204 lines
18 KiB
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204 lines
18 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.869.33997" ID-GBIF-Dataset="5b500434-09e3-4370-82a1-1eb789d9d58a" ID-PMC="PMC6690892" ID-Pensoft-Pub="1313-2970-869-19" ID-Pensoft-UUID="C44C602AB0235FB2922DC2A4915C3288" ID-PubMed="31413659" ID-ZooBank="B7C88A64C06E4B67A352F2F9C8FB0D1C" ModsDocID="1313-2970-869-19" checkinTime="1565056339550" checkinUser="pensoft" docAuthor="Ranarilalatiana, Tolotra, Raveloson Ravaomanarivo, Lala Harivelo & Bergsten, Johannes" docDate="2019" docId="0DA77899943B5226BBA57EB36328B122" docLanguage="en" docName="ZooKeys 869: 19-90" docOrigin="ZooKeys 869" docSource="http://dx.doi.org/10.3897/zookeys.869.33997" docTitle="Copelatus baculiformis Guignot 1955" docType="treatment" docVersion="4" lastPageNumber="19" masterDocId="5E52FF84FFE5FF9193131A454057613C" masterDocTitle="Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species" masterLastPageNumber="90" masterPageNumber="19" pageNumber="19" updateTime="1668167671827" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic revision of the genus Copelatus of Madagascar (Coleoptera, Dytiscidae, Copelatinae): the non- erichsonii group species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ranarilalatiana, Tolotra</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Raveloson Ravaomanarivo, Lala Harivelo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bergsten, Johannes</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>869</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>19</mods:start>
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<mods:end>90</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.869.33997</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.869.33997</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-869-19</mods:identifier>
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<mods:identifier type="ZooBank">B7C88A64C06E4B67A352F2F9C8FB0D1C</mods:identifier>
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<mods:identifier type="Pensoft-UUID">C44C602AB0235FB2922DC2A4915C3288</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="158995558" LSID="urn:lsid:plazi:treatment:0DA77899943B5226BBA57EB36328B122" httpUri="http://treatment.plazi.org/id/0DA77899943B5226BBA57EB36328B122" lastPageNumber="19" pageId="0" pageNumber="19">
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<subSubSection pageId="0" pageNumber="19" type="nomenclature">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName LSID="0da77899-943b-5226-bba5-7eb36328b122" authority="Guignot, 1955" authorityName="Guignot" authorityYear="1955" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">Copelatus baculiformis Guignot, 1955</taxonomicName>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7B</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="reference_group">
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<paragraph pageId="0" pageNumber="19">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">Copelatus baculiformis</taxonomicName>
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Guignot, 1955b: 193.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="type locality">
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<paragraph pageId="0" pageNumber="19">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="19">Madagascar, Massif Ankaratra, Manjakatompo, alt. 1700-1800 m.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">Type information from original description.</paragraph>
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<paragraph pageId="0" pageNumber="19">based on a single female specimen (holotype), collected December 1951 by R. Benoist.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="type material">
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<paragraph pageId="0" pageNumber="19">Type material studied.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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<emphasis bold="true" pageId="0" pageNumber="19">Antananarivo. Vakinankaratra: Ambatolampy</emphasis>
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: -HT ♀ (MNHN "coll. Guignot"): // Data in NHRS | JLKB | 000030226 // Madagascar: Massif An- | karatra 1700/1800 Man- | jakatompo XII-51 Benoist // Type [red label] // Guignot det., 1954 |
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
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<emphasis italics="true" pageId="0" pageNumber="19">baculiformis</emphasis>
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n. sp. | Type ♀ //
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="diagnosis">
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<paragraph pageId="0" pageNumber="19">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Small size (4 mm). Elytra uniformly dark brown ferrugineous, without a basal testaceous area (
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7B</figureCitation>
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) which separates the species from small species of
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
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</taxonomicName>
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. The absence of deep impressed elytral striae (remnants of four elytral striae present) separates this species from all other
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
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</taxonomicName>
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of Madagascar except
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus peridinus</emphasis>
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</taxonomicName>
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, a larger species (5.7-6.6 mm).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="description">
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<paragraph pageId="0" pageNumber="19">Description.</paragraph>
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<paragraph pageId="0" pageNumber="19">(based on holotype ♀):</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Body length 4 mm. Body shape elongate oval and dark brown to blackish ferrugineous. Head uniformly dark brown ferrugineous to slightly darker posteriorly inside eyes, with thin sparse punctation. Pronotum dark brown ferrugineous, same colour medially and laterally but darker along anterior and posterior third. Disc of pronotum less densely punctuated, posterolateral corners with dense superficial strioles. The entire dorsal surface covered with a microsculpture. Elytra uniformly coloured in same dark brown to blackish ferrugineous colour as anterior and posterior parts of pronotum (
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Habitus, dorsal view. A Male: Copelatus peridinus B Female: Copelatus baculiformis C Female: Copelatus sp. 2 (Andasibe) D Female: Copelatus distinguendus." figureDoi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19">Fig. 7B</figureCitation>
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). Impressed striae absent but four elytral rows of impressed points present. Innermost, a presumed first row completely lacking, row 2, 3, and 4 visible from base but row 3 less distinct than 2 and 4. Row 5 very vague and only visible posteriorly. Elytra densely covered with punctures which laterally of approximately the third row of points is replaced by strioles (probably only in the female). Appendages testaceous.
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</paragraph>
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<caption doi="10.3897/zookeys.869.33997.figure7" httpUri="https://binary.pensoft.net/fig/322932" pageId="0" pageNumber="19" start="Figure 7" startId="F7">
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<paragraph pageId="0" pageNumber="19">
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Figure 7. Habitus, dorsal view.
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<emphasis bold="true" pageId="0" pageNumber="19">A</emphasis>
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Male:
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus peridinus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="peridinus">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus peridinus</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="19">B</emphasis>
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Female:
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus baculiformis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="19">C</emphasis>
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Female:
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
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</taxonomicName>
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sp. 2 (Andasibe)
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<emphasis bold="true" pageId="0" pageNumber="19">D</emphasis>
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Female:
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus distinguendus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="distinguendus">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus distinguendus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="19">
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Ventral side testaceous to weakly infuscated. Prosternal process carinate also onto apical process. Lateral parts of metaventrite ("metasternal wings") rather broad. Metacoxal lines anteriorly diverging and abbreviated before metaventral margin. Metacoxa with fine and long longitudinal strioles, continuing onto abdominal ventrites, and with 6-7 transverse
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<normalizedToken originalValue="“wrinkles”">"wrinkles"</normalizedToken>
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anterolaterally.
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</paragraph>
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<paragraph pageId="0" pageNumber="19">Male: unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="distribution">
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<paragraph pageId="0" pageNumber="19">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Madagascar, central highlands, only known from type locality Manjakatompo, Ankaratra Massif (
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution maps of Copelatus species. A C. marginipennis (circle), C. baculiformis (square) B C. mahajanga (circle), C. sp. 2 (square) C C. pulchellus (circle), C. peridinus (square) D C. distinguendus (circle), C. sp. 1 (square)." figureDoi="10.3897/zookeys.869.33997.figure11" httpUri="https://binary.pensoft.net/fig/322936" pageId="0" pageNumber="19">Fig. 11A</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="habitat">
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<paragraph pageId="0" pageNumber="19">Habitat and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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Unknown, but according to original description collected at at an altitude of 1700-1800 m. See
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.350.6127" author="Hjalmarsson, AE" journalOrPublisher="ZooKeys" pageId="0" pageNumber="19" pagination="21 - 45" refId="B38" refString="Hjalmarsson, AE, Bukontaite, R, Ranarilalatiana, T, Randriamihaja, JH, Bergsten, J, 2013. Taxonomic revision of Madagascan Rhantus (Coleoptera, Dytiscidae, Colymbetinae) with an emphasis on Manjakatompo as a conservation priority. . ZooKeys 350: 21 - 45" title="Taxonomic revision of Madagascan Rhantus (Coleoptera, Dytiscidae, Colymbetinae) with an emphasis on Manjakatompo as a conservation priority." url="https://doi.org/10.3897/zookeys.350.6127" volume="350" year="2013">Hjalmarsson et al. (2013)</bibRefCitation>
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for a description of the locality Manjakatompo and its conservation priority.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="19" type="comments">
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<paragraph pageId="0" pageNumber="19">Comments.</paragraph>
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<paragraph pageId="0" pageNumber="19">
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No other specimen than the female holotype is known of this species and it is a bit of a "mystery species". We have conducted fieldwork at the type locality Manjakatompo multiple times (2011, 2012, 2014, and 2016) but never found any specimens resembling this species. The species belongs in the
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<taxonomicName lsidName="hydroporoides" pageId="0" pageNumber="19" rank="species" species="hydroporoides">
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<emphasis italics="true" pageId="0" pageNumber="19">hydroporoides</emphasis>
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</taxonomicName>
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species group of
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus</emphasis>
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</taxonomicName>
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, but two other Malagasy species placed in this group have turned out to be misidentified
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Madaglymbus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madaglymbus" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="19">Madaglymbus</emphasis>
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</taxonomicName>
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or
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="19">Exocelina</emphasis>
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</taxonomicName>
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(see above).
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus baculiformis" order="Coleoptera" pageId="0" pageNumber="19" phylum="Arthropoda" rank="species" species="baculiformis">
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<emphasis italics="true" pageId="0" pageNumber="19">Copelatus baculiformis</emphasis>
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</taxonomicName>
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was described by Guignot the same year (1955) that he described
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<taxonomicName lsidName="C. bilunatus" pageId="0" pageNumber="19" rank="species" species="bilunatus">
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<emphasis italics="true" pageId="0" pageNumber="19">C. bilunatus</emphasis>
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</taxonomicName>
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, considered mislabeled (
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<bibRefCitation author="Balke, M" editor="Guilbert, E" journalOrPublisher="Memoires du Museum national d'Histoire naturelle" pageId="0" pageNumber="19" pagination="1 - 524" refId="B6" refString="Balke, M, Hajek, J, Hendrich, L, Wewalka, G, 2014. Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183-192]. . In: Guilbert, E, Robillardt, T, Jourdan, H, Grancolas, P, Eds., Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia. . Memoires du Museum national d'Histoire naturelle 206: 1 - 524" title="Exocelina nehoue n. sp. from New Caledonia, with a new synonym and new collecting records for other species in the genus (Coleoptera: Dytiscidae) [183 - 192]." volume="206" volumeTitle="Zoologica Neocaledonica 8. Biodiversity Studies in New Caledonia." year="2014">Balke et al. 2014</bibRefCitation>
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) from the Zimmermann collection. It is certainly possible that also
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<taxonomicName lsidName="C. baculiformis" pageId="0" pageNumber="19" rank="species" species="baculiformis">
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<emphasis italics="true" pageId="0" pageNumber="19">C. baculiformis</emphasis>
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</taxonomicName>
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is based on a mislabeled specimen, but in contrast to
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<taxonomicName lsidName="C. bilunatus" pageId="0" pageNumber="19" rank="species" species="bilunatus">
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<emphasis italics="true" pageId="0" pageNumber="19">C. bilunatus</emphasis>
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</taxonomicName>
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the type locality data is more exact; Massif Ankaratra (mountain), Manjakatompo (locality), 1700-1800 m (altitude in meters), XII-51 (collecting month and year), and R. Benoist (collector), speaking against such a mistake. Based on general external morphology, body shape and lack of striae the species resembles
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<taxonomicName lsidName="C. peridinus" pageId="0" pageNumber="19" rank="species" species="peridinus">
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<emphasis italics="true" pageId="0" pageNumber="19">C. peridinus</emphasis>
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</taxonomicName>
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. It is much smaller and therefore likely not conspecific, but they could be closely related, as well as with the female sequenced from Andasibe (NHRS-JLKB000065698).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |