831 lines
107 KiB
XML
831 lines
107 KiB
XML
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<mods:title id="272B949193277085E6E730323F44B98B">Two new species of Eleutherodactylus (Anura: Eleutherodactylidae) from Southern Mexico, with comments on the taxonomy of related species and their advertisement calls</mods:title>
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<mods:namePart id="CC300A4593B30E8F9732B78191C94412">Grünwald, Christoph I.</mods:namePart>
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<mods:affiliation id="B5DD21CA7FA09EB2B168F8261DACB49F">Biencom Real Estate, Carretera Chapala-Jocotepec # 57 - 1, C. P. 45920, Ajijic, Jalisco, MEXICO 2 Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, MEXICO 3 Herp. mx A. C., Villa de Alvarez, Colima, MEXICO 4 Los Angeles Zoo and Botanical Gardens, 45333 Zoo Drive, Los Angeles, California 90027, USA</mods:affiliation>
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<mods:nameIdentifier id="7130BBA288E7F725A02E0DC5A689F0EA" type="email">cgruenwald@switaki.com</mods:nameIdentifier>
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<mods:namePart id="C766ECA91018907BD87F81DA540C5FD5">Reyes-Velasco, Jacobo</mods:namePart>
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<mods:namePart id="ECC935DC5CE72D180847487D8B14C194">Franz-Chávez, Héctor</mods:namePart>
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<mods:namePart id="D343765A93CAC0F7A44C918BA759A1EB">Morales-Flores, Karen I.</mods:namePart>
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<mods:namePart id="0AF799FD8A5119635C055BDC8B19D33F">Ahumada-Carrillo, Ivan T.</mods:namePart>
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<mods:namePart id="19AE0D75C5A5D2E93DA6C989CB3391AC">Rodriguez, Christopher M.</mods:namePart>
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<mods:namePart id="CF40EB6C3DFD304FA728BD2AF03C970F">Jones, Jason M.</mods:namePart>
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<mods:title id="16FC62AB6719527F8455401A4C9EA020">Amphibian & Reptile Conservation</mods:title>
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<mods:date id="47025A72C35C7DE12834A1683ADF4397">2021</mods:date>
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<treatment id="670387A7FFB1FF9DFF0FF8B2FC55F9BC" ID-DOI="http://doi.org/10.5281/zenodo.11265062" ID-GBIF-Taxon="230931092" ID-Zenodo-Dep="11265062" LSID="urn:lsid:plazi:treatment:670387A7FFB1FF9DFF0FF8B2FC55F9BC" httpUri="http://treatment.plazi.org/id/670387A7FFB1FF9DFF0FF8B2FC55F9BC" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
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<subSubSection id="A7B0653AFFB1FF9FFF0FF8B2FE78F885" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="EF1536B1FFB1FF9FFF0FF8B2FE78F885" blockId="13.[141,746,1842,1904]" pageId="13" pageNumber="14">
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<heading id="B45D81DDFFB1FF9FFF0FF8B2FE78F885" bold="true" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="6">
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<emphasis id="DDDEEAA3FFB1FF9FFF0FF8B2FE78F885" bold="true" pageId="13" pageNumber="14">
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Summary Data for
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<taxonomicName id="28AA4D32FFB1FF9FFE1EF8B2FD5DF8B8" ID-CoL="399J5" authority="(Peters, 1870)" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[412,746,1842,1869]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="nitidus">
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<emphasis id="DDDEEAA3FFB1FF9FFE1EF8B2FD5DF8B8" bold="true" box="[412,746,1842,1869]" italics="true" pageId="13" pageNumber="14">Eleutherodactylus nitidus</emphasis>
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</taxonomicName>
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Species Group Members
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="A7B0653AFFB1FF9CFF0FF818FB1EF86B" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="description">
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<paragraph id="EF1536B1FFB1FF9FFF0FF818FBD2FE73" blockId="13.[141,744,1943,1967]" lastBlockId="13.[791,1394,142,390]" pageId="13" pageNumber="14">
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To help differentiate between the two new species described above, and the related species in the
|
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<taxonomicName id="28AA4D32FFB1FF9FFC95FF2EFB88FF33" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[791,1087,174,198]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="nitidus">
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<emphasis id="DDDEEAA3FFB1FF9FFC95FF2EFB88FF33" box="[791,1087,174,198]" italics="true" pageId="13" pageNumber="14">Eleutherodactylus nitidus</emphasis>
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</taxonomicName>
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species group, important morphological, mensural, and call differences of all of the species in this group are collated into a single table (
|
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<tableCitation id="A228030AFFB1FF9FFCDFFE8EFC03FED3" box="[861,948,270,294]" captionStart="Table 3" captionStartId="16.[124,180,451,475]" captionTargetPageId="16" captionText="Table 3. Key comparative characters of members of the Eleutherodactylus nitidus species group." httpUri="http://table.plazi.org/id/BBD56639FFACFF82FFFEFE43FBA8FE2F" pageId="13" pageNumber="14" tableUuid="BBD56639FFACFF82FFFEFE43FBA8FE2F">Table 3</tableCitation>
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), and photographs of related species in the
|
||
<taxonomicName id="28AA4D32FFB1FF9FFCBDFEAEFC1CFEB3" authority="(Peters, 1870)" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[831,939,302,326]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="nitidus">
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<emphasis id="DDDEEAA3FFB1FF9FFCBDFEAEFC1CFEB3" box="[831,939,302,326]" italics="true" pageId="13" pageNumber="14">E. nitidus</emphasis>
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</taxonomicName>
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||
species group are provided (
|
||
<figureCitation id="77912A34FFB1FF9FFB69FEAEFAD1FEB3" box="[1259,1382,302,326]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="18.[141,176,1763,1785]" captionStartId-1="19.[128,163,1763,1785]" captionTargetBox-0="[135,1395,231,1706]" captionTargetBox-1="[129,1414,232,1739]" captionTargetId-0="figure-31@18.[135,1395,724,974]" captionTargetId-1="figure-28@19.[129,986,736,991]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Fig. 9. Comparison photos of Eleutherodactylus species related to E. nitidus in life. (A–I) E. syristes from the vicinity of Agua de Obispo, Municipality of Chilpancingo, Guerrero, Mexico. (J–O) E. albolabris from Agua de Obispo, Municipality of Chilpancingo, Guerrero, Mexico. (P–R) E. nitidus from Yerba Santa, Municipality of General Heliodoro Castillo, Guerrero, Mexico." captionText-1="Fig. 10. Comparison photos of Eleutherodactylus species related to E. nitidus in life. (A–I) E. dilatus from Omiltemi, Municipality of Chilpancingo, Guerrero, Mexico. (J–O) E. petersi from Puerto El Balsamo, Municipality of José Azueta, Guerrero, Mexico. (P–R) E. pipilans from Acahuizotla, Municipality of Chilpancingo, Guerrero, Mexico." figureDoi-0="http://doi.org/10.5281/zenodo.11265074" figureDoi-1="http://doi.org/10.5281/zenodo.11265076" httpUri-0="https://zenodo.org/record/11265074/files/figure.png" httpUri-1="https://zenodo.org/record/11265076/files/figure.png" pageId="13" pageNumber="14">Figs. 9–10</figureCitation>
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). The distributions of the species in the
|
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<taxonomicName id="28AA4D32FFB1FF9FFB42FECEFA9CFE93" authority="(Peters, 1870)" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[1216,1323,334,358]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="nitidus">
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<emphasis id="DDDEEAA3FFB1FF9FFB42FECEFA9CFE93" box="[1216,1323,334,358]" italics="true" pageId="13" pageNumber="14">E. nitidus</emphasis>
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</taxonomicName>
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group are mapped in
|
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<figureCitation id="77912A34FFB1FF9FFC3CFEEEFBBDFE70" box="[958,1034,366,390]" captionStart="Fig" captionStartId="13.[791,826,1658,1680]" captionTargetBox="[791,1395,909,1631]" captionTargetId="figure-532@13.[791,1395,909,1631]" captionTargetPageId="13" captionText="Fig. 7. (A) Map showing the type localities and distribution of Eleutherodactylus species related to E. nitidus in southern Mexico. The stars represent type localities and circles represent additional localities, with each color coded for the species: E. sentinelus sp. nov. (purple), E. maculabialis sp. nov. (green), E. dilatus (pink), E. maurus (yellow), and E. syristes (blue). (B) Map showing the type localities and distribution of Eleutherodactylus species related to E. pipilans in southern Mexico. The red star represents the type locality of E. pipilans and red circles represent additional localities. The black star represents the type locality of E. nebulosus and black circles represent additional localities." figureDoi="http://doi.org/10.5281/zenodo.11265068" httpUri="https://zenodo.org/record/11265068/files/figure.png" pageId="13" pageNumber="14">Figs. 7</figureCitation>
|
||
and
|
||
<figureCitation id="77912A34FFB1FF9FFBC0FEEEFBD2FE73" box="[1090,1125,366,390]" captionStart="Fig" captionStartId="23.[141,176,622,644]" captionTargetBox="[141,745,154,609]" captionTargetId="figure-730@23.[141,745,154,609]" captionTargetPageId="23" captionText="Fig. 13. Map of type localities and distributions of members of the Eleutherodactylus nitidus species group. Purple circles represent localities of an undescribed species related to E. petersi. For the other four species, stars represent the type localities and circles represent additional localities, which are color coded for species: E. orarius (red), E. albolabris (blue), E. nitidus (green), and E. petersi (orange)." figureDoi="http://doi.org/10.5281/zenodo.11265082" httpUri="https://zenodo.org/record/11265082/files/figure.png" pageId="13" pageNumber="14">13.</figureCitation>
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</paragraph>
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<paragraph id="EF1536B1FFB1FF9FFC95FE2FFBEFFDE5" blockId="13.[791,1395,431,528]" pageId="13" pageNumber="14">
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<emphasis id="DDDEEAA3FFB1FF9FFC95FE2FFBEFFDE5" bold="true" pageId="13" pageNumber="14">
|
||
Differentiating the Advertisement Calls of Closely Related
|
||
<taxonomicName id="28AA4D32FFB1FF9FFB94FE52FAC4FE18" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[1046,1395,466,493]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="nitidus">
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||
<emphasis id="DDDEEAA3FFB1FF9FFB94FE52FAC4FE18" bold="true" box="[1046,1395,466,493]" italics="true" pageId="13" pageNumber="14">Eleutherodactylus nitidus</emphasis>
|
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</taxonomicName>
|
||
Species Group Members
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB1FF9CFC95FDB7FE69FA4E" blockId="13.[791,1395,567,879]" lastBlockId="14.[141,746,1123,1467]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
Recordings were made from 120 calling males of all 24 species of
|
||
<taxonomicName id="28AA4D32FFB1FF9FFC2EFDD7FBCFFD9A" authorityName="Dumeril and Bibron" authorityYear="1841" box="[940,1144,599,623]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="DDDEEAA3FFB1FF9FFC2EFDD7FBCFFD9A" box="[940,1144,599,623]" italics="true" pageId="13" pageNumber="14">Eleutherodactylus</emphasis>
|
||
</taxonomicName>
|
||
from western
|
||
<collectingCountry id="97BD7621FFB1FF9FFA95FDD7FAD8FD9A" box="[1303,1391,599,623]" name="Mexico" pageId="13" pageNumber="14">Mexico</collectingCountry>
|
||
. The calls of the species analyzed were found to fall into five different categories of a rapid burst whistle (trill), a drawn-out whistle (whistle), a strong high-pitched chirp (peep), a soft high-pitched chirp (chirp), and a drawn-out chirp (pipe). Of the species analyzed herein, four (
|
||
<emphasis id="DDDEEAA3FFB1FF9FFC3AFC98FC1DFCBA" italics="true" pageId="13" pageNumber="14">
|
||
<taxonomicName id="28AA4D32FFB1FF9FFC3AFC98FB99FCDA" baseAuthorityName="Davis and Dixon" baseAuthorityYear="1955" box="[952,1070,791,815]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="dilatus">E. dilatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB1FF9FFBC2FC98FB6EFCDA" baseAuthorityName="Taylor" baseAuthorityYear="1943" box="[1088,1241,791,815]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="nebulosus">E. nebulosus</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB1FF9FFB69FC98FAD9FCDA" baseAuthorityName="Taylor" baseAuthorityYear="1940" box="[1259,1390,791,815]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="pipilans">E. pipilans</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB1FF9FFC95FCB8FC1DFCBA" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[791,938,823,847]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="sentinelus">E. sentinelus</taxonomicName>
|
||
</emphasis>
|
||
) produce a strongly high-pitched chirp (peep), four (
|
||
<emphasis id="DDDEEAA3FFB1FF9CFC38FCD8FEB6FB8E" italics="true" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
<taxonomicName id="28AA4D32FFB1FF9FFC38FCD8FBEFFC9A" baseAuthorityName="Taylor" baseAuthorityYear="1943" box="[954,1112,855,879]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="albolabris">E. albolabris</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB1FF9FFBE8FCD8FB57FC9A" authority="(Peters, 1870)" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[1130,1248,855,879]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="nitidus">E. nitidus</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB1FF9FFB71FCD8FAD9FC9A" baseAuthorityName="Duellman" baseAuthorityYear="1958" box="[1267,1390,856,879]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="orarius">E. orarius</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB2FF9CFF0FFBE4FEB6FB8E" baseAuthorityName="Duellman" baseAuthorityYear="1954" box="[141,257,1124,1147]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="petersi">E. petersi</taxonomicName>
|
||
</emphasis>
|
||
) produce a drawn-out whistle (whistle), two (
|
||
<taxonomicName id="28AA4D32FFB2FF9CFF48FB04FECBFB6E" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[202,380,1155,1179]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maculabialis">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFF48FB04FECBFB6E" box="[202,380,1155,1179]" italics="true" pageId="14" pageNumber="15">E. maculabialis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="28AA4D32FFB2FF9CFE3AFB04FD99FB6E" baseAuthorityName="Hoyt" baseAuthorityYear="1965" box="[440,558,1156,1179]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="syristes">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFE3AFB04FD99FB6E" box="[440,558,1156,1179]" italics="true" pageId="14" pageNumber="15">E. syristes</emphasis>
|
||
</taxonomicName>
|
||
) produce multi-note or pulsed calls (trill) of various rates and notes, and one (
|
||
<taxonomicName id="28AA4D32FFB2FF9CFF79FB44FED8FB2E" authorityName="Hedges" authorityYear="1989" box="[251,367,1220,1243]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maurus">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFF79FB44FED8FB2E" box="[251,367,1220,1243]" italics="true" pageId="14" pageNumber="15">E. maurus</emphasis>
|
||
</taxonomicName>
|
||
) produces a slow and drawn-out chirp (pipe). While a detailed account is given of the calls of the species we describe here, as well as some of their relatives, we refrain from giving detailed descriptions of the calls of
|
||
<emphasis id="DDDEEAA3FFB2FF9CFE38FAC4FD3BFA8E" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="28AA4D32FFB2FF9CFE38FAC4FD90FAAE" authority="(Peters, 1870)" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[442,551,1347,1371]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="nitidus">E. nitidus</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB2FF9CFDB1FAC4FD71FAAE" baseAuthorityName="Taylor" baseAuthorityYear="1943" box="[563,710,1347,1371]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="albolabris">E. albolabris</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB2FF9CFD50FAC4FF55FA8E" baseAuthorityName="Duellman" baseAuthorityYear="1958" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="orarius">E. orarius</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB2FF9CFF6DFAE4FEE8FA8E" baseAuthorityName="Duellman" baseAuthorityYear="1954" box="[239,351,1380,1403]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="petersi">E. petersi</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB2FF9CFEE9FAE4FE5EFA8E" baseAuthorityName="Taylor" baseAuthorityYear="1940" box="[363,489,1379,1403]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="pipilans">E. pipilans</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB2FF9CFE74FAE4FD3FFA8E" baseAuthorityName="Taylor" baseAuthorityYear="1943" box="[502,648,1379,1403]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="nebulosus">E. nebulosus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and “
|
||
<taxonomicName id="28AA4D32FFB2FF9CFD50FAE4FEF7FA6E" baseAuthorityName="Taylor & Smith" baseAuthorityYear="1945" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="rubrimactulatus">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFD50FAE4FEF7FA6E" italics="true" pageId="14" pageNumber="15">E. rubrimactulatus</emphasis>
|
||
</taxonomicName>
|
||
” pending a more detailed taxonomic study of this species complex.
|
||
</paragraph>
|
||
<caption id="BBD56639FFB2FF9CFF0FFC4AFD61FBBC" ID-DOI="http://doi.org/10.5281/zenodo.11265070" ID-Zenodo-Dep="11265070" httpUri="https://zenodo.org/record/11265070/files/figure.png" pageId="14" pageNumber="15" startId="14.[141,176,970,992]" targetBox="[149,744,157,938]" targetPageId="14" targetType="figure">
|
||
<paragraph id="EF1536B1FFB2FF9CFF0FFC4AFD61FBBC" blockId="14.[141,746,970,1098]" pageId="14" pageNumber="15">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFF0FFC4AFF49FC15" bold="true" box="[141,254,970,992]" pageId="14" pageNumber="15">Fig. 8. (A)</emphasis>
|
||
Type locality of
|
||
<taxonomicName id="28AA4D32FFB2FF9CFE37FC4AFD5EFC2A" authority="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones, 2021" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[437,745,970,991]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maculabialis" status="sp. nov.">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFE37FC4AFD5EFC2A" box="[437,745,970,991]" italics="true" pageId="14" pageNumber="15">Eleutherodactylus maculabialis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DDDEEAA3FFB2FF9CFF0FFC64FF56FC0C" bold="true" box="[141,225,996,1017]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="C6ED57D8FFB2FF9CFF0FFC64FF56FC0C" box="[141,225,996,1017]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
at 11.4 km S of Puerto de Gallo, Municipio de Atoyac de Álvarez, Guerrero, Mexico.
|
||
<emphasis id="DDDEEAA3FFB2FF9CFD9FFC7EFD8AFBE1" bold="true" box="[541,573,1022,1044]" pageId="14" pageNumber="15">(B)</emphasis>
|
||
Type locality of
|
||
<taxonomicName id="28AA4D32FFB2FF9CFF0FFB99FE14FBDB" authority="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones, 2021" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[141,419,1049,1070]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="sentinelus" status="sp. nov.">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFF0FFB99FE14FBDB" box="[141,419,1049,1070]" italics="true" pageId="14" pageNumber="15">Eleutherodactylus sentinelus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DDDEEAA3FFB2FF9CFE28FB99FE4FFBDB" bold="true" box="[426,504,1049,1070]" pageId="14" pageNumber="15">
|
||
<taxonomicNameLabel id="C6ED57D8FFB2FF9CFE28FB99FE4FFBDB" box="[426,504,1049,1070]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
at 8.9 km SW of Puerto El Balsamo, Municipality of José Azueta, Guerrero, Mexico.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EF1536B1FFB2FF9CFF0FFA63FD39F96E" blockId="14.[141,745,1507,1691]" pageId="14" pageNumber="15">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFF0FFA63FE73FA0E" bold="true" box="[141,452,1507,1531]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="28AA4D32FFB2FF9CFF0FFA63FE77FA0E" baseAuthorityName="Davis and Dixon" baseAuthorityYear="1955" box="[141,448,1507,1531]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="dilatus">Eleutherodactylus dilatus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The advertisement call of
|
||
<taxonomicName id="28AA4D32FFB2FF9CFF31F984FE92F9EE" baseAuthorityName="Davis and Dixon" baseAuthorityYear="1955" box="[179,293,1539,1563]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="dilatus">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFF31F984FE92F9EE" box="[179,293,1539,1563]" italics="true" pageId="14" pageNumber="15">E. dilatus</emphasis>
|
||
</taxonomicName>
|
||
is a single high-pitched “chirp” best described as a “peep” (
|
||
<figureCitation id="77912A34FFB2FF9CFE0EF9A3FE47F9CE" box="[396,496,1571,1595]" captionStart="Fig" captionStartId="20.[141,176,1334,1356]" captionTargetBox="[181,1389,156,1293]" captionTargetId="figure-331@20.[181,1389,113,1309]" captionTargetPageId="20" captionText="Fig. 11. Oscillograms and spectrograms of the advertisement calls of adult males of Eleutherodactylus species related to E. maculabialis sp. nov. and E. sentinelus sp. nov. (A) E. syristes fromAgua del Obispo, Guerrero, Mexico.(B) E. syristes from east of Hwy. 95, near Acahuizotla, Guerrero, Mexico. (C) E. dilatus from Municipality of Chilpancingo, Guerrero, Mexico. (D) E. maurus from Municipality of Ocuilán, Estado de México, Mexico. (E) E. albolabris from Municipality of Agua de Obispo, Guerrero, Mexico. (F) E. albolabris from Vallecitos, Guerrero, Mexico." figureDoi="http://doi.org/10.5281/zenodo.11265078" httpUri="https://zenodo.org/record/11265078/files/figure.png" pageId="14" pageNumber="15">Fig. 11C</figureCitation>
|
||
). Calls recorded at the
|
||
<typeStatus id="30118813FFB2FF9CFF0FF9C4FF0AF9A9" box="[141,189,1604,1628]" pageId="14" pageNumber="15">type</typeStatus>
|
||
locality consist of a rapid chirp, with an average duration of 110 ms, and a dominant frequency ranging between starting at 2,550 kHz and 2,780 kHz.
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB2FF9CFF0FF943FB7EFEDE" blockId="14.[141,745,1731,1947]" lastBlockId="14.[790,1394,147,299]" pageId="14" pageNumber="15">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFF0FF943FDB1F92E" bold="true" box="[141,518,1731,1755]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="28AA4D32FFB2FF9CFF0FF943FDB5F92E" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[141,514,1731,1755]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maculabialis">Eleutherodactylus maculabialis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The advertisement call of
|
||
<taxonomicName id="28AA4D32FFB2FF9CFF5CF964FE39F90E" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[222,398,1763,1787]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maculabialis">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFF5CF964FE39F90E" box="[222,398,1763,1787]" italics="true" pageId="14" pageNumber="15">E. maculabialis</emphasis>
|
||
</taxonomicName>
|
||
is a relatively slow multi-note “trill” (
|
||
<figureCitation id="77912A34FFB2FF9CFF64F883FE87F8EE" box="[230,304,1795,1819]" captionStart="Fig" captionStartId="6.[140,175,1829,1851]" captionTargetBox="[150,1368,1125,1810]" captionTargetPageId="6" captionText="Fig. 3. Oscillograms and spectrograms of the advertisement calls of Eleutherodactylus maculabialis sp. nov. and Eleutherodactylus sentinelus sp. nov. (A) Adult male E. maculabialis sp. nov. from type locality. (B) Adult male E. maculabialis sp. nov. from 1.0 km S of bridge between Nueva Delhi and El Paraíso, Municipality of Atoyac de Álvarez, Guerrero, Mexico. (C) Adult male Eleutherodactylus sentinelus sp. nov. from Puerto El Balsamo, Municipality of José Azueta, Guerrero, Mexico. (D) Adult male Eleutherodactylus sentinelus sp. nov. from type locality." figureDoi="http://doi.org/10.5281/zenodo.11265052" httpUri="https://zenodo.org/record/11265052/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||
). Calls recorded at the
|
||
<typeStatus id="30118813FFB2FF9CFDDAF884FD3FF8E9" box="[600,648,1796,1820]" pageId="14" pageNumber="15">type</typeStatus>
|
||
locality consist of slow six-note trills with a duration of 180–210 ms, and a dominant frequency starting at
|
||
<geoCoordinate id="8A9E5076FFB2FF9CFF0FF8E3FF7CF88E" box="[141,203,1891,1915]" degrees="2,600" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="55" value="2.6">2,600</geoCoordinate>
|
||
<geoCoordinate id="8A9E5076FFB2FF9CFF4EF8E3FEAFF88E" box="[204,280,1891,1915]" degrees="2,900" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="55" value="-2.9">–2,900</geoCoordinate>
|
||
kHz and rising to
|
||
<geoCoordinate id="8A9E5076FFB2FF9CFE6CF8E3FD9BF88E" box="[494,556,1891,1915]" degrees="3,050" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="55" value="3.05">3,050</geoCoordinate>
|
||
<geoCoordinate id="8A9E5076FFB2FF9CFDAEF8E3FDCEF88E" box="[556,633,1891,1915]" degrees="3,200" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="55" value="-3.2">–3,200</geoCoordinate>
|
||
kHz. The call interval is about 40–55 ms (
|
||
<figureCitation id="77912A34FFB2FF9CFE7BF803FDE4F86E" box="[505,595,1923,1947]" captionStart="Fig" captionStartId="6.[140,175,1829,1851]" captionTargetBox="[150,1368,1125,1810]" captionTargetPageId="6" captionText="Fig. 3. Oscillograms and spectrograms of the advertisement calls of Eleutherodactylus maculabialis sp. nov. and Eleutherodactylus sentinelus sp. nov. (A) Adult male E. maculabialis sp. nov. from type locality. (B) Adult male E. maculabialis sp. nov. from 1.0 km S of bridge between Nueva Delhi and El Paraíso, Municipality of Atoyac de Álvarez, Guerrero, Mexico. (C) Adult male Eleutherodactylus sentinelus sp. nov. from Puerto El Balsamo, Municipality of José Azueta, Guerrero, Mexico. (D) Adult male Eleutherodactylus sentinelus sp. nov. from type locality." figureDoi="http://doi.org/10.5281/zenodo.11265052" httpUri="https://zenodo.org/record/11265052/files/figure.png" pageId="14" pageNumber="15">Fig. 3A</figureCitation>
|
||
). At a nearby locality at
|
||
<quantity id="28529B54FFB2FF9CFC13FF13FC5AFF5E" box="[913,1005,147,171]" metricMagnitude="3" metricUnit="m" metricValue="1.3199999999999998" pageId="14" pageNumber="15" unit="m" value="1320.0">1,320 m</quantity>
|
||
asl (
|
||
<figureCitation id="77912A34FFB2FF9CFBA4FF13FBC8FF5E" box="[1062,1151,147,171]" captionStart="Fig" captionStartId="6.[140,175,1829,1851]" captionTargetBox="[150,1368,1125,1810]" captionTargetPageId="6" captionText="Fig. 3. Oscillograms and spectrograms of the advertisement calls of Eleutherodactylus maculabialis sp. nov. and Eleutherodactylus sentinelus sp. nov. (A) Adult male E. maculabialis sp. nov. from type locality. (B) Adult male E. maculabialis sp. nov. from 1.0 km S of bridge between Nueva Delhi and El Paraíso, Municipality of Atoyac de Álvarez, Guerrero, Mexico. (C) Adult male Eleutherodactylus sentinelus sp. nov. from Puerto El Balsamo, Municipality of José Azueta, Guerrero, Mexico. (D) Adult male Eleutherodactylus sentinelus sp. nov. from type locality." figureDoi="http://doi.org/10.5281/zenodo.11265052" httpUri="https://zenodo.org/record/11265052/files/figure.png" pageId="14" pageNumber="15">Fig. 3B</figureCitation>
|
||
), calls were recorded of
|
||
<specimenCount id="F9ACFD38FFB2FF9CFCBAFF34FC1BFF3E" box="[824,940,179,203]" count="2" pageId="14" pageNumber="15" type="male">two males</specimenCount>
|
||
which varied from the
|
||
<typeStatus id="30118813FFB2FF9CFB46FF34FB43FF39" box="[1220,1268,180,204]" pageId="14" pageNumber="15">type</typeStatus>
|
||
locality in consisting of five-note trills with an average duration of 230 ms and a dominant frequency starting at 3,630 kHz and reaching 4,000 kHz (
|
||
<tableCitation id="A228030AFFB2FF9CFBE7FE93FB0BFEDE" box="[1125,1212,275,299]" captionStart="Table 4" captionStartId="17.[141,197,142,163]" captionTargetPageId="17" captionText="Table 4. Advertisement call data of Elutherodactylus maculabialis sp. nov. and Eleutherodactylus sentinelus sp. nov." httpUri="http://table.plazi.org/id/BBD56639FFADFF83FF0FFF0EFB47FF56" pageId="14" pageNumber="15" tableUuid="BBD56639FFADFF83FF0FFF0EFB47FF56">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB2FF9CFC94FED3FAC7FD3E" blockId="14.[790,1395,339,715]" pageId="14" pageNumber="15">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFC94FED3FBE2FE9E" bold="true" box="[790,1109,339,363]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="28AA4D32FFB2FF9CFC94FED3FBE6FE9E" authorityName="Hedges" authorityYear="1989" box="[790,1105,339,363]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maurus">Eleutherodactylus maurus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The advertisement call of
|
||
<taxonomicName id="28AA4D32FFB2FF9CFCBCFEF4FC0FFE7E" authorityName="Hedges" authorityYear="1989" box="[830,952,372,395]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maurus">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFCBCFEF4FC0FFE7E" box="[830,952,372,395]" italics="true" pageId="14" pageNumber="15">E. maurus</emphasis>
|
||
</taxonomicName>
|
||
is a single, drawn-out “chirp” best described as a “pipe.” Calls recorded near Ocuilan, in
|
||
<collectingRegion id="2D6EF853FFB2FF9CFC94FE33FC51FE3E" box="[790,998,435,459]" country="Mexico" name="Mexico" pageId="14" pageNumber="15">Estado de México</collectingRegion>
|
||
, consisted of a single chirp with a duration of 205 ms, and a dominant frequency starting at 2,760 kHz and rising to 3,150 kHz. The amplitude was strongest at the middle of a call (
|
||
<figureCitation id="77912A34FFB2FF9CFB47FD93FA99FDDE" box="[1221,1326,531,555]" captionStart="Fig" captionStartId="20.[141,176,1334,1356]" captionTargetBox="[181,1389,156,1293]" captionTargetId="figure-331@20.[181,1389,113,1309]" captionTargetPageId="20" captionText="Fig. 11. Oscillograms and spectrograms of the advertisement calls of adult males of Eleutherodactylus species related to E. maculabialis sp. nov. and E. sentinelus sp. nov. (A) E. syristes fromAgua del Obispo, Guerrero, Mexico.(B) E. syristes from east of Hwy. 95, near Acahuizotla, Guerrero, Mexico. (C) E. dilatus from Municipality of Chilpancingo, Guerrero, Mexico. (D) E. maurus from Municipality of Ocuilán, Estado de México, Mexico. (E) E. albolabris from Municipality of Agua de Obispo, Guerrero, Mexico. (F) E. albolabris from Vallecitos, Guerrero, Mexico." figureDoi="http://doi.org/10.5281/zenodo.11265078" httpUri="https://zenodo.org/record/11265078/files/figure.png" pageId="14" pageNumber="15">Fig. 11D</figureCitation>
|
||
). At a locality near Taxco,
|
||
<collectingRegion id="2D6EF853FFB2FF9CFC78FDB3FBD6FDBE" box="[1018,1121,563,587]" country="Mexico" name="Guerrero" pageId="14" pageNumber="15">Guerrero</collectingRegion>
|
||
, a call of
|
||
<taxonomicName id="28AA4D32FFB2FF9CFB4CFDB4FAF6FDBE" authorityName="Hedges" authorityYear="1989" box="[1230,1345,564,587]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="maurus">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFB4CFDB4FAF6FDBE" box="[1230,1345,564,587]" italics="true" pageId="14" pageNumber="15">E. maurus</emphasis>
|
||
</taxonomicName>
|
||
was recorded which consisted of a single, drawn-out chirp. However, the call was longer, averaging 275 ms and the amplitude was distributed in three sub-pulses. This indicates a further need to investigate this population.
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB2FF9CFC94FD73FC0FFBBE" blockId="14.[790,1394,755,1099]" pageId="14" pageNumber="15">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFC94FD73FBDDFCFE" bold="true" box="[790,1130,755,779]" italics="true" pageId="14" pageNumber="15">
|
||
<taxonomicName id="28AA4D32FFB2FF9CFC94FD73FBD0FCFE" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[790,1127,755,779]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="sentinelus">Eleutherodactylus sentinelus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The advertisement call of
|
||
<taxonomicName id="28AA4D32FFB2FF9CFCBAFC94FC7FFCDE" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[824,968,787,811]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="sentinelus">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFCBAFC94FC7FFCDE" box="[824,968,787,811]" italics="true" pageId="14" pageNumber="15">E. sentinelus</emphasis>
|
||
</taxonomicName>
|
||
is a single high-pitched “chirp” best described as a “peep” (
|
||
<figureCitation id="77912A34FFB2FF9CFBA1FCB3FBDDFCBE" box="[1059,1130,819,843]" captionStart="Fig" captionStartId="6.[140,175,1829,1851]" captionTargetBox="[150,1368,1125,1810]" captionTargetPageId="6" captionText="Fig. 3. Oscillograms and spectrograms of the advertisement calls of Eleutherodactylus maculabialis sp. nov. and Eleutherodactylus sentinelus sp. nov. (A) Adult male E. maculabialis sp. nov. from type locality. (B) Adult male E. maculabialis sp. nov. from 1.0 km S of bridge between Nueva Delhi and El Paraíso, Municipality of Atoyac de Álvarez, Guerrero, Mexico. (C) Adult male Eleutherodactylus sentinelus sp. nov. from Puerto El Balsamo, Municipality of José Azueta, Guerrero, Mexico. (D) Adult male Eleutherodactylus sentinelus sp. nov. from type locality." figureDoi="http://doi.org/10.5281/zenodo.11265052" httpUri="https://zenodo.org/record/11265052/files/figure.png" pageId="14" pageNumber="15">Fig. 3</figureCitation>
|
||
). Calls recorded at the
|
||
<typeStatus id="30118813FFB2FF9CFC94FCD4FCF1FC99" box="[790,838,852,876]" pageId="14" pageNumber="15">type</typeStatus>
|
||
locality consist of a rapid chirp, with an average duration of 160 ms, and a dominant frequency starting at 2,560 kHz and rising to 3,100 kHz (
|
||
<figureCitation id="77912A34FFB2FF9CFB48FC13FA93FC5E" box="[1226,1316,915,939]" captionStart="Fig" captionStartId="6.[140,175,1829,1851]" captionTargetBox="[150,1368,1125,1810]" captionTargetPageId="6" captionText="Fig. 3. Oscillograms and spectrograms of the advertisement calls of Eleutherodactylus maculabialis sp. nov. and Eleutherodactylus sentinelus sp. nov. (A) Adult male E. maculabialis sp. nov. from type locality. (B) Adult male E. maculabialis sp. nov. from 1.0 km S of bridge between Nueva Delhi and El Paraíso, Municipality of Atoyac de Álvarez, Guerrero, Mexico. (C) Adult male Eleutherodactylus sentinelus sp. nov. from Puerto El Balsamo, Municipality of José Azueta, Guerrero, Mexico. (D) Adult male Eleutherodactylus sentinelus sp. nov. from type locality." figureDoi="http://doi.org/10.5281/zenodo.11265052" httpUri="https://zenodo.org/record/11265052/files/figure.png" pageId="14" pageNumber="15">Fig. 3D</figureCitation>
|
||
). Calls recorded above the
|
||
<typeStatus id="30118813FFB2FF9CFC7BFC34FB9EFC39" box="[1017,1065,948,972]" pageId="14" pageNumber="15">type</typeStatus>
|
||
locality at
|
||
<quantity id="28529B54FFB2FF9CFB2CFC33FABCFC3E" box="[1198,1291,947,971]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="15" unit="m" value="1900.0">1,900 m</quantity>
|
||
asl (
|
||
<figureCitation id="77912A34FFB2FF9CFAC7FC33FC80FC1E" captionStart="Fig" captionStartId="6.[140,175,1829,1851]" captionTargetBox="[150,1368,1125,1810]" captionTargetPageId="6" captionText="Fig. 3. Oscillograms and spectrograms of the advertisement calls of Eleutherodactylus maculabialis sp. nov. and Eleutherodactylus sentinelus sp. nov. (A) Adult male E. maculabialis sp. nov. from type locality. (B) Adult male E. maculabialis sp. nov. from 1.0 km S of bridge between Nueva Delhi and El Paraíso, Municipality of Atoyac de Álvarez, Guerrero, Mexico. (C) Adult male Eleutherodactylus sentinelus sp. nov. from Puerto El Balsamo, Municipality of José Azueta, Guerrero, Mexico. (D) Adult male Eleutherodactylus sentinelus sp. nov. from type locality." figureDoi="http://doi.org/10.5281/zenodo.11265052" httpUri="https://zenodo.org/record/11265052/files/figure.png" pageId="14" pageNumber="15">Fig. 3C</figureCitation>
|
||
) are slightly shorter and of higher frequency, with an average duration of 120 ms, and a dominant frequency starting at 2,700 kHz and finishing at 3,300 kHz (
|
||
<tableCitation id="A228030AFFB2FF9CFCD1FBB3FC1CFBBE" box="[851,939,1075,1099]" captionStart="Table 4" captionStartId="17.[141,197,142,163]" captionTargetPageId="17" captionText="Table 4. Advertisement call data of Elutherodactylus maculabialis sp. nov. and Eleutherodactylus sentinelus sp. nov." httpUri="http://table.plazi.org/id/BBD56639FFADFF83FF0FFF0EFB47FF56" pageId="14" pageNumber="15" tableUuid="BBD56639FFADFF83FF0FFF0EFB47FF56">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB2FF9CFC94FBEDFC71FA3C" blockId="14.[790,1394,1133,1481]" pageId="14" pageNumber="15">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFC94FBEDFBE4FB73" bold="true" box="[790,1107,1133,1158]" pageId="14" pageNumber="15">
|
||
<taxonomicName id="28AA4D32FFB2FF9CFC94FBEDFBFCFB73" baseAuthorityName="Hoyt" baseAuthorityYear="1965" box="[790,1099,1133,1158]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="syristes">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFC94FBEDFBFCFB73" bold="true" box="[790,1099,1133,1158]" italics="true" pageId="14" pageNumber="15">Eleutherodactylus syristes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<collectingCounty id="06744E3DFFB2FF9CFBE1FBEEFB39FB73" box="[1123,1166,1134,1158]" pageId="14" pageNumber="15">The</collectingCounty>
|
||
advertisement call of
|
||
<taxonomicName id="28AA4D32FFB2FF9CFCB4FB0CFC1DFB56" baseAuthorityName="Hoyt" baseAuthorityYear="1965" box="[822,938,1164,1187]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="syristes">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFCB4FB0CFC1DFB56" box="[822,938,1164,1187]" italics="true" pageId="14" pageNumber="15">E. syristes</emphasis>
|
||
</taxonomicName>
|
||
is a rapid but long multiple-pulse “trill” (
|
||
<figureCitation id="77912A34FFB2FF9CFC9DFB29FC12FB34" box="[799,933,1193,1217]" captionStart="Fig" captionStartId="20.[141,176,1334,1356]" captionTargetBox="[181,1389,156,1293]" captionTargetId="figure-331@20.[181,1389,113,1309]" captionTargetPageId="20" captionText="Fig. 11. Oscillograms and spectrograms of the advertisement calls of adult males of Eleutherodactylus species related to E. maculabialis sp. nov. and E. sentinelus sp. nov. (A) E. syristes fromAgua del Obispo, Guerrero, Mexico.(B) E. syristes from east of Hwy. 95, near Acahuizotla, Guerrero, Mexico. (C) E. dilatus from Municipality of Chilpancingo, Guerrero, Mexico. (D) E. maurus from Municipality of Ocuilán, Estado de México, Mexico. (E) E. albolabris from Municipality of Agua de Obispo, Guerrero, Mexico. (F) E. albolabris from Vallecitos, Guerrero, Mexico." figureDoi="http://doi.org/10.5281/zenodo.11265078" httpUri="https://zenodo.org/record/11265078/files/figure.png" pageId="14" pageNumber="15">Fig. 11A–B</figureCitation>
|
||
).
|
||
<collectingMunicipality id="0F71ACCBFFB2FF9CFC3AFB29FC46FB34" box="[952,1009,1193,1217]" pageId="14" pageNumber="15">Calls</collectingMunicipality>
|
||
recorded at two different localities in
|
||
<collectingRegion id="2D6EF853FFB2FF9CFCB5FB46FC2CFB2B" box="[823,923,1222,1246]" country="Mexico" name="Guerrero" pageId="14" pageNumber="15">Guerrero</collectingRegion>
|
||
consisted of 49–60 pulses, about 10 ms apart, with a total duration of the call averaging 500– 560 ms.
|
||
<location id="EA75606AFFB2FF9CFCFBFA81FC13FAEC" LSID="urn:lsid:plazi:treatment:670387A7FFB1FF9DFF0FF8B2FC55F9BC:EA75606AFFB2FF9CFCFBFA81FC13FAEC" box="[889,932,1281,1305]" county="The" latitude="3.0" longLatPrecision="78" longitude="-3.39" municipality="Calls" name="The" pageId="14" pageNumber="15" stateProvince="Guerrero">The</location>
|
||
dominant frequency at
|
||
<location id="EA75606AFFB2FF9CFB31FA81FAD9FAEC" LSID="urn:lsid:plazi:treatment:670387A7FFB1FF9DFF0FF8B2FC55F9BC:EA75606AFFB2FF9CFB31FA81FAD9FAEC" box="[1203,1390,1281,1305]" county="The" latitude="3.0" longLatPrecision="78" longitude="-3.39" municipality="Calls" name="Agua de Obispo" pageId="14" pageNumber="15" stateProvince="Guerrero">
|
||
Agua
|
||
<collectorName id="425F5367FFB2FF9CFB7AFA81FAD9FAEC" box="[1272,1390,1281,1305]" pageId="14" pageNumber="15">de Obispo</collectorName>
|
||
</location>
|
||
,
|
||
<collectingRegion id="2D6EF853FFB2FF9CFC94FA9EFCCAFAC3" box="[790,893,1310,1334]" country="Mexico" name="Guerrero" pageId="14" pageNumber="15">Guerrero</collectingRegion>
|
||
, was between
|
||
<geoCoordinate id="8A9E5076FFB2FF9CFBBFFA9EFBCCFAC3" box="[1085,1147,1310,1334]" degrees="2,700" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="55" value="2.7">2,700</geoCoordinate>
|
||
<geoCoordinate id="8A9E5076FFB2FF9CFBF9FA9EFB70FAC3" box="[1147,1223,1310,1334]" degrees="3,170" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="55" value="-3.17">–3,170</geoCoordinate>
|
||
kHz, starting lower, peaking, and then lowering again (
|
||
<figureCitation id="77912A34FFB2FF9CFB7EFABBFAD2FAA6" box="[1276,1381,1339,1363]" captionStart="Fig" captionStartId="20.[141,176,1334,1356]" captionTargetBox="[181,1389,156,1293]" captionTargetId="figure-331@20.[181,1389,113,1309]" captionTargetPageId="20" captionText="Fig. 11. Oscillograms and spectrograms of the advertisement calls of adult males of Eleutherodactylus species related to E. maculabialis sp. nov. and E. sentinelus sp. nov. (A) E. syristes fromAgua del Obispo, Guerrero, Mexico.(B) E. syristes from east of Hwy. 95, near Acahuizotla, Guerrero, Mexico. (C) E. dilatus from Municipality of Chilpancingo, Guerrero, Mexico. (D) E. maurus from Municipality of Ocuilán, Estado de México, Mexico. (E) E. albolabris from Municipality of Agua de Obispo, Guerrero, Mexico. (F) E. albolabris from Vallecitos, Guerrero, Mexico." figureDoi="http://doi.org/10.5281/zenodo.11265078" httpUri="https://zenodo.org/record/11265078/files/figure.png" pageId="14" pageNumber="15">Fig. 11A</figureCitation>
|
||
). At a second locality, slightly lower than
|
||
<location id="EA75606AFFB2FF9CFA8CFAD9FCDEFA7B" LSID="urn:lsid:plazi:treatment:670387A7FFB1FF9DFF0FF8B2FC55F9BC:EA75606AFFB2FF9CFA8CFAD9FCDEFA7B" county="The" latitude="3.0" longLatPrecision="78" longitude="-3.39" municipality="Calls" name="Agua de Obispo" pageId="14" pageNumber="15" stateProvince="Guerrero">
|
||
Agua
|
||
<collectorName id="425F5367FFB2FF9CFADBFAD9FCDEFA7B" pageId="14" pageNumber="15">de Obispo</collectorName>
|
||
</location>
|
||
, east of
|
||
<location id="EA75606AFFB2FF9CFC49FAF6FB87FA7B" LSID="urn:lsid:plazi:treatment:670387A7FFB1FF9DFF0FF8B2FC55F9BC:EA75606AFFB2FF9CFC49FAF6FB87FA7B" box="[971,1072,1398,1422]" county="The" latitude="3.0" longLatPrecision="78" longitude="-3.39" municipality="Calls" name="Highway" pageId="14" pageNumber="15" stateProvince="Guerrero">Highway</location>
|
||
95, the calls tended to have a dominant frequency ranging between
|
||
<geoCoordinate id="8A9E5076FFB2FF9CFB6AFA13FA91FA5E" box="[1256,1318,1427,1451]" degrees="3,000" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="55" value="3.0">3,000</geoCoordinate>
|
||
<geoCoordinate id="8A9E5076FFB2FF9CFAA4FA13FAC5FA5E" box="[1318,1394,1427,1451]" degrees="3,390" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="55" value="-3.39">–3,390</geoCoordinate>
|
||
kHz (
|
||
<figureCitation id="77912A34FFB2FF9CFCD6FA31FC0FFA3C" box="[852,952,1457,1481]" captionStart="Fig" captionStartId="20.[141,176,1334,1356]" captionTargetBox="[181,1389,156,1293]" captionTargetId="figure-331@20.[181,1389,113,1309]" captionTargetPageId="20" captionText="Fig. 11. Oscillograms and spectrograms of the advertisement calls of adult males of Eleutherodactylus species related to E. maculabialis sp. nov. and E. sentinelus sp. nov. (A) E. syristes fromAgua del Obispo, Guerrero, Mexico.(B) E. syristes from east of Hwy. 95, near Acahuizotla, Guerrero, Mexico. (C) E. dilatus from Municipality of Chilpancingo, Guerrero, Mexico. (D) E. maurus from Municipality of Ocuilán, Estado de México, Mexico. (E) E. albolabris from Municipality of Agua de Obispo, Guerrero, Mexico. (F) E. albolabris from Vallecitos, Guerrero, Mexico." figureDoi="http://doi.org/10.5281/zenodo.11265078" httpUri="https://zenodo.org/record/11265078/files/figure.png" pageId="14" pageNumber="15">Fig. 11B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB2FF9CFC94FA6BFB1EF86B" blockId="14.[790,1394,1515,1950]" pageId="14" pageNumber="15">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFC94FA6BFBCDF9F6" bold="true" box="[790,1146,1515,1539]" pageId="14" pageNumber="15">
|
||
<taxonomicName id="28AA4D32FFB2FF9CFC94FA6BFBC5F9F6" baseAuthorityName="Taylor" baseAuthorityYear="1943" box="[790,1138,1515,1539]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="albolabris">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFC94FA6BFBC5F9F6" bold="true" box="[790,1138,1515,1539]" italics="true" pageId="14" pageNumber="15">Eleutherodactylus albolabris</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The advertisement call of
|
||
<taxonomicName id="28AA4D32FFB2FF9CFCE5F989FC4DF9D4" baseAuthorityName="Taylor" baseAuthorityYear="1943" box="[871,1018,1545,1569]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="albolabris">
|
||
<emphasis id="DDDEEAA3FFB2FF9CFCE5F989FC4DF9D4" box="[871,1018,1545,1569]" italics="true" pageId="14" pageNumber="15">E. albolabris</emphasis>
|
||
</taxonomicName>
|
||
is a single, multi-pulsed whistle, with the call continuous amongst the pulses and with a duration of approximately 150–250 ms (
|
||
<figureCitation id="77912A34FFB2FF9CFA82F9C3FAD2F9AE" box="[1280,1381,1603,1627]" captionStart="Fig" captionStartId="20.[141,176,1334,1356]" captionTargetBox="[181,1389,156,1293]" captionTargetId="figure-331@20.[181,1389,113,1309]" captionTargetPageId="20" captionText="Fig. 11. Oscillograms and spectrograms of the advertisement calls of adult males of Eleutherodactylus species related to E. maculabialis sp. nov. and E. sentinelus sp. nov. (A) E. syristes fromAgua del Obispo, Guerrero, Mexico.(B) E. syristes from east of Hwy. 95, near Acahuizotla, Guerrero, Mexico. (C) E. dilatus from Municipality of Chilpancingo, Guerrero, Mexico. (D) E. maurus from Municipality of Ocuilán, Estado de México, Mexico. (E) E. albolabris from Municipality of Agua de Obispo, Guerrero, Mexico. (F) E. albolabris from Vallecitos, Guerrero, Mexico." figureDoi="http://doi.org/10.5281/zenodo.11265078" httpUri="https://zenodo.org/record/11265078/files/figure.png" pageId="14" pageNumber="15">Fig. 11F</figureCitation>
|
||
). Calls recorded at the
|
||
<typeStatus id="30118813FFB2FF9CFB88F9E1FB8DF98C" box="[1034,1082,1633,1657]" pageId="14" pageNumber="15">type</typeStatus>
|
||
locality at
|
||
<collectingMunicipality id="0F71ACCBFFB2FF9CFB39F9E1FAC5F98C" box="[1211,1394,1633,1657]" pageId="14" pageNumber="15">
|
||
Agua
|
||
<collectorName id="425F5367FFB2FF9CFA82F9E1FAC5F98C" box="[1280,1394,1633,1657]" pageId="14" pageNumber="15">de Obispo</collectorName>
|
||
</collectingMunicipality>
|
||
had a duration of 160 ms, and started at a dominant frequency of 2,840 kHz rising to 3,015 kHz. Calls varied between 7–10 continuous pulses. At a second locality, near Vallecitos in the
|
||
<location id="EA75606AFFB2FF9CFB01F956FCD0F8FE" LSID="urn:lsid:plazi:treatment:670387A7FFB1FF9DFF0FF8B2FC55F9BC:EA75606AFFB2FF9CFB01F956FCD0F8FE" municipality="Agua de Obispo" name="Municipality of Jose Azueta" pageId="14" pageNumber="15" stateProvince="Guerrero">Municipality of José Azueta</location>
|
||
,
|
||
<collectingRegion id="2D6EF853FFB2FF9CFCF7F973FC6BF8FE" box="[885,988,1779,1803]" country="Mexico" name="Guerrero" pageId="14" pageNumber="15">Guerrero</collectingRegion>
|
||
, the calls had a duration of 240 ms, and started at a dominant frequency of 2,740 kHz which rose to 2,915 kHz. Calls started with a multi-pulsed trill and then ended with several distinct pulses. This population’s call is best described as a combination of a “trill” with a “whistle” (
|
||
<figureCitation id="77912A34FFB2FF9CFBBAF806FB2BF86B" box="[1080,1180,1926,1950]" captionStart="Fig" captionStartId="20.[141,176,1334,1356]" captionTargetBox="[181,1389,156,1293]" captionTargetId="figure-331@20.[181,1389,113,1309]" captionTargetPageId="20" captionText="Fig. 11. Oscillograms and spectrograms of the advertisement calls of adult males of Eleutherodactylus species related to E. maculabialis sp. nov. and E. sentinelus sp. nov. (A) E. syristes fromAgua del Obispo, Guerrero, Mexico.(B) E. syristes from east of Hwy. 95, near Acahuizotla, Guerrero, Mexico. (C) E. dilatus from Municipality of Chilpancingo, Guerrero, Mexico. (D) E. maurus from Municipality of Ocuilán, Estado de México, Mexico. (E) E. albolabris from Municipality of Agua de Obispo, Guerrero, Mexico. (F) E. albolabris from Vallecitos, Guerrero, Mexico." figureDoi="http://doi.org/10.5281/zenodo.11265078" httpUri="https://zenodo.org/record/11265078/files/figure.png" pageId="14" pageNumber="15">Fig. 11E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="BBD56639FFB3FF9DFF0CFF0EFF45FF4B" ID-Table-UUID="BBD56639FFB3FF9DFF0CFF0EFF45FF4B" httpUri="http://table.plazi.org/id/BBD56639FFB3FF9DFF0CFF0EFF45FF4B" pageId="15" pageNumber="16" startId="15.[142,198,142,163]" targetBox="[152,1377,210,1184]" targetIsTable="true" targetPageId="15" targetType="table">
|
||
<paragraph id="EF1536B1FFB3FF9DFF0CFF0EFF45FF4B" blockId="15.[142,1395,142,190]" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF0CFF0EFF69FF56" bold="true" box="[142,222,142,163]" pageId="15" pageNumber="16">Table 2.</emphasis>
|
||
Morphological measurements of
|
||
<taxonomicName id="28AA4D32FFB3FF9DFDA2FF0EFC83FF56" authority="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones, 2021" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[544,820,142,163]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="sentinelus" status="sp. nov.">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFDA2FF0EFC83FF56" box="[544,820,142,163]" italics="true" pageId="15" pageNumber="16">Eleutherodactylus sentinelus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="DDDEEAA3FFB3FF9DFCBBFF0EFC32FF56" bold="true" box="[825,901,142,163]" pageId="15" pageNumber="16">
|
||
<taxonomicNameLabel id="C6ED57D8FFB3FF9DFCBBFF0EFC32FF56" box="[825,901,142,163]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
specimens. See text for definitions of measurement acronyms.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="EF1536B1FFB3FF9DFE13FF52FAFEFB6A" pageId="15" pageNumber="16">
|
||
<table id="9DAAC411FFB3006DFF1AFF52FAD6FB55" box="[152,1377,210,1184]" gridcols="9" gridrows="24" pageId="15" pageNumber="16">
|
||
<tr id="519A34F3FFB3006DFF1AFF52FAD6FF1D" box="[152,1377,210,232]" gridrow="0" pageId="15" pageNumber="16" rowspan-0="1">
|
||
<th id="124B5D8FFFB3006DFE13FF52FE5CFF1D" box="[401,491,210,232]" gridcol="1" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFE13FF52FE5CFF1D" bold="true" box="[401,491,210,232]" pageId="15" pageNumber="16">CIG-333</emphasis>
|
||
</th>
|
||
<th id="124B5D8FFFB3006DFD8EFF52FDD2FF1D" box="[524,613,210,232]" gridcol="2" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFD8EFF52FDD2FF1D" bold="true" box="[524,613,210,232]" pageId="15" pageNumber="16">CIG-334</emphasis>
|
||
</th>
|
||
<th id="124B5D8FFFB3006DFD08FF52FD54FF1D" box="[650,739,210,232]" gridcol="3" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFD08FF52FD54FF1D" bold="true" box="[650,739,210,232]" pageId="15" pageNumber="16">CIG-335</emphasis>
|
||
</th>
|
||
<th id="124B5D8FFFB3006DFC83FF52FCEDFF1D" box="[769,858,210,232]" gridcol="4" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFC83FF52FCEDFF1D" bold="true" box="[769,858,210,232]" pageId="15" pageNumber="16">CIG-907</emphasis>
|
||
</th>
|
||
<th id="124B5D8FFFB3006DFCFBFF52FC65FF1D" box="[889,978,210,232]" gridcol="5" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFCFBFF52FC65FF1D" bold="true" box="[889,978,210,232]" pageId="15" pageNumber="16">CIG-908</emphasis>
|
||
</th>
|
||
<th id="124B5D8FFFB3006DFC7EFF52FBE2FF1D" box="[1020,1109,210,232]" gridcol="6" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFC7EFF52FBE2FF1D" bold="true" box="[1020,1109,210,232]" pageId="15" pageNumber="16">CIG-909</emphasis>
|
||
</th>
|
||
<th id="124B5D8FFFB3006DFB01FF52FB6BFF1D" box="[1155,1244,210,232]" gridcol="7" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFB01FF52FB6BFF1D" bold="true" box="[1155,1244,210,232]" pageId="15" pageNumber="16">CIG-910</emphasis>
|
||
</th>
|
||
<th id="124B5D8FFFB3006DFA8AFF52FAD6FF1D" box="[1288,1377,210,232]" gridcol="8" gridrow="0" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFA8AFF52FAD6FF1D" bold="true" box="[1288,1377,210,232]" pageId="15" pageNumber="16">CIG-913</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFF7AFAD6FEE7" box="[152,1377,250,274]" gridrow="1" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFF7AFEA4FEE7" box="[152,275,250,274]" gridcol="0" gridrow="1" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFF7AFF70FEE5" bold="true" box="[152,199,250,272]" pageId="15" pageNumber="16">SVL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FF7AFE5CFEE7" box="[401,491,250,274]" gridcol="1" gridrow="1" pageId="15" pageNumber="16">24.37</td>
|
||
<td id="124B5D8FFFB3006DFD8EFF7AFDD2FEE7" box="[524,613,250,274]" gridcol="2" gridrow="1" pageId="15" pageNumber="16">24.48</td>
|
||
<td id="124B5D8FFFB3006DFD08FF7AFD54FEE7" box="[650,739,250,274]" gridcol="3" gridrow="1" pageId="15" pageNumber="16">25.30</td>
|
||
<td id="124B5D8FFFB3006DFC83FF7AFCEDFEE7" box="[769,858,250,274]" gridcol="4" gridrow="1" pageId="15" pageNumber="16">25.1</td>
|
||
<td id="124B5D8FFFB3006DFCFBFF7AFC65FEE7" box="[889,978,250,274]" gridcol="5" gridrow="1" pageId="15" pageNumber="16">23.31</td>
|
||
<td id="124B5D8FFFB3006DFC7EFF7AFBE2FEE7" box="[1020,1109,250,274]" gridcol="6" gridrow="1" pageId="15" pageNumber="16">23.41</td>
|
||
<td id="124B5D8FFFB3006DFB01FF7AFB6BFEE7" box="[1155,1244,250,274]" gridcol="7" gridrow="1" pageId="15" pageNumber="16">23.25</td>
|
||
<td id="124B5D8FFFB3006DFA8AFF7AFAD6FEE7" box="[1288,1377,250,274]" gridcol="8" gridrow="1" pageId="15" pageNumber="16">24.54</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFEA4FAD6FEC9" box="[152,1377,292,316]" gridrow="2" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFEA4FEA4FEC9" box="[152,275,292,316]" gridcol="0" gridrow="2" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFEA4FF0CFECC" bold="true" box="[152,187,292,313]" pageId="15" pageNumber="16">HL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FEA4FE5CFEC9" box="[401,491,292,316]" gridcol="1" gridrow="2" pageId="15" pageNumber="16">8.58</td>
|
||
<td id="124B5D8FFFB3006DFD8EFEA4FDD2FEC9" box="[524,613,292,316]" gridcol="2" gridrow="2" pageId="15" pageNumber="16">8.70</td>
|
||
<td id="124B5D8FFFB3006DFD08FEA4FD54FEC9" box="[650,739,292,316]" gridcol="3" gridrow="2" pageId="15" pageNumber="16">8.67</td>
|
||
<td id="124B5D8FFFB3006DFC83FEA4FCEDFEC9" box="[769,858,292,316]" gridcol="4" gridrow="2" pageId="15" pageNumber="16">7.88</td>
|
||
<td id="124B5D8FFFB3006DFCFBFEA4FC65FEC9" box="[889,978,292,316]" gridcol="5" gridrow="2" pageId="15" pageNumber="16">7.1</td>
|
||
<td id="124B5D8FFFB3006DFC7EFEA4FBE2FEC9" box="[1020,1109,292,316]" gridcol="6" gridrow="2" pageId="15" pageNumber="16">7.15</td>
|
||
<td id="124B5D8FFFB3006DFB01FEA4FB6BFEC9" box="[1155,1244,292,316]" gridcol="7" gridrow="2" pageId="15" pageNumber="16">7.1</td>
|
||
<td id="124B5D8FFFB3006DFA8AFEA4FAD6FEC9" box="[1288,1377,292,316]" gridcol="8" gridrow="2" pageId="15" pageNumber="16">8.03</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFECDFAD6FE96" box="[152,1377,333,355]" gridrow="3" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFECDFEA4FE96" box="[152,275,333,355]" gridcol="0" gridrow="3" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFECDFF77FE97" bold="true" box="[152,192,333,354]" pageId="15" pageNumber="16">TW</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FECDFE5CFE96" box="[401,491,333,355]" gridcol="1" gridrow="3" pageId="15" pageNumber="16">0.74</td>
|
||
<td id="124B5D8FFFB3006DFD8EFECDFDD2FE96" box="[524,613,333,355]" gridcol="2" gridrow="3" pageId="15" pageNumber="16">0.76</td>
|
||
<td id="124B5D8FFFB3006DFD08FECDFD54FE96" box="[650,739,333,355]" gridcol="3" gridrow="3" pageId="15" pageNumber="16">0.80</td>
|
||
<td id="124B5D8FFFB3006DFC83FECDFCEDFE96" box="[769,858,333,355]" gridcol="4" gridrow="3" pageId="15" pageNumber="16">0.76</td>
|
||
<td id="124B5D8FFFB3006DFCFBFECDFC65FE96" box="[889,978,333,355]" gridcol="5" gridrow="3" pageId="15" pageNumber="16">0.74</td>
|
||
<td id="124B5D8FFFB3006DFC7EFECDFBE2FE96" box="[1020,1109,333,355]" gridcol="6" gridrow="3" pageId="15" pageNumber="16">0.74</td>
|
||
<td id="124B5D8FFFB3006DFB01FECDFB6BFE96" box="[1155,1244,333,355]" gridcol="7" gridrow="3" pageId="15" pageNumber="16">0.74</td>
|
||
<td id="124B5D8FFFB3006DFA8AFECDFAD6FE96" box="[1288,1377,333,355]" gridcol="8" gridrow="3" pageId="15" pageNumber="16">0.81</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFEF5FAD6FE78" box="[152,1377,373,397]" gridrow="4" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFEF5FEA4FE78" box="[152,275,373,397]" gridcol="0" gridrow="4" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFEF5FF0EFE7F" bold="true" box="[152,185,373,394]" pageId="15" pageNumber="16">ED</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FEF5FE5CFE78" box="[401,491,373,397]" gridcol="1" gridrow="4" pageId="15" pageNumber="16">2.63</td>
|
||
<td id="124B5D8FFFB3006DFD8EFEF5FDD2FE78" box="[524,613,373,397]" gridcol="2" gridrow="4" pageId="15" pageNumber="16">2.68</td>
|
||
<td id="124B5D8FFFB3006DFD08FEF5FD54FE78" box="[650,739,373,397]" gridcol="3" gridrow="4" pageId="15" pageNumber="16">2.77</td>
|
||
<td id="124B5D8FFFB3006DFC83FEF5FCEDFE78" box="[769,858,373,397]" gridcol="4" gridrow="4" pageId="15" pageNumber="16">2.67</td>
|
||
<td id="124B5D8FFFB3006DFCFBFEF5FC65FE78" box="[889,978,373,397]" gridcol="5" gridrow="4" pageId="15" pageNumber="16">2.77</td>
|
||
<td id="124B5D8FFFB3006DFC7EFEF5FBE2FE78" box="[1020,1109,373,397]" gridcol="6" gridrow="4" pageId="15" pageNumber="16">2.8</td>
|
||
<td id="124B5D8FFFB3006DFB01FEF5FB6BFE78" box="[1155,1244,373,397]" gridcol="7" gridrow="4" pageId="15" pageNumber="16">2.73</td>
|
||
<td id="124B5D8FFFB3006DFA8AFEF5FAD6FE78" box="[1288,1377,373,397]" gridcol="8" gridrow="4" pageId="15" pageNumber="16">2.85</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFE1FFAD6FE4D" box="[152,1377,415,440]" gridrow="5" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFE1FFEA4FE4D" box="[152,275,415,440]" gridcol="0" gridrow="5" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFE1FFF70FE41" bold="true" box="[152,199,415,436]" pageId="15" pageNumber="16">ElW</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FE1FFE5CFE4D" box="[401,491,415,440]" gridcol="1" gridrow="5" pageId="15" pageNumber="16">1.55</td>
|
||
<td id="124B5D8FFFB3006DFD8EFE1FFDD2FE4D" box="[524,613,415,440]" gridcol="2" gridrow="5" pageId="15" pageNumber="16">1.54</td>
|
||
<td id="124B5D8FFFB3006DFD08FE1FFD54FE4D" box="[650,739,415,440]" gridcol="3" gridrow="5" pageId="15" pageNumber="16">1.6</td>
|
||
<td id="124B5D8FFFB3006DFC83FE1FFCEDFE4D" box="[769,858,415,440]" gridcol="4" gridrow="5" pageId="15" pageNumber="16">1.58</td>
|
||
<td id="124B5D8FFFB3006DFCFBFE1FFC65FE4D" box="[889,978,415,440]" gridcol="5" gridrow="5" pageId="15" pageNumber="16">1.53</td>
|
||
<td id="124B5D8FFFB3006DFC7EFE1FFBE2FE4D" box="[1020,1109,415,440]" gridcol="6" gridrow="5" pageId="15" pageNumber="16">1.54</td>
|
||
<td id="124B5D8FFFB3006DFB01FE1FFB6BFE4D" box="[1155,1244,415,440]" gridcol="7" gridrow="5" pageId="15" pageNumber="16">1.54</td>
|
||
<td id="124B5D8FFFB3006DFA8AFE1FFAD6FE4D" box="[1288,1377,415,440]" gridcol="8" gridrow="5" pageId="15" pageNumber="16">1.6</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFE49FAD6FE17" box="[152,1377,457,482]" gridrow="6" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFE49FEA4FE17" box="[152,275,457,482]" gridcol="0" gridrow="6" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFE49FF72FE2A" bold="true" box="[152,197,457,479]" pageId="15" pageNumber="16">IOD</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FE49FE5CFE17" box="[401,491,457,482]" gridcol="1" gridrow="6" pageId="15" pageNumber="16">4.80</td>
|
||
<td id="124B5D8FFFB3006DFD8EFE49FDD2FE17" box="[524,613,457,482]" gridcol="2" gridrow="6" pageId="15" pageNumber="16">4.83</td>
|
||
<td id="124B5D8FFFB3006DFD08FE49FD54FE17" box="[650,739,457,482]" gridcol="3" gridrow="6" pageId="15" pageNumber="16">4.87</td>
|
||
<td id="124B5D8FFFB3006DFC83FE49FCEDFE17" box="[769,858,457,482]" gridcol="4" gridrow="6" pageId="15" pageNumber="16">4.73</td>
|
||
<td id="124B5D8FFFB3006DFCFBFE49FC65FE17" box="[889,978,457,482]" gridcol="5" gridrow="6" pageId="15" pageNumber="16">4.54</td>
|
||
<td id="124B5D8FFFB3006DFC7EFE49FBE2FE17" box="[1020,1109,457,482]" gridcol="6" gridrow="6" pageId="15" pageNumber="16">4.5</td>
|
||
<td id="124B5D8FFFB3006DFB01FE49FB6BFE17" box="[1155,1244,457,482]" gridcol="7" gridrow="6" pageId="15" pageNumber="16">4.5</td>
|
||
<td id="124B5D8FFFB3006DFA8AFE49FAD6FE17" box="[1288,1377,457,482]" gridcol="8" gridrow="6" pageId="15" pageNumber="16">4.8</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFE74FAD6FDF9" box="[152,1377,500,524]" gridrow="7" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFE74FEA4FDF9" box="[152,275,500,524]" gridcol="0" gridrow="7" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFE74FF73FDFC" bold="true" box="[152,196,500,521]" pageId="15" pageNumber="16">IND</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FE74FE5CFDF9" box="[401,491,500,524]" gridcol="1" gridrow="7" pageId="15" pageNumber="16">2.58</td>
|
||
<td id="124B5D8FFFB3006DFD8EFE74FDD2FDF9" box="[524,613,500,524]" gridcol="2" gridrow="7" pageId="15" pageNumber="16">2.37</td>
|
||
<td id="124B5D8FFFB3006DFD08FE74FD54FDF9" box="[650,739,500,524]" gridcol="3" gridrow="7" pageId="15" pageNumber="16">2.55</td>
|
||
<td id="124B5D8FFFB3006DFC83FE74FCEDFDF9" box="[769,858,500,524]" gridcol="4" gridrow="7" pageId="15" pageNumber="16">2.34</td>
|
||
<td id="124B5D8FFFB3006DFCFBFE74FC65FDF9" box="[889,978,500,524]" gridcol="5" gridrow="7" pageId="15" pageNumber="16">2.31</td>
|
||
<td id="124B5D8FFFB3006DFC7EFE74FBE2FDF9" box="[1020,1109,500,524]" gridcol="6" gridrow="7" pageId="15" pageNumber="16">2.33</td>
|
||
<td id="124B5D8FFFB3006DFB01FE74FB6BFDF9" box="[1155,1244,500,524]" gridcol="7" gridrow="7" pageId="15" pageNumber="16">2.28</td>
|
||
<td id="124B5D8FFFB3006DFA8AFE74FAD6FDF9" box="[1288,1377,500,524]" gridcol="8" gridrow="7" pageId="15" pageNumber="16">2.44</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFD9EFAD6FDC3" box="[152,1377,542,566]" gridrow="8" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFD9EFEA4FDC3" box="[152,275,542,566]" gridcol="0" gridrow="8" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFD9EFF7CFDC6" bold="true" box="[152,203,542,563]" pageId="15" pageNumber="16">END</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FD9EFE5CFDC3" box="[401,491,542,566]" gridcol="1" gridrow="8" pageId="15" pageNumber="16">2.80</td>
|
||
<td id="124B5D8FFFB3006DFD8EFD9EFDD2FDC3" box="[524,613,542,566]" gridcol="2" gridrow="8" pageId="15" pageNumber="16">2.74</td>
|
||
<td id="124B5D8FFFB3006DFD08FD9EFD54FDC3" box="[650,739,542,566]" gridcol="3" gridrow="8" pageId="15" pageNumber="16">2.78</td>
|
||
<td id="124B5D8FFFB3006DFC83FD9EFCEDFDC3" box="[769,858,542,566]" gridcol="4" gridrow="8" pageId="15" pageNumber="16">2.59</td>
|
||
<td id="124B5D8FFFB3006DFCFBFD9EFC65FDC3" box="[889,978,542,566]" gridcol="5" gridrow="8" pageId="15" pageNumber="16">2.4</td>
|
||
<td id="124B5D8FFFB3006DFC7EFD9EFBE2FDC3" box="[1020,1109,542,566]" gridcol="6" gridrow="8" pageId="15" pageNumber="16">2.41</td>
|
||
<td id="124B5D8FFFB3006DFB01FD9EFB6BFDC3" box="[1155,1244,542,566]" gridcol="7" gridrow="8" pageId="15" pageNumber="16">2.38</td>
|
||
<td id="124B5D8FFFB3006DFA8AFD9EFAD6FDC3" box="[1288,1377,542,566]" gridcol="8" gridrow="8" pageId="15" pageNumber="16">2.52</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFDC8FAD6FD95" box="[152,1377,584,608]" gridrow="9" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFDC8FEA4FD95" box="[152,275,584,608]" gridcol="0" gridrow="9" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFDC8FF7EFDA8" bold="true" box="[152,201,584,605]" pageId="15" pageNumber="16">ETD</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FDC8FE5CFD95" box="[401,491,584,608]" gridcol="1" gridrow="9" pageId="15" pageNumber="16">1.03</td>
|
||
<td id="124B5D8FFFB3006DFD8EFDC8FDD2FD95" box="[524,613,584,608]" gridcol="2" gridrow="9" pageId="15" pageNumber="16">1.00</td>
|
||
<td id="124B5D8FFFB3006DFD08FDC8FD54FD95" box="[650,739,584,608]" gridcol="3" gridrow="9" pageId="15" pageNumber="16">1.01</td>
|
||
<td id="124B5D8FFFB3006DFC83FDC8FCEDFD95" box="[769,858,584,608]" gridcol="4" gridrow="9" pageId="15" pageNumber="16">1.03</td>
|
||
<td id="124B5D8FFFB3006DFCFBFDC8FC65FD95" box="[889,978,584,608]" gridcol="5" gridrow="9" pageId="15" pageNumber="16">0.97</td>
|
||
<td id="124B5D8FFFB3006DFC7EFDC8FBE2FD95" box="[1020,1109,584,608]" gridcol="6" gridrow="9" pageId="15" pageNumber="16">0.95</td>
|
||
<td id="124B5D8FFFB3006DFB01FDC8FB6BFD95" box="[1155,1244,584,608]" gridcol="7" gridrow="9" pageId="15" pageNumber="16">0.96</td>
|
||
<td id="124B5D8FFFB3006DFA8AFDC8FAD6FD95" box="[1288,1377,584,608]" gridcol="8" gridrow="9" pageId="15" pageNumber="16">1.07</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFDF0FAD6FD73" box="[152,1377,624,646]" gridrow="10" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFDF0FEA4FD73" box="[152,275,624,646]" gridcol="0" gridrow="10" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFDF0FF70FD70" bold="true" box="[152,199,624,645]" pageId="15" pageNumber="16">UpL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FDF0FE5CFD73" box="[401,491,624,646]" gridcol="1" gridrow="10" pageId="15" pageNumber="16">5.76</td>
|
||
<td id="124B5D8FFFB3006DFD8EFDF0FDD2FD73" box="[524,613,624,646]" gridcol="2" gridrow="10" pageId="15" pageNumber="16">6.26</td>
|
||
<td id="124B5D8FFFB3006DFD08FDF0FD54FD73" box="[650,739,624,646]" gridcol="3" gridrow="10" pageId="15" pageNumber="16">6.40</td>
|
||
<td id="124B5D8FFFB3006DFC83FDF0FCEDFD73" box="[769,858,624,646]" gridcol="4" gridrow="10" pageId="15" pageNumber="16">4.52</td>
|
||
<td id="124B5D8FFFB3006DFCFBFDF0FC65FD73" box="[889,978,624,646]" gridcol="5" gridrow="10" pageId="15" pageNumber="16">5.32</td>
|
||
<td id="124B5D8FFFB3006DFC7EFDF0FBE2FD73" box="[1020,1109,624,646]" gridcol="6" gridrow="10" pageId="15" pageNumber="16">5.4</td>
|
||
<td id="124B5D8FFFB3006DFB01FDF0FB6BFD73" box="[1155,1244,624,646]" gridcol="7" gridrow="10" pageId="15" pageNumber="16">5.37</td>
|
||
<td id="124B5D8FFFB3006DFA8AFDF0FAD6FD73" box="[1288,1377,624,646]" gridcol="8" gridrow="10" pageId="15" pageNumber="16">6.2</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFD18FAD6FD45" box="[152,1377,664,688]" gridrow="11" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFD18FEA4FD45" box="[152,275,664,688]" gridcol="0" gridrow="11" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFD18FF74FD58" bold="true" box="[152,195,664,685]" pageId="15" pageNumber="16">FoL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FD18FE5CFD45" box="[401,491,664,688]" gridcol="1" gridrow="11" pageId="15" pageNumber="16">6.74</td>
|
||
<td id="124B5D8FFFB3006DFD8EFD18FDD2FD45" box="[524,613,664,688]" gridcol="2" gridrow="11" pageId="15" pageNumber="16">7.43</td>
|
||
<td id="124B5D8FFFB3006DFD08FD18FD54FD45" box="[650,739,664,688]" gridcol="3" gridrow="11" pageId="15" pageNumber="16">7.56</td>
|
||
<td id="124B5D8FFFB3006DFC83FD18FCEDFD45" box="[769,858,664,688]" gridcol="4" gridrow="11" pageId="15" pageNumber="16">7.15</td>
|
||
<td id="124B5D8FFFB3006DFCFBFD18FC65FD45" box="[889,978,664,688]" gridcol="5" gridrow="11" pageId="15" pageNumber="16">6.8</td>
|
||
<td id="124B5D8FFFB3006DFC7EFD18FBE2FD45" box="[1020,1109,664,688]" gridcol="6" gridrow="11" pageId="15" pageNumber="16">6.93</td>
|
||
<td id="124B5D8FFFB3006DFB01FD18FB6BFD45" box="[1155,1244,664,688]" gridcol="7" gridrow="11" pageId="15" pageNumber="16">6.87</td>
|
||
<td id="124B5D8FFFB3006DFA8AFD18FAD6FD45" box="[1288,1377,664,688]" gridcol="8" gridrow="11" pageId="15" pageNumber="16">7.92</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFD41FAD6FD22" box="[152,1377,705,727]" gridrow="12" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFD41FEA4FD22" box="[152,275,705,727]" gridcol="0" gridrow="12" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFD41FF70FD23" bold="true" box="[152,199,705,726]" pageId="15" pageNumber="16">HaL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FD41FE5CFD22" box="[401,491,705,727]" gridcol="1" gridrow="12" pageId="15" pageNumber="16">5.5</td>
|
||
<td id="124B5D8FFFB3006DFD8EFD41FDD2FD22" box="[524,613,705,727]" gridcol="2" gridrow="12" pageId="15" pageNumber="16">5.4</td>
|
||
<td id="124B5D8FFFB3006DFD08FD41FD54FD22" box="[650,739,705,727]" gridcol="3" gridrow="12" pageId="15" pageNumber="16">6.2</td>
|
||
<td id="124B5D8FFFB3006DFC83FD41FCEDFD22" box="[769,858,705,727]" gridcol="4" gridrow="12" pageId="15" pageNumber="16">5.25</td>
|
||
<td id="124B5D8FFFB3006DFCFBFD41FC65FD22" box="[889,978,705,727]" gridcol="5" gridrow="12" pageId="15" pageNumber="16">5.11</td>
|
||
<td id="124B5D8FFFB3006DFC7EFD41FBE2FD22" box="[1020,1109,705,727]" gridcol="6" gridrow="12" pageId="15" pageNumber="16">5</td>
|
||
<td id="124B5D8FFFB3006DFB01FD41FB6BFD22" box="[1155,1244,705,727]" gridcol="7" gridrow="12" pageId="15" pageNumber="16">5.05</td>
|
||
<td id="124B5D8FFFB3006DFA8AFD41FAD6FD22" box="[1288,1377,705,727]" gridcol="8" gridrow="12" pageId="15" pageNumber="16">6.25</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFD69FAD6FCF4" box="[152,1377,745,769]" gridrow="13" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFD69FEA4FCF4" box="[152,275,745,769]" gridcol="0" gridrow="13" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFD69FEA4FD0A" bold="true" box="[152,275,745,767]" pageId="15" pageNumber="16">F3PW/F3W</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FD69FE5CFCF4" box="[401,491,745,769]" gridcol="1" gridrow="13" pageId="15" pageNumber="16">1.70</td>
|
||
<td id="124B5D8FFFB3006DFD8EFD69FDD2FCF4" box="[524,613,745,769]" gridcol="2" gridrow="13" pageId="15" pageNumber="16">1.80</td>
|
||
<td id="124B5D8FFFB3006DFD08FD69FD54FCF4" box="[650,739,745,769]" gridcol="3" gridrow="13" pageId="15" pageNumber="16">1.80</td>
|
||
<td id="124B5D8FFFB3006DFC83FD69FCEDFCF4" box="[769,858,745,769]" gridcol="4" gridrow="13" pageId="15" pageNumber="16">2.2</td>
|
||
<td id="124B5D8FFFB3006DFCFBFD69FC65FCF4" box="[889,978,745,769]" gridcol="5" gridrow="13" pageId="15" pageNumber="16">1.7</td>
|
||
<td id="124B5D8FFFB3006DFC7EFD69FBE2FCF4" box="[1020,1109,745,769]" gridcol="6" gridrow="13" pageId="15" pageNumber="16">1.7</td>
|
||
<td id="124B5D8FFFB3006DFB01FD69FB6BFCF4" box="[1155,1244,745,769]" gridcol="7" gridrow="13" pageId="15" pageNumber="16">1.6</td>
|
||
<td id="124B5D8FFFB3006DFA8AFD69FAD6FCF4" box="[1288,1377,745,769]" gridcol="8" gridrow="13" pageId="15" pageNumber="16">2.3</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFC93FAD6FCD9" box="[152,1377,787,812]" gridrow="14" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFC93FEA4FCD9" box="[152,275,787,812]" gridcol="0" gridrow="14" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFC93FEA4FCDC" bold="true" box="[152,275,787,809]" pageId="15" pageNumber="16">F4PW/F4W</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FC93FE5CFCD9" box="[401,491,787,812]" gridcol="1" gridrow="14" pageId="15" pageNumber="16">1.60</td>
|
||
<td id="124B5D8FFFB3006DFD8EFC93FDD2FCD9" box="[524,613,787,812]" gridcol="2" gridrow="14" pageId="15" pageNumber="16">1.90</td>
|
||
<td id="124B5D8FFFB3006DFD08FC93FD54FCD9" box="[650,739,787,812]" gridcol="3" gridrow="14" pageId="15" pageNumber="16">1.70</td>
|
||
<td id="124B5D8FFFB3006DFC83FC93FCEDFCD9" box="[769,858,787,812]" gridcol="4" gridrow="14" pageId="15" pageNumber="16">2.3</td>
|
||
<td id="124B5D8FFFB3006DFCFBFC93FC65FCD9" box="[889,978,787,812]" gridcol="5" gridrow="14" pageId="15" pageNumber="16">1.4</td>
|
||
<td id="124B5D8FFFB3006DFC7EFC93FBE2FCD9" box="[1020,1109,787,812]" gridcol="6" gridrow="14" pageId="15" pageNumber="16">1.5</td>
|
||
<td id="124B5D8FFFB3006DFB01FC93FB6BFCD9" box="[1155,1244,787,812]" gridcol="7" gridrow="14" pageId="15" pageNumber="16">1.5</td>
|
||
<td id="124B5D8FFFB3006DFA8AFC93FAD6FCD9" box="[1288,1377,787,812]" gridcol="8" gridrow="14" pageId="15" pageNumber="16">2.3</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFCBEFAD6FCA3" box="[152,1377,830,854]" gridrow="15" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFCBEFEA4FCA3" box="[152,275,830,854]" gridcol="0" gridrow="15" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFCBEFF76FCA6" bold="true" box="[152,193,830,851]" pageId="15" pageNumber="16">FeL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FCBEFE5CFCA3" box="[401,491,830,854]" gridcol="1" gridrow="15" pageId="15" pageNumber="16">10.26</td>
|
||
<td id="124B5D8FFFB3006DFD8EFCBEFDD2FCA3" box="[524,613,830,854]" gridcol="2" gridrow="15" pageId="15" pageNumber="16">10.58</td>
|
||
<td id="124B5D8FFFB3006DFD08FCBEFD54FCA3" box="[650,739,830,854]" gridcol="3" gridrow="15" pageId="15" pageNumber="16">11.00</td>
|
||
<td id="124B5D8FFFB3006DFC83FCBEFCEDFCA3" box="[769,858,830,854]" gridcol="4" gridrow="15" pageId="15" pageNumber="16">9.62</td>
|
||
<td id="124B5D8FFFB3006DFCFBFCBEFC65FCA3" box="[889,978,830,854]" gridcol="5" gridrow="15" pageId="15" pageNumber="16">9.35</td>
|
||
<td id="124B5D8FFFB3006DFC7EFCBEFBE2FCA3" box="[1020,1109,830,854]" gridcol="6" gridrow="15" pageId="15" pageNumber="16">9.26</td>
|
||
<td id="124B5D8FFFB3006DFB01FCBEFB6BFCA3" box="[1155,1244,830,854]" gridcol="7" gridrow="15" pageId="15" pageNumber="16">9.3</td>
|
||
<td id="124B5D8FFFB3006DFA8AFCBEFAD6FCA3" box="[1288,1377,830,854]" gridcol="8" gridrow="15" pageId="15" pageNumber="16">11.28</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFCE6FAD6FC88" box="[152,1377,870,893]" gridrow="16" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFCE6FEA4FC88" box="[152,275,870,893]" gridcol="0" gridrow="16" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFCE6FF0FFC8E" bold="true" box="[152,184,870,891]" pageId="15" pageNumber="16">TL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FCE6FE5CFC88" box="[401,491,870,893]" gridcol="1" gridrow="16" pageId="15" pageNumber="16">11.03</td>
|
||
<td id="124B5D8FFFB3006DFD8EFCE6FDD2FC88" box="[524,613,870,893]" gridcol="2" gridrow="16" pageId="15" pageNumber="16">11.65</td>
|
||
<td id="124B5D8FFFB3006DFD08FCE6FD54FC88" box="[650,739,870,893]" gridcol="3" gridrow="16" pageId="15" pageNumber="16">11.33</td>
|
||
<td id="124B5D8FFFB3006DFC83FCE6FCEDFC88" box="[769,858,870,893]" gridcol="4" gridrow="16" pageId="15" pageNumber="16">10.31</td>
|
||
<td id="124B5D8FFFB3006DFCFBFCE6FC65FC88" box="[889,978,870,893]" gridcol="5" gridrow="16" pageId="15" pageNumber="16">9.95</td>
|
||
<td id="124B5D8FFFB3006DFC7EFCE6FBE2FC88" box="[1020,1109,870,893]" gridcol="6" gridrow="16" pageId="15" pageNumber="16">10.12</td>
|
||
<td id="124B5D8FFFB3006DFB01FCE6FB6BFC88" box="[1155,1244,870,893]" gridcol="7" gridrow="16" pageId="15" pageNumber="16">9.9</td>
|
||
<td id="124B5D8FFFB3006DFA8AFCE6FAD6FC88" box="[1288,1377,870,893]" gridcol="8" gridrow="16" pageId="15" pageNumber="16">12.33</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFC0FFAD6FC52" box="[152,1377,911,935]" gridrow="17" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFC0FFEA4FC52" box="[152,275,911,935]" gridcol="0" gridrow="17" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFC0FFF6FFC51" bold="true" box="[152,216,911,932]" pageId="15" pageNumber="16">TotFL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FC0FFE5CFC52" box="[401,491,911,935]" gridcol="1" gridrow="17" pageId="15" pageNumber="16">16.4</td>
|
||
<td id="124B5D8FFFB3006DFD8EFC0FFDD2FC52" box="[524,613,911,935]" gridcol="2" gridrow="17" pageId="15" pageNumber="16">16.3</td>
|
||
<td id="124B5D8FFFB3006DFD08FC0FFD54FC52" box="[650,739,911,935]" gridcol="3" gridrow="17" pageId="15" pageNumber="16">17.5</td>
|
||
<td id="124B5D8FFFB3006DFC83FC0FFCEDFC52" box="[769,858,911,935]" gridcol="4" gridrow="17" pageId="15" pageNumber="16">15.4</td>
|
||
<td id="124B5D8FFFB3006DFCFBFC0FFC65FC52" box="[889,978,911,935]" gridcol="5" gridrow="17" pageId="15" pageNumber="16">15.51</td>
|
||
<td id="124B5D8FFFB3006DFC7EFC0FFBE2FC52" box="[1020,1109,911,935]" gridcol="6" gridrow="17" pageId="15" pageNumber="16">15.35</td>
|
||
<td id="124B5D8FFFB3006DFB01FC0FFB6BFC52" box="[1155,1244,911,935]" gridcol="7" gridrow="17" pageId="15" pageNumber="16">15.4</td>
|
||
<td id="124B5D8FFFB3006DFA8AFC0FFAD6FC52" box="[1288,1377,911,935]" gridcol="8" gridrow="17" pageId="15" pageNumber="16">18.05</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFC37FAD6FC3B" box="[152,1377,951,974]" gridrow="18" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFC37FEA4FC3B" box="[152,275,951,974]" gridcol="0" gridrow="18" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFC37FF77FC39" bold="true" box="[152,192,951,972]" pageId="15" pageNumber="16">IPT</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FC37FE5CFC3B" box="[401,491,951,974]" gridcol="1" gridrow="18" pageId="15" pageNumber="16">0.58</td>
|
||
<td id="124B5D8FFFB3006DFD8EFC37FDD2FC3B" box="[524,613,951,974]" gridcol="2" gridrow="18" pageId="15" pageNumber="16">0.70</td>
|
||
<td id="124B5D8FFFB3006DFD08FC37FD54FC3B" box="[650,739,951,974]" gridcol="3" gridrow="18" pageId="15" pageNumber="16">0.74</td>
|
||
<td id="124B5D8FFFB3006DFC83FC37FCEDFC3B" box="[769,858,951,974]" gridcol="4" gridrow="18" pageId="15" pageNumber="16">0.5</td>
|
||
<td id="124B5D8FFFB3006DFCFBFC37FC65FC3B" box="[889,978,951,974]" gridcol="5" gridrow="18" pageId="15" pageNumber="16">0.58</td>
|
||
<td id="124B5D8FFFB3006DFC7EFC37FBE2FC3B" box="[1020,1109,951,974]" gridcol="6" gridrow="18" pageId="15" pageNumber="16">0.6</td>
|
||
<td id="124B5D8FFFB3006DFB01FC37FB6BFC3B" box="[1155,1244,951,974]" gridcol="7" gridrow="18" pageId="15" pageNumber="16">0.59</td>
|
||
<td id="124B5D8FFFB3006DFA8AFC37FAD6FC3B" box="[1288,1377,951,974]" gridcol="8" gridrow="18" pageId="15" pageNumber="16">0.65</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFC60FAD6FC0D" box="[152,1377,992,1016]" gridrow="19" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFC60FEA4FC0D" box="[152,275,992,1016]" gridcol="0" gridrow="19" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFC60FF7AFC00" bold="true" box="[152,205,992,1013]" pageId="15" pageNumber="16">MPT</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FC60FE5CFC0D" box="[401,491,992,1016]" gridcol="1" gridrow="19" pageId="15" pageNumber="16">0.84</td>
|
||
<td id="124B5D8FFFB3006DFD8EFC60FDD2FC0D" box="[524,613,992,1016]" gridcol="2" gridrow="19" pageId="15" pageNumber="16">1.00</td>
|
||
<td id="124B5D8FFFB3006DFD08FC60FD54FC0D" box="[650,739,992,1016]" gridcol="3" gridrow="19" pageId="15" pageNumber="16">1.07</td>
|
||
<td id="124B5D8FFFB3006DFC83FC60FCEDFC0D" box="[769,858,992,1016]" gridcol="4" gridrow="19" pageId="15" pageNumber="16">0.77</td>
|
||
<td id="124B5D8FFFB3006DFCFBFC60FC65FC0D" box="[889,978,992,1016]" gridcol="5" gridrow="19" pageId="15" pageNumber="16">0.88</td>
|
||
<td id="124B5D8FFFB3006DFC7EFC60FBE2FC0D" box="[1020,1109,992,1016]" gridcol="6" gridrow="19" pageId="15" pageNumber="16">0.86</td>
|
||
<td id="124B5D8FFFB3006DFB01FC60FB6BFC0D" box="[1155,1244,992,1016]" gridcol="7" gridrow="19" pageId="15" pageNumber="16">0.87</td>
|
||
<td id="124B5D8FFFB3006DFA8AFC60FAD6FC0D" box="[1288,1377,992,1016]" gridcol="8" gridrow="19" pageId="15" pageNumber="16">0.86</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFB8AFAD6FBD7" box="[152,1377,1034,1058]" gridrow="20" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFB8AFEA4FBD7" box="[152,275,1034,1058]" gridcol="0" gridrow="20" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFB8AFF7EFBD5" bold="true" box="[152,201,1034,1056]" pageId="15" pageNumber="16">OPT</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FB8AFE5CFBD7" box="[401,491,1034,1058]" gridcol="1" gridrow="20" pageId="15" pageNumber="16">0.4</td>
|
||
<td id="124B5D8FFFB3006DFD8EFB8AFDD2FBD7" box="[524,613,1034,1058]" gridcol="2" gridrow="20" pageId="15" pageNumber="16">0.45</td>
|
||
<td id="124B5D8FFFB3006DFD08FB8AFD54FBD7" box="[650,739,1034,1058]" gridcol="3" gridrow="20" pageId="15" pageNumber="16">0.45</td>
|
||
<td id="124B5D8FFFB3006DFC83FB8AFCEDFBD7" box="[769,858,1034,1058]" gridcol="4" gridrow="20" pageId="15" pageNumber="16">0.32</td>
|
||
<td id="124B5D8FFFB3006DFCFBFB8AFC65FBD7" box="[889,978,1034,1058]" gridcol="5" gridrow="20" pageId="15" pageNumber="16">0.39</td>
|
||
<td id="124B5D8FFFB3006DFC7EFB8AFBE2FBD7" box="[1020,1109,1034,1058]" gridcol="6" gridrow="20" pageId="15" pageNumber="16">0.42</td>
|
||
<td id="124B5D8FFFB3006DFB01FB8AFB6BFBD7" box="[1155,1244,1034,1058]" gridcol="7" gridrow="20" pageId="15" pageNumber="16">0.42</td>
|
||
<td id="124B5D8FFFB3006DFA8AFB8AFAD6FBD7" box="[1288,1377,1034,1058]" gridcol="8" gridrow="20" pageId="15" pageNumber="16">0.4</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFBB4FAD6FBB9" box="[152,1377,1076,1100]" gridrow="21" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFBB4FEA4FBB9" box="[152,275,1076,1100]" gridcol="0" gridrow="21" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFBB4FF6FFBBC" bold="true" box="[152,216,1076,1097]" pageId="15" pageNumber="16">IMTL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FBB4FE5CFBB9" box="[401,491,1076,1100]" gridcol="1" gridrow="21" pageId="15" pageNumber="16">0.50</td>
|
||
<td id="124B5D8FFFB3006DFD8EFBB4FDD2FBB9" box="[524,613,1076,1100]" gridcol="2" gridrow="21" pageId="15" pageNumber="16">0.58</td>
|
||
<td id="124B5D8FFFB3006DFD08FBB4FD54FBB9" box="[650,739,1076,1100]" gridcol="3" gridrow="21" pageId="15" pageNumber="16">0.46</td>
|
||
<td id="124B5D8FFFB3006DFC83FBB4FCEDFBB9" box="[769,858,1076,1100]" gridcol="4" gridrow="21" pageId="15" pageNumber="16">0.52</td>
|
||
<td id="124B5D8FFFB3006DFCFBFBB4FC65FBB9" box="[889,978,1076,1100]" gridcol="5" gridrow="21" pageId="15" pageNumber="16">0.47</td>
|
||
<td id="124B5D8FFFB3006DFC7EFBB4FBE2FBB9" box="[1020,1109,1076,1100]" gridcol="6" gridrow="21" pageId="15" pageNumber="16">0.46</td>
|
||
<td id="124B5D8FFFB3006DFB01FBB4FB6BFBB9" box="[1155,1244,1076,1100]" gridcol="7" gridrow="21" pageId="15" pageNumber="16">0.47</td>
|
||
<td id="124B5D8FFFB3006DFA8AFBB4FAD6FBB9" box="[1288,1377,1076,1100]" gridcol="8" gridrow="21" pageId="15" pageNumber="16">0.64</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFBDCFAD6FB86" box="[152,1377,1116,1139]" gridrow="22" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFBDCFEA4FB86" box="[152,275,1116,1139]" gridcol="0" gridrow="22" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFBDCFF56FB87" bold="true" box="[152,225,1116,1138]" pageId="15" pageNumber="16">OMTL</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FBDCFE5CFB86" box="[401,491,1116,1139]" gridcol="1" gridrow="22" pageId="15" pageNumber="16">0.90</td>
|
||
<td id="124B5D8FFFB3006DFD8EFBDCFDD2FB86" box="[524,613,1116,1139]" gridcol="2" gridrow="22" pageId="15" pageNumber="16">0.92</td>
|
||
<td id="124B5D8FFFB3006DFD08FBDCFD54FB86" box="[650,739,1116,1139]" gridcol="3" gridrow="22" pageId="15" pageNumber="16">1.04</td>
|
||
<td id="124B5D8FFFB3006DFC83FBDCFCEDFB86" box="[769,858,1116,1139]" gridcol="4" gridrow="22" pageId="15" pageNumber="16">0.74</td>
|
||
<td id="124B5D8FFFB3006DFCFBFBDCFC65FB86" box="[889,978,1116,1139]" gridcol="5" gridrow="22" pageId="15" pageNumber="16">0.84</td>
|
||
<td id="124B5D8FFFB3006DFC7EFBDCFBE2FB86" box="[1020,1109,1116,1139]" gridcol="6" gridrow="22" pageId="15" pageNumber="16">0.85</td>
|
||
<td id="124B5D8FFFB3006DFB01FBDCFB6BFB86" box="[1155,1244,1116,1139]" gridcol="7" gridrow="22" pageId="15" pageNumber="16">0.83</td>
|
||
<td id="124B5D8FFFB3006DFA8AFBDCFAD6FB86" box="[1288,1377,1116,1139]" gridcol="8" gridrow="22" pageId="15" pageNumber="16">0.99</td>
|
||
</tr>
|
||
<tr id="519A34F3FFB3006DFF1AFB06FAD6FB55" box="[152,1377,1158,1184]" gridrow="23" pageId="15" pageNumber="16">
|
||
<th id="124B5D8FFFB3006DFF1AFB06FEA4FB55" box="[152,275,1158,1184]" gridcol="0" gridrow="23" pageId="15" pageNumber="16">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF1AFB06FF5FFB69" bold="true" box="[152,232,1158,1180]" pageId="15" pageNumber="16">TW/ED</emphasis>
|
||
</th>
|
||
<td id="124B5D8FFFB3006DFE13FB06FE5CFB55" box="[401,491,1158,1184]" gridcol="1" gridrow="23" pageId="15" pageNumber="16">0.28</td>
|
||
<td id="124B5D8FFFB3006DFD8EFB06FDD2FB55" box="[524,613,1158,1184]" gridcol="2" gridrow="23" pageId="15" pageNumber="16">0.28</td>
|
||
<td id="124B5D8FFFB3006DFD08FB06FD54FB55" box="[650,739,1158,1184]" gridcol="3" gridrow="23" pageId="15" pageNumber="16">0.29</td>
|
||
<td id="124B5D8FFFB3006DFC83FB06FCEDFB55" box="[769,858,1158,1184]" gridcol="4" gridrow="23" pageId="15" pageNumber="16">0.28</td>
|
||
<td id="124B5D8FFFB3006DFCFBFB06FC65FB55" box="[889,978,1158,1184]" gridcol="5" gridrow="23" pageId="15" pageNumber="16">0.27</td>
|
||
<td id="124B5D8FFFB3006DFC7EFB06FBE2FB55" box="[1020,1109,1158,1184]" gridcol="6" gridrow="23" pageId="15" pageNumber="16">0.26</td>
|
||
<td id="124B5D8FFFB3006DFB01FB06FB6BFB55" box="[1155,1244,1158,1184]" gridcol="7" gridrow="23" pageId="15" pageNumber="16">0.27</td>
|
||
<td id="124B5D8FFFB3006DFA8AFB06FAD6FB55" box="[1288,1377,1158,1184]" gridcol="8" gridrow="23" pageId="15" pageNumber="16">0.28</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<subSubSection id="A7B0653AFFB3FF9DFF0FFB50FC55F9BC" pageId="15" pageNumber="16" type="discussion">
|
||
<paragraph id="EF1536B1FFB3FF9DFF0FFB50FE1BFAFF" blockId="15.[141,745,1232,1291]" pageId="15" pageNumber="16">
|
||
<heading id="B45D81DDFFB3FF9DFF0FFB50FE1BFAFF" bold="true" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="6">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF0FFB50FE1BFAFF" bold="true" pageId="15" pageNumber="16">
|
||
Molecular Analysis of the
|
||
<taxonomicName id="28AA4D32FFB3FF9DFD80FB50FF52FAFE" ID-CoL="399J5" baseAuthorityName="Peters" baseAuthorityYear="1870" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nitidus">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFD80FB50FF52FAFE" bold="true" italics="true" pageId="15" pageNumber="16">Eleutherodactylus nitidus</emphasis>
|
||
</taxonomicName>
|
||
Species Group
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB3FF9DFF0FFAADFDDEF9C5" blockId="15.[141,746,1325,1965]" pageId="15" pageNumber="16">
|
||
The molecular phylogeny based on the mitochondrial rRNA 16S recovered a well-supported
|
||
<taxonomicName id="28AA4D32FFB3FF9DFF0FFAE8FE00FA75" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[141,439,1384,1408]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nitidus">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF0FFAE8FE00FA75" box="[141,439,1384,1408]" italics="true" pageId="15" pageNumber="16">Eleutherodactylus nitidus</emphasis>
|
||
</taxonomicName>
|
||
species group (posterior probability (pp) 1;
|
||
<figureCitation id="77912A34FFB3FF9DFEECFA05FE7CFA68" box="[366,459,1413,1437]" captionStart="Fig" captionStartId="21.[139,174,1854,1876]" captionTargetBox="[240,1267,156,1823]" captionTargetId="figure-22@21.[165,1360,141,1832]" captionTargetPageId="21" captionText="Fig. 12. Bayesian phylogenetic inference of members of the Eleutherodactylus subgenus Syrrhophus, with a focus on the E. nitidus species group, based on the mitochondrial loci 16S rRNA. Black circles represent nodes with a posterior support of 1. All nodes with support of less than 0.5 are collapsed." figureDoi="http://doi.org/10.5281/zenodo.11265080" httpUri="https://zenodo.org/record/11265080/files/figure.png" pageId="15" pageNumber="16">Figs. 12</figureCitation>
|
||
and S
|
||
<figureCitation id="77912A34FFB3FF9DFD99FA05FD91FA68" box="[539,550,1413,1437]" captionStart="Fig" captionStartId="4.[141,176,1744,1766]" captionTargetBox="[144,1416,192,1728]" captionTargetId="figure-73@4.[144,1416,192,1322]" captionTargetPageId="4" captionText="Fig. 1. Holotype of Eleutherodactylus maculabialis sp. nov., MZFC 33312 (CIG 00921) from 11.4 km S of Puerto de Gallo, Municipio de Atoyac de Álvarez, Guerrero, Mexico. (A) Dorsolateral perspective in life. (B) Lateral perspective in life. (C) Ventral perspective in life. (D) Dorsal perspective in preservative. (E) Ventral perspective in preservative." figureDoi="http://doi.org/10.5281/zenodo.11265044" httpUri="https://zenodo.org/record/11265044/files/figure.png" pageId="15" pageNumber="16">1</figureCitation>
|
||
), which is sister to the
|
||
<taxonomicName id="28AA4D32FFB3FF9DFF5BFA23FED4FA4F" baseAuthorityName="Taylor" baseAuthorityYear="1942" box="[217,355,1442,1466]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF5BFA23FED4FA4F" box="[217,355,1442,1466]" italics="true" pageId="15" pageNumber="16">E. modestus</emphasis>
|
||
</taxonomicName>
|
||
group as defined by
|
||
<bibRefCitation id="8B3B4B40FFB3FF9DFDDEFA22FF49FA2D" author="Grunwald CI & Reyes-Velasco J & Franz-Chavez H & Morales-Flores KI & Ahumada-Carrillo IT & Jones JM & Boissinot S." pageId="15" pageNumber="16" pagination="6 - 83" refId="ref15970" refString="Grunwald CI, Reyes-Velasco J, Franz-Chavez H, Morales-Flores KI, Ahumada-Carrillo IT, Jones JM, Boissinot S. 2018. Six new species of Eleutherodactylus (Anura: Eleutherodactylidae: subgenus Syrrhophus) from Mexico, with a discussion of their systematic relationships and the validity of related species. Mesoamerican Herpetology 5 (1): 6 - 83." type="journal article" year="2018">Grünwald et al. (2018)</bibRefCitation>
|
||
. As with previous phylogenies of the group based on 16S, some of the intermediate nodes in the phylogeny are not well supported (pp <0.5), and we have collapsed all nodes below this value.
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB3FF9DFF36F9B5FF52F8A0" blockId="15.[141,746,1325,1965]" pageId="15" pageNumber="16">
|
||
In the
|
||
<taxonomicName id="28AA4D32FFB3FF9DFF79F9B5FDACF9B8" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[251,539,1589,1613]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nitidus">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF79F9B5FDACF9B8" box="[251,539,1589,1613]" italics="true" pageId="15" pageNumber="16">Eleutherodactylus nitidus</emphasis>
|
||
</taxonomicName>
|
||
species group, we recovered two main clades. The first one is composed of
|
||
<emphasis id="DDDEEAA3FFB3FF9DFF2DF9F0FDA6F97D" box="[175,529,1648,1672]" italics="true" pageId="15" pageNumber="16">
|
||
<taxonomicName id="28AA4D32FFB3FF9DFF2DF9F0FE87F97D" baseAuthorityName="Taylor" baseAuthorityYear="1940" box="[175,304,1648,1672]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="pipilans">E. pipilans</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB3FF9DFEC2F9F0FDBBF97D" baseAuthorityName="Palacios-Aguilar and Santos-Bibiano" baseAuthorityYear="2020" box="[320,524,1648,1672]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="erythrochomus">E. erythrochomus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis id="DDDEEAA3FFB3FF9DFDCDF9F0FD5FF97D" box="[591,744,1648,1672]" italics="true" pageId="15" pageNumber="16">
|
||
<taxonomicName id="28AA4D32FFB3FF9DFDCDF9F0FD53F97D" baseAuthorityName="Taylor" baseAuthorityYear="1943" box="[591,740,1648,1672]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nebulosus">E. nebulosus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
while the second group includes all the other species. In the second group, many of the intermediate nodes have little to no support. However, individual species do show strong support (pp = 1) in most cases, including
|
||
<emphasis id="DDDEEAA3FFB3FF9DFE82F883FD2FF8EF" box="[256,664,1794,1818]" italics="true" pageId="15" pageNumber="16">
|
||
<taxonomicName id="28AA4D32FFB3FF9DFE82F883FEC7F8EF" baseAuthorityName="Davis and Dixon" baseAuthorityYear="1955" box="[256,368,1794,1818]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="dilatus">E. dilatus</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB3FF9DFEFFF883FDA6F8EF" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[381,529,1794,1818]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="sentinelus">E. sentinelus</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB3FF9DFD9CF883FD22F8EF" authorityName="Hedges" authorityYear="1989" box="[542,661,1795,1818]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="maurus">E. maurus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<emphasis id="DDDEEAA3FFB3FF9DFD50F883FF5EF8C2" italics="true" pageId="15" pageNumber="16">
|
||
<taxonomicName id="28AA4D32FFB3FF9DFD50F883FF52F8C2" baseAuthorityName="Hoyt" baseAuthorityYear="1965" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="syristes">E. syristes</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
while
|
||
<taxonomicName id="28AA4D32FFB3FF9DFEB7F8A0FE52F8CD" authorityName="Grünwald & Reyes-Velasco & Franz-Chávez & Morales-Flores & Ahumada-Carrillo & Rodriguez & Jones" authorityYear="2021" box="[309,485,1824,1848]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="maculabialis">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFEB7F8A0FE52F8CD" box="[309,485,1824,1848]" italics="true" pageId="15" pageNumber="16">E. maculabialis</emphasis>
|
||
</taxonomicName>
|
||
has a posterior support of 0.86.
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB3FF9DFF2FF8DAFAADFA6C" blockId="15.[141,746,1325,1965]" lastBlockId="15.[790,1395,1233,1609]" pageId="15" pageNumber="16">
|
||
All the remaining species in the group form a second, well-supported clade (pp =0.99). In this clade, we recovered an early split between
|
||
<taxonomicName id="28AA4D32FFB3FF9DFC94FB51FBD4FB1C" baseAuthorityName="Taylor" baseAuthorityYear="1943" box="[790,1123,1233,1257]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="albolabris">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFC94FB51FBD4FB1C" box="[790,1123,1233,1257]" italics="true" pageId="15" pageNumber="16">Eleutherodactylus albolabris</emphasis>
|
||
</taxonomicName>
|
||
and the other species, which include
|
||
<taxonomicName id="28AA4D32FFB3FF9DFC40FB6FFB80FAF3" baseAuthorityName="Duellman" baseAuthorityYear="1958" box="[962,1079,1263,1286]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="orarius">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFC40FB6FFB80FAF3" box="[962,1079,1263,1286]" italics="true" pageId="15" pageNumber="16">E. orarius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="DDDEEAA3FFB3FF9DFBCBFB6FFAF7FAF3" box="[1097,1344,1262,1286]" italics="true" pageId="15" pageNumber="16">
|
||
<taxonomicName id="28AA4D32FFB3FF9DFBCBFB6FFB0CFAF3" baseAuthorityName="Duellman" baseAuthorityYear="1954" box="[1097,1211,1263,1286]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="petersi">E. petersi</taxonomicName>
|
||
,
|
||
<taxonomicName id="28AA4D32FFB3FF9DFB48FB6FFA8BFAF3" authority="(Peters, 1870)" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[1226,1340,1262,1286]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nitidus">E. nitidus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and a hitherto undescribed form from
|
||
<collectingRegion id="2D6EF853FFB3FF9DFB1BFA8BFB51FAD6" box="[1177,1254,1291,1315]" country="Mexico" name="Jalisco" pageId="15" pageNumber="16">Jalisco</collectingRegion>
|
||
and
|
||
<collectingRegion id="2D6EF853FFB3FF9DFA9DFA8BFAC5FAD6" box="[1311,1394,1291,1315]" country="Mexico" name="Nayarit" pageId="15" pageNumber="16">Nayarit</collectingRegion>
|
||
(
|
||
<figureCitation id="77912A34FFB3FF9DFC9DFAA9FCC6FAB4" box="[799,881,1321,1345]" captionStart="Fig" captionStartId="23.[141,176,622,644]" captionTargetBox="[141,745,154,609]" captionTargetId="figure-730@23.[141,745,154,609]" captionTargetPageId="23" captionText="Fig. 13. Map of type localities and distributions of members of the Eleutherodactylus nitidus species group. Purple circles represent localities of an undescribed species related to E. petersi. For the other four species, stars represent the type localities and circles represent additional localities, which are color coded for species: E. orarius (red), E. albolabris (blue), E. nitidus (green), and E. petersi (orange)." figureDoi="http://doi.org/10.5281/zenodo.11265082" httpUri="https://zenodo.org/record/11265082/files/figure.png" pageId="15" pageNumber="16">Fig. 13</figureCitation>
|
||
). We refrain from describing the western-most form at this time, as material from the
|
||
<typeStatus id="30118813FFB3FF9DFB61FAC6FAA4FAAB" box="[1251,1299,1350,1374]" pageId="15" pageNumber="16">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="28AA4D32FFB3FF9DFCB4FAE4FC14FA8E" baseAuthorityName="Duellman" baseAuthorityYear="1954" box="[822,931,1380,1403]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="petersi">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFCB4FAE4FC14FA8E" box="[822,931,1380,1403]" italics="true" pageId="15" pageNumber="16">E. petersi</emphasis>
|
||
</taxonomicName>
|
||
is unavailable, so the relationship of the western form with
|
||
<taxonomicName id="28AA4D32FFB3FF9DFC6FFA01FBEEFA6D" baseAuthorityName="Duellman" baseAuthorityYear="1954" box="[1005,1113,1409,1432]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="petersi">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFC6FFA01FBEEFA6D" box="[1005,1113,1409,1432]" italics="true" pageId="15" pageNumber="16">E. petersi</emphasis>
|
||
</taxonomicName>
|
||
remains unclear.
|
||
</paragraph>
|
||
<paragraph id="EF1536B1FFB3FF9DFCB4FA1EFC55F9BC" blockId="15.[790,1395,1233,1609]" pageId="15" pageNumber="16">
|
||
We believe that a more detailed study of the taxa related to
|
||
<taxonomicName id="28AA4D32FFB3FF9DFC0FFA3CFC4BFA26" authority="(Peters, 1870)" baseAuthorityName="Peters" baseAuthorityYear="1870" box="[909,1020,1467,1491]" class="Amphibia" family="Eleutherodactylidae" genus="Eleutherodactylus" kingdom="Animalia" order="Anura" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="nitidus">
|
||
<emphasis id="DDDEEAA3FFB3FF9DFC0FFA3CFC4BFA26" box="[909,1020,1467,1491]" italics="true" pageId="15" pageNumber="16">E. nitidus</emphasis>
|
||
</taxonomicName>
|
||
is needed, when more thorough sampling is available and employing more populationlevel specific tools, such as genome-wide SNP data, in order to better understand the patterns of gene-flow and introgression.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |