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<document id="7A0D8428E665CE7A748E6CC9D84A9963" ID-DOI="10.11646/zootaxa.4742.2.6" ID-GBIF-Dataset="a741a81e-10e2-4068-904d-58a7b1ca9fea" ID-ISSN="1175-5326" ID-Zenodo-Dep="3677845" ID-ZooBank="FB7C6E62-8F89-44E6-967D-1049BBD03C7E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582272800686" checkinUser="plazi" docAuthor="Zhu, Wenhui, Yao, Zhiyuan, Zheng, Guo &amp; Li, Shuqiang" docDate="2020" docId="0385475AFFC5FFD172D48D2EFCBCFEC8" docLanguage="en" docName="zootaxa.4742.2.6.pdf" docOrigin="Zootaxa 4742 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana mengla Zhu &amp; Yao &amp; Zheng &amp; Li 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="334" masterDocId="FFBC3F22FFC4FFD372438923FFB6FFD7" masterDocTitle="Three new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from Xishuangbanna, Yunnan, China" masterLastPageNumber="342" masterPageNumber="332" pageNumber="333" updateTime="1698720804479" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D5948FB19EE71B3321F677CEE9F40013">Three new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from Xishuangbanna, Yunnan, China</mods:title>
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<mods:namePart id="F6A5A0C19048DCDB01EC74B69E47AC79">Zhu, Wenhui</mods:namePart>
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<mods:namePart id="31B8459B8A0E75BD08BA9B119E6F0C8F">Yao, Zhiyuan</mods:namePart>
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<mods:namePart id="4B73D0EB11E6AF534BC8DBA72ADD0A59">Zheng, Guo</mods:namePart>
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<mods:namePart id="6C18126C3768E9C4234E1EEE70D09FEC">Li, Shuqiang</mods:namePart>
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<mods:date id="A6988132B21F263F246FDA5486B6F37A">2020</mods:date>
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<treatment id="0385475AFFC5FFD172D48D2EFCBCFEC8" ID-DOI="http://doi.org/10.5281/zenodo.3684927" ID-GBIF-Taxon="162244871" ID-Zenodo-Dep="3684927" LSID="urn:lsid:plazi:treatment:0385475AFFC5FFD172D48D2EFCBCFEC8" httpUri="http://treatment.plazi.org/id/0385475AFFC5FFD172D48D2EFCBCFEC8" lastPageId="2" lastPageNumber="334" pageId="1" pageNumber="333">
<subSubSection id="C336A5C7FFC5FFD272D48D2EFDF6FBF0" box="[151,576,1037,1064]" pageId="1" pageNumber="333" type="nomenclature">
<paragraph id="8B93F64CFFC5FFD272D48D2EFDF6FBF0" blockId="1.[151,576,1037,1099]" box="[151,576,1037,1064]" pageId="1" pageNumber="333">
<heading id="D0DB4120FFC5FFD272D48D2EFDF6FBF0" bold="true" box="[151,576,1037,1064]" fontSize="11" level="1" pageId="1" pageNumber="333" reason="1">
<emphasis id="B9582A5EFFC5FFD272D48D2EFDF6FBF0" bold="true" box="[151,576,1037,1064]" pageId="1" pageNumber="333">
<taxonomicName id="4C2C8DCFFFC5FFD272D48D2EFE6AFBF0" ID-CoL="98ZVL" authority="Yao &amp; Li" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[151,476,1037,1064]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="333" phylum="Arthropoda" rank="species" species="mengla" status="sp. nov.">
<emphasis id="B9582A5EFFC5FFD272D48D2EFED4FBFF" bold="true" box="[151,354,1037,1064]" italics="true" pageId="1" pageNumber="333">Belisana mengla</emphasis>
Yao &amp; Li
</taxonomicName>
<taxonomicNameLabel id="A26B9725FFC5FFD273A08D2EFDF6FBF0" box="[483,576,1037,1063]" pageId="1" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C336A5C7FFC5FFD272D48D12FF4EFB9C" box="[151,248,1073,1099]" pageId="1" pageNumber="333" type="description">
<paragraph id="8B93F64CFFC5FFD272D48D12FF4EFB9C" blockId="1.[151,576,1037,1099]" box="[151,248,1073,1099]" pageId="1" pageNumber="333">
<figureCitation id="1317EAC9FFC5FFD272D48D12FF4EFB9C" box="[151,248,1073,1099]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="3.[151,250,1894,1919]" captionStartId-1="4.[151,250,1886,1911]" captionTargetBox-0="[218,1369,181,1874]" captionTargetBox-1="[221,1366,181,1866]" captionTargetId-0="figure@3.[218,1369,181,1874]" captionTargetId-1="figure@4.[221,1366,181,1866]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." captionText-1="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.3677847" figureDoi-1="http://doi.org/10.5281/zenodo.3677849" httpUri-0="https://zenodo.org/record/3677847/files/figure.png" httpUri-1="https://zenodo.org/record/3677849/files/figure.png" pageId="1" pageNumber="333">Figs 12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C336A5C7FFC5FFD272D48D5AFD6EFB0C" pageId="1" pageNumber="333" type="materials_examined">
<paragraph id="8B93F64CFFC5FFD272D48D5AFD6EFB0C" blockId="1.[151,1437,1144,2035]" pageId="1" pageNumber="333">
<emphasis id="B9582A5EFFC5FFD272D48D5AFEF3FB44" bold="true" box="[151,325,1144,1171]" pageId="1" pageNumber="333">Type material.</emphasis>
<materialsCitation id="3B44FC11FFC5FFD273088D5AFA96FB60" ID-GBIF-Occurrence="2570055983" collectingDate="2012-06-30" collectionCode="IZCAS" collectorName="Q. Zhao &amp; Z. Chen" country="China" county="Mengla County" elevation="22" latitude="21.4055" location="Xiaolongha Village" longLatPrecision="1" longitude="101.61703" municipality="Xishuangbanna" pageId="1" pageNumber="333" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B9582A5EFFC5FFD273088D5AFE77FB44" bold="true" box="[331,449,1145,1171]" pageId="1" pageNumber="333">
<typeStatus id="549748EEFFC5FFD273088D5AFE0AFB44" box="[331,444,1145,1171]" pageId="1" pageNumber="333" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male (
<collectionCode id="ED3D6E89FFC5FFD270538D5AFDD5FB44" box="[528,611,1145,1171]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="333">IZCAS</collectionCode>
Ar39745),
<location id="8EF3A097FFC5FFD270A18D5AFC09FB44" LSID="urn:lsid:plazi:treatment:0385475AFFC5FFD172D48D2EFCBCFEC8:8EF3A097FFC5FFD270A18D5AFC09FB44" box="[738,959,1145,1171]" country="China" county="Mengla County" latitude="21.4055" longLatPrecision="1" longitude="101.61703" municipality="Xishuangbanna" name="Xiaolongha Village" pageId="1" pageNumber="333" stateProvince="Yunnan">Xiaolongha Village</location>
, (
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,
<geoCoordinate id="EE18908BFFC5FFD2763C8D5AFA9CFB44" box="[1151,1322,1145,1171]" degrees="101" direction="east" minutes="37" orientation="longitude" pageId="1" pageNumber="333" precision="1" seconds="01.3" value="101.61703">101°3701.3”E</geoCoordinate>
, elevation 801 ±
<quantity id="4CD45BA9FFC5FFD2729D8DBEFEA2FB60" box="[222,276,1181,1207]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="333" unit="m" value="22.0">
<elevation id="0001117FFFC5FFD2729D8DBEFEA2FB60" box="[222,276,1181,1207]" metricMagnitude="1" metricUnit="m" metricValue="2.2" pageId="1" pageNumber="333" unit="m" value="22.0">22m</elevation>
</quantity>
),
<collectingCounty id="62F28EC0FFC5FFD273648DBEFE6EFB60" box="[295,472,1181,1207]" pageId="1" pageNumber="333">Mengla County</collectingCounty>
,
<collectingMunicipality id="6BF76C36FFC5FFD273A18DBEFD25FB60" box="[482,659,1181,1207]" pageId="1" pageNumber="333">Xishuangbanna</collectingMunicipality>
,
<emphasis id="B9582A5EFFC5FFD270DE8DBEFCE4FB60" bold="true" box="[669,850,1180,1207]" pageId="1" pageNumber="333">
<collectingRegion id="49E838AEFFC5FFD270DE8DBEFD4BFB60" box="[669,765,1181,1207]" country="China" name="Yunnan" pageId="1" pageNumber="333">Yunnan</collectingRegion>
,
<collectingCountry id="F33BB6DCFFC5FFD2714A8DBFFCE4FB60" box="[777,850,1180,1207]" name="China" pageId="1" pageNumber="333">China</collectingCountry>
</emphasis>
,
<date id="FF92D08CFFC5FFD271238DBEFC4EFB60" box="[864,1016,1181,1207]" pageId="1" pageNumber="333" value="2012-06-30">
<collectingDate id="EFD62964FFC5FFD271238DBEFC4EFB60" box="[864,1016,1181,1207]" pageId="1" pageNumber="333" value="2012-06-30">30 June 2012</collectingDate>
</date>
,
<collectorName id="26D9939AFFC5FFD276408DBEFBE9FB60" box="[1027,1119,1181,1207]" pageId="1" pageNumber="333">Q. Zhao</collectorName>
and
<collectorName id="26D9939AFFC5FFD276D68DBEFB59FB60" box="[1173,1263,1181,1207]" pageId="1" pageNumber="333">Z. Chen</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B44FC11FFC5FFD277658DBEFD62FB0C" ID-GBIF-Occurrence="2570055982" collectingDate="2012-06-30" collectionCode="IZCAS" collectorName="Q. Zhao &amp; Z. Chen" country="China" county="Mengla County" elevation="22" latitude="21.4055" location="Xiaolongha Village" longLatPrecision="1" longitude="101.61703" municipality="Xishuangbanna" pageId="1" pageNumber="333" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B9582A5EFFC5FFD277658DBEFA2AFB60" bold="true" box="[1318,1436,1181,1207]" pageId="1" pageNumber="333">
<typeStatus id="549748EEFFC5FFD277658DBEFA21FB60" box="[1318,1431,1181,1207]" pageId="1" pageNumber="333" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D2A3DC5FFC5FFD272D48DE2FF4EFB0C" box="[151,248,1217,1243]" pageId="1" pageNumber="333" type="female">1 female</specimenCount>
(
<collectionCode id="ED3D6E89FFC5FFD273448DE2FEECFB0C" box="[263,346,1217,1243]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="333">IZCAS</collectionCode>
Ar39746), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C336A5C7FFC5FFD272848DC6FBCAFB28" box="[199,1148,1253,1279]" pageId="1" pageNumber="333" type="etymology">
<paragraph id="8B93F64CFFC5FFD272848DC6FBCAFB28" blockId="1.[151,1437,1144,2035]" box="[199,1148,1253,1279]" pageId="1" pageNumber="333">
<emphasis id="B9582A5EFFC5FFD272848DC6FEF8FB28" bold="true" box="[199,334,1253,1279]" pageId="1" pageNumber="333">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="549748EEFFC5FFD270F68DC6FD53FB28" box="[693,741,1253,1279]" pageId="1" pageNumber="333">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C336A5C7FFC5FFD272848C2BFD3AFA00" pageId="1" pageNumber="333" type="diagnosis">
<paragraph id="8B93F64CFFC5FFD272848C2BFD3AFA00" blockId="1.[151,1437,1144,2035]" pageId="1" pageNumber="333">
<emphasis id="B9582A5EFFC5FFD272848C2BFEF4FAF4" bold="true" box="[199,322,1288,1315]" pageId="1" pageNumber="333">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="4C2C8DCFFFC5FFD2700C8C29FC7FFAF4" authority="Yao &amp; Li, 2018" authorityName="Yao &amp; Li" authorityYear="2018" box="[591,969,1289,1315]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="333" phylum="Arthropoda" rank="species" species="yangxiaodongi">
<emphasis id="B9582A5EFFC5FFD2700C8C29FCA2FAF4" box="[591,788,1289,1315]" italics="true" pageId="1" pageNumber="333">B. yangxiaodongi</emphasis>
<bibRefCitation id="EFBD8BBDFFC5FFD271588C2AFC7FFAF4" author="Yao, Z. &amp; Zhu, K. &amp; Du, Z. &amp; Li, S." box="[795,969,1289,1315]" pageId="1" pageNumber="333" pagination="243 - 262" refId="ref4560" refString="Yao, Z., Zhu, K., Du, Z. &amp; Li, S. (2018) The Belisana spiders (Araneae: Pholcidae) from Xishuangbanna Tropical Botanical Garden, Yunnan, China. Zootaxa, 4425 (2), 243 - 262. https: // doi. org / 10.11646 / zootaxa. 4425.2.3" type="journal article" year="2018">Yao &amp; Li, 2018</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFBD8BBDFFC5FFD276418C2AFB5EFAF4" author="Yao, Z. &amp; Zhu, K. &amp; Du, Z. &amp; Li, S." box="[1026,1256,1288,1315]" pageId="1" pageNumber="333" pagination="243 - 262" refId="ref4560" refString="Yao, Z., Zhu, K., Du, Z. &amp; Li, S. (2018) The Belisana spiders (Araneae: Pholcidae) from Xishuangbanna Tropical Botanical Garden, Yunnan, China. Zootaxa, 4425 (2), 243 - 262. https: // doi. org / 10.11646 / zootaxa. 4425.2.3" type="journal article" year="2018">
Yao
<emphasis id="B9582A5EFFC5FFD276768C29FBDBFAF4" box="[1077,1133,1289,1315]" italics="true" pageId="1" pageNumber="333">et al.</emphasis>
2018: 255
</bibRefCitation>
, figs 9
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, 12E) with similar male chelicerae (
<figureCitation id="1317EAC9FFC5FFD2706C8C0EFD3DFA90" box="[559,651,1325,1351]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="1" pageNumber="333">Fig. 2D</figureCitation>
), bulbal apophyses (
<figureCitation id="1317EAC9FFC5FFD2713A8C0EFC64FA90" box="[889,978,1325,1351]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="1" pageNumber="333">Fig. 2C</figureCitation>
) and vulva (
<figureCitation id="1317EAC9FFC5FFD276248C0EFB76FA90" box="[1127,1216,1325,1351]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="1" pageNumber="333">Fig. 2B</figureCitation>
) but can be distinguished by presence of dorsal apophysis proximally on pedipalpal femur (arrow in
<figureCitation id="1317EAC9FFC5FFD276018C72FB2DFABC" box="[1090,1179,1361,1387]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Fig. 1A</figureCitation>
), absence of spine distally and membranous flap retrolaterally on procursus (
<figureCitation id="1317EAC9FFC5FFD270BE8C56FC88FA58" box="[765,830,1397,1423]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Figs 1</figureCitation>
AD), presence of angular distal sclerite retrolaterally (arrow
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<figureCitation id="1317EAC9FFC5FFD273588CBAFEC7FA64" box="[283,369,1433,1459]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Fig. 1D</figureCitation>
) and large, flat, distal membranous lamella (arrow
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<figureCitation id="1317EAC9FFC5FFD271A08CBAFB8FFA64" box="[995,1081,1433,1459]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Fig. 1D</figureCitation>
) on procursus, and by presence of subcuticular teeth on epigynum (
<figureCitation id="1317EAC9FFC5FFD270668C9EFDC9FA00" box="[549,639,1469,1495]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="1" pageNumber="333">Fig. 2A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C336A5C7FFC5FFD172848CC3FAB8FF2C" lastPageId="2" lastPageNumber="334" pageId="1" pageNumber="333" type="description">
<paragraph id="8B93F64CFFC5FFD272848CC3FCCDF87C" blockId="1.[151,1437,1144,2035]" pageId="1" pageNumber="333">
<emphasis id="B9582A5EFFC5FFD272848CC3FD91FA2C" bold="true" box="[199,551,1504,1531]" pageId="1" pageNumber="333">
Description. Male (
<typeStatus id="549748EEFFC5FFD273EE8CC3FDADFA2C" box="[429,539,1504,1531]" pageId="1" pageNumber="333" type="holotype">holotype</typeStatus>
):
</emphasis>
Total length 1.50 (1.61 with clypeus), carapace 0.73 long, 0.63 wide, opisthosoma 0.77 long, 0.56 wide. Leg I: 11.32 (3.80 + 0.40 + 2.64 + 3.36 + 1.12), leg II: — (2.19 + 0.31 + 2.00 + 2.91 + ), leg III missing, leg IV: — (2.10 + 0.50 + 1.75 + 2.35 + ); tibia I L/d: 42. Habitus as in
<figureCitation id="1317EAC9FFC5FFD276F08F0AFB4EF994" box="[1203,1272,1577,1603]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="1" pageNumber="333">Figs 2</figureCitation>
EF. Carapace and sternum pale, without marks. Legs pale, without darker rings. Opisthosoma pale, without spots. Distance PME- PME 0.13, diameter PME 0.08, distance PME-ALE 0.03, AME absent. Ocular area not elevated. Thoracic furrow absent. Clypeus unmodified. Sternum approximately as wide as long (0.64). Chelicerae (
<figureCitation id="1317EAC9FFC5FFD276318FB6FB7FF978" box="[1138,1225,1685,1711]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="1" pageNumber="333">Fig. 2D</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses each provided with modified seta on its tip (distance between tips: 0.42). Pedipalps as in
<figureCitation id="1317EAC9FFC5FFD270D88FFEFD6BF920" box="[667,733,1757,1783]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Figs 1</figureCitation>
AB; coxa unmodified; trochanter with very short retrolateroventral apophysis (arrow in
<figureCitation id="1317EAC9FFC5FFD273978E22FD9CF8CC" box="[468,554,1793,1819]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Fig. 1B</figureCitation>
); femur with small dorsal apophysis proximally (arrow in
<figureCitation id="1317EAC9FFC5FFD276838E22FAAFF8CC" box="[1216,1305,1793,1819]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Fig. 1A</figureCitation>
); procursus (
<figureCitation id="1317EAC9FFC5FFD272DC8E06FF57F8E8" box="[159,225,1829,1855]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Figs 1</figureCitation>
AD) simple proximally but complex distally, with angular distal sclerite retrolaterally (arrow
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<figureCitation id="1317EAC9FFC5FFD2777E8E06FA25F8E8" box="[1341,1427,1829,1855]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Fig. 1D</figureCitation>
) and large, flat, distal membranous lamella (arrow
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<figureCitation id="1317EAC9FFC5FFD2714B8E6AFCD7F8B4" box="[776,865,1865,1891]" captionStart="FIGURE 1" captionStartId="3.[151,250,1894,1919]" captionTargetBox="[218,1369,181,1874]" captionTargetId="figure@3.[218,1369,181,1874]" captionTargetPageId="3" captionText="FIGURE 1. Belisana mengla sp. nov., holotype male. AB. Pedipalp (A. Prolateral view, arrow points at dorsal apophysis on pedipalpal femur; B. Retrolateral view, arrow points at retrolatero-ventral apophysis on pedipalpal trochanter); CD. Distal part of procursus (C. Prolateral view; D. Retrolateral view, arrow 1 points at angular distal sclerite, arrow 2 points at large, flat, distal membranous lamella). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.02 (AB), 0.01 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3677847" httpUri="https://zenodo.org/record/3677847/files/figure.png" pageId="1" pageNumber="333">Fig. 1D</figureCitation>
); bulb (
<figureCitation id="1317EAC9FFC5FFD271FE8E6AFBA2F8B4" box="[957,1044,1865,1891]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="1" pageNumber="333">Fig. 2C</figureCitation>
) with hooked apophysis and wide embolus. Retrolateral trichobothria of tibia I at 13% proximally; legs with short vertical setae on metatarsi, without spines and curved setae; tarsus I with 17 distinct pseudosegments.
</paragraph>
<paragraph id="8B93F64CFFC5FFD172848E96FAB8FF2C" blockId="1.[151,1437,1144,2035]" lastBlockId="2.[151,1437,152,287]" lastPageId="2" lastPageNumber="334" pageId="1" pageNumber="333">
<emphasis id="B9582A5EFFC5FFD272848E96FEA8F818" bold="true" box="[199,286,1973,1999]" pageId="1" pageNumber="333">Female</emphasis>
(IZCAS Ar39746): Similar to male, habitus as in
<figureCitation id="1317EAC9FFC5FFD271238E96FC11F818" box="[864,935,1973,1999]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="1" pageNumber="333">Figs 2</figureCitation>
GH. Total length 1.40 (1.52 with clypeus), carapace 0.68 long, 0.56 wide, opisthosoma 0.72 long, 0.68 wide; tibia I: 1.42; tibia I L/d: 37. Distance PME-PME 0.14, diameter PME 0.08, distance PME-ALE 0.03, AME absent. Sternum approximately as wide as long (0.58). Epigynum (
<figureCitation id="1317EAC9FFC6FFD1735E899EFECEFF00" box="[285,376,189,215]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="2" pageNumber="334">Fig. 2A</figureCitation>
) simple and flat, with pair of pockets 0.47 apart (ep in
<figureCitation id="1317EAC9FFC6FFD171B9899EFBF4FF00" box="[1018,1090,189,215]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="2" pageNumber="334">Figs 2</figureCitation>
AB) and several subcuticular teeth. Vulva (
<figureCitation id="1317EAC9FFC6FFD1736E89C2FE35FF2C" box="[301,387,225,251]" captionStart="FIGURE 2" captionStartId="4.[151,250,1886,1911]" captionTargetBox="[221,1366,181,1866]" captionTargetId="figure@4.[221,1366,181,1866]" captionTargetPageId="4" captionText="FIGURE 2. Belisana mengla sp. nov., holotype male (CF) and paratype female (AB, GH). A. Epigynum, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 (AB), 0.05 (CD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3677849" httpUri="https://zenodo.org/record/3677849/files/figure.png" pageId="2" pageNumber="334">Fig. 2B</figureCitation>
) with ridge-shaped anterior arch and pair of narrow, curved pore plates far apart.
</paragraph>
</subSubSection>
<subSubSection id="C336A5C7FFC6FFD172848827FCBCFEC8" box="[199,778,260,287]" pageId="2" pageNumber="334" type="distribution">
<paragraph id="8B93F64CFFC6FFD172848827FCBCFEC8" blockId="2.[151,1437,152,287]" box="[199,778,260,287]" pageId="2" pageNumber="334">
<emphasis id="B9582A5EFFC6FFD172848827FED7FEC8" bold="true" box="[199,353,260,287]" pageId="2" pageNumber="334">Distribution.</emphasis>
<collectingCountry id="F33BB6DCFFC6FFD173248826FE1CFEC8" box="[359,426,261,287]" name="China" pageId="2" pageNumber="334">China</collectingCountry>
(
<collectingRegion id="49E838AEFFC6FFD173FA8826FDA5FEC8" box="[441,531,261,287]" country="China" name="Yunnan" pageId="2" pageNumber="334">Yunnan</collectingRegion>
,
<typeStatus id="549748EEFFC6FFD1705D8826FDF8FEC8" box="[542,590,261,287]" pageId="2" pageNumber="334">type</typeStatus>
locality;
<figureCitation id="1317EAC9FFC6FFD170FB8826FD4AFEC8" box="[696,764,261,287]" captionStart="FIGURE 7" captionStartId="9.[151,250,1983,2008]" captionTargetBox="[212,1376,741,1963]" captionTargetId="figure@9.[212,1376,741,1963]" captionTargetPageId="9" captionText="FIGURE 7. Distribution records of three new Belisana species. 1. B. mengla Yao &amp; Li sp. nov.; 2. B. menglun Yao &amp; Li sp. nov.; 3. B. mengyang Yao &amp; Li sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3677859" httpUri="https://zenodo.org/record/3677859/files/figure.png" pageId="2" pageNumber="334">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>