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<document id="B8E8FF018C19C8C0EF5A6D8569E1BD04" ID-CLB-Dataset="9574" ID-DOI="10.5281/zenodo.4272127" ID-GBIF-Dataset="724cb358-6f18-4816-afc7-bb42cb9b6942" ID-ISSN="0374-1036" ID-Zenodo-Dep="4272127" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1605298182318" checkinUser="felipe" docAuthor="Lackner, Tomáš" docDate="2010" docId="0385915EFF38099A600AFF3DCF34FCCD" docLanguage="en" docName="ActaEntMusNatPra.50.suppl.1-254.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (suppl.) 50" docStyle="DocumentStyle:CE2601992B79F4B08764287CD82450D1.3:ActaEntMusNatPra.2010-2016.monograph" docStyleId="CE2601992B79F4B08764287CD82450D1" docStyleName="ActaEntMusNatPra.2010-2016.monograph" docStyleVersion="3" docTitle="Philothis (Atavinus) atavus" docType="treatment" docVersion="18" lastPageNumber="172" masterDocId="FFBCE926FF9709296157FFE7CE2AFFFE" masterDocTitle="Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)" masterLastPageNumber="254" masterPageNumber="1" pageNumber="168" updateTime="1698846217483" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B6705782C1E628301FA5883A6DDC5D36">Review of the Palaearctic genera of Saprininae (Coleoptera: Histeridae)</mods:title>
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<taxonomicName id="4C2C5BCBFF380986600AFF3DCD7CFF0D" ID-CoL="7WD7Q" authority="(Reichardt, 1931)" baseAuthorityName="Reichardt" baseAuthorityYear="1931" box="[349,854,218,243]" class="Insecta" family="Histeridae" genus="Philothis" kingdom="Animalia" order="Coleoptera" pageId="175" pageNumber="168" phylum="Arthropoda" rank="species" species="atavus" subGenus="Atavinus">
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<emphasis id="B958FC5AFF380986600AFF3DCFEAFF0D" bold="true" box="[349,448,218,243]" italics="true" pageId="175" pageNumber="168">Philothis</emphasis>
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<emphasis id="B958FC5AFF3809866087FF3CCC18FF0D" bold="true" box="[464,562,219,243]" italics="true" pageId="175" pageNumber="168">Atavinus</emphasis>
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<emphasis id="B958FC5AFF3809866316FF3CCCA2FF0C" bold="true" box="[577,648,219,242]" italics="true" pageId="175" pageNumber="168">atavus</emphasis>
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<bibRefCitation id="EFBD5DB9FF38098663C1FF3CCD64FF0D" author="REICHARDT A." box="[662,846,219,243]" pageId="175" pageNumber="168" pagination="308 - 311" refId="ref120322" refString="REICHARDT A. 1931: Sur une nouveau representant du genre Philothis Rchdt. (Coleoptera: Histeridae). Doklady Akademii Nauk SSSR A: 308 - 311." type="book chapter" year="1931">Reichardt, 1931</bibRefCitation>
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<paragraph id="8B932048FF38098660E2FF1ECCD7FEF2" blockId="175.[349,854,218,269]" box="[437,765,249,269]" pageId="175" pageNumber="168">
(
<figureCitation id="13173CCDFF38098660ECFF1ECFDDFEF2" box="[443,503,249,268]" captionStart="Figs" captionStartId="22.[151,190,757,776]" captionTargetBox="[151,1126,216,722]" captionTargetId="figure-310@22.[151,1126,213,724]" captionTargetPageId="22" captionText="Figs. 57. Saprininae, 8th antennomere: 5 Paravolvulus ovillum (Solskij, 1876); 6 Philothis (Atavinus) atavus (Reichardt, 1931); 7 Turanostyphrus ignoratus Tishechkin, 2005." figureDoi="http://doi.org/10.5281/zenodo.4272135" httpUri="https://zenodo.org/record/4272135/files/figure.png" pageId="175" pageNumber="168">Figs. 6</figureCitation>
,
<figureCitation id="13173CCDFF3809866356FF1DCC32FEF3" box="[513,536,250,269]" captionStart="Figs" captionStartId="26.[150,189,1085,1104]" captionTargetBox="[202,1075,219,1057]" captionTargetId="figure-156@26.[202,1076,219,1058]" captionTargetPageId="26" captionText="Figs. 2938. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 29 Xenophilothis choumovitchi (Thérond, 1965); 30 Zorius funereus (Schmidt, 1890); 31 Alienocacculus neftensis (Olexa, 1984); 32 Philothis (Philothis) arcanus Reichardt, 1930; 33 Philothis (Atavinus) atavus (Reichardt, 1931); 34 Philothis (Farabius) hexeris Reichardt, 1930; 35 Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 36 Gnathoncus rotundatus (Kugelann, 1792); 37 Eremosaprinus vlasovi (Reichardt, 1941); 38 Microsaprinus therondianus (Dahlgren, 1973)." figureDoi="http://doi.org/10.5281/zenodo.4272141" httpUri="https://zenodo.org/record/4272141/files/figure.png" pageId="175" pageNumber="168">33</figureCitation>
,
<figureCitation id="13173CCDFF3809866376FF1DCC12FEF3" box="[545,568,250,269]" captionStart="Figs" captionStartId="30.[152,191,1491,1510]" captionTargetBox="[229,1051,894,1466]" captionTargetId="figure-295@30.[229,1051,894,1510]" captionTargetPageId="30" captionText="Figs. 3944. Saprininae, labrum, lateral view: 39 Philothis (Farabius) hexeris Reichardt, 1930; 40 Philothis (Atavinus) atavus (Reichardt, 1931); 41 Saprinus (Phaonius) pharao (Marseul, 1855); 42 Eremosaprinus vlasovi (Reichardt, 1941); 43 Ctenophilothis chobauti (Théry, 1900); 44 Philothis (Philothis) arcanus Reichardt, 1930." figureDoi="http://doi.org/10.5281/zenodo.4272143" httpUri="https://zenodo.org/record/4272143/files/figure.png" pageId="175" pageNumber="168">40</figureCitation>
,
<figureCitation id="13173CCDFF3809866316FF1ECC72FEF2" box="[577,600,249,268]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 4773. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 Alienocacculus neftensis (Olexa, 1984); 48 Ammostyphrus cerberus Reichardt, 1924; 49 Chalcionellus amoenus (Erichson, 1834); 50 Chivaenius kryzhanovskii Olexa, 1980; 51 Eopachylopus ripae (Lewis, 1885); 52 Eremosaprinus vlasovi (Reichardt, 1941); 53 Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 Exaesiopus grossipes grossipes (Marseul, 55); 55 Gnathoncus rotundatus (Kugelann, 1792); 56 Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 Microsaprinus therondianus (Dahlgren, 1973); 61 Myrmetes paykulli Kanaar, 1979; 62 Paravolvulus lateristriatus (Solskij, 1876); 63 Philothis (Philothis) arcanus Reichardt, 1930; 64 Philothis (Atavinus) atavus (Reichardt, 1931); 65 Philothis (Farabius) hexeris Reichardt, 1930; 66 Pholioxenus phoenix (Reichardt, 1930); 67 Reichardtiolus duriculus (Reitter, 1904); 68 Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 Saprinus (Phaonius) pharao (Marseul, 1855); 71 Styphrus corpulentus Motschulsky, 1845; 72 Xenonychus tridens (Jacquelin-Duval, 1852); 73 Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="175" pageNumber="168">64</figureCitation>
,
<figureCitation id="13173CCDFF3809866336FF1DCC52FEF3" box="[609,632,250,269]" captionStart="Figs" captionStartId="35.[114,153,1183,1202]" captionTargetBox="[113,1088,219,1152]" captionTargetId="figure-95@35.[113,1088,219,1153]" captionTargetPageId="35" captionText="Figs. 9098. Saprininae, mandibles, ventral view: 90 Gnathoncus rotundatus (Kugelann, 1792); 91 Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 92 Hypocacculus (Hypocacculus) metalescens (Erichson, 1834); 93 Hypocacculus (Colpellus) praecox (Erichson, 1834); 94 Hypocacculus (Nessus) rubripes (Erichson, 1834); 95 Myrmetes paykulli Kanaar, 1979; 96 Paravolvulus lateristrius (Solskij, 1876); 97 Philothis (Philothis) arcanus Reichardt, 1930; 98 Philothis (Atavinus) atavus (Reichardt, 1931)." figureDoi="http://doi.org/10.5281/zenodo.4272153" httpUri="https://zenodo.org/record/4272153/files/figure.png" pageId="175" pageNumber="168">98</figureCitation>
,
<figureCitation id="13173CCDFF38098663D6FF1DCC89FEF3" box="[641,675,250,269]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109141. Saprininae, anterior margin of mentum, ventral view: 109 Alienocacculus neftensis (Olexa, 1984); 110 Ammostyphrus cerberus Reichardt, 1924; 111 Axelinus ghilarovi Kryzhanovskij, 1976; 112 Chalcionellus amoenus (Erichson, 1834); 113 Chivaenius kryzhanovskii Olexa, 1980; 114 Ctenophilotis chobauti (Théry, 1900); 115 Eopachylopus ripae (Lewis, 1885); 116 Eremosaprinus vlasovi (Reichardt, 1941); 117 Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 Exaesiopus grossipes grossipes (Marseul, 1855); 119 Gnathoncus rotundatus (Kugelann, 1792); 120 Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 Microsaprinus therondianus (Dahlgren, 1973); 126 Myrmetes paykulli Kanaar, 1979; 127 Paravolvulus ovillum (Solskij, 1876); 128 Philothis (Philothis) arcanus Reichardt, 1930; 129 Philothis (Atavinus) atavus (Reichardt, 1931); 130 Philothis (Farabius) hexeris Reichardt, 1930; 131 Pholioxenus phoenix (Reichardt, 1930); 132 Reichardtiolus duriculus (Reitter, 1904); 133 Saprinillus paromaloides Kryzhanovskij, 1974; 134 Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 Saprinus (Phaonius) pharao (Marseul, 1855); 137 Styphrus corpulentus Motschulsky, 1845; 138 Turanostyphrus ignoratus Tishechkin, 2005; 139 Xenonychus tridens (Jacquelin-Duval, 1852); 140 Xenophilotis choumovitchi (Thérond, 1965); 141 Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="175" pageNumber="168">129</figureCitation>
,
<figureCitation id="13173CCDFF38098663FCFF1ECCDDFEF2" box="[683,759,249,268]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="176.[151,191,1487,1506]" captionStartId-1="177.[115,154,1197,1216]" captionTargetBox-0="[245,1034,219,1471]" captionTargetBox-1="[256,940,223,1169]" captionTargetId-0="figure-11@176.[245,1034,219,1482]" captionTargetId-1="figure-107@177.[224,977,219,1184]" captionTargetPageId-0="176" captionTargetPageId-1="177" captionText-0="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." captionText-1="Figs. 552558.Philothis (Atavinus) atavus (Reichardt, 1931), male terminalia: 552 8th sternite and tergite, ventral view; 553 ditto, dorsal view; 554 ditto, lateral view; 555 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 556 9th tergite, 10th tergite and spiculum gastrale, lateral view; 557 aedeagus, dorsal view; 558 ditto, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.4272270" figureDoi-1="http://doi.org/10.5281/zenodo.4272272" httpUri-0="https://zenodo.org/record/4272270/files/figure.png" httpUri-1="https://zenodo.org/record/4272272/files/figure.png" pageId="175" pageNumber="168">543558</figureCitation>
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<treatmentCitation id="0A8D0659FF3809866126FEFBCF9CFED1" author="REICHARDT A." box="[113,438,284,303]" page="308" pageId="175" pageNumber="168" year="1931">
<taxonomicName id="4C2C5BCBFF3809866126FEFBCF9CFED1" ID-CoL="7WD7Q" authority="Reichardt, 1931: 308" authorityName="Reichardt" authorityPageNumber="308" authorityYear="1931" box="[113,438,284,303]" class="Insecta" family="Histeridae" genus="Philothis" kingdom="Animalia" order="Coleoptera" pageId="175" pageNumber="168" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="B958FC5AFF3809866126FEFBCED6FED0" box="[113,252,284,303]" italics="true" pageId="175" pageNumber="168">Philothis atavus</emphasis>
<bibRefCitation id="EFBD5DB9FF3809866056FEFBCF9CFED1" author="REICHARDT A." box="[257,438,284,303]" pageId="175" pageNumber="168" pagination="308 - 311" refId="ref120322" refString="REICHARDT A. 1931: Sur une nouveau representant du genre Philothis Rchdt. (Coleoptera: Histeridae). Doklady Akademii Nauk SSSR A: 308 - 311." type="book chapter" year="1931">Reichardt, 1931: 308</bibRefCitation>
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, Figs. 14.
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<treatmentCitationGroup id="AB3C0766FF3809866126FED1CF4CFE9A" pageId="175" pageNumber="168">
<treatmentCitation id="0A8D0659FF3809866126FED1CFE4FEB7" author="REICHARDT A." box="[113,462,310,330]" page="338" pageId="175" pageNumber="168" year="1941">
<taxonomicName id="4C2C5BCBFF3809866126FED1CF89FEB7" ID-CoL="7WD7Q" authority=": REICHARDT (1941)" authorityName="REICHARDT" authorityYear="1941" box="[113,419,310,330]" class="Insecta" family="Histeridae" genus="Philothis" kingdom="Animalia" order="Coleoptera" pageId="175" pageNumber="168" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="B958FC5AFF3809866126FED1CED6FEB7" box="[113,252,310,329]" italics="true" pageId="175" pageNumber="168">Philothis atavus</emphasis>
:
<bibRefCitation id="EFBD5DB9FF380986605FFED0CF89FEB7" author="REICHARDT A." box="[264,419,310,330]" pageId="175" pageNumber="168" refId="ref120382" refString="REICHARDT A. 1941: Semeystva Sphaeritidae i Histeridae (Vol. 1). [Families Sphaeritidae and Histeridae]. Fauna SSSR, Nasekomye Zhestokrylye, V, 3. Nauka, Moskva-Leningrad, xiii + 419 pp." type="book" year="1941">REICHARDT (1941)</bibRefCitation>
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: 338
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, 340, 409,
<figureCitation id="13173CCDFF380986637AFED1CD19FEB6" box="[557,819,310,330]" captionStart="Figs" captionStartId="59.[113,152,494,513]" captionTargetBox="[182,1019,223,464]" captionTargetId="figure-200@59.[182,1020,223,470]" captionTargetPageId="59" captionText="Figs. 174178. Variability of the caudal end of spiculum gastrale:174 Philothis (Farabius) hexeris Reichardt,1930; 175 Hypocacculus (Nessus) rubripes (Erichson, 1834); 176 Microsaprinus therondianus (Dahlgren, 1973); 177 Saprinus (Saprinus) semistriatus (Scriba, 1790); 178 Euspilotus (Neosaprinus) perrisi (Marseul, 1972)." figureDoi="http://doi.org/10.5281/zenodo.4272186" httpUri="https://zenodo.org/record/4272186/files/figure.png" pageId="175" pageNumber="168">
Figs. 176 A,B;
<treatmentCitation id="0A8D0659FF38098663E5FED0CD19FEB6" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[690,819,311,330]" page="248" pageId="175" pageNumber="168" year="1976">
<bibRefCitation id="EFBD5DB9FF38098663E5FED0CD19FEB6" author="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N." box="[690,819,311,330]" pageId="175" pageNumber="168" refId="ref117064" refString="KRYZHANOVSKIJ O. L. &amp; REICHARDT A. N. 1976: Zhuki Nadsemeystva Histeroidea (semeystva Sphaeritidae, Histeridae, Synteliidae). [Beetles of the superfamily Histeroidea (families Sphaeritidae, Histeridae, Syntelidae)]. Fauna SSSR, Zhestokrylye, Vyp. 4. Nauka, Leningrad, 434 pp (in Russian)." type="book" year="1976">KRYZHANOVSKIJ</bibRefCitation>
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&amp; REICHARDT (1976): 248,
<figureCitation id="13173CCDFF3809866548FED1CE99FE9A" captionStart="Figs" captionStartId="157.[114,153,986,1005]" captionTargetBox="[200,991,252,941]" captionTargetId="figure-171@157.[180,1022,219,965]" captionTargetPageId="157" captionText="Figs. 482490. Microsaprinus therondianus (Dahlgren 1973), male terminalia: 482 8th sternite and tergite, ventral view; 483 ditto, dorsal view; 484 ditto, lateral view; 485 9th tergite, 10th tergite, dorsal view; 486 ditto, lateral view; 487 aedeagus, dorsal view; 488 ditto, lateral view; 489 spiculum gastrale, ventral view; 490 ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272254" httpUri="https://zenodo.org/record/4272254/files/figure.png" pageId="175" pageNumber="168">Fig. 484</figureCitation>
;
<treatmentCitation id="0A8D0659FF38098661ECFEB6CF48FE9A" author="MAZUR S." box="[187,354,337,356]" page="109" pageId="175" pageNumber="168" year="1984">
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: 109
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B932048FF3809866126FE8BCDCDFE81" blockId="175.[113,1090,284,383]" box="[113,999,364,383]" pageId="175" pageNumber="168">
<treatmentCitationGroup id="AB3C0766FF3809866126FE8BCDCDFE81" box="[113,999,364,383]" pageId="175" pageNumber="168">
<treatmentCitation id="0A8D0659FF3809866126FE8BCC22FE81" author="OLEXA A." box="[113,520,364,383]" page="148" pageId="175" pageNumber="168" year="1990">
<taxonomicName id="4C2C5BCBFF3809866126FE8BCFF7FE81" ID-CoL="7WD7Q" authority=": OLEXA (1990)" authorityName=": OLEXA" authorityYear="1990" box="[113,477,364,383]" class="Insecta" family="Histeridae" genus="Philothis" kingdom="Animalia" order="Coleoptera" pageId="175" pageNumber="168" phylum="Arthropoda" rank="species" species="atavus" subGenus="Atavinus">
<emphasis id="B958FC5AFF3809866126FE8BCE94FE81" box="[113,190,364,383]" italics="true" pageId="175" pageNumber="168">Philothis</emphasis>
(
<emphasis id="B958FC5AFF380986619CFE8BCF3CFE81" box="[203,278,364,383]" italics="true" pageId="175" pageNumber="168">Atavinus</emphasis>
)
<emphasis id="B958FC5AFF3809866075FE8BCF70FE80" box="[290,346,364,382]" italics="true" pageId="175" pageNumber="168">atavus</emphasis>
:
<bibRefCitation id="EFBD5DB9FF3809866032FE8BCFF7FE81" author="OLEXA A." box="[357,477,364,383]" pageId="175" pageNumber="168" pagination="141 - 155" refId="ref119556" refString="OLEXA A. 1990: On the genus Philothis and related genera (Coleoptera: Histeridae). Acta Entomologica Bohemoslovaca 87: 141 - 155." type="journal article" year="1990">OLEXA (1990)</bibRefCitation>
</taxonomicName>
: 148
</treatmentCitation>
,
<figureCitation id="13173CCDFF3809866347FE8BCC66FE81" box="[528,588,364,383]" captionStart="Fig" captionStartId="18.[151,182,1516,1535]" captionTargetBox="[151,1051,549,1543]" captionTargetId="figure-124@18.[200,1051,549,1544]" captionTargetPageId="18" captionText="Fig. 2. Saprininae, habitus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272129" httpUri="https://zenodo.org/record/4272129/files/figure.png" pageId="175" pageNumber="168">Figs. 2</figureCitation>
,
<figureCitation id="13173CCDFF3809866302FE8BCCA6FE81" box="[597,652,364,383]" captionStart="Figs" captionStartId="26.[150,189,1085,1104]" captionTargetBox="[202,1075,219,1057]" captionTargetId="figure-156@26.[202,1076,219,1058]" captionTargetPageId="26" captionText="Figs. 2938. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 29 Xenophilothis choumovitchi (Thérond, 1965); 30 Zorius funereus (Schmidt, 1890); 31 Alienocacculus neftensis (Olexa, 1984); 32 Philothis (Philothis) arcanus Reichardt, 1930; 33 Philothis (Atavinus) atavus (Reichardt, 1931); 34 Philothis (Farabius) hexeris Reichardt, 1930; 35 Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 36 Gnathoncus rotundatus (Kugelann, 1792); 37 Eremosaprinus vlasovi (Reichardt, 1941); 38 Microsaprinus therondianus (Dahlgren, 1973)." figureDoi="http://doi.org/10.5281/zenodo.4272141" httpUri="https://zenodo.org/record/4272141/files/figure.png" pageId="175" pageNumber="168">3233</figureCitation>
;
<treatmentCitation id="0A8D0659FF38098663C1FE8BCD14FE81" author="MAZUR S." box="[662,830,364,383]" page="269" pageId="175" pageNumber="168" year="1997">
<bibRefCitation id="EFBD5DB9FF38098663C1FE8BCD39FE81" author="MAZUR S." box="[662,787,364,383]" pageId="175" pageNumber="168" pagination="1 - 373" refId="ref118633" refString="MAZUR S. 1997: A world catalogue of the Histeridae (Coleoptera: Histeridae). Genus, Supplement, pp. 1 - 373." type="book chapter" year="1997">MAZUR (1997)</bibRefCitation>
: 269
</treatmentCitation>
;
<treatmentCitation id="0A8D0659FF3809866211FE8BCDC9FE81" author="MAZUR S." box="[838,995,364,383]" page="96" pageId="175" pageNumber="168" year="2004">
<bibRefCitation id="EFBD5DB9FF3809866211FE8BCDE9FE81" author="MAZUR S." box="[838,963,364,383]" pageId="175" pageNumber="168" pagination="68 - 102" refId="ref118682" refString="MAZUR S. 2004: Family Histeridae. Pp. 68 - 102. In: LOBL I. &amp; SMETANA A. (eds.): Catalogue of Palaearctic Coleoptera, Vol. 2, Hydrophiloidea-Histeroidea-Staphylinidea. Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">MAZUR (2004)</bibRefCitation>
: 96
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF3809866126FE71CECAFDB7" pageId="175" pageNumber="168" type="materials_examined">
<paragraph id="8B932048FF3809866126FE71CF8FFE57" blockId="175.[113,1089,405,585]" box="[113,421,405,425]" pageId="175" pageNumber="168">
<emphasis id="B958FC5AFF3809866126FE71CEC7FE57" bold="true" box="[113,237,405,425]" pageId="175" pageNumber="168">
<typeStatus id="54979EEAFF3809866126FE71CEB5FE57" box="[113,159,406,425]" pageId="175" pageNumber="168">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F33B60D8FF38098661A5FE72CF7EFE56" box="[242,340,405,424]" name="Uzbekistan" pageId="175" pageNumber="168">Uzbekistan</collectingCountry>
, Termez.
</paragraph>
<paragraph id="8B932048FF3809866126FE57CFCEFE07" blockId="175.[113,1089,405,585]" pageId="175" pageNumber="168">
<emphasis id="B958FC5AFF3809866126FE57CF70FE3D" bold="true" box="[113,346,431,451]" pageId="175" pageNumber="168">Type material examined.</emphasis>
<typeStatus id="54979EEAFF3809866035FE56CF9FFE3C" box="[354,437,433,452]" pageId="175" pageNumber="168" type="holotype">HOLOTYPE</typeStatus>
: spec., Termes, Buchara /
<date id="FF920688FF38098663F9FE57CD29FE3D" box="[686,771,432,451]" pageId="175" pageNumber="168" value="1926-05-31">31.v.1926</date>
/ Shestoperov [written] // Termez / 31.v.[19]26 [written] //
<taxonomicName id="4C2C5BCBFF380986606DFE2CCFADFE20" authorityName=": REICHARDT" authorityYear="1941" box="[314,391,459,478]" class="Insecta" family="Histeridae" genus="Philothis" kingdom="Animalia" order="Coleoptera" pageId="175" pageNumber="168" phylum="Arthropoda" rank="genus">
<emphasis id="B958FC5AFF380986606DFE2CCFADFE20" box="[314,391,459,478]" italics="true" pageId="175" pageNumber="168">Philothis</emphasis>
</taxonomicName>
/
<taxonomicName id="4C2C5BCBFF38098660C0FE2CCFE5FE23" authorityName=": OLEXA" authorityYear="1990" box="[407,463,459,477]" class="Insecta" family="Histeridae" genus="Philothis" kingdom="Animalia" order="Coleoptera" pageId="175" pageNumber="168" phylum="Arthropoda" rank="species" species="atavus">
<emphasis id="B958FC5AFF38098660C0FE2CCFE5FE23" box="[407,463,459,477]" italics="true" pageId="175" pageNumber="168">atavus</emphasis>
</taxonomicName>
/ sp. n. / Monotypus / A. Reichardt det [printed-written] //
<typeStatus id="54979EEAFF3809866297FE2CCA33FE20" box="[960,1049,459,478]" pageId="175" pageNumber="168" type="holotype">Holotypus</typeStatus>
[red label, printed] // golden round label (
<collectionCode id="ED3DB88DFF38098660E5FE01CFF0FE07" box="[434,474,486,505]" country="Russia" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="175" pageNumber="168" type="Museum">ZIN</collectionCode>
).
</paragraph>
<paragraph id="8B932048FF3809866126FE18CECAFDB7" blockId="175.[113,1089,405,585]" pageId="175" pageNumber="168">
<emphasis id="B958FC5AFF3809866126FE18CFA9FDED" bold="true" box="[113,387,511,531]" pageId="175" pageNumber="168">Additional material examined.</emphasis>
<materialsCitation id="3B442A15FF38098660D1FE18CD82FDED" ID-GBIF-Occurrence="2981920381" box="[390,936,511,532]" collectingDate="1980-04-16" collectorName="A. Olexa" country="Turkmenistan" location="Ashgabat" pageId="175" pageNumber="168" specimenCount="1" stateProvince="Ashgabat">
<emphasis id="B958FC5AFF38098660D1FE18CC17FDED" bold="true" box="[390,573,511,531]" pageId="175" pageNumber="168">
<collectingCountry id="F33B60D8FF38098660D1FE18CC13FDED" box="[390,569,511,531]" name="Turkmenistan" pageId="175" pageNumber="168">TURKMENISTAN</collectingCountry>
:
</emphasis>
<collectingRegion id="49E8EEAAFF3809866316FDE7CCB9FDED" box="[577,659,512,531]" country="Turkmenistan" name="Ashgabat" pageId="175" pageNumber="168">Ashgabat</collectingRegion>
,
<date id="FF920688FF38098663CDFDE7CCDCFDED" box="[666,758,512,531]" pageId="175" pageNumber="168" value="1980-04-16">
<collectingDate id="EFD6FF60FF38098663CDFDE7CCDCFDED" box="[666,758,512,531]" pageId="175" pageNumber="168" value="1980-04-16">16.iv.1980</collectingDate>
</date>
,
<specimenCount id="9D2AEBC1FF38098663AAFDE7CD10FDEA" box="[765,826,512,532]" pageId="175" pageNumber="168" type="generic">1 spec.</specimenCount>
,
<collectorName id="26D9459EFF3809866217FDE7CDA1FDED" box="[832,907,512,531]" pageId="175" pageNumber="168">A.Olexa</collectorName>
lgt
</materialsCitation>
.
<materialsCitation id="3B442A15FF38098662F9FE18CC10FDD0" ID-GBIF-Occurrence="2981920329" collectingDate="1978-04-28" collectorName="A. Olexa" country="Uzbekistan" location="Kyzylkum" municipality="Buchara" pageId="175" pageNumber="168" specimenCount="1">
<emphasis id="B958FC5AFF38098662F9FE18CA6BFDED" bold="true" box="[942,1089,511,531]" pageId="175" pageNumber="168">
<collectingCountry id="F33B60D8FF38098662F9FE18CA16FDED" box="[942,1084,511,531]" name="Uzbekistan" pageId="175" pageNumber="168">UZBEKISTAN</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6BF7BA32FF3809866126FDFCCE90FDD0" box="[113,186,539,558]" pageId="175" pageNumber="168">Buchara</collectingMunicipality>
,
<location id="8EF37693FF3809866193FDFCCF0AFDD0" LSID="urn:lsid:plazi:treatment:0385915EFF38099A600AFF3DCF34FCCD:8EF37693FF3809866193FDFCCF0AFDD0" box="[196,288,539,558]" country="Uzbekistan" municipality="Buchara" name="Kyzylkum" pageId="175" pageNumber="168">Kyzylkum</location>
,
<date id="FF920688FF380986607EFDFCCFACFDD0" box="[297,390,539,558]" pageId="175" pageNumber="168" value="1978-04-28">
<collectingDate id="EFD6FF60FF380986607EFDFCCFACFDD0" box="[297,390,539,558]" pageId="175" pageNumber="168" value="1978-04-28">28.iv.1978</collectingDate>
</date>
, 2 ♁♁,
<collectorName id="26D9459EFF380986609CFDFCCC33FDD0" box="[459,537,539,558]" pageId="175" pageNumber="168">A. Olexa</collectorName>
lgt
</materialsCitation>
.;
<materialsCitation id="3B442A15FF3809866310FDFCCEF6FDB7" ID-GBIF-Occurrence="2981920310" collectingDate="1979-05-01" collectingDateMax="1979-05-05" collectingDateMin="1979-05-01" collectionCode="TLAN" collectorName="A. Olexa" country="Uzbekistan" location="Karakum" municipality="Chiva" pageId="175" pageNumber="168" specimenCount="1">
<collectingMunicipality id="6BF7BA32FF3809866310FDFCCC50FDD0" box="[583,634,539,558]" pageId="175" pageNumber="168">Chiva</collectingMunicipality>
[= Khiva],
<location id="8EF37693FF38098663B7FDFCCD18FDD0" LSID="urn:lsid:plazi:treatment:0385915EFF38099A600AFF3DCF34FCCD:8EF37693FF38098663B7FDFCCD18FDD0" box="[736,818,539,558]" country="Uzbekistan" municipality="Chiva" name="Karakum" pageId="175" pageNumber="168">Karakum</location>
,
<date id="FF920688FF380986626CFDFCCD88FDD0" box="[827,930,539,558]" pageId="175" pageNumber="168" value="1979-05-01" valueMax="1979-05-05" valueMin="1979-05-01">
<collectingDate id="EFD6FF60FF380986626CFDFCCD88FDD0" box="[827,930,539,558]" pageId="175" pageNumber="168" value="1979-05-01" valueMax="1979-05-05" valueMin="1979-05-01">1.5.v.1979</collectingDate>
</date>
,
<specimenCount id="9D2AEBC1FF38098662FCFDFCCDC0FDD0" box="[939,1002,539,558]" pageId="175" pageNumber="168" type="generic">1 spec.</specimenCount>
,
<collectorName id="26D9459EFF38098662A4FDFCCA6BFDD0" box="[1011,1089,539,558]" pageId="175" pageNumber="168">A. Olexa</collectorName>
lgt. (
<collectionCode id="ED3DB88DFF38098661CEFDD1CEFDFDB7" box="[153,215,566,585]" pageId="175" pageNumber="168">TLAN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33673C3FF38099A6126FD86CF34FCCD" lastPageId="179" lastPageNumber="172" pageId="175" pageNumber="168" type="description">
<paragraph id="8B932048FF3809866126FD86CC5FFD66" blockId="175.[113,1089,609,1593]" pageId="175" pageNumber="168">
<emphasis id="B958FC5AFF3809866126FD86CF3DFD87" bold="true" box="[113,279,609,633]" pageId="175" pageNumber="168">Redescription.</emphasis>
Body length: PEL:
<quantity id="4CD48DADFF38098660A7FD86CCB7FD86" box="[496,669,609,633]" metricMagnitude="-3" metricUnit="m" metricValue="2.0625" metricValueMax="2.25" metricValueMin="1.875" pageId="175" pageNumber="168" unit="mm" value="2.0625" valueMax="2.25" valueMin="1.875">1.8752.25 mm</quantity>
; APW: 0.751.0 mm; PPW:
<quantity id="4CD48DADFF3809866281FD86CEB4FD66" metricMagnitude="-3" metricUnit="m" metricValue="1.7875" metricValueMax="1.875" metricValueMin="1.7" pageId="175" pageNumber="168" unit="mm" value="1.7875" valueMax="1.875" valueMin="1.7">1.71.875 mm</quantity>
; EL:
<quantity id="4CD48DADFF380986618FFD66CF5CFD66" box="[216,374,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="1.4375" metricValueMax="1.5" metricValueMin="1.375" pageId="175" pageNumber="168" unit="mm" value="1.4375" valueMax="1.5" valueMin="1.375">1.3751.5 mm</quantity>
; EW:
<quantity id="4CD48DADFF38098660EFFD66CC5BFD66" box="[440,625,641,665]" metricMagnitude="-3" metricUnit="m" metricValue="1.975" metricValueMax="2.075" metricValueMin="1.875" pageId="175" pageNumber="168" unit="mm" value="1.975" valueMax="2.075" valueMin="1.875">
<geoCoordinate id="EE18468FFF38098660EFFD66CFD8FD67" box="[440,498,641,665]" degrees="1.875" direction="north" orientation="latitude" pageId="175" pageNumber="168" precision="55" value="1.875">1.875</geoCoordinate>
<geoCoordinate id="EE18468FFF38098660A4FD66CC14FD67" box="[499,574,641,665]" degrees="2.075" direction="west" orientation="longitude" pageId="175" pageNumber="168" precision="55" value="-2.075">2.075</geoCoordinate>
mm
</quantity>
.
</paragraph>
<paragraph id="8B932048FF38098661C7FD46CF82FD07" blockId="175.[113,1089,609,1593]" pageId="175" pageNumber="168">
Body (
<figureCitation id="13173CCDFF380986618FFD46CF59FD47" box="[216,371,673,697]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">Figs. 543544</figureCitation>
) roundly oval, moderately convex from above, underside very convex, cuticle dark brown with conspicuous greenish-bronze metallic luster; legs, mouthparts rufous, antennal club yellow.
</paragraph>
<paragraph id="8B932048FF38098661C7FCE6CCE3FC27" blockId="175.[113,1089,609,1593]" pageId="175" pageNumber="168">
Antennal scape (
<figureCitation id="13173CCDFF3809866013FCE6CF89FCE7" box="[324,419,769,793]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">Fig. 546</figureCitation>
), with numerous long setae; 8
<superScript id="7C598D00FF38098663B0FD18CCD9FCF0" attach="left" box="[743,755,767,782]" fontSize="6" pageId="175" pageNumber="168">th</superScript>
segment (
<figureCitation id="13173CCDFF3809866234FCE6CD9AFCE7" box="[867,944,769,793]" captionStart="Figs" captionStartId="22.[151,190,757,776]" captionTargetBox="[151,1126,216,722]" captionTargetId="figure-310@22.[151,1126,213,724]" captionTargetPageId="22" captionText="Figs. 57. Saprininae, 8th antennomere: 5 Paravolvulus ovillum (Solskij, 1876); 6 Philothis (Atavinus) atavus (Reichardt, 1931); 7 Turanostyphrus ignoratus Tishechkin, 2005." figureDoi="http://doi.org/10.5281/zenodo.4272135" httpUri="https://zenodo.org/record/4272135/files/figure.png" pageId="175" pageNumber="168">Figs. 6</figureCitation>
,
<figureCitation id="13173CCDFF38098662EBFCE6CDCFFCE7" box="[956,997,769,793]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">545</figureCitation>
) cupuliform, surrounding basal half of the club; club (
<figureCitation id="13173CCDFF380986630EFCC6CC98FCC7" box="[601,690,801,825]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">Fig. 545</figureCitation>
) comparatively small, without visible articulation, apical third (roughly) imbricate with dense curved short sensilla intermingled with somewhat sparser straight thicker sensilla, concealing two round apical sensory areas; basal two-thirds (roughly) glabrous; sensory structures of antennal club (
<figureCitation id="13173CCDFF38098662D9FC66CDF4FC67" box="[910,990,897,921]" captionStart="Figs" captionStartId="26.[150,189,1085,1104]" captionTargetBox="[202,1075,219,1057]" captionTargetId="figure-156@26.[202,1076,219,1058]" captionTargetPageId="26" captionText="Figs. 2938. Saprininae, sensory structures of the antennal club (vesicles marked with grey; sensory areas shaded): 29 Xenophilothis choumovitchi (Thérond, 1965); 30 Zorius funereus (Schmidt, 1890); 31 Alienocacculus neftensis (Olexa, 1984); 32 Philothis (Philothis) arcanus Reichardt, 1930; 33 Philothis (Atavinus) atavus (Reichardt, 1931); 34 Philothis (Farabius) hexeris Reichardt, 1930; 35 Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 36 Gnathoncus rotundatus (Kugelann, 1792); 37 Eremosaprinus vlasovi (Reichardt, 1941); 38 Microsaprinus therondianus (Dahlgren, 1973)." figureDoi="http://doi.org/10.5281/zenodo.4272141" httpUri="https://zenodo.org/record/4272141/files/figure.png" pageId="175" pageNumber="168">Fig. 33</figureCitation>
) in form of one apical sensory area and one stipe-shaped vesicle situated under it, supplemented by another vague sensory area under apical surface of club.
</paragraph>
<paragraph id="8B932048FF38098661C7FC06CF8DFAC7" blockId="175.[113,1089,609,1593]" pageId="175" pageNumber="168">
Mouthparts. Mandibles (
<figureCitation id="13173CCDFF38098660C4FC06CFF5FC07" box="[403,479,993,1017]" captionStart="Figs" captionStartId="35.[114,153,1183,1202]" captionTargetBox="[113,1088,219,1152]" captionTargetId="figure-95@35.[113,1088,219,1153]" captionTargetPageId="35" captionText="Figs. 9098. Saprininae, mandibles, ventral view: 90 Gnathoncus rotundatus (Kugelann, 1792); 91 Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 92 Hypocacculus (Hypocacculus) metalescens (Erichson, 1834); 93 Hypocacculus (Colpellus) praecox (Erichson, 1834); 94 Hypocacculus (Nessus) rubripes (Erichson, 1834); 95 Myrmetes paykulli Kanaar, 1979; 96 Paravolvulus lateristrius (Solskij, 1876); 97 Philothis (Philothis) arcanus Reichardt, 1930; 98 Philothis (Atavinus) atavus (Reichardt, 1931)." figureDoi="http://doi.org/10.5281/zenodo.4272153" httpUri="https://zenodo.org/record/4272153/files/figure.png" pageId="175" pageNumber="168">Fig. 98</figureCitation>
) with straight outer margin strongly curved inwards; mandibular apex acute; sub-apical tooth on left mandible moderately large, almost perpendicular; disc of labrum (
<figureCitation id="13173CCDFF3809866075FBC6CF58FBC7" box="[290,370,1057,1081]" captionStart="Figs" captionStartId="31.[122,161,1196,1215]" captionTargetId="figure-11@32.[151,1129,219,1531]" captionTargetPageId="32" captionText="Figs. 4773. Saprininae, labral structures, dorsal view (left half showing external and right half showing internal structures): 47 Alienocacculus neftensis (Olexa, 1984); 48 Ammostyphrus cerberus Reichardt, 1924; 49 Chalcionellus amoenus (Erichson, 1834); 50 Chivaenius kryzhanovskii Olexa, 1980; 51 Eopachylopus ripae (Lewis, 1885); 52 Eremosaprinus vlasovi (Reichardt, 1941); 53 Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 54 Exaesiopus grossipes grossipes (Marseul, 55); 55 Gnathoncus rotundatus (Kugelann, 1792); 56 Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 57 Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 58 Hypocacculus (Colpellus) praecox (Erichson, 1834); 59 Hypocacculus (Nessus) rubripes (Erichson, 1834); 60 Microsaprinus therondianus (Dahlgren, 1973); 61 Myrmetes paykulli Kanaar, 1979; 62 Paravolvulus lateristriatus (Solskij, 1876); 63 Philothis (Philothis) arcanus Reichardt, 1930; 64 Philothis (Atavinus) atavus (Reichardt, 1931); 65 Philothis (Farabius) hexeris Reichardt, 1930; 66 Pholioxenus phoenix (Reichardt, 1930); 67 Reichardtiolus duriculus (Reitter, 1904); 68 Saprinus (Saprinus) semistriatus (Scriba, 1790); 69 Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 70 Saprinus (Phaonius) pharao (Marseul, 1855); 71 Styphrus corpulentus Motschulsky, 1845; 72 Xenonychus tridens (Jacquelin-Duval, 1852); 73 Xenophilothis choumovitchi (Thérond, 1965)." figureDoi="http://doi.org/10.5281/zenodo.4288743" httpUri="https://zenodo.org/record/4288743/files/figure.png" pageId="175" pageNumber="168">Fig. 64</figureCitation>
) slightly emarginate medially, almost smooth; epipharynx almost completely hidden under labral fold (
<figureCitation id="13173CCDFF38098660AEFBA6CC62FBA7" box="[505,584,1089,1113]" captionStart="Figs" captionStartId="30.[152,191,1491,1510]" captionTargetBox="[229,1051,894,1466]" captionTargetId="figure-295@30.[229,1051,894,1510]" captionTargetPageId="30" captionText="Figs. 3944. Saprininae, labrum, lateral view: 39 Philothis (Farabius) hexeris Reichardt, 1930; 40 Philothis (Atavinus) atavus (Reichardt, 1931); 41 Saprinus (Phaonius) pharao (Marseul, 1855); 42 Eremosaprinus vlasovi (Reichardt, 1941); 43 Ctenophilothis chobauti (Théry, 1900); 44 Philothis (Philothis) arcanus Reichardt, 1930." figureDoi="http://doi.org/10.5281/zenodo.4272143" httpUri="https://zenodo.org/record/4272143/files/figure.png" pageId="175" pageNumber="168">Fig. 40</figureCitation>
); terminal labial palpomere elongated, its width about one-third its length; mentum (
<figureCitation id="13173CCDFF38098660B1FB86CC15FB87" box="[486,575,1121,1145]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">Fig. 548</figureCitation>
) sub-trapezoid, finely imbricate with tiny median notch (
<figureCitation id="13173CCDFF38098661EDFB66CF3DFB67" box="[186,279,1153,1177]" captionStart="Figs" captionStartId="38.[151,190,1224,1243]" captionTargetBox="[229,1046,222,1195]" captionTargetId="figure-11@38.[229,1046,219,1196]" captionTargetPageId="38" captionText="Figs. 109141. Saprininae, anterior margin of mentum, ventral view: 109 Alienocacculus neftensis (Olexa, 1984); 110 Ammostyphrus cerberus Reichardt, 1924; 111 Axelinus ghilarovi Kryzhanovskij, 1976; 112 Chalcionellus amoenus (Erichson, 1834); 113 Chivaenius kryzhanovskii Olexa, 1980; 114 Ctenophilotis chobauti (Théry, 1900); 115 Eopachylopus ripae (Lewis, 1885); 116 Eremosaprinus vlasovi (Reichardt, 1941); 117 Euspilotus (Neosaprinus) perrisi (Marseul, 1872); 118 Exaesiopus grossipes grossipes (Marseul, 1855); 119 Gnathoncus rotundatus (Kugelann, 1792); 120 Hypocaccus (Hypocaccus) rugiceps (Duftschmid, 1805); 121 Hypocaccus (Baeckmanniolus) dimidiatus dimidiatus (Illiger, 1807); 122 Hypocacculus (Hypocacculus) metallescens (Erichson, 1834); 123 Hypocacculus (Colpellus) praecox (Erichson, 1834); 124 Hypocacculus (Nessus) rubripes (Erichson, 1834); 125 Microsaprinus therondianus (Dahlgren, 1973); 126 Myrmetes paykulli Kanaar, 1979; 127 Paravolvulus ovillum (Solskij, 1876); 128 Philothis (Philothis) arcanus Reichardt, 1930; 129 Philothis (Atavinus) atavus (Reichardt, 1931); 130 Philothis (Farabius) hexeris Reichardt, 1930; 131 Pholioxenus phoenix (Reichardt, 1930); 132 Reichardtiolus duriculus (Reitter, 1904); 133 Saprinillus paromaloides Kryzhanovskij, 1974; 134 Saprinus (Saprinus) semistriatus (Scriba, 1790); 135 Saprinus (Hemisaprinus) subvirescens (Ménétries, 1832); 136 Saprinus (Phaonius) pharao (Marseul, 1855); 137 Styphrus corpulentus Motschulsky, 1845; 138 Turanostyphrus ignoratus Tishechkin, 2005; 139 Xenonychus tridens (Jacquelin-Duval, 1852); 140 Xenophilotis choumovitchi (Thérond, 1965); 141 Zorius funereus (Schmidt, 1890)." figureDoi="http://doi.org/10.5281/zenodo.4272160" httpUri="https://zenodo.org/record/4272160/files/figure.png" pageId="175" pageNumber="168">Fig. 129</figureCitation>
); anterior margin with dense brush of long ramose setae, lateral margins with a row of slightly shorter setae, disc of mentum with several ramose setae, approximately as long as those of anterior margin; cardo of maxilla (
<figureCitation id="13173CCDFF38098663D3FB26CCF7FB27" box="[644,733,1217,1241]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">Fig. 548</figureCitation>
) on lateral margin with numerous short ramose setae; stipes (
<figureCitation id="13173CCDFF38098660C2FB06CFDBFB07" box="[405,497,1249,1273]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">Fig. 548</figureCitation>
) triangular, with numerous much longer ramose setae; terminal maxillary palpomere elongated, its width about one-third its length, approximately twice as long as penultimate.
</paragraph>
<paragraph id="8B932048FF38098661C7FAA6CED9FA47" blockId="175.[113,1089,609,1593]" pageId="175" pageNumber="168">
Clypeus (
<figureCitation id="13173CCDFF38098661A2FAA6CF7AFAA7" box="[245,336,1345,1369]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">Fig. 546</figureCitation>
) rectangular, rounded laterally, smooth; frontal stria complete, slightly weakened medially, continued as complete and carinate supraorbital stria; frontal disc (
<figureCitation id="13173CCDFF3809866541FA86CEB0FA67" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">Fig. 546</figureCitation>
) almost smooth, only with scattered microscopic punctation; eyes very flattened, invisible from above.
</paragraph>
<paragraph id="8B932048FF38099861C7FA26CD37FA6C" blockId="175.[113,1089,609,1593]" lastBlockId="177.[113,1089,1338,1586]" lastPageId="177" lastPageNumber="170" pageId="175" pageNumber="168">
Pronotal sides (
<figureCitation id="13173CCDFF3809866064FA26CFA7FA27" box="[307,397,1473,1497]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="175" pageNumber="168">Fig. 543</figureCitation>
) strongly convergent anteriorly, apical angles acute, anterior pronotal margin with a shallow incision for head, straight almost medially; marginal pronotal stria well impressed, carinate laterally, broadly interrupted behind head; pronotal disc moderately convex, with irregular wrinkles and curved lines becoming coarser laterally, wrinkles intermingled with shallow scattered punctures; lateral and anterior margins of pronotum with narrow smooth band; pronotal base with a row of dense elongate punctures; pronotal hypomeron with long yellow setae; scutellum small, but visible.
</paragraph>
<caption id="DF5370C0FF27099961C0FA28CD7AF9CC" ID-DOI="http://doi.org/10.5281/zenodo.4272270" ID-Zenodo-Dep="4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="176" pageNumber="173" startId="176.[151,191,1487,1506]" targetBox="[245,1034,219,1471]" targetPageId="176">
<paragraph id="8B932048FF27099961C0FA28CD7AF9CC" blockId="176.[151,1129,1487,1587]" pageId="176" pageNumber="173">
Figs. 543551.
<taxonomicName id="4C2C5BCBFF2709996074FA36CC90FA1D" authority="(Reichardt, 1931)" baseAuthorityName="Reichardt" baseAuthorityYear="1931" box="[291,698,1487,1507]" class="Insecta" family="Histeridae" genus="Philothis" kingdom="Animalia" order="Coleoptera" pageId="176" pageNumber="173" phylum="Arthropoda" rank="species" species="atavus" subGenus="Atavinus">
<emphasis id="B958FC5AFF2709996074FA36CF5BFA1D" box="[291,369,1489,1507]" italics="true" pageId="176" pageNumber="173">Philothis</emphasis>
(
<emphasis id="B958FC5AFF27099960D6FA36CFE7FA1D" box="[385,461,1489,1507]" italics="true" pageId="176" pageNumber="173">Atavinus</emphasis>
)
<emphasis id="B958FC5AFF270999608BFA36CC3EFA1D" box="[476,532,1489,1507]" italics="true" pageId="176" pageNumber="173">atavus</emphasis>
(
<bibRefCitation id="EFBD5DB9FF2709996375FA28CC99FA1D" author="REICHARDT A." box="[546,691,1487,1507]" pageId="176" pageNumber="173" pagination="308 - 311" refId="ref120322" refString="REICHARDT A. 1931: Sur une nouveau representant du genre Philothis Rchdt. (Coleoptera: Histeridae). Doklady Akademii Nauk SSSR A: 308 - 311." type="book chapter" year="1931">Reichardt, 1931</bibRefCitation>
)
</taxonomicName>
, SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view.
</paragraph>
</caption>
<caption id="DF5370C0FF2609986124FB4ACF52FAEE" ID-DOI="http://doi.org/10.5281/zenodo.4272272" ID-Zenodo-Dep="4272272" httpUri="https://zenodo.org/record/4272272/files/figure.png" pageId="177" pageNumber="170" startId="177.[115,154,1197,1216]" targetBox="[256,940,223,1169]" targetPageId="177">
<paragraph id="8B932048FF2609986124FB4ACF52FAEE" blockId="177.[113,1087,1196,1297]" pageId="177" pageNumber="170">
Figs. 552558.
<taxonomicName id="4C2C5BCBFF26099861A2FB49CC5DFB3E" authority="(Reichardt, 1931)" baseAuthorityName="Reichardt" baseAuthorityYear="1931" box="[245,631,1197,1216]" class="Insecta" family="Histeridae" genus="Philothis" kingdom="Animalia" order="Coleoptera" pageId="177" pageNumber="170" phylum="Arthropoda" rank="species" species="atavus" subGenus="Atavinus">
<emphasis id="B958FC5AFF26099861A2FB49CF68FB3E" box="[245,322,1198,1216]" italics="true" pageId="177" pageNumber="170">Philothis</emphasis>
(
<emphasis id="B958FC5AFF2609986019FB49CFB3FB3E" box="[334,409,1198,1216]" italics="true" pageId="177" pageNumber="170">Atavinus</emphasis>
)
<emphasis id="B958FC5AFF26099860F3FB49CFF6FB3E" box="[420,476,1198,1216]" italics="true" pageId="177" pageNumber="170">atavus</emphasis>
(
<bibRefCitation id="EFBD5DB9FF26099860B1FB4ACC5BFB3E" author="REICHARDT A." box="[486,625,1197,1216]" pageId="177" pageNumber="170" pagination="308 - 311" refId="ref120322" refString="REICHARDT A. 1931: Sur une nouveau representant du genre Philothis Rchdt. (Coleoptera: Histeridae). Doklady Akademii Nauk SSSR A: 308 - 311." type="book chapter" year="1931">Reichardt, 1931</bibRefCitation>
)
</taxonomicName>
, male terminalia: 552 8
<superScript id="7C598D00FF2609986219FB4BCD72FB46" attach="left" box="[846,856,1196,1208]" fontSize="5" pageId="177" pageNumber="170">th</superScript>
sternite and tergite, ventral view; 553 ditto, dorsal view; 554 ditto, lateral view; 555 9
<superScript id="7C598D00FF26099863C8FB21CC83FB2C" attach="left" box="[671,681,1222,1234]" fontSize="5" pageId="177" pageNumber="170">th</superScript>
tergite, 10
<superScript id="7C598D00FF2609986251FB21CD3AFB2C" attach="left" box="[774,784,1222,1234]" fontSize="5" pageId="177" pageNumber="170">th</superScript>
tergite (dorsal view) and spiculum gastrale (ventral view); 556 9
<superScript id="7C598D00FF26099860D2FB06CFA5FB13" attach="left" box="[389,399,1249,1261]" fontSize="5" pageId="177" pageNumber="170">th</superScript>
tergite, 10
<superScript id="7C598D00FF26099860BAFB06CFDDFB13" attach="left" box="[493,503,1249,1261]" fontSize="5" pageId="177" pageNumber="170">th</superScript>
tergite and spiculum gastrale, lateral view; 557 aedeagus, dorsal view; 558 ditto, lateral view.
</paragraph>
</caption>
<paragraph id="8B932048FF26099B61C7FA7DCF8EFDED" blockId="177.[113,1089,1338,1586]" lastBlockId="178.[151,1127,218,1587]" lastPageId="178" lastPageNumber="175" pageId="177" pageNumber="170">
Elytral humeri weakly developed, glabrous, elytral epipleura with sparse fine punctures, almost smooth; marginal epipleural stria complete, thin; marginal elytral stria well impressed, slightly carinate on outer side, continued as complete apical elytral stria. Humeral elytral stria finely impressed on basal third, occasionally doubled; inner subhumeral stria short, present medially, sometimes basally joining humeral elytral stria; elytral base without bulge; disc with five developed dorsal elytral striae 15, striae (except for the 5
<superScript id="7C598D00FF25099B6232FF3DCD5BFF17" attach="left" box="[869,881,218,233]" fontSize="6" pageId="178" pageNumber="175">th</superScript>
stria) in shallow round punctures; first dorsal elytral stria the longest, reaching about three-fourths of elytral length apically, striae 25 about the same length, reaching about elytral half apically; fourth dorsal elytral stria basally connected with complete sutural elytral stria, which is apically connected with the apical elytral stria and on basal half somewhat distanced from elytral suture. Elytral disc with dense round, shallow punctation, punctures separated by about their own to twice their diameter, becoming confluent apically, punctation becomes sparser anteriorly and laterally, elytral flanks almost smooth; interspace between sutural elytral stria and elytral suture impunctate; a complementary thin stria runs along elytral suture, reaching about half of elytral length apically.
</paragraph>
<paragraph id="8B932048FF25099B61E2FDFCCCFEFD6D" blockId="178.[151,1127,218,1587]" pageId="178" pageNumber="175">Propygidium completely exposed, about half as long as broad, with dense, large, confluent shallow punctures becoming sparser and smaller laterally; pygidium slightly convex, about as long as broad, medially with confluent shallow punctation, similar to that of propygidium, punctures become much sparser and smaller laterally.</paragraph>
<paragraph id="8B932048FF25099B61E2FD7CCCA6FD0D" blockId="178.[151,1127,218,1587]" pageId="178" pageNumber="175">
Anterior margin of prosternum (
<figureCitation id="13173CCDFF25099B6374FD7CCCAEFD4D" box="[547,644,667,691]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="178" pageNumber="175">Fig. 549</figureCitation>
) obtuse-angulate; marginal prosternal stria anteriorly absent; prosternal process substrigulate, densely setose, gradually sloping down anteriorly; both sets of prosternal striae absent.
</paragraph>
<paragraph id="8B932048FF25099B61E2FD1CCD93FC0D" blockId="178.[151,1127,218,1587]" pageId="178" pageNumber="175">
Anterior marginal stria of mesoventrite slightly emarginate medially, discal marginal mesoventral stria complete, carinate, slightly weakened medially; disc of mesoventrite with scattered shallow punctures; meso-metaventral sutural stria well impressed, carinate, undulate; intercoxal disc of metaventrite convex, with scattered microscopic punctures, along posterior margin a band of coarser and denser punctation appears; lateral metaventral stria (
<figureCitation id="13173CCDFF25099B6553FC9CCA4AFC6D" box="[1028,1120,891,915]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="178" pageNumber="175">Fig. 550</figureCitation>
) shortened, carinate, almost straight; lateral disc of metaventrite concave, with dense round moderately deep punctures, fringed with long setae; metepisternum + fused metepimeron (
<figureCitation id="13173CCDFF25099B61C8FC3CCED1FC0D" box="[159,251,987,1011]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="178" pageNumber="175">Fig. 550</figureCitation>
) very setose, punctation almost unrecognizable beneath the setae.
</paragraph>
<paragraph id="8B932048FF25099B61E2FC1CCCF8FBCD" blockId="178.[151,1127,218,1587]" pageId="178" pageNumber="175">Intercoxal disc of first abdominal sternite completely striate laterally; disc smooth; lateral disc of all visible abdominal sternites setose laterally.</paragraph>
<paragraph id="8B932048FF25099B61E2FBDCCC31FA8D" blockId="178.[151,1127,218,1587]" pageId="178" pageNumber="175">
Protibia (
<figureCitation id="13173CCDFF25099B6041FBDCCF45FBAD" box="[278,367,1083,1107]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="178" pageNumber="175">Fig. 551</figureCitation>
) moderately dilated, anterior margin truncate, formed by anterior margin of distal-most large triangular tooth; outer margin of protibia apart from this tooth with another, somewhat shorter triangular tooth, both teeth topped with short rounded blunt denticle; setae of outer row confined to basal third, thick, sparse and moderately long; setae of median row much shorter than those of outer row but more regular; protarsal groove inconspicuous; apical margin of protibia posteriorly without denticles; outer part of posterior surface of protibia with carinate rugae, distinctly separated from smooth median part of posterior surface by costiform stria; posterior protibial stria shortened apically; inner margin of protibia with long lamelliform dense setae; protibial spur rudimentary, almost invisible, entombed in apical margin of protibia; protarsus absent.
</paragraph>
<paragraph id="8B932048FF25099A61E2FA9CCD43FDCD" blockId="178.[151,1127,218,1587]" lastBlockId="179.[113,1089,219,819]" lastPageId="179" lastPageNumber="172" pageId="178" pageNumber="175">Mesotibia somewhat dilated, not particularly thickened, outer margin with one double row of sparse, morphologically different short denticles significantly growing in size and density near tarsal insertion, denticles continuous ventrally along apical margin; setae of outer row moderately long, dense, thickened; setae of median row dense and long, covering almost entire posterior mesotibial surface; posterior mesotibial stria shortened, present only on basal half; mesotibial spur conspicuously long and stout; outer part of median surface of mesotibia almost smooth, only with scattered microscopic punctation; anterior mesotibial stria carinate, almost complete; inner margin of mesotibia with moderately dense row of long lamelliform setae; mesotarsus telescope-like, diameter of tarsomeres diminishing apically, each tarsomere with two long lamelliform strongly sclerotized setae, one posteriorly and one anteriorly; claws of apical mesotarsomeres thin, hair-like, several times as long as mesotarsomeres itself. Metatibia conspicuously triangularly dilated, outer margin with double row (short, blunt, dense and sparser, much longer denticles) of morphologically different denticles, another row of short sparser round denticles markedly shifted from it, present on anterior surface of metatibia and only observable from ventral view. Posterior surface of metatibia with dense rows of short setae, distinction between outer and median row unclear; posterior metatibial stria inconspicuous (absent?); metatibia otherwise similar to mesotibia.</paragraph>
<paragraph id="8B932048FF24099A61C7FDDCCF34FCCD" blockId="179.[113,1089,219,819]" pageId="179" pageNumber="172">
Male genitalia. Eighth sternite (
<figureCitation id="13173CCDFF24099A60B5FDDCCC57FDAD" box="[482,637,571,595]" captionStart="Figs" captionStartId="177.[115,154,1197,1216]" captionTargetBox="[256,940,223,1169]" captionTargetId="figure-107@177.[224,977,219,1184]" captionTargetPageId="177" captionText="Figs. 552558.Philothis (Atavinus) atavus (Reichardt, 1931), male terminalia: 552 8th sternite and tergite, ventral view; 553 ditto, dorsal view; 554 ditto, lateral view; 555 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 556 9th tergite, 10th tergite and spiculum gastrale, lateral view; 557 aedeagus, dorsal view; 558 ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272272" httpUri="https://zenodo.org/record/4272272/files/figure.png" pageId="179" pageNumber="172">Figs. 552553</figureCitation>
) conspicuously narrowing apically, longitudinally fused medially, apically with tiny inflatable membrane (velum); fringed with few short setae; eighth tergite and eighth sternite fused laterally (
<figureCitation id="13173CCDFF24099A6250FD9CCD4DFD6D" box="[775,871,635,659]" captionStart="Figs" captionStartId="176.[151,191,1487,1506]" captionTargetBox="[245,1034,219,1471]" captionTargetId="figure-11@176.[245,1034,219,1482]" captionTargetPageId="176" captionText="Figs. 543551. Philothis (Atavinus) atavus (Reichardt, 1931), SEM micrographs: 543 habitus, dorsal view; 544 ditto, ventral view; 545 antennal club, ventral view; 546 head, dorsal view; 547 ditto, ventral view; 548 mentum, submentum, cardines and stipites of maxilla, ventral view; 549 prosternum; 550 lateral disc of metaventrite, metepisternum and fused metepimeron; 551 protibia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4272270" httpUri="https://zenodo.org/record/4272270/files/figure.png" pageId="179" pageNumber="172">Fig. 546</figureCitation>
). Morphology of 9
<superScript id="7C598D00FF24099A6563FD9DCA6AFD77" attach="none" box="[1076,1088,634,649]" fontSize="6" pageId="179" pageNumber="172">th</superScript>
tergite (
<figureCitation id="13173CCDFF24099A6193FD7CCF4AFD4D" box="[196,352,667,691]" captionStart="Figs" captionStartId="177.[115,154,1197,1216]" captionTargetBox="[256,940,223,1169]" captionTargetId="figure-107@177.[224,977,219,1184]" captionTargetPageId="177" captionText="Figs. 552558.Philothis (Atavinus) atavus (Reichardt, 1931), male terminalia: 552 8th sternite and tergite, ventral view; 553 ditto, dorsal view; 554 ditto, lateral view; 555 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 556 9th tergite, 10th tergite and spiculum gastrale, lateral view; 557 aedeagus, dorsal view; 558 ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272272" httpUri="https://zenodo.org/record/4272272/files/figure.png" pageId="179" pageNumber="172">Figs. 555556</figureCitation>
) typical for the subfamily; spiculum gastrale (
<figureCitation id="13173CCDFF24099A6218FD7CCD80FD4D" box="[847,938,667,691]" captionStart="Figs" captionStartId="177.[115,154,1197,1216]" captionTargetBox="[256,940,223,1169]" captionTargetId="figure-107@177.[224,977,219,1184]" captionTargetPageId="177" captionText="Figs. 552558.Philothis (Atavinus) atavus (Reichardt, 1931), male terminalia: 552 8th sternite and tergite, ventral view; 553 ditto, dorsal view; 554 ditto, lateral view; 555 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 556 9th tergite, 10th tergite and spiculum gastrale, lateral view; 557 aedeagus, dorsal view; 558 ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272272" httpUri="https://zenodo.org/record/4272272/files/figure.png" pageId="179" pageNumber="172">Fig. 555</figureCitation>
) expanded on both ends; tenth tergite conspicuously small. Basal piece of aedeagus (
<figureCitation id="13173CCDFF24099A620CFD5CCDDFFD2D" box="[859,1013,699,723]" captionStart="Figs" captionStartId="177.[115,154,1197,1216]" captionTargetBox="[256,940,223,1169]" captionTargetId="figure-107@177.[224,977,219,1184]" captionTargetPageId="177" captionText="Figs. 552558.Philothis (Atavinus) atavus (Reichardt, 1931), male terminalia: 552 8th sternite and tergite, ventral view; 553 ditto, dorsal view; 554 ditto, lateral view; 555 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 556 9th tergite, 10th tergite and spiculum gastrale, lateral view; 557 aedeagus, dorsal view; 558 ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272272" httpUri="https://zenodo.org/record/4272272/files/figure.png" pageId="179" pageNumber="172">Figs. 557558</figureCitation>
) rather short, ratio of its length: length of parameres ca 1: 5; aedeagus slender along its entire length; parameres fused along their basal four-fifths; aedeagus gently sinuate from lateral view (
<figureCitation id="13173CCDFF24099A61E3FCFCCF38FCCD" box="[180,274,795,819]" captionStart="Figs" captionStartId="177.[115,154,1197,1216]" captionTargetBox="[256,940,223,1169]" captionTargetId="figure-107@177.[224,977,219,1184]" captionTargetPageId="177" captionText="Figs. 552558.Philothis (Atavinus) atavus (Reichardt, 1931), male terminalia: 552 8th sternite and tergite, ventral view; 553 ditto, dorsal view; 554 ditto, lateral view; 555 9th tergite, 10th tergite (dorsal view) and spiculum gastrale (ventral view); 556 9th tergite, 10th tergite and spiculum gastrale, lateral view; 557 aedeagus, dorsal view; 558 ditto, lateral view." figureDoi="http://doi.org/10.5281/zenodo.4272272" httpUri="https://zenodo.org/record/4272272/files/figure.png" pageId="179" pageNumber="172">Fig. 558</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>