treatments-xml/data/0D/4A/4E/0D4A4E06FFEFA73AFF08FB85FA1274BA.xml

<document id="5E6C5B63505856BAD80FF9898C6C560A" ID-DOI="10.11646/zootaxa.4750.4.3" ID-GBIF-Dataset="140b7cbb-b422-42a3-95cb-04062d0453e5" ID-ISSN="1175-5326" ID-Zenodo-Dep="3708838" ID-ZooBank="5FE0773B-3C35-4D30-8CE7-F1691ABA5F83" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584084501875" checkinUser="plazi" docAuthor="Acevedo, Aldemar A., Armesto, Orlando &amp; Palma, R. Eduardo" docDate="2020" docId="0D4A4E06FFEFA73AFF08FB85FA1274BA" docLanguage="en" docName="zootaxa.4750.4.3.pdf" docOrigin="Zootaxa 4750 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pristimantis ardilae Acevedo &amp; Armesto &amp; Palma 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="510" masterDocId="F173367EFFECA731FF9FFF98FF8B7301" masterDocTitle="Two new species of Pristimantis (Anura: Craugastoridae) with notes on the distribution of the genus in northeastern Colombia" masterLastPageNumber="523" masterPageNumber="499" pageNumber="502" updateTime="1702002986539" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="61C115A27AA3C711ABC979A769A271F6">Two new species of Pristimantis (Anura: Craugastoridae) with notes on the distribution of the genus in northeastern Colombia</mods:title>
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<mods:namePart id="8C4C5BE8573A5F4DD7A1053C0476F4E5">Acevedo, Aldemar A.</mods:namePart>
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<mods:namePart id="21AD6DA441D13088C22F01C0F2BF3ED2">Armesto, Orlando</mods:namePart>
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<mods:namePart id="02A81C36B305FB638C78D6F618794078">Palma, R. Eduardo</mods:namePart>
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<treatment id="0D4A4E06FFEFA73AFF08FB85FA1274BA" ID-DOI="http://doi.org/10.5281/zenodo.3717962" ID-GBIF-Taxon="162486207" ID-Zenodo-Dep="3717962" LSID="urn:lsid:plazi:treatment:0D4A4E06FFEFA73AFF08FB85FA1274BA" httpUri="http://treatment.plazi.org/id/0D4A4E06FFEFA73AFF08FB85FA1274BA" lastPageId="11" lastPageNumber="510" pageId="3" pageNumber="502">
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<paragraph id="855CFF10FFEFA732FF08FB85FE797739" blockId="3.[151,498,1053,1115]" box="[151,498,1053,1080]" pageId="3" pageNumber="502">
<heading id="DE14487CFFEFA732FF08FB85FE797739" bold="true" box="[151,498,1053,1080]" fontSize="11" level="1" pageId="3" pageNumber="502" reason="1">
<emphasis id="B7972302FFEFA732FF08FB85FE797739" bold="true" box="[151,498,1053,1080]" pageId="3" pageNumber="502">
<emphasis id="B7972302FFEFA732FF08FB85FE067739" bold="true" box="[151,397,1053,1080]" italics="true" pageId="3" pageNumber="502">
<taxonomicName id="42E38493FFEFA732FF08FB85FE0C7739" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[151,391,1053,1080]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">Pristimantis ardilae</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="ACA49E79FFEFA732FE0AFB86FE797739" box="[405,498,1054,1080]" pageId="3" pageNumber="502" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="CDF9AC9BFFEFA732FF08FBD9FE97775A" box="[151,284,1089,1115]" pageId="3" pageNumber="502" type="description">
<paragraph id="855CFF10FFEFA732FF08FBD9FE97775A" blockId="3.[151,498,1053,1115]" box="[151,284,1089,1115]" pageId="3" pageNumber="502">
(
<figureCitation id="1DD8E395FFEFA732FF00FBD9FE9F775A" box="[159,276,1089,1115]" captionStart="FIGURE 4" captionStartId="7.[151,250,1926,1951]" captionTargetBox="[155,1432,991,1899]" captionTargetId="figure@7.[152,1435,989,1900]" captionTargetPageId="7" captionText="FIGURE 4. Photos of the specimens in life. A. Pristimantis ardilae sp. nov., uncollected animals. B. Pristimantis bowara sp. nov., holoytpe, MCNUP-H 165. Lower zone of the buffer zone of PNN Tamá, Colombia." figureDoi="http://doi.org/10.5281/zenodo.3708846" httpUri="https://zenodo.org/record/3708846/files/figure.png" pageId="3" pageNumber="502">Figure 4A</figureCitation>
)
</paragraph>
</subSubSection>
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<paragraph id="855CFF10FFEFA732FF08FB12FC2F77EA" blockId="3.[151,1437,1161,2016]" pageId="3" pageNumber="502">
<materialsCitation id="358BF54DFFEFA732FF08FB12FC2F77EA" ID-GBIF-Occurrence="2575874647" collectingDate="2010-09-10" collectionCode="PNNT" collectorName="An &amp; Aldemar A. Acevedo &amp; Karen Silva &amp; Rosmery Franco &amp; San Antonio &amp; Toledo &amp; Tama National Natural Park" elevation="612" latitude="7.162172" longLatPrecision="1" longitude="-72.25254" pageId="3" pageNumber="502" specimenCode="MCNUP-H 117" specimenCount="612" typeStatus="holotype">
<emphasis id="B7972302FFEFA732FF08FB12FE8F77A5" bold="true" box="[151,260,1162,1188]" pageId="3" pageNumber="502">
<typeStatus id="5A5841B2FFEFA732FF08FB12FE8F77A5" box="[151,260,1162,1188]" pageId="3" pageNumber="502" type="holotype">Holotype</typeStatus>
</emphasis>
. 
<collectorName id="28169AC6FFEFA732FE90FB12FEBA77A5" box="[271,305,1162,1188]" pageId="3" pageNumber="502">An</collectorName>
adult female (
<specimenCode id="D545576BFFEFA732FE51FB11FD0977A2" box="[462,642,1161,1187]" collectionCode="MCNUP-H" pageId="3" pageNumber="502">MCNUP-H 117</specimenCode>
, 
<figureCitation id="1DD8E395FFEFA732FD12FB11FC8A77A5" box="[653,769,1161,1188]" captionStart="FIGURE 5" captionStartId="10.[151,250,1680,1705]" captionTargetBox="[160,1419,196,1645]" captionTargetId="figure@10.[151,1436,181,1655]" captionTargetPageId="10" captionText="FIGURE 5. Photos of the holotype specimens. A. Pristimantis ardilae sp. nov. (MCNUP-H 117) B. Pristimantis bowara sp. nov. (MCNUP-H 165)." figureDoi="http://doi.org/10.5281/zenodo.3708848" httpUri="https://zenodo.org/record/3708848/files/figure.png" pageId="3" pageNumber="502">Figure 5A</figureCitation>
), collected on 
<date id="F15DD9D0FFEFA732FC3EFB11FB0977A2" box="[929,1154,1161,1187]" pageId="3" pageNumber="502" value="2010-09-10">
<collectingDate id="E1192038FFEFA732FC3EFB11FB0977A2" box="[929,1154,1161,1187]" pageId="3" pageNumber="502" value="2010-09-10">September 10, 2010</collectingDate>
</date>
, by 
<collectorName id="28169AC6FFEFA732FB33FB11FA1377A2" box="[1196,1432,1161,1188]" pageId="3" pageNumber="502">Aldemar A. Acevedo</collectorName>
, 
<collectorName id="28169AC6FFEFA732FF08FB36FE9577C6" box="[151,286,1197,1224]" pageId="3" pageNumber="502">Karen Silva</collectorName>
, and 
<collectorName id="28169AC6FFEFA732FEC8FB36FD9B77C9" box="[343,528,1198,1224]" pageId="3" pageNumber="502">Rosmery Franco</collectorName>
in the locality of 
<collectorName id="28169AC6FFEFA732FD4DFB35FCD677C6" box="[722,861,1197,1223]" pageId="3" pageNumber="502">San Antonio</collectorName>
(
<geoCoordinate id="E0D799D7FFEFA732FCF5FB35FC5D77C9" box="[874,982,1197,1224]" degrees="7.162172" direction="north" orientation="latitude" pageId="3" pageNumber="502" precision="1" value="7.162172">7.162172</geoCoordinate>
<geoCoordinate id="E0D799D7FFEFA732FC4AFB35FBDB77C9" box="[981,1104,1197,1224]" degrees="72.25254" direction="west" orientation="longitude" pageId="3" pageNumber="502" precision="1" value="-72.25254">–72.25254</geoCoordinate>
; 
<specimenCount id="93E53499FFEFA732FBC5FB35FB2077C9" box="[1114,1195,1197,1224]" pageId="3" pageNumber="502" type="generic">
<quantity id="421B52F5FFEFA732FBC5FB35FB2C77C9" box="[1114,1191,1197,1224]" metricMagnitude="2" metricUnit="m" metricValue="6.12" pageId="3" pageNumber="502" unit="m" value="612.0">
<elevation id="0ECE1823FFEFA732FBC5FB35FB2C77C9" box="[1114,1191,1197,1224]" metricMagnitude="2" metricUnit="m" metricValue="6.12" pageId="3" pageNumber="502" unit="m" value="612.0">612 m</elevation>
</quantity>
.
</specimenCount>
a.s.l.), municipality of 
<collectorName id="28169AC6FFEFA732FF08FB49FF6F77EA" box="[151,228,1233,1259]" pageId="3" pageNumber="502">Toledo</collectorName>
in the buffer zone of the 
<collectorName id="28169AC6FFEFA732FE60FB49FCB277EA" box="[511,825,1233,1259]" pageId="3" pageNumber="502">Tamá National Natural Park</collectorName>
(
<collectionCode id="E3F267D5FFEFA732FCD6FB49FC1277EA" box="[841,921,1233,1259]" pageId="3" pageNumber="502">PNNT</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="855CFF10FFEFA732FF58FB6EFEBE769E" blockId="3.[151,1437,1161,2016]" pageId="3" pageNumber="502">
<materialsCitation id="358BF54DFFEFA732FF58FB6EFB7E7635" ID-GBIF-Occurrence="2575874645" collectionCode="MCNUP-H" pageId="3" pageNumber="502" specimenCode="MCNUP-H 121, MCNUP-H 166, MCNUP-H 163" specimenCount="2" typeStatus="paratype">
<emphasis id="B7972302FFEFA732FF58FB6EFEB47611" bold="true" box="[199,319,1270,1296]" pageId="3" pageNumber="502">
<typeStatus id="5A5841B2FFEFA732FF58FB6EFEB47611" box="[199,319,1270,1296]" pageId="3" pageNumber="502" type="paratype">Paratypes</typeStatus>
</emphasis>
. Two adult females (
<specimenCode id="D545576BFFEFA732FDA6FB6DFD7A760E" box="[569,753,1269,1295]" collectionCode="MCNUP-H" pageId="3" pageNumber="502">MCNUP-H 121</specimenCode>
and 157), and two adult males (
<specimenCode id="D545576BFFEFA732FBECFB6DFAA0760E" box="[1139,1323,1269,1295]" collectionCode="MCNUP-H" pageId="3" pageNumber="502">MCNUP-H 166</specimenCode>
and 253) collected at the type locality on the same date of the 
<typeStatus id="5A5841B2FFEFA732FD6FFA81FCDD7632" box="[752,854,1305,1331]" pageId="3" pageNumber="502" type="holotype">holotype</typeStatus>
. One adult female (
<specimenCode id="D545576BFFEFA732FBABFA81FB677635" box="[1076,1260,1305,1332]" collectionCode="MCNUP-H" pageId="3" pageNumber="502">MCNUP-H 163</specimenCode>
)
</materialsCitation>
, 
<materialsCitation id="358BF54DFFEFA732FA9EFA81FEBA769E" ID-GBIF-Occurrence="2575874644" collectingDate="2013-05-03" collectionCode="PNNT" collectorName="Aldemar A. Acevedo &amp; La Vega" elevation="465" latitude="7.13612" longLatPrecision="5" longitude="-72.1843" pageId="3" pageNumber="502" specimenCode="MCNUP-H 673" specimenCount="465" typeStatus="paratype">
and two adult females (
<specimenCode id="D545576BFFEFA732FF63FAA5FE247656" box="[252,431,1341,1367]" collectionCode="MCNUP-H" pageId="3" pageNumber="502">MCNUP-H 673</specimenCode>
and 247), Collected on 
<date id="F15DD9D0FFEFA732FD28FAA6FCCD7656" box="[695,838,1341,1368]" pageId="3" pageNumber="502" value="2013-05-03">
<collectingDate id="E1192038FFEFA732FD28FAA6FCCD7656" box="[695,838,1341,1368]" pageId="3" pageNumber="502" value="2013-05-03">May 3, 2013</collectingDate>
</date>
, by 
<collectorName id="28169AC6FFEFA732FCEFFAA5FBD07656" box="[880,1115,1341,1368]" pageId="3" pageNumber="502">Aldemar A. Acevedo</collectorName>
in the low area of the 
<collectionCode id="E3F267D5FFEFA732FAD1FAA6FA137659" box="[1358,1432,1342,1368]" pageId="3" pageNumber="502">PNNT</collectionCode>
, 
<collectorName id="28169AC6FFEFA732FF08FAFAFF7F767D" box="[151,244,1378,1404]" pageId="3" pageNumber="502">La Vega</collectorName>
(
<geoCoordinate id="E0D799D7FFEFA732FE9BFAF9FEE9767A" box="[260,354,1377,1403]" degrees="7.13612" direction="north" orientation="latitude" pageId="3" pageNumber="502" precision="1" value="7.13612">7.13612</geoCoordinate>
<geoCoordinate id="E0D799D7FFEFA732FEFDFAF9FE45767A" box="[354,462,1377,1403]" degrees="72.1843" direction="west" orientation="longitude" pageId="3" pageNumber="502" precision="5" value="-72.1843">–72.1843</geoCoordinate>
; 
<specimenCount id="93E53499FFEFA732FE45FAF9FDA6767D" box="[474,557,1377,1404]" pageId="3" pageNumber="502" type="generic">
<quantity id="421B52F5FFEFA732FE45FAF9FDA2767D" box="[474,553,1377,1404]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="3" pageNumber="502" unit="m" value="465.0">
<elevation id="0ECE1823FFEFA732FE45FAF9FDA2767D" box="[474,553,1377,1404]" metricMagnitude="2" metricUnit="m" metricValue="4.65" pageId="3" pageNumber="502" unit="m" value="465.0">465 m</elevation>
</quantity>
.
</specimenCount>
a.s.l.), municipality of Toledo in the buffer zone of the Tamá National Natural Park (
<collectionCode id="E3F267D5FFEFA732FF46FA1DFEA2769E" box="[217,297,1413,1439]" pageId="3" pageNumber="502">PNNT</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="855CFF10FFEFA737FF58FA31FE237266" blockId="3.[151,1437,1161,2016]" lastBlockId="6.[151,1437,153,359]" lastPageId="6" lastPageNumber="505" pageId="3" pageNumber="502">
<emphasis id="B7972302FFEFA732FF58FA31FE6D76C5" bold="true" box="[199,486,1449,1476]" pageId="3" pageNumber="502">Definition and diagnosis</emphasis>
. 
<taxonomicName id="42E38493FFEFA732FE6BFA32FD5876C2" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[500,723,1449,1475]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFEFA732FE6BFA32FD5876C2" box="[500,723,1449,1475]" italics="true" pageId="3" pageNumber="502">Pristimantis ardilae</emphasis>
</taxonomicName>
, 
<emphasis id="B7972302FFEFA732FD7FFA32FCB276C5" bold="true" box="[736,825,1450,1476]" pageId="3" pageNumber="502">
<taxonomicNameLabel id="ACA49E79FFEFA732FD7FFA32FCB276C5" box="[736,825,1450,1476]" pageId="3" pageNumber="502" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is assigned to the genus 
<taxonomicName id="42E38493FFEFA732FBD5FA32FB5876C2" baseAuthorityName="Frost" baseAuthorityYear="2019" box="[1098,1235,1450,1475]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="genus">
<emphasis id="B7972302FFEFA732FBD5FA32FB5876C2" box="[1098,1235,1450,1475]" italics="true" pageId="3" pageNumber="502">Pristimantis</emphasis>
</taxonomicName>
based on molecular phylogenetic relationships (
<figureCitation id="1DD8E395FFEFA732FE6FFA55FDDB76E6" box="[496,592,1485,1511]" captionStart="FIGURE 2" captionStartId="5.[151,250,1908,1933]" captionTargetBox="[175,1423,192,1879]" captionTargetId="figure@5.[160,1427,181,1882]" captionTargetPageId="5" captionText="FIGURE 2. Phylogenetic relationships for some Pristimantis species in South America obtained from a maximum likelihood analysis. Numbers to the left of slash are bootstrap values and to the right are posterior probability are the highest posterior probability. Pristimantis ardilae sp. nov. (red clade), P. bowara sp. nov. (blue clade). Additional species of Pristimantis from Norte de Santander appear as green labels." figureDoi="http://doi.org/10.5281/zenodo.3708842" httpUri="https://zenodo.org/record/3708842/files/figure.png" pageId="3" pageNumber="502">Figure 2</figureCitation>
) and its morphological characteristics (
<bibRefCitation id="E17282E1FFEFA732FB98FA55FAA976E6" author="Lynch, J. D. &amp; Duellman, W. E." box="[1031,1314,1485,1511]" pageId="3" pageNumber="502" pagination="1 - 236" refId="ref11808" refString="Lynch, J. D. &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador: systematics, ecology and biogeography. University of Kansas, Natural History Museum Special Publication, 23, 1 - 236. https: // doi. org / 10.5962 / bhl. title. 7951" type="journal article" year="1997">Lynch &amp; Duellman 1997</bibRefCitation>
; 
<bibRefCitation id="E17282E1FFEFA732FAB1FA55FEA4750A" author="Duellman, W. E. &amp; Lehr, E." pageId="3" pageNumber="502" refId="ref10651" refString="Duellman, W. E. &amp; Lehr, E. (2009) Terrestrial Breeding Frogs (Strabomantidae) in Peru. Natur und Tier-Verlag, Munster, 382 pp." type="book" year="2009">Duellman &amp; Lehr 2009</bibRefCitation>
). We assign the new species to the 
<taxonomicName id="42E38493FFEFA732FD20FA6AFCF3750A" baseAuthorityName="Günther" baseAuthorityYear="1858" box="[703,888,1521,1547]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="3" pageNumber="502" phylum="Chordata" rank="species" species="conspicillatus">
<emphasis id="B7972302FFEFA732FD20FA6AFCF3750A" box="[703,888,1521,1547]" italics="true" pageId="3" pageNumber="502">P. conspicillatus</emphasis>
</taxonomicName>
species group based on position obtained in our phylogenetic analyses (
<figureCitation id="1DD8E395FFEFA732FE3EF98DFD8D752E" box="[417,518,1557,1583]" captionStart="FIGURE 2" captionStartId="5.[151,250,1908,1933]" captionTargetBox="[175,1423,192,1879]" captionTargetId="figure@5.[160,1427,181,1882]" captionTargetPageId="5" captionText="FIGURE 2. Phylogenetic relationships for some Pristimantis species in South America obtained from a maximum likelihood analysis. Numbers to the left of slash are bootstrap values and to the right are posterior probability are the highest posterior probability. Pristimantis ardilae sp. nov. (red clade), P. bowara sp. nov. (blue clade). Additional species of Pristimantis from Norte de Santander appear as green labels." figureDoi="http://doi.org/10.5281/zenodo.3708842" httpUri="https://zenodo.org/record/3708842/files/figure.png" pageId="3" pageNumber="502">Figure 2</figureCitation>
), and following 
<bibRefCitation id="E17282E1FFEFA732FD59F98DFC7C752E" author="Lynch, J. D. &amp; Duellman, W. E." box="[710,1015,1557,1583]" pageId="3" pageNumber="502" pagination="1 - 236" refId="ref11808" refString="Lynch, J. D. &amp; Duellman, W. E. (1997) Frogs of the genus Eleutherodactylus (Leptodactylidae) in western Ecuador: systematics, ecology and biogeography. University of Kansas, Natural History Museum Special Publication, 23, 1 - 236. https: // doi. org / 10.5962 / bhl. title. 7951" type="journal article" year="1997">Lynch &amp; Duellman (1997)</bibRefCitation>
and 
<bibRefCitation id="E17282E1FFEFA732FBADF98DFAF9752E" author="Padial, J. M. &amp; De la Riva, I." box="[1074,1394,1557,1584]" pageId="3" pageNumber="502" pagination="97 - 122" refId="ref11972" refString="Padial, J. M. &amp; De la Riva, I. (2009). Integrative taxonomy reveals cryptic Amazonian species of Pristimantis (Anura: Strabomantidae). Zoological Journal of the Linnean Society, 155 (1), 97 - 122. https: // doi. org / 10.1111 / j. 1096 - 3642.2008.00424. x" type="journal article" year="2009">Padial &amp; De la Riva (2009)</bibRefCitation>
for having: (1) skin on dorsum shagreen; venter smooth to finely areolate whitish or yellowish coloration, and some black spots on the throat; short dorsolateral folds in scapular region. (2) tympanum distinct, round, ~1/2–1/3 of the eye diameter; tympanic membrane and annulus present; supratympanic fold evident. (3) snout subacuminate in a dorsal view; rounded in profile (
<figureCitation id="1DD8E395FFEFA732FD9BF93DFDF675C1" box="[516,637,1701,1728]" captionStart="FIGURE 6" captionStartId="12.[151,250,1653,1678]" captionTargetBox="[168,1424,534,1620]" captionTargetId="figure@12.[151,1436,525,1628]" captionTargetPageId="12" captionText="FIGURE 6. Illustration of the morphological characteristics of palmar surface, the foot and the head (lateral and dorsal view): A. Pristimantis ardilae sp. nov. B. Pristimantis bowara sp. nov. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3708850" httpUri="https://zenodo.org/record/3708850/files/figure.png" pageId="3" pageNumber="502">Figure 6A</figureCitation>
). (4) canthus rostralis defined, straight in dorsal view, slightly rounded in cross section; loreal region weakly concave. Nostrils slightly distinct, directed laterally. (5) upper eyelid without tubercles; cranial crests absent. (6) choanae small, round; dentigerous processes of vomers evident, with 5–6 teeth on the posterior margin; 12–16 premaxillary teeth. (7) vocal slits and nuptial pads present. (8) ED: 89% (females) and 96% (males) of the END. IOD: 81 % (females) and 78 % (males) of the ED. (9) THL ~51% (females) and 46% (males) of the SVL. TL ~ 58% (females) and 56% (males) of the SVL. FL ~ 47 % (females) and 46 % (males) of the SVL. HW ~ 37% of the SVL in both females and males. (10) ulnar tubercles absent. (11) relative lenght of fingers: II ≤ I &lt;
<collectionCode id="E3F267D5FFEFA732FF4EF83AFF6574BD" box="[209,238,1954,1980]" pageId="3" pageNumber="502">IV</collectionCode>
&lt;
<collectionCode id="E3F267D5FFEFA732FE96F83AFEAE74BD" box="[265,293,1954,1980]" pageId="3" pageNumber="502">III</collectionCode>
(
<figureCitation id="1DD8E395FFEFA732FEADF839FE2274BD" box="[306,425,1953,1980]" captionStart="FIGURE 6" captionStartId="12.[151,250,1653,1678]" captionTargetBox="[168,1424,534,1620]" captionTargetId="figure@12.[151,1436,525,1628]" captionTargetPageId="12" captionText="FIGURE 6. Illustration of the morphological characteristics of palmar surface, the foot and the head (lateral and dorsal view): A. Pristimantis ardilae sp. nov. B. Pristimantis bowara sp. nov. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3708850" httpUri="https://zenodo.org/record/3708850/files/figure.png" pageId="3" pageNumber="502">Figure 6A</figureCitation>
). Subarticular tubercles are round and upraised. The palmar tubercle is bifid, shaped similar to a heart; supernumerary tubercles is rounded and pronounced below the proximal subarticular tubercle of 
<collectionCode id="E3F267D5FFEFA732FAF6F85EFA1374E1" box="[1385,1432,1990,2016]" pageId="3" pageNumber="502">FIV</collectionCode>
; the thenar tubercle 80% of palmar tubercle. Finger fringes absent. 
<collectionCode id="E3F267D5FFEAA737FC02FF01FC4273B2" box="[925,969,153,179]" pageId="6" pageNumber="505">FIII</collectionCode>
~30% longer than FI. Disks on 
<collectionCode id="E3F267D5FFEAA737FAD9FF01FA1673B2" box="[1350,1437,153,179]" pageId="6" pageNumber="505">FIII–IV</collectionCode>
moderately expanded (state 2 according to 
<bibRefCitation id="E17282E1FFEAA737FDE7FF25FCB273D9" author="Grant, T. &amp; Frost, D. R. &amp; Caldwell, J. P. &amp; Gagliardo, R. &amp; Haddad, C. F. B. &amp; Kok, P. J. R. &amp; Means, D. B. &amp; Noonan, B. P. &amp; Schargel, W. E. &amp; Wheeler, W." box="[632,825,189,216]" pageId="6" pageNumber="505" pagination="1 - 262" refId="ref10824" refString="Grant, T., Frost, D. R., Caldwell, J. P., Gagliardo, R., Haddad, C. F. B., Kok, P. J. R., Means, D. B., Noonan, B. P., Schargel, W. E. &amp; Wheeler, W. (2006) Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia, Athesphatanura, Dendrobatidae). Bulletin of the American Museum of Natural History, 299, 1 - 262. https: // doi. org / 10.1206 / 0003 - 0090 (2006) 299 [1: PSODFA] 2.0. CO; 2" type="journal article" year="2006">
Grant 
<emphasis id="B7972302FFEAA737FD21FF26FD7D73D6" box="[702,758,189,215]" italics="true" pageId="6" pageNumber="505">et al.</emphasis>
2006
</bibRefCitation>
); disks on FI–II weakly expanded (state 1 according to 
<bibRefCitation id="E17282E1FFEAA737FF08FF79FED373FD" author="Grant, T. &amp; Frost, D. R. &amp; Caldwell, J. P. &amp; Gagliardo, R. &amp; Haddad, C. F. B. &amp; Kok, P. J. R. &amp; Means, D. B. &amp; Noonan, B. P. &amp; Schargel, W. E. &amp; Wheeler, W." box="[151,344,225,252]" pageId="6" pageNumber="505" pagination="1 - 262" refId="ref10824" refString="Grant, T., Frost, D. R., Caldwell, J. P., Gagliardo, R., Haddad, C. F. B., Kok, P. J. R., Means, D. B., Noonan, B. P., Schargel, W. E. &amp; Wheeler, W. (2006) Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia, Athesphatanura, Dendrobatidae). Bulletin of the American Museum of Natural History, 299, 1 - 262. https: // doi. org / 10.1206 / 0003 - 0090 (2006) 299 [1: PSODFA] 2.0. CO; 2" type="journal article" year="2006">
Grant 
<emphasis id="B7972302FFEAA737FF42FF7AFE8573FA" box="[221,270,225,251]" italics="true" pageId="6" pageNumber="505">et al</emphasis>
. 2006
</bibRefCitation>
). (12) inner metatarsal tubercle ovoid; outer metatarsal tubercle small; tarsus without fold; relative lenght of toes: I &lt;
<collectionCode id="E3F267D5FFEAA737FEF9FE9DFEF2721E" box="[358,377,261,287]" pageId="6" pageNumber="505">II</collectionCode>
&lt;
<collectionCode id="E3F267D5FFEAA737FE09FE9DFE39721E" box="[406,434,261,287]" pageId="6" pageNumber="505">III</collectionCode>
&lt;
<collectionCode id="E3F267D5FFEAA737FE4FFE9DFE6F721E" box="[464,484,261,287]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="505" type="Museum">V</collectionCode>
&lt;
<collectionCode id="E3F267D5FFEAA737FD9EFE9DFDAB721E" box="[513,544,261,287]" pageId="6" pageNumber="505">IV</collectionCode>
; Toe disks elliptical; Toe webbing absent (
<figureCitation id="1DD8E395FFEAA737FC64FE9DFBF87221" box="[1019,1139,261,288]" captionStart="FIGURE 6" captionStartId="12.[151,250,1653,1678]" captionTargetBox="[168,1424,534,1620]" captionTargetId="figure@12.[151,1436,525,1628]" captionTargetPageId="12" captionText="FIGURE 6. Illustration of the morphological characteristics of palmar surface, the foot and the head (lateral and dorsal view): A. Pristimantis ardilae sp. nov. B. Pristimantis bowara sp. nov. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3708850" httpUri="https://zenodo.org/record/3708850/files/figure.png" pageId="6" pageNumber="505">Figure 6A</figureCitation>
). (13) 
<taxonomicName id="42E38493FFEAA737FB23FE9EFA17721E" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[1212,1436,261,287]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="505" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFEAA737FB23FE9EFA17721E" box="[1212,1436,261,287]" italics="true" pageId="6" pageNumber="505">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFEAA737FF08FEB1FF7B7242" bold="true" box="[151,240,297,323]" pageId="6" pageNumber="505">
<taxonomicNameLabel id="ACA49E79FFEAA737FF08FEB1FF7B7242" box="[151,240,297,323]" pageId="6" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is apparently sexually dimorphic in size, with a mean SVL of 
<quantity id="421B52F5FFEAA737FC36FEB0FB867242" box="[937,1037,296,323]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="6" pageNumber="505" unit="mm" value="27.5">27.5 mm</quantity>
(n = 4) for males and 
<quantity id="421B52F5FFEAA737FA9BFEB1FAE37242" box="[1284,1384,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="3.8299999999999996" pageId="6" pageNumber="505" unit="mm" value="38.3">38.3 mm</quantity>
(n = 4) for females (
<tableCitation id="C861CAABFFEAA737FEDCFED5FE117266" box="[323,410,333,359]" captionStart="TABLE 2" captionStartId="7.[151,239,153,177]" captionText="TABLE 2. Morphological variation (mm) of adults specimens of Pristimantis ardilae sp. nov. (range, average ± SD)." pageId="6" pageNumber="505">Table 2</tableCitation>
).
</paragraph>
<paragraph id="855CFF10FFE8A735FF08FDFFFE817180" blockId="4.[151,266,615,641]" box="[151,266,615,641]" pageId="4" pageNumber="503">
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<emphasis id="B7972302FFE8A735FF08FDFFFE887180" bold="true" box="[151,259,615,641]" pageId="4" pageNumber="503">TABLE 1</emphasis>
.
</tableNote>
</paragraph>
<paragraph id="855CFF10FFE8A735FF3AFD08F86E7537" pageId="4" pageNumber="503">
<table id="F7E30DB0FFE858CEFF3AFD08F86E7537" box="[165,2021,656,1590]" gridcols="9" gridrows="20" pageId="4" pageNumber="503">
<tr id="3BD3FD52FFE858CEFF3AFD08F86E71CC" box="[165,2021,656,717]" gridrow="0" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFD08FEFE71CC" box="[165,373,656,717]" gridcol="0" gridrow="0" pageId="4" pageNumber="503">Species</th>
<th id="7802942EFFE858CEFE37FD08FC9E71CC" box="[424,789,656,717]" gridcol="1" gridrow="0" pageId="4" pageNumber="503">Locality</th>
<th id="7802942EFFE858CEFCDFFD08FBAB71CC" box="[832,1056,656,717]" gridcol="2" gridrow="0" pageId="4" pageNumber="503">Geographical coordinates</th>
<th id="7802942EFFE858CEFBD9FD08FB6A71CC" box="[1094,1249,656,717]" gridcol="3" gridrow="0" pageId="4" pageNumber="503">Voucher</th>
<th id="7802942EFFE858CEFA98FD08FAF771CC" box="[1287,1404,656,717]" gridcol="4" gridrow="0" pageId="4" pageNumber="503">12S</th>
<th id="7802942EFFE858CEFA3DFD08F99C71CC" box="[1442,1559,656,717]" gridcol="5" gridrow="0" pageId="4" pageNumber="503">16S</th>
<th id="7802942EFFE858CEF9A2FD08F93971CC" box="[1597,1714,656,717]" gridcol="6" gridrow="0" pageId="4" pageNumber="503">CYTB</th>
<th id="7802942EFFE858CEF947FD08F8C671CC" box="[1752,1869,656,717]" gridcol="7" gridrow="0" pageId="4" pageNumber="503">RAG1</th>
<th id="7802942EFFE858CEF8EFFD08F86E71CC" box="[1904,2021,656,717]" gridcol="8" gridrow="0" pageId="4" pageNumber="503">TYR</th>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFD44F86E7018" box="[165,2021,732,793]" gridrow="1" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFD44FEFE7018" box="[165,373,732,793]" gridcol="0" gridrow="1" pageId="4" pageNumber="503">
<taxonomicName id="42E38493FFE8A735FF3AFD44FEFB71F4" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[165,368,732,757]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="503" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE8A735FF3AFD44FEFB71F4" box="[165,368,732,757]" italics="true" pageId="4" pageNumber="503">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE8A735FF3AFC98FF7D7018" bold="true" box="[165,246,768,793]" pageId="4" pageNumber="503">
<taxonomicNameLabel id="ACA49E79FFE8A735FF3AFC98FF7D7018" box="[165,246,768,793]" pageId="4" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="7802942EFFE858CEFE37FD44FC9E7018" box="[424,789,732,793]" gridcol="1" gridrow="1" pageId="4" pageNumber="503">San Antonio</td>
<td id="7802942EFFE858CEFCDFFD44FBAB7018" box="[832,1056,732,793]" gridcol="2" gridrow="1" pageId="4" pageNumber="503">7.162172, -72.25254</td>
<td id="7802942EFFE858CEFBD9FD44FB6A7018" box="[1094,1249,732,793]" gridcol="3" gridrow="1" pageId="4" pageNumber="503">MCNUP-H117</td>
<td id="7802942EFFE858CEFA98FD44FAF77018" box="[1287,1404,732,793]" gridcol="4" gridrow="1" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FD44FAF771F4" box="[1287,1404,732,757]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215410" pageId="4" pageNumber="503">MN215410</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFD44F99C7018" box="[1442,1559,732,793]" gridcol="5" gridrow="1" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFD44F99D71F4" box="[1442,1558,732,757]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215424" pageId="4" pageNumber="503">MN215424</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FD44F9397018" box="[1597,1714,732,793]" gridcol="6" gridrow="1" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FD44F93971F4" box="[1597,1714,732,757]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218376" pageId="4" pageNumber="503">MN218376</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FD44F8C67018" box="[1752,1869,732,793]" gridcol="7" gridrow="1" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FD44F8C671F4" box="[1752,1869,732,757]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225943" pageId="4" pageNumber="503">MN225943</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFD44F86E7018" box="[1904,2021,732,793]" gridcol="8" gridrow="1" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFD44F86E71F4" box="[1904,2021,732,757]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218361" pageId="4" pageNumber="503">MN218361</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFCB0F86E7040" box="[165,2021,808,833]" gridrow="2" pageId="4" pageNumber="503" rowspan-0="1">
<td id="7802942EFFE858CEFE37FCB0FC9E7040" box="[424,789,808,833]" gridcol="1" gridrow="2" pageId="4" pageNumber="503">San Antonio</td>
<td id="7802942EFFE858CEFCDFFCB0FBAB7040" box="[832,1056,808,833]" gridcol="2" gridrow="2" pageId="4" pageNumber="503">7.162172, -72.25254</td>
<td id="7802942EFFE858CEFBD9FCB0FB6A7040" box="[1094,1249,808,833]" gridcol="3" gridrow="2" pageId="4" pageNumber="503">MCNUP-H121</td>
<td id="7802942EFFE858CEFA98FCB0FAF77040" box="[1287,1404,808,833]" gridcol="4" gridrow="2" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FCB0FAF77040" box="[1287,1404,808,833]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215412" pageId="4" pageNumber="503">MN215412</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFCB0F99C7040" box="[1442,1559,808,833]" gridcol="5" gridrow="2" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFCB0F99C7040" box="[1442,1559,808,833]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215427" pageId="4" pageNumber="503">MN215427</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FCB0F9397040" box="[1597,1714,808,833]" gridcol="6" gridrow="2" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FCB0F9397040" box="[1597,1714,808,833]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218378" pageId="4" pageNumber="503">MN218378</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FCB0F8C67040" box="[1752,1869,808,833]" gridcol="7" gridrow="2" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FCB0F8C67040" box="[1752,1869,808,833]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225945" pageId="4" pageNumber="503">MN225945</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFCB0F86E7040" box="[1904,2021,808,833]" gridcol="8" gridrow="2" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFCB0F86E7040" box="[1904,2021,808,833]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218365" pageId="4" pageNumber="503">MN218365</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFCC9F86E706B" box="[165,2021,849,874]" gridrow="3" pageId="4" pageNumber="503" rowspan-0="1">
<td id="7802942EFFE858CEFE37FCC9FC9E706B" box="[424,789,849,874]" gridcol="1" gridrow="3" pageId="4" pageNumber="503">San Antonio</td>
<td id="7802942EFFE858CEFCDFFCC9FBAB706B" box="[832,1056,849,874]" gridcol="2" gridrow="3" pageId="4" pageNumber="503">7.162172, -72.25254</td>
<td id="7802942EFFE858CEFBD9FCC9FB6A706B" box="[1094,1249,849,874]" gridcol="3" gridrow="3" pageId="4" pageNumber="503">MCNUP-H157</td>
<td id="7802942EFFE858CEFA98FCC9FAF7706B" box="[1287,1404,849,874]" gridcol="4" gridrow="3" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA97FCC9FAF7706B" box="[1288,1404,849,874]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215411" pageId="4" pageNumber="503">MN215411</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFCC9F99C706B" box="[1442,1559,849,874]" gridcol="5" gridrow="3" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFCC9F99C706B" box="[1442,1559,849,874]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215425" pageId="4" pageNumber="503">MN215425</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FCC9F939706B" box="[1597,1714,849,874]" gridcol="6" gridrow="3" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FCC9F939706B" box="[1597,1714,849,874]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218377" pageId="4" pageNumber="503">MN218377</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FCC9F8C6706B" box="[1752,1869,849,874]" gridcol="7" gridrow="3" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FCC9F8C7706B" box="[1752,1868,849,874]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225944" pageId="4" pageNumber="503">MN225944</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFCC9F86E706B" box="[1904,2021,849,874]" gridcol="8" gridrow="3" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFCC9F86E706B" box="[1904,2021,849,874]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218362" pageId="4" pageNumber="503">MN218362</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFCE1F86E7093" box="[165,2021,889,914]" gridrow="4" pageId="4" pageNumber="503" rowspan-0="1">
<td id="7802942EFFE858CEFE37FCE1FC9E7093" box="[424,789,889,914]" gridcol="1" gridrow="4" pageId="4" pageNumber="503">San Antonio</td>
<td id="7802942EFFE858CEFCDFFCE1FBAB7093" box="[832,1056,889,914]" gridcol="2" gridrow="4" pageId="4" pageNumber="503">7.162172, -72.25254</td>
<td id="7802942EFFE858CEFBD9FCE1FB6A7093" box="[1094,1249,889,914]" gridcol="3" gridrow="4" pageId="4" pageNumber="503">MCNUP-H166</td>
<td id="7802942EFFE858CEFA98FCE1FAF77093" box="[1287,1404,889,914]" gridcol="4" gridrow="4" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FCE1FAF77093" box="[1287,1404,889,914]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215414" pageId="4" pageNumber="503">MN215414</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFCE1F99C7093" box="[1442,1559,889,914]" gridcol="5" gridrow="4" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFCE1F99C7093" box="[1442,1559,889,914]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215429" pageId="4" pageNumber="503">MN215429</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FCE1F9397093" box="[1597,1714,889,914]" gridcol="6" gridrow="4" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FCE1F9397093" box="[1597,1714,889,914]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218380" pageId="4" pageNumber="503">MN218380</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FCE1F8C67093" box="[1752,1869,889,914]" gridcol="7" gridrow="4" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FCE1F8C67093" box="[1752,1869,889,914]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225947" pageId="4" pageNumber="503">MN225947</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFCE1F86E7093" box="[1904,2021,889,914]" gridcol="8" gridrow="4" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFCE1F86E7093" box="[1904,2021,889,914]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218363" pageId="4" pageNumber="503">MN218363</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFC39F86E70BB" box="[165,2021,929,954]" gridrow="5" pageId="4" pageNumber="503" rowspan-0="1">
<td id="7802942EFFE858CEFE37FC39FC9E70BB" box="[424,789,929,954]" gridcol="1" gridrow="5" pageId="4" pageNumber="503">San Antonio</td>
<td id="7802942EFFE858CEFCDFFC39FBAB70BB" box="[832,1056,929,954]" gridcol="2" gridrow="5" pageId="4" pageNumber="503">7.162172, -72.25254</td>
<td id="7802942EFFE858CEFBD9FC39FB6A70BB" box="[1094,1249,929,954]" gridcol="3" gridrow="5" pageId="4" pageNumber="503">MCNUP-H253</td>
<td id="7802942EFFE858CEFA98FC39FAF770BB" box="[1287,1404,929,954]" gridcol="4" gridrow="5" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEFA3DFC39F99C70BB" box="[1442,1559,929,954]" gridcol="5" gridrow="5" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFC39F99D70BB" box="[1442,1558,929,954]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215426" pageId="4" pageNumber="503">MN215426</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FC39F93970BB" box="[1597,1714,929,954]" gridcol="6" gridrow="5" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEF947FC39F8C670BB" box="[1752,1869,929,954]" gridcol="7" gridrow="5" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEF8EFFC39F86E70BB" box="[1904,2021,929,954]" gridcol="8" gridrow="5" pageId="4" pageNumber="503">-</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFC51F86E70E3" box="[165,2021,969,994]" gridrow="6" pageId="4" pageNumber="503" rowspan-0="1">
<td id="7802942EFFE858CEFE37FC51FC9E70E3" box="[424,789,969,994]" gridcol="1" gridrow="6" pageId="4" pageNumber="503">La Vega</td>
<td id="7802942EFFE858CEFCDFFC51FBAB70E3" box="[832,1056,969,994]" gridcol="2" gridrow="6" pageId="4" pageNumber="503">7.13612, -72.1843</td>
<td id="7802942EFFE858CEFBD9FC51FB6A70E3" box="[1094,1249,969,994]" gridcol="3" gridrow="6" pageId="4" pageNumber="503">MCNUP-H163</td>
<td id="7802942EFFE858CEFA98FC51FAF770E3" box="[1287,1404,969,994]" gridcol="4" gridrow="6" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FC51FAF770E3" box="[1287,1404,969,994]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215413" pageId="4" pageNumber="503">MN215413</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFC51F99C70E3" box="[1442,1559,969,994]" gridcol="5" gridrow="6" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFC51F99C70E3" box="[1442,1559,969,994]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215428" pageId="4" pageNumber="503">MN215428</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FC51F93970E3" box="[1597,1714,969,994]" gridcol="6" gridrow="6" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FC51F93970E3" box="[1597,1714,969,994]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218379" pageId="4" pageNumber="503">MN218379</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FC51F8C670E3" box="[1752,1869,969,994]" gridcol="7" gridrow="6" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FC51F8C770E3" box="[1752,1868,969,994]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225946" pageId="4" pageNumber="503">MN225946</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFC51F86E70E3" box="[1904,2021,969,994]" gridcol="8" gridrow="6" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFC51F86E70E3" box="[1904,2021,969,994]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218364" pageId="4" pageNumber="503">MN218364</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFC6AF86E772E" box="[165,2021,1010,1071]" gridrow="7" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFC6AFEFE772E" box="[165,373,1010,1071]" gridcol="0" gridrow="7" pageId="4" pageNumber="503">
<taxonomicName id="42E38493FFE8A735FF3AFC6AFEFE770A" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[165,373,1010,1035]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="503" phylum="Chordata" rank="species" species="bowara" status="sp. nov.">
<emphasis id="B7972302FFE8A735FF3AFC6AFEFE770A" box="[165,373,1010,1035]" italics="true" pageId="4" pageNumber="503">Pristimantis bowara</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE8A735FF3AFB8EFF7D772E" bold="true" box="[165,246,1046,1071]" pageId="4" pageNumber="503">
<taxonomicNameLabel id="ACA49E79FFE8A735FF3AFB8EFF7D772E" box="[165,246,1046,1071]" pageId="4" pageNumber="503" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<td id="7802942EFFE858CEFE37FC6AFC9E772E" box="[424,789,1010,1071]" gridcol="1" gridrow="7" pageId="4" pageNumber="503">San Antonio</td>
<td id="7802942EFFE858CEFCDFFC6AFBAB772E" box="[832,1056,1010,1071]" gridcol="2" gridrow="7" pageId="4" pageNumber="503">7.16866, -72.2655</td>
<td id="7802942EFFE858CEFBD9FC6AFB6A772E" box="[1094,1249,1010,1071]" gridcol="3" gridrow="7" pageId="4" pageNumber="503">MCNUP-H165</td>
<td id="7802942EFFE858CEFA98FC6AFAF7772E" box="[1287,1404,1010,1071]" gridcol="4" gridrow="7" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FC6AFAF7770A" box="[1287,1404,1010,1035]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215406" pageId="4" pageNumber="503">MN215406</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFC6AF99C772E" box="[1442,1559,1010,1071]" gridcol="5" gridrow="7" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFC6AF99D770A" box="[1442,1558,1010,1035]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215431" pageId="4" pageNumber="503">MN215431</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FC6AF939772E" box="[1597,1714,1010,1071]" gridcol="6" gridrow="7" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FC6AF939770A" box="[1597,1714,1010,1035]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218372" pageId="4" pageNumber="503">MN218372</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FC6AF8C6772E" box="[1752,1869,1010,1071]" gridcol="7" gridrow="7" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FC6AF8C7770A" box="[1752,1868,1010,1035]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225939" pageId="4" pageNumber="503">MN225939</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFC6AF86E772E" box="[1904,2021,1010,1071]" gridcol="8" gridrow="7" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFC6AF86E770A" box="[1904,2021,1010,1035]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218357" pageId="4" pageNumber="503">MN218357</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFBA6F86E7756" box="[165,2021,1086,1111]" gridrow="8" pageId="4" pageNumber="503" rowspan-0="1">
<td id="7802942EFFE858CEFE37FBA6FC9E7756" box="[424,789,1086,1111]" gridcol="1" gridrow="8" pageId="4" pageNumber="503">San Antonio</td>
<td id="7802942EFFE858CEFCDFFBA6FBAB7756" box="[832,1056,1086,1111]" gridcol="2" gridrow="8" pageId="4" pageNumber="503">7.16866, -72.2655</td>
<td id="7802942EFFE858CEFBD9FBA6FB6A7756" box="[1094,1249,1086,1111]" gridcol="3" gridrow="8" pageId="4" pageNumber="503">MCNUP-H160</td>
<td id="7802942EFFE858CEFA98FBA6FAF77756" box="[1287,1404,1086,1111]" gridcol="4" gridrow="8" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FBA6FAF77756" box="[1287,1404,1086,1111]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215407" pageId="4" pageNumber="503">MN215407</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFBA6F99C7756" box="[1442,1559,1086,1111]" gridcol="5" gridrow="8" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFBA6F99D7756" box="[1442,1558,1086,1111]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215432" pageId="4" pageNumber="503">MN215432</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FBA6F9397756" box="[1597,1714,1086,1111]" gridcol="6" gridrow="8" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FBA6F9397756" box="[1597,1714,1086,1111]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218373" pageId="4" pageNumber="503">MN218373</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FBA6F8C67756" box="[1752,1869,1086,1111]" gridcol="7" gridrow="8" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FBA6F8C77756" box="[1752,1868,1086,1111]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225940" pageId="4" pageNumber="503">MN225940</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFBA6F86E7756" box="[1904,2021,1086,1111]" gridcol="8" gridrow="8" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFBA6F86E7756" box="[1904,2021,1086,1111]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218358" pageId="4" pageNumber="503">MN218358</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFBFEF86E777E" box="[165,2021,1126,1151]" gridrow="9" pageId="4" pageNumber="503" rowspan-0="1">
<td id="7802942EFFE858CEFE37FBFEFC9E777E" box="[424,789,1126,1151]" gridcol="1" gridrow="9" pageId="4" pageNumber="503">San Antonio</td>
<td id="7802942EFFE858CEFCDFFBFEFBAB777E" box="[832,1056,1126,1151]" gridcol="2" gridrow="9" pageId="4" pageNumber="503">7.16866, -72.2655</td>
<td id="7802942EFFE858CEFBD9FBFEFB6A777E" box="[1094,1249,1126,1151]" gridcol="3" gridrow="9" pageId="4" pageNumber="503">MCNUP-H230</td>
<td id="7802942EFFE858CEFA98FBFEFAF7777E" box="[1287,1404,1126,1151]" gridcol="4" gridrow="9" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FBFEFAF7777E" box="[1287,1404,1126,1151]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215408" pageId="4" pageNumber="503">MN215408</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFBFEF99C777E" box="[1442,1559,1126,1151]" gridcol="5" gridrow="9" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFBFEF99C777E" box="[1442,1559,1126,1151]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215433" pageId="4" pageNumber="503">MN215433</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FBFEF939777E" box="[1597,1714,1126,1151]" gridcol="6" gridrow="9" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FBFEF939777E" box="[1597,1714,1126,1151]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218374" pageId="4" pageNumber="503">MN218374</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FBFEF8C6777E" box="[1752,1869,1126,1151]" gridcol="7" gridrow="9" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FBFEF8C7777E" box="[1752,1868,1126,1151]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225941" pageId="4" pageNumber="503">MN225941</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFBFEF86E777E" box="[1904,2021,1126,1151]" gridcol="8" gridrow="9" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFBFEF86E777E" box="[1904,2021,1126,1151]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218360" pageId="4" pageNumber="503">MN218360</accessionNumber>
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</tr>
<tr id="3BD3FD52FFE858CEFF3AFB16F86E77A6" box="[165,2021,1166,1191]" gridrow="10" pageId="4" pageNumber="503" rowspan-0="1">
<td id="7802942EFFE858CEFE37FB16FC9E77A6" box="[424,789,1166,1191]" gridcol="1" gridrow="10" pageId="4" pageNumber="503">San Lorenzo</td>
<td id="7802942EFFE858CEFCDFFB16FBAB77A6" box="[832,1056,1166,1191]" gridcol="2" gridrow="10" pageId="4" pageNumber="503">7.16615, -72.2336</td>
<td id="7802942EFFE858CEFBD9FB16FB6A77A6" box="[1094,1249,1166,1191]" gridcol="3" gridrow="10" pageId="4" pageNumber="503">MCNUP-H303</td>
<td id="7802942EFFE858CEFA98FB16FAF777A6" box="[1287,1404,1166,1191]" gridcol="4" gridrow="10" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEFA3DFB16F99C77A6" box="[1442,1559,1166,1191]" gridcol="5" gridrow="10" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFB16F99D77A6" box="[1442,1558,1166,1191]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215430" pageId="4" pageNumber="503">MN215430</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FB16F93977A6" box="[1597,1714,1166,1191]" gridcol="6" gridrow="10" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEF947FB16F8C677A6" box="[1752,1869,1166,1191]" gridcol="7" gridrow="10" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEF8EFFB16F86E77A6" box="[1904,2021,1166,1191]" gridcol="8" gridrow="10" pageId="4" pageNumber="503">-</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFB2FF86E77D1" box="[165,2021,1207,1232]" gridrow="11" pageId="4" pageNumber="503" rowspan-0="1">
<td id="7802942EFFE858CEFE37FB2FFC9E77D1" box="[424,789,1207,1232]" gridcol="1" gridrow="11" pageId="4" pageNumber="503">San Lorenzo</td>
<td id="7802942EFFE858CEFCDFFB2FFBAB77D1" box="[832,1056,1207,1232]" gridcol="2" gridrow="11" pageId="4" pageNumber="503">7.16615, -72.2336</td>
<td id="7802942EFFE858CEFBD9FB2FFB6A77D1" box="[1094,1249,1207,1232]" gridcol="3" gridrow="11" pageId="4" pageNumber="503">MCNUP-H304</td>
<td id="7802942EFFE858CEFA98FB2FFAF777D1" box="[1287,1404,1207,1232]" gridcol="4" gridrow="11" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FB2FFAF777D1" box="[1287,1404,1207,1232]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215409" pageId="4" pageNumber="503">MN215409</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFB2FF99C77D1" box="[1442,1559,1207,1232]" gridcol="5" gridrow="11" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFB2FF99D77D1" box="[1442,1558,1207,1232]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215434" pageId="4" pageNumber="503">MN215434</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FB2FF93977D1" box="[1597,1714,1207,1232]" gridcol="6" gridrow="11" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FB2FF93977D1" box="[1597,1714,1207,1232]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218375" pageId="4" pageNumber="503">MN218375</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FB2FF8C677D1" box="[1752,1869,1207,1232]" gridcol="7" gridrow="11" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FB2FF8C777D1" box="[1752,1868,1207,1232]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225942" pageId="4" pageNumber="503">MN225942</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFB2FF86E77D1" box="[1904,2021,1207,1232]" gridcol="8" gridrow="11" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFB2FF86E77D1" box="[1904,2021,1207,1232]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218359" pageId="4" pageNumber="503">MN218359</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFA9BF86E761D" box="[165,2021,1283,1308]" gridrow="12" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFA9BFEFE761D" box="[165,373,1283,1308]" gridcol="0" gridrow="12" pageId="4" pageNumber="503">
<taxonomicName id="42E38493FFE8A735FF3AFA9BFE86761D" baseAuthorityName="Cochran &amp; Goin" baseAuthorityYear="1970" box="[165,269,1283,1308]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="503" phylum="Chordata" rank="species" species="nicefori">
<emphasis id="B7972302FFE8A735FF3AFA9BFE86761D" box="[165,269,1283,1308]" italics="true" pageId="4" pageNumber="503">P. nicefori</emphasis>
</taxonomicName>
</th>
<td id="7802942EFFE858CEFE37FA9BFC9E761D" box="[424,789,1283,1308]" gridcol="1" gridrow="12" pageId="4" pageNumber="503">Presidente, Páramo de Almorzadero</td>
<td id="7802942EFFE858CEFCDFFA9BFBAB761D" box="[832,1056,1283,1308]" gridcol="2" gridrow="12" pageId="4" pageNumber="503">7.39008, -72.3945</td>
<td id="7802942EFFE858CEFBD9FA9BFB6A761D" box="[1094,1249,1283,1308]" gridcol="3" gridrow="12" pageId="4" pageNumber="503">MCNUP-H48</td>
<td id="7802942EFFE858CEFA98FA9BFAF7761D" box="[1287,1404,1283,1308]" gridcol="4" gridrow="12" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FA9BFAF7761D" box="[1287,1404,1283,1308]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215418" pageId="4" pageNumber="503">MN215418</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFA9BF99C761D" box="[1442,1559,1283,1308]" gridcol="5" gridrow="12" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFA9BF99D761D" box="[1442,1558,1283,1308]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215436" pageId="4" pageNumber="503">MN215436</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FA9BF939761D" box="[1597,1714,1283,1308]" gridcol="6" gridrow="12" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FA9BF939761D" box="[1597,1714,1283,1308]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218383" pageId="4" pageNumber="503">MN218383</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FA9BF8C6761D" box="[1752,1869,1283,1308]" gridcol="7" gridrow="12" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FA9BF8C7761D" box="[1752,1868,1283,1308]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225949" pageId="4" pageNumber="503">MN225949</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFA9BF86E761D" box="[1904,2021,1283,1308]" gridcol="8" gridrow="12" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFA9BF86E761D" box="[1904,2021,1283,1308]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218368" pageId="4" pageNumber="503">MN218368</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFAB3F86E7645" box="[165,2021,1323,1348]" gridrow="13" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFAB3FEFE7645" box="[165,373,1323,1348]" gridcol="0" gridrow="13" pageId="4" pageNumber="503">
<taxonomicName id="42E38493FFE8A735FF3AFAB3FE9A7645" baseAuthorityName="Lynch" baseAuthorityYear="1983" box="[165,273,1323,1348]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="503" phylum="Chordata" rank="species" species="anolirex">
<emphasis id="B7972302FFE8A735FF3AFAB3FE9A7645" box="[165,273,1323,1348]" italics="true" pageId="4" pageNumber="503">P. anolirex</emphasis>
</taxonomicName>
</th>
<td id="7802942EFFE858CEFE37FAB3FC9E7645" box="[424,789,1323,1348]" gridcol="1" gridrow="13" pageId="4" pageNumber="503">Chitagá, Páramo de Almorzadero</td>
<td id="7802942EFFE858CEFCDFFAB3FBAB7645" box="[832,1056,1323,1348]" gridcol="2" gridrow="13" pageId="4" pageNumber="503">7.150928, -72.6335</td>
<td id="7802942EFFE858CEFBD9FAB3FB6A7645" box="[1094,1249,1323,1348]" gridcol="3" gridrow="13" pageId="4" pageNumber="503">MCNUP-H67</td>
<td id="7802942EFFE858CEFA98FAB3FAF77645" box="[1287,1404,1323,1348]" gridcol="4" gridrow="13" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FAB3FAF77645" box="[1287,1404,1323,1348]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215421" pageId="4" pageNumber="503">MN215421</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFAB3F99C7645" box="[1442,1559,1323,1348]" gridcol="5" gridrow="13" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFAB3F99D7645" box="[1442,1558,1323,1348]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215441" pageId="4" pageNumber="503">MN215441</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FAB3F9397645" box="[1597,1714,1323,1348]" gridcol="6" gridrow="13" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FAB3F9397645" box="[1597,1714,1323,1348]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218386" pageId="4" pageNumber="503">MN218386</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FAB3F8C67645" box="[1752,1869,1323,1348]" gridcol="7" gridrow="13" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FAB3F8C77645" box="[1752,1868,1323,1348]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225954" pageId="4" pageNumber="503">MN225954</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFAB3F86E7645" box="[1904,2021,1323,1348]" gridcol="8" gridrow="13" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFAB3F86E7645" box="[1904,2021,1323,1348]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218371" pageId="4" pageNumber="503">MN218371</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFACBF86E766C" box="[165,2021,1363,1389]" gridrow="14" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFACBFEFE766C" box="[165,373,1363,1389]" gridcol="0" gridrow="14" pageId="4" pageNumber="503">
<taxonomicName id="42E38493FFE8A735FF3AFACCFE94766C" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[165,287,1364,1389]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="503" phylum="Chordata" rank="species" species="mondolfii">
<emphasis id="B7972302FFE8A735FF3AFACCFE94766C" box="[165,287,1364,1389]" italics="true" pageId="4" pageNumber="503">P. mondolfii</emphasis>
</taxonomicName>
</th>
<td id="7802942EFFE858CEFE37FACBFC9E766C" box="[424,789,1363,1389]" gridcol="1" gridrow="14" pageId="4" pageNumber="503">Asiria de Belen, PNN Tamá</td>
<td id="7802942EFFE858CEFCDFFACBFBAB766C" box="[832,1056,1363,1389]" gridcol="2" gridrow="14" pageId="4" pageNumber="503">7.3223888, -72.36497</td>
<td id="7802942EFFE858CEFBD9FACBFB6A766C" box="[1094,1249,1363,1389]" gridcol="3" gridrow="14" pageId="4" pageNumber="503">MCNUP-H457</td>
<td id="7802942EFFE858CEFA98FACBFAF7766C" box="[1287,1404,1363,1389]" gridcol="4" gridrow="14" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FACBFAF7766D" box="[1287,1404,1363,1388]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215420" pageId="4" pageNumber="503">MN215420</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFACBF99C766C" box="[1442,1559,1363,1389]" gridcol="5" gridrow="14" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFACBF99D766D" box="[1442,1558,1363,1388]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215440" pageId="4" pageNumber="503">MN215440</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FACBF939766C" box="[1597,1714,1363,1389]" gridcol="6" gridrow="14" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FACBF939766D" box="[1597,1714,1363,1388]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218385" pageId="4" pageNumber="503">MN218385</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FACBF8C6766C" box="[1752,1869,1363,1389]" gridcol="7" gridrow="14" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FACBF8C6766D" box="[1752,1869,1363,1388]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225953" pageId="4" pageNumber="503">MN225953</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFACBF86E766C" box="[1904,2021,1363,1389]" gridcol="8" gridrow="14" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFACBF86E766D" box="[1904,2021,1363,1388]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218370" pageId="4" pageNumber="503">MN218370</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFAE4F86E7694" box="[165,2021,1404,1429]" gridrow="15" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFAE4FEFE7694" box="[165,373,1404,1429]" gridcol="0" gridrow="15" pageId="4" pageNumber="503">
<taxonomicName id="42E38493FFE8A735FF3AFAE4FF717694" box="[165,250,1404,1429]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="503" phylum="Chordata" rank="species" species="yukpa">
<emphasis id="B7972302FFE8A735FF3AFAE4FF717694" box="[165,250,1404,1429]" italics="true" pageId="4" pageNumber="503">P. yukpa</emphasis>
</taxonomicName>
</th>
<td id="7802942EFFE858CEFE37FAE4FC9E7694" box="[424,789,1404,1429]" gridcol="1" gridrow="15" pageId="4" pageNumber="503">Bochalena</td>
<td id="7802942EFFE858CEFCDFFAE4FBAB7694" box="[832,1056,1404,1429]" gridcol="2" gridrow="15" pageId="4" pageNumber="503">7.6845, -72.6194</td>
<td id="7802942EFFE858CEFBD9FAE4FB6A7694" box="[1094,1249,1404,1429]" gridcol="3" gridrow="15" pageId="4" pageNumber="503">MCNUP-H28</td>
<td id="7802942EFFE858CEFA98FAE4FAF77694" box="[1287,1404,1404,1429]" gridcol="4" gridrow="15" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FAE4FAF77694" box="[1287,1404,1404,1429]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215416" pageId="4" pageNumber="503">MN215416</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFAE4F99C7694" box="[1442,1559,1404,1429]" gridcol="5" gridrow="15" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFAE4F99C7694" box="[1442,1559,1404,1429]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215438" pageId="4" pageNumber="503">MN215438</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FAE4F9397694" box="[1597,1714,1404,1429]" gridcol="6" gridrow="15" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEF947FAE4F8C67694" box="[1752,1869,1404,1429]" gridcol="7" gridrow="15" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FAE4F8C77694" box="[1752,1868,1404,1429]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225950" pageId="4" pageNumber="503">MN225950</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFAE4F86E7694" box="[1904,2021,1404,1429]" gridcol="8" gridrow="15" pageId="4" pageNumber="503">-</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFA3CF86E76BC" box="[165,2021,1444,1469]" gridrow="16" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFA3CFEFE76BC" box="[165,373,1444,1469]" gridcol="0" gridrow="16" pageId="4" pageNumber="503">
<taxonomicName id="42E38493FFE8A735FF3AFA3CFF7176BC" box="[165,250,1444,1469]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="4" pageNumber="503" phylum="Chordata" rank="species" species="yukpa">
<emphasis id="B7972302FFE8A735FF3AFA3CFF7176BC" box="[165,250,1444,1469]" italics="true" pageId="4" pageNumber="503">P. yukpa</emphasis>
</taxonomicName>
</th>
<td id="7802942EFFE858CEFE37FA3CFC9E76BC" box="[424,789,1444,1469]" gridcol="1" gridrow="16" pageId="4" pageNumber="503">Bochalena</td>
<td id="7802942EFFE858CEFCDFFA3CFBAB76BC" box="[832,1056,1444,1469]" gridcol="2" gridrow="16" pageId="4" pageNumber="503">7.6845, -72.6194</td>
<td id="7802942EFFE858CEFBD9FA3CFB6A76BC" box="[1094,1249,1444,1469]" gridcol="3" gridrow="16" pageId="4" pageNumber="503">MCNUP-H29</td>
<td id="7802942EFFE858CEFA98FA3CFAF776BC" box="[1287,1404,1444,1469]" gridcol="4" gridrow="16" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FA3CFAF776BC" box="[1287,1404,1444,1469]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215417" pageId="4" pageNumber="503">MN215417</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFA3CF99C76BC" box="[1442,1559,1444,1469]" gridcol="5" gridrow="16" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFA3CF99C76BC" box="[1442,1559,1444,1469]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215439" pageId="4" pageNumber="503">MN215439</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FA3CF93976BC" box="[1597,1714,1444,1469]" gridcol="6" gridrow="16" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEF947FA3CF8C676BC" box="[1752,1869,1444,1469]" gridcol="7" gridrow="16" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FA3CF8C776BC" box="[1752,1868,1444,1469]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225951" pageId="4" pageNumber="503">MN225951</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFA3CF86E76BC" box="[1904,2021,1444,1469]" gridcol="8" gridrow="16" pageId="4" pageNumber="503">-</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFA54F86E76E4" box="[165,2021,1484,1509]" gridrow="17" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFA54FEFE76E4" box="[165,373,1484,1509]" gridcol="0" gridrow="17" pageId="4" pageNumber="503">
<emphasis id="B7972302FFE8A735FF3AFA54FF3F76E4" box="[165,180,1484,1509]" italics="true" pageId="4" pageNumber="503">P</emphasis>
. sp1
</th>
<td id="7802942EFFE858CEFE37FA54FC9E76E4" box="[424,789,1484,1509]" gridcol="1" gridrow="17" pageId="4" pageNumber="503">Orocué, PNN Tamá</td>
<td id="7802942EFFE858CEFCDFFA54FBAB76E4" box="[832,1056,1484,1509]" gridcol="2" gridrow="17" pageId="4" pageNumber="503">7.429583, -72.44533</td>
<td id="7802942EFFE858CEFBD9FA54FB6A76E4" box="[1094,1249,1484,1509]" gridcol="3" gridrow="17" pageId="4" pageNumber="503">MCNUP-H100</td>
<td id="7802942EFFE858CEFA98FA54FAF776E4" box="[1287,1404,1484,1509]" gridcol="4" gridrow="17" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98FA54FAF776E4" box="[1287,1404,1484,1509]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215415" pageId="4" pageNumber="503">MN215415</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DFA54F99C76E4" box="[1442,1559,1484,1509]" gridcol="5" gridrow="17" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DFA54F99C76E4" box="[1442,1559,1484,1509]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215435" pageId="4" pageNumber="503">MN215435</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2FA54F93976E4" box="[1597,1714,1484,1509]" gridcol="6" gridrow="17" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FA54F93976E4" box="[1597,1714,1484,1509]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218381" pageId="4" pageNumber="503">MN218381</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FA54F8C676E4" box="[1752,1869,1484,1509]" gridcol="7" gridrow="17" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947FA54F8C676E4" box="[1752,1869,1484,1509]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225948" pageId="4" pageNumber="503">MN225948</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFFA54F86E76E4" box="[1904,2021,1484,1509]" gridcol="8" gridrow="17" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFA54F86E76E4" box="[1904,2021,1484,1509]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218366" pageId="4" pageNumber="503">MN218366</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AFA6DF86E750F" box="[165,2021,1525,1550]" gridrow="18" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AFA6DFEFE750F" box="[165,373,1525,1550]" gridcol="0" gridrow="18" pageId="4" pageNumber="503">
<emphasis id="B7972302FFE8A735FF3AFA6DFF3F750F" box="[165,180,1525,1550]" italics="true" pageId="4" pageNumber="503">P</emphasis>
. sp1
</th>
<td id="7802942EFFE858CEFE37FA6DFC9E750F" box="[424,789,1525,1550]" gridcol="1" gridrow="18" pageId="4" pageNumber="503">Orocué, PNN Tamá</td>
<td id="7802942EFFE858CEFCDFFA6DFBAB750F" box="[832,1056,1525,1550]" gridcol="2" gridrow="18" pageId="4" pageNumber="503">7.429583, -72.44533</td>
<td id="7802942EFFE858CEFBD9FA6DFB6A750F" box="[1094,1249,1525,1550]" gridcol="3" gridrow="18" pageId="4" pageNumber="503">MCNUP-H75</td>
<td id="7802942EFFE858CEFA98FA6DFAF7750F" box="[1287,1404,1525,1550]" gridcol="4" gridrow="18" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEFA3DFA6DF99C750F" box="[1442,1559,1525,1550]" gridcol="5" gridrow="18" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEF9A2FA6DF939750F" box="[1597,1714,1525,1550]" gridcol="6" gridrow="18" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2FA6DF939750F" box="[1597,1714,1525,1550]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218382" pageId="4" pageNumber="503">MN218382</accessionNumber>
</td>
<td id="7802942EFFE858CEF947FA6DF8C6750F" box="[1752,1869,1525,1550]" gridcol="7" gridrow="18" pageId="4" pageNumber="503">-</td>
<td id="7802942EFFE858CEF8EFFA6DF86E750F" box="[1904,2021,1525,1550]" gridcol="8" gridrow="18" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFFA6DF86E750F" box="[1904,2021,1525,1550]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218367" pageId="4" pageNumber="503">MN218367</accessionNumber>
</td>
</tr>
<tr id="3BD3FD52FFE858CEFF3AF985F86E7537" box="[165,2021,1565,1590]" gridrow="19" pageId="4" pageNumber="503">
<th id="7802942EFFE858CEFF3AF985FEFE7537" box="[165,373,1565,1590]" gridcol="0" gridrow="19" pageId="4" pageNumber="503">
<emphasis id="B7972302FFE8A735FF3AF985FF3F7537" box="[165,180,1565,1590]" italics="true" pageId="4" pageNumber="503">P</emphasis>
. sp2
</th>
<td id="7802942EFFE858CEFE37F985FC9E7537" box="[424,789,1565,1590]" gridcol="1" gridrow="19" pageId="4" pageNumber="503">Páramo del Tamá, PNN Tamá</td>
<td id="7802942EFFE858CEFCDFF985FBAB7537" box="[832,1056,1565,1590]" gridcol="2" gridrow="19" pageId="4" pageNumber="503">7.390086, -72.3945</td>
<td id="7802942EFFE858CEFBD9F985FB6A7537" box="[1094,1249,1565,1590]" gridcol="3" gridrow="19" pageId="4" pageNumber="503">MCNUP-H600</td>
<td id="7802942EFFE858CEFA98F985FAF77537" box="[1287,1404,1565,1590]" gridcol="4" gridrow="19" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA98F985FAF77537" box="[1287,1404,1565,1590]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215419" pageId="4" pageNumber="503">MN215419</accessionNumber>
</td>
<td id="7802942EFFE858CEFA3DF985F99C7537" box="[1442,1559,1565,1590]" gridcol="5" gridrow="19" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735FA3DF985F99C7537" box="[1442,1559,1565,1590]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN215437" pageId="4" pageNumber="503">MN215437</accessionNumber>
</td>
<td id="7802942EFFE858CEF9A2F985F9397537" box="[1597,1714,1565,1590]" gridcol="6" gridrow="19" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F9A2F985F9397537" box="[1597,1714,1565,1590]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218384" pageId="4" pageNumber="503">MN218384</accessionNumber>
</td>
<td id="7802942EFFE858CEF947F985F8C67537" box="[1752,1869,1565,1590]" gridcol="7" gridrow="19" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F947F985F8C77537" box="[1752,1868,1565,1590]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN225952" pageId="4" pageNumber="503">MN225952</accessionNumber>
</td>
<td id="7802942EFFE858CEF8EFF985F86E7537" box="[1904,2021,1565,1590]" gridcol="8" gridrow="19" pageId="4" pageNumber="503">
<accessionNumber id="9AB062F3FFE8A735F8EFF985F86E7537" box="[1904,2021,1565,1590]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN218369" pageId="4" pageNumber="503">MN218369</accessionNumber>
</td>
</tr>
</table>
</paragraph>
<caption id="D19CAF98FFE9A734FF08F8ECFDCC74F8" ID-DOI="http://doi.org/10.5281/zenodo.3708842" ID-Zenodo-Dep="3708842" httpUri="https://zenodo.org/record/3708842/files/figure.png" pageId="5" pageNumber="504" startId="5.[151,250,1908,1933]" targetBox="[175,1423,192,1879]" targetPageId="5">
<paragraph id="855CFF10FFE9A734FF08F8ECFDCC74F8" blockId="5.[151,1436,1908,2041]" pageId="5" pageNumber="504">
<emphasis id="B7972302FFE9A734FF08F8ECFE9F748C" bold="true" box="[151,276,1908,1933]" pageId="5" pageNumber="504">FIGURE 2.</emphasis>
Phylogenetic relationships for some 
<taxonomicName id="42E38493FFE9A734FD0EF8EDFC86748D" baseAuthorityName="Frost" baseAuthorityYear="2019" box="[657,781,1909,1932]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="504" phylum="Chordata" rank="genus">
<emphasis id="B7972302FFE9A734FD0EF8EDFC86748D" box="[657,781,1909,1932]" italics="true" pageId="5" pageNumber="504">Pristimantis</emphasis>
</taxonomicName>
species in South America obtained from a maximum likelihood analysis. Numbers to the left of slash are bootstrap values and to the right are posterior probability are the highest posterior probability. 
<taxonomicName id="42E38493FFE9A734FE8DF825FE5574D5" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[274,478,1980,2004]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="504" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE9A734FE8DF825FE5574D5" box="[274,478,1980,2004]" italics="true" pageId="5" pageNumber="504">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE9A734FE7AF824FDBC74D5" bold="true" box="[485,567,1980,2004]" pageId="5" pageNumber="504">
<taxonomicNameLabel id="ACA49E79FFE9A734FE7AF824FDBC74D5" box="[485,567,1980,2004]" pageId="5" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(red clade), 
<taxonomicName id="42E38493FFE9A734FD25F825FCAA74D5" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[698,801,1980,2004]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="504" phylum="Chordata" rank="species" species="bowara" status="sp. nov.">
<emphasis id="B7972302FFE9A734FD25F825FCAA74D5" box="[698,801,1980,2004]" italics="true" pageId="5" pageNumber="504">P. bowara</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE9A734FCB6F824FCF174D5" bold="true" box="[809,890,1980,2004]" pageId="5" pageNumber="504">
<taxonomicNameLabel id="ACA49E79FFE9A734FCB6F824FCF174D5" box="[809,890,1980,2004]" pageId="5" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(blue clade). Additional species of 
<taxonomicName id="42E38493FFE9A734FB77F825FAEF74D5" baseAuthorityName="Frost" baseAuthorityYear="2019" box="[1256,1380,1981,2004]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="5" pageNumber="504" phylum="Chordata" rank="genus">
<emphasis id="B7972302FFE9A734FB77F825FAEF74D5" box="[1256,1380,1981,2004]" italics="true" pageId="5" pageNumber="504">Pristimantis</emphasis>
</taxonomicName>
from Norte de Santander appear as green labels.
</paragraph>
</caption>
<caption id="D19CAF98FFEAA737FF08F8EBFBA374D5" ID-DOI="http://doi.org/10.5281/zenodo.3708844" ID-Zenodo-Dep="3708844" httpUri="https://zenodo.org/record/3708844/files/figure.png" pageId="6" pageNumber="505" startId="6.[151,250,1907,1932]" targetBox="[151,1435,384,1876]" targetPageId="6">
<paragraph id="855CFF10FFEAA737FF08F8EBFBA374D5" blockId="6.[151,1437,1907,2004]" pageId="6" pageNumber="505">
<emphasis id="B7972302FFEAA737FF08F8EBFE9D748D" bold="true" box="[151,278,1907,1932]" pageId="6" pageNumber="505">FIGURE 3.</emphasis>
Phylogenetic relationships subset for some 
<taxonomicName id="42E38493FFEAA737FD75F8ECFCED748A" baseAuthorityName="Frost" baseAuthorityYear="2019" box="[746,870,1908,1931]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="505" phylum="Chordata" rank="genus">
<emphasis id="B7972302FFEAA737FD75F8ECFCED748A" box="[746,870,1908,1931]" italics="true" pageId="6" pageNumber="505">Pristimantis</emphasis>
</taxonomicName>
species obtained from a Bayesian Inference analysis. Numbers above branches are the highest posterior probability. 
<taxonomicName id="42E38493FFEAA737FC94F800FC5E74B1" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[779,981,1944,1968]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="505" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFEAA737FC94F800FC5E74B1" box="[779,981,1944,1968]" italics="true" pageId="6" pageNumber="505">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFEAA737FC45F800FBA174B1" bold="true" box="[986,1066,1944,1968]" pageId="6" pageNumber="505">
<taxonomicNameLabel id="ACA49E79FFEAA737FC45F800FBA174B1" box="[986,1066,1944,1968]" pageId="6" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(red clade), 
<taxonomicName id="42E38493FFEAA737FB37F800FA8674B1" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[1192,1293,1944,1968]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="505" phylum="Chordata" rank="species" species="bowara" status="sp. nov.">
<emphasis id="B7972302FFEAA737FB37F800FA8674B1" box="[1192,1293,1944,1968]" italics="true" pageId="6" pageNumber="505">P. bowara</emphasis>
</taxonomicName>
<emphasis id="B7972302FFEAA737FA8CF800FAE974B1" bold="true" box="[1299,1378,1944,1968]" pageId="6" pageNumber="505">
<taxonomicNameLabel id="ACA49E79FFEAA737FA8CF800FAE974B1" box="[1299,1378,1944,1968]" pageId="6" pageNumber="505" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(blue clade). Additional species of 
<taxonomicName id="42E38493FFEAA737FE21F824FDB174D2" baseAuthorityName="Frost" baseAuthorityYear="2019" box="[446,570,1980,2003]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="6" pageNumber="505" phylum="Chordata" rank="genus">
<emphasis id="B7972302FFEAA737FE21F824FDB174D2" box="[446,570,1980,2003]" italics="true" pageId="6" pageNumber="505">Pristimantis</emphasis>
</taxonomicName>
from Norte de Santander appear as green labels.
</paragraph>
</caption>
<caption id="D19CAF98FFEBA736FF08FF01FAF673B0" box="[151,1405,153,178]" pageId="7" pageNumber="506">
<paragraph id="855CFF10FFEBA736FF08FF01FAF673B0" blockId="7.[151,1405,153,178]" box="[151,1405,153,178]" pageId="7" pageNumber="506">
<emphasis id="B7972302FFEBA736FF08FF01FE8873B0" bold="true" box="[151,259,153,177]" pageId="7" pageNumber="506">TABLE 2</emphasis>
. Morphological variation (mm) of adults specimens of 
<taxonomicName id="42E38493FFEBA736FCCCFF02FBA373B0" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[851,1064,153,177]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="506" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFEBA736FCCCFF02FBA373B0" box="[851,1064,153,177]" italics="true" pageId="7" pageNumber="506">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFEBA736FBB0FF01FB0E73B0" bold="true" box="[1071,1157,153,177]" pageId="7" pageNumber="506">
<taxonomicNameLabel id="ACA49E79FFEBA736FBB0FF01FB0E73B0" box="[1071,1157,153,177]" pageId="7" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(range, average ± SD).
</paragraph>
</caption>
<paragraph id="855CFF10FFEBA736FF39FF5AFAD97034" pageId="7" pageNumber="506">
<table id="F7E30DB0FFEB58CEFF39FF59FAED7034" box="[166,1382,193,821]" gridcols="3" gridrows="16" pageId="7" pageNumber="506">
<tr id="3BD3FD52FFEB58CEFF39FF59FAED73DB" box="[166,1382,193,218]" gridrow="0" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FF59FDB373DB" box="[166,568,193,218]" gridcol="0" gridrow="0" pageId="7" pageNumber="506">Measurements</th>
<th id="7802942EFFEB58CEFCB4FF59FB8473DB" box="[811,1039,193,218]" gridcol="1" gridrow="0" pageId="7" pageNumber="506">Female (5)</th>
<th id="7802942EFFEB58CEFB1DFF59FAED73DB" box="[1154,1382,193,218]" gridcol="2" gridrow="0" pageId="7" pageNumber="506">Males (5)</th>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FF71FAED7203" box="[166,1382,233,258]" gridrow="1" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FF71FDB37203" box="[166,568,233,258]" gridcol="0" gridrow="1" pageId="7" pageNumber="506">Snout-vent length (SVL)</th>
<td id="7802942EFFEB58CEFCB4FF71FB847203" box="[811,1039,233,258]" gridcol="1" gridrow="1" pageId="7" pageNumber="506">36.3–39.7 (38.3 ± 1.6)</td>
<td id="7802942EFFEB58CEFB1DFF71FAED7203" box="[1154,1382,233,258]" gridcol="2" gridrow="1" pageId="7" pageNumber="506">25.1–29.1 (27.5 ± 1.8)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FE89FAED722B" box="[166,1382,273,298]" gridrow="2" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FE89FDB3722B" box="[166,568,273,298]" gridcol="0" gridrow="2" pageId="7" pageNumber="506">Head length (HL)</th>
<td id="7802942EFFEB58CEFCB4FE89FB84722B" box="[811,1039,273,298]" gridcol="1" gridrow="2" pageId="7" pageNumber="506">12.1–15.7 (13.9 ± 1.7)</td>
<td id="7802942EFFEB58CEFB1DFE89FAED722B" box="[1154,1382,273,298]" gridcol="2" gridrow="2" pageId="7" pageNumber="506">9.0–9.7 (9.1 ± 0.6)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FEA1FAED7253" box="[166,1382,313,338]" gridrow="3" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FEA1FDB37253" box="[166,568,313,338]" gridcol="0" gridrow="3" pageId="7" pageNumber="506">Head width (HW)</th>
<td id="7802942EFFEB58CEFCB4FEA1FB847253" box="[811,1039,313,338]" gridcol="1" gridrow="3" pageId="7" pageNumber="506">13.0–15.3 (14.2 ± 1.0)</td>
<td id="7802942EFFEB58CEFB1DFEA1FAED7253" box="[1154,1382,313,338]" gridcol="2" gridrow="3" pageId="7" pageNumber="506">9.4–10.8 (10.2 ± 0.6)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FEFAFAED727A" box="[166,1382,354,379]" gridrow="4" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FEFAFDB3727A" box="[166,568,354,379]" gridcol="0" gridrow="4" pageId="7" pageNumber="506">Eye diameter (ED)</th>
<td id="7802942EFFEB58CEFCB4FEFAFB84727A" box="[811,1039,354,379]" gridcol="1" gridrow="4" pageId="7" pageNumber="506">3.4–5.1 (4.2 ± 0.7)</td>
<td id="7802942EFFEB58CEFB1DFEFAFAED727A" box="[1154,1382,354,379]" gridcol="2" gridrow="4" pageId="7" pageNumber="506">3.2–3.6 (3.2 ± 0.4)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FE12FAED72A2" box="[166,1382,394,419]" gridrow="5" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FE12FDB372A2" box="[166,568,394,419]" gridcol="0" gridrow="5" pageId="7" pageNumber="506">Eye to nostril distance (END)</th>
<td id="7802942EFFEB58CEFCB4FE12FB8472A2" box="[811,1039,394,419]" gridcol="1" gridrow="5" pageId="7" pageNumber="506">4.3–5.1 (4.7 ± 0.4)</td>
<td id="7802942EFFEB58CEFB1DFE12FAED72A2" box="[1154,1382,394,419]" gridcol="2" gridrow="5" pageId="7" pageNumber="506">2.9–3.3 (3.1 ± 0.2)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FE2AFAED72CA" box="[166,1382,434,459]" gridrow="6" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FE2AFDB372CA" box="[166,568,434,459]" gridcol="0" gridrow="6" pageId="7" pageNumber="506">Inter Nostril distance (IND)</th>
<td id="7802942EFFEB58CEFCB4FE2AFB8472CA" box="[811,1039,434,459]" gridcol="1" gridrow="6" pageId="7" pageNumber="506">2.9–3.6 (3.3 ± 0.3)</td>
<td id="7802942EFFEB58CEFB1DFE2AFAED72CA" box="[1154,1382,434,459]" gridcol="2" gridrow="6" pageId="7" pageNumber="506">2.3–2.7 (2.4 ± 0.2)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FE42FAED72F2" box="[166,1382,474,499]" gridrow="7" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FE42FDB372F2" box="[166,568,474,499]" gridcol="0" gridrow="7" pageId="7" pageNumber="506">Inter orbital diameter (IOD)</th>
<td id="7802942EFFEB58CEFCB4FE42FB8472F2" box="[811,1039,474,499]" gridcol="1" gridrow="7" pageId="7" pageNumber="506">3.3–3.8 (3.4 ± 0.3)</td>
<td id="7802942EFFEB58CEFB1DFE42FAED72F2" box="[1154,1382,474,499]" gridcol="2" gridrow="7" pageId="7" pageNumber="506">2.3–2.7 (2.5 ± 0.2)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FD9BFAED711D" box="[166,1382,515,540]" gridrow="8" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FD9BFDB3711D" box="[166,568,515,540]" gridcol="0" gridrow="8" pageId="7" pageNumber="506">Tympanum diameter (TD)</th>
<td id="7802942EFFEB58CEFCB4FD9BFB84711D" box="[811,1039,515,540]" gridcol="1" gridrow="8" pageId="7" pageNumber="506">1.3–1.7 (1.5 ± 0.1)</td>
<td id="7802942EFFEB58CEFB1DFD9BFAED711D" box="[1154,1382,515,540]" gridcol="2" gridrow="8" pageId="7" pageNumber="506">0.9–1.0 (0.9 ± 0.1)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FDB3FAED7145" box="[166,1382,555,580]" gridrow="9" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FDB3FDB37145" box="[166,568,555,580]" gridcol="0" gridrow="9" pageId="7" pageNumber="506">Forearm length (FOL)</th>
<td id="7802942EFFEB58CEFCB4FDB3FB847145" box="[811,1039,555,580]" gridcol="1" gridrow="9" pageId="7" pageNumber="506">7.2–8.2 (7.8 ± 0.4)</td>
<td id="7802942EFFEB58CEFB1DFDB3FAED7145" box="[1154,1382,555,580]" gridcol="2" gridrow="9" pageId="7" pageNumber="506">5.0–6.2 (5.7 ± 0.6)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FDCBFAED716D" box="[166,1382,595,620]" gridrow="10" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FDCBFDB3716D" box="[166,568,595,620]" gridcol="0" gridrow="10" pageId="7" pageNumber="506">Length of the humerus (LHU)</th>
<td id="7802942EFFEB58CEFCB4FDCBFB84716D" box="[811,1039,595,620]" gridcol="1" gridrow="10" pageId="7" pageNumber="506">5.1–5.7 (5.4 ± 0.2)</td>
<td id="7802942EFFEB58CEFB1DFDCBFAED716D" box="[1154,1382,595,620]" gridcol="2" gridrow="10" pageId="7" pageNumber="506">4.1–5.0 (4.6 ± 0.5)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FDE3FAED7195" box="[166,1382,635,660]" gridrow="11" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FDE3FDB37195" box="[166,568,635,660]" gridcol="0" gridrow="11" pageId="7" pageNumber="506">Foof length (FL)</th>
<td id="7802942EFFEB58CEFCB4FDE3FB847195" box="[811,1039,635,660]" gridcol="1" gridrow="11" pageId="7" pageNumber="506">17.5–19.0 (18.0 ± 0.7)</td>
<td id="7802942EFFEB58CEFB1DFDE3FAED7195" box="[1154,1382,635,660]" gridcol="2" gridrow="11" pageId="7" pageNumber="506">11.8–13.3 (12.6 ± 0.9)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FD3CFAED71BC" box="[166,1382,676,701]" gridrow="12" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FD3CFDB371BC" box="[166,568,676,701]" gridcol="0" gridrow="12" pageId="7" pageNumber="506">Thigh length (THL)</th>
<td id="7802942EFFEB58CEFCB4FD3CFB8471BC" box="[811,1039,676,701]" gridcol="1" gridrow="12" pageId="7" pageNumber="506">18.7–19.9 (19.5 ± 0.6)</td>
<td id="7802942EFFEB58CEFB1DFD3CFAED71BC" box="[1154,1382,676,701]" gridcol="2" gridrow="12" pageId="7" pageNumber="506">11.4–13.4 (12.6 ± 1.0)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FD54FAED71E4" box="[166,1382,716,741]" gridrow="13" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FD54FDB371E4" box="[166,568,716,741]" gridcol="0" gridrow="13" pageId="7" pageNumber="506">Tibia length (TL)</th>
<td id="7802942EFFEB58CEFCB4FD54FB8471E4" box="[811,1039,716,741]" gridcol="1" gridrow="13" pageId="7" pageNumber="506">21.6–23.1 (22.5 ± 0.7)</td>
<td id="7802942EFFEB58CEFB1DFD54FAED71E4" box="[1154,1382,716,741]" gridcol="2" gridrow="13" pageId="7" pageNumber="506">13.4–16.8 (15.3 ± 1.7)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FD6CFAED700C" box="[166,1382,756,781]" gridrow="14" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FD6CFDB3700C" box="[166,568,756,781]" gridcol="0" gridrow="14" pageId="7" pageNumber="506">Disc diameter of the finger three (DD3)</th>
<td id="7802942EFFEB58CEFCB4FD6CFB84700C" box="[811,1039,756,781]" gridcol="1" gridrow="14" pageId="7" pageNumber="506">1.0–1.2 (1.1 ± 0.1)</td>
<td id="7802942EFFEB58CEFB1DFD6CFAED700C" box="[1154,1382,756,781]" gridcol="2" gridrow="14" pageId="7" pageNumber="506">0.7–1.1 (0.8 ± 0.2)</td>
</tr>
<tr id="3BD3FD52FFEB58CEFF39FC84FAED7034" box="[166,1382,796,821]" gridrow="15" pageId="7" pageNumber="506">
<th id="7802942EFFEB58CEFF39FC84FDB37034" box="[166,568,796,821]" gridcol="0" gridrow="15" pageId="7" pageNumber="506">Disc diameter toe four (DF4)</th>
<td id="7802942EFFEB58CEFCB4FC84FB847034" box="[811,1039,796,821]" gridcol="1" gridrow="15" pageId="7" pageNumber="506">1.1–1.3 (1.2 ± 0.1)</td>
<td id="7802942EFFEB58CEFB1DFC84FAED7034" box="[1154,1382,796,821]" gridcol="2" gridrow="15" pageId="7" pageNumber="506">0.7–0.9 (0.8 ± 0.1)</td>
</tr>
</table>
</paragraph>
<paragraph id="855CFF10FFEBA736FF58FCFCFB6770C6" blockId="7.[151,1437,868,967]" pageId="7" pageNumber="506">
<emphasis id="B7972302FFEBA736FF58FCFCFDCA707E" bold="true" box="[199,577,868,895]" pageId="7" pageNumber="506">Comparisons with other species</emphasis>
. 
<taxonomicName id="42E38493FFEBA736FDCFFCFEFCBB707E" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[592,816,869,895]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="506" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFEBA736FDCFFCFEFCBB707E" box="[592,816,869,895]" italics="true" pageId="7" pageNumber="506">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFEBA736FCA8FCFDFC1B707E" bold="true" box="[823,912,869,895]" pageId="7" pageNumber="506">
<taxonomicNameLabel id="ACA49E79FFEBA736FCA8FCFDFC1B707E" box="[823,912,869,895]" pageId="7" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be further distinguished from other 
<taxonomicName id="42E38493FFEBA736FACAFCFEFF6970A2" baseAuthorityName="Frost" baseAuthorityYear="2019" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="B7972302FFEBA736FACAFCFEFF6970A2" italics="true" pageId="7" pageNumber="506">Pristimantis</emphasis>
</taxonomicName>
species from the northeastern region of 
<collectingCountry id="FDF4BF80FFEBA736FD38FC11FC9370A2" box="[679,792,905,931]" name="Colombia" pageId="7" pageNumber="506">Colombia</collectingCountry>
, the Venezuelan Andes, and phylogenetically related species of the 
<taxonomicName id="42E38493FFEBA736FE8EFC36FE4270C6" baseAuthorityName="Günther" baseAuthorityYear="1858" box="[273,457,941,967]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="506" phylum="Chordata" rank="species" species="conspicillatus">
<emphasis id="B7972302FFEBA736FE8EFC36FE4270C6" box="[273,457,941,967]" italics="true" pageId="7" pageNumber="506">P. conspicillatus</emphasis>
</taxonomicName>
species group by the combination of characters summarized in 
<tableCitation id="C861CAABFFEBA736FB0DFC35FB6D70C6" box="[1170,1254,941,967]" captionStart="TABLE 3" captionStartId="8.[151,239,645,671]" captionText="TABLE 3. Comparison of diagnostic characters of some species of the Pristimantis conspicillatus group and species from northeastern Colombia, including the new species:" pageId="7" pageNumber="506">Table 3</tableCitation>
.
</paragraph>
<caption id="D19CAF98FFEBA736FF08F81EFBAB74C2" ID-DOI="http://doi.org/10.5281/zenodo.3708846" ID-Zenodo-Dep="3708846" httpUri="https://zenodo.org/record/3708846/files/figure.png" pageId="7" pageNumber="506" startId="7.[151,250,1926,1951]" targetBox="[155,1432,991,1899]" targetPageId="7">
<paragraph id="855CFF10FFEBA736FF08F81EFBAB74C2" blockId="7.[151,1436,1926,1987]" pageId="7" pageNumber="506">
<emphasis id="B7972302FFEBA736FF08F81EFE98749E" bold="true" box="[151,275,1926,1951]" pageId="7" pageNumber="506">FIGURE 4.</emphasis>
Photos of the specimens in life. A. 
<taxonomicName id="42E38493FFEBA736FD1FF81FFCC0749E" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[640,843,1927,1951]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="506" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFEBA736FD1FF81FFCC0749E" box="[640,843,1927,1951]" italics="true" pageId="7" pageNumber="506">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFEBA736FCCDF81FFC21749E" bold="true" box="[850,938,1927,1951]" pageId="7" pageNumber="506">
<taxonomicNameLabel id="ACA49E79FFEBA736FCCDF81FFC2D749E" box="[850,934,1927,1951]" pageId="7" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
uncollected animals. B. 
<taxonomicName id="42E38493FFEBA736FB38F81FFAFC749E" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[1191,1399,1927,1951]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="7" pageNumber="506" phylum="Chordata" rank="species" species="bowara" status="sp. nov.">
<emphasis id="B7972302FFEBA736FB38F81FFAFC749E" box="[1191,1399,1927,1951]" italics="true" pageId="7" pageNumber="506">Pristimantis bowara</emphasis>
</taxonomicName>
<emphasis id="B7972302FFEBA736FAE1F81FFF4174C2" bold="true" pageId="7" pageNumber="506">
<taxonomicNameLabel id="ACA49E79FFEBA736FAE1F81FFF4E74C2" pageId="7" pageNumber="506" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
holoytpe, MCNUP-H 165. Lower zone of the buffer zone of PNN Tamá, Colombia.
</paragraph>
</caption>
<caption id="D19CAF98FFE4A739FF08FD1DF871719E" box="[151,2042,645,671]" pageId="8" pageNumber="507">
<paragraph id="855CFF10FFE4A739FF08FD1DF871719E" blockId="8.[151,2042,645,779]" box="[151,2042,645,671]" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF08FD1DFE87719E" bold="true" box="[151,268,645,671]" pageId="8" pageNumber="507">TABLE 3.</emphasis>
Comparison of diagnostic characters of some species of the 
<taxonomicName id="42E38493FFE4A739FC39FD1DFB4F719E" baseAuthorityName="Günther" baseAuthorityYear="1858" box="[934,1220,645,671]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="conspicillatus">
<emphasis id="B7972302FFE4A739FC39FD1DFB4F719E" box="[934,1220,645,671]" italics="true" pageId="8" pageNumber="507">Pristimantis conspicillatus</emphasis>
</taxonomicName>
group and species from northeastern Colombia, including the new species:
</paragraph>
</caption>
<tableNote id="7805FE9EFFE4A739FF08FD31FE08700A" pageId="8" pageNumber="507" targetBox="[165,1984,798,1492]" targetPageId="8">
<paragraph id="855CFF10FFE4A739FF08FD31F87271C2" blockId="8.[151,2042,645,779]" box="[151,2041,681,707]" pageId="8" pageNumber="507">(1) Length of first and second fingers. (2) Belly skin texture. (3) Dorsal skin texture. (4) Dorsal tubercles. (5) Dorsolateral folds. (6) Finger fringes. (7) Toe fringes. (8) Basal toe</paragraph>
<paragraph id="855CFF10FFE4A739FF08FD55F87171E6" blockId="8.[151,2042,645,779]" box="[151,2042,717,743]" pageId="8" pageNumber="507">webbing. (9) Throat color pattern. (10) Tarsal fold. (11) Colour pattern of posterior surfaces of thighs. (12) Nuptial pads. (13) SVL (males). (14) SVL (females). Character states:</paragraph>
<paragraph id="855CFF10FFE4A739FF08FD69FE08700A" blockId="8.[151,2042,645,779]" box="[151,387,753,779]" pageId="8" pageNumber="507">1= present, 0= absent.</paragraph>
</tableNote>
<paragraph id="855CFF10FFE4A739FF3AFC86F8F976D5" pageId="8" pageNumber="507">
<table id="F7E30DB0FFE458CEFF3AFC86F84B76D5" box="[165,1984,798,1492]" gridcols="8" gridrows="15" pageId="8" pageNumber="507">
<tr id="3BD3FD52FFE458CEFF3AFC86F84B7036" box="[165,1984,798,823]" gridrow="0" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFC86FE357036" box="[165,446,798,823]" gridcol="0" gridrow="0" pageId="8" pageNumber="507">Species</th>
<th id="7802942EFFE458CEFDECFC86FD307036" box="[627,699,798,823]" gridcol="1" gridrow="0" pageId="8" pageNumber="507">1</th>
<th id="7802942EFFE458CEFD69FC86FBCD7036" box="[758,1094,798,823]" gridcol="2" gridrow="0" pageId="8" pageNumber="507">2</th>
<th id="7802942EFFE458CEFB10FC86FB627036" box="[1167,1257,798,823]" gridcol="3" gridrow="0" pageId="8" pageNumber="507">3</th>
<th id="7802942EFFE458CEFABCFC86F9AA7036" box="[1315,1569,798,823]" gridcol="4" gridrow="0" pageId="8" pageNumber="507">4</th>
<th id="7802942EFFE458CEF915FC86F91C7036" box="[1674,1687,798,823]" gridcol="5" gridrow="0" pageId="8" pageNumber="507">5</th>
<th id="7802942EFFE458CEF95EFC86F8977036" box="[1729,1820,798,823]" gridcol="6" gridrow="0" pageId="8" pageNumber="507">6</th>
<th id="7802942EFFE458CEF8FAFC86F84B7036" box="[1893,1984,798,823]" gridcol="7" gridrow="0" pageId="8" pageNumber="507">7</th>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFCD6F84B7066" box="[165,1984,846,871]" gridrow="1" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFCD6FE357066" box="[165,446,846,871]" gridcol="0" gridrow="1" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFCD6FEF27066" bold="true" box="[165,377,846,871]" italics="true" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFCD6FEE47066" baseAuthorityName="Lynch" baseAuthorityYear="1994" box="[165,367,846,871]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="carranguerorum">P. carranguerorum</taxonomicName>
*
</emphasis>
(N=3)
</th>
<td id="7802942EFFE458CEFDECFCD6FD307066" box="[627,699,846,871]" gridcol="1" gridrow="1" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FCD6FBCD7066" box="[758,1094,846,871]" gridcol="2" gridrow="1" pageId="8" pageNumber="507">smooth</td>
<td id="7802942EFFE458CEFB10FCD6FB627066" box="[1167,1257,846,871]" gridcol="3" gridrow="1" pageId="8" pageNumber="507">shagreen</td>
<td id="7802942EFFE458CEFABCFCD6F9AA7066" box="[1315,1569,846,871]" gridcol="4" gridrow="1" pageId="8" pageNumber="507">short folds</td>
<td id="7802942EFFE458CEF915FCD6F91C7066" box="[1674,1687,846,871]" gridcol="5" gridrow="1" pageId="8" pageNumber="507">1</td>
<td id="7802942EFFE458CEF95EFCD6F8977066" box="[1729,1820,846,871]" gridcol="6" gridrow="1" pageId="8" pageNumber="507">weak</td>
<td id="7802942EFFE458CEF8FAFCD6F84B7066" box="[1893,1984,846,871]" gridcol="7" gridrow="1" pageId="8" pageNumber="507">weak</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFCE5F84B7097" box="[165,1984,893,918]" gridrow="2" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFCE5FE357097" box="[165,446,893,918]" gridcol="0" gridrow="2" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFCE5FE967097" bold="true" box="[165,285,893,918]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFCE5FE9B7097" baseAuthorityName="Lynch" baseAuthorityYear="1994" box="[165,272,893,918]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="medemi">
<emphasis id="B7972302FFE4A739FF3AFCE5FE9B7097" bold="true" box="[165,272,893,918]" italics="true" pageId="8" pageNumber="507">P. medemi</emphasis>
</taxonomicName>
*
</emphasis>
(N=3)
</th>
<td id="7802942EFFE458CEFDECFCE5FD307097" box="[627,699,893,918]" gridcol="1" gridrow="2" pageId="8" pageNumber="507">F I&gt; II</td>
<td id="7802942EFFE458CEFD69FCE5FBCD7097" box="[758,1094,893,918]" gridcol="2" gridrow="2" pageId="8" pageNumber="507">slightly granular</td>
<td id="7802942EFFE458CEFB10FCE5FB627097" box="[1167,1257,893,918]" gridcol="3" gridrow="2" pageId="8" pageNumber="507">shagreen</td>
<td id="7802942EFFE458CEFABCFCE5F9AA7097" box="[1315,1569,893,918]" gridcol="4" gridrow="2" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FCE5F91C7097" box="[1674,1687,893,918]" gridcol="5" gridrow="2" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF95EFCE5F8977097" box="[1729,1820,893,918]" gridcol="6" gridrow="2" pageId="8" pageNumber="507">weak</td>
<td id="7802942EFFE458CEF8FAFCE5F84B7097" box="[1893,1984,893,918]" gridcol="7" gridrow="2" pageId="8" pageNumber="507">weak</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFC35F84B70C7" box="[165,1984,941,966]" gridrow="3" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFC35FE3570C7" box="[165,446,941,966]" gridcol="0" gridrow="3" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFC35FED470C7" bold="true" box="[165,351,941,966]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFC35FED970C7" baseAuthorityName="La Marca" baseAuthorityYear="2004" box="[165,338,941,966]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="pedimontanus">
<emphasis id="B7972302FFE4A739FF3AFC35FED970C7" bold="true" box="[165,338,941,966]" italics="true" pageId="8" pageNumber="507">P. pedimontanus</emphasis>
</taxonomicName>
*
</emphasis>
(N=7)
</th>
<td id="7802942EFFE458CEFDECFC35FD3070C7" box="[627,699,941,966]" gridcol="1" gridrow="3" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FC35FBCD70C7" box="[758,1094,941,966]" gridcol="2" gridrow="3" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFB10FC35FB6270C7" box="[1167,1257,941,966]" gridcol="3" gridrow="3" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFABCFC35F9AA70C7" box="[1315,1569,941,966]" gridcol="4" gridrow="3" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FC35F91C70C7" box="[1674,1687,941,966]" gridcol="5" gridrow="3" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF95EFC35F89770C7" box="[1729,1820,941,966]" gridcol="6" gridrow="3" pageId="8" pageNumber="507">prominet</td>
<td id="7802942EFFE458CEF8FAFC35F84B70C7" box="[1893,1984,941,966]" gridcol="7" gridrow="3" pageId="8" pageNumber="507">prominet</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFC45F84B70F7" box="[165,1984,989,1014]" gridrow="4" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFC45FE3570F7" box="[165,446,989,1014]" gridcol="0" gridrow="4" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFC45FE9E70F7" baseAuthorityName="Lynch" baseAuthorityYear="1983" box="[165,277,989,1014]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="anolirex">
<emphasis id="B7972302FFE4A739FF3AFC45FE9E70F7" bold="true" box="[165,277,989,1014]" italics="true" pageId="8" pageNumber="507">P. anolirex</emphasis>
</taxonomicName>
(N=8)
</th>
<td id="7802942EFFE458CEFDECFC45FD3070F7" box="[627,699,989,1014]" gridcol="1" gridrow="4" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FC45FBCD70F7" box="[758,1094,989,1014]" gridcol="2" gridrow="4" pageId="8" pageNumber="507">slightly granular</td>
<td id="7802942EFFE458CEFB10FC45FB6270F7" box="[1167,1257,989,1014]" gridcol="3" gridrow="4" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFABCFC45F9AA70F7" box="[1315,1569,989,1014]" gridcol="4" gridrow="4" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FC45F91C70F7" box="[1674,1687,989,1014]" gridcol="5" gridrow="4" pageId="8" pageNumber="507">1</td>
<td id="7802942EFFE458CEF95EFC45F89770F7" box="[1729,1820,989,1014]" gridcol="6" gridrow="4" pageId="8" pageNumber="507">prominet</td>
<td id="7802942EFFE458CEF8FAFC45F84B70F7" box="[1893,1984,989,1014]" gridcol="7" gridrow="4" pageId="8" pageNumber="507">prominet</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFB95F84B7727" box="[165,1984,1037,1062]" gridrow="5" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFB95FE357727" box="[165,446,1037,1062]" gridcol="0" gridrow="5" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFB95FEB47727" bold="true" box="[165,319,1037,1062]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFB95FEBA7727" baseAuthorityName="Lynch" baseAuthorityYear="2013" box="[165,305,1037,1062]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="batrachites">
<emphasis id="B7972302FFE4A739FF3AFB95FEBA7727" bold="true" box="[165,305,1037,1062]" italics="true" pageId="8" pageNumber="507">P. batrachites</emphasis>
</taxonomicName>
*
</emphasis>
(N=3)
</th>
<td id="7802942EFFE458CEFDECFB95FD307727" box="[627,699,1037,1062]" gridcol="1" gridrow="5" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FB95FBCD7727" box="[758,1094,1037,1062]" gridcol="2" gridrow="5" pageId="8" pageNumber="507">slightly granular</td>
<td id="7802942EFFE458CEFB10FB95FB627727" box="[1167,1257,1037,1062]" gridcol="3" gridrow="5" pageId="8" pageNumber="507">smooth</td>
<td id="7802942EFFE458CEFABCFB95F9AA7727" box="[1315,1569,1037,1062]" gridcol="4" gridrow="5" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FB95F91C7727" box="[1674,1687,1037,1062]" gridcol="5" gridrow="5" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF95EFB95F8977727" box="[1729,1820,1037,1062]" gridcol="6" gridrow="5" pageId="8" pageNumber="507">prominet</td>
<td id="7802942EFFE458CEF8FAFB95F84B7727" box="[1893,1984,1037,1062]" gridcol="7" gridrow="5" pageId="8" pageNumber="507">prominet</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFBA5F84B7757" box="[165,1984,1085,1110]" gridrow="6" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFBA5FE357757" box="[165,446,1085,1110]" gridcol="0" gridrow="6" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFBA5FEED7757" bold="true" box="[165,358,1085,1110]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFBA5FED27757" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[165,345,1085,1110]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="melanoproctus">
<emphasis id="B7972302FFE4A739FF3AFBA5FED27757" bold="true" box="[165,345,1085,1110]" italics="true" pageId="8" pageNumber="507">P. melanoproctus</emphasis>
</taxonomicName>
*
</emphasis>
(N=1)
</th>
<td id="7802942EFFE458CEFDECFBA5FD307757" box="[627,699,1085,1110]" gridcol="1" gridrow="6" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FBA5FBCD7757" box="[758,1094,1085,1110]" gridcol="2" gridrow="6" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFB10FBA5FB627757" box="[1167,1257,1085,1110]" gridcol="3" gridrow="6" pageId="8" pageNumber="507">shagreen</td>
<td id="7802942EFFE458CEFABCFBA5F9AA7757" box="[1315,1569,1085,1110]" gridcol="4" gridrow="6" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FBA5F91C7757" box="[1674,1687,1085,1110]" gridcol="5" gridrow="6" pageId="8" pageNumber="507">1</td>
<td id="7802942EFFE458CEF95EFBA5F8977757" box="[1729,1820,1085,1110]" gridcol="6" gridrow="6" pageId="8" pageNumber="507">weak</td>
<td id="7802942EFFE458CEF8FAFBA5F84B7757" box="[1893,1984,1085,1110]" gridcol="7" gridrow="6" pageId="8" pageNumber="507">weak</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFBF4F84B7784" box="[165,1984,1132,1157]" gridrow="7" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFBF4FE357784" box="[165,446,1132,1157]" gridcol="0" gridrow="7" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFBF4FEBA7784" bold="true" box="[165,305,1132,1157]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFBF4FEA87784" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[165,291,1132,1157]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="mondolfii">
<emphasis id="B7972302FFE4A739FF3AFBF4FEA87784" bold="true" box="[165,291,1132,1157]" italics="true" pageId="8" pageNumber="507">P. mondolfii</emphasis>
</taxonomicName>
*
</emphasis>
(N=2)
</th>
<td id="7802942EFFE458CEFDECFBF4FD307784" box="[627,699,1132,1157]" gridcol="1" gridrow="7" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FBF4FBCD7784" box="[758,1094,1132,1157]" gridcol="2" gridrow="7" pageId="8" pageNumber="507">smooth, granular posterolaterally</td>
<td id="7802942EFFE458CEFB10FBF4FB627784" box="[1167,1257,1132,1157]" gridcol="3" gridrow="7" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFABCFBF4F9AA7784" box="[1315,1569,1132,1157]" gridcol="4" gridrow="7" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FBF4F91C7784" box="[1674,1687,1132,1157]" gridcol="5" gridrow="7" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF95EFBF4F8977784" box="[1729,1820,1132,1157]" gridcol="6" gridrow="7" pageId="8" pageNumber="507">weak</td>
<td id="7802942EFFE458CEF8FAFBF4F84B7784" box="[1893,1984,1132,1157]" gridcol="7" gridrow="7" pageId="8" pageNumber="507">weak</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFB04F84B77B4" box="[165,1984,1180,1205]" gridrow="8" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFB04FE3577B4" box="[165,446,1180,1205]" gridcol="0" gridrow="8" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFB04FE9B77B4" baseAuthorityName="Cochran &amp; Goin" baseAuthorityYear="1970" box="[165,272,1180,1205]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="nicefori">
<emphasis id="B7972302FFE4A739FF3AFB04FE9B77B4" bold="true" box="[165,272,1180,1205]" italics="true" pageId="8" pageNumber="507">P. nicefori</emphasis>
</taxonomicName>
(N=4)
</th>
<td id="7802942EFFE458CEFDECFB04FD3077B4" box="[627,699,1180,1205]" gridcol="1" gridrow="8" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FB04FBCD77B4" box="[758,1094,1180,1205]" gridcol="2" gridrow="8" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFB10FB04FB6277B4" box="[1167,1257,1180,1205]" gridcol="3" gridrow="8" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFABCFB04F9AA77B4" box="[1315,1569,1180,1205]" gridcol="4" gridrow="8" pageId="8" pageNumber="507">short folds</td>
<td id="7802942EFFE458CEF915FB04F91C77B4" box="[1674,1687,1180,1205]" gridcol="5" gridrow="8" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF95EFB04F89777B4" box="[1729,1820,1180,1205]" gridcol="6" gridrow="8" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF8FAFB04F84B77B4" box="[1893,1984,1180,1205]" gridcol="7" gridrow="8" pageId="8" pageNumber="507">0</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFB54F84B77E4" box="[165,1984,1228,1253]" gridrow="9" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFB54FE3577E4" box="[165,446,1228,1253]" gridcol="0" gridrow="9" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFB54FEB477E4" bold="true" box="[165,319,1228,1253]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFB54FEBA77E4" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[165,305,1228,1253]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="tubernasus">
<emphasis id="B7972302FFE4A739FF3AFB54FEBA77E4" bold="true" box="[165,305,1228,1253]" italics="true" pageId="8" pageNumber="507">P. tubernasus</emphasis>
</taxonomicName>
*
</emphasis>
(N=1)
</th>
<td id="7802942EFFE458CEFDECFB54FD3077E4" box="[627,699,1228,1253]" gridcol="1" gridrow="9" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FB54FBCD77E4" box="[758,1094,1228,1253]" gridcol="2" gridrow="9" pageId="8" pageNumber="507">coarsely shagreen</td>
<td id="7802942EFFE458CEFB10FB54FB6277E4" box="[1167,1257,1228,1253]" gridcol="3" gridrow="9" pageId="8" pageNumber="507">shagreen</td>
<td id="7802942EFFE458CEFABCFB54F9AA77E4" box="[1315,1569,1228,1253]" gridcol="4" gridrow="9" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FB54F91C77E4" box="[1674,1687,1228,1253]" gridcol="5" gridrow="9" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF95EFB54F89777E4" box="[1729,1820,1228,1253]" gridcol="6" gridrow="9" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF8FAFB54F84B77E4" box="[1893,1984,1228,1253]" gridcol="7" gridrow="9" pageId="8" pageNumber="507">0</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFB64F84B7614" box="[165,1984,1276,1301]" gridrow="10" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFB64FE357614" box="[165,446,1276,1301]" gridcol="0" gridrow="10" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFB64FE857614" bold="true" box="[165,270,1276,1301]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFB64FE8B7614" baseAuthorityName="Lynch" baseAuthorityYear="1984" box="[165,256,1276,1301]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="lutitus">
<emphasis id="B7972302FFE4A739FF3AFB64FE8B7614" bold="true" box="[165,256,1276,1301]" italics="true" pageId="8" pageNumber="507">P. lutitus</emphasis>
</taxonomicName>
*
</emphasis>
(N=2)
</th>
<td id="7802942EFFE458CEFDECFB64FD307614" box="[627,699,1276,1301]" gridcol="1" gridrow="10" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FB64FBCD7614" box="[758,1094,1276,1301]" gridcol="2" gridrow="10" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFB10FB64FB627614" box="[1167,1257,1276,1301]" gridcol="3" gridrow="10" pageId="8" pageNumber="507">shagreen</td>
<td id="7802942EFFE458CEFABCFB64F9AA7614" box="[1315,1569,1276,1301]" gridcol="4" gridrow="10" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FB64F91C7614" box="[1674,1687,1276,1301]" gridcol="5" gridrow="10" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF95EFB64F8977614" box="[1729,1820,1276,1301]" gridcol="6" gridrow="10" pageId="8" pageNumber="507">-</td>
<td id="7802942EFFE458CEF8FAFB64F84B7614" box="[1893,1984,1276,1301]" gridcol="7" gridrow="10" pageId="8" pageNumber="507">-</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFAB4F84B7644" box="[165,1984,1324,1349]" gridrow="11" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFAB4FE357644" box="[165,446,1324,1349]" gridcol="0" gridrow="11" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFAB4FEC87644" bold="true" box="[165,323,1324,1349]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFAB4FEBD7644" baseAuthorityName="Lynch" baseAuthorityYear="1984" box="[165,310,1324,1349]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="merostictus">
<emphasis id="B7972302FFE4A739FF3AFAB4FEBD7644" bold="true" box="[165,310,1324,1349]" italics="true" pageId="8" pageNumber="507">P. merostictus</emphasis>
</taxonomicName>
*
</emphasis>
(N=3)
</th>
<td id="7802942EFFE458CEFDECFAB4FD307644" box="[627,699,1324,1349]" gridcol="1" gridrow="11" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FAB4FBCD7644" box="[758,1094,1324,1349]" gridcol="2" gridrow="11" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFB10FAB4FB627644" box="[1167,1257,1324,1349]" gridcol="3" gridrow="11" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFABCFAB4F9AA7644" box="[1315,1569,1324,1349]" gridcol="4" gridrow="11" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FAB4F91C7644" box="[1674,1687,1324,1349]" gridcol="5" gridrow="11" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF95EFAB4F8977644" box="[1729,1820,1324,1349]" gridcol="6" gridrow="11" pageId="8" pageNumber="507">weak</td>
<td id="7802942EFFE458CEF8FAFAB4F84B7644" box="[1893,1984,1324,1349]" gridcol="7" gridrow="11" pageId="8" pageNumber="507">weak</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFAC4F84B7674" box="[165,1984,1372,1397]" gridrow="12" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFAC4FE357674" box="[165,446,1372,1397]" gridcol="0" gridrow="12" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFAC4FEAA7674" bold="true" box="[165,289,1372,1397]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFAC4FE8C7674" authorityName="Rivera-Correa, Lamadrid-Feris &amp; Crawford" authorityYear="2016" box="[165,263,1372,1397]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="dorado">
<emphasis id="B7972302FFE4A739FF3AFAC4FE8C7674" bold="true" box="[165,263,1372,1397]" italics="true" pageId="8" pageNumber="507">P. dorado</emphasis>
</taxonomicName>
**
</emphasis>
(N=7)
</th>
<td id="7802942EFFE458CEFDECFAC4FD307674" box="[627,699,1372,1397]" gridcol="1" gridrow="12" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FAC4FBCD7674" box="[758,1094,1372,1397]" gridcol="2" gridrow="12" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFB10FAC4FB627674" box="[1167,1257,1372,1397]" gridcol="3" gridrow="12" pageId="8" pageNumber="507">shagreen</td>
<td id="7802942EFFE458CEFABCFAC4F9AA7674" box="[1315,1569,1372,1397]" gridcol="4" gridrow="12" pageId="8" pageNumber="507">-</td>
<td id="7802942EFFE458CEF915FAC4F91C7674" box="[1674,1687,1372,1397]" gridcol="5" gridrow="12" pageId="8" pageNumber="507">1</td>
<td id="7802942EFFE458CEF95EFAC4F8977674" box="[1729,1820,1372,1397]" gridcol="6" gridrow="12" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF8FAFAC4F84B7674" box="[1893,1984,1372,1397]" gridcol="7" gridrow="12" pageId="8" pageNumber="507">0</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFA13F84B76A5" box="[165,1984,1419,1444]" gridrow="13" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFA13FE3576A5" box="[165,446,1419,1444]" gridcol="0" gridrow="13" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFA13FEEE76A5" bold="true" box="[165,357,1419,1444]" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFA13FE8C76A5" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[165,263,1419,1444]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE4A739FF3AFA13FE8C76A5" bold="true" box="[165,263,1419,1444]" italics="true" pageId="8" pageNumber="507">P. ardilae</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="ACA49E79FFE4A739FE8BFA13FEEE76A5" box="[276,357,1419,1444]" pageId="8" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(N=10)
</th>
<td id="7802942EFFE458CEFDECFA13FD3076A5" box="[627,699,1419,1444]" gridcol="1" gridrow="13" pageId="8" pageNumber="507">F I&gt; II</td>
<td id="7802942EFFE458CEFD69FA13FBCD76A5" box="[758,1094,1419,1444]" gridcol="2" gridrow="13" pageId="8" pageNumber="507">Smooth, slightly granular</td>
<td id="7802942EFFE458CEFB10FA13FB6276A5" box="[1167,1257,1419,1444]" gridcol="3" gridrow="13" pageId="8" pageNumber="507">shagreen</td>
<td id="7802942EFFE458CEFABCFA13F9AA76A5" box="[1315,1569,1419,1444]" gridcol="4" gridrow="13" pageId="8" pageNumber="507">presence dorsal tubercles</td>
<td id="7802942EFFE458CEF915FA13F91C76A5" box="[1674,1687,1419,1444]" gridcol="5" gridrow="13" pageId="8" pageNumber="507">1</td>
<td id="7802942EFFE458CEF95EFA13F89776A5" box="[1729,1820,1419,1444]" gridcol="6" gridrow="13" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF8FAFA13F84B76A5" box="[1893,1984,1419,1444]" gridcol="7" gridrow="13" pageId="8" pageNumber="507">0</td>
</tr>
<tr id="3BD3FD52FFE458CEFF3AFA23F84B76D5" box="[165,1984,1467,1492]" gridrow="14" pageId="8" pageNumber="507">
<th id="7802942EFFE458CEFF3AFA23FE3576D5" box="[165,446,1467,1492]" gridcol="0" gridrow="14" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFA23FEE276D5" bold="true" box="[165,361,1467,1492]" pageId="8" pageNumber="507">
<emphasis id="B7972302FFE4A739FF3AFA23FE9976D5" bold="true" box="[165,274,1467,1492]" italics="true" pageId="8" pageNumber="507">
<taxonomicName id="42E38493FFE4A739FF3AFA23FE8776D5" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[165,268,1467,1492]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="8" pageNumber="507" phylum="Chordata" rank="species" species="bowara" status="sp. nov.">P. bowara</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="ACA49E79FFE4A739FE87FA23FEE276D5" box="[280,361,1467,1492]" pageId="8" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(N=5)
</th>
<td id="7802942EFFE458CEFDECFA23FD3076D5" box="[627,699,1467,1492]" gridcol="1" gridrow="14" pageId="8" pageNumber="507">F I &lt;II</td>
<td id="7802942EFFE458CEFD69FA23FBCD76D5" box="[758,1094,1467,1492]" gridcol="2" gridrow="14" pageId="8" pageNumber="507">granular</td>
<td id="7802942EFFE458CEFB10FA23FB6276D5" box="[1167,1257,1467,1492]" gridcol="3" gridrow="14" pageId="8" pageNumber="507">smooth</td>
<td id="7802942EFFE458CEFABCFA23F9AA76D5" box="[1315,1569,1467,1492]" gridcol="4" gridrow="14" pageId="8" pageNumber="507">short folds</td>
<td id="7802942EFFE458CEF915FA23F91C76D5" box="[1674,1687,1467,1492]" gridcol="5" gridrow="14" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF95EFA23F89776D5" box="[1729,1820,1467,1492]" gridcol="6" gridrow="14" pageId="8" pageNumber="507">0</td>
<td id="7802942EFFE458CEF8FAFA23F84B76D5" box="[1893,1984,1467,1492]" gridcol="7" gridrow="14" pageId="8" pageNumber="507">0</td>
</tr>
</table>
</paragraph>
<paragraph id="855CFF10FFE4A739F92BFA73F8607505" blockId="8.[1716,2027,1515,1540]" box="[1716,2027,1515,1540]" pageId="8" pageNumber="507">
<tableNote id="7805FE9EFFE4A739F92BFA73F8607505" box="[1716,2027,1515,1540]" pageId="8" pageNumber="507" targetBox="[165,1984,798,1492]" targetPageId="8">......continued on the next page</tableNote>
</paragraph>
<caption id="D19CAF98FFE5A738FF08FD27FE1171D8" box="[151,410,703,729]" pageId="9" pageNumber="508">
<paragraph id="855CFF10FFE5A738FF08FD27FE1171D8" blockId="9.[151,410,703,729]" box="[151,410,703,729]" pageId="9" pageNumber="508">TABLE 3. (Continued)</paragraph>
</caption>
<paragraph id="855CFF10FFE5A738FF3AFD74F82A76A3" pageId="9" pageNumber="508">
<table id="F7E30DB0FFE558CEFF3AFD74F85076A3" box="[165,2011,748,1442]" gridcols="8" gridrows="15" pageId="9" pageNumber="508">
<tr id="3BD3FD52FFE558CEFF3AFD74F8507004" box="[165,2011,748,773]" gridrow="0" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFD74FE357004" box="[165,446,748,773]" gridcol="0" gridrow="0" pageId="9" pageNumber="508">Species</th>
<th id="7802942EFFE558CEFD20FD74FD477004" box="[703,716,748,773]" gridcol="1" gridrow="0" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FD20FD74FD477004" bold="true" box="[703,716,748,773]" pageId="9" pageNumber="508">8</emphasis>
</th>
<th id="7802942EFFE558CEFD67FD74FC3F7004" box="[760,948,748,773]" gridcol="2" gridrow="0" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FD67FD74FC8E7004" bold="true" box="[760,773,748,773]" pageId="9" pageNumber="508">9</emphasis>
</th>
<th id="7802942EFFE558CEFB1AFD74FB157004" box="[1157,1182,748,773]" gridcol="3" gridrow="0" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FB1AFD74FB157004" bold="true" box="[1157,1182,748,773]" pageId="9" pageNumber="508">10</emphasis>
</th>
<th id="7802942EFFE558CEFB5DFD74F9937004" box="[1218,1560,748,773]" gridcol="4" gridrow="0" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FB5DFD74FB517004" bold="true" box="[1218,1242,748,773]" pageId="9" pageNumber="508">11</emphasis>
</th>
<th id="7802942EFFE558CEF93BFD74F9367004" box="[1700,1725,748,773]" gridcol="5" gridrow="0" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738F93BFD74F9367004" bold="true" box="[1700,1725,748,773]" pageId="9" pageNumber="508">12</emphasis>
</th>
<th id="7802942EFFE558CEF97AFD74F8C07004" box="[1765,1867,748,773]" gridcol="6" gridrow="0" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738F97AFD74F9757004" bold="true" box="[1765,1790,748,773]" pageId="9" pageNumber="508">13</emphasis>
</th>
<th id="7802942EFFE558CEF8EAFD74F8507004" box="[1909,2011,748,773]" gridcol="7" gridrow="0" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738F8EAFD74F8057004" bold="true" box="[1909,1934,748,773]" pageId="9" pageNumber="508">14</emphasis>
</th>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFC84F8507034" box="[165,2011,796,821]" gridrow="1" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFC84FE357034" box="[165,446,796,821]" gridcol="0" gridrow="1" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFC84FEF27034" bold="true" box="[165,377,796,821]" italics="true" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFC84FEE47034" baseAuthorityName="Lynch" baseAuthorityYear="1994" box="[165,367,796,821]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="carranguerorum">P. carranguerorum</taxonomicName>
*
</emphasis>
(N=3)
</th>
<td id="7802942EFFE558CEFD20FC84FD477034" box="[703,716,796,821]" gridcol="1" gridrow="1" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FC84FC3F7034" box="[760,948,796,821]" gridcol="2" gridrow="1" pageId="9" pageNumber="508">stained or variable</td>
<td id="7802942EFFE558CEFB1AFC84FB157034" box="[1157,1182,796,821]" gridcol="3" gridrow="1" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFB5DFC84F9937034" box="[1218,1560,796,821]" gridcol="4" gridrow="1" pageId="9" pageNumber="508">barely visible</td>
<td id="7802942EFFE558CEF93BFC84F9367034" box="[1700,1725,796,821]" gridcol="5" gridrow="1" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEF97AFC84F8C07034" box="[1765,1867,796,821]" gridcol="6" gridrow="1" pageId="9" pageNumber="508">32.7</td>
<td id="7802942EFFE558CEF8EAFC84F8507034" box="[1909,2011,796,821]" gridcol="7" gridrow="1" pageId="9" pageNumber="508">41.7–50.7</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFCD3F8507064" box="[165,2011,843,869]" gridrow="2" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFCD3FE357064" box="[165,446,843,869]" gridcol="0" gridrow="2" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFCD3FE967065" bold="true" box="[165,285,843,868]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFCD3FE9B7065" baseAuthorityName="Lynch" baseAuthorityYear="1994" box="[165,272,843,868]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="medemi">
<emphasis id="B7972302FFE5A738FF3AFCD3FE9B7065" bold="true" box="[165,272,843,868]" italics="true" pageId="9" pageNumber="508">P. medemi</emphasis>
</taxonomicName>
*
</emphasis>
</th>
<td id="7802942EFFE558CEFD20FCD3FD477064" box="[703,716,843,869]" gridcol="1" gridrow="2" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FCD3FC3F7064" box="[760,948,843,869]" gridcol="2" gridrow="2" pageId="9" pageNumber="508">stained or variable</td>
<td id="7802942EFFE558CEFB1AFCD3FB157064" box="[1157,1182,843,869]" gridcol="3" gridrow="2" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFB5DFCD3F9937064" box="[1218,1560,843,869]" gridcol="4" gridrow="2" pageId="9" pageNumber="508">barely visible and Weakly spotted</td>
<td id="7802942EFFE558CEF93BFCD3F9367064" box="[1700,1725,843,869]" gridcol="5" gridrow="2" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEF97AFCD3F8C07064" box="[1765,1867,843,869]" gridcol="6" gridrow="2" pageId="9" pageNumber="508">29.4–32.9</td>
<td id="7802942EFFE558CEF8EAFCD3F8507064" box="[1909,2011,843,869]" gridcol="7" gridrow="2" pageId="9" pageNumber="508">39.8–43.1</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFCE3F8507095" box="[165,2011,891,916]" gridrow="3" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFCE3FE357095" box="[165,446,891,916]" gridcol="0" gridrow="3" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFCE3FED47095" bold="true" box="[165,351,891,916]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFCE3FED97095" baseAuthorityName="La Marca" baseAuthorityYear="2004" box="[165,338,891,916]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="pedimontanus">
<emphasis id="B7972302FFE5A738FF3AFCE3FED97095" bold="true" box="[165,338,891,916]" italics="true" pageId="9" pageNumber="508">P. pedimontanus</emphasis>
</taxonomicName>
*
</emphasis>
(N=7)
</th>
<td id="7802942EFFE558CEFD20FCE3FD477095" box="[703,716,891,916]" gridcol="1" gridrow="3" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFD67FCE3FC3F7095" box="[760,948,891,916]" gridcol="2" gridrow="3" pageId="9" pageNumber="508">stained or variable</td>
<td id="7802942EFFE558CEFB1AFCE3FB157095" box="[1157,1182,891,916]" gridcol="3" gridrow="3" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFB5DFCE3F9937095" box="[1218,1560,891,916]" gridcol="4" gridrow="3" pageId="9" pageNumber="508">plain</td>
<td id="7802942EFFE558CEF93BFCE3F9367095" box="[1700,1725,891,916]" gridcol="5" gridrow="3" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEF97AFCE3F8C07095" box="[1765,1867,891,916]" gridcol="6" gridrow="3" pageId="9" pageNumber="508">26.2–33.6</td>
<td id="7802942EFFE558CEF8EAFCE3F8507095" box="[1909,2011,891,916]" gridcol="7" gridrow="3" pageId="9" pageNumber="508">38.8–44.8</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFC33F85070C5" box="[165,2011,939,964]" gridrow="4" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFC33FE3570C5" box="[165,446,939,964]" gridcol="0" gridrow="4" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFC33FE9E70C5" baseAuthorityName="Lynch" baseAuthorityYear="1983" box="[165,277,939,964]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="anolirex">
<emphasis id="B7972302FFE5A738FF3AFC33FE9E70C5" bold="true" box="[165,277,939,964]" italics="true" pageId="9" pageNumber="508">P. anolirex</emphasis>
</taxonomicName>
(N=8)
</th>
<td id="7802942EFFE558CEFD20FC33FD4770C5" box="[703,716,939,964]" gridcol="1" gridrow="4" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FC33FC3F70C5" box="[760,948,939,964]" gridcol="2" gridrow="4" pageId="9" pageNumber="508">stained</td>
<td id="7802942EFFE558CEFB1AFC33FB1570C5" box="[1157,1182,939,964]" gridcol="3" gridrow="4" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFB5DFC33F99370C5" box="[1218,1560,939,964]" gridcol="4" gridrow="4" pageId="9" pageNumber="508">barely visible</td>
<td id="7802942EFFE558CEF93BFC33F93670C5" box="[1700,1725,939,964]" gridcol="5" gridrow="4" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEF97AFC33F8C070C5" box="[1765,1867,939,964]" gridcol="6" gridrow="4" pageId="9" pageNumber="508">24–31.8</td>
<td id="7802942EFFE558CEF8EAFC33F85070C5" box="[1909,2011,939,964]" gridcol="7" gridrow="4" pageId="9" pageNumber="508">35.5–40</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFC43F85070F5" box="[165,2011,987,1012]" gridrow="5" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFC43FE3570F5" box="[165,446,987,1012]" gridcol="0" gridrow="5" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFC43FEB470F5" bold="true" box="[165,319,987,1012]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFC43FEBA70F5" baseAuthorityName="Lynch" baseAuthorityYear="2013" box="[165,305,987,1012]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="batrachites">
<emphasis id="B7972302FFE5A738FF3AFC43FEBA70F5" bold="true" box="[165,305,987,1012]" italics="true" pageId="9" pageNumber="508">P. batrachites</emphasis>
</taxonomicName>
*
</emphasis>
(N=3)
</th>
<td id="7802942EFFE558CEFD20FC43FD4770F5" box="[703,716,987,1012]" gridcol="1" gridrow="5" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FC43FC3F70F5" box="[760,948,987,1012]" gridcol="2" gridrow="5" pageId="9" pageNumber="508">stained</td>
<td id="7802942EFFE558CEFB1AFC43FB1570F5" box="[1157,1182,987,1012]" gridcol="3" gridrow="5" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFB5DFC43F99370F5" box="[1218,1560,987,1012]" gridcol="4" gridrow="5" pageId="9" pageNumber="508">barely visible and Weakly spotted</td>
<td id="7802942EFFE558CEF93BFC43F93670F5" box="[1700,1725,987,1012]" gridcol="5" gridrow="5" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEF97AFC43F8C070F5" box="[1765,1867,987,1012]" gridcol="6" gridrow="5" pageId="9" pageNumber="508">14–17</td>
<td id="7802942EFFE558CEF8EAFC43F85070F5" box="[1909,2011,987,1012]" gridcol="7" gridrow="5" pageId="9" pageNumber="508">17.9–21.7</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFB93F8507725" box="[165,2011,1035,1060]" gridrow="6" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFB93FE357725" box="[165,446,1035,1060]" gridcol="0" gridrow="6" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFB93FEED7725" bold="true" box="[165,358,1035,1060]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFB93FED27725" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[165,345,1035,1060]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="melanoproctus">
<emphasis id="B7972302FFE5A738FF3AFB93FED27725" bold="true" box="[165,345,1035,1060]" italics="true" pageId="9" pageNumber="508">P. melanoproctus</emphasis>
</taxonomicName>
*
</emphasis>
(N=1)
</th>
<td id="7802942EFFE558CEFD20FB93FD477725" box="[703,716,1035,1060]" gridcol="1" gridrow="6" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFD67FB93FC3F7725" box="[760,948,1035,1060]" gridcol="2" gridrow="6" pageId="9" pageNumber="508">stained</td>
<td id="7802942EFFE558CEFB1AFB93FB157725" box="[1157,1182,1035,1060]" gridcol="3" gridrow="6" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFB5DFB93F9937725" box="[1218,1560,1035,1060]" gridcol="4" gridrow="6" pageId="9" pageNumber="508">barely visible and Weakly spotted</td>
<td id="7802942EFFE558CEF93BFB93F9367725" box="[1700,1725,1035,1060]" gridcol="5" gridrow="6" pageId="9" pageNumber="508">-</td>
<td id="7802942EFFE558CEF97AFB93F8C07725" box="[1765,1867,1035,1060]" gridcol="6" gridrow="6" pageId="9" pageNumber="508">12–14.3</td>
<td id="7802942EFFE558CEF8EAFB93F8507725" box="[1909,2011,1035,1060]" gridcol="7" gridrow="6" pageId="9" pageNumber="508">20–28.2</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFBA3F8507755" box="[165,2011,1083,1108]" gridrow="7" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFBA3FE357755" box="[165,446,1083,1108]" gridcol="0" gridrow="7" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFBA3FEBA7755" bold="true" box="[165,305,1083,1108]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFBA3FEA87755" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[165,291,1083,1108]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="mondolfii">
<emphasis id="B7972302FFE5A738FF3AFBA3FEA87755" bold="true" box="[165,291,1083,1108]" italics="true" pageId="9" pageNumber="508">P. mondolfii</emphasis>
</taxonomicName>
*
</emphasis>
(N=2)
</th>
<td id="7802942EFFE558CEFD20FBA3FD477755" box="[703,716,1083,1108]" gridcol="1" gridrow="7" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFD67FBA3FC3F7755" box="[760,948,1083,1108]" gridcol="2" gridrow="7" pageId="9" pageNumber="508">stained</td>
<td id="7802942EFFE558CEFB1AFBA3FB157755" box="[1157,1182,1083,1108]" gridcol="3" gridrow="7" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFB5DFBA3F9937755" box="[1218,1560,1083,1108]" gridcol="4" gridrow="7" pageId="9" pageNumber="508">well-defined and spotted</td>
<td id="7802942EFFE558CEF93BFBA3F9367755" box="[1700,1725,1083,1108]" gridcol="5" gridrow="7" pageId="9" pageNumber="508">-</td>
<td id="7802942EFFE558CEF97AFBA3F8C07755" box="[1765,1867,1083,1108]" gridcol="6" gridrow="7" pageId="9" pageNumber="508">24.5–37.2</td>
<td id="7802942EFFE558CEF8EAFBA3F8507755" box="[1909,2011,1083,1108]" gridcol="7" gridrow="7" pageId="9" pageNumber="508">31.2–38.5</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFBF2F8507782" box="[165,2011,1130,1155]" gridrow="8" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFBF2FE357782" box="[165,446,1130,1155]" gridcol="0" gridrow="8" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFBF2FE9B7782" baseAuthorityName="Cochran &amp; Goin" baseAuthorityYear="1970" box="[165,272,1130,1155]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="nicefori">
<emphasis id="B7972302FFE5A738FF3AFBF2FE9B7782" bold="true" box="[165,272,1130,1155]" italics="true" pageId="9" pageNumber="508">P. nicefori</emphasis>
</taxonomicName>
(N=4)
</th>
<td id="7802942EFFE558CEFD20FBF2FD477782" box="[703,716,1130,1155]" gridcol="1" gridrow="8" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFD67FBF2FC3F7782" box="[760,948,1130,1155]" gridcol="2" gridrow="8" pageId="9" pageNumber="508">weakly spotted</td>
<td id="7802942EFFE558CEFB1AFBF2FB157782" box="[1157,1182,1130,1155]" gridcol="3" gridrow="8" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFB5DFBF2F9937782" box="[1218,1560,1130,1155]" gridcol="4" gridrow="8" pageId="9" pageNumber="508">barely visible</td>
<td id="7802942EFFE558CEF93BFBF2F9367782" box="[1700,1725,1130,1155]" gridcol="5" gridrow="8" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEF97AFBF2F8C07782" box="[1765,1867,1130,1155]" gridcol="6" gridrow="8" pageId="9" pageNumber="508">22.5–26.5</td>
<td id="7802942EFFE558CEF8EAFBF2F8507782" box="[1909,2011,1130,1155]" gridcol="7" gridrow="8" pageId="9" pageNumber="508">29.2–31.2</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFB02F85077B2" box="[165,2011,1178,1203]" gridrow="9" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFB02FE3577B2" box="[165,446,1178,1203]" gridcol="0" gridrow="9" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFB02FEB477B2" bold="true" box="[165,319,1178,1203]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFB02FEBA77B2" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[165,305,1178,1203]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="tubernasus">
<emphasis id="B7972302FFE5A738FF3AFB02FEBA77B2" bold="true" box="[165,305,1178,1203]" italics="true" pageId="9" pageNumber="508">P. tubernasus</emphasis>
</taxonomicName>
*
</emphasis>
(N=1)
</th>
<td id="7802942EFFE558CEFD20FB02FD4777B2" box="[703,716,1178,1203]" gridcol="1" gridrow="9" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FB02FC3F77B2" box="[760,948,1178,1203]" gridcol="2" gridrow="9" pageId="9" pageNumber="508">stained</td>
<td id="7802942EFFE558CEFB1AFB02FB1577B2" box="[1157,1182,1178,1203]" gridcol="3" gridrow="9" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFB5DFB02F99377B2" box="[1218,1560,1178,1203]" gridcol="4" gridrow="9" pageId="9" pageNumber="508">plain</td>
<td id="7802942EFFE558CEF93BFB02F93677B2" box="[1700,1725,1178,1203]" gridcol="5" gridrow="9" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEF97AFB02F8C077B2" box="[1765,1867,1178,1203]" gridcol="6" gridrow="9" pageId="9" pageNumber="508">26</td>
<td id="7802942EFFE558CEF8EAFB02F85077B2" box="[1909,2011,1178,1203]" gridcol="7" gridrow="9" pageId="9" pageNumber="508">22.5–24</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFB52F85077E2" box="[165,2011,1226,1251]" gridrow="10" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFB52FE3577E2" box="[165,446,1226,1251]" gridcol="0" gridrow="10" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFB52FE8577E2" bold="true" box="[165,270,1226,1251]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFB52FE8B77E2" baseAuthorityName="Lynch" baseAuthorityYear="1984" box="[165,256,1226,1251]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="lutitus">
<emphasis id="B7972302FFE5A738FF3AFB52FE8B77E2" bold="true" box="[165,256,1226,1251]" italics="true" pageId="9" pageNumber="508">P. lutitus</emphasis>
</taxonomicName>
*
</emphasis>
(N=2)
</th>
<td id="7802942EFFE558CEFD20FB52FD4777E2" box="[703,716,1226,1251]" gridcol="1" gridrow="10" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FB52FC3F77E2" box="[760,948,1226,1251]" gridcol="2" gridrow="10" pageId="9" pageNumber="508">stained</td>
<td id="7802942EFFE558CEFB1AFB52FB1577E2" box="[1157,1182,1226,1251]" gridcol="3" gridrow="10" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFB5DFB52F99377E2" box="[1218,1560,1226,1251]" gridcol="4" gridrow="10" pageId="9" pageNumber="508">barely visible</td>
<td id="7802942EFFE558CEF93BFB52F93677E2" box="[1700,1725,1226,1251]" gridcol="5" gridrow="10" pageId="9" pageNumber="508">-</td>
<td id="7802942EFFE558CEF97AFB52F8C077E2" box="[1765,1867,1226,1251]" gridcol="6" gridrow="10" pageId="9" pageNumber="508">-</td>
<td id="7802942EFFE558CEF8EAFB52F85077E2" box="[1909,2011,1226,1251]" gridcol="7" gridrow="10" pageId="9" pageNumber="508">23.7–28</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFB62F8507612" box="[165,2011,1274,1299]" gridrow="11" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFB62FE357612" box="[165,446,1274,1299]" gridcol="0" gridrow="11" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFB62FEC87612" bold="true" box="[165,323,1274,1299]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFB62FEBD7612" baseAuthorityName="Lynch" baseAuthorityYear="1984" box="[165,310,1274,1299]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="merostictus">
<emphasis id="B7972302FFE5A738FF3AFB62FEBD7612" bold="true" box="[165,310,1274,1299]" italics="true" pageId="9" pageNumber="508">P. merostictus</emphasis>
</taxonomicName>
*
</emphasis>
(N=3)
</th>
<td id="7802942EFFE558CEFD20FB62FD477612" box="[703,716,1274,1299]" gridcol="1" gridrow="11" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FB62FC3F7612" box="[760,948,1274,1299]" gridcol="2" gridrow="11" pageId="9" pageNumber="508">stained or variable</td>
<td id="7802942EFFE558CEFB1AFB62FB157612" box="[1157,1182,1274,1299]" gridcol="3" gridrow="11" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEFB5DFB62F9937612" box="[1218,1560,1274,1299]" gridcol="4" gridrow="11" pageId="9" pageNumber="508">Well-defined and spotted</td>
<td id="7802942EFFE558CEF93BFB62F9367612" box="[1700,1725,1274,1299]" gridcol="5" gridrow="11" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEF97AFB62F8C07612" box="[1765,1867,1274,1299]" gridcol="6" gridrow="11" pageId="9" pageNumber="508">16.5–22.5</td>
<td id="7802942EFFE558CEF8EAFB62F8507612" box="[1909,2011,1274,1299]" gridcol="7" gridrow="11" pageId="9" pageNumber="508">31</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFAB2F8507642" box="[165,2011,1322,1347]" gridrow="12" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFAB2FE357642" box="[165,446,1322,1347]" gridcol="0" gridrow="12" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFAB2FEAA7642" bold="true" box="[165,289,1322,1347]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFAB2FE8C7642" authorityName="Rivera-Correa, Lamadrid-Feris &amp; Crawford" authorityYear="2016" box="[165,263,1322,1347]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="dorado">
<emphasis id="B7972302FFE5A738FF3AFAB2FE8C7642" bold="true" box="[165,263,1322,1347]" italics="true" pageId="9" pageNumber="508">P. dorado</emphasis>
</taxonomicName>
**
</emphasis>
(N=7)
</th>
<td id="7802942EFFE558CEFD20FAB2FD477642" box="[703,716,1322,1347]" gridcol="1" gridrow="12" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FAB2FC3F7642" box="[760,948,1322,1347]" gridcol="2" gridrow="12" pageId="9" pageNumber="508">weakly spotted</td>
<td id="7802942EFFE558CEFB1AFAB2FB157642" box="[1157,1182,1322,1347]" gridcol="3" gridrow="12" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFB5DFAB2F9937642" box="[1218,1560,1322,1347]" gridcol="4" gridrow="12" pageId="9" pageNumber="508">plain or Weakly spotted</td>
<td id="7802942EFFE558CEF93BFAB2F9367642" box="[1700,1725,1322,1347]" gridcol="5" gridrow="12" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEF97AFAB2F8C07642" box="[1765,1867,1322,1347]" gridcol="6" gridrow="12" pageId="9" pageNumber="508">14.2–17.9</td>
<td id="7802942EFFE558CEF8EAFAB2F8507642" box="[1909,2011,1322,1347]" gridcol="7" gridrow="12" pageId="9" pageNumber="508">25.6</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFAC1F8507673" box="[165,2011,1369,1394]" gridrow="13" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFAC1FE357673" box="[165,446,1369,1394]" gridcol="0" gridrow="13" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFAC1FEEE7673" bold="true" box="[165,357,1369,1394]" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFAC1FE8C7673" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[165,263,1369,1394]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE5A738FF3AFAC1FE8C7673" bold="true" box="[165,263,1369,1394]" italics="true" pageId="9" pageNumber="508">P. ardilae</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="ACA49E79FFE5A738FE8BFAC1FEEE7673" box="[276,357,1369,1394]" pageId="9" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(N=10)
</th>
<td id="7802942EFFE558CEFD20FAC1FD477673" box="[703,716,1369,1394]" gridcol="1" gridrow="13" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FAC1FC3F7673" box="[760,948,1369,1394]" gridcol="2" gridrow="13" pageId="9" pageNumber="508">weakly spotted</td>
<td id="7802942EFFE558CEFB1AFAC1FB157673" box="[1157,1182,1369,1394]" gridcol="3" gridrow="13" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFB5DFAC1F9937673" box="[1218,1560,1369,1394]" gridcol="4" gridrow="13" pageId="9" pageNumber="508">Well-defined</td>
<td id="7802942EFFE558CEF93BFAC1F9367673" box="[1700,1725,1369,1394]" gridcol="5" gridrow="13" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEF97AFAC1F8C07673" box="[1765,1867,1369,1394]" gridcol="6" gridrow="13" pageId="9" pageNumber="508">25.1–29.1</td>
<td id="7802942EFFE558CEF8EAFAC1F8507673" box="[1909,2011,1369,1394]" gridcol="7" gridrow="13" pageId="9" pageNumber="508">36.3–39.7</td>
</tr>
<tr id="3BD3FD52FFE558CEFF3AFA11F85076A3" box="[165,2011,1417,1442]" gridrow="14" pageId="9" pageNumber="508">
<th id="7802942EFFE558CEFF3AFA11FE3576A3" box="[165,446,1417,1442]" gridcol="0" gridrow="14" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFA11FEE276A3" bold="true" box="[165,361,1417,1442]" pageId="9" pageNumber="508">
<emphasis id="B7972302FFE5A738FF3AFA11FE9976A3" bold="true" box="[165,274,1417,1442]" italics="true" pageId="9" pageNumber="508">
<taxonomicName id="42E38493FFE5A738FF3AFA11FE8776A3" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[165,268,1417,1442]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="9" pageNumber="508" phylum="Chordata" rank="species" species="bowara" status="sp. nov.">P. bowara</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="ACA49E79FFE5A738FE87FA11FEE276A3" box="[280,361,1417,1442]" pageId="9" pageNumber="508" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(N=5)
</th>
<td id="7802942EFFE558CEFD20FA11FD4776A3" box="[703,716,1417,1442]" gridcol="1" gridrow="14" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFD67FA11FC3F76A3" box="[760,948,1417,1442]" gridcol="2" gridrow="14" pageId="9" pageNumber="508">heavily spotted</td>
<td id="7802942EFFE558CEFB1AFA11FB1576A3" box="[1157,1182,1417,1442]" gridcol="3" gridrow="14" pageId="9" pageNumber="508">0</td>
<td id="7802942EFFE558CEFB5DFA11F99376A3" box="[1218,1560,1417,1442]" gridcol="4" gridrow="14" pageId="9" pageNumber="508">Well-defined</td>
<td id="7802942EFFE558CEF93BFA11F93676A3" box="[1700,1725,1417,1442]" gridcol="5" gridrow="14" pageId="9" pageNumber="508">1</td>
<td id="7802942EFFE558CEF97AFA11F8C076A3" box="[1765,1867,1417,1442]" gridcol="6" gridrow="14" pageId="9" pageNumber="508">28.4–25.5</td>
<td id="7802942EFFE558CEF8EAFA11F85076A3" box="[1909,2011,1417,1442]" gridcol="7" gridrow="14" pageId="9" pageNumber="508">39.7</td>
</tr>
</table>
</paragraph>
<tableNote id="7805FE9EFFE5A738FF08FA29FC8276EE" pageId="9" pageNumber="508" targetBox="[165,2011,748,1442]" targetPageId="9">
<paragraph id="855CFF10FFE5A738FF08FA29FC0B76CA" blockId="9.[151,896,1457,1519]" box="[151,896,1457,1483]" pageId="9" pageNumber="508">*complementary information was taken from the original description.</paragraph>
<paragraph id="855CFF10FFE5A738FF08FA4DFC8276EE" blockId="9.[151,896,1457,1519]" box="[151,777,1493,1519]" pageId="9" pageNumber="508">**All information was taken from the original description.</paragraph>
</tableNote>
<caption id="D19CAF98FFE6A73BFF08F908FE0875CC" ID-DOI="http://doi.org/10.5281/zenodo.3708848" ID-Zenodo-Dep="3708848" httpUri="https://zenodo.org/record/3708848/files/figure.png" pageId="10" pageNumber="509" startId="10.[151,250,1680,1705]" targetBox="[160,1419,196,1645]" targetPageId="10">
<paragraph id="855CFF10FFE6A73BFF08F908FE0875CC" blockId="10.[151,1436,1680,1741]" pageId="10" pageNumber="509">
<emphasis id="B7972302FFE6A73BFF08F908FE9F75A8" bold="true" box="[151,276,1680,1705]" pageId="10" pageNumber="509">FIGURE 5.</emphasis>
Photos of the holotype specimens. A. 
<taxonomicName id="42E38493FFE6A73BFD3EF90AFCE775A8" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[673,876,1681,1705]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="509" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE6A73BFD3EF90AFCE775A8" box="[673,876,1681,1705]" italics="true" pageId="10" pageNumber="509">Pristimantis ardilae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACA49E79FFE6A73BFCEBF909FC4D75A8" box="[884,966,1681,1705]" pageId="10" pageNumber="509" rank="species">
<emphasis id="B7972302FFE6A73BFCEBF909FC4D75A8" bold="true" box="[884,966,1681,1705]" pageId="10" pageNumber="509">sp. nov.</emphasis>
</taxonomicNameLabel>
(MCNUP-H 117) B. 
<taxonomicName id="42E38493FFE6A73BFB39F90AFAFD75A8" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[1190,1398,1681,1705]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="509" phylum="Chordata" rank="species" species="bowara" status="sp. nov.">
<emphasis id="B7972302FFE6A73BFB39F90AFAFD75A8" box="[1190,1398,1681,1705]" italics="true" pageId="10" pageNumber="509">Pristimantis bowara</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE6A73BFAE1F909FF4875CC" bold="true" pageId="10" pageNumber="509">
<taxonomicNameLabel id="ACA49E79FFE6A73BFAE1F909FF4875CC" pageId="10" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(MCNUP-H 165).
</paragraph>
</caption>
<paragraph id="855CFF10FFE6A73AFF58F963FBA072F6" blockId="10.[151,1437,1787,2030]" lastBlockId="11.[151,1437,152,1979]" lastPageId="11" lastPageNumber="510" pageId="10" pageNumber="509">
Of the 33 species that are part of the 
<taxonomicName id="42E38493FFE6A73BFDF0F964FCAC7414" baseAuthorityName="Günther" baseAuthorityYear="1858" box="[623,807,1787,1813]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="509" phylum="Chordata" rank="species" species="conspicillatus">
<emphasis id="B7972302FFE6A73BFDF0F964FCAC7414" box="[623,807,1787,1813]" italics="true" pageId="10" pageNumber="509">P. conspicillatus</emphasis>
</taxonomicName>
species group, only 
<taxonomicName id="42E38493FFE6A73BFB8DF964FB627414" baseAuthorityName="Lynch" baseAuthorityYear="1994" box="[1042,1257,1788,1813]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="509" phylum="Chordata" rank="species" species="carranguerorum">
<emphasis id="B7972302FFE6A73BFB8DF964FB627414" box="[1042,1257,1788,1813]" italics="true" pageId="10" pageNumber="509">P. carranguerorum</emphasis>
</taxonomicName>
, and 
<taxonomicName id="42E38493FFE6A73BFAB7F964FEBB743B" authority="(Lynch 1994)" baseAuthorityName="Lynch" baseAuthorityYear="1994" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="509" phylum="Chordata" rank="species" species="medemi">
<emphasis id="B7972302FFE6A73BFAB7F964FA177414" box="[1320,1436,1787,1813]" italics="true" pageId="10" pageNumber="509">P. medemi</emphasis>
(Lynch 1994)
</taxonomicName>
are found in the Eastern Cordillera of 
<collectingCountry id="FDF4BF80FFE6A73BFD77F887FCD27438" box="[744,857,1823,1849]" name="Colombia" pageId="10" pageNumber="509">Colombia</collectingCountry>
, the remaining species are found in distant biogeographic areas (Amazon and Cerrado, the Guianas and in the Atlantic Forest). Therefore, the comparison was based on 
<taxonomicName id="42E38493FFE6A73BFF25F8F0FEC87480" baseAuthorityName="Frost" baseAuthorityYear="2019" box="[186,323,1896,1921]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="509" phylum="Chordata" rank="genus">
<emphasis id="B7972302FFE6A73BFF25F8F0FEC87480" box="[186,323,1896,1921]" italics="true" pageId="10" pageNumber="509">Pristimantis</emphasis>
</taxonomicName>
species present in surrounding areas of the Northeast of 
<collectingCountry id="FDF4BF80FFE6A73BFC25F8FFFBA37480" box="[954,1064,1895,1921]" name="Colombia" pageId="10" pageNumber="509">Colombia</collectingCountry>
and the Andes of 
<collectingCountry id="FDF4BF80FFE6A73BFB6EF8FFFAE37480" box="[1265,1384,1895,1921]" name="Venezuela" pageId="10" pageNumber="509">Venezuela</collectingCountry>
. The character state of the compared species is between parentheses. 
<taxonomicName id="42E38493FFE6A73BFCF9F814FBCD74A4" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[870,1094,1931,1957]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="509" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE6A73BFCF9F814FBCD74A4" box="[870,1094,1931,1957]" italics="true" pageId="10" pageNumber="509">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE6A73BFBD1F814FB2374A7" bold="true" box="[1102,1192,1932,1958]" pageId="10" pageNumber="509">
<taxonomicNameLabel id="ACA49E79FFE6A73BFBD1F814FB2374A7" box="[1102,1192,1932,1958]" pageId="10" pageNumber="509" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from 
<taxonomicName id="42E38493FFE6A73BFF08F828FEE574C8" baseAuthorityName="Lynch" baseAuthorityYear="1994" box="[151,366,1968,1993]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="509" phylum="Chordata" rank="species" species="carranguerorum">
<emphasis id="B7972302FFE6A73BFF08F828FEE574C8" box="[151,366,1968,1993]" italics="true" pageId="10" pageNumber="509">P. carranguerorum</emphasis>
</taxonomicName>
for having the first finger slightly longer than second (Finger I &lt;
<collectionCode id="E3F267D5FFE6A73BFBC1F828FBFF74CB" box="[1118,1140,1968,1994]" pageId="10" pageNumber="509">II</collectionCode>
); fingers and toes without lateral fringes or keels (fingers and toes bearing fleshy lateral keels); Throat weakly spotted (heavily marble with brown). Tarsal fold absent (presente); 
<taxonomicName id="42E38493FFE7A73AFDDEFF02FD2773B2" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[577,684,153,179]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE7A73AFDDEFF02FD2773B2" box="[577,684,153,179]" italics="true" pageId="11" pageNumber="510">P. ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE7A73AFD2DFF01FC8773B2" bold="true" box="[690,780,153,179]" pageId="11" pageNumber="510">
<taxonomicNameLabel id="ACA49E79FFE7A73AFD2DFF01FC8773B2" box="[690,780,153,179]" pageId="11" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is smaller (male SVL: 25.1–29.1, female SVL: 36.3–39.7) than 
<taxonomicName id="42E38493FFE7A73AFF51FF26FE2873D6" baseAuthorityName="Lynch" baseAuthorityYear="1994" box="[206,419,190,215]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="carranguerorum">
<emphasis id="B7972302FFE7A73AFF51FF26FE2873D6" box="[206,419,190,215]" italics="true" pageId="11" pageNumber="510">P. carranguerorum</emphasis>
</taxonomicName>
(male SVL: 32.7 and female SVL: 41.7–50.7). 
<taxonomicName id="42E38493FFE7A73AFC29FF26FB1773D6" baseAuthorityName="Lynch" baseAuthorityYear="1994" box="[950,1180,189,215]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="medemi">
<emphasis id="B7972302FFE7A73AFC29FF26FB1773D6" box="[950,1180,189,215]" italics="true" pageId="11" pageNumber="510">Pristimantis medemi</emphasis>
</taxonomicName>
is superficially similar to 
<taxonomicName id="42E38493FFE7A73AFF2AFF7AFEAA73FA" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[181,289,225,251]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE7A73AFF2AFF7AFEAA73FA" box="[181,289,225,251]" italics="true" pageId="11" pageNumber="510">P. ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE7A73AFEB7FF79FE0873FA" bold="true" box="[296,387,225,251]" pageId="11" pageNumber="510">
<taxonomicNameLabel id="ACA49E79FFE7A73AFEB7FF79FE0873FA" box="[296,387,225,251]" pageId="11" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but differ in the states of the following characters: short dorsolateral folds in scapular region (absent); fingers and toes without lateral keels (fingers and toes bearing lateral keels). 
<taxonomicName id="42E38493FFE7A73AFBC3FE9EFAB7721E" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[1116,1340,261,287]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE7A73AFBC3FE9EFAB7721E" box="[1116,1340,261,287]" italics="true" pageId="11" pageNumber="510">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE7A73AFADCFE9DFA17721E" bold="true" box="[1347,1436,261,287]" pageId="11" pageNumber="510">
<taxonomicNameLabel id="ACA49E79FFE7A73AFADCFE9DFA17721E" box="[1347,1436,261,287]" pageId="11" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has no marks on the groin and hidden surfaces of the hind limbs (present in 
<taxonomicName id="42E38493FFE7A73AFC44FEB2FB107242" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[987,1179,297,323]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="melanoproctus">
<emphasis id="B7972302FFE7A73AFC44FEB2FB107242" box="[987,1179,297,323]" italics="true" pageId="11" pageNumber="510">P. melanoproctus</emphasis>
</taxonomicName>
(
<bibRefCitation id="E17282E1FFE7A73AFB36FEB1FABE7242" author="Rivero, J. A." box="[1193,1333,297,323]" pageId="11" pageNumber="510" pagination="57 - 132" refId="ref12568" refString="Rivero, J. A. (1984 &quot; 1982 &quot;) Los Eleutherodactylus (Amphibia, Leptodactylidae) de los Andes Venezolanos II. Especies subparameras. Memoria Sociedad Ciencias Naturales La Salle, 42, 57 - 132." type="journal article" year="1984">Rivero 1984</bibRefCitation>
“1982”), 
<taxonomicName id="42E38493FFE7A73AFF08FED6FE957266" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[151,286,333,359]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="mondolfii">
<emphasis id="B7972302FFE7A73AFF08FED6FE957266" box="[151,286,333,359]" italics="true" pageId="11" pageNumber="510">P. mondolfii</emphasis>
</taxonomicName>
, 
<taxonomicName id="42E38493FFE7A73AFEB2FED6FE2E7266" baseAuthorityName="Lynch" baseAuthorityYear="1983" box="[301,421,333,359]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="anolirex">
<emphasis id="B7972302FFE7A73AFEB2FED6FE2E7266" box="[301,421,333,359]" italics="true" pageId="11" pageNumber="510">P. anolirex</emphasis>
</taxonomicName>
). It is a larger species (with a maximum SVL of 
<quantity id="421B52F5FFE7A73AFC51FED5FBCA7266" box="[974,1089,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="3.973" pageId="11" pageNumber="510" unit="mm" value="39.73">39.73 mm</quantity>
females; 
<quantity id="421B52F5FFE7A73AFB31FED5FA987266" box="[1198,1299,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="2.91" pageId="11" pageNumber="510" unit="mm" value="29.1">29.1 mm</quantity>
males) than species that are geographically and phylogenetically close, (
<tableCitation id="C861CAABFFE7A73AFCDDFEE9FC11728A" box="[834,922,369,395]" captionStart="TABLE 3" captionStartId="8.[151,239,645,671]" captionText="TABLE 3. Comparison of diagnostic characters of some species of the Pristimantis conspicillatus group and species from northeastern Colombia, including the new species:" pageId="11" pageNumber="510">Table 3</tableCitation>
). 
<taxonomicName id="42E38493FFE7A73AFC2EFEEAFB18728A" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[945,1171,369,395]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE7A73AFC2EFEEAFB18728A" box="[945,1171,369,395]" italics="true" pageId="11" pageNumber="510">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE7A73AFB04FEE9FB7D728A" bold="true" box="[1179,1270,369,395]" pageId="11" pageNumber="510">
<taxonomicNameLabel id="ACA49E79FFE7A73AFB04FEE9FB7D728A" box="[1179,1270,369,395]" pageId="11" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
does not have cranial crests (present in 
<taxonomicName id="42E38493FFE7A73AFE34FE0EFDA972AE" baseAuthorityName="Lynch" baseAuthorityYear="1983" box="[427,546,405,431]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="anolirex">
<emphasis id="B7972302FFE7A73AFE34FE0EFDA972AE" box="[427,546,405,431]" italics="true" pageId="11" pageNumber="510">P. anolirex</emphasis>
</taxonomicName>
). Its first finger is longer than the second (the first finger is shorter than or about the same size as the second in 
<taxonomicName id="42E38493FFE7A73AFE6CFE22FDE772D2" baseAuthorityName="Lynch" baseAuthorityYear="1983" box="[499,620,441,467]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="anolirex">
<emphasis id="B7972302FFE7A73AFE6CFE22FDE772D2" box="[499,620,441,467]" italics="true" pageId="11" pageNumber="510">P. anolirex</emphasis>
</taxonomicName>
, 
<taxonomicName id="42E38493FFE7A73AFDE4FE22FD7772D2" box="[635,764,441,467]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="mondolfi">
<emphasis id="B7972302FFE7A73AFDE4FE22FD7772D2" box="[635,764,441,467]" italics="true" pageId="11" pageNumber="510">P. mondolfi</emphasis>
</taxonomicName>
, 
<taxonomicName id="42E38493FFE7A73AFC95FE22FC4772D2" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[778,972,441,467]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="melanoproctus">
<emphasis id="B7972302FFE7A73AFC95FE22FC4772D2" box="[778,972,441,467]" italics="true" pageId="11" pageNumber="510">P. melanoproctus</emphasis>
</taxonomicName>
). Inner and outer edges of tarsus lacking tubercles (present in 
<taxonomicName id="42E38493FFE7A73AFE1EFE46FD9372F6" baseAuthorityName="Rivero" baseAuthorityYear="1984" box="[385,536,477,503]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="tubernasus">
<emphasis id="B7972302FFE7A73AFE1EFE46FD9372F6" box="[385,536,477,503]" italics="true" pageId="11" pageNumber="510">P. tubernasus</emphasis>
</taxonomicName>
(
<bibRefCitation id="E17282E1FFE7A73AFDB9FE45FD3872F6" author="Rivero, J. A." box="[550,691,477,503]" pageId="11" pageNumber="510" pagination="57 - 132" refId="ref12568" refString="Rivero, J. A. (1984 &quot; 1982 &quot;) Los Eleutherodactylus (Amphibia, Leptodactylidae) de los Andes Venezolanos II. Especies subparameras. Memoria Sociedad Ciencias Naturales La Salle, 42, 57 - 132." type="journal article" year="1984">Rivero 1984</bibRefCitation>
“1982”), 
<taxonomicName id="42E38493FFE7A73AFCBDFE46FC1172F6" baseAuthorityName="Lynch" baseAuthorityYear="1983" box="[802,922,477,503]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="anolirex">
<emphasis id="B7972302FFE7A73AFCBDFE46FC1172F6" box="[802,922,477,503]" italics="true" pageId="11" pageNumber="510">P. anolirex</emphasis>
</taxonomicName>
, 
<taxonomicName id="42E38493FFE7A73AFC37FE46FB9072F6" baseAuthorityName="Cochran &amp; Goin" baseAuthorityYear="1970" box="[936,1051,477,503]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="nicefori">
<emphasis id="B7972302FFE7A73AFC37FE46FB9072F6" box="[936,1051,477,503]" italics="true" pageId="11" pageNumber="510">P. nicefori</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="CDF9AC9BFFE7A73AFF58FD98FDC0750A" pageId="11" pageNumber="510" type="description">
<paragraph id="855CFF10FFE7A73AFF58FD98FAB07082" blockId="11.[151,1437,152,1979]" pageId="11" pageNumber="510">
<emphasis id="B7972302FFE7A73AFF58FD98FD85711A" bold="true" box="[199,526,512,539]" pageId="11" pageNumber="510">
Description of the 
<typeStatus id="5A5841B2FFE7A73AFE3EFD99FD85711A" box="[417,526,513,539]" pageId="11" pageNumber="510" type="holotype">Holotype</typeStatus>
</emphasis>
. SVL: 
<quantity id="421B52F5FFE7A73AFDC1FD98FD59711A" box="[606,722,512,539]" metricMagnitude="-2" metricUnit="m" metricValue="3.9509999999999996" pageId="11" pageNumber="510" unit="mm" value="39.51">39.51 mm</quantity>
; upper eyelid tubercles absent in preservative (slightly visible in life); HL 38% of the SVL; HW 37% of the SVL; snout subacuminate in a dorsal view and rounded in profile; END 34% of the HW; tongue longer than wide. ED slightly shorter (88%) than the END; canthus rostralis distinguishable, straight in dorsal view and slightly rounded in cross-section; IND 70% of the END; IOD 96% of the IND. Cranial crests absent; supratympanic fold evident, tympanic annulus and tympanic membrane present, TD 34% of the ED; distance tympanum-eye: 
<quantity id="421B52F5FFE7A73AFE31FD2DFD9871CE" box="[430,531,693,719]" metricMagnitude="-3" metricUnit="m" metricValue="2.96" pageId="11" pageNumber="510" unit="mm" value="2.96">2.96 mm</quantity>
. TD 9.8% of the HL. Choanae small, round; dentigerous processes of vomers with 6 teeth. Premaxilla with 16 teeth. Ulnar tubercles absent. Fingers and toes without lateral fringes. FI&gt; FII; disks on FIII and FIV moderately expanded (state 2 according to 
<bibRefCitation id="E17282E1FFE7A73AFC99FD65FC4E7019" author="Grant, T. &amp; Frost, D. R. &amp; Caldwell, J. P. &amp; Gagliardo, R. &amp; Haddad, C. F. B. &amp; Kok, P. J. R. &amp; Means, D. B. &amp; Noonan, B. P. &amp; Schargel, W. E. &amp; Wheeler, W." box="[774,965,765,792]" pageId="11" pageNumber="510" pagination="1 - 262" refId="ref10824" refString="Grant, T., Frost, D. R., Caldwell, J. P., Gagliardo, R., Haddad, C. F. B., Kok, P. J. R., Means, D. B., Noonan, B. P., Schargel, W. E. &amp; Wheeler, W. (2006) Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia, Athesphatanura, Dendrobatidae). Bulletin of the American Museum of Natural History, 299, 1 - 262. https: // doi. org / 10.1206 / 0003 - 0090 (2006) 299 [1: PSODFA] 2.0. CO; 2" type="journal article" year="2006">
Grant 
<emphasis id="B7972302FFE7A73AFCD4FD66FCF77016" box="[843,892,765,791]" italics="true" pageId="11" pageNumber="510">et al</emphasis>
. 2006
</bibRefCitation>
); relative size of the yuxtaposed fingers: II &lt;I &lt;IV &lt;III. Subarticular tubercles round and upraised, the largest one on FI. Palmar tubercle bifid; supernumerary tubercles distinct, protruding; thenar tubercle 80% the size of palmar tubercle. Tarsal and calcar tubercles absent. Relative size of toes: I &lt;II &lt;III &lt;V &lt;IV; toe disks elliptical; toe webbing absent. LHU 66 % of the FOL.
</paragraph>
<paragraph id="855CFF10FFE7A73AFF58FC14FE5F775A" blockId="11.[151,1437,152,1979]" pageId="11" pageNumber="510">
<emphasis id="B7972302FFE7A73AFF58FC14FED070A6" bold="true" box="[199,347,908,935]" pageId="11" pageNumber="510">Color in life.</emphasis>
The dorsum usually intergrades yellowish brown tones against a predominantly dark brown background. Head lighter cream and copper color than the dorsal part, with a large, dark, nearly triangular, interorbital spot. The upper lip has pale stripes. The flanks are predominantly lighter, often crossed by oblique brown bands, more evident toward the inguinal area. Ventral surface light cream; throat speckled with small dark brown spots. Upper arms and forearms cream color with brown to reddish transverse bands. The inguinal area is lighter with slightly mottled with cream.
</paragraph>
<paragraph id="855CFF10FFE7A73AFF58FBFCFB0F760E" blockId="11.[151,1437,152,1979]" pageId="11" pageNumber="510">
<emphasis id="B7972302FFE7A73AFF58FBFCFE46777E" bold="true" box="[199,461,1124,1151]" pageId="11" pageNumber="510">Color in preservative.</emphasis>
Specimens fixed with 10% formalin and stored in alcohol at 75% presented a dorsum with a dark brown coloration, with the head darker than the body, the lips maintained a pattern of light and brown stripes, although less defined than in life.The ventral part maintained a clear cream coloration, with a weak pattern towards the neck identifiable by small dark brown mottles. Slightly creamy extremities, but with a predominance of a dark copper color, the arms and hind limbs displayed weak traces of dark brown stripes.
</paragraph>
<paragraph id="855CFF10FFE7A73AFF58FA80FDC0750A" blockId="11.[151,1437,152,1979]" pageId="11" pageNumber="510">
<emphasis id="B7972302FFE7A73AFF58FA80FEB47632" bold="true" box="[199,319,1304,1331]" pageId="11" pageNumber="510">Variation.</emphasis>
<taxonomicName id="42E38493FFE7A73AFED8FA82FDA37632" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[327,552,1305,1331]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE7A73AFED8FA82FDA37632" box="[327,552,1305,1331]" italics="true" pageId="11" pageNumber="510">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE7A73AFDAFFA81FD017632" bold="true" box="[560,650,1305,1331]" pageId="11" pageNumber="510">
<taxonomicNameLabel id="ACA49E79FFE7A73AFDAFFA81FD017632" box="[560,650,1305,1331]" pageId="11" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
exhibits sexual dimorphism in body size with adult females reaching 
<quantity id="421B52F5FFE7A73AFF08FAA5FF707656" box="[151,251,1341,1367]" metricMagnitude="-2" metricUnit="m" metricValue="3.97" pageId="11" pageNumber="510" unit="mm" value="39.7">39.7 mm</quantity>
SVL (MCNUP-H 163), adult males up to 
<quantity id="421B52F5FFE7A73AFD4BFAA5FCB37656" box="[724,824,1341,1367]" metricMagnitude="-2" metricUnit="m" metricValue="2.91" pageId="11" pageNumber="510" unit="mm" value="29.1">29.1 mm</quantity>
SVL (MCNUP-H 253). In addition, the texture of skin dorsally is roughened and is bedecked with scattered tubercles, unlike females, whose back is usually smooth. Dorsal color pattern also differs between sexes; females tend to have a more uniform cream background on the dorsal surface and light copper stripes arranged towards the sides of the body. In contrast, the males typically have a copper background on the dorsal with the dorsolateral folds with a strong reddish marking that extended from posterior edge of the upper eyelid to the sacrum.
</paragraph>
</subSubSection>
<subSubSection id="CDF9AC9BFFE7A73AFF58F98CFEFA744E" pageId="11" pageNumber="510" type="distribution">
<paragraph id="855CFF10FFE7A73AFF58F98CFEFA744E" blockId="11.[151,1437,152,1979]" pageId="11" pageNumber="510">
<emphasis id="B7972302FFE7A73AFF58F98CFDC1752E" bold="true" box="[199,586,1556,1583]" pageId="11" pageNumber="510">Distribution and natural history</emphasis>
. 
<taxonomicName id="42E38493FFE7A73AFDC6F98EFCB2752E" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[601,825,1557,1583]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE7A73AFDC6F98EFCB2752E" box="[601,825,1557,1583]" italics="true" pageId="11" pageNumber="510">Pristimantis ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE7A73AFCDEF98DFC10752E" bold="true" box="[833,923,1557,1583]" pageId="11" pageNumber="510">
<taxonomicNameLabel id="ACA49E79FFE7A73AFCDEF98DFC10752E" box="[833,923,1557,1583]" pageId="11" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from two localities: the settlements of San Antonio and La Vega, 
<quantity id="421B52F5FFE7A73AFDA3F9A1FD2B7552" box="[572,672,1593,1619]" metricMagnitude="4" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="11" pageNumber="510" unit="km" value="10.0" valueMax="11.0" valueMin="9.0">9–11 km</quantity>
from the road Toledo-Saravena, near the Margua river. These two localities correspond to tropical humid forest and riparian zones of the lowlands (400 and 
<specimenCount id="93E53499FFE7A73AFB3BF9C5FB737579" box="[1188,1272,1629,1656]" pageId="11" pageNumber="510" type="generic">
<quantity id="421B52F5FFE7A73AFB3BF9C5FB7F7579" box="[1188,1268,1629,1656]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="11" pageNumber="510" unit="m" value="700.0">700 m</quantity>
.
</specimenCount>
a.s.l.) from the westen flank of the Eastern Cordillera, in the Tamá National Natural Park and its buffer zone, department 
<collectingRegion id="472731F2FFE7A73AFAA4F919FE8275BE" country="Colombia" name="Norte de Santander" pageId="11" pageNumber="510">Norte de Santander</collectingRegion>
, 
<collectingCountry id="FDF4BF80FFE7A73AFE8BF93DFE0975BE" box="[276,386,1701,1727]" name="Colombia" pageId="11" pageNumber="510">Colombia</collectingCountry>
(
<figureCitation id="1DD8E395FFE7A73AFE0FF93DFE6475BE" box="[400,495,1701,1727]" captionStart="FIGURE 1" captionStartId="2.[151,250,1389,1414]" captionTargetBox="[160,1429,424,1358]" captionTargetId="figure@2.[153,1434,417,1364]" captionTargetPageId="2" captionText="FIGURE 1. Map of the geographical records of the new species. Norte de Santander, Colombia." figureDoi="http://doi.org/10.5281/zenodo.3708840" httpUri="https://zenodo.org/record/3708840/files/figure.png" pageId="11" pageNumber="510">Figure 1</figureCitation>
). The individuals of 
<taxonomicName id="42E38493FFE7A73AFD46F93EFCCF75BE" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[729,836,1701,1727]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="ardilae" status="sp. nov.">
<emphasis id="B7972302FFE7A73AFD46F93EFCCF75BE" box="[729,836,1701,1727]" italics="true" pageId="11" pageNumber="510">P. ardilae</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE7A73AFCD5F93DFC2F75BE" bold="true" box="[842,932,1701,1727]" pageId="11" pageNumber="510">
<taxonomicNameLabel id="ACA49E79FFE7A73AFCD5F93DFC2F75BE" box="[842,932,1701,1727]" pageId="11" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were found among the riparian zone perched at a height between 1 and 2.5 meters from the ground. The highest activity was recorded between 8 and 11 at night. The species was found sympatrically with other amphibians such as 
<taxonomicName id="42E38493FFE7A73AFC0EF976FB897406" authority="Acevedo &amp; Armesto &amp; Palma, 2020" authorityName="Acevedo &amp; Armesto &amp; Palma" authorityYear="2020" box="[913,1026,1773,1799]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="bowara" status="sp. nov.">
<emphasis id="B7972302FFE7A73AFC0EF976FB897406" box="[913,1026,1773,1799]" italics="true" pageId="11" pageNumber="510">P. bowara</emphasis>
</taxonomicName>
<emphasis id="B7972302FFE7A73AFB97F975FBEA7406" bold="true" box="[1032,1121,1773,1799]" pageId="11" pageNumber="510">
<taxonomicNameLabel id="ACA49E79FFE7A73AFB97F975FBEA7406" box="[1032,1121,1773,1799]" pageId="11" pageNumber="510" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="42E38493FFE7A73AFBF1F975FF44742D" authority="(Gunther)" baseAuthorityName="Gunther" class="Amphibia" family="Bufonidae" genus="Rhaebo" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="glaberrimus">
<emphasis id="B7972302FFE7A73AFBF1F975FAD87406" box="[1134,1363,1773,1799]" italics="true" pageId="11" pageNumber="510">Rhaebo glaberrimus</emphasis>
(Günther)
</taxonomicName>
, 
<taxonomicName id="42E38493FFE7A73AFF43F889FDBB742A" authority="(Dunn)" baseAuthorityName="Dunn" box="[220,560,1809,1835]" class="Amphibia" family="Caeciliidae" genus="Caecilia" kingdom="Animalia" order="Gymnophiona" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="subnigricans">
<emphasis id="B7972302FFE7A73AFF43F889FE5F742A" box="[220,468,1809,1835]" italics="true" pageId="11" pageNumber="510">Caecilia subnigricans</emphasis>
(Dunn)
</taxonomicName>
, 
<taxonomicName id="42E38493FFE7A73AFDA2F889FBB6742D" authority="(Lynch &amp; Duellman)" baseAuthorityName="Lynch &amp; Duellman" baseAuthorityYear="1973" box="[573,1085,1809,1836]" class="Amphibia" family="Centrolenidae" genus="Rulyrana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="flavopunctata">
<emphasis id="B7972302FFE7A73AFDA2F889FCCC742A" box="[573,839,1809,1835]" italics="true" pageId="11" pageNumber="510">Rulyrana flavopunctata</emphasis>
(Lynch &amp; Duellman)
</taxonomicName>
and 
<taxonomicName id="42E38493FFE7A73AFBE8F889FAB1742A" authorityName="Barrio-Amoros &amp; Santos" authorityYear="2009" box="[1143,1338,1809,1835]" class="Amphibia" family="Aromobatidae" genus="Allobates" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="species" species="algorei">
<emphasis id="B7972302FFE7A73AFBE8F889FAB1742A" box="[1143,1338,1809,1835]" italics="true" pageId="11" pageNumber="510">Allobates algorei</emphasis>
</taxonomicName>
(Barrio- Amorós &amp; Santos).
</paragraph>
</subSubSection>
<subSubSection id="CDF9AC9BFFE7A73AFF58F8C1FA1274BA" pageId="11" pageNumber="510" type="etymology">
<paragraph id="855CFF10FFE7A73AFF58F8C1FA1274BA" blockId="11.[151,1437,152,1979]" pageId="11" pageNumber="510">
<emphasis id="B7972302FFE7A73AFF58F8C1FEC37472" bold="true" box="[199,328,1881,1907]" pageId="11" pageNumber="510">Etymology</emphasis>
. The specific epithet refers to the professor and herpetologist María Cristina Ardila-Robayo (
<date id="F15DD9D0FFE7A73AFA1FF8C0FEB17496" pageId="11" pageNumber="510" value="1947-02-15">15 February 1947</date>
– 
<date id="F15DD9D0FFE7A73AFED6F8E5FDAE7496" box="[329,549,1917,1943]" pageId="11" pageNumber="510" value="2017-11-24">24 November 2017</date>
), who was a pioneer of herpetological studies in 
<collectingCountry id="FDF4BF80FFE7A73AFBD8F8E5FB3E7496" box="[1095,1205,1917,1943]" name="Colombia" pageId="11" pageNumber="510">Colombia</collectingCountry>
and contributed significantly to the knowledge of the diversity of Colombian amphibians, describing 15 species of 
<taxonomicName id="42E38493FFE7A73AFB57F83AFADA74BA" baseAuthorityName="Frost" baseAuthorityYear="2019" box="[1224,1361,1954,1979]" class="Amphibia" family="Strabomantidae" genus="Pristimantis" kingdom="Animalia" order="Anura" pageId="11" pageNumber="510" phylum="Chordata" rank="genus">
<emphasis id="B7972302FFE7A73AFB57F83AFADA74BA" box="[1224,1361,1954,1979]" italics="true" pageId="11" pageNumber="510">Pristimantis</emphasis>
</taxonomicName>
frogs.
</paragraph>
</subSubSection>
</treatment>
</document>