420 lines
53 KiB
XML
420 lines
53 KiB
XML
<document ID-DOI="10.11646/zootaxa.5264.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="7836798" IM.bibliography_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1681728995984" checkinUser="plazi" docAuthor="Heppner, John B. & Bae, Yang-Seop" docDate="2023" docId="3A0C87F6DD19A71C51D9FF23FCE3F80B" docLanguage="en" docName="zootaxa.5264.3.5.pdf" docOrigin="Zootaxa 5264 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5264.3.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Vangatinea Heppner & Bae 2023, gen. nov." docType="treatment" docVersion="4" lastPageNumber="372" masterDocId="C635FF8EDD18A71F514EFFB4FFD3FF9E" masterDocTitle="New species of colorful tineids from Cambodia, Thailand, and Vietnam in the new genus Vangatinea (Lepidoptera: Tineidae: Erechthiinae)" masterLastPageNumber="380" masterPageNumber="369" pageNumber="370" updateTime="1681825662450" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species of colorful tineids from Cambodia, Thailand, and Vietnam in the new genus Vangatinea (Lepidoptera: Tineidae: Erechthiinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Heppner, John B.</mods:namePart>
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<mods:affiliation>McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, Florida 32611, USA. & jheppner @ flmnh. ufl. edu</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bae, Yang-Seop</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-7356-5633</mods:nameIdentifier>
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<mods:affiliation>Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. & io ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. & Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea. & baeys @ inu. ac. kr; https: // orcid. org / 0000 - 0001 - 7356 - 5633</mods:affiliation>
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<mods:nameIdentifier type="email">baeys@inu.ac.kr</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-14</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5264</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>3</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>369</mods:start>
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<mods:end>380</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5264.3.5</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5264.3.5</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7836798</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7841998" ID-Zenodo-Dep="7841998" LSID="urn:lsid:plazi:treatment:3A0C87F6DD19A71C51D9FF23FCE3F80B" httpUri="http://treatment.plazi.org/id/3A0C87F6DD19A71C51D9FF23FCE3F80B" lastPageId="3" lastPageNumber="372" pageId="1" pageNumber="370">
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<subSubSection box="[151,667,151,177]" pageId="1" pageNumber="370" type="nomenclature">
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<paragraph blockId="1.[151,979,151,250]" box="[151,667,151,177]" pageId="1" pageNumber="370">
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<heading bold="true" box="[151,667,151,177]" fontSize="11" level="1" pageId="1" pageNumber="370" reason="1">
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<emphasis bold="true" box="[151,667,151,177]" pageId="1" pageNumber="370">
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<taxonomicName authority="Heppner & Bae" authorityName="Heppner & Bae" authorityYear="2023" box="[151,544,151,177]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[151,338,151,177]" italics="true" pageId="1" pageNumber="370">VANGATINEA</emphasis>
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Heppner & Bae
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</taxonomicName>
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,
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<taxonomicNameLabel box="[557,667,151,177]" pageId="1" pageNumber="370" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="370" type="description">
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<paragraph blockId="1.[151,979,151,250]" box="[151,274,187,213]" pageId="1" pageNumber="370">
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(
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<figureCitation box="[159,265,187,213]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[151,250,1449,1473]" captionStartId-1="2.[151,250,1799,1823]" captionTargetBox-0="[267,1315,717,1425]" captionTargetBox-1="[238,1359,1149,1768]" captionTargetId-0="figure-172@1.[255,1332,700,1425]" captionTargetId-1="figure-510@2.[229,1359,1137,1776]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Vangatinea gen. nov., wing venation (V. sontraensis sp. nov., male paratype) (11.4mm wingspan) (wing slide JBH- 4257) (MGCL)." captionText-1="FIGURE 2. Vangatinea gen. nov., schematic of head morphology (V. sontraensis sp. nov., male paratype) (head slide JBH- 4257) (MGCL)." figureDoi-0="http://doi.org/10.5281/zenodo.7836800" figureDoi-1="http://doi.org/10.5281/zenodo.7836802" httpUri-0="https://zenodo.org/record/7836800/files/figure.png" httpUri-1="https://zenodo.org/record/7836802/files/figure.png" pageId="1" pageNumber="370">Figs. 1–2</figureCitation>
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)
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</paragraph>
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<paragraph blockId="1.[151,979,151,250]" box="[151,979,223,250]" pageId="1" pageNumber="370">
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<uri box="[151,979,223,250]" pageId="1" pageNumber="370">
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urn:lsid:zoobank.org:act:
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<uuid box="[425,979,223,250]" pageId="1" pageNumber="370">7F849327-272A-4D3B-AC32-5D4ED7D5CA33</uuid>
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</uri>
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</paragraph>
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<paragraph blockId="1.[151,789,292,316]" box="[151,789,292,316]" pageId="1" pageNumber="370">
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<typeStatus box="[151,201,292,316]" pageId="1" pageNumber="370">Type</typeStatus>
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species:
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<taxonomicName authority="Heppner & Bae" authorityName="Heppner & Bae" authorityYear="2023" box="[294,698,292,316]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="sontraensis" status="sp. nov.">
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<emphasis box="[294,530,292,315]" italics="true" pageId="1" pageNumber="370">Vangatinea sontraensis</emphasis>
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Heppner & Bae
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</taxonomicName>
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,
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<emphasis bold="true" box="[708,789,292,316]" pageId="1" pageNumber="370">
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<taxonomicNameLabel box="[708,789,292,316]" pageId="1" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="370" type="diagnosis">
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<paragraph blockId="1.[151,1437,361,676]" pageId="1" pageNumber="370">
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<emphasis bold="true" box="[151,266,361,387]" pageId="1" pageNumber="370">Diagnosis</emphasis>
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. The genus is distinguished by the apically acute wings, with the hindwing costal margin abruptly arcuate at midwing (
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<figureCitation box="[302,371,398,424]" captionStart="FIGURE 1" captionStartId="1.[151,250,1449,1473]" captionTargetBox="[267,1315,717,1425]" captionTargetId="figure-172@1.[255,1332,700,1425]" captionTargetPageId="1" captionText="FIGURE 1. Vangatinea gen. nov., wing venation (V. sontraensis sp. nov., male paratype) (11.4mm wingspan) (wing slide JBH- 4257) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.7836800" httpUri="https://zenodo.org/record/7836800/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
