372 lines
48 KiB
XML
372 lines
48 KiB
XML
<document id="C31D84167260F5283EBBF5035E32C68F" ID-CLB-Dataset="6542" ID-DOI="10.1080/00222933.2013.840399" ID-GBIF-Dataset="a7ef425a-8153-40bb-9190-009cc1316c60" ID-ISSN="1464-5262" ID-Zenodo-Dep="4631325" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1616520776157" checkinUser="carolina" docAuthor="Hoch, Hannelore, Bourgoin, Thierry, Stelbrink, Björn & Wessel, Andreas" docDate="2013" docId="147287CE4B681E04FE27FA8FFCDADB20" docLanguage="en" docName="JNATHIST.48.15-16.919-935.pdf" docOrigin="Journal of Natural History 48 (15 - 16)" docSource="http://dx.doi.org/10.1080/00222933.2013.840399" docStyle="DocumentStyle:CD936EBDDF5274E0936345450CC2DAE4.5:JNatHist.2013-2015.journal_article.1cover" docStyleId="CD936EBDDF5274E0936345450CC2DAE4" docStyleName="JNatHist.2013-2015.journal_article.1cover" docStyleVersion="5" docTitle="Tsingya clarkei Hoch and Wessel 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="930" masterDocId="E84BFFB64B611E08FFE3FFADFFADDA68" masterDocTitle="Small giants from Madagascan caves: autapomorphic giantism in the new cave-dwelling planthopper Tsingya clarkei gen. nov., sp. nov. (Hemiptera: Fulgoroidea: Meenoplidae)" masterLastPageNumber="935" masterPageNumber="919" pageNumber="927" updateTime="1698934576448" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FF50D16A961A9F2D09ADC5B51D231B99">Small giants from Madagascan caves: autapomorphic giantism in the new cave-dwelling planthopper Tsingya clarkei gen. nov., sp. nov. (Hemiptera: Fulgoroidea: Meenoplidae)</mods:title>
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<mods:roleTerm id="88F4FB95736EDDDA5334D62FFCCC6C5A">Author</mods:roleTerm>
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<mods:namePart id="AF7E77218294B623B6BDB59DE7180CC8">Hoch, Hannelore</mods:namePart>
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<mods:affiliation id="9FDC17D0F53CD69319755968373B5742">Museum für Naturkunde - Leibniz-Institut für Evolutions- und Biodiversitätsforschung an der Humboldt-Universität zu Berlin, Berlin, Germany;</mods:affiliation>
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<mods:namePart id="788C404276697D7CB43A56FB3CC88FBB">Bourgoin, Thierry</mods:namePart>
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<mods:affiliation id="900BD019E00E11A27992A8CCF69B9EA6">Museum national d'Histoire naturelle, Département Systématique et Evolution, Laboratoire d’entomologie, Paris, France</mods:affiliation>
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<mods:roleTerm id="AF5C83ADF86F9F10154C40C44B6617CE">Author</mods:roleTerm>
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<mods:namePart id="ACF78C765AAEA351E4CB62DA2778426E">Stelbrink, Björn</mods:namePart>
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<mods:affiliation id="06CD6BF26EC4D0812A757C7762DC0FB8">Museum für Naturkunde - Leibniz-Institut für Evolutions- und Biodiversitätsforschung an der Humboldt-Universität zu Berlin, Berlin, Germany;</mods:affiliation>
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<mods:namePart id="0910B0B09F5CA183D363AE3CCB7281CF">Wessel, Andreas</mods:namePart>
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<mods:affiliation id="9AFB59D9A4144AB07BC556C969FF217F">Museum für Naturkunde - Leibniz-Institut für Evolutions- und Biodiversitätsforschung an der Humboldt-Universität zu Berlin, Berlin, Germany;</mods:affiliation>
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<mods:title id="67E8D4D0079F953CE7839A8E5514AB3F">Journal of Natural History</mods:title>
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<mods:part id="414A8406CCDCCE48CE6654E457F4AEAB">
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<mods:date id="C4EB08A745EED7CA5DD40AF7F0E2ABA2">2013</mods:date>
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<mods:number id="1AC6B6BAE3A693217C487F40E7C21500">2014-01-30</mods:number>
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<mods:number id="FB9DB6D458FDA0DA3C839F5FD1050E74">48</mods:number>
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<mods:number id="0C3C42FB024DC8690A043BC294E9E9D9">15 - 16</mods:number>
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<mods:url id="51B081A880FB6AA259F54EF60FE78B0B">http://dx.doi.org/10.1080/00222933.2013.840399</mods:url>
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<treatment id="147287CE4B681E04FE27FA8FFCDADB20" ID-DOI="http://doi.org/10.5281/zenodo.4783512" ID-GBIF-Taxon="182383660" ID-Zenodo-Dep="4783512" LSID="urn:lsid:plazi:treatment:147287CE4B681E04FE27FA8FFCDADB20" httpUri="http://treatment.plazi.org/id/147287CE4B681E04FE27FA8FFCDADB20" lastPageId="12" lastPageNumber="930" pageId="9" pageNumber="927">
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<subSubSection id="D4C165534B681E01FE27FA8FFC0EDF52" box="[452,931,1314,1338]" pageId="9" pageNumber="927" type="nomenclature">
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<paragraph id="9C6436D84B681E01FE27FA8FFC0EDF52" blockId="9.[452,931,1314,1370]" box="[452,931,1314,1338]" pageId="9" pageNumber="927">
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<heading id="C72C81B44B681E01FE27FA8FFC0EDF52" box="[452,931,1314,1338]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="927" reason="2">
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<taxonomicName id="5BDB4D5B4B681E01FE27FA8FFCECDF52" ID-CoL="59HKM" authority="Hoch and Wessel" authorityName="Hoch and Wessel" authorityYear="2013" box="[452,833,1314,1338]" class="Insecta" family="Meenoplidae" genus="Tsingya" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="927" phylum="Arthropoda" rank="species" species="clarkei" status="sp. nov.">
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<emphasis id="AEAFEACA4B681E01FE27FA8FFDDDDF52" bold="true" box="[452,624,1314,1338]" italics="true" pageId="9" pageNumber="927">Tsingya clarkei</emphasis>
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Hoch and Wessel
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</taxonomicName>
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<taxonomicNameLabel id="B59C57B14B681E01FCAAFA8FFC0EDF52" box="[841,931,1314,1338]" pageId="9" pageNumber="927" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="D4C165534B681E02FD8AFAECFCCFD8E1" lastPageId="10" lastPageNumber="928" pageId="9" pageNumber="927" type="description">
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<paragraph id="9C6436D84B681E01FD8AFAECFD52DF31" blockId="9.[452,931,1314,1370]" box="[617,767,1345,1370]" pageId="9" pageNumber="927">
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(
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<figureCitation id="04E02A5D4B681E01FD92FAECFD5BDF31" box="[625,758,1345,1370]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStartId-0="3.[212,279,1421,1443]" captionStartId-1="5.[212,279,627,649]" captionStartId-2="5.[212,279,1666,1688]" captionStartId-3="6.[144,211,754,776]" captionStartId-4="7.[212,279,693,715]" captionStartId-5="7.[213,280,1193,1215]" captionStartId-6="7.[209,276,1695,1717]" captionStartId-7="8.[144,211,776,798]" captionStartId-8="8.[148,215,1356,1378]" captionTargetBox-0="[212,1172,168,1389]" captionTargetBox-1="[220,1162,168,595]" captionTargetBox-2="[324,1060,1358,1634]" captionTargetBox-3="[158,1104,177,718]" captionTargetBox-4="[531,852,168,661]" captionTargetBox-5="[536,851,745,1161]" captionTargetBox-6="[546,834,1248,1664]" captionTargetBox-7="[144,1104,171,744]" captionTargetBox-8="[340,916,894,1325]" captionTargetId-0="figure-1@3.[212,1172,168,1389]" captionTargetId-1="figure-215@5.[220,1165,168,596]" captionTargetId-2="figure-233@5.[324,1060,1355,1634]" captionTargetId-3="figure-268@6.[144,1105,168,722]" captionTargetId-4="figure-1@7.[531,852,168,661]" captionTargetId-5="figure-17@7.[536,851,745,1161]" captionTargetId-6="figure-35@7.[546,834,1248,1664]" captionTargetId-7="figure-71@8.[144,1104,168,744]" captionTargetId-8="figure-92@8.[340,916,894,1325]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="7" captionTargetPageId-6="7" captionTargetPageId-7="8" captionTargetPageId-8="8" captionText-0="Figure 1. Map of Madagascar (composed and modified from satellite photos provided by Google Earth™), the simplified geology shows the limestone areas (red = Tertiary limestones + marls and chalks; yellow = Mesozoic limestones + marls (incl. “tsingy”), modified after Du Puy and Moat 1996: Figure 1), area enlarged (bottom right): southernmost Petit Tsingy (“Little Tsingy”) section of the Tsingy de Bemaraha National Park east of Bekopaka, just north of the Manambolo River. Upper right corner: Tsingya clarkei sp. nov. (still images from the wildlife documentary “Voyage to the center of the stone”: Gabriel 2003), see also supplementary movie." captionText-1="Figure 2. (A) Head and thorax, dorsal aspect; (B) head, ventral aspect. Scale bar 0.5 mm." captionText-2="Figure 3. Right tegmen, maximal aspect. Scale bar 1 mm." captionText-3="Figure 4. (A) Male genital segment, caudal aspect; (B) male genitalia, in situ, left lateral aspect. Scale bar 0.5 mm." captionText-4="Figure 5. Male genitalia, dissected. Male anal segment, dorsal aspect. Scale bar 0.1 mm." captionText-5="Figure 6. Malegenitalia, dissected.Aedeagus (A)dorsal aspect,(B) ventral aspect.Scale bar 0.1 mm." captionText-6="Figure 7. Male genitalia, dissected. Parameres, ventral aspect. Scale bar 0.1 mm." captionText-7="Figure 8. External female genitalia, in situ. (A) Right lateral aspect; (B) ventral aspect. Scale bar 0.5 mm." captionText-8="Figure 9. Internal female genitalia, dissected, right lateral aspect. Scale bar 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4631329" figureDoi-1="http://doi.org/10.5281/zenodo.4631333" figureDoi-2="http://doi.org/10.5281/zenodo.4631337" figureDoi-3="http://doi.org/10.5281/zenodo.4631339" figureDoi-4="http://doi.org/10.5281/zenodo.4631343" figureDoi-5="http://doi.org/10.5281/zenodo.4631345" figureDoi-6="http://doi.org/10.5281/zenodo.4631349" figureDoi-7="http://doi.org/10.5281/zenodo.4631351" figureDoi-8="http://doi.org/10.5281/zenodo.4631355" httpUri-0="https://zenodo.org/record/4631329/files/figure.png" httpUri-1="https://zenodo.org/record/4631333/files/figure.png" httpUri-2="https://zenodo.org/record/4631337/files/figure.png" httpUri-3="https://zenodo.org/record/4631339/files/figure.png" httpUri-4="https://zenodo.org/record/4631343/files/figure.png" httpUri-5="https://zenodo.org/record/4631345/files/figure.png" httpUri-6="https://zenodo.org/record/4631349/files/figure.png" httpUri-7="https://zenodo.org/record/4631351/files/figure.png" httpUri-8="https://zenodo.org/record/4631355/files/figure.png" pageId="9" pageNumber="927">Figures 1–9</figureCitation>
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)
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</paragraph>
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<paragraph id="9C6436D84B681E01FF37FA31FEFEDFDC" blockId="9.[212,1171,1436,1691]" box="[212,339,1436,1460]" pageId="9" pageNumber="927">
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<heading id="C72C81B44B681E01FF37FA31FEFEDFDC" box="[212,339,1436,1460]" fontSize="10" level="3" pageId="9" pageNumber="927" reason="5">
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<emphasis id="AEAFEACA4B681E01FF37FA31FEFEDFDC" box="[212,339,1436,1460]" italics="true" pageId="9" pageNumber="927">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="9C6436D84B681E01FF37FA69FD2ADF94" blockId="9.[212,1171,1436,1691]" pageId="9" pageNumber="927">
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<emphasis id="AEAFEACA4B681E01FF37FA69FE9EDFB4" box="[212,307,1476,1500]" italics="true" pageId="9" pageNumber="927">Habitus.</emphasis>
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Unusually large meenoplid, tegmina long, steeply tectiform, small compound eyes, pale bodily pigmentation.
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</paragraph>
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<paragraph id="9C6436D84B681E01FF1DF9A9FDD5DC33" blockId="9.[212,1171,1436,1691]" pageId="9" pageNumber="927">
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Body length. Anterior margin of head to distal margin of tegmina. Males. 10.0–
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<quantity id="5B239B3D4B681E01FF37F989FE97DC54" box="[212,314,1572,1596]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="9" pageNumber="927" unit="mm" value="10.6">10.6 mm</quantity>
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(
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<emphasis id="AEAFEACA4B681E01FEAEF989FEF7DC54" box="[333,346,1572,1596]" italics="true" pageId="9" pageNumber="927">n</emphasis>
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= 2). Female.
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<quantity id="5B239B3D4B681E01FDF3F989FDDBDC54" box="[528,630,1572,1596]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="9" pageNumber="927" unit="mm" value="11.5">11.5 mm</quantity>
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(
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<emphasis id="AEAFEACA4B681E01FD69F989FD3ADC54" box="[650,663,1572,1596]" italics="true" pageId="9" pageNumber="927">n</emphasis>
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= 1). Tip of head to distal margin of anal segment. Males. 6.9–7.0 mm (
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<emphasis id="AEAFEACA4B681E01FDC8F9E9FD95DC34" box="[555,568,1604,1628]" italics="true" pageId="9" pageNumber="927">n</emphasis>
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= 2).
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</paragraph>
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<paragraph id="9C6436D84B681E01FF1DF9CEFC18DCF3" blockId="9.[212,1171,1436,1691]" pageId="9" pageNumber="927">
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Colouration (
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<figureCitation id="04E02A5D4B681E01FE74F9CEFE5BDC13" box="[407,502,1635,1659]" captionStart="Figure 1" captionStartId="3.[212,279,1421,1443]" captionTargetBox="[212,1172,168,1389]" captionTargetId="figure-1@3.[212,1172,168,1389]" captionTargetPageId="3" captionText="Figure 1. Map of Madagascar (composed and modified from satellite photos provided by Google Earth™), the simplified geology shows the limestone areas (red = Tertiary limestones + marls and chalks; yellow = Mesozoic limestones + marls (incl. “tsingy”), modified after Du Puy and Moat 1996: Figure 1), area enlarged (bottom right): southernmost Petit Tsingy (“Little Tsingy”) section of the Tsingy de Bemaraha National Park east of Bekopaka, just north of the Manambolo River. Upper right corner: Tsingya clarkei sp. nov. (still images from the wildlife documentary “Voyage to the center of the stone”: Gabriel 2003), see also supplementary movie." figureDoi="http://doi.org/10.5281/zenodo.4631329" httpUri="https://zenodo.org/record/4631329/files/figure.png" pageId="9" pageNumber="927">Figure 1</figureCitation>
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). Body pale (white) throughout, except for compound eyes, these pigmented lightly red; tegmina translucent, venation white.
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</paragraph>
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<paragraph id="9C6436D84B6B1E02FF59FF05FD26DA88" blockId="10.[144,1104,168,224]" pageId="10" pageNumber="928">
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Configuration and proportions of head and thorax incl. legs, tegmina and wings as described for the genus (
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<figureCitation id="04E02A5D4B6B1E02FE2BFF65FD9FDA88" box="[456,562,200,224]" captionStart="Figure 2" captionStartId="5.[212,279,627,649]" captionTargetBox="[220,1162,168,595]" captionTargetId="figure-215@5.[220,1165,168,596]" captionTargetPageId="5" captionText="Figure 2. (A) Head and thorax, dorsal aspect; (B) head, ventral aspect. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4631333" httpUri="https://zenodo.org/record/4631333/files/figure.png" pageId="10" pageNumber="928">Figures 2</figureCitation>
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and
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<figureCitation id="04E02A5D4B6B1E02FD8DFF65FDD6DA88" box="[622,635,200,224]" captionStart="Figure 3" captionStartId="5.[212,279,1666,1688]" captionTargetBox="[324,1060,1358,1634]" captionTargetId="figure-233@5.[324,1060,1355,1634]" captionTargetPageId="5" captionText="Figure 3. Right tegmen, maximal aspect. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4631337" httpUri="https://zenodo.org/record/4631337/files/figure.png" pageId="10" pageNumber="928">3</figureCitation>
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).
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</paragraph>
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<paragraph id="9C6436D84B6B1E02FF73FEA0FE76D82C" blockId="10.[144,1104,269,580]" pageId="10" pageNumber="928">
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<emphasis id="AEAFEACA4B6B1E02FF73FEA0FE97DB4D" box="[144,314,269,293]" italics="true" pageId="10" pageNumber="928">Male genitalia.</emphasis>
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(
|
||
<figureCitation id="04E02A5D4B6B1E02FEA8FEA0FE7DDB4D" box="[331,464,269,293]" captionStart-0="Figure 4" captionStart-1="Figure 5" captionStart-2="Figure 6" captionStart-3="Figure 7" captionStartId-0="6.[144,211,754,776]" captionStartId-1="7.[212,279,693,715]" captionStartId-2="7.[213,280,1193,1215]" captionStartId-3="7.[209,276,1695,1717]" captionTargetBox-0="[158,1104,177,718]" captionTargetBox-1="[531,852,168,661]" captionTargetBox-2="[536,851,745,1161]" captionTargetBox-3="[546,834,1248,1664]" captionTargetId-0="figure-268@6.[144,1105,168,722]" captionTargetId-1="figure-1@7.[531,852,168,661]" captionTargetId-2="figure-17@7.[536,851,745,1161]" captionTargetId-3="figure-35@7.[546,834,1248,1664]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionTargetPageId-3="7" captionText-0="Figure 4. (A) Male genital segment, caudal aspect; (B) male genitalia, in situ, left lateral aspect. Scale bar 0.5 mm." captionText-1="Figure 5. Male genitalia, dissected. Male anal segment, dorsal aspect. Scale bar 0.1 mm." captionText-2="Figure 6. Malegenitalia, dissected.Aedeagus (A)dorsal aspect,(B) ventral aspect.Scale bar 0.1 mm." captionText-3="Figure 7. Male genitalia, dissected. Parameres, ventral aspect. Scale bar 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4631339" figureDoi-1="http://doi.org/10.5281/zenodo.4631343" figureDoi-2="http://doi.org/10.5281/zenodo.4631345" figureDoi-3="http://doi.org/10.5281/zenodo.4631349" httpUri-0="https://zenodo.org/record/4631339/files/figure.png" httpUri-1="https://zenodo.org/record/4631343/files/figure.png" httpUri-2="https://zenodo.org/record/4631345/files/figure.png" httpUri-3="https://zenodo.org/record/4631349/files/figure.png" pageId="10" pageNumber="928">Figures 4–7</figureCitation>
|
||
). Genital and anal segments as described for the genus. Parameres with ventral branch more strongly developed, dorsal branch differentiated into a strong, finger-shaped process bent medially, and a short, stout, apically blunt appendage, densely beset with setae, directed laterodorsally. Aedeagus with dorsal and ventral processes arising from a short, wide common base: dorsal trilobate process forming a half-pipe, with median part hood-shaped, distally subacute with phallotreme located medially at apex, phallotreme exposed ventrally; lateral lobes apically rounded, in lateral aspect slightly surpassing median lobe; ventral aedeagal process tongue-shaped, dorsally at base concave, distally taeniform, narrow, apically acute, its tip curved dorsally.
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6B1E02FF73FDDCFCCFD8E1" blockId="10.[144,866,625,649]" box="[144,866,625,649]" pageId="10" pageNumber="928">
|
||
<emphasis id="AEAFEACA4B6B1E02FF73FDDCFEE2D8E1" box="[144,335,625,649]" italics="true" pageId="10" pageNumber="928">Female genitalia.</emphasis>
|
||
(
|
||
<figureCitation id="04E02A5D4B6B1E02FE83FDDCFE67D8E1" box="[352,458,625,649]" captionStart="Figure 8" captionStartId="8.[144,211,776,798]" captionTargetBox="[144,1104,171,744]" captionTargetId="figure-71@8.[144,1104,168,744]" captionTargetPageId="8" captionText="Figure 8. External female genitalia, in situ. (A) Right lateral aspect; (B) ventral aspect. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4631351" httpUri="https://zenodo.org/record/4631351/files/figure.png" pageId="10" pageNumber="928">Figures 8</figureCitation>
|
||
and
|
||
<figureCitation id="04E02A5D4B6B1E02FDE5FDDCFDBED8E1" box="[518,531,625,649]" captionStart="Figure 9" captionStartId="8.[148,215,1356,1378]" captionTargetBox="[340,916,894,1325]" captionTargetId="figure-92@8.[340,916,894,1325]" captionTargetPageId="8" captionText="Figure 9. Internal female genitalia, dissected, right lateral aspect. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4631355" httpUri="https://zenodo.org/record/4631355/files/figure.png" pageId="10" pageNumber="928">9</figureCitation>
|
||
). As described for the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4C165534B6B1E02FF73FD7BFDACD9FD" pageId="10" pageNumber="928" type="diagnosis">
|
||
<paragraph id="9C6436D84B6B1E02FF73FD7BFF51D886" blockId="10.[144,1104,726,917]" box="[144,252,726,750]" pageId="10" pageNumber="928">
|
||
<heading id="C72C81B44B6B1E02FF73FD7BFF51D886" box="[144,252,726,750]" fontSize="10" level="3" pageId="10" pageNumber="928" reason="5">
|
||
<emphasis id="AEAFEACA4B6B1E02FF73FD7BFF51D886" box="[144,252,726,750]" italics="true" pageId="10" pageNumber="928">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6B1E02FF73FD53FDACD9FD" blockId="10.[144,1104,726,917]" pageId="10" pageNumber="928">
|
||
<taxonomicName id="5BDB4D5B4B6B1E02FF73FD53FE98D97E" authority="Hoch & Bourgoin & Stelbrink & Wessel, 2013" authorityName="Hoch & Bourgoin & Stelbrink & Wessel" authorityYear="2013" box="[144,309,766,790]" class="Insecta" family="Meenoplidae" genus="Tsingy" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="928" phylum="Arthropoda" rank="species" species="clarkei" status="sp. nov.">
|
||
<emphasis id="AEAFEACA4B6B1E02FF73FD53FE98D97E" box="[144,309,766,790]" italics="true" pageId="10" pageNumber="928">Tsingy clarkei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B59C57B14B6B1E02FEA7FD53FE09D97E" box="[324,420,766,790]" pageId="10" pageNumber="928" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is characterized by a unique combination of characters pertaining to the male genital structures: parameres bifurcate with ventral branch more strongly developed, aedeagus with a dorsal, trilobate process forming a halfpipe, and phallotreme situated medially at apex and a ventral, tongue-shaped, taeniform and apically acute process.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4C165534B6B1E02FF73FC4EFD8ADEEA" pageId="10" pageNumber="928" type="etymology">
|
||
<paragraph id="9C6436D84B6B1E02FF73FC4EFEA4D993" blockId="10.[144,1103,995,1154]" box="[144,265,995,1019]" pageId="10" pageNumber="928">
|
||
<heading id="C72C81B44B6B1E02FF73FC4EFEA4D993" box="[144,265,995,1019]" fontSize="10" level="3" pageId="10" pageNumber="928" reason="5">
|
||
<emphasis id="AEAFEACA4B6B1E02FF73FC4EFEA4D993" box="[144,265,995,1019]" italics="true" pageId="10" pageNumber="928">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6B1E02FF73FBA7FD8ADEEA" blockId="10.[144,1103,995,1154]" pageId="10" pageNumber="928">
|
||
<materialsCitation id="2CB33C854B6B1E02FF73FBA7FD8EDEEA" ID-GBIF-Occurrence="3117066301" country="Australia" location="School of Zoology" pageId="10" pageNumber="928" specimenCount="1" stateProvince="Tasmania">
|
||
The species is named in honour of the collector, Arthur Clarke (Honorary Research Associate in the
|
||
<location id="990460034B6B1E02FEB1FB87FD81DE2A" LSID="urn:lsid:plazi:treatment:147287CE4B681E04FE27FA8FFCDADB20:990460034B6B1E02FEB1FB87FD81DE2A" box="[338,556,1066,1090]" country="Australia" name="School of Zoology" pageId="10" pageNumber="928" stateProvince="Tasmania">School of Zoology</location>
|
||
, University of
|
||
<collectingRegion id="5E1FF83A4B6B1E02FD38FB87FCE6DE2A" box="[731,843,1066,1090]" country="Australia" name="Tasmania" pageId="10" pageNumber="928">Tasmania</collectingRegion>
|
||
) in Hobart,
|
||
<collectingRegion id="5E1FF83A4B6B1E02FC03FB87FBE2DE2A" box="[992,1103,1066,1090]" country="Australia" name="Tasmania" pageId="10" pageNumber="928">Tasmania</collectingRegion>
|
||
(
|
||
<collectingCountry id="E4CC76484B6B1E02FF74FBE7FEA9DE0A" box="[151,260,1098,1122]" name="Australia" pageId="10" pageNumber="928">Australia</collectingCountry>
|
||
), ardent cave explorer, dedicated cave ecologist and biospeleologist, reliable caving buddy and good friend
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4C165534B6B1E02FF73FB62FEC4DFE6" pageId="10" pageNumber="928" type="distribution">
|
||
<paragraph id="9C6436D84B6B1E02FF73FB62FEB9DE8F" blockId="10.[144,1104,1231,1422]" box="[144,276,1231,1255]" pageId="10" pageNumber="928">
|
||
<heading id="C72C81B44B6B1E02FF73FB62FEB9DE8F" box="[144,276,1231,1255]" fontSize="10" level="3" pageId="10" pageNumber="928" reason="5">
|
||
<emphasis id="AEAFEACA4B6B1E02FF73FB62FEB9DE8F" box="[144,276,1231,1255]" italics="true" pageId="10" pageNumber="928">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6B1E02FF73FB5AFEC4DFE6" blockId="10.[144,1104,1231,1422]" pageId="10" pageNumber="928">
|
||
Known only from Anjohy Manitsy (translates as “Cold Cave”) in the southernmost Petit
|
||
<taxonomicName id="5BDB4D5B4B6B1E02FEF0FABAFECFDF47" box="[275,354,1303,1327]" class="Insecta" family="Meenoplidae" genus="Tsingy" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="928" phylum="Arthropoda" rank="genus">Tsingy</taxonomicName>
|
||
(“Little
|
||
<taxonomicName id="5BDB4D5B4B6B1E02FE2EFABAFDB1DF47" box="[461,540,1303,1327]" class="Insecta" family="Meenoplidae" genus="Tsingy" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="928" phylum="Arthropoda" rank="genus">Tsingy</taxonomicName>
|
||
”) section of the
|
||
<taxonomicName id="5BDB4D5B4B6B1E02FD0BFABAFC70DF47" authority="de Bemaraha" authorityName="de Bemaraha" box="[744,989,1303,1327]" class="Insecta" family="Meenoplidae" genus="Tsingy" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="928" phylum="Arthropoda" rank="genus">Tsingy de Bemaraha</taxonomicName>
|
||
National Park east of Bekopaka, just north of the Manambolo River, in western
|
||
<collectingCountry id="E4CC76484B6B1E02FF73FAFBFE89DF06" box="[144,292,1366,1390]" name="Madagascar" pageId="10" pageNumber="928">Madagascar</collectingCountry>
|
||
. Likely to be a narrow range endemic,
|
||
<taxonomicName id="5BDB4D5B4B6B1E02FCFEFAFAFC3FDF07" authorityName="Hoch & Bourgoin & Stelbrink & Wessel" authorityYear="2013" box="[797,914,1367,1391]" class="Insecta" family="Meenoplidae" genus="Tsingy" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="928" phylum="Arthropoda" rank="species" species="clarkei">
|
||
<emphasis id="AEAFEACA4B6B1E02FCFEFAFAFC3FDF07" box="[797,914,1367,1391]" italics="true" pageId="10" pageNumber="928">T. clarkei</emphasis>
|
||
</taxonomicName>
|
||
is known only from
|
||
<collectingCountry id="E4CC76484B6B1E02FF32FADBFEC8DFE6" box="[209,357,1398,1422]" name="Madagascar" pageId="10" pageNumber="928">Madagascar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4C165534B6B1E03FF73FA71FE29DF93" lastPageId="11" lastPageNumber="929" pageId="10" pageNumber="928" type="biology_ecology">
|
||
<paragraph id="9C6436D84B6B1E02FF73FA71FF47DF9C" blockId="10.[144,1104,1500,1627]" box="[144,234,1500,1524]" pageId="10" pageNumber="928">
|
||
<heading id="C72C81B44B6B1E02FF73FA71FF47DF9C" box="[144,234,1500,1524]" fontSize="10" level="3" pageId="10" pageNumber="928" reason="5">
|
||
<emphasis id="AEAFEACA4B6B1E02FF73FA71FF47DF9C" box="[144,234,1500,1524]" italics="true" pageId="10" pageNumber="928">Ecology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6B1E03FF73F9AEFE4ED877" blockId="10.[144,1104,1500,1627]" lastBlockId="11.[212,1172,168,1531]" lastPageId="11" lastPageNumber="929" pageId="10" pageNumber="928">
|
||
<bibRefCitation id="F84A4B294B6B1E02FF73F9AEFE82DC73" author="Clarke A" box="[144,303,1539,1563]" pageId="10" pageNumber="928" pagination="33 - 42" refId="ref6660" refString="Clarke A. 2003. Cultural diversity, karst ecosystems and karst management in the Tsingy de Bemaraha Parc of western Madagascar. Australas Cave Karst Manage Ass J. 52: 33 - 42." type="journal article" year="2003">Clarke (2003)</bibRefCitation>
|
||
reported that some substantial trees occur in the dense thickets of vegetation formed in the canyons and fault grabens between the elevated blocks of pinnacle karst. However, above the caves the karst surface is generally devoid of large trees and almost barren. Among the perilously razor sharp karren flutes, the surface vegetation in the tsingy itself is unique with many endemic xerophytic and/or water storage plants. Although the Bemaraha tsingy receives torrential downpours in the wet season, very little water remains on the surface and in the dry season the only moisture for 6–7 months is the nightly dew, so the karst is essentially an arid environment for much of the year. The “arid” flora of the Bemaraha karst surface typically includes cactiform succulents such as
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FD06FEC5FCF5DBE8" box="[741,856,360,384]" class="Magnoliopsida" family="Euphorbiaceae" genus="Euphorbia" kingdom="Plantae" order="Malpighiales" pageId="11" pageNumber="929" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="AEAFEACA4B6A1E03FD06FEC5FCF5DBE8" box="[741,856,360,384]" italics="true" pageId="11" pageNumber="929">Euphorbia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FC71FEC5FBB8DBE8" box="[914,1045,360,384]" class="Magnoliopsida" family="Apocynaceae" genus="Cynanchum" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="929" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="AEAFEACA4B6A1E03FC71FEC5FBB8DBE8" box="[914,1045,360,384]" italics="true" pageId="11" pageNumber="929">Cynanchum</emphasis>
|
||
</taxonomicName>
|
||
, numerous bulbous stemmed plants or trees that hold water in their internal pith (e.g.
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FF37FE05FE61DBA8" authority=", Adenia" authorityName="Adenia" box="[212,460,424,448]" class="Magnoliopsida" family="Apocynaceae" genus="Pachypodium" kingdom="Plantae" order="Gentianales" pageId="11" pageNumber="929" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="AEAFEACA4B6A1E03FF37FE05FE61DBA8" box="[212,460,424,448]" italics="true" pageId="11" pageNumber="929">Pachypodium, Adenia</emphasis>
|
||
</taxonomicName>
|
||
and the occasional baobab), the many thorned, narrow or spiny leafed xerophytic or xerophilous species such as
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FCA8FE6AFC1BDBB7" box="[843,950,455,479]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="11" pageNumber="929" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="AEAFEACA4B6A1E03FCA8FE6AFC1BDBB7" box="[843,950,455,479]" italics="true" pageId="11" pageNumber="929">Pandanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FC17FE6AFBF2DBB7" box="[1012,1119,455,479]" class="Reptilia" family="Teiidae" genus="Dracaena" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="929" phylum="Chordata" rank="genus">
|
||
<emphasis id="AEAFEACA4B6A1E03FC17FE6AFBF2DBB7" box="[1012,1119,455,479]" italics="true" pageId="11" pageNumber="929">Dracaena</emphasis>
|
||
</taxonomicName>
|
||
and the unusual leafless plants such as
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FDBBFE4AFD45DB97" authorityName="N.J.Jacquin" authorityYear="1797" box="[600,744,487,511]" class="Magnoliopsida" family="Burseraceae" genus="Commiphora" kingdom="Plantae" order="Sapindales" pageId="11" pageNumber="929" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="AEAFEACA4B6A1E03FDBBFE4AFD45DB97" box="[600,744,487,511]" italics="true" pageId="11" pageNumber="929">Commiphora</emphasis>
|
||
</taxonomicName>
|
||
with its photosynthesizing trunks and branches (
|
||
<bibRefCitation id="F84A4B294B6A1E03FEABFDAAFE7FD877" author="Clarke A" box="[328,466,519,543]" pageId="11" pageNumber="929" pagination="33 - 42" refId="ref6660" refString="Clarke A. 2003. Cultural diversity, karst ecosystems and karst management in the Tsingy de Bemaraha Parc of western Madagascar. Australas Cave Karst Manage Ass J. 52: 33 - 42." type="journal article" year="2003">Clarke 2003</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6A1E03FF1DFD8AFE79D9B5" blockId="11.[212,1172,168,1531]" pageId="11" pageNumber="929">
|
||
Access to the enclosed cave collection site involved passing through a lushly forested canyon, before commencing a traverse of
|
||
<quantity id="5B239B3D4B6A1E03FCECFDEAFCDDD837" box="[783,880,583,607]" metricMagnitude="1" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="11" pageNumber="929" unit="m" value="45.0" valueMax="50.0" valueMin="40.0">40–50 m</quantity>
|
||
along a narrow 1.0–
|
||
<quantity id="5B239B3D4B6A1E03FBBFFDEAFB3ED837" box="[1116,1171,583,607]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="929" unit="m" value="1.5">1.5m</quantity>
|
||
wide,
|
||
<quantity id="5B239B3D4B6A1E03FEC3FDCAFED5D817" box="[288,376,615,639]" metricMagnitude="1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="11" pageNumber="929" unit="m" value="35.0" valueMax="40.0" valueMin="30.0">30–40m</quantity>
|
||
high diaclase (roofless cave) rift with abundant tree roots. The entrance elevation (altitude) of the diaclase is approximately
|
||
<quantity id="5B239B3D4B6A1E03FC21FD2BFBA2D8F6" box="[962,1039,646,670]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="11" pageNumber="929" unit="m" value="130.0">130 m</quantity>
|
||
above sea level. Among the green-leafed foliage in the canyon and in the outer extremities of the diaclase, a number of vertebrate species were observed, including three smaller species of lemur: the common brown lemur
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FC8BFD4BFBBAD896" baseAuthorityName="E.Geoffroy Saint-Hilaire" baseAuthorityYear="1796" box="[872,1047,742,766]" class="Mammalia" family="Lemuridae" genus="Eulemur" kingdom="Animalia" order="Primates" pageId="11" pageNumber="929" phylum="Chordata" rank="species" species="fulvus">
|
||
<emphasis id="AEAFEACA4B6A1E03FC8BFD4BFBBAD896" box="[872,1047,742,766]" italics="true" pageId="11" pageNumber="929">Eulemur fulvus</emphasis>
|
||
</taxonomicName>
|
||
, the grey mouse lemur
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FE66FCABFDDCD976" baseAuthorityName="J.F.Miller" baseAuthorityYear="1777" box="[389,625,774,798]" class="Mammalia" family="Cheirogaleidae" genus="Microcebus" kingdom="Animalia" order="Primates" pageId="11" pageNumber="929" phylum="Chordata" rank="species" species="murinus">
|
||
<emphasis id="AEAFEACA4B6A1E03FE66FCABFDDCD976" box="[389,625,774,798]" italics="true" pageId="11" pageNumber="929">Microcebus murinus</emphasis>
|
||
</taxonomicName>
|
||
and a large nocturnal species the Milne-Edwards’ weasel lemur
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FE0BFC8BFD6ED956" baseAuthorityName="Forsyth Major" baseAuthorityYear="1894" box="[488,707,806,830]" class="Mammalia" family="Lepilemuridae" genus="Lepilemur" kingdom="Animalia" order="Primates" pageId="11" pageNumber="929" phylum="Chordata" rank="species" species="edwardsi">
|
||
<emphasis id="AEAFEACA4B6A1E03FE0BFC8BFD6ED956" box="[488,707,806,830]" italics="true" pageId="11" pageNumber="929">Lepilemur edwardsi</emphasis>
|
||
</taxonomicName>
|
||
and several reptiles. Along with one of the leaf-tailed
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FE94FCEBFE49D936" authorityName="A.M.C.Dumeril" authorityYear="1806" box="[375,484,838,862]" class="Reptilia" family="Gekkonidae" genus="Uroplatus" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="929" phylum="Chordata" rank="genus">
|
||
<emphasis id="AEAFEACA4B6A1E03FE94FCEBFE49D936" box="[375,484,838,862]" italics="true" pageId="11" pageNumber="929">Uroplatus</emphasis>
|
||
</taxonomicName>
|
||
geckos, three species of chameleons were noted; two species of
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FF1AFCCBFECAD916" box="[249,359,870,894]" class="Reptilia" family="Chamaeleonidae" genus="Brookesia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="929" phylum="Chordata" rank="genus">
|
||
<emphasis id="AEAFEACA4B6A1E03FF1AFCCBFECAD916" box="[249,359,870,894]" italics="true" pageId="11" pageNumber="929">Brookesia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FE9EFCCBFDE0D916" baseAuthorityName="Boettger" baseAuthorityYear="1880" box="[381,589,870,894]" class="Reptilia" family="Chamaeleonidae" genus="Brookesia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="929" phylum="Chordata" rank="species" species="ebenaui">
|
||
<emphasis id="AEAFEACA4B6A1E03FE9EFCCBFDE0D916" box="[381,589,870,894]" italics="true" pageId="11" pageNumber="929">Brookesia ebenaui</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FD70FCCBFC2FD916" baseAuthorityName="Angel" baseAuthorityYear="1933" box="[659,898,870,894]" class="Reptilia" family="Chamaeleonidae" genus="Brookesia" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="929" phylum="Chordata" rank="species" species="perarmata">
|
||
<emphasis id="AEAFEACA4B6A1E03FD70FCCBFC2FD916" box="[659,898,870,894]" italics="true" pageId="11" pageNumber="929">Brookesia perarmata</emphasis>
|
||
</taxonomicName>
|
||
, and the much longer Malagasy Giant Chameleon
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FDC4FC2BFD45D9F6" box="[551,744,902,926]" class="Reptilia" family="Chamaeleonidae" genus="Furcifer" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="929" phylum="Chordata" rank="species" species="oustelati">
|
||
<emphasis id="AEAFEACA4B6A1E03FDC4FC2BFD45D9F6" box="[551,744,902,926]" italics="true" pageId="11" pageNumber="929">Furcifer oustelati</emphasis>
|
||
</taxonomicName>
|
||
. A colubrid snake
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FC32FC2BFE63D9D6" authorityName="Domergue" authorityYear="1987" class="Reptilia" family="Lamprophiidae" genus="Madagascarophis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="929" phylum="Chordata" rank="subSpecies" species="colubrinus" subSpecies="occidentalis">
|
||
<emphasis id="AEAFEACA4B6A1E03FC32FC2BFE63D9D6" italics="true" pageId="11" pageNumber="929">Madagascarophis colubrinus occidentalis</emphasis>
|
||
</taxonomicName>
|
||
was also spotted in a
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FD35FC0BFCECD9D6" box="[726,833,934,958]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="11" pageNumber="929" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="AEAFEACA4B6A1E03FD35FC0BFCECD9D6" box="[726,833,934,958]" italics="true" pageId="11" pageNumber="929">Pandanus</emphasis>
|
||
</taxonomicName>
|
||
tree (A. Clarke, unpublished records, pers. comm.).
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6A1E03FF1DFC48FE29DF93" blockId="11.[212,1172,168,1531]" pageId="11" pageNumber="929">
|
||
In the subdued daylight/twilight inner reaches of the Anjohy Manitsy diaclase rift heading into the roofed cave, several specimens of a locally known so-called nocturnal butterfly (a noctuid moth) were found on the rift walls. In the outer twilight section of the cave, two species of land snails were seen among tree roots and rootlets on the moist sandy floor:
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FD4EFBC8FB39DE15" authorityName="Emberton" authorityYear="2002" box="[685,1172,1125,1149]" class="Gastropoda" family="Ariophantidae" genus="Kalidos" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="929" phylum="Mollusca" rank="species" species="griffithshauchleri">
|
||
<emphasis id="AEAFEACA4B6A1E03FD4EFBC8FC63DE15" box="[685,974,1125,1149]" italics="true" pageId="11" pageNumber="929">Kalidos griffithshauchleri</emphasis>
|
||
(Ariophantidae)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FEE8FB28FD15DEF5" authorityName="Fischer-Piette & Bedoucha" authorityYear="1965" box="[267,696,1157,1181]" class="Gastropoda" family="Cyclophoridae" genus="Acroptychia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Architaenioglossa" pageId="11" pageNumber="929" phylum="Mollusca" rank="species" species="bathiei">
|
||
<emphasis id="AEAFEACA4B6A1E03FEE8FB28FE40DEF5" box="[267,493,1157,1181]" italics="true" pageId="11" pageNumber="929">Acroptychia bathiei</emphasis>
|
||
(Cyclophoridae)
|
||
</taxonomicName>
|
||
. Two frogs were observed in this same outer part of cave: the grey-legged, red and brown-backed
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FC24FB08FEC5DEB4" baseAuthorityName="Grandidier" baseAuthorityYear="1872" class="Amphibia" family="Mantellidae" genus="Mantella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="929" phylum="Chordata" rank="species" species="betsileo">
|
||
<emphasis id="AEAFEACA4B6A1E03FC24FB08FB39DED5" box="[967,1172,1189,1213]" italics="true" pageId="11" pageNumber="929">Mantella betsileo</emphasis>
|
||
(Mantelidae)
|
||
</taxonomicName>
|
||
and a speckled brown frog, possibly the Mascarene Ridged Frog
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FF37FB48FD2ADE94" baseAuthorityName="Dumeril & Bibron" baseAuthorityYear="1841" box="[212,647,1252,1277]" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" kingdom="Animalia" order="Anura" pageId="11" pageNumber="929" phylum="Chordata" rank="species" species="mascareniensis">
|
||
<emphasis id="AEAFEACA4B6A1E03FF37FB48FDA5DE95" box="[212,520,1253,1277]" italics="true" pageId="11" pageNumber="929">Ptychadena mascareniensis</emphasis>
|
||
(Ranidae)
|
||
</taxonomicName>
|
||
. In this outer twilight zone, there was also a dark brown coloured flatworm
|
||
<quantity id="5B239B3D4B6A1E03FDB4FAA9FD67DF74" box="[599,714,1284,1308]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="11" pageNumber="929" unit="cm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 cm</quantity>
|
||
long, with broadly flattened bulbous head, which may possibly predate on the cavernicolous species including the planthoppers. All three specimens of
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FD46FAE9FCF7DF34" authorityName="Hoch and Wessel" authorityYear="2013" box="[677,858,1348,1372]" class="Insecta" family="Meenoplidae" genus="Tsingya" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="929" phylum="Arthropoda" rank="species" species="clarkei">
|
||
<emphasis id="AEAFEACA4B6A1E03FD46FAE9FCF7DF34" box="[677,858,1348,1372]" italics="true" pageId="11" pageNumber="929">Tsingya clarkei</emphasis>
|
||
</taxonomicName>
|
||
were found in the inner high humidity dark transition zone of the cave, collected from the subterranean roots of an unidentified surface tree and from smaller rootlets in clay soil adjacent to the cave wall. Among the other cave-dwelling invertebrates in Anjohy Manitsy, there were unidentified species of depigmented isopods, entomobryid springtails and millipedes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4C165534B6A1E03FF37F991FE86DCF3" pageId="11" pageNumber="929" type="discussion">
|
||
<paragraph id="9C6436D84B6A1E03FF37F991FE9BDC3C" blockId="11.[212,1171,1596,1691]" box="[212,310,1596,1620]" pageId="11" pageNumber="929">
|
||
<heading id="C72C81B44B6A1E03FF37F991FE9BDC3C" box="[212,310,1596,1620]" fontSize="10" level="3" pageId="11" pageNumber="929" reason="5">
|
||
<emphasis id="AEAFEACA4B6A1E03FF37F991FE9BDC3C" box="[212,310,1596,1620]" italics="true" pageId="11" pageNumber="929">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6A1E03FF37F9CEFE86DCF3" blockId="11.[212,1171,1596,1691]" pageId="11" pageNumber="929">
|
||
With 10.0–
|
||
<quantity id="5B239B3D4B6A1E03FEB3F9CEFE1FDC13" box="[336,434,1635,1659]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="929" unit="mm" value="11.5">11.5 mm</quantity>
|
||
in length,
|
||
<taxonomicName id="5BDB4D5B4B6A1E03FDCAF9CEFD7EDC13" authority="Hoch & Bourgoin & Stelbrink & Wessel, 2013" authorityName="Hoch & Bourgoin & Stelbrink & Wessel" authorityYear="2013" box="[553,723,1635,1659]" class="Insecta" family="Meenoplidae" genus="Tsingya" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="929" phylum="Arthropoda" rank="species" species="clarkei" status="sp. nov.">
|
||
<emphasis id="AEAFEACA4B6A1E03FDCAF9CEFD7EDC13" box="[553,723,1635,1659]" italics="true" pageId="11" pageNumber="929">Tsingya clarkei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B59C57B14B6A1E03FD3AF9CEFC9FDC13" box="[729,818,1635,1659]" pageId="11" pageNumber="929" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the largest known meenoplid to date.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D4C165534B6D1E04FF73FF05FCDADB20" pageId="12" pageNumber="930" type="materials_examined">
|
||
<paragraph id="9C6436D84B6D1E04FF73FF05FECEDAA8" blockId="12.[144,1104,168,328]" box="[144,355,168,192]" pageId="12" pageNumber="930">
|
||
<heading id="C72C81B44B6D1E04FF73FF05FECEDAA8" box="[144,355,168,192]" fontSize="10" level="3" pageId="12" pageNumber="930" reason="5">
|
||
<emphasis id="AEAFEACA4B6D1E04FF73FF05FECEDAA8" box="[144,355,168,192]" italics="true" pageId="12" pageNumber="930">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6D1E04FF73FF7DFF5EDB40" blockId="12.[144,1104,168,328]" pageId="12" pageNumber="930">
|
||
<materialsCitation id="2CB33C854B6D1E04FF73FF7DFF5EDB40" ID-GBIF-Occurrence="3117066302" collectingDate="2003-05" collectionCode="MNHN" collectorName="Arthur Clarke" country="Madagascar" county="western Madagascar" location="Anjohy Manitsy" municipality="Bekopaka" pageId="12" pageNumber="930" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="4360887A4B6D1E04FF73FF7DFF57DA80" box="[144,250,208,232]" pageId="12" pageNumber="930" type="holotype">Holotype</typeStatus>
|
||
male:
|
||
<collectingCountry id="E4CC76484B6D1E04FEBDFF7DFD82DA80" box="[350,559,208,232]" name="Madagascar" pageId="12" pageNumber="930">MADAGASCAR</collectingCountry>
|
||
,
|
||
<location id="990460034B6D1E04FDA4FF7DFCA2DA80" LSID="urn:lsid:plazi:treatment:147287CE4B681E04FE27FA8FFCDADB20:990460034B6D1E04FDA4FF7DFCA2DA80" box="[583,783,208,232]" country="Madagascar" county="western Madagascar" municipality="Bekopaka" name="Anjohy Manitsy" pageId="12" pageNumber="930">Anjohy Manitsy</location>
|
||
,
|
||
<location id="990460034B6D1E04FCC4FF7DFC13DA80" LSID="urn:lsid:plazi:treatment:147287CE4B681E04FE27FA8FFCDADB20:990460034B6D1E04FCC4FF7DFC13DA80" box="[807,958,208,232]" country="Madagascar" county="western Madagascar" municipality="Bekopaka" name="Petit Tsingy" pageId="12" pageNumber="930">Petit Tsingy</location>
|
||
,
|
||
<taxonomicName id="5BDB4D5B4B6D1E04FC35FF7DFEAADB60" authority="de Bemaraha" authorityName="de Bemaraha" class="Insecta" family="Meenoplidae" genus="Tsingy" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="930" phylum="Arthropoda" rank="genus">Tsingy de Bemaraha</taxonomicName>
|
||
, east of
|
||
<collectingMunicipality id="7C00ACA24B6D1E04FE8FFF5DFE4CDB60" box="[364,481,240,264]" pageId="12" pageNumber="930">Bekopaka</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="75054E544B6D1E04FE13FF5DFD73DB60" box="[496,734,240,264]" pageId="12" pageNumber="930">western Madagascar</collectingCounty>
|
||
,
|
||
<date id="E86510184B6D1E04FD0FFF5DFCC9DB60" box="[748,868,240,264]" pageId="12" pageNumber="930" value="2003-05">
|
||
<collectingDate id="F821E9F04B6D1E04FD0FFF5DFCC9DB60" box="[748,868,240,264]" pageId="12" pageNumber="930" value="2003-05">May 2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="312E530E4B6D1E04FC91FF5DFBBADB60" box="[882,1047,240,264]" pageId="12" pageNumber="930">Arthur Clarke</collectorName>
|
||
leg.,
|
||
<collectionCode id="FACAAE1D4B6D1E04FF73FEBDFF5DDB40" box="[144,240,272,296]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="930">MNHN</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="9C6436D84B6D1E04FF59FE9DFCDADB20" blockId="12.[144,1104,168,328]" box="[186,887,304,328]" pageId="12" pageNumber="930">
|
||
<materialsCitation id="2CB33C854B6D1E04FF59FE9DFCDEDB20" ID-GBIF-Occurrence="3117066303" box="[186,883,304,328]" collectingDate="2003-05" collectionCode="MNHN" collectorName="Arthur Clarke" country="Madagascar" county="western Madagascar" location="Anjohy Manitsy" municipality="Bekopaka" pageId="12" pageNumber="930" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
|
||
<typeStatus id="4360887A4B6D1E04FF59FE9DFE81DB20" box="[186,300,304,328]" pageId="12" pageNumber="930" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
<specimenCount id="8ADDFD514B6D1E04FEDAFE9DFE25DB20" box="[313,392,304,328]" pageId="12" pageNumber="930" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="8ADDFD514B6D1E04FE76FE9DFE54DB20" box="[405,505,304,328]" pageId="12" pageNumber="930" type="female">1 female</specimenCount>
|
||
, same data as holotype,
|
||
<collectionCode id="FACAAE1D4B6D1E04FCF0FE9DFCDEDB20" box="[787,883,304,328]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="930">MNHN</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |