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<document id="FB0B5B4A6DB4FA118B8F3C6E6611B56C" ID-CLB-Dataset="82886" ID-DOI="10.1080/00222933.2021.2017499" ID-GBIF-Dataset="5cd01cfe-52bb-4092-97b2-804c3c7d145d" ID-ISSN="1464-5262" ID-Zenodo-Dep="6758313" ID-ZooBank="D766FC5E-CEC8-4184-BE90-C25350B90612" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656311158681" checkinUser="plazi" docAuthor="Costa, Leidiane O., Duarte, Paulo R. M., Iannuzzi, Luciana &amp; Grossi, Paschoal C." docDate="2022" docId="CF46879EFFBCFF88FF55706CFBC62848" docLanguage="en" docName="JNATHIST.56.1-4.191-225.pdf" docOrigin="Journal of Natural History 56 (1 - 4)" docSource="http://dx.doi.org/10.1080/00222933.2021.2017499" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Gibboryctes endroedii Duarte and Grossi 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="214" masterDocId="337FFFE6FFAAFF90FFF57271FF852A10" masterDocTitle="Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)" masterLastPageNumber="225" masterPageNumber="191" pageNumber="212" updateTime="1699352275692" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="586C7FCC44DFDF7B7FE67CD07BFE0B71" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B419C24E4520670B37A68BD128F52088">Taxonomic revision and notes on natural history of the enigmatic beetle genus Gibboryctes Endrödi (Coleoptera: Melolonthidae: Dynastinae)</mods:title>
</mods:titleInfo>
<mods:name id="D14AE821A8F92071159823555AFD6F2B" type="personal">
<mods:role id="5325DBD2349E39F7218BC5BA49F0723A">
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<mods:namePart id="316A44AD3C45FD1B820F7E86335EF4DE">Costa, Leidiane O.</mods:namePart>
</mods:name>
<mods:name id="AE2E459B4DE2FA0AD1C2C441C52EE07B" type="personal">
<mods:role id="D60654F08F53D64D305DD1DB473C2D02">
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<mods:namePart id="7CE7584303403207D0B4A5E49CB0DCEA">Duarte, Paulo R. M.</mods:namePart>
</mods:name>
<mods:name id="547D2D4106C91982D8B92961CF11882F" type="personal">
<mods:role id="FFFC9294A9E29817A4AA583F0A2B3C30">
<mods:roleTerm id="AE99A101E3C6DE75BD994938F97CF1A5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B10C12A086DF342D3252C2013B0EAEAC">Iannuzzi, Luciana</mods:namePart>
</mods:name>
<mods:name id="86646B8AE35AA1EDB4D0681E28CB51BA" type="personal">
<mods:role id="95336F47B956F77143477FC7B9D42F81">
<mods:roleTerm id="6A1A14E39071238C83E06E52DCB636EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B2FACCBEB2DB77FFFC9BAAE317C2E563">Grossi, Paschoal C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="14407478FA2F3DF55ADFA556F0F394F2">text</mods:typeOfResource>
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<mods:title id="DBF51B590C2C18F34BDC347A1CB0E713">Journal of Natural History</mods:title>
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<mods:part id="745A4D2B3B57F40A3C25BE96DE3AC2E3">
<mods:date id="537637AB47A88144FD222AB0FE70B578">2022</mods:date>
<mods:detail id="B9249E735577999369C0F36443367AF4" type="pubDate">
<mods:number id="8BEAFC4DDC59F765FB98979562CAD711">2022-05-17</mods:number>
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<mods:detail id="63AA4A83F3F15784F12D18572A243F92" type="volume">
<mods:number id="993C5DA836AAF93149D321330F201CAB">56</mods:number>
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<mods:detail id="AF0F68C3C3326AC1E9655F1A3270B972" type="issue">
<mods:number id="24F0B8EB6DCA1385982FDFE4A17E758E">1 - 4</mods:number>
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<mods:start id="9ABCFBFF9379FF6C26FA1B0F02EA7031">191</mods:start>
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<mods:classification id="F58DA090631C6ACB0B01D8642B211215">journal article</mods:classification>
<mods:identifier id="EE81C97D6F085DE3CF505AD270EDF51A" type="CLB-Dataset">82886</mods:identifier>
<mods:identifier id="695FA4E2E3767135B68724038294674A" type="DOI">10.1080/00222933.2021.2017499</mods:identifier>
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<treatment id="CF46879EFFBCFF88FF55706CFBC62848" ID-DOI="http://doi.org/10.5281/zenodo.6761928" ID-GBIF-Taxon="196400127" ID-Zenodo-Dep="6761928" LSID="urn:lsid:plazi:treatment:CF46879EFFBCFF88FF55706CFBC62848" httpUri="http://treatment.plazi.org/id/CF46879EFFBCFF88FF55706CFBC62848" lastPageId="24" lastPageNumber="214" pageId="22" pageNumber="212">
<subSubSection id="0FF56503FFBCFF86FF55706CFD6A2827" box="[160,751,541,567]" pageId="22" pageNumber="212" type="nomenclature">
<paragraph id="47503688FFBCFF86FF55706CFD6A2827" blockId="22.[160,1155,541,602]" box="[160,751,541,567]" pageId="22" pageNumber="212">
<taxonomicName id="80EF4D0BFFBCFF86FF55706CFD032827" authority="Duarte and Grossi" authorityName="Duarte and Grossi" authorityYear="2022" box="[160,646,541,567]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="212" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">Gibboryctes endroedii Duarte and Grossi</taxonomicName>
,
<taxonomicNameLabel id="6EA857E1FFBCFF86FD67706CFD6A2827" box="[658,751,541,567]" pageId="22" pageNumber="212" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBCFF86FD00706CFCD4284A" pageId="22" pageNumber="212" type="description">
<paragraph id="47503688FFBCFF86FD00706CFCD4284A" blockId="22.[160,1155,541,602]" pageId="22" pageNumber="212">
(
<figureCitation id="DFD42A0DFFBCFF86FD0B706CFCE92827" box="[766,876,541,567]" captionStart="Figure 3" captionStartId="6.[160,225,1740,1763]" captionTargetBox="[178,1132,148,1706]" captionTargetId="figure-3@6.[166,1149,142,1708]" captionTargetPageId="6" captionText="Figure 3. Gibboryctes endroedii sp. nov. (a,b) Habitus of holotype male in dorsal and lateral views, respectively; (c) habitus of female paratype in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6758321" httpUri="https://zenodo.org/record/6758321/files/figure.png" pageId="22" pageNumber="212">Figures 3</figureCitation>
;
<figureCitation id="DFD42A0DFFBCFF86FC8C706CFC032827" box="[889,902,541,567]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="22" pageNumber="212">6</figureCitation>
(c); 7(c); 8(c); 9(b); 10 (c,d); 11(c); 12(e,f); 13(c); 14(b))
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBCFF86FF5570E7FE6B2990" pageId="22" pageNumber="212" type="diagnosis">
<paragraph id="47503688FFBCFF86FF5570E7FE9128A0" blockId="22.[160,1155,662,896]" box="[160,276,662,688]" pageId="22" pageNumber="212">
<emphasis id="759BEA9AFFBCFF86FF5570E7FE9128A0" bold="true" box="[160,276,662,688]" italics="true" pageId="22" pageNumber="212">Diagnosis</emphasis>
</paragraph>
<paragraph id="47503688FFBCFF86FF5570CBFE6B2990" blockId="22.[160,1155,662,896]" pageId="22" pageNumber="212">
<taxonomicName id="80EF4D0BFFBCFF86FF5570CBFE0F28C3" authorityName="Duarte and Grossi" authorityYear="2022" box="[160,394,698,723]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="212" phylum="Arthropoda" rank="species" species="endroedii">
<emphasis id="759BEA9AFFBCFF86FF5570CBFE0F28C3" box="[160,394,698,723]" italics="true" pageId="22" pageNumber="212">Gibboryctes endroedii</emphasis>
</taxonomicName>
differs from other
<taxonomicName id="80EF4D0BFFBCFF86FDAE70CBFCB428C3" box="[603,817,698,723]" pageId="22" pageNumber="212">
<emphasis id="759BEA9AFFBCFF86FDAE70CBFD5F28C3" box="[603,730,698,723]" italics="true" pageId="22" pageNumber="212">Gibboryctes</emphasis>
species
</taxonomicName>
by the following combination of characters: body colouration reddish brown or dark reddish brown (
<figureCitation id="DFD42A0DFFBCFF86FC3770ADFB9B28E5" box="[962,1054,732,757]" captionStart="Figure 3" captionStartId="6.[160,225,1740,1763]" captionTargetBox="[178,1132,148,1706]" captionTargetId="figure-3@6.[166,1149,142,1708]" captionTargetPageId="6" captionText="Figure 3. Gibboryctes endroedii sp. nov. (a,b) Habitus of holotype male in dorsal and lateral views, respectively; (c) habitus of female paratype in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6758321" httpUri="https://zenodo.org/record/6758321/files/figure.png" pageId="22" pageNumber="212">Figure 3</figureCitation>
); labrum subtrapezoidal with broadly rounded apex (
<figureCitation id="DFD42A0DFFBCFF86FD6C708EFD722908" box="[665,759,767,792]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="22" pageNumber="212">Figure 6</figureCitation>
(c)); maxillae with lobed galea and stipe with a triangular lateral lobe (
<figureCitation id="DFD42A0DFFBCFF86FDD37153FD07292B" box="[550,642,802,827]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="22" pageNumber="212">Figure 8</figureCitation>
(c)); pronotal sides almost completely covered with large and coalescent punctures (
<figureCitation id="DFD42A0DFFBCFF86FDCA7134FD2D294E" box="[575,680,837,862]" captionStart="Figure 12" captionStartId="15.[160,225,1189,1212]" captionTargetBox="[167,1149,146,1154]" captionTargetId="figure-100@15.[166,1149,142,1157]" captionTargetPageId="15" captionText="Figure 12. Females of Gibboryctes species with detail of head, pronotum and prothoracic legs in lateral view and left elytra in dorsal view, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes ebeninus sp. nov.; (e,f) Gibboryctes endroedii sp. nov. Jsi, juxtasutural interstria. Arrow indicates coalescent punctures in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758343" httpUri="https://zenodo.org/record/6758343/files/figure.png" pageId="22" pageNumber="212">Figure 12</figureCitation>
(e)); elytral interstriae densely covered with large punctures (
<figureCitation id="DFD42A0DFFBCFF86FEAA7116FE4E2990" box="[351,459,871,896]" captionStart="Figure 12" captionStartId="15.[160,225,1189,1212]" captionTargetBox="[167,1149,146,1154]" captionTargetId="figure-100@15.[166,1149,142,1157]" captionTargetPageId="15" captionText="Figure 12. Females of Gibboryctes species with detail of head, pronotum and prothoracic legs in lateral view and left elytra in dorsal view, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes ebeninus sp. nov.; (e,f) Gibboryctes endroedii sp. nov. Jsi, juxtasutural interstria. Arrow indicates coalescent punctures in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758343" httpUri="https://zenodo.org/record/6758343/files/figure.png" pageId="22" pageNumber="212">Figure 12</figureCitation>
(f)).
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBCFF86FF5571DDFEC129D6" box="[160,324,940,966]" pageId="22" pageNumber="212" type="description">
<paragraph id="47503688FFBCFF86FF5571DDFEC129D6" blockId="22.[160,1157,940,1278]" box="[160,324,940,966]" pageId="22" pageNumber="212">
<emphasis id="759BEA9AFFBCFF86FF5571DDFEC129D6" bold="true" box="[160,324,940,966]" italics="true" pageId="22" pageNumber="212">
<typeStatus id="9854882AFFBCFF86FF5571DDFF5D29D6" box="[160,216,940,966]" pageId="22" pageNumber="212">Type</typeStatus>
material
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBCFF86FF5571BEFE132F53" pageId="22" pageNumber="212" type="materials_examined">
<paragraph id="47503688FFBCFF86FF5571BEFE292EEE" blockId="22.[160,1157,940,1278]" pageId="22" pageNumber="212">
<materialsCitation id="F7873CD5FFBCFF86FF5571BEFC552E3E" ID-GBIF-Occurrence="3819562306" collectingDate="2012-11-20" collectionCode="CERPE" country="Brazil" elevation="1060" location="Grossi &amp; Parizotto" municipality="Lavras / Poco Bonito" pageId="22" pageNumber="212" specimenCount="2" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="holotype">
<typeStatus id="9854882AFFBCFF86FF5571BEFE8229F8" box="[160,263,975,1000]" pageId="22" pageNumber="212" type="holotype">Holotype</typeStatus>
<specimenCount id="51E9FD01FFBCFF86FEF871BEFEC129F8" box="[269,324,975,1000]" pageId="22" pageNumber="212" type="male">male</specimenCount>
dissected, labelled: (a) ‘
<collectingCountry id="3FF87618FFBCFF86FDB271BEFD0D29F8" box="[583,648,975,1000]" name="Brazil" pageId="22" pageNumber="212">Brasil</collectingCountry>
,
<collectingRegion id="852BF86AFFBCFF86FD6571BEFCA529F8" box="[656,800,975,1000]" country="Brazil" name="Minas Gerais" pageId="22" pageNumber="212">Minas Gerais</collectingRegion>
,
<collectingMunicipality id="A734ACF2FFBCFF86FCDD71BEFB8429F8" box="[808,1025,975,1000]" pageId="22" pageNumber="212">Lavras/Poço Bonito</collectingMunicipality>
,
<date id="33511048FFBCFF86FBFF71BEFBFF29F8" box="[1034,1146,975,1000]" pageId="22" pageNumber="212" value="2012-11-20">
<collectingDate id="2315E9A0FFBCFF86FBFF71BEFBFF29F8" box="[1034,1146,975,1000]" pageId="22" pageNumber="212" value="2012-11-20">20.xi.2012</collectingDate>
</date>
/
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<elevation id="CCC2D1BBFFBCFF86FF557183FF7D2E1B" box="[160,248,1010,1035]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="22" pageNumber="212" unit="m" value="1060.0">1060 m</elevation>
</quantity>
,
<location id="42306053FFBCFF86FEF07183FE4A2E1B" LSID="urn:lsid:plazi:treatment:CF46879EFFBCFF88FF55706CFBC62848:42306053FFBCFF86FEF07183FE4A2E1B" box="[261,463,1010,1035]" country="Brazil" municipality="Lavras / Poco Bonito" name="Grossi &amp; Parizotto" pageId="22" pageNumber="212" stateProvince="Minas Gerais">
<taxonomicName id="80EF4D0BFFBCFF86FEF07183FECC2E1B" authority="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi, 2022" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" box="[261,329,1010,1035]" genus="Grossi" pageId="22" pageNumber="222" rank="genus" status="gen. nov.">Grossi</taxonomicName>
&amp; Parizotto
</location>
/criação ninhos de cupim’ [white label]; (b) ‘
<taxonomicName id="80EF4D0BFFBCFF86FC3E7183FF582E3E" authority="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi, 2022" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="212" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
<emphasis id="759BEA9AFFBCFF86FC3E7183FF582E3E" italics="true" pageId="22" pageNumber="212">
<location id="42306053FFBCFF86FC3E7183FBCF2E1B" LSID="urn:lsid:plazi:treatment:CF46879EFFBCFF88FF55706CFBC62848:42306053FFBCFF86FC3E7183FBCF2E1B" box="[971,1098,1010,1035]" country="Brazil" municipality="Lavras / Poco Bonito" name="Gibboryctes" pageId="22" pageNumber="212" stateProvince="Minas Gerais">Gibboryctes</location>
endroedii
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EA857E1FFBCFF86FF107664FEB22E3E" box="[229,311,1045,1070]" pageId="22" pageNumber="212" rank="species">
<specimenCount id="51E9FD01FFBCFF86FF107664FE802E3E" box="[229,261,1045,1070]" pageId="22" pageNumber="212" type="generic">sp.</specimenCount>
nov.
</taxonomicNameLabel>
/
<typeStatus id="9854882AFFBCFF86FEC87664FE402E3E" box="[317,453,1045,1070]" pageId="22" pageNumber="212" type="holotype">HOLOTYPE</typeStatus>
/
<location id="42306053FFBCFF86FE387664FDFC2E3E" LSID="urn:lsid:plazi:treatment:CF46879EFFBCFF88FF55706CFBC62848:42306053FFBCFF86FE387664FDFC2E3E" box="[461,633,1045,1070]" country="Brazil" municipality="Lavras / Poco Bonito" name="Duarte &amp; Grossi" pageId="22" pageNumber="212" stateProvince="Minas Gerais">
Duarte &amp;
<taxonomicName id="80EF4D0BFFBCFF86FDC07664FDFC2E3E" authority="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi, 2022" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" box="[565,633,1045,1070]" genus="Grossi" pageId="22" pageNumber="222" rank="genus" status="gen. nov.">Grossi</taxonomicName>
</location>
det. 2021’ [red label] (
<collectionCode id="21FEAE4DFFBCFF86FC8C7664FC422E3E" box="[889,967,1045,1070]" collectionName="CERPE" pageId="22" pageNumber="212">CERPE</collectionCode>
).
</materialsCitation>
<materialsCitation id="F7873CD5FFBCFF86FC227664FCEA2E40" ID-GBIF-Occurrence="3819562309" collectionCode="CERPE" location="Two" pageId="22" pageNumber="212" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<location id="42306053FFBCFF86FC227664FB822E3E" LSID="urn:lsid:plazi:treatment:CF46879EFFBCFF88FF55706CFBC62848:42306053FFBCFF86FC227664FB822E3E" box="[983,1031,1045,1070]" name="Two" pageId="22" pageNumber="212">Two</location>
<specimenCount id="51E9FD01FFBCFF86FBFA7664FBD52E3E" box="[1039,1104,1045,1070]" pageId="22" pageNumber="212" type="male">males</specimenCount>
and
<specimenCount id="51E9FD01FFBCFF86FF557646FEB52E40" box="[160,304,1079,1104]" pageId="22" pageNumber="212" type="female">two females</specimenCount>
<typeStatus id="9854882AFFBCFF86FECE7646FE2E2E40" box="[315,427,1079,1104]" pageId="22" pageNumber="212" type="paratype">paratypes</typeStatus>
with same data as
<typeStatus id="9854882AFFBCFF86FD6A7646FC802E40" box="[671,773,1079,1104]" pageId="22" pageNumber="212" type="holotype">holotype</typeStatus>
(
<collectionCode id="21FEAE4DFFBCFF86FCE37646FCE02E40" box="[790,869,1079,1104]" collectionName="CERPE" pageId="22" pageNumber="212">CERPE</collectionCode>
).
</materialsCitation>
<materialsCitation id="F7873CD5FFBCFF86FC8C7646FE242E86" ID-GBIF-Occurrence="3819562320" collectingDate="2008-03" collectionCode="CERPE" country="Brazil" location="Two" municipality="Ingai" pageId="22" pageNumber="212" specimenCount="1" specimenCount-female="1" stateProvince="Minas Gerais" typeStatus="paratype">
<location id="42306053FFBCFF86FC8C7646FC2C2E40" LSID="urn:lsid:plazi:treatment:CF46879EFFBCFF88FF55706CFBC62848:42306053FFBCFF86FC8C7646FC2C2E40" box="[889,937,1079,1104]" country="Brazil" municipality="Ingai" name="Two" pageId="22" pageNumber="212" stateProvince="Minas Gerais">Two</location>
<specimenCount id="51E9FD01FFBCFF86FC407646FB872E40" box="[949,1026,1079,1104]" pageId="22" pageNumber="212" type="female">female</specimenCount>
<typeStatus id="9854882AFFBCFF86FBFB7646FB042E40" box="[1038,1153,1079,1104]" pageId="22" pageNumber="212" type="paratype">paratypes</typeStatus>
, labelled: (a) ‘
<collectingCountry id="3FF87618FFBCFF86FEC7762BFEF62E63" box="[306,371,1114,1139]" name="Brazil" pageId="22" pageNumber="212">Brasil</collectingCountry>
,
<collectingRegion id="852BF86AFFBCFF86FE75762BFD912E63" box="[384,532,1114,1139]" country="Brazil" name="Minas Gerais" pageId="22" pageNumber="212">Minas Gerais</collectingRegion>
,
<collectingMunicipality id="A734ACF2FFBCFF86FDD4762BFDDE2E63" box="[545,603,1114,1139]" pageId="22" pageNumber="212">Ingaí</collectingMunicipality>
/
<date id="33511048FFBCFF86FD91762BFD362E63" box="[612,691,1114,1139]" pageId="22" pageNumber="212" value="2008-03">
<collectingDate id="2315E9A0FFBCFF86FD91762BFD362E63" box="[612,691,1114,1139]" pageId="22" pageNumber="212" value="2008-03">iii.2008</collectingDate>
</date>
, termiteiro chão/Vaz-de-Mello, ab. larva’ [white label]; (
<collectionCode id="21FEAE4DFFBCFF86FEBC760CFE1D2E86" box="[329,408,1149,1174]" collectionName="CERPE" pageId="22" pageNumber="212">CERPE</collectionCode>
).
</materialsCitation>
<materialsCitation id="F7873CD5FFBCFF86FE5E760CFE292EEE" ID-GBIF-Occurrence="3819562303" collectingDate="1947-01" collectingDateMax="2007-11" collectingDateMin="1947-01" collectionCode="CEMT, CERPE" country="Brazil" location="Mello" municipality="Ingai" pageId="22" pageNumber="212" specimenCount="4" specimenCount-female="1" specimenCount-male="1" stateProvince="Minas Gerais" typeStatus="paratype">
One
<specimenCount id="51E9FD01FFBCFF86FE13760CFDB62E86" box="[486,563,1149,1174]" pageId="22" pageNumber="212" type="female">female</specimenCount>
<typeStatus id="9854882AFFBCFF86FDCA760CFD222E86" box="[575,679,1149,1174]" pageId="22" pageNumber="212" type="paratype">paratype</typeStatus>
, labelled: (a) ‘
<collectingCountry id="3FF87618FFBCFF86FCB9760CFC0B2E86" box="[844,910,1149,1174]" name="Brazil" pageId="22" pageNumber="212">Brasil</collectingCountry>
,
<collectingRegion id="852BF86AFFBCFF86FC69760CFBB12E86" box="[924,1076,1149,1174]" country="Brazil" name="Minas Gerais" pageId="22" pageNumber="212">Minas Gerais</collectingRegion>
,
<collectingMunicipality id="A734ACF2FFBCFF86FBB7760CFBFE2E86" box="[1090,1147,1149,1174]" pageId="22" pageNumber="212">Ingaí</collectingMunicipality>
/
<date id="33511048FFBCFF86FF5576EEFF772EA8" box="[160,242,1183,1208]" pageId="22" pageNumber="212" value="2007-11">
<collectingDate id="2315E9A0FFBCFF86FF5576EEFF772EA8" box="[160,242,1183,1208]" pageId="22" pageNumber="212" value="2007-11">xi.2007</collectingDate>
</date>
, breed, Vaz-de -
<location id="42306053FFBCFF86FE5276EEFE632EA8" LSID="urn:lsid:plazi:treatment:CF46879EFFBCFF88FF55706CFBC62848:42306053FFBCFF86FE5276EEFE632EA8" box="[423,486,1183,1208]" country="Brazil" municipality="Ingai" name="Mello" pageId="22" pageNumber="212" stateProvince="Minas Gerais">Mello</location>
leg’. [white label]; (
<collectionCode id="21FEAE4DFFBCFF86FD3676EEFC8C2EA8" box="[707,777,1183,1208]" pageId="22" pageNumber="212">CEMT</collectionCode>
).
<location id="42306053FFBCFF86FCEF76EEFCCC2EA8" LSID="urn:lsid:plazi:treatment:CF46879EFFBCFF88FF55706CFBC62848:42306053FFBCFF86FCEF76EEFCCC2EA8" box="[794,841,1183,1208]" country="Brazil" municipality="Ingai" name="One" pageId="22" pageNumber="212" stateProvince="Minas Gerais">One</location>
<specimenCount id="51E9FD01FFBCFF86FCA476EEFC0D2EA8" box="[849,904,1183,1208]" pageId="22" pageNumber="212" type="male">male</specimenCount>
<typeStatus id="9854882AFFBCFF86FC6576EEFC732EA8" box="[912,1014,1183,1208]" pageId="22" pageNumber="212" type="paratype">paratype</typeStatus>
, labelled: (a) ‘
<collectingCountry id="3FF87618FFBCFF86FF5676B3FF602ECB" box="[163,229,1218,1243]" name="Brazil" pageId="22" pageNumber="212">Brasil</collectingCountry>
,
<collectingRegion id="852BF86AFFBCFF86FF0676B3FE242ECB" box="[243,417,1218,1243]" country="Brazil" name="Rio de Janeiro" pageId="22" pageNumber="212">Rio de Janeiro</collectingRegion>
/
<location id="42306053FFBCFF86FE5C76B3FE702ECB" LSID="urn:lsid:plazi:treatment:CF46879EFFBCFF88FF55706CFBC62848:42306053FFBCFF86FE5C76B3FE702ECB" box="[425,501,1218,1243]" country="Brazil" municipality="Ingai" name="Itatiaia" pageId="22" pageNumber="212" stateProvince="Minas Gerais">Itatiaia</location>
,
<date id="33511048FFBCFF86FDF176B3FDCF2ECB" box="[516,586,1218,1243]" pageId="22" pageNumber="212" value="1947-01">
<collectingDate id="2315E9A0FFBCFF86FDF176B3FDCF2ECB" box="[516,586,1218,1243]" pageId="22" pageNumber="212" value="1947-01">I.1947</collectingDate>
</date>
, A. Englir’ [white label]; (
<collectionCode id="21FEAE4DFFBCFF86FC8076B3FC412ECB" box="[885,964,1218,1243]" collectionName="CERPE" pageId="22" pageNumber="212">CERPE</collectionCode>
).
<typeStatus id="9854882AFFBCFF86FC2276B3FBC02ECB" box="[983,1093,1218,1243]" pageId="22" pageNumber="212" type="paratype">Paratypes</typeStatus>
with a yellow
<typeStatus id="9854882AFFBCFF86FEF07694FEEC2EEE" box="[261,361,1253,1278]" pageId="22" pageNumber="212" type="paratype">paratype</typeStatus>
label.
</materialsCitation>
</paragraph>
<paragraph id="47503688FFBCFF86FF557758FE132F53" blockId="22.[160,1156,1321,1728]" box="[160,406,1321,1347]" pageId="22" pageNumber="212">
<emphasis id="759BEA9AFFBCFF86FF557758FE132F53" bold="true" box="[160,406,1321,1347]" italics="true" pageId="22" pageNumber="212">
<typeStatus id="9854882AFFBCFF86FF557758FE8E2F53" box="[160,267,1321,1347]" pageId="22" pageNumber="212" type="holotype">Holotype</typeStatus>
description
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBCFF87FF55773DFE8B2EFA" lastPageId="23" lastPageNumber="213" pageId="22" pageNumber="212" type="description">
<paragraph id="47503688FFBCFF87FF55773DFEBF2E84" blockId="22.[160,1156,1321,1728]" lastBlockId="23.[160,1156,142,1172]" lastPageId="23" lastPageNumber="213" pageId="22" pageNumber="212">
<emphasis id="759BEA9AFFBCFF86FF55773DFE3A2F76" bold="true" box="[160,447,1356,1382]" pageId="22" pageNumber="212">
Male
<emphasis id="759BEA9AFFBCFF86FF11773DFED82F76" bold="true" box="[228,349,1356,1382]" italics="true" pageId="22" pageNumber="212">
(
<figureCitation id="DFD42A0DFFBCFF86FF19773DFECA2F76" box="[236,335,1356,1382]" captionStart="Figure 3" captionStartId="6.[160,225,1740,1763]" captionTargetBox="[178,1132,148,1706]" captionTargetId="figure-3@6.[166,1149,142,1708]" captionTargetPageId="6" captionText="Figure 3. Gibboryctes endroedii sp. nov. (a,b) Habitus of holotype male in dorsal and lateral views, respectively; (c) habitus of female paratype in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6758321" httpUri="https://zenodo.org/record/6758321/files/figure.png" pageId="22" pageNumber="212">Figure 3</figureCitation>
).
</emphasis>
Length
</emphasis>
:
<quantity id="80179B6DFFBCFF86FE3B773CFDB72F76" box="[462,562,1357,1382]" metricMagnitude="-2" metricUnit="m" metricValue="2.46" pageId="22" pageNumber="212" unit="mm" value="24.6">24.6 mm</quantity>
.
<emphasis id="759BEA9AFFBCFF86FDCB773CFD002F76" bold="true" box="[574,645,1357,1382]" pageId="22" pageNumber="212">Width</emphasis>
:
<quantity id="80179B6DFFBCFF86FD66773CFD7D2F76" box="[659,760,1357,1382]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="22" pageNumber="212" unit="mm" value="11.7">11.7 mm</quantity>
.
<emphasis id="759BEA9AFFBCFF86FCF1773CFCD72F76" bold="true" box="[772,850,1357,1382]" pageId="22" pageNumber="212">Colour</emphasis>
: Nearly completely reddish brown; protibial teeth black.
<emphasis id="759BEA9AFFBCFF86FE28771EFD9F2F98" bold="true" box="[477,538,1391,1416]" pageId="22" pageNumber="212">Head</emphasis>
: Clypeus triangular, transverse, 2 times wider than long, acuminate anteriorly, weakly narrowed laterally; lateral margin slightly raised; surface rugose, densely setose on sides separated by a glabrous middle area. Frontoclypeal suture with a transverse, flattened, short, subtrapezoidal tubercle. Frontal surface glabrous, transversely rugopunctate; punctures large, coalescent, C-shaped; interocular width equals 4.1 transverse eye diameter. Ocular canthus transverse, subtriangular, glabrous, with a small notch at lateroposterior outer corner.
<emphasis id="759BEA9AFFBCFF86FD9B744EFD7E2C48" bold="true" box="[622,763,1599,1624]" pageId="22" pageNumber="212">Mouthparts</emphasis>
: Labrum subtrapezoidal, broadly rounded on apical margin (
<figureCitation id="DFD42A0DFFBCFF86FE2C7413FDB32C6B" box="[473,566,1634,1659]" captionStart="Figure 6" captionStartId="9.[160,225,1090,1113]" captionTargetBox="[166,1145,146,1050]" captionTargetId="figure-188@9.[166,1149,142,1058]" captionTargetPageId="9" captionText="Figure 6. Labrum in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758331" httpUri="https://zenodo.org/record/6758331/files/figure.png" pageId="22" pageNumber="212">Figure 6</figureCitation>
(c)). Mandibles with 2 teeth on outer margin; apical tooth subtriangular; basal tooth subrectangular, rounded apically, clearly larger in size compared to apical tooth (
<figureCitation id="DFD42A0DFFBCFF86FE2F74D6FDBC2CD0" box="[474,569,1703,1728]" captionStart="Figure 7" captionStartId="10.[160,225,1156,1179]" captionTargetBox="[169,1149,148,1090]" captionTargetId="figure-156@10.[166,1149,142,1125]" captionTargetPageId="10" captionText="Figure 7. Right mandible in dorsal view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicate the apical tooth in (a–e), striate molar in (d)." figureDoi="http://doi.org/10.5281/zenodo.6758333" httpUri="https://zenodo.org/record/6758333/files/figure.png" pageId="22" pageNumber="212">Figure 7</figureCitation>
(c)). Maxillary galea lobed, widely rounded at apex (
<figureCitation id="DFD42A0DFFBDFF87FF5272FFFE852AB7" box="[167,256,142,167]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="23" pageNumber="213">Figure 8</figureCitation>
(c)); inner margin with 3 subapical teeth increasing in size towards apex; apical and basal teeth triangular; medial tooth lobed; inner margin rounded just below basal tooth; stipe expanded laterally in triangular shape (
<figureCitation id="DFD42A0DFFBDFF87FD6172A5FD752AFD" box="[660,752,212,237]" captionStart="Figure 8" captionStartId="11.[160,225,1161,1184]" captionTargetBox="[167,1127,143,1126]" captionTargetId="figure-61@11.[166,1149,142,1129]" captionTargetPageId="11" captionText="Figure 8. Right maxilla in ventral view. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedii sp. nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Red arrows indicating triangular galea in (a), elongate maxillary palpomere II in (b), lobed inner margin of galea and stipe with triangular lobe in (c), pointed and triangular teeth on galea in (d), scythe-shape d teeth and rounded inner margin of galea in (e)." figureDoi="http://doi.org/10.5281/zenodo.6758335" httpUri="https://zenodo.org/record/6758335/files/figure.png" pageId="23" pageNumber="213">Figure 8</figureCitation>
(c)). Maxillary palpomere II 1.6 times longer than width at middle. Labium subtriangular, slightly rounded laterally, becoming narrow towards apex; surface densely punctate; sides densely covered with long bristles; disc with scarce bristles, shorter than lateral compared to lateral bristles.
<emphasis id="759BEA9AFFBDFF87FC42734DFB8C2B45" bold="true" box="[951,1033,316,341]" pageId="23" pageNumber="213">Thorax</emphasis>
: Pronotum rounded laterally in dorsal view, weakly convex in lateral view, longitudinal middle area slightly concave; anterior area with a small tubercle separated from anterior border by a deep groove; posterior pronotal border incomplete; pronotal surface almost entirely covered with ocellate punctures (
<figureCitation id="DFD42A0DFFBDFF87FDE873B7FDFE2BCF" box="[541,635,454,479]" captionStart="Figure 9" captionStartId="12.[160,225,1146,1169]" captionTargetBox="[184,1146,142,1069]" captionTargetId="figure-59@12.[166,1149,142,1114]" captionTargetPageId="12" captionText="Figure 9. Detail of head, pronotum and prothoracic legs of males in dorsal view: (a) Gibboryctes szelenyii (Endrödi); (b) Gibboryctes endroedii sp. nov.; (c) Heterogomphus gracilicornis Prell; (d) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758337" httpUri="https://zenodo.org/record/6758337/files/figure.png" pageId="23" pageNumber="213">Figure 9</figureCitation>
(b)); punctures on sides deep, large, predominantly coalescent; longitudinal pronotal mid-line with a row of coalescent punctures; posterior areas on each side with shallow punctures scattered about 2 diameters of punctures. Scutellar plate subtriangular, scarcely punctate; punctures small, confined to anterior area. Elytra with 9 well-marked striae (1 sutural, 4 discal, 4 lateral); striae covered with a row of ocellate, deep, oval punctures; discal striae with contiguous punctures, gradually becoming smaller and sparser (about 1 diameter of punctures) towards posterior area; lateral striae with punctures scattered about 2 diameters of punctures, smaller compared to the discal striae; interstriae with punctures irregularly scattered, with mixed large and fine punctures.
<emphasis id="759BEA9AFFBDFF87FE4D708FFE6B2907" bold="true" box="[440,494,766,791]" pageId="23" pageNumber="213">Legs</emphasis>
: Mesotibial outer carinae with 11 stout spinules like setae (3 on basal carina, 8 on medial carina). Metatibia with 16 stout spinules like setae (6 on basal carina, 10 on medial carina).
<emphasis id="759BEA9AFFBDFF87FE107132FDDD294C" bold="true" box="[485,600,835,860]" pageId="23" pageNumber="213">Abdomen</emphasis>
: Tergite VIII with glabrous surface, strongly rugopunctate on lateral corners, densely punctate on discal area; punctures large, deep, from coalescent to contiguous on sides, becoming scattered about 1 diameter of punctures on disc. Ventrites I–VI rugopunctate on sides, finely punctate on discal area; ventrites II–V with an incomplete row of setigerous punctures near to posterior margin; ventrite VI glabrous, emarginate posteriorly at middle.
<emphasis id="759BEA9AFFBDFF87FDE37180FD0F2E1A" bold="true" box="[534,650,1009,1034]" pageId="23" pageNumber="213">Aedeagus</emphasis>
: Parameres, in dorsal view, wide at basal half, becoming convergent towards a narrow apical half, covered with scarce bristles on inner edge of apex (
<figureCitation id="DFD42A0DFFBDFF87FEBA7647FE3E2E5F" box="[335,443,1078,1103]" captionStart="Figure 10" captionStartId="13.[160,226,1164,1187]" captionTargetBox="[169,1145,145,1132]" captionTargetId="figure-89@13.[166,1149,142,1133]" captionTargetPageId="13" captionText="Figure 10. Aedeagus with parameres in dorsal and lateral views, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes endroedii sp. nov.; (e,f) Heterogomphus gracilicornis Prell; (g,h) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758339" httpUri="https://zenodo.org/record/6758339/files/figure.png" pageId="23" pageNumber="213">Figure 10</figureCitation>
(c)). Parameres, in lateral view, arched dorsally, ventrally with a longitudinal carina at middle, apex rounded, strongly constricted dorsoventrally (
<figureCitation id="DFD42A0DFFBDFF87FF52760AFE942E84" box="[167,273,1147,1172]" captionStart="Figure 10" captionStartId="13.[160,226,1164,1187]" captionTargetBox="[169,1145,145,1132]" captionTargetId="figure-89@13.[166,1149,142,1133]" captionTargetPageId="13" captionText="Figure 10. Aedeagus with parameres in dorsal and lateral views, respectively. (a,b) Gibboryctes szelenyii Endrödi; (c,d) Gibboryctes endroedii sp. nov.; (e,f) Heterogomphus gracilicornis Prell; (g,h) Tetragus waldenfelsi (Endrödi)." figureDoi="http://doi.org/10.5281/zenodo.6758339" httpUri="https://zenodo.org/record/6758339/files/figure.png" pageId="23" pageNumber="213">Figure 10</figureCitation>
(d)).
</paragraph>
<paragraph id="47503688FFBDFF87FF5576A1FE8B2EFA" blockId="23.[160,1156,1232,1535]" box="[160,270,1232,1258]" pageId="23" pageNumber="213">
<emphasis id="759BEA9AFFBDFF87FF5576A1FE8B2EFA" bold="true" box="[160,270,1232,1258]" italics="true" pageId="23" pageNumber="213">Variation</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBDFF87FF557685FC192FEF" pageId="23" pageNumber="213" type="materials_examined">
<paragraph id="47503688FFBDFF87FF557685FC192FEF" blockId="23.[160,1156,1232,1535]" pageId="23" pageNumber="213">
Male
<typeStatus id="9854882AFFBDFF87FF287685FED52F1D" box="[221,336,1268,1293]" pageId="23" pageNumber="213" type="paratype">paratypes</typeStatus>
. Length:
<quantity id="80179B6DFFBDFF87FE407685FDD32F1D" box="[437,598,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="2.525" metricValueMax="2.94" metricValueMin="2.11" pageId="23" pageNumber="213" unit="mm" value="25.25" valueMax="29.4" valueMin="21.1">21.1–29.4 mm</quantity>
; Width: 11.0–
<quantity id="80179B6DFFBDFF87FD1B7685FCD42F1D" box="[750,849,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="23" pageNumber="213" unit="mm" value="11.1">11.1 mm</quantity>
. As for
<typeStatus id="9854882AFFBDFF87FC557685FB832F1D" box="[928,1030,1268,1293]" pageId="23" pageNumber="213" type="holotype">holotype</typeStatus>
except the dark reddish brown colour; frontoclypeal tubercle bilobed; mandibles with lobed apical teeth; maxillae with 4 teeth at inner margin of galea; parameres widely divergent at basal half in dorsal view and with straight ventral surface in lateral view. Female
<typeStatus id="9854882AFFBDFF87FF55770FFE962F87" box="[160,275,1406,1431]" pageId="23" pageNumber="213" type="paratype">paratypes</typeStatus>
. Length: 21.0–
<quantity id="80179B6DFFBDFF87FE49770FFDB02F87" box="[444,565,1406,1431]" metricMagnitude="-2" metricUnit="m" metricValue="2.337" pageId="23" pageNumber="213" unit="mm" value="23.37">23.37 mm</quantity>
; Width:
<quantity id="80179B6DFFBDFF87FD62770FFCB82F87" box="[663,829,1406,1431]" metricMagnitude="-2" metricUnit="m" metricValue="1.11" metricValueMax="1.13" metricValueMin="1.09" pageId="23" pageNumber="213" unit="mm" value="11.1" valueMax="11.3" valueMin="10.9">10.9–11.3 mm</quantity>
. As
<typeStatus id="9854882AFFBDFF87FC84770FFC522F87" box="[881,983,1406,1431]" pageId="23" pageNumber="213" type="holotype">holotype</typeStatus>
except by the clypeus slightly rounded anteriorly (
<figureCitation id="DFD42A0DFFBDFF87FDBA77D0FD382FAA" box="[591,701,1441,1466]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="23" pageNumber="213">Figure 11</figureCitation>
(c)); pronotum densely punctate compared to males (
<figureCitation id="DFD42A0DFFBDFF87FE9277B5FE642FCD" box="[359,481,1476,1501]" captionStart="Figure 11" captionStartId="14.[160,225,1441,1464]" captionTargetBox="[168,1146,144,1407]" captionTargetId="figure-42@14.[166,1149,142,1410]" captionTargetPageId="14" captionText="Figure 11. Detail of head, pronotum and prothoracic legs of females in dorsal view: (a) Gibboryctes szelenyii Endrödi;(b) Gibboryctes ebeninus sp. nov.; (c) Gibboryctes endroedi sp.nov.; (d) Heterogomphus gracilicornis Prell; (e) Tetragus waldenfelsi (Endrödi). Arrow indicates punctate frons in (b)." figureDoi="http://doi.org/10.5281/zenodo.6758341" httpUri="https://zenodo.org/record/6758341/files/figure.png" pageId="23" pageNumber="213">Figures 11</figureCitation>
(c) and 12(e)); tergite VIII slightly convex in lateral view; ventrite VI parabolic, lacking posterior emargination (
<figureCitation id="DFD42A0DFFBDFF87FCF37797FCF72FEF" box="[774,882,1510,1535]" captionStart="Figure 13" captionStartId="16.[160,225,1058,1081]" captionTargetBox="[166,1146,145,1023]" captionTargetId="figure-155@16.[166,1149,142,1026]" captionTargetPageId="16" captionText="Figure 13. Detail of ventrites I–VI of females. (a) Gibboryctes szelenyii Endrödi; (b) Gibboryctes ebeninus sp. nov; (c) Gibboryctes endroedii sp. nov.; (d) Tetragus waldenfelsi (Endrödi). Red arrows indicate the proportions of ventrites V and VI." figureDoi="http://doi.org/10.5281/zenodo.6758345" httpUri="https://zenodo.org/record/6758345/files/figure.png" pageId="23" pageNumber="213">Figure 13</figureCitation>
(c)).
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFBDFF87FF55744AFBD62C8A" pageId="23" pageNumber="213" type="etymology">
<paragraph id="47503688FFBDFF87FF55744AFE982C45" blockId="23.[160,1155,1595,1690]" box="[160,285,1595,1621]" pageId="23" pageNumber="213">
<emphasis id="759BEA9AFFBDFF87FF55744AFE982C45" bold="true" box="[160,285,1595,1621]" italics="true" pageId="23" pageNumber="213">Etymology</emphasis>
</paragraph>
<paragraph id="47503688FFBDFF87FF55742FFBD62C8A" blockId="23.[160,1155,1595,1690]" pageId="23" pageNumber="213">
The specific epithet ‘
<taxonomicName id="80EF4D0BFFBDFF87FE61742FFE7C2C67" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" box="[404,505,1630,1655]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="213" phylum="Arthropoda" rank="species" species="endroedii">
<emphasis id="759BEA9AFFBDFF87FE61742FFE7C2C67" box="[404,505,1630,1655]" italics="true" pageId="23" pageNumber="213">endroedii</emphasis>
</taxonomicName>
’ is a homage to Dr Sebö Endrödi, author of the genus
<taxonomicName id="80EF4D0BFFBDFF87FF5574F0FE9A2C8A" box="[160,287,1665,1690]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="213" phylum="Arthropoda" rank="genus">
<emphasis id="759BEA9AFFBDFF87FF5574F0FE9A2C8A" box="[160,287,1665,1690]" italics="true" pageId="23" pageNumber="213">Gibboryctes</emphasis>
</taxonomicName>
and one of the greatest experts on Dynastinae in the twentieth century.
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFB2FF88FF5572FFFDB12ADB" pageId="24" pageNumber="214" type="description">
<paragraph id="47503688FFB2FF88FF5572FFFDE82AB8" blockId="24.[160,621,142,203]" box="[160,621,142,168]" pageId="24" pageNumber="214">
<heading id="1C1881E4FFB2FF88FF5572FFFDE82AB8" bold="true" box="[160,621,142,168]" centered="true" fontSize="11" level="2" pageId="24" pageNumber="214" reason="3">
<emphasis id="759BEA9AFFB2FF88FF5572FFFDE82AB8" bold="true" box="[160,621,142,168]" pageId="24" pageNumber="214">
Geographic distribution
<emphasis id="759BEA9AFFB2FF88FE3972FFFDE82AB8" bold="true" box="[460,621,142,168]" italics="true" pageId="24" pageNumber="214">
(
<figureCitation id="DFD42A0DFFB2FF88FE2072FFFDC72AB8" box="[469,578,142,168]" captionStart="Figure 14" captionStartId="17.[160,227,1388,1411]" captionTargetBox="[168,1147,145,1355]" captionTargetId="figure-89@17.[166,1149,142,1357]" captionTargetPageId="17" captionText="Figure 14. Geographic distribution maps. (a) Tetragus species; (b) Gibboryctes species; (c) Heterogomphus gracilicornis Prell." figureDoi="http://doi.org/10.5281/zenodo.6758347" httpUri="https://zenodo.org/record/6758347/files/figure.png" pageId="24" pageNumber="214">Figure 14</figureCitation>
(b))
</emphasis>
</emphasis>
</heading>
</paragraph>
<paragraph id="47503688FFB2FF88FF5572C3FDB12ADB" blockId="24.[160,621,142,203]" box="[160,564,178,203]" pageId="24" pageNumber="214">
<collectingCountry id="3FF87618FFB2FF88FF5572C3FF592ADB" box="[160,220,178,203]" name="Brazil" pageId="24" pageNumber="214">Brazil</collectingCountry>
(
<collectingRegion id="852BF86AFFB2FF88FF1C72C3FEF92ADB" box="[233,380,178,203]" country="Brazil" name="Minas Gerais" pageId="24" pageNumber="214">Minas Gerais</collectingRegion>
,
<collectingRegion id="852BF86AFFB2FF88FE7372C3FDAF2ADB" box="[390,554,178,203]" country="Brazil" name="Rio de Janeiro" pageId="24" pageNumber="214">Rio de Janeiro</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="0FF56503FFB2FF88FF557377FBC62848" pageId="24" pageNumber="214" type="discussion">
<paragraph id="47503688FFB2FF88FF557377FE832B30" blockId="24.[160,1156,262,600]" box="[160,262,262,288]" pageId="24" pageNumber="214">
<emphasis id="759BEA9AFFB2FF88FF557377FE832B30" bold="true" box="[160,262,262,288]" italics="true" pageId="24" pageNumber="214">Remarks</emphasis>
</paragraph>
<paragraph id="47503688FFB2FF88FF55735BFBC62848" blockId="24.[160,1156,262,600]" pageId="24" pageNumber="214">
<taxonomicName id="80EF4D0BFFB2FF88FF55735BFE0C2B53" authority="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi, 2022" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" box="[160,393,298,323]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="214" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
<emphasis id="759BEA9AFFB2FF88FF55735BFE0C2B53" box="[160,393,298,323]" italics="true" pageId="24" pageNumber="214">Gibboryctes endroedii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EA857E1FFB2FF88FE7A735BFE602B53" box="[399,485,298,323]" pageId="24" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="80EF4D0BFFB2FF88FD94735BFD532B53" authorityName="Endrodi" authorityYear="1974" box="[609,726,298,323]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="214" phylum="Arthropoda" rank="species" species="szelenyii">
<emphasis id="759BEA9AFFB2FF88FD94735BFD532B53" box="[609,726,298,323]" italics="true" pageId="24" pageNumber="214">G. szelenyii</emphasis>
</taxonomicName>
in the reddish-brown body colour and elytral interstriae densely covered with punctures mixed among small and shallow, and large and deep. Besides this, males of
<taxonomicName id="80EF4D0BFFB2FF88FD83731EFC852B98" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" box="[630,768,367,392]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="214" phylum="Arthropoda" rank="species" species="endroedii">
<emphasis id="759BEA9AFFB2FF88FD83731EFC852B98" box="[630,768,367,392]" italics="true" pageId="24" pageNumber="214">G. endroedii</emphasis>
</taxonomicName>
have variations in the shape of parameres that sometimes overlap those observed in
<taxonomicName id="80EF4D0BFFB2FF88FCF273E3FCFA2BBB" authorityName="Endrodi" authorityYear="1974" box="[775,895,402,427]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="214" phylum="Arthropoda" rank="species" species="szelenyii">
<emphasis id="759BEA9AFFB2FF88FCF273E3FCFA2BBB" box="[775,895,402,427]" italics="true" pageId="24" pageNumber="214">G. szelenyii</emphasis>
</taxonomicName>
. However,
<taxonomicName id="80EF4D0BFFB2FF88FC0A73E3FB062BBB" authority="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi, 2022" authorityName="Costa &amp; Duarte &amp; Iannuzzi &amp; Grossi" authorityYear="2022" box="[1023,1155,402,427]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="214" phylum="Arthropoda" rank="species" species="endroedii" status="sp. nov.">
<emphasis id="759BEA9AFFB2FF88FC0A73E3FB062BBB" box="[1023,1155,402,427]" italics="true" pageId="24" pageNumber="214">G. endroedii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6EA857E1FFB2FF88FF5573C5FF7C2BDD" box="[160,249,436,461]" pageId="24" pageNumber="214" rank="species">sp. nov.</taxonomicNameLabel>
is clearly distinct by the labrum parabolic in shape; maxillary galea rounded at apex with all teeth located subapically on the inner margin; anterior corners of pronotum covered with large and coalescent punctures.
<taxonomicName id="80EF4D0BFFB2FF88FD68738BFCFF2803" authorityName="Endrodi" authorityYear="1974" box="[669,890,506,531]" class="Insecta" family="Dynastidae" genus="Gibboryctes" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="214" phylum="Arthropoda" rank="species" species="szelenyii">
<emphasis id="759BEA9AFFB2FF88FD68738BFCFF2803" box="[669,890,506,531]" italics="true" pageId="24" pageNumber="214">Gibboryctes szelenyii</emphasis>
</taxonomicName>
have triangular labrum; triangular maxillary galea with all teeth located at middle of inner margin; anterior corners of pronotum with contiguous punctures or spaced about 1 diameter of punctures.
</paragraph>
</subSubSection>
</treatment>
</document>
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