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), and the colorful orange (to golden-orange) forewing maculation, with dark blue-black to purple-brown basal and distal patches, pale brown hindwings, golden-yellow head, and slightly thickened filiform antennae. The male genitalia are strongly sclerotized in compact and rounded tegumen-vinculum conical shell, with very short saccus (or undeveloped), with complicated and small mandible-like valvae; aedeagus with single tubular cornutus as long as aedeagus; male abdomen with small lateral pregenital coremata of spiculate setae (or lacking). The female genitalia have an extremely long and thin ovipositor with simple papillae anales (small and setose), and a strongly sclerotized acute bifurcate sterigma that seems to be a piercing structure; corethrogyne absent; bursa small and ovate; and signum absent.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7836800" ID-Zenodo-Dep="7836800" httpUri="https://zenodo.org/record/7836800/files/figure.png" pageId="1" pageNumber="370" startId="1.[151,250,1449,1473]" targetBox="[267,1315,717,1425]" targetPageId="1">
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<paragraph blockId="1.[151,1436,1449,1509]" pageId="1" pageNumber="370">
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<emphasis bold="true" box="[151,274,1449,1473]" pageId="1" pageNumber="370">FIGURE 1.</emphasis>
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<taxonomicName authority="Heppner & Bae, 2023" authorityName="Heppner & Bae" authorityYear="2023" box="[280,394,1450,1473]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis box="[280,394,1450,1473]" italics="true" pageId="1" pageNumber="370">Vangatinea</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[399,493,1449,1473]" pageId="1" pageNumber="370">
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<taxonomicNameLabel box="[399,493,1449,1473]" pageId="1" pageNumber="370" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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, wing venation (
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<taxonomicName authority="Heppner & Bae, 2023" authorityName="Heppner & Bae" authorityYear="2023" box="[663,802,1450,1473]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="sontraensis" status="sp. nov.">
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<emphasis box="[663,802,1450,1473]" italics="true" pageId="1" pageNumber="370">V. sontraensis</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[808,888,1449,1473]" pageId="1" pageNumber="370">
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<taxonomicNameLabel box="[808,888,1449,1473]" pageId="1" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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, male paratype) (11.4mm wingspan) (wing slide JBH- 4257) (MGCL).
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</paragraph>
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</caption>
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<subSubSection lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370" type="description">
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<paragraph blockId="1.[151,1437,1555,2014]" lastBlockId="2.[151,1437,150,1113]" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370">
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<emphasis bold="true" box="[199,333,1555,1581]" pageId="1" pageNumber="370">Description</emphasis>
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. Wing expanse:
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<quantity box="[523,654,1555,1581]" metricMagnitude="-2" metricUnit="m" metricValue="1.335" metricValueMax="1.6" metricValueMin="1.0699999999999998" pageId="1" pageNumber="370" unit="mm" value="13.35" valueMax="16.0" valueMin="10.7">10.7–16mm</quantity>
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(n = 5).
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<emphasis box="[760,819,1555,1581]" italics="true" pageId="1" pageNumber="370">Head</emphasis>
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(
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<figureCitation box="[836,907,1555,1581]" captionStart="FIGURE 2" captionStartId="2.[151,250,1799,1823]" captionTargetBox="[238,1359,1149,1768]" captionTargetId="figure-510@2.[229,1359,1137,1776]" captionTargetPageId="2" captionText="FIGURE 2. Vangatinea gen. nov., schematic of head morphology (V. sontraensis sp. nov., male paratype) (head slide JBH- 4257) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.7836802" httpUri="https://zenodo.org/record/7836802/files/figure.png" pageId="1" pageNumber="370">Fig. 2</figureCitation>
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): vertex medially conical, with short epicranial suture; transverse suture indistinct; long golden-yellow to orange piliform scales on vertex and frons; eyes medium to large (smaller in female), with simple facets; ocellus absent; chaetosema absent; maxillary palpus short, 5-segmented (total length subequal to basal segment of labial palpus); labial palpus drooping (more upturned in female), segments slender and distal two segments nearly subequal in length, basal segment shorter, with only 2–3 ventral or lateral bristles on middle segment; vom Rath's organ on apical segment; haustellum short (galeae free, length subequal to apical labial palpus segment), held straight ventrally (with minute lateral filaments); mandible minute stub; pilifer minute (with single long seta); antenna filiform, average length (ca. 2/3 forewing length), with single row of scales per barrel-shaped segment; scape of median length (ca. 3–4 antennal segments in length), with indistinct narrow pecten field of 20–22 minute stubby bristles (6–8 of which are stouter); pedicel barrel-shaped (ca. 1.5x in width and length of remaining antennal segments).
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<emphasis box="[521,599,1915,1941]" italics="true" pageId="1" pageNumber="370">Thorax</emphasis>
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: average for body size, smooth-scaled; tegula average; patagia undeveloped; legs average length for body.
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<emphasis box="[490,595,1952,1977]" italics="true" pageId="1" pageNumber="370">Forewing</emphasis>
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(
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<figureCitation box="[613,681,1951,1977]" captionStart="FIGURE 1" captionStartId="1.[151,250,1449,1473]" captionTargetBox="[267,1315,717,1425]" captionTargetId="figure-172@1.[255,1332,700,1425]" captionTargetPageId="1" captionText="FIGURE 1. Vangatinea gen. nov., wing venation (V. sontraensis sp. nov., male paratype) (11.4mm wingspan) (wing slide JBH- 4257) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.7836800" httpUri="https://zenodo.org/record/7836800/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
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): very narrow elongated (4x longer than wide), having acute apex (female forewing slightly less narrow, see
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<figureCitation box="[628,695,1987,2014]" captionStart="FIGURES 3–5" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[199,1398,1421,1873]" captionTargetId="figure-539@4.[190,1398,1412,1875]" captionTargetPageId="4" captionText="FIGURES 3–5. Vangatinea species: 3) V. sontraensis sp. nov. (Vietnam: Danang), male holotpe (11.4mm wingspan) (MGCL); 4) V. panghapha sp. nov. (Thailand: Mai Hong Son), male holotype (13.5mm wingspan) (MGCL); 5) V. cambodiensis sp. nov. (Cambodia: Pursat), female holotype (16mm wingspan) (INUC)." figureDoi="http://doi.org/10.5281/zenodo.7836804" httpUri="https://zenodo.org/record/7836804/files/figure.png" pageId="1" pageNumber="370">Fig. 5</figureCitation>
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); Sc very close to costal wing margin, to midwing; radius basally vestigial; R1 long, from basal 1/3 of wing; R2–5 evenly spaced from discal cell, with R2 somewhat more distant from R3, and with R5 to before apex; discal cell long, with chorda present but indistinct; median veins evenly spaced, but M3 slightly more approximate to M2 at cell than M1 to M2; cubitus basally vestigial; CuA1 and CuA2 short and parallel to tornus; CuP short at tornus, but stronger from base to near tornus; A1+2 strong, with convex bulge of A1 after basal segment; A2 weak in basal fork; A3 indistinct; tornus straight (slightly rounded after termen); dorsal margin convex; anal margin short-rounded; maculation bright orange with golden iridescence, and black-brown wing base and distal patches; fringes long on termen; retinaculum normal (elongated triangle with rolled apex).
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<emphasis box="[1147,1254,367,393]" italics="true" pageId="2" pageNumber="371">Hindwing</emphasis>
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(
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<figureCitation box="[1270,1335,367,393]" captionStart="FIGURE 1" captionStartId="1.[151,250,1449,1473]" captionTargetBox="[267,1315,717,1425]" captionTargetId="figure-172@1.[255,1332,700,1425]" captionTargetPageId="1" captionText="FIGURE 1. Vangatinea gen. nov., wing venation (V. sontraensis sp. nov., male paratype) (11.4mm wingspan) (wing slide JBH- 4257) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.7836800" httpUri="https://zenodo.org/record/7836800/files/figure.png" pageId="2" pageNumber="371">Fig. 1</figureCitation>
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): narrow (4x longer than wide), with acute attenuated apex and costal margin concave from midwing to apex (female hindwing slightly less narrow, see
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<figureCitation box="[423,490,438,465]" captionStart="FIGURES 3–5" captionStartId="4.[151,264,1898,1922]" captionTargetBox="[199,1398,1421,1873]" captionTargetId="figure-539@4.[190,1398,1412,1875]" captionTargetPageId="4" captionText="FIGURES 3–5. Vangatinea species: 3) V. sontraensis sp. nov. (Vietnam: Danang), male holotpe (11.4mm wingspan) (MGCL); 4) V. panghapha sp. nov. (Thailand: Mai Hong Son), male holotype (13.5mm wingspan) (MGCL); 5) V. cambodiensis sp. nov. (Cambodia: Pursat), female holotype (16mm wingspan) (INUC)." figureDoi="http://doi.org/10.5281/zenodo.7836804" httpUri="https://zenodo.org/record/7836804/files/figure.png" pageId="2" pageNumber="371">Fig. 5</figureCitation>
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); Sc strong, to post-midwing arcuate costal margin; Rs weak until before intersection with transverse vein, then stronger to before apex; M1 to just dorsad of apex; M2 divergent of M1 at termen; M3 and CuA1 nearly parallel; CuA1 and CuA2 short, parallel to tornus; CuP strong to wing base; A1+2 short, to basal dorsal margin, medially concave, with indistinct or vestigial basal fork;A3 short; tornus convex and evenly merging to termen; dorsal margin relatively straight; anal margin turned abruptly from dorsal margin before wing base; maculation pale to brown, with long fringes on termen to anal margin; frenulum of average length for wing size.
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<emphasis box="[1184,1287,619,645]" italics="true" pageId="2" pageNumber="371">Abdomen</emphasis>
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: average size for body; normal scaling; with or without small lateral pregenital coremata in males; female corethrogyne absent.
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<emphasis italics="true" pageId="2" pageNumber="371">Male genitalia</emphasis>
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(
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<figureCitation box="[263,329,691,717]" captionStart="FIGURE 9" captionStartId="6.[151,250,1271,1295]" captionTargetBox="[155,1417,184,1247]" captionTargetId="figure-29@6.[151,1436,181,1247]" captionTargetPageId="6" captionText="FIGURE 9. Vangatinea sontraensis sp. nov., holotype male genitalia (Vietnam: Danang) (valvae shown ventrally), with details, a) aedeagus (in situ) and juxta-anellus, and b) pre-genital segment of abdomen with small coremata (gen. slide JBH-2997) (MGCL)." figureDoi="http://doi.org/10.5281/zenodo.7836814" httpUri="https://zenodo.org/record/7836814/files/figure.png" pageId="2" pageNumber="371">Fig. 9</figureCitation>
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): complicated structure, with compact tegumen-vinculum conical cylinder strongly sclerotized; uncus projected as bifurcate arms, or reduced; gnathos absent; tuba analis short or undeveloped; subscaphium absent; saccus very small or absent (or with internal spur); valva short, quadratic, and strongly sclerotized (sclerotized more along the saccular margin); valval shape mandible-like with saccular tooth-like spines, or with setose field and more dense setae on saccular margin, and with or without basal appendage; juxta-anellus a short tubular structure, with or without basally lateral pad-like termini; aedeagus long and slender; cornutus a single very long tubule (subequal to length of aedeagus and nearly as wide as aedeagus); phallobase undeveloped; seminal vesica short.
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<emphasis box="[1025,1211,907,933]" italics="true" pageId="2" pageNumber="371">Female genitalia</emphasis>
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(
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<figureCitation box="[1225,1305,907,933]" captionStart="FIGURE 11" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1434,181,1920]" captionTargetId="figure-29@8.[154,1434,181,1920]" captionTargetPageId="8" captionText="FIGURE 11. Vangatinea cambodiensis sp. nov., holotype female genitalia (Cambodia: Pursat) (ovipositor not completely extended, thus even longer than shown), with details, a) sterigma and ostium (gen. slide JBH-2998) (INUC)." figureDoi="http://doi.org/10.5281/zenodo.7836820" httpUri="https://zenodo.org/record/7836820/files/figure.png" pageId="2" pageNumber="371">Fig. 11</figureCitation>
|
||
): ovipositor extremely long and thin (ca. 3x segment VI, and longer when ovipositor is fully extended); papillae anales small, slightly setose (not formed strongly for piercing); apophyses anteriores twice thickness of long apophyses posteriores (and 2.3x length of apophyses posteriores); ostium simple, membranous; sterigma divided into proximate and extremely acute asymmetrical caudal knife-like projections, with ventro-lateral carinate edges; ductus bursae a membranous long tube (ca. 1.5x bursa length); corpus bursae small and ovate (ca. 1/2x length of segment VI); signum absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7836802" ID-Zenodo-Dep="7836802" httpUri="https://zenodo.org/record/7836802/files/figure.png" pageId="2" pageNumber="371" startId="2.[151,250,1799,1823]" targetBox="[238,1359,1149,1768]" targetPageId="2">
|
||
<paragraph blockId="2.[151,1436,1799,1859]" pageId="2" pageNumber="371">
|
||
<emphasis bold="true" box="[151,277,1799,1823]" pageId="2" pageNumber="371">FIGURE 2.</emphasis>
|
||
<taxonomicName authority="Heppner & Bae, 2023" authorityName="Heppner & Bae" authorityYear="2023" box="[285,399,1800,1823]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis box="[285,399,1800,1823]" italics="true" pageId="2" pageNumber="371">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[407,504,1799,1823]" pageId="2" pageNumber="371">
|
||
<taxonomicNameLabel box="[407,504,1799,1823]" pageId="2" pageNumber="371" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, schematic of head morphology (
|
||
<taxonomicName authority="Heppner & Bae, 2023" authorityName="Heppner & Bae" authorityYear="2023" box="[854,995,1800,1823]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="sontraensis" status="sp. nov.">
|
||
<emphasis box="[854,995,1800,1823]" italics="true" pageId="2" pageNumber="371">V. sontraensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1004,1086,1799,1823]" pageId="2" pageNumber="371">
|
||
<taxonomicNameLabel box="[1004,1086,1799,1823]" pageId="2" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male paratype) (head slide JBH- 4257) (MGCL).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection pageId="2" pageNumber="371" type="etymology">
|
||
<paragraph blockId="2.[151,1437,1905,2040]" pageId="2" pageNumber="371">
|
||
<emphasis bold="true" box="[199,328,1906,1932]" pageId="2" pageNumber="371">Etymology</emphasis>
|
||
. This genus is named for the colorful golden-orange adult moths, thus "vang" in Vietnamese ("golden" in English); the moths having orange forewings with golden iridescence, and golden-yellow to orange head scaling.
|
||
<emphasis bold="true" box="[199,289,2013,2039]" pageId="2" pageNumber="371">Biology</emphasis>
|
||
. Unknown. Adults are likely day-fliers, although all specimens were collected at lights.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1421,150,177]" pageId="3" pageNumber="372" type="distribution">
|
||
<paragraph blockId="3.[151,1437,150,1941]" box="[199,1421,150,177]" pageId="3" pageNumber="372">
|
||
<emphasis bold="true" box="[199,345,150,176]" pageId="3" pageNumber="372">Distribution</emphasis>
|
||
. Only known from mainland Southeast Asia (
|
||
<collectingCountry box="[859,979,151,177]" name="Cambodia" pageId="3" pageNumber="372">Cambodia</collectingCountry>
|
||
,
|
||
<collectingCountry box="[989,1091,151,177]" name="Thailand" pageId="3" pageNumber="372">Thailand</collectingCountry>
|
||
, and
|
||
<collectingCountry box="[1149,1246,151,177]" name="Vietnam" pageId="3" pageNumber="372">Vietnam</collectingCountry>
|
||
) (
|
||
<figureCitation box="[1269,1349,151,177]" captionStart="FIGURE 12" captionStartId="9.[151,250,1934,1958]" captionTargetBox="[311,1277,930,1911]" captionTargetId="figure-367@9.[311,1277,930,1911]" captionTargetPageId="9" captionText="FIGURE 12. Southeast Asia distribution map of Vangatinea species: V. cambodiensis sp. nov. (white square, Cambodia), V. panghapha sp. nov. (black star, Thailand), and V. sontraensis sp. nov. (black square, Vietnam) (base map courtesy of J. Bartholomew & Son Maps, Edinburgh, Scotland; Times Atlas, 1988. London, UK)." figureDoi="http://doi.org/10.5281/zenodo.7836822" httpUri="https://zenodo.org/record/7836822/files/figure.png" pageId="3" pageNumber="372">Fig. 12</figureCitation>
|
||
map).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="372" type="discussion">
|
||
<paragraph blockId="3.[151,1437,150,1941]" pageId="3" pageNumber="372">
|
||
<emphasis bold="true" box="[199,325,186,212]" pageId="3" pageNumber="372">Discussion</emphasis>
|
||
. Pending further studies, based on the character spectrum of
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[1055,1181,187,212]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1055,1181,187,212]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
we tentatively assign this new genus to the subfamily
|
||
<taxonomicName authorityName="Meyrick" authorityYear="1880" box="[519,663,223,249]" class="Insecta" family="Tineidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="subFamily" subFamily="Erechthiinae">Erechthiinae</taxonomicName>
|
||
, with possible relationship to
|
||
<taxonomicName authority="Meyrick 1880" authorityName="Meyrick" authorityYear="1880" box="[1003,1290,223,249]" class="Insecta" family="Tineidae" genus="Erechthias" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1003,1123,223,249]" italics="true" pageId="3" pageNumber="372">Erechthias</emphasis>
|
||
Meyrick 1880
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authority="Stainton 1859" authorityName="Stainton" authorityYear="1859" class="Insecta" family="Tineidae" genus="Pyloetis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1346,1436,223,249]" italics="true" pageId="3" pageNumber="372">Pyloetis</emphasis>
|
||
Stainton 1859
|
||
</taxonomicName>
|
||
, but lacking some relational characters like the female signum of the latter two genera (
|
||
<bibRefCitation author="Common, I. F. B." pageId="3" pageNumber="372" pagination="181 - 186" refId="ref5708" refString="Common, I. F. B. (1990) Family Tineidae. In: Moths of Australia. Melbourne University Press, Melbourne, pp. 181 - 186." type="book chapter" year="1990">Common 1990</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Davis, D. R. & Robinson, G. S." box="[221,512,295,321]" pageId="3" pageNumber="372" pagination="91 - 117" refId="ref5783" refString="Davis, D. R. & Robinson, G. S. (1998) The Tineoidea and Gracillarioidea. In: Kristensen, N. P. (Ed.), Lepidoptera, Moths and Butterflies. Vol. 1. Evolution, Systematics, and Biogeography. In Handbuch der Zoologie. Band IV. Arthropoda: Insecta. Teilband 35. W. De Gruyter, Berlin, pp. 91 - 117. https: // doi. org / 10.1515 / 9783110804744.91" type="book chapter" year="1998">Davis and Robinson 1998</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Miyamoto, Y. & Huang, G. - H. & Hirowatari, T." box="[522,766,295,321]" pageId="3" pageNumber="372" pagination="363 - 371" refId="ref6066" refString="Miyamoto, Y., Huang, G. - H. & Hirowatari, T. (2007) Systematic position of Pyloetis mimosae (Stainton) (Lepidoptera: Tineidae), with description of the adults and immature stages. Entomological Science, 10, 363 - 371. https: // doi. org / 10.1111 / j. 1479 - 8298.2007.00235. x" type="journal article" year="2007">
|
||
Miyamoto
|
||
<emphasis box="[645,701,295,321]" italics="true" pageId="3" pageNumber="372">et al.</emphasis>
|
||
2007
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Moriuti, S. & Kadohara, T." box="[776,1089,295,321]" pageId="3" pageNumber="372" pagination="565 - 584" refId="ref6197" refString="Moriuti, S. & Kadohara, T. (1994) Erechthiinae (Lepidoptera, Tineidae) of Japan. Japanese Journal of Entomology, 62, 565 - 584." type="journal article" year="1994">Moriuti and Kadohara 1994</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Robinson, G. S." box="[1099,1272,295,321]" pageId="3" pageNumber="372" refId="ref6388" refString="Robinson, G. S. (2009) Biology, Distribution and Diversity of Tineid Moths. Southdene, Kuala Lumpur, 143 pp., 16 pls." type="book" year="2009">Robinson 2009</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Robinson, G. S. & Nielsen, E. S." pageId="3" pageNumber="372" refId="ref6418" refString="Robinson, G. S. & Nielsen, E. S. (1993) Tineid Genera of Australia (Lepidoptera). Monographs on Australian Lepidoptera. Vol. 2. CSIRO, East Melbourne, 344 pp. https: // doi. org / 10.1071 / 9780643105102" type="book" year="1993">Robinson and Nielsen 1993</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Robinson, G. S. & Tuck, K. R. & Shaffer, M." box="[314,550,331,357]" pageId="3" pageNumber="372" refId="ref6511" refString="Robinson, G. S., Tuck, K. R. & Shaffer, M. (1994) A Field Guide to the Smaller Moths of South-East Asia. Malaysian Nature Society, Kuala Lumpur, 309 pp., 32 pls." type="book" year="1994">
|
||
Robinson
|
||
<emphasis box="[428,477,331,357]" italics="true" pageId="3" pageNumber="372">et al</emphasis>
|
||
. 1994
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Zimmerman, E. C." box="[561,766,331,357]" pageId="3" pageNumber="372" refId="ref6794" refString="Zimmerman, E. C. (1978) Tineidae. In: Insects of Hawaii. Vol. 9. Microlepidoptera, University of Hawaii Press, Honolulu, pp. 254 - 296." type="book" year="1978">Zimmerman 1978</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,150,1941]" pageId="3" pageNumber="372">
|
||
Typical erechthiine genera have an elongated antennal scape, acute narrow wings, lateral male coremata on sternum VI, and females lacking a corethrogyne, as does
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[785,911,403,428]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[785,911,403,428]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
. The most comprehensive and detailed modern review of tineid genera thus far is by
|
||
<bibRefCitation author="Robinson, G. S. & Nielsen, E. S." box="[582,910,439,465]" pageId="3" pageNumber="372" refId="ref6418" refString="Robinson, G. S. & Nielsen, E. S. (1993) Tineid Genera of Australia (Lepidoptera). Monographs on Australian Lepidoptera. Vol. 2. CSIRO, East Melbourne, 344 pp. https: // doi. org / 10.1071 / 9780643105102" type="book" year="1993">Robinson and Nielsen (1993</bibRefCitation>
|
||
; slightly updated for subfamilies in
|
||
<bibRefCitation author="Davis, D. R. & Robinson, G. S." pageId="3" pageNumber="372" pagination="91 - 117" refId="ref5783" refString="Davis, D. R. & Robinson, G. S. (1998) The Tineoidea and Gracillarioidea. In: Kristensen, N. P. (Ed.), Lepidoptera, Moths and Butterflies. Vol. 1. Evolution, Systematics, and Biogeography. In Handbuch der Zoologie. Band IV. Arthropoda: Insecta. Teilband 35. W. De Gruyter, Berlin, pp. 91 - 117. https: // doi. org / 10.1515 / 9783110804744.91" type="book chapter" year="1998">Davis and Robinson 1998</bibRefCitation>
|
||
), but this is already from 30 years ago and encompasses only the limited Australian fauna (only 44 genera present out of a world fauna of 320 genera); so, in this work
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[920,1046,511,536]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[920,1046,511,536]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
keys out best to
|
||
<taxonomicName authorityName="Meyrick" authorityYear="1880" box="[1237,1382,511,537]" class="Insecta" family="Tineidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="subFamily" subFamily="Erechthiinae">Erechthiinae</taxonomicName>
|
||
. The more recent paper by
|
||
<bibRefCitation author="Regier, J. C. & Mitter, C. & Davis, D. R. & Harrison, T. L. & Sohn, J. - C. & Cummins, M. P. & Zwick, A. & Mitter, K. T." box="[395,615,547,573]" pageId="3" pageNumber="372" pagination="409 - 432" refId="ref6283" refString="Regier, J. C., Mitter, C., Davis, D. R., Harrison, T. L., Sohn, J. - C., Cummins, M. P., Zwick, A. & Mitter, K. T. (2014) A molecular phylogeny and revised classification for the oldest ditrysian moth lineages (Lepidoptera: Tineoidea), with implication for ancestral feeding habits of the mega-diverse Ditrysia. Systematic Entomology, 40, 409 - 432. https: // doi. org / 10.1111 / syen. 12110" type="journal article" year="2014">
|
||
Regier
|
||
<emphasis box="[477,526,547,573]" italics="true" pageId="3" pageNumber="372">et al</emphasis>
|
||
. (2014)
|
||
</bibRefCitation>
|
||
likewise has no solution for these various unassociated genera.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,150,1941]" pageId="3" pageNumber="372">
|
||
The short maxillary palpi, the minute pilifers with single setae, the radically complex yet compact male genitalia, and the elongated acute forewings, as well as the costal arcuation of the hindwings, would also indicate a possible placement in
|
||
<taxonomicName box="[401,527,655,681]" class="Insecta" family="Meessiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="family">Meessiinae</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Davis, D. R. & Robinson, G. S." box="[542,837,655,681]" pageId="3" pageNumber="372" pagination="91 - 117" refId="ref5783" refString="Davis, D. R. & Robinson, G. S. (1998) The Tineoidea and Gracillarioidea. In: Kristensen, N. P. (Ed.), Lepidoptera, Moths and Butterflies. Vol. 1. Evolution, Systematics, and Biogeography. In Handbuch der Zoologie. Band IV. Arthropoda: Insecta. Teilband 35. W. De Gruyter, Berlin, pp. 91 - 117. https: // doi. org / 10.1515 / 9783110804744.91" type="book chapter" year="1998">Davis and Robinson 1998</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Robinson, G. S. & Nielsen, E. S." box="[849,1163,655,681]" pageId="3" pageNumber="372" refId="ref6418" refString="Robinson, G. S. & Nielsen, E. S. (1993) Tineid Genera of Australia (Lepidoptera). Monographs on Australian Lepidoptera. Vol. 2. CSIRO, East Melbourne, 344 pp. https: // doi. org / 10.1071 / 9780643105102" type="book" year="1993">Robinson and Nielsen 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Zagulajev, A. K." box="[1175,1353,655,681]" pageId="3" pageNumber="372" pagination="124 - 129" refId="ref6626" refString="Zagulajev, A. K. (1977) The classification and phylogeny of the subfamily Meessiinae (Lepidoptera, Tineidae). Entomologicheskoe Obozrenie, 56, 662 - 669. [in Russian, English translation: Entomological Review, 56 (3), 124 - 129 (1978)]" type="journal article" year="1977">Zagulajev 1977</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Zagulajev, A. K." box="[1364,1424,655,681]" pageId="3" pageNumber="372" pagination="1 - 408" refId="ref6680" refString="Zagulajev, A. K. (1979) Tineidae. Subfamily Meessiinae. In: Fauna USSR. Lepidoptera. Part 4 (6). Akademie Nauk USSR, St. Petersburg, pp. 1 - 408. [in Russian]" type="book chapter" year="1979">1979</bibRefCitation>
|
||
). However, meessiine tineids are mostly dull marked to white in forewing maculation. One African meessiine genus (
|
||
<taxonomicName authority="Gozmany 1969" authorityName="Gozmany" authorityYear="1969" box="[161,487,727,753]" class="Insecta" family="Tineidae" genus="Merunympha" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[161,310,727,753]" italics="true" pageId="3" pageNumber="372">Merunympha</emphasis>
|
||
Gozmány 1969
|
||
</taxonomicName>
|
||
) has genitalia like
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[694,820,727,752]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[694,820,727,752]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
, with a female ovipositor that also is very long and with a strongly sclerotized and caudally acute sternum VII sterigmal plate (
|
||
<bibRefCitation author="Gozmany, L. A. & Vari, L." box="[950,1231,763,789]" pageId="3" pageNumber="372" refId="ref5924" refString="Gozmany, L. A. & Vari, L. (1973) The Tineidae of the Ethiopian Region. Transvaal Museum, Pretoria, (Transvaal Museum Memoirs, 18), 238 pp." type="book" year="1973">Gozmány and Varí 1973</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Janse, A. J. T." box="[1243,1372,763,789]" pageId="3" pageNumber="372" pagination="1 - 127" refId="ref6026" refString="Janse, A. J. T. (1968) On the types of South African Microlepidoptera. Vol. 1. Tineidae. Transvaal Museum Memoirs, Pretoria, 16, 1 - 127, 118 pls." type="journal article" year="1968">Janse 1968</bibRefCitation>
|
||
). The tineid generic keys of
|
||
<bibRefCitation author="Zagulajev, A. K." box="[398,590,799,825]" pageId="3" pageNumber="372" pagination="26 - 122" refId="ref6725" refString="Zagulajev, A. K. (1981) Fam. Tineidae. In: Identification Keys to Insects of European Russia. Vol. 4. Lepidoptera. Part 2. Akad. Nauk., St. Petersburg, pp. 20 - 93. [in Russian, English translation: pp. 26 - 122, Amerind Publishing, New Delhi (1989)]" type="journal article" year="1981">Zagulajev (1981)</bibRefCitation>
|
||
for
|
||
<collectingCountry box="[636,711,799,825]" name="Russia" pageId="3" pageNumber="372">Russia</collectingCountry>
|
||
key out to
|
||
<taxonomicName box="[835,964,799,825]" class="Insecta" family="Meessiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="family">Meessiinae</taxonomicName>
|
||
, but again it covers only a limited fauna in European
|
||
<collectingCountry box="[266,341,835,861]" name="Russia" pageId="3" pageNumber="372">Russia</collectingCountry>
|
||
of only 38 genera, and no tropical genera but only temperate Palearctic genera (see also
|
||
<bibRefCitation author="Petersen, G." pageId="3" pageNumber="372" pagination="55 - 176" refId="ref6229" refString="Petersen, G. (1957 - 1958) Die Genitalien der palaarktischen Tineiden (Lepidoptera: Tineidae). Beitrage zur Entomologie, 7 & 8, 55 - 176 + 338 - 379 + 557 - 595 (1957) & 111 - 118 + 398 - 430 (1958)." type="journal article" year="1957">Petersen 1957</bibRefCitation>
|
||
-58).
|
||
<bibRefCitation author="Regier, J. C. & Mitter, C. & Davis, D. R. & Harrison, T. L. & Sohn, J. - C. & Cummins, M. P. & Zwick, A. & Mitter, K. T." box="[268,488,871,897]" pageId="3" pageNumber="372" pagination="409 - 432" refId="ref6283" refString="Regier, J. C., Mitter, C., Davis, D. R., Harrison, T. L., Sohn, J. - C., Cummins, M. P., Zwick, A. & Mitter, K. T. (2014) A molecular phylogeny and revised classification for the oldest ditrysian moth lineages (Lepidoptera: Tineoidea), with implication for ancestral feeding habits of the mega-diverse Ditrysia. Systematic Entomology, 40, 409 - 432. https: // doi. org / 10.1111 / syen. 12110" type="journal article" year="2014">
|
||
Regier
|
||
<emphasis box="[350,406,871,897]" italics="true" pageId="3" pageNumber="372">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
provisionally elevated the subfamily
|
||
<taxonomicName box="[910,1036,871,897]" class="Insecta" family="Meessiinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="family">Meessiinae</taxonomicName>
|
||
to family status, as
|
||
<taxonomicName box="[1260,1389,871,897]" class="Insecta" family="Meessiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="family">Meessiidae</taxonomicName>
|
||
, but they suggest further studies are needed and this 'family' status is not followed herein.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,150,1941]" pageId="3" pageNumber="372">
|
||
There is superficial resemblance in wing form to the Southeast Asian genus
|
||
<taxonomicName authority="Moriuti (1982)" authorityName="Moriuti" authorityYear="1982" box="[1120,1436,943,969]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1120,1251,943,968]" italics="true" pageId="3" pageNumber="372">Tineovertex</emphasis>
|
||
<bibRefCitation author="Moriuti, S." box="[1263,1436,943,969]" pageId="3" pageNumber="372" pagination="162 - 171" refId="ref6134" refString="Moriuti, S. (1982) Tineidae. In: Inoue, h., Sugi, S., Kuroko, H., Moriuti, S., Kawabe, A. & Owada, M. (Eds.), Moths of Japan. Kodansha, Tokyo, pp. 162 - 171. [in Japanese]" type="book chapter" year="1982">Moriuti (1982)</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName box="[159,354,979,1005]" class="Insecta" family="Tineidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="subFamily" subFamily="Myrmecozelinae">Myrmecozelinae</taxonomicName>
|
||
), but basic characters differ greatly (maxillary palpi long and male genitalia more typically of tineid form in
|
||
<taxonomicName authorityName="Moriuti" authorityYear="1982" box="[313,444,1015,1040]" class="Insecta" family="Tineidae" genus="Tineovertex" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[313,444,1015,1040]" italics="true" pageId="3" pageNumber="372">Tineovertex</emphasis>
|
||
</taxonomicName>
|
||
, with vinculum having a long saccus) (
|
||
<bibRefCitation author="Huang, G. - H. & Hirowatari, T. & Wang, M." box="[887,1094,1015,1041]" pageId="3" pageNumber="372" pagination="1 - 12" refId="ref5961" refString="Huang, G. - H., Hirowatari, T. & Wang, M. (2011) A revision of the genus Tineovertex Moriuti (Insecta: Lepidoptera: Tineidae), with descriptions of five new species. Zootaxa, 2991 (1), 1 - 12. https: // doi. org / 10.11646 / zootaxa. 2991.1.1" type="journal article" year="2011">
|
||
Huang
|
||
<emphasis box="[971,1021,1015,1041]" italics="true" pageId="3" pageNumber="372">et al</emphasis>
|
||
. 2011
|
||
</bibRefCitation>
|
||
); however, both genera have a greatly elongated female ovipositor with setose papillae anales.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,150,1941]" pageId="3" pageNumber="372">
|
||
There is some similarity to the Perissomasticinae as well, for their lustrous forewings, the ubiquitous yellow head vertex of erect piliform setae, the compact and strongly sclerotized male genitalia with bifurcate uncus, and the small lateral coremata with spiculate scales in the male abdomen segment VI (
|
||
<bibRefCitation author="Gozmany, L. A. & Vari, L." box="[1026,1306,1159,1185]" pageId="3" pageNumber="372" refId="ref5924" refString="Gozmany, L. A. & Vari, L. (1973) The Tineidae of the Ethiopian Region. Transvaal Museum, Pretoria, (Transvaal Museum Memoirs, 18), 238 pp." type="book" year="1973">Gozmány and Vári 1973</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Yang, L. - L. & Wang, S. - X. & Li, H. - H." pageId="3" pageNumber="372" pagination="1 - 102" refId="ref6557" refString="Yang, L. - L., Wang, S. - X. & Li, H. - H. (2014) A taxonomic revision of the genus Edosa Walker, 1886 from China (Lepidoptera, Tineidae, Perissomasticinae). Zootaxa, 3777 (1), 1 - 102. https: // doi. org / 10.11646 / zootaxa. 3777.1.1" type="journal article" year="2014">
|
||
Yang
|
||
<emphasis box="[1380,1429,1159,1185]" italics="true" pageId="3" pageNumber="372">et al</emphasis>
|
||
. 2014
|
||
</bibRefCitation>
|
||
), characters especially evident in some African genera like
|
||
<taxonomicName authority="Gozmany 1959" authorityName="Gozmany" authorityYear="1959" box="[883,1227,1194,1221]" class="Insecta" family="Tineidae" genus="Sphallesthasis" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[883,1043,1195,1221]" italics="true" pageId="3" pageNumber="372">Sphallesthasis</emphasis>
|
||
Gozmány 1959
|
||
</taxonomicName>
|
||
, but the subfamily characteristic corethrogyne of the females and the typical stubby labial palpus are lacking in
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[1229,1355,1231,1256]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1229,1355,1231,1256]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
(labial palpus slender and long in
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[452,578,1267,1292]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[452,578,1267,1292]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,150,1941]" pageId="3" pageNumber="372">
|
||
Wing venation and female genitalia have resemblance also to the Philippine, Sulawesi and Solomons tineid genus
|
||
<taxonomicName authority="Sauber 1902" authorityName="Sauber" authorityYear="1902" box="[151,425,1339,1365]" class="Insecta" family="Tineidae" genus="Ischnuridia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[151,276,1339,1365]" italics="true" pageId="3" pageNumber="372">Ischnuridia</emphasis>
|
||
Sauber 1902
|
||
</taxonomicName>
|
||
, except the ovipositor has a spear-like cutting papillae anales (setose in
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[1224,1346,1339,1364]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1224,1346,1339,1364]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
) and an otherwise typical very long male saccus as in most tineids (versus the short or obsolete saccus in
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[1225,1347,1375,1400]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1225,1347,1375,1400]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation author="Davis, D. R. & Heppner, J. B." pageId="3" pageNumber="372" pagination="145 - 150" refId="ref5740" refString="Davis, D. R. & Heppner, J. B. (1987) New discoveries concerning Ischnuridia Sauber, a remarkable genus of Indo-Australian Tineidae (Lepidoptera). Tinea, 12 (Supplement), 145 - 150." type="journal article" year="1987">Davis and Heppner 1987</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Diakonoff, A. N." box="[363,538,1411,1437]" pageId="3" pageNumber="372" pagination="398 - 411" refId="ref5872" refString="Diakonoff, A. N. ([1968]) Tineidae. In: Microlepidoptera of the Philippine Islands. Bulletin of the United States National Museum, 257, pp. 266 - 293 + 396, 398 - 411 + 413 + 448 - 452. [1967]" type="journal article" year="1968">Diakonoff 1968</bibRefCitation>
|
||
). While female
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[708,830,1411,1436]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[708,830,1411,1436]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
have what appears to be a non-piercing ovipositor, in its place the sterigma is extremely sharp (
|
||
<figureCitation box="[558,646,1447,1473]" captionStart="FIGURE 11" captionStartId="8.[151,250,1943,1967]" captionTargetBox="[178,1434,181,1920]" captionTargetId="figure-29@8.[154,1434,181,1920]" captionTargetPageId="8" captionText="FIGURE 11. Vangatinea cambodiensis sp. nov., holotype female genitalia (Cambodia: Pursat) (ovipositor not completely extended, thus even longer than shown), with details, a) sterigma and ostium (gen. slide JBH-2998) (INUC)." figureDoi="http://doi.org/10.5281/zenodo.7836820" httpUri="https://zenodo.org/record/7836820/files/figure.png" pageId="3" pageNumber="372">Fig. 11a</figureCitation>
|
||
) (paired knife-like projections but asymmetrical in size), and likely serves as the piercing structure for egg deposition.
|
||
<taxonomicName authorityName="Sauber" authorityYear="1902" box="[625,750,1483,1509]" class="Insecta" family="Tineidae" genus="Ischnuridia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[625,750,1483,1509]" italics="true" pageId="3" pageNumber="372">Ischnuridia</emphasis>
|
||
</taxonomicName>
|
||
females may pierce into a soft hostplant like banana (
|
||
<taxonomicName box="[1330,1428,1483,1509]" class="Liliopsida" family="Musaceae" genus="Musa" kingdom="Plantae" order="Zingiberales" pageId="3" pageNumber="372" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis box="[1330,1390,1483,1508]" italics="true" pageId="3" pageNumber="372">Musa</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
) since an unnamed
|
||
<taxonomicName authorityName="Sauber" authorityYear="1902" box="[356,481,1519,1545]" class="Insecta" family="Tineidae" genus="Ischnuridia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[356,481,1519,1545]" italics="true" pageId="3" pageNumber="372">Ischnuridia</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<collectingCountry box="[591,775,1519,1545]" name="Solomon Islands" pageId="3" pageNumber="372">Solomon Islands</collectingCountry>
|
||
was captured on a banana plant (
|
||
<bibRefCitation author="Davis, D. R. & Heppner, J. B." box="[1144,1424,1519,1545]" pageId="3" pageNumber="372" pagination="145 - 150" refId="ref5740" refString="Davis, D. R. & Heppner, J. B. (1987) New discoveries concerning Ischnuridia Sauber, a remarkable genus of Indo-Australian Tineidae (Lepidoptera). Tinea, 12 (Supplement), 145 - 150." type="journal article" year="1987">Davis and Heppner 1987</bibRefCitation>
|
||
).
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[151,273,1555,1580]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[151,273,1555,1580]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
likewise has a long ovipositor and what appears to be a piercing structure on the sterigma, so possibly also for oviposition into a relatively soft hostplant.
|
||
<taxonomicName authorityName="Sauber" authorityYear="1902" box="[657,782,1591,1617]" class="Insecta" family="Tineidae" genus="Ischnuridia" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[657,782,1591,1617]" italics="true" pageId="3" pageNumber="372">Ischnuridia</emphasis>
|
||
</taxonomicName>
|
||
is one of many tineid genera unplaced to subfamily.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,150,1941]" pageId="3" pageNumber="372">
|
||
The characters for the new genus description may need revision, since only a single female from the three species is available for the female characters, and likewise the male characters are from only two of the species: however, the overlap of characters of several subfamilies, as noted above, makes subfamily placement in
|
||
<taxonomicName authorityName="Meyrick" authorityYear="1880" box="[1244,1386,1699,1725]" class="Insecta" family="Tineidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="subFamily" subFamily="Erechthiinae">Erechthiinae</taxonomicName>
|
||
still tentative.
|
||
<taxonomicName authorityName="Latreille" authorityYear="1810" box="[260,357,1735,1761]" class="Insecta" family="Tineidae" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="family">Tineidae</taxonomicName>
|
||
as a whole still require considerable further study on a world basis, and as noted above, such odd new discoveries as
|
||
<taxonomicName authorityName="Heppner & Bae" authorityYear="2023" box="[368,494,1771,1796]" class="Insecta" family="Tineidae" genus="Vangatinea" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[368,494,1771,1796]" italics="true" pageId="3" pageNumber="372">Vangatinea</emphasis>
|
||
</taxonomicName>
|
||
do not always fit neatly into existing subfamilies, such that current subfamilies may need revision.As also noted by
|
||
<bibRefCitation author="Robinson, G. S." box="[500,691,1807,1833]" pageId="3" pageNumber="372" refId="ref6388" refString="Robinson, G. S. (2009) Biology, Distribution and Diversity of Tineid Moths. Southdene, Kuala Lumpur, 143 pp., 16 pls." type="book" year="2009">Robinson (2009)</bibRefCitation>
|
||
, out of 340 named tineid genera worldwide over 200 genera remain of uncertain subfamily placement, especially so for tropical genera. Thus, the family requires a thorough revision of the subfamilies to account for all the unassociated genera, but such a task will take decades yet given the few taxonomists who could even venture to tackle such a quest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |