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<document id="629C53A2A6C7450645B510E7E9D9BB6D" ID-CLB-Dataset="9574" ID-DOI="10.5281/zenodo.4272127" ID-GBIF-Dataset="724cb358-6f18-4816-afc7-bb42cb9b6942" ID-ISSN="0374-1036" ID-Zenodo-Dep="4272127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1605298182318" checkinUser="felipe" docAuthor="Lackner, Tomáš" docDate="2010" docId="0385915EFF7209C06036FF3DCECFFCAD" docLanguage="en" docName="ActaEntMusNatPra.50.suppl.1-254.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl.) 50" docStyle="DocumentStyle:CE2601992B79F4B08764287CD82450D1.3:ActaEntMusNatPra.2010-2016.monograph" docStyleId="CE2601992B79F4B08764287CD82450D1" docStyleName="ActaEntMusNatPra.2010-2016.monograph" docStyleVersion="3" docTitle="Xenonychus tridens : GANGLBAUER 1899" docType="treatment" docVersion="18" lastPageNumber="226" masterDocId="FFBCE926FF9709296157FFE7CE2AFFFE" masterDocTitle="Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)" masterLastPageNumber="254" masterPageNumber="1" pageNumber="222" updateTime="1698846217483" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5E82C0426F256C9D4262B473291BA185">Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)</mods:title>
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<mods:title id="B5F9DDE4C6CFD85A6880697AC1682F5C">suppl.</mods:title>
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<mods:number id="013C6EDD16EDABCD9E1E2FF73BAE9A9F">2010-10-20</mods:number>
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<treatment id="0385915EFF7209C06036FF3DCECFFCAD" ID-DOI="http://doi.org/10.5281/zenodo.4341991" ID-GBIF-Taxon="169092645" ID-Zenodo-Dep="4341991" LSID="urn:lsid:plazi:treatment:0385915EFF7209C06036FF3DCECFFCAD" httpUri="http://treatment.plazi.org/id/0385915EFF7209C06036FF3DCECFFCAD" lastPageId="233" lastPageNumber="226" pageId="229" pageNumber="222">
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<taxonomicName id="4C2C5BCBFF7209CC6036FF3DCD7BFF0D" ID-CoL="7FVVL" authority="(Jacquelin-Duval, 1852)" authorityName=": GANGLBAUER" authorityYear="1899" baseAuthorityName="Jacquelin-Duval" baseAuthorityYear="1852" box="[353,849,218,243]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens">
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<emphasis id="B958FC5AFF7209CC6036FF3DCC16FF0D" bold="true" box="[353,572,218,243]" italics="true" pageId="229" pageNumber="222">Xenonychus tridens</emphasis>
(
<bibRefCitation id="EFBD5DB9FF7209CC631EFF3CCD60FF0D" author="JACQUELIN-DUVAL M." box="[585,842,219,243]" pageId="229" pageNumber="222" pagination="695 - 718" refId="ref115897" refString="JACQUELIN-DUVAL M. 1852: Descriptions de deux genres nouveaux et de plusieurs especes nouvelles de Coleo / pteres propers a la faune francaise. Annales de la Societe Entomologique de France 10: 695 - 718." type="journal article" year="1852">Jacquelin-Duval, 1852</bibRefCitation>
)
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<subSubSection id="C33673C3FF7209CC609DFF1ECCC2FEF2" box="[458,744,249,269]" pageId="229" pageNumber="222" type="description">
<paragraph id="8B932048FF7209CC609DFF1ECCC2FEF2" blockId="229.[353,849,218,269]" box="[458,744,249,269]" pageId="229" pageNumber="222">
(
<figureCitation id="13173CCDFF7209CC6087FF1ECC32FEF3" box="[464,536,249,269]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="229" pageNumber="222">Figs. 28</figureCitation>
,
<figureCitation id="13173CCDFF7209CC6376FF1DCC12FEF3" box="[545,568,250,269]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 47–73. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 – Alienocacculus neftensis (Olexa, 1984); 48 – Ammostyphrus cerberus Reichardt, 1924; 49 – Chalcionellus amoenus (Erichson, 1834); 50 – Chivaenius kryzhanovskii Olexa, 1980; 51 – Eopachylopus ripae (Lewis, 1885); 52 – Eremosaprinus vlasovi (Reichardt, 1941); 53 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 – Exaesiopus grossipes grossipes (Marseul, 55); 55 – Gnathoncus rotundatus (Kugelann, 1792); 56 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 – Microsaprinus therondianus (Dahlgren, 1973); 61 – Myrmetes paykulli Kanaar, 1979; 62 – Paravolvulus lateristriatus (Solskij, 1876); 63 – Philothis (Philothis) arcanus Reichardt, 1930; 64 – Philothis (Atavinus) atavus (Reichardt, 1931); 65 – Philothis (Farabius) hexeris Reichardt, 1930; 66 – Pholioxenus phoenix (Reichardt, 1930); 67 – Reichardtiolus duriculus (Reitter, 1904); 68 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 – Saprinus (Phaonius) pharao (Marseul, 1855); 71 – Styphrus corpulentus Motschulsky, 1845; 72 – Xenonychus tridens (Jacquelin-Duval, 1852); 73 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="229" pageNumber="222">72</figureCitation>
,
<figureCitation id="13173CCDFF7209CC6316FF1ECC49FEF2" box="[577,611,249,268]" captionStart="Figs" captionStartId="36.[151,190,1161,1180]" captionTargetBox="[151,1126,219,1130]" captionTargetId="figure-100@36.[151,1126,219,1132]" captionTargetPageId="36" captionText="Figs.99–107.Saprininae, mandibles,ventral view:99 – Philothis (Farabius) hexeris Reichardt, 1930; 100 – Pholioxenus phoenix (Reichardt,1930); 101 – Reichardtiolus duriculus (Reitter, 1904); 102 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 103 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 104 – Saprinus (Phaonius) pharao (Marseul, 1855); 105 – Styphrus corpulentus Motschulsky, 1845; 106 – Xenonychus tridens (Jacquelin-Duval, 1852); 107 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4272155" httpUri="https://zenodo.org/record/4272155/files/figure.png" pageId="229" pageNumber="222">106</figureCitation>
,
<figureCitation id="13173CCDFF7209CC633CFF1DCCA7FEF3" box="[619,653,250,269]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="229" pageNumber="222">139</figureCitation>
,
<figureCitation id="13173CCDFF7209CC63C1FF1ECCC8FEF2" box="[662,738,249,268]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="230.[151,189,1558,1577]" captionStartId-1="231.[113,152,1157,1176]" captionTargetBox-0="[228,1051,219,1536]" captionTargetBox-1="[194,1001,225,1129]" captionTargetId-0="figure-11@230.[228,1051,219,1537]" captionTargetId-1="figure-99@231.[151,1054,216,1136]" captionTargetPageId-0="230" captionTargetPageId-1="231" captionText-0="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." captionText-1="Figs. 721–729. Xenonychus tridens (Jacquelin-Duval, 1852), male terminalia: 721 – 8th sternite and tergite, ventral view; 722 – ditto, dorsal view; 723 – ditto, lateral view; 724 – 9th tergite and 10th tergite, dorsal view; 725 – ditto, lateral view; 726 – spiculum gastrale, ventral view; 727 – ditto, lateral view; 728 – aedeagus, dorsal view; 729 – ditto, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4272310" figureDoi-1="http://doi.org/10.5281/zenodo.4272312" httpUri-0="https://zenodo.org/record/4272310/files/figure.png" httpUri-1="https://zenodo.org/record/4272312/files/figure.png" pageId="229" pageNumber="222">711–729</figureCitation>
)
</paragraph>
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<treatmentCitation id="0A8D0659FF7209CC6126FEC4CFD9FEC8" author="JACQUELIN-DUVAL M." box="[113,499,291,310]" page="703" pageId="229" pageNumber="222" year="1852">
<taxonomicName id="4C2C5BCBFF7209CC6126FEC4CFD9FEC8" ID-CoL="9N3CR" authority="Jacquelin-Duval, 1852: 703" authorityName="Jacquelin-Duval" authorityPageNumber="703" authorityYear="1852" box="[113,499,291,310]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7209CC6126FEC4CED4FEC8" box="[113,254,291,310]" italics="true" pageId="229" pageNumber="222">Saprinus tridens</emphasis>
<bibRefCitation id="EFBD5DB9FF7209CC6054FEC4CFD9FEC8" author="JACQUELIN-DUVAL M." box="[259,499,291,310]" pageId="229" pageNumber="222" pagination="695 - 718" refId="ref115897" refString="JACQUELIN-DUVAL M. 1852: Descriptions de deux genres nouveaux et de plusieurs especes nouvelles de Coleo / pteres propers a la faune francaise. Annales de la Societe Entomologique de France 10: 695 - 718." type="journal article" year="1852">Jacquelin-Duval, 1852: 703</bibRefCitation>
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</paragraph>
<paragraph id="8B932048FF7209CC6126FED9CD34FEAF" blockId="229.[113,1089,291,577]" box="[113,798,318,337]" pageId="229" pageNumber="222">
<treatmentCitation id="0A8D0659FF7209CC6126FED9CFE8FEAF" author="MARSEUL S. A." box="[113,450,318,337]" page="501" pageId="229" pageNumber="222" year="1855">
<taxonomicName id="4C2C5BCBFF7209CC6126FED9CFB2FEAF" ID-CoL="9N3CR" authority=": MARSEUL (1855)" authorityName="MARSEUL" authorityYear="1855" box="[113,408,318,337]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7209CC6126FED9CED4FEAF" box="[113,254,318,337]" italics="true" pageId="229" pageNumber="222">Saprinus tridens</emphasis>
:
<bibRefCitation id="EFBD5DB9FF7209CC605EFED9CFB2FEAF" author="MARSEUL S. A." box="[265,408,318,337]" pageId="229" pageNumber="222" pagination="677 - 758" refId="ref117906" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Serie 3 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">MARSEUL (1855)</bibRefCitation>
</taxonomicName>
: 501
</treatmentCitation>
, t. XIX,
<figureCitation id="13173CCDFF7209CC635CFED9CC7FFEAF" box="[523,597,318,337]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="229" pageNumber="222">Fig. 118</figureCitation>
;
<treatmentCitation id="0A8D0659FF7209CC6309FED9CD31FEAF" author="SCHMIDT J." box="[606,795,318,337]" page="309" pageId="229" pageNumber="222" year="1885">
<bibRefCitation id="EFBD5DB9FF7209CC6309FED9CCC5FEAF" author="SCHMIDT J." box="[606,751,318,337]" pageId="229" pageNumber="222" pagination="279 - 330" refId="ref120947" refString="SCHMIDT J. 1885 a: Bestimmungs-Tabellen der europaischen Coleopteren. XIV. Histeridae. Berliner Entomologische Zeitung 29: 279 - 330." type="journal article" year="1885">SCHMIDT (1885a)</bibRefCitation>
: 309
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.
</paragraph>
<paragraph id="8B932048FF7209CC6126FEBECD05FE92" blockId="229.[113,1089,291,577]" box="[113,815,344,364]" pageId="229" pageNumber="222">
<treatmentCitation id="0A8D0659FF7209CC6126FEBECFCAFE92" author="MULSANT E. &amp; REY C." box="[113,480,344,364]" page="99" pageId="229" pageNumber="222" year="1853">
<taxonomicName id="4C2C5BCBFF7209CC6126FEBECFCAFE92" ID-CoL="9N38N" authority="Mulsant &amp; Rey, 1853: 99" authorityName="Mulsant &amp; Rey" authorityPageNumber="99" authorityYear="1853" box="[113,480,344,364]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="ciliaris">
<emphasis id="B958FC5AFF7209CC6126FEBECED5FE95" box="[113,255,344,364]" italics="true" pageId="229" pageNumber="222">Saprinus ciliaris</emphasis>
<bibRefCitation id="EFBD5DB9FF7209CC6053FEBFCFCAFE92" author="MULSANT E. &amp; REY C." box="[260,480,344,364]" pageId="229" pageNumber="222" pagination="97 - 100" refId="ref119161" refString="MULSANT E. &amp; REY C. 1853: Description des deux Coleopteres nouveaux de la tribu des Fracticornes. Opuscules Entomologiques 2: 97 - 100." type="journal article" year="1853">Mulsant &amp; Rey, 1853: 99</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="0A8D0659FF7209CC6329FEBECD00FE92" author="KRAATZ G." box="[638,810,344,364]" page="131" pageId="229" pageNumber="222" year="1858">
<bibRefCitation id="EFBD5DB9FF7209CC6329FEBECD2BFE95" author="KRAATZ G." box="[638,769,344,364]" pageId="229" pageNumber="222" pagination="123 - 148" refId="ref116665" refString="KRAATZ G. 1858: Beitrag zur Kaferfauna Griechenlands. Drittes Stuck: Staphylinidae (Schluss), Trichopterygia, Histeridae, Phalacridae, Nitidulariae, Trogositarii, Colydii, Cucujidae, Cryptophagidae, Thorictidae, Mycetophagidae, Dermestini, Byrhii. Berliner Entomologische Zeitschrift 12: 123 - 148." type="journal article" year="1858">KRAATZ (1858)</bibRefCitation>
: 131
</treatmentCitation>
.
</paragraph>
<paragraph id="8B932048FF7209CC6126FE94CD3AFE78" blockId="229.[113,1089,291,577]" box="[113,784,371,391]" pageId="229" pageNumber="222">
<treatmentCitation id="0A8D0659FF7209CC6126FE94CF9CFE78" author="MARSEUL S. A." box="[113,438,371,390]" page="677" pageId="229" pageNumber="222" year="1855">
<taxonomicName id="4C2C5BCBFF7209CC6126FE94CF9CFE78" ID-CoL="9N3BZ" authority="Marseul, 1855: 677" authorityName="Marseul" authorityPageNumber="677" authorityYear="1855" box="[113,438,371,390]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="serripes">
<emphasis id="B958FC5AFF7209CC6126FE94CF22FE78" box="[113,264,371,390]" italics="true" pageId="229" pageNumber="222">Saprinus serripes</emphasis>
<bibRefCitation id="EFBD5DB9FF7209CC605AFE94CF9CFE78" author="MARSEUL S. A." box="[269,438,371,390]" pageId="229" pageNumber="222" pagination="677 - 758" refId="ref117906" refString="MARSEUL S. A. 1855: Essai monographique sur la familie des Histerides. (Suite). Annales de Societe Entomologique de France, Serie 3 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1855">Marseul, 1855: 677</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="0A8D0659FF7209CC6304FE93CD27FE78" author="MARSEUL S. A." box="[595,781,371,391]" page="482" pageId="229" pageNumber="222" year="1862">
<bibRefCitation id="EFBD5DB9FF7209CC6304FE93CCCBFE78" author="MARSEUL S. A." box="[595,737,371,391]" pageId="229" pageNumber="222" pagination="677 - 758" refId="ref118029" refString="MARSEUL S. A. 1862: Supplement a la monographie des Histerides. (Suite). Annales de la Societe Entomologique de France, Serie 4 3: 83 - 165, 327 - 506, 677 - 758." type="journal article" year="1862">MARSEUL (1862)</bibRefCitation>
: 482
</treatmentCitation>
.
</paragraph>
<paragraph id="8B932048FF7209CC6126FE69CD0CFE5F" blockId="229.[113,1089,291,577]" box="[113,806,398,417]" pageId="229" pageNumber="222">
<treatmentCitation id="0A8D0659FF7209CC6126FE69CFF8FE5F" author="WOLLASTON T. V." box="[113,466,398,417]" page="181" pageId="229" pageNumber="222" year="1864">
<taxonomicName id="4C2C5BCBFF7209CC6126FE69CFF8FE5F" ID-CoL="9N7LD" authority="Wollaston, 1864: 181" authorityName="Wollaston" authorityPageNumber="181" authorityYear="1864" box="[113,466,398,417]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="fossor">
<emphasis id="B958FC5AFF7209CC6126FE69CF38FE5F" box="[113,274,398,417]" italics="true" pageId="229" pageNumber="222">Xenonychus fossor</emphasis>
<bibRefCitation id="EFBD5DB9FF7209CC6040FE69CFF8FE5F" author="WOLLASTON T. V." box="[279,466,398,417]" pageId="229" pageNumber="222" refId="ref122598" refString="WOLLASTON T. V. 1864: Catalogue of the coleopterous insects of the Canaries in the collection of the British Museum. Taylor and Francis, London, xiii + 648 pp." type="book" year="1864">Wollaston, 1864: 181</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. Synonymized by
<treatmentCitation id="0A8D0659FF7209CC6327FE69CD09FE5F" author="SCHMIDT J." box="[624,803,398,417]" page="354" pageId="229" pageNumber="222" year="1887">
<bibRefCitation id="EFBD5DB9FF7209CC6327FE69CCD2FE5F" author="SCHMIDT J." box="[624,760,398,417]" pageId="229" pageNumber="222" pagination="353 - 356" refId="ref120992" refString="SCHMIDT J. 1887: Verzeichniss der von Herrn Premierleutenant M. Quedenfeldt in Marokko 1885 - 86 gesammelten Histeriden. Entomologische Nachrichten 13: 353 - 356." type="journal article" year="1887">SCHMIDT (1887)</bibRefCitation>
: 354
</treatmentCitation>
.
</paragraph>
<paragraph id="8B932048FF7209CC6126FE4ECC6FFE42" blockId="229.[113,1089,291,577]" box="[113,581,424,444]" pageId="229" pageNumber="222">
<treatmentCitation id="0A8D0659FF7209CC6126FE4ECC68FE42" author="SCHMIDT J." box="[113,578,424,444]" page="309" pageId="229" pageNumber="222" year="1885">
<taxonomicName id="4C2C5BCBFF7209CC6126FE4ECC3CFE45" ID-CoL="9N3CR" authority=": SCHMIDT (1885 a)" authorityName=": SCHMIDT" authorityYear="1885" box="[113,534,424,444]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens" subGenus="Hypocaccus">
<emphasis id="B958FC5AFF7209CC6126FE4ECE97FE42" box="[113,189,425,444]" italics="true" pageId="229" pageNumber="222">Saprinus</emphasis>
(
<emphasis id="B958FC5AFF7209CC619DFE4ECF18FE45" box="[202,306,425,443]" italics="true" pageId="229" pageNumber="222">Hypocaccus</emphasis>
)
<emphasis id="B958FC5AFF7209CC6069FE4FCF53FE45" box="[318,377,424,443]" italics="true" pageId="229" pageNumber="222">tridens</emphasis>
:
<bibRefCitation id="EFBD5DB9FF7209CC60D2FE4ECC3CFE45" author="SCHMIDT J." box="[389,534,424,444]" pageId="229" pageNumber="222" pagination="279 - 330" refId="ref120947" refString="SCHMIDT J. 1885 a: Bestimmungs-Tabellen der europaischen Coleopteren. XIV. Histeridae. Berliner Entomologische Zeitung 29: 279 - 330." type="journal article" year="1885">SCHMIDT (1885a)</bibRefCitation>
</taxonomicName>
: 309
</treatmentCitation>
.
</paragraph>
<paragraph id="8B932048FF7209CC6126FE24CC8FFE28" blockId="229.[113,1089,291,577]" box="[113,677,451,471]" pageId="229" pageNumber="222">
<treatmentCitation id="0A8D0659FF7209CC6126FE24CFE5FE28" author="BICKHARDT H." box="[113,463,451,471]" page="107" pageId="229" pageNumber="222" year="1910">
<taxonomicName id="4C2C5BCBFF7209CC6126FE24CF8EFE28" authority=": BICKHARDT (1910)" authorityName="BICKHARDT" authorityYear="1910" box="[113,420,451,471]" class="Insecta" family="Histeridae" genus="Styphrus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7209CC6126FE24CED6FE28" box="[113,252,451,470]" italics="true" pageId="229" pageNumber="222">Styphrus tridens</emphasis>
:
<bibRefCitation id="EFBD5DB9FF7209CC605FFE23CF8EFE28" author="BICKHARDT H." box="[264,420,451,471]" pageId="229" pageNumber="222" refId="ref112602" refString="BICKHARDT H. 1910: Histeridae. In: JUNK W. &amp; SCHENKLING S. (eds.): Coleopterorum Catalogus, volume 24. W. Junk, Berlin, 137 pp." type="book" year="1910">BICKHARDT (1910)</bibRefCitation>
</taxonomicName>
: 107
</treatmentCitation>
; G.
<treatmentCitation id="0A8D0659FF7209CC60A5FE23CC88FE28" author="MULLER G." box="[498,674,451,471]" page="102" pageId="229" pageNumber="222" year="1931">
<bibRefCitation id="EFBD5DB9FF7209CC60A5FE23CC5DFE28" author="MULLER G." box="[498,631,451,471]" pageId="229" pageNumber="222" pagination="93 - 104" refId="ref119189" refString="MULLER G. 1931: Note sugli Histeridae della Tripolitania e Tunisia raccolti durante i viaggi organizzati da S. A. Serenissima il Principe Alessandro della torre tasso. Memorie de la Societa Entomologica Italiana 10: 93 - 104." type="journal article" year="1931">MÜLLER (1931)</bibRefCitation>
: 102
</treatmentCitation>
.
</paragraph>
<paragraph id="8B932048FF7209CC6126FE39CDFEFDD8" blockId="229.[113,1089,291,577]" pageId="229" pageNumber="222">
<treatmentCitation id="0A8D0659FF7209CC6126FE39CC28FE0F" author="GANGLBAUER L." box="[113,514,478,497]" page="394" pageId="229" pageNumber="222" year="1899">
<taxonomicName id="4C2C5BCBFF7209CC6126FE39CFFCFE0F" ID-CoL="7FVVL" authority=": GANGLBAUER (1899)" authorityName=": GANGLBAUER" authorityYear="1899" baseAuthorityName="Jacquelin-Duval" baseAuthorityYear="1852" box="[113,470,478,497]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7209CC6126FE39CF30FE0F" box="[113,282,478,497]" italics="true" pageId="229" pageNumber="222">Xenonychus tridens</emphasis>
:
<bibRefCitation id="EFBD5DB9FF7209CC6070FE39CFFCFE0F" author="GANGLBAUER L." box="[295,470,478,497]" pageId="229" pageNumber="222" refId="ref114798" refString="GANGLBAUER L. 1899: Die Kafer von Mitteleuropa. Die Kafer der osterreichisch-ungarischen Monarchie, Deutschland, der Schweiz, sowie des franzosischen und italienischen Alpengebiets. III, 1. Familienreihe Staphylinoidea. 2 Theil: Scydmaenidae, Silphidae, Clambidae, Leptinidae, Platypsyllidae, Corylophidae, Sphaeritidae, Trichopterigydae, Hydroscaphidae, Scaphidiidae, Histeridae. Carl Gerolds Sohn, Wien, 408 pp." type="book" year="1899">GANGLBAUER (1899)</bibRefCitation>
</taxonomicName>
: 394
</treatmentCitation>
;
<treatmentCitation id="0A8D0659FF7209CC635CFE39CC9AFE0F" author="REITTER E." box="[523,688,478,497]" page="13" pageId="229" pageNumber="222" year="1910">
<bibRefCitation id="EFBD5DB9FF7209CC635CFE39CCA5FE0F" author="REITTER E." box="[523,655,478,497]" pageId="229" pageNumber="222" pagination="13" refId="ref120599" refString="REITTER E. 1910: Uebersicht der Arten der Coleopterengattung Xenonychus Woll. Entomologische Blatter 6: 13." type="journal article" year="1910">REITTER (1910)</bibRefCitation>
: 13
</treatmentCitation>
;
<treatmentCitation id="0A8D0659FF7209CC63ECFE39CDAEFE0F" author="REICHARDT A." box="[699,900,478,497]" page="334" pageId="229" pageNumber="222" year="1941">
<bibRefCitation id="EFBD5DB9FF7209CC63ECFE39CD72FE0F" author="REICHARDT A." box="[699,856,478,497]" pageId="229" pageNumber="222" refId="ref120382" refString="REICHARDT A. 1941: Semeystva Sphaeritidae i Histeridae (Vol. 1). [Families Sphaeritidae and Histeridae]. Fauna SSSR, Nasekomye Zhestokrylye, V, 3. Nauka, Moskva-Leningrad, xiii + 419 pp." type="book" year="1941">REICHARDT (1941)</bibRefCitation>
: 334
</treatmentCitation>
; THÉROND (1963): 69;
<bibRefCitation id="EFBD5DB9FF7209CC61C7FE1ECFEFFDF5" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[144,453,504,524]" pageId="229" pageNumber="222" refId="ref117064" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae)]. Fauna SSSR, Zhestokrylye, Vyp. 4. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ &amp; REICHARDT (1976)</bibRefCitation>
: 112, 242,
<figureCitation id="13173CCDFF7209CC6375FE1FCC41FDF2" box="[546,619,504,524]" captionStart="Figs" captionStartId="156.[151,190,1489,1508]" captionTargetBox="[250,1029,219,1466]" captionTargetId="figure-11@156.[250,1030,219,1467]" captionTargetPageId="156" captionText="Figs. 471–481. Microsaprinus therondianus (Dahlgren, 1973), SEM micrographs: 471 – habitus, dorsal view; 472 – ditto, ventral view; 473 – antennal club, ventral view; 474 – ditto, dorsal view; 475 – head, dorsal view; 476 – protibia, dorsal view; 477 – metatibia, ventral view; 478 – mentum, stipites and cardines, ventral view; 479 – prosternum; 480 – lateral disc of metaventrite, metepisternum and fused metepimeron; 481 – protibia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272252" httpUri="https://zenodo.org/record/4272252/files/figure.png" pageId="229" pageNumber="222">Fig. 471</figureCitation>
;
<bibRefCitation id="EFBD5DB9FF7209CC6323FE1ECCD8FDF2" author="VIENNA P." box="[628,754,505,524]" pageId="229" pageNumber="222" refId="ref122194" refString="VIENNA P. 1980: Fauna d'Italia. Vol. XVI. Coleoptera Histeridae. Bologna, Calderini Press, ix + 386 pp." type="book" year="1980">
<collectingRegion id="49E8EEAAFF7209CC6323FE1ECC9BFDF4" box="[628,689,505,524]" country="Austria" name="Wien" pageId="229" pageNumber="222">VIENNA</collectingRegion>
(1980)
</bibRefCitation>
: 115, 198,
<figureCitation id="13173CCDFF7209CC6218FE1FCDA4FDF2" box="[847,910,504,524]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 47–73. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 – Alienocacculus neftensis (Olexa, 1984); 48 – Ammostyphrus cerberus Reichardt, 1924; 49 – Chalcionellus amoenus (Erichson, 1834); 50 – Chivaenius kryzhanovskii Olexa, 1980; 51 – Eopachylopus ripae (Lewis, 1885); 52 – Eremosaprinus vlasovi (Reichardt, 1941); 53 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 – Exaesiopus grossipes grossipes (Marseul, 55); 55 – Gnathoncus rotundatus (Kugelann, 1792); 56 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 – Microsaprinus therondianus (Dahlgren, 1973); 61 – Myrmetes paykulli Kanaar, 1979; 62 – Paravolvulus lateristriatus (Solskij, 1876); 63 – Philothis (Philothis) arcanus Reichardt, 1930; 64 – Philothis (Atavinus) atavus (Reichardt, 1931); 65 – Philothis (Farabius) hexeris Reichardt, 1930; 66 – Pholioxenus phoenix (Reichardt, 1930); 67 – Reichardtiolus duriculus (Reitter, 1904); 68 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 – Saprinus (Phaonius) pharao (Marseul, 1855); 71 – Styphrus corpulentus Motschulsky, 1845; 72 – Xenonychus tridens (Jacquelin-Duval, 1852); 73 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="229" pageNumber="222">Fig. 70</figureCitation>
;
<bibRefCitation id="EFBD5DB9FF7209CC62C1FE1ECA38FDF2" author="MAZUR S." box="[918,1042,505,524]" pageId="229" pageNumber="222" pagination="1 - 376" refId="ref118519" refString="MAZUR S. 1984: A world catalogue of Histeridae. Polskie Pismo Entomologiczne 54 (3 - 4): 1 - 376." type="journal article" year="1984">MAZUR (1984)</bibRefCitation>
: 108;
<bibRefCitation id="EFBD5DB9FF7209CC61C7FDF3CF27FDD8" author="MAZUR S." box="[144,269,531,551]" pageId="229" pageNumber="222" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR (1997)</bibRefCitation>
: 267;
<bibRefCitation id="EFBD5DB9FF7209CC6017FDF3CFE4FDD8" author="YELAMOS T." box="[320,462,531,551]" pageId="229" pageNumber="222" refId="ref122718" refString="YELAMOS T. 2002: Coleoptera, Histeridae. In: RAMOS M. A., TERCEDOR J. A, BELLES-ROS X., GOSAL- BEZ-NOGUERA J., SIERRA A. G., MAYOL E. M., PIERA F. M., MARINO J. S. &amp; GONZALEZ J. T. (eds.): Fauna Iberica. Vol. 17. Museo Nacional de Ciencias Naturales, CSCI, Madrid, 411 pp." type="book" year="2002">YÉLAMOS (2002)</bibRefCitation>
: 245, 340,
<figureCitation id="13173CCDFF7209CC637BFDF4CCABFDD8" box="[556,641,531,550]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="229" pageNumber="222">Figs. 12A</figureCitation>
,
<figureCitation id="13173CCDFF7209CC63DDFDF4CC96FDD8" box="[650,700,531,550]" captionStart="Figs" captionStartId="54.[152,191,1567,1586]" captionTargetBox="[184,1092,915,1541]" captionTargetId="figure-311@54.[184,1093,915,1542]" captionTargetPageId="54" captionText="Figs. 160–161. Myrmetes paykulli Kanaar, 1979, mesotibia: 160 – dorsal view; 161 – ventral view." figureDoi="http://doi.org/10.5281/zenodo.4272179" httpUri="https://zenodo.org/record/4272179/files/figure.png" pageId="229" pageNumber="222">161H</figureCitation>
,
<figureCitation id="13173CCDFF7209CC6393FDF4CD35FDD8" box="[708,799,531,550]" captionStart="Figs" captionStartId="58.[151,190,1544,1563]" captionTargetBox="[178,1096,461,1510]" captionTargetId="figure-11@58.[178,1096,459,1512]" captionTargetPageId="58" captionText="Figs. 168–173. Saprininae, male terminalia, schematic: 168 – 8th sternite, ventral view; 169 – 8th sternite and tergite, dorsal view; 170 – ditto, lateral view; 171 – ninth tergites and tenth tergite, dorsal view; 172 – spiculum gastrale (10th sternite), ventral view; 173 – aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272183" httpUri="https://zenodo.org/record/4272183/files/figure.png" pageId="229" pageNumber="222">170B, 171</figureCitation>
;
<bibRefCitation id="EFBD5DB9FF7209CC627EFDF3CD8FFDD8" author="MAZUR S." box="[809,933,531,551]" pageId="229" pageNumber="222" pagination="68 - 102" refId="ref118682" refString="MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. &amp; SMETANA A. (eds.): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">MAZUR (2004)</bibRefCitation>
: 101.
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF7209CC61C7FDC9CA0DFDBF" box="[144,1063,558,577]" pageId="229" pageNumber="222" type="description">
<paragraph id="8B932048FF7209CC61C7FDC9CA0DFDBF" blockId="229.[113,1089,291,577]" box="[144,1063,558,577]" pageId="229" pageNumber="222">
<emphasis id="B958FC5AFF7209CC61C7FDC9CEEAFDBF" bold="true" box="[144,192,558,577]" pageId="229" pageNumber="222">Note.</emphasis>
Sensory structures of antennal club and spermatheca were studied by
<bibRefCitation id="EFBD5DB9FF7209CC6240FDC9CA24FDBF" author="DE MARZO L. &amp; VIENNA P." box="[791,1038,558,577]" pageId="229" pageNumber="222" pagination="79 - 89" refId="ref114178" refString="DE MARZO L. &amp; VIENNA P. 1982 a: Osservazioni morfologiche e ultrastrutturali su particolari organi di senso delle clave antennali in Isteridi della subf. Saprininae e considerazioni sistematiche. Entomologica 17: 79 - 89." type="journal article" year="1982">
DE MARZO &amp;
<collectingRegion id="49E8EEAAFF7209CC62DCFDC9CDE3FDBE" box="[907,969,558,577]" country="Austria" name="Wien" pageId="229" pageNumber="222">VIENNA</collectingRegion>
(1982a
</bibRefCitation>
,b).
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF7209CC6126FDBFCFC4FC6E" pageId="229" pageNumber="222" type="materials_examined">
<paragraph id="8B932048FF7209CC6126FDBFCFEBFD95" blockId="229.[113,1090,599,913]" box="[113,449,599,619]" pageId="229" pageNumber="222">
<emphasis id="B958FC5AFF7209CC6126FDBFCEC7FD95" bold="true" box="[113,237,599,619]" pageId="229" pageNumber="222">
<typeStatus id="54979EEAFF7209CC6126FDBFCEB5FD95" box="[113,159,600,619]" pageId="229" pageNumber="222">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33B60D8FF7209CC61A5FDBFCF04FD95" box="[242,302,600,619]" name="France" pageId="229" pageNumber="222">France</collectingCountry>
, Le Grau de Roi.
</paragraph>
<paragraph id="8B932048FF7209CC6126FD95CA00FD45" blockId="229.[113,1090,599,913]" pageId="229" pageNumber="222">
<emphasis id="B958FC5AFF7209CC6126FD95CED2FD7B" bold="true" box="[113,248,625,645]" pageId="229" pageNumber="222">Type material.</emphasis>
<emphasis id="B958FC5AFF7209CC61AAFD95CF86FD7B" box="[253,428,626,645]" italics="true" pageId="229" pageNumber="222">
<taxonomicName id="4C2C5BCBFF7209CC61AAFD95CF82FD7B" authorityName=": GANGLBAUER" authorityYear="1899" baseAuthorityName="Jacquelin-Duval" baseAuthorityYear="1852" box="[253,424,626,645]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens">Xenonychus tridens</taxonomicName>
.
</emphasis>
<typeStatus id="54979EEAFF7209CC60E7FD94CFD2FD7A" box="[432,504,627,646]" pageId="229" pageNumber="222" type="neotype">NEOTYPE</typeStatus>
(here designated): ♁, ‘
<collectingCountry id="F33B60D8FF7209CC63ECFD95CD39FD7B" box="[699,787,626,645]" name="France" pageId="229" pageNumber="222">FRANCE</collectingCountry>
/ Le Grau de Roi / (13)
<date id="FF920688FF7209CC62B3FD95CA6BFD7B" box="[996,1089,626,645]" pageId="229" pageNumber="222" value="1962-08-08">8.viii.1962</date>
[written] // tamissage au / pied des / Graminées [written] // Y. Gomy [printed] // Collection Y. Gomy [printed] // NEO- TYPUS /
<taxonomicName id="4C2C5BCBFF7209CC6192FD4FCF06FD45" authorityName=": MARSEUL" authorityYear="1864" box="[197,300,680,699]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF7209CC6192FD4FCF06FD45" box="[197,300,680,699]" italics="true" pageId="229" pageNumber="222">Xenonychus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C2C5BCBFF7209CC606AFD4FCF52FD45" authorityName=": GANGLBAUER" authorityYear="1899" baseAuthorityName="Jacquelin-Duval" baseAuthorityYear="1852" box="[317,376,680,699]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7209CC606AFD4FCF52FD45" box="[317,376,680,699]" italics="true" pageId="229" pageNumber="222">tridens</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EFBD5DB9FF7209CC60D8FD4FCC79FD45" author="JACQUELIN-DUVAL M." box="[399,595,680,699]" pageId="229" pageNumber="222" pagination="695 - 718" refId="ref115897" refString="JACQUELIN-DUVAL M. 1852: Descriptions de deux genres nouveaux et de plusieurs especes nouvelles de Coleo / pteres propers a la faune francaise. Annales de la Societe Entomologique de France 10: 695 - 718." type="journal article" year="1852">Jacquelin-Duval, 1852</bibRefCitation>
) / Des. T. Lackner 2009 [red label, written]’ (
<collectionCode id="ED3DB88DFF7209CC628DFD4FCA0AFD45" box="[986,1056,680,699]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="229" pageNumber="222">MNHN</collectionCode>
).
</paragraph>
<paragraph id="8B932048FF7209CC6126FD26CC31FCBE" blockId="229.[113,1090,599,913]" pageId="229" pageNumber="222">
<emphasis id="B958FC5AFF7209CC6126FD26CEFBFD2B" bold="true" box="[113,209,705,725]" pageId="229" pageNumber="222">Comment.</emphasis>
The type specimen of this species was not found in the collection of Jacquelin-Duval housed in
<collectingRegion id="49E8EEAAFF7209CC62ADFD24CA14FD28" box="[1018,1086,707,726]" country="Ireland" name="Monaghan" pageId="229" pageNumber="222">MNHN</collectingRegion>
. According to Mrs. Azadeh Taghavian, the type specimen of this species is probably lost, since there is an empty mounting card with the label ‘
<taxonomicName id="4C2C5BCBFF7209CC603BFD1FCFDDFCF5" authorityName=": SCHMIDT" authorityYear="1885" box="[364,503,760,779]" class="Insecta" family="Histeridae" genus="Saprinus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7209CC603BFD1FCFDDFCF5" box="[364,503,760,779]" italics="true" pageId="229" pageNumber="222">Saprinus tridens</emphasis>
</taxonomicName>
’. Although the author has inspected the fragments of various histerids on the bottom of the box, none of them contained pieces of
<taxonomicName id="4C2C5BCBFF7209CC6338FCF5CD3DFCDB" authorityName=": GANGLBAUER" authorityYear="1899" baseAuthorityName="Jacquelin-Duval" baseAuthorityYear="1852" box="[623,791,786,805]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7209CC6338FCF5CD3DFCDB" box="[623,791,786,805]" italics="true" pageId="229" pageNumber="222">Xenonychus tridens</emphasis>
</taxonomicName>
. The
<typeStatus id="54979EEAFF7209CC621EFCF4CDA7FCD8" box="[841,909,787,806]" pageId="229" pageNumber="222" type="neotype">neotype</typeStatus>
is designated here to fix the identity of the type species of
<taxonomicName id="4C2C5BCBFF7209CC60F8FCCACC3CFCBE" authorityName=": MARSEUL" authorityYear="1864" box="[431,534,813,832]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="229" pageNumber="222" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF7209CC60F8FCCACC3CFCBE" box="[431,534,813,832]" italics="true" pageId="229" pageNumber="222">Xenonychus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B932048FF7209CC6126FCA0CFC4FC6E" blockId="229.[113,1090,599,913]" pageId="229" pageNumber="222">
<emphasis id="B958FC5AFF7209CC6126FCA0CFA8FCA5" bold="true" box="[113,386,839,859]" pageId="229" pageNumber="222">Additional material examined.</emphasis>
<materialsCitation id="3B442A15FF7209CC60D1FCA0CD05FCA5" ID-GBIF-Occurrence="2981920302" box="[390,815,837,860]" collectingDate="1987-05-01" collectorName="A. Olexa" country="Algeria" location="Ghardaia" pageId="229" pageNumber="222" specimenCount="1" stateProvince="Ghardaia">
<emphasis id="B958FC5AFF7209CC60D1FCA0CFDAFCA5" bold="true" box="[390,496,839,859]" pageId="229" pageNumber="222">
<collectingCountry id="F33B60D8FF7209CC60D1FCA0CFC6FCA5" box="[390,492,839,859]" name="Algeria" pageId="229" pageNumber="222">ALGERIA</collectingCountry>
:
</emphasis>
<collectingRegion id="49E8EEAAFF7209CC60A3FCAFCC6EFCA5" box="[500,580,840,859]" country="Algeria" name="Ghardaia" pageId="229" pageNumber="222">Ghardaia</collectingRegion>
,
<date id="FF920688FF7209CC631DFCAFCCBCFCA5" box="[586,662,840,859]" pageId="229" pageNumber="222" value="1987-05-01">
<collectingDate id="EFD6FF60FF7209CC631DFCAFCCBCFCA5" box="[586,662,840,859]" pageId="229" pageNumber="222" value="1987-05-01">1.v.1987</collectingDate>
</date>
, 1♁,
<collectorName id="26D9459EFF7209CC6392FCAFCD3AFCA5" box="[709,784,840,859]" pageId="229" pageNumber="222">A.Olexa</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="3B442A15FF7209CC626EFCAFCF23FC8B" ID-GBIF-Occurrence="2981920323" collectingDate="1980-10-20" collectorName="A. Olexa" country="Algeria" location="Beni Abbes" municipality="Sahara" pageId="229" pageNumber="222" specimenCount="1" specimenCount-female="1">
<collectingMunicipality id="6BF7BA32FF7209CC626EFCAFCD5FFCA5" box="[825,885,840,859]" pageId="229" pageNumber="222">Sahara</collectingMunicipality>
,
<location id="8EF37693FF7209CC622BFCAFCDF5FCA5" LSID="urn:lsid:plazi:treatment:0385915EFF7209C06036FF3DCECFFCAD:8EF37693FF7209CC622BFCAFCDF5FCA5" box="[892,991,840,859]" country="Algeria" municipality="Sahara" name="Beni Abbes" pageId="229" pageNumber="222">Béni Abbès</location>
,
<date id="FF920688FF7209CC62B0FCAFCA17FCA5" box="[999,1085,840,859]" pageId="229" pageNumber="222" value="1980-10-20">
<collectingDate id="EFD6FF60FF7209CC62B0FCAFCA17FCA5" box="[999,1085,840,859]" pageId="229" pageNumber="222" value="1980-10-20">20.x.1980</collectingDate>
</date>
,
<specimenCount id="9D2AEBC1FF7209CC6126FC85CEBBFC88" box="[113,145,863,886]" pageId="229" pageNumber="222" type="female">1 ♀</specimenCount>
,
<collectorName id="26D9459EFF7209CC61CBFC85CEC3FC8B" box="[156,233,866,885]" pageId="229" pageNumber="222">A. Olexa</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="3B442A15FF7209CC6042FC85CFDAFC8B" ID-GBIF-Occurrence="2981920349" box="[277,496,863,886]" collectingDate="1987-04-27" collectorName="A. Olexa" country="Algeria" location="Beni Abbes" municipality="Sahara" pageId="229" pageNumber="222" specimenCount="1">
1 ♁, ditto, but
<date id="FF920688FF7209CC60C5FC85CFDAFC8B" box="[402,496,866,885]" pageId="229" pageNumber="222" value="1987-04-27">
<collectingDate id="EFD6FF60FF7209CC60C5FC85CFDAFC8B" box="[402,496,866,885]" pageId="229" pageNumber="222" value="1987-04-27">27.iv.1987</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B442A15FF7209CC60AEFC85CD78FC8B" ID-GBIF-Occurrence="2981920363" box="[505,850,863,886]" collectingDate="1987-04-26" collectorName="A. Olexa" country="Algeria" location="Ain Sefra" pageId="229" pageNumber="222" specimenCount="1">
<location id="8EF37693FF7209CC60AEFC85CC67FC8B" LSID="urn:lsid:plazi:treatment:0385915EFF7209C06036FF3DCECFFCAD:8EF37693FF7209CC60AEFC85CC67FC8B" box="[505,589,866,885]" country="Algeria" name="Ain Sefra" pageId="229" pageNumber="222">Ain Sefra</location>
,
<date id="FF920688FF7209CC6301FC85CC99FC8B" box="[598,691,866,885]" pageId="229" pageNumber="222" value="1987-04-26">
<collectingDate id="EFD6FF60FF7209CC6301FC85CC99FC8B" box="[598,691,866,885]" pageId="229" pageNumber="222" value="1987-04-26">26.iv.1987</collectingDate>
</date>
, 1 ♁,
<collectorName id="26D9459EFF7209CC63B0FC85CD1EFC8B" box="[743,820,866,885]" pageId="229" pageNumber="222">A. Olexa</collectorName>
lgt
</materialsCitation>
.
<materialsCitation id="3B442A15FF7209CC620DFC86CFC0FC6E" ID-GBIF-Occurrence="2981920388" collectionCode="TLAN" collectorName="Quedenfeldt" country="Morocco" location="Mogador" pageId="229" pageNumber="222" specimenCount="2" specimenCount-female="2">
<emphasis id="B958FC5AFF7209CC620DFC86CDE4FC8B" bold="true" box="[858,974,865,885]" pageId="229" pageNumber="222">
<collectingCountry id="F33B60D8FF7209CC620DFC86CDE4FC8B" box="[858,974,865,885]" name="Morocco" pageId="229" pageNumber="222">MOROCCO</collectingCountry>
</emphasis>
:
<location id="8EF37693FF7209CC628DFC85CA02FC8B" LSID="urn:lsid:plazi:treatment:0385915EFF7209C06036FF3DCECFFCAD:8EF37693FF7209CC628DFC85CA02FC8B" box="[986,1064,866,885]" country="Morocco" name="Mogador" pageId="229" pageNumber="222">Mogador</location>
[= Essaouira],
<specimenCount id="9D2AEBC1FF7209CC6181FC9ACF2FFC6F" box="[214,261,890,913]" pageId="229" pageNumber="222" type="female">2 ♀♀</specimenCount>
,
<collectorName id="26D9459EFF7209CC6047FC9ACF53FC6E" box="[272,377,893,912]" pageId="229" pageNumber="222">Quedenfeldt</collectorName>
lgt. (
<collectionCode id="ED3DB88DFF7209CC60F1FC9ACFCEFC6E" box="[422,484,893,912]" pageId="229" pageNumber="222">TLAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF7209C06126FC4FCA03FD0A" lastPageId="233" lastPageNumber="226" pageId="229" pageNumber="222" type="description">
<paragraph id="8B932048FF7209CC6126FC4FCCEFFC1E" blockId="229.[113,1089,936,1632]" pageId="229" pageNumber="222">
<emphasis id="B958FC5AFF7209CC6126FC4FCF3DFC3E" bold="true" box="[113,279,936,960]" pageId="229" pageNumber="222">Redescription.</emphasis>
Body length: PEL:
<quantity id="4CD48DADFF7209CC60BAFC4FCCB3FC3E" box="[493,665,936,960]" metricMagnitude="-3" metricUnit="m" metricValue="2.4875" metricValueMax="2.8" metricValueMin="2.175" pageId="229" pageNumber="222" unit="mm" value="2.4875" valueMax="2.8" valueMin="2.175">2.175–2.80 mm</quantity>
; APW:
<quantity id="4CD48DADFF7209CC63B9FC4FCD8DFC3E" box="[750,935,936,960]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="10.75" metricValueMin="8.25" pageId="229" pageNumber="222" unit="mm" value="0.95" valueMax="1.075" valueMin="0.825">
<geoCoordinate id="EE18468FFF7209CC63B9FC4FCD01FC3E" box="[750,811,936,960]" degrees="0.825" direction="north" orientation="latitude" pageId="229" pageNumber="222" precision="55" value="0.825">0.825</geoCoordinate>
<geoCoordinate id="EE18468FFF7209CC627CFC4FCD59FC3E" box="[811,883,936,960]" degrees="1.075" direction="west" orientation="longitude" pageId="229" pageNumber="222" precision="55" value="-1.075">–1.075</geoCoordinate>
mm
</quantity>
; PPW:
<quantity id="4CD48DADFF7209CC62A0FC4FCECBFC1E" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.125" metricValueMin="1.775" pageId="229" pageNumber="222" unit="mm" value="1.95" valueMax="2.125" valueMin="1.775">
<geoCoordinate id="EE18468FFF7209CC62A0FC4FCA18FC3E" box="[1015,1074,936,960]" degrees="1.775" direction="north" orientation="latitude" pageId="229" pageNumber="222" precision="55" value="1.775">1.775</geoCoordinate>
<geoCoordinate id="EE18468FFF7209CC6565FC4FCE87FC1E" degrees="2.125" direction="west" orientation="longitude" pageId="229" pageNumber="222" precision="55" value="-2.125">– 2.125</geoCoordinate>
mm
</quantity>
; EL:
<quantity id="4CD48DADFF7209CC604DFC2FCFECFC1E" box="[282,454,968,992]" metricMagnitude="-3" metricUnit="m" metricValue="1.5625" metricValueMax="1.7" metricValueMin="1.425" pageId="229" pageNumber="222" unit="mm" value="1.5625" valueMax="1.7" valueMin="1.425">1.425–1.70 mm</quantity>
; EW:
<quantity id="4CD48DADFF7209CC635FFC2FCCEBFC1E" box="[520,705,968,992]" metricMagnitude="-3" metricUnit="m" metricValue="2.225" metricValueMax="2.475" metricValueMin="1.975" pageId="229" pageNumber="222" unit="mm" value="2.225" valueMax="2.475" valueMin="1.975">
<geoCoordinate id="EE18468FFF7209CC635FFC2FCC68FC1E" box="[520,578,968,992]" degrees="1.975" direction="north" orientation="latitude" pageId="229" pageNumber="222" precision="55" value="1.975">1.975</geoCoordinate>
<geoCoordinate id="EE18468FFF7209CC6314FC2FCCA4FC1E" box="[579,654,968,992]" degrees="2.475" direction="west" orientation="longitude" pageId="229" pageNumber="222" precision="55" value="-2.475">–2.475</geoCoordinate>
mm
</quantity>
.
</paragraph>
<paragraph id="8B932048FF7209CC61C7FC0FCC76FBBE" blockId="229.[113,1089,936,1632]" pageId="229" pageNumber="222">
Body (
<figureCitation id="13173CCDFF7209CC618FFC0FCF58FBFF" box="[216,370,1000,1025]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="229" pageNumber="222">Figs. 711–712</figureCitation>
) ovoid, moderately convex from above, underside strongly convex, cuticle light to dark brown with feeble bronze metallic tinge; legs, mouthparts and antennal scape rufopiceous, antennal flagellum yellow.
</paragraph>
<paragraph id="8B932048FF7209CC61C7FBAFCDA8FB1E" blockId="229.[113,1089,936,1632]" pageId="229" pageNumber="222">
Antennal scape (
<figureCitation id="13173CCDFF7209CC6017FBAFCFB0FB9E" box="[320,410,1096,1120]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="229" pageNumber="222">Fig. 715</figureCitation>
) with long numerous setae; club (
<figureCitation id="13173CCDFF7209CC63A9FBAFCD73FB9F" box="[766,857,1096,1121]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="229" pageNumber="222">Fig. 714</figureCitation>
) rather small, without visible articulation, surface (apart from two sensory areas on apical part of club furnished with thick short sensilla, intermingled with thicker sparse erect sensilla) glabrous; sensory structures of antennal club (
<figureCitation id="13173CCDFF7209CC60C9FB4FCFC7FB3F" box="[414,493,1192,1217]" captionStart="Figs" captionStartId="25.[113,152,1384,1403]" captionTargetBox="[140,1062,219,1365]" captionTargetId="figure-12@25.[140,1062,219,1366]" captionTargetPageId="25" captionText="Figs. 9–28. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 9 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 10 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 11 – Myrmetes paykulli Kanaar, 1979; 12 – Styphrus corpulentus Motschulsky, 1845; 13 – Saprinus (Phaonius) pharao (Marseul, 1855); 14 – Ammostyphrus cerberus Reichardt, 1924; 15 – Chalcionellus amoenus (Erichson, 1834); 16 – Chivaenius kryzhanovskii Olexa,1980; 17 – Ctenophilothis chobauti (Théry,1900); 18 – Eopachylopus ripae (Lewis, 1885); 19 – Exaesiopus grossipes grossipes (Marseul, 1855); 20 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 21 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 22 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 23 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 24 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 25 – Paravolvulus lateristrius (Solskij, 1876); 26 – Pholioxenus phoenix (Reichardt, 1930); 27 – Reichardtiolus duriculus (Reitter, 1904); 28 – Xenonychus tridens (Jacquelin-Duval, 1852)." figureDoi="http://doi.org/10.5281/zenodo.4272139" httpUri="https://zenodo.org/record/4272139/files/figure.png" pageId="229" pageNumber="222">Fig. 28</figureCitation>
) in form of stipe-shaped vesicle situated under circular sensory area on internal distal margin of the ventral side of antennal club.
</paragraph>
<paragraph id="8B932048FF7209CC61C7FB0FCCB2F9DE" blockId="229.[113,1089,936,1632]" pageId="229" pageNumber="222">
Mouthparts. Mandibles (
<figureCitation id="13173CCDFF7209CC60C9FB0FCFD6FAFE" box="[414,508,1256,1280]" captionStart="Figs" captionStartId="36.[151,190,1161,1180]" captionTargetBox="[151,1126,219,1130]" captionTargetId="figure-100@36.[151,1126,219,1132]" captionTargetPageId="36" captionText="Figs.99–107.Saprininae, mandibles,ventral view:99 – Philothis (Farabius) hexeris Reichardt, 1930; 100 – Pholioxenus phoenix (Reichardt,1930); 101 – Reichardtiolus duriculus (Reitter, 1904); 102 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 103 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 104 – Saprinus (Phaonius) pharao (Marseul, 1855); 105 – Styphrus corpulentus Motschulsky, 1845; 106 – Xenonychus tridens (Jacquelin-Duval, 1852); 107 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4272155" httpUri="https://zenodo.org/record/4272155/files/figure.png" pageId="229" pageNumber="222">Fig. 106</figureCitation>
) with straight carinate outer margin strongly curved inwardly; mandibular apex acutely pointed; sub-apical tooth on left mandible prominent, acute; labrum (
<figureCitation id="13173CCDFF7209CC6058FACFCF76FABF" box="[271,348,1320,1345]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 47–73. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 – Alienocacculus neftensis (Olexa, 1984); 48 – Ammostyphrus cerberus Reichardt, 1924; 49 – Chalcionellus amoenus (Erichson, 1834); 50 – Chivaenius kryzhanovskii Olexa, 1980; 51 – Eopachylopus ripae (Lewis, 1885); 52 – Eremosaprinus vlasovi (Reichardt, 1941); 53 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 – Exaesiopus grossipes grossipes (Marseul, 55); 55 – Gnathoncus rotundatus (Kugelann, 1792); 56 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 – Microsaprinus therondianus (Dahlgren, 1973); 61 – Myrmetes paykulli Kanaar, 1979; 62 – Paravolvulus lateristriatus (Solskij, 1876); 63 – Philothis (Philothis) arcanus Reichardt, 1930; 64 – Philothis (Atavinus) atavus (Reichardt, 1931); 65 – Philothis (Farabius) hexeris Reichardt, 1930; 66 – Pholioxenus phoenix (Reichardt, 1930); 67 – Reichardtiolus duriculus (Reitter, 1904); 68 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 – Saprinus (Phaonius) pharao (Marseul, 1855); 71 – Styphrus corpulentus Motschulsky, 1845; 72 – Xenonychus tridens (Jacquelin-Duval, 1852); 73 – Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="229" pageNumber="222">Fig. 72</figureCitation>
) almost smooth, slightly convex; labral pits with single weakly sclerotized seta; epipharynx almost completely hidden under labral fold; terminal labial palpomere elongated, its width about one-third its length; mentum sub-trapezoid, anterior margin (
<figureCitation id="13173CCDFF7209CC6542FA8FCEB0FA5F" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109–141. Saprininae, anterior margin of mentum, ventral view: 109 – Alienocacculus neftensis (Olexa, 1984); 110 – Ammostyphrus cerberus Reichardt, 1924; 111 – Axelinus ghilarovi Kryzhanovskij, 1976; 112 – Chalcionellus amoenus (Erichson, 1834); 113 – Chivaenius kryzhanovskii Olexa, 1980; 114 – Ctenophilotis chobauti (Théry, 1900); 115 – Eopachylopus ripae (Lewis, 1885); 116 – Eremosaprinus vlasovi (Reichardt, 1941); 117 – Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 – Exaesiopus grossipes grossipes (Marseul, 1855); 119 – Gnathoncus rotundatus (Kugelann, 1792); 120 – Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 – Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 – Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 – Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 – Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 – Microsaprinus therondianus (Dahlgren, 1973); 126 – Myrmetes paykulli Kanaar, 1979; 127 – Paravolvulus ovillum (Solskij, 1876); 128 – Philothis (Philothis) arcanus Reichardt, 1930; 129 – Philothis (Atavinus) atavus (Reichardt, 1931); 130 – Philothis (Farabius) hexeris Reichardt, 1930; 131 – Pholioxenus phoenix (Reichardt, 1930); 132 – Reichardtiolus duriculus (Reitter, 1904); 133 – Saprinillus paromaloides Kryzhanovskij, 1974; 134 – Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 – Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 – Saprinus (Phaonius) pharao (Marseul, 1855); 137 – Styphrus corpulentus Motschulsky, 1845; 138 – Turanostyphrus ignoratus Tishechkin, 2005; 139 – Xenonychus tridens (Jacquelin-Duval, 1852); 140 – Xenophilotis choumovitchi (Thérond, 1965); 141 – Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="229" pageNumber="222">Fig. 139</figureCitation>
) with shallow median notch, surrounded with several moderately long ramose setae, disc of mentum with few similar setae, lateral margins with single row of shorter ramose setae; cardo of maxilla on lateral margin with several long setae; stipes triangular, with numerous much longer setae; terminal maxillary palpomere elongated, its width about one-third its length, approximately twice as long as penultimate.
</paragraph>
<paragraph id="8B932048FF7209CE61C7F9CFCA2EFA70" blockId="229.[113,1089,936,1632]" lastBlockId="231.[113,1089,1334,1582]" lastPageId="231" lastPageNumber="224" pageId="229" pageNumber="222">
Clypeus (
<figureCitation id="13173CCDFF7209CC61ACF9CFCF76F9BE" box="[251,348,1576,1600]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="229" pageNumber="222">Fig. 715</figureCitation>
) rectangular, broader than long, slightly depressed laterally, with scattered microscopic punctation, almost smooth; frontal stria well impressed, thin, almost straight (occasionally weakened medially or interrupted), continued as well impressed carinate supraorbital stria; frontal disc (
<figureCitation id="13173CCDFF7009CE60B0FAB1CC6AFA90" box="[487,576,1366,1390]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="231" pageNumber="224">Fig. 715</figureCitation>
) flat, almost smooth, broader than long, laterally exceptionally with vague interrupted rugae; eyes very flattened, invisible from above.
</paragraph>
<caption id="DF5370C0FF7109CF61C0F9F1CA4BF9A0" ID-DOI="http://doi.org/10.5281/zenodo.4272310" ID-Zenodo-Dep="4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="230" pageNumber="227" startId="230.[151,189,1558,1577]" targetBox="[228,1051,219,1536]" targetPageId="230">
<paragraph id="8B932048FF7109CF61C0F9F1CA4BF9A0" blockId="230.[151,1129,1558,1631]" pageId="230" pageNumber="227">
Figs.711–720.
<taxonomicName id="4C2C5BCBFF7109CF6041F9F0CCACF9D7" authority="(Jacquelin-Duval, 1852)" authorityName=": GANGLBAUER" authorityYear="1899" baseAuthorityName="Jacquelin-Duval" baseAuthorityYear="1852" box="[278,646,1558,1577]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="230" pageNumber="227" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7109CF6041F9F0CF92F9D7" box="[278,440,1559,1577]" italics="true" pageId="230" pageNumber="227">Xenonychus tridens</emphasis>
(
<bibRefCitation id="EFBD5DB9FF7109CF6096F9F1CC55F9D7" author="JACQUELIN-DUVAL M." box="[449,639,1558,1577]" pageId="230" pageNumber="227" pagination="695 - 718" refId="ref115897" refString="JACQUELIN-DUVAL M. 1852: Descriptions de deux genres nouveaux et de plusieurs especes nouvelles de Coleo / pteres propers a la faune francaise. Annales de la Societe Entomologique de France 10: 695 - 718." type="journal article" year="1852">Jacquelin-Duval,1852</bibRefCitation>
)
</taxonomicName>
, SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view.
</paragraph>
</caption>
<caption id="DF5370C0FF7009CE6126FB62CF34FB16" ID-DOI="http://doi.org/10.5281/zenodo.4272312" ID-Zenodo-Dep="4272312" httpUri="https://zenodo.org/record/4272312/files/figure.png" pageId="231" pageNumber="224" startId="231.[113,152,1157,1176]" targetBox="[194,1001,225,1129]" targetPageId="231">
<paragraph id="8B932048FF7009CE6126FB62CF34FB16" blockId="231.[113,1089,1155,1256]" pageId="231" pageNumber="224">
Figs. 721–729.
<taxonomicName id="4C2C5BCBFF7009CE61A1FB61CC5EFB66" authority="(Jacquelin-Duval, 1852)" authorityName=": GANGLBAUER" authorityYear="1899" baseAuthorityName="Jacquelin-Duval" baseAuthorityYear="1852" box="[246,628,1157,1176]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="231" pageNumber="224" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7009CE61A1FB61CFB7FB66" box="[246,413,1158,1176]" italics="true" pageId="231" pageNumber="224">Xenonychus tridens</emphasis>
(
<bibRefCitation id="EFBD5DB9FF7009CE60FEFB62CC47FB66" author="JACQUELIN-DUVAL M." box="[425,621,1157,1176]" pageId="231" pageNumber="224" pagination="695 - 718" refId="ref115897" refString="JACQUELIN-DUVAL M. 1852: Descriptions de deux genres nouveaux et de plusieurs especes nouvelles de Coleo / pteres propers a la faune francaise. Annales de la Societe Entomologique de France 10: 695 - 718." type="journal article" year="1852">Jacquelin-Duval, 1852</bibRefCitation>
)
</taxonomicName>
, male terminalia: 721 – 8
<superScript id="7C598D00FF7009CE621AFB64CD7DFB71" attach="left" box="[845,855,1155,1167]" fontSize="5" pageId="231" pageNumber="224">th</superScript>
sternite and tergite, ventral view; 722 – ditto, dorsal view; 723 – ditto, lateral view; 724 – 9
<superScript id="7C598D00FF7009CE63CFFB79CC88FB54" attach="left" box="[664,674,1182,1194]" fontSize="5" pageId="231" pageNumber="224">th</superScript>
tergite and 10
<superScript id="7C598D00FF7009CE6249FB79CD02FB54" attach="left" box="[798,808,1182,1194]" fontSize="5" pageId="231" pageNumber="224">th</superScript>
tergite, dorsal view; 725 – ditto, lateral view; 726 – spiculum gastrale, ventral view; 727 – ditto, lateral view; 728 – aedeagus, dorsal view; 729 – ditto, lateral view.
</paragraph>
</caption>
<paragraph id="8B932048FF7009CE61C7FA71CFA5F9D0" blockId="231.[113,1089,1334,1582]" pageId="231" pageNumber="224">
Pronotal sides (
<figureCitation id="13173CCDFF7009CE606AFA71CFB7FA50" box="[317,413,1430,1454]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="231" pageNumber="224">Fig. 711</figureCitation>
) moderately convergent forwardly; apical angles prominent; anterior incision for head shallow; marginal pronotal stria well impressed, complete; disc entirely covered with shallow round regular punctation, becoming denser laterally; pronotal base with a row of oval dense punctures; pronotal hypomeron with long yellow setae; scutellum small, conspicuous.
</paragraph>
<paragraph id="8B932048FF7F09C161E2FF3CCC76FDED" blockId="232.[151,1127,219,1587]" pageId="232" pageNumber="229">Elytral epipleuron with round, rather deep punctures fringed with long setae; marginal epipleural stria complete; marginal elytral stria deeply impressed, continued as weakly impressed (at times intermittent) apical elytral stria. Humeral elytral stria distinctly impressed on basal third; inner subhumeral stria present medially, deep, rather long, rarely joining marginal elytral stria; elytra with four dorsal elytral striae 1–4, almost reaching elytral apex (seldom fourth dorsal elytral stria slightly shortened apically), fourth dorsal elytral stria basally connected with sutural elytral stria; sutural elytral stria well impressed, apically usually connected with apical elytral stria. Elytral punctation confined to interspace between fourth dorsal and sutural elytral striae, basally along elytral suture almost reaching elytral base, punctation irregular and shallow, becoming confluent apically.</paragraph>
<paragraph id="8B932048FF7F09C161E2FDFCCF6FFD8D" blockId="232.[151,1127,219,1587]" pageId="232" pageNumber="229">
Propygidium (
<figureCitation id="13173CCDFF7F09C16006FDFCCF84FDCA" box="[337,430,539,564]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="232" pageNumber="229">Fig. 718</figureCitation>
) completely exposed, with shallow confluent punctures of various sizes; pygidium (
<figureCitation id="13173CCDFF7F09C1602CFDDCCFFCFDAA" box="[379,470,571,596]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="232" pageNumber="229">Fig. 718</figureCitation>
) long, with similar punctation, punctures becoming finer and sparser apically.
</paragraph>
<paragraph id="8B932048FF7F09C161E2FD9CCD50FCAD" blockId="232.[151,1127,219,1587]" pageId="232" pageNumber="229">
Anterior margin of median portion of prosternum (
<figureCitation id="13173CCDFF7F09C163B0FD9CCD6FFD6A" box="[743,837,635,660]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="232" pageNumber="229">Fig. 719</figureCitation>
) almost straight; marginal prosternal stria present laterally and as short anterior fragment; pre-apical foveae deep; prosternal process slightly concave, compressed, surface between lateral prosternal striae substrigulate-punctate, punctures shallow, surface laterad to lateral prosternal striae substrigulate, impunctate; carinal prosternal striae divergent on prosternal apophysis, thence strongly convergent, intermittent, vaguely attaining united lateral prosternal striae; lateral prosternal striae well impressed, carinate, convergent anteriorly, united in front.
</paragraph>
<paragraph id="8B932048FF7F09C161E2FCBCCC64FB8D" blockId="232.[151,1127,219,1587]" pageId="232" pageNumber="229">
Anterior margin of mesoventrite (
<figureCitation id="13173CCDFF7F09C16372FCBCCCABFC8A" box="[549,641,859,884]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="232" pageNumber="229">Fig. 713</figureCitation>
) shallowly emarginate medially; discal marginal mesoventral stria well impressed, carinate, weakened anteriorly; disc with scattered deep punctures separated by about 2–3 times their diameter; meso-metaventral sutural stria (
<figureCitation id="13173CCDFF7F09C161C8FC5CCED5FC2A" box="[159,255,955,980]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="232" pageNumber="229">Fig. 713</figureCitation>
) well impressed, almost straight, undulate, exposing meso-metaventral suture; intercoxal disc of metaventrite almost smooth, only with microscopic punctation, along base with a band of irregular deep punctation; lateral metaventral stria well impressed, carinate, obliquely arcuate, shortened apically; lateral disc of metaventrite concave, with deep setiferous punctures of various sizes; metepisternum + fused metepimeron with very dense setae, punctation unrecognizable beneath them.
</paragraph>
<paragraph id="8B932048FF7F09C161E2FB9CCA04FB0D" blockId="232.[151,1127,219,1587]" pageId="232" pageNumber="229">Intercoxal disc of the first abdominal sternite almost completely striate laterally; basal half of disc with punctures of varied sizes, separated by about their own to twice their own diameter, apically replaced by very fine scattered microscopic punctation; along apical margin a row of larger punctures appears; lateral disc of all visible abdominal sternites setose laterally.</paragraph>
<paragraph id="8B932048FF7F09C161E2FB1CCA0CF9CD" blockId="232.[151,1127,219,1587]" pageId="232" pageNumber="229">
Protibia (
<figureCitation id="13173CCDFF7F09C1604CFB1CCF52FAEA" box="[283,376,1275,1300]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="232" pageNumber="229">Fig. 720</figureCitation>
) flattened and dilated, outer margin with three large triangular teeth topped with moderately large triangular denticle, followed by five short rounded denticles diminishing in size in proximal direction; setae of outer row long, confined to basal third of protibia; setae of median row much shorter than those of outer row, but strongly sclerotized and present almost along entire length of protibia; protarsal groove rather deep; anterior protibial stria inconspicuous; single, short tarsal denticle present apically; protibial spur tiny, bent, growing out from apical protibial margin; apical margin of protibia posteriorly with two tiny apical denticles; outer part of posterior surface of protibia areolate-rugose; distinctly separated from glabrous median part of posterior surface; posterior protibial stria complete; inner-ventral denticles absent; inner margin with row of strongly sclerotized lamelliform long setae.
</paragraph>
<paragraph id="8B932048FF7E09C061C7FF3CCF84FDCD" blockId="233.[113,1089,219,851]" pageId="233" pageNumber="226">
Mesotibia (
<figureCitation id="13173CCDFF7E09C06050FF3CCF75FF0D" box="[263,351,219,243]" captionStart="Figs" captionStartId="230.[151,189,1558,1577]" captionTargetBox="[228,1051,219,1536]" captionTargetId="figure-11@230.[228,1051,219,1537]" captionTargetPageId="230" captionText="Figs.711–720.Xenonychus tridens (Jacquelin-Duval,1852), SEM micrographs:711 – habitus,dorsal view; 712 – ditto, ventral view; 713 – mesoventrite; 714 – antennal club, ventral view; 715 – head, dorsal view; 716 – mesotibia, dorsal view; 717 – head, ventral view; 718 – propygidium and pygidium; 719 – prosternum; 720 – protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272310" httpUri="https://zenodo.org/record/4272310/files/figure.png" pageId="233" pageNumber="226">Fig. 716</figureCitation>
) slightly thickened, outer margin with two dense rows of thin denticles growing in size apically, abutting each other; posterior mesotibial surface with dense, strongly sclerotized long setae growing out near inner mesotibial margin and covering almost entire posterior surface of mesotibia, their affiliation with either dorsal or median row dubious; posterior mesotibial stria inconspicuous; anterior surface of mesotibia glabrous; anterior mesotibial stria almost complete, shortened apically; mesotibial spur long and thin; apical margin with several short denticles; claws of apical tarsomere almost straight, almost as long as apical-most mesotarsomeres itself; each tarsomere with single long strongly sclerotized seta posteriorly and anteriorly; metatibia basically similar to mesotibia, but more thickened and dilated and denticles on outer margin much longer, three rows of denticles present (as opposed to two in mesotibia).
</paragraph>
<paragraph id="8B932048FF7E09C061C7FDDCCA03FD0A" blockId="233.[113,1089,219,851]" pageId="233" pageNumber="226">
Male genitalia. Eighth sternite (
<figureCitation id="13173CCDFF7E09C0608FFDDCCC5BFDAA" box="[472,625,571,596]" captionStart="Figs" captionStartId="231.[113,152,1157,1176]" captionTargetBox="[194,1001,225,1129]" captionTargetId="figure-99@231.[151,1054,216,1136]" captionTargetPageId="231" captionText="Figs. 721–729. Xenonychus tridens (Jacquelin-Duval, 1852), male terminalia: 721 – 8th sternite and tergite, ventral view; 722 – ditto, dorsal view; 723 – ditto, lateral view; 724 – 9th tergite and 10th tergite, dorsal view; 725 – ditto, lateral view; 726 – spiculum gastrale, ventral view; 727 – ditto, lateral view; 728 – aedeagus, dorsal view; 729 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272312" httpUri="https://zenodo.org/record/4272312/files/figure.png" pageId="233" pageNumber="226">Figs. 721–722</figureCitation>
) segment longitudinally separated medially, apically with tiny inflatable membrane (velum); apex laterally fringed with short brush of setae; eighth tergite and eighth sternite fused laterally (
<figureCitation id="13173CCDFF7E09C06397FD9CCD35FD6A" box="[704,799,635,660]" captionStart="Figs" captionStartId="231.[113,152,1157,1176]" captionTargetBox="[194,1001,225,1129]" captionTargetId="figure-99@231.[151,1054,216,1136]" captionTargetPageId="231" captionText="Figs. 721–729. Xenonychus tridens (Jacquelin-Duval, 1852), male terminalia: 721 – 8th sternite and tergite, ventral view; 722 – ditto, dorsal view; 723 – ditto, lateral view; 724 – 9th tergite and 10th tergite, dorsal view; 725 – ditto, lateral view; 726 – spiculum gastrale, ventral view; 727 – ditto, lateral view; 728 – aedeagus, dorsal view; 729 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272312" httpUri="https://zenodo.org/record/4272312/files/figure.png" pageId="233" pageNumber="226">Fig. 723</figureCitation>
). Morphology of 9
<superScript id="7C598D00FF7E09C062BEFD9DCDDFFD77" attach="left" box="[1001,1013,634,649]" fontSize="6" pageId="233" pageNumber="226">th</superScript>
tergite (
<figureCitation id="13173CCDFF7E09C0612EFD7CCF3AFD4D" box="[121,272,667,691]" captionStart="Figs" captionStartId="231.[113,152,1157,1176]" captionTargetBox="[194,1001,225,1129]" captionTargetId="figure-99@231.[151,1054,216,1136]" captionTargetPageId="231" captionText="Figs. 721–729. Xenonychus tridens (Jacquelin-Duval, 1852), male terminalia: 721 – 8th sternite and tergite, ventral view; 722 – ditto, dorsal view; 723 – ditto, lateral view; 724 – 9th tergite and 10th tergite, dorsal view; 725 – ditto, lateral view; 726 – spiculum gastrale, ventral view; 727 – ditto, lateral view; 728 – aedeagus, dorsal view; 729 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272312" httpUri="https://zenodo.org/record/4272312/files/figure.png" pageId="233" pageNumber="226">Figs. 724–725</figureCitation>
) typical for the subfamily; spiculum gastrale (
<figureCitation id="13173CCDFF7E09C063BEFD7CCD6BFD4D" box="[745,833,667,691]" captionStart="Figs" captionStartId="231.[113,152,1157,1176]" captionTargetBox="[194,1001,225,1129]" captionTargetId="figure-99@231.[151,1054,216,1136]" captionTargetPageId="231" captionText="Figs. 721–729. Xenonychus tridens (Jacquelin-Duval, 1852), male terminalia: 721 – 8th sternite and tergite, ventral view; 722 – ditto, dorsal view; 723 – ditto, lateral view; 724 – 9th tergite and 10th tergite, dorsal view; 725 – ditto, lateral view; 726 – spiculum gastrale, ventral view; 727 – ditto, lateral view; 728 – aedeagus, dorsal view; 729 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272312" httpUri="https://zenodo.org/record/4272312/files/figure.png" pageId="233" pageNumber="226">Fig. 726</figureCitation>
) expanded on both ends. Basal piece of aedeagus (
<figureCitation id="13173CCDFF7E09C0602CFD5CCC3CFD2A" box="[379,534,699,724]" captionStart="Figs" captionStartId="231.[113,152,1157,1176]" captionTargetBox="[194,1001,225,1129]" captionTargetId="figure-99@231.[151,1054,216,1136]" captionTargetPageId="231" captionText="Figs. 721–729. Xenonychus tridens (Jacquelin-Duval, 1852), male terminalia: 721 – 8th sternite and tergite, ventral view; 722 – ditto, dorsal view; 723 – ditto, lateral view; 724 – 9th tergite and 10th tergite, dorsal view; 725 – ditto, lateral view; 726 – spiculum gastrale, ventral view; 727 – ditto, lateral view; 728 – aedeagus, dorsal view; 729 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272312" httpUri="https://zenodo.org/record/4272312/files/figure.png" pageId="233" pageNumber="226">Figs. 728–729</figureCitation>
) rather short, ratio of its length: length of parameres 1: 4; parameres fused along their basal half; aedeagus slightly curved ventrad (
<figureCitation id="13173CCDFF7E09C062E8FD3CCA37FD0A" box="[959,1053,731,756]" captionStart="Figs" captionStartId="231.[113,152,1157,1176]" captionTargetBox="[194,1001,225,1129]" captionTargetId="figure-99@231.[151,1054,216,1136]" captionTargetPageId="231" captionText="Figs. 721–729. Xenonychus tridens (Jacquelin-Duval, 1852), male terminalia: 721 – 8th sternite and tergite, ventral view; 722 – ditto, dorsal view; 723 – ditto, lateral view; 724 – 9th tergite and 10th tergite, dorsal view; 725 – ditto, lateral view; 726 – spiculum gastrale, ventral view; 727 – ditto, lateral view; 728 – aedeagus, dorsal view; 729 – ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272312" httpUri="https://zenodo.org/record/4272312/files/figure.png" pageId="233" pageNumber="226">Fig. 729</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF7E09C06126FD1BCECFFCAD" pageId="233" pageNumber="226" type="discussion">
<paragraph id="8B932048FF7E09C06126FD1BCECFFCAD" blockId="233.[113,1089,219,851]" pageId="233" pageNumber="226">
<emphasis id="B958FC5AFF7E09C06126FD1BCEF3FCEA" bold="true" box="[113,217,764,788]" pageId="233" pageNumber="226">Remarks</emphasis>
.
<taxonomicName id="4C2C5BCBFF7E09C061BFFD1ACF91FCEA" authorityName=": GANGLBAUER" authorityYear="1899" baseAuthorityName="Jacquelin-Duval" baseAuthorityYear="1852" box="[232,443,765,788]" class="Insecta" family="Histeridae" genus="Xenonychus" kingdom="Animalia" order="Coleoptera" pageId="233" pageNumber="226" phylum="Arthropoda" rank="species" species="tridens">
<emphasis id="B958FC5AFF7E09C061BFFD1ACF91FCEA" box="[232,443,765,788]" italics="true" pageId="233" pageNumber="226">Xenonychus tridens</emphasis>
</taxonomicName>
is a widely distributed species and as such exhibits a large degree of variation, especially regarding punctation of dorsal surface, frontal stria and other characters.
</paragraph>
</subSubSection>
</treatment>
</document>
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