates" municipality="Museum of Amber Inclusions" name="University of Gdansk." pageId="6" pageNumber="119" stateProvince="Biskra">University of Gdańsk.</location>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="119" type="etymology">
|
||
<paragraph blockId="6.[816,1445,180,1125]" pageId="6" pageNumber="119">
|
||
<emphasis bold="true" box="[854,983,241,263]" italics="true" pageId="6" pageNumber="119">Etymology</emphasis>
|
||
. The specific name refers to the Mediterranean region where the species was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="6" pageNumber="119" type="discussion">
|
||
<paragraph blockId="6.[816,1445,180,1125]" pageId="6" pageNumber="119">
|
||
<emphasis bold="true" box="[854,988,303,324]" italics="true" pageId="6" pageNumber="119">Discussion</emphasis>
|
||
. The new species is very similar to
|
||
<taxonomicName authority="Wirth & Grogan, 1981" authorityName="Wirth & Grogan" authorityYear="1981" box="[816,1376,334,356]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="andersoni">
|
||
<emphasis box="[816,1093,334,355]" italics="true" pageId="6" pageNumber="119">Monohelea andersoni</emphasis>
|
||
<bibRefCitation author="Wirth, W. W. & W. L. Grogan" box="[1107,1376,334,356]" pageId="6" pageNumber="119" pagination="1 - 102" refId="ref2756" refString="Wirth, W. W. and W. L. Grogan. 1981. Natural history of Plummers Island, Maryland XXV. Biting midges (Diptera: Ceratopogonidae). 3. The species of the tribe Stilobezziini. Bulletin of the Biological Society of Washington, 5: 1 - 102." type="journal article" year="1981">Wirth & Grogan, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
from North America (
|
||
<bibRefCitation author="Wirth, W. W. & W. L. Grogan" box="[1011,1300,364,387]" pageId="6" pageNumber="119" pagination="1 - 102" refId="ref2756" refString="Wirth, W. W. and W. L. Grogan. 1981. Natural history of Plummers Island, Maryland XXV. Biting midges (Diptera: Ceratopogonidae). 3. The species of the tribe Stilobezziini. Bulletin of the Biological Society of Washington, 5: 1 - 102." type="journal article" year="1981">Wirth and Grogan 1981</bibRefCitation>
|
||
). The latter species however, is smaller (wing length
|
||
<quantity box="[1337,1432,395,417]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="119" unit="mm" value="0.9">0.9 mm</quantity>
|
||
), has darker legs, the aedeagus is longer and apices of parameres abruptly expanded in a cap-like lobe bent ventrally. The aedeagus and parameres of
|
||
<taxonomicName authority="Szadziewski & Dominiak & Withers, 2020" authorityName="Szadziewski & Dominiak & Withers" authorityYear="2020" box="[816,1028,518,540]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="mediterranea" status="sp. nov.">
|
||
<emphasis box="[816,1028,518,540]" italics="true" pageId="6" pageNumber="119">M. mediterranea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1037,1126,519,540]" pageId="6" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
|
||
are also somewhat similar to those of
|
||
<taxonomicName authority="Clastrier & Delecolle, 1990" authorityName="Clastrier & Delecolle" authorityYear="1990" box="[943,1383,549,571]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="pallida">
|
||
<emphasis box="[943,1072,549,571]" italics="true" pageId="6" pageNumber="119">M. pallida</emphasis>
|
||
<bibRefCitation author="Clastrier, J. & J. - C. Delecolle" box="[1080,1383,549,571]" pageId="6" pageNumber="119" pagination="129 - 157" refId="ref2195" refString="Clastrier, J. and J. - C. Delecolle. 1990. Description d'un nouveau genre et de nouvelles especes africaines des genres Allohelea Kieffer, Monohelea Kieffer, Downeshelea Wirth & Grogan, Boreohelea nov. gen. (Diptera: Ceratopogonidae). Annales de la Societe Entomologique de France (N. S.), 26: 129 - 157." type="journal article" year="1990">Clastrier & Delécolle, 1990</bibRefCitation>
|
||
</taxonomicName>
|
||
from tropical Africa (
|
||
<bibRefCitation author="Clastrier, J. & J. - C. Delecolle" box="[995,1320,580,602]" pageId="6" pageNumber="119" pagination="129 - 157" refId="ref2195" refString="Clastrier, J. and J. - C. Delecolle. 1990. Description d'un nouveau genre et de nouvelles especes africaines des genres Allohelea Kieffer, Monohelea Kieffer, Downeshelea Wirth & Grogan, Boreohelea nov. gen. (Diptera: Ceratopogonidae). Annales de la Societe Entomologique de France (N. S.), 26: 129 - 157." type="journal article" year="1990">Clastrier and Delécolle 1990</bibRefCitation>
|
||
). However, the latter species has pale legs and second radial cell with dark spot.
|
||
</paragraph>
|
||
<paragraph blockId="6.[816,1445,180,1125]" pageId="6" pageNumber="119">
|
||
<taxonomicName authorityName="Kieffer" authorityYear="1917" box="[854,987,672,693]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[854,987,672,693]" italics="true" pageId="6" pageNumber="119">Monohelea</emphasis>
|
||
</taxonomicName>
|
||
is a worldwide distributed genus, which includes 88 recent species (
|
||
<bibRefCitation author="Borkent, A." box="[1175,1346,703,725]" pageId="6" pageNumber="119" refId="ref2156" refString="Borkent, A. 2016. World species of biting midges (Diptera: Ceratopogonidae). Available from: http: // www. inhs. uiuc. edu / research / FLYTREE / CeratopogonidaeCatalog. pdf." type="url" year="2016">Borkent 2016</bibRefCitation>
|
||
). Three species are currently known from western Palaearctic. In addition to the new species described above they are
|
||
<taxonomicName authority="Remm, 1965" authorityName="Remm" authorityYear="1965" box="[816,1111,795,817]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="estonica">
|
||
<emphasis box="[816,960,796,817]" italics="true" pageId="6" pageNumber="119">M. estonica</emphasis>
|
||
<bibRefCitation author="Remm, H." box="[969,1111,795,817]" pageId="6" pageNumber="119" pagination="182 - 188" refId="ref2617" refString="Remm, H. 1965. New species of biting midges (Diptera, Heleidae) from the European part of the USSR. Entomologitscheskoe Obozrenie, 44: 182 - 188." type="journal article" year="1965">Remm, 1965</bibRefCitation>
|
||
</taxonomicName>
|
||
(North and Central Europe) and
|
||
<taxonomicName authority="Remm, 1980" authorityName="Remm" authorityYear="1980" box="[870,1184,826,848]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="hissarica">
|
||
<emphasis box="[870,1029,826,848]" italics="true" pageId="6" pageNumber="119">M. hissarica</emphasis>
|
||
<bibRefCitation author="Remm, H." box="[1040,1184,826,848]" pageId="6" pageNumber="119" pagination="85 - 128" refId="ref2650" refString="Remm, H. 1980. New species of the family Ceratopogonidae (Diptera) from the Middle Asia [in Russian]. Tartu Riikliku Ulikooli Toimetised, 516: 85 - 128." type="journal article" year="1980">Remm, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
(Tajikstan in Central Asia, only female). The latter two species have dark spot in the second radial cell which in
|
||
<taxonomicName authority="Szadziewski & Dominiak & Withers, 2020" authorityName="Szadziewski & Dominiak & Withers" authorityYear="2020" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="mediterranea" status="sp. nov.">
|
||
<emphasis italics="true" pageId="6" pageNumber="119">M. mediterranea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[901,992,919,940]" pageId="6" pageNumber="119" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is totally pale (
|
||
<figureCitation box="[1182,1252,918,941]" captionStart="Figure 5" captionStartId="5.[406,465,1881,1898]" captionTargetBox="[202,1385,706,1852]" captionTargetId="figure@5.[202,1385,706,1853]" captionTargetPageId="5" captionText="Figure 5. Monohelea mediterranea sp. nov., wing of male; a – holotype, b – paratype." figureDoi="http://doi.org/10.5281/zenodo.3776625" httpUri="https://zenodo.org/record/3776625/files/figure.png" pageId="6" pageNumber="119">Fig. 5</figureCitation>
|
||
). The European
|
||
<taxonomicName authority="Remm" authorityName="Remm" authorityYear="1965" box="[816,1160,949,971]" class="Insecta" family="Ceratopogonidae" genus="Monohelea" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="estonica">
|
||
<emphasis box="[816,1071,949,971]" italics="true" pageId="6" pageNumber="119">Monohelea estonica</emphasis>
|
||
Remm
|
||
</taxonomicName>
|
||
reported from
|
||
<collectingCountry box="[1359,1440,949,971]" name="Russia" pageId="6" pageNumber="119">Russia</collectingCountry>
|
||
,
|
||
<collectingCountry box="[816,906,980,1002]" name="Estonia" pageId="6" pageNumber="119">Estonia</collectingCountry>
|
||
,
|
||
<collectingCountry box="[924,1006,980,1002]" name="Poland" pageId="6" pageNumber="119">Poland</collectingCountry>
|
||
,
|
||
<collectingCountry box="[1024,1117,980,1002]" name="Czech Republic" pageId="6" pageNumber="119">Czechia</collectingCountry>
|
||
, N.
|
||
<collectingCountry box="[1172,1253,981,1002]" name="France" pageId="6" pageNumber="119">France</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Dominiak, P. & W. Michalczuk" pageId="6" pageNumber="119" pagination="8 - 13" refId="ref2336" refString="Dominiak, P. and W. Michalczuk. 2009. Dwa nowe dla fauny Polski gatunki kuczmanow (Diptera: Ceratopogonidae). Dipteron, Bulletin of the Dipterological Section of the Polish Entomological Society, 25: 8 - 13." type="journal article" year="2009">Dominiak and Michalczuk 2009</bibRefCitation>
|
||
) differs also in having gonocoxites with a tubercle on mesal margin and simple pointed apices of parameres (
|
||
<bibRefCitation author="Remm, H." box="[1071,1211,1072,1094]" pageId="6" pageNumber="119" pagination="182 - 188" refId="ref2617" refString="Remm, H. 1965. New species of biting midges (Diptera, Heleidae) from the European part of the USSR. Entomologitscheskoe Obozrenie, 44: 182 - 188." type="journal article" year="1965">Remm 1965</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Delecolle, J. - C. & J. - P. Rieb" pageId="6" pageNumber="119" pagination="17 - 45" refId="ref2262" refString="Delecolle, J. - C. and J. - P. Rieb. 1995. Redescription de Monohelea estonica Remm, 1965, de M. macfiei Wirth, 1953, et de M. floridensis Wirth & Williams, 1964. Description d'une forme et d'une espece nouvelle affines (Diptera, Ceratopogonidae). Nouvelle Revue d'Entomologie (N. S.), 12: 17 - 45." type="journal article" year="1995">Delécolle and Rieb 1995</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |