511 lines
75 KiB
XML
511 lines
75 KiB
XML
<document id="4D3DFE23AE93987A194D0CF7EF3CA5FD" ID-CLB-Dataset="9100" ID-DOI="10.11646/zootaxa.4858.3.8" ID-GBIF-Dataset="129edbf2-2a5a-4c7f-84eb-13334fca6c19" ID-ISSN="1175-5326" ID-Zenodo-Dep="4412537" ID-ZooBank="B48283E5-D2D7-4134-B7C5-0FF74C736BFA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609690113455" checkinUser="plazi" docAuthor="Tchemeris, Aleksey N." docDate="2020" docId="9A4D87FE73706766FF10FD803E02FC04" docLanguage="en" docName="zootaxa.4858.3.8.pdf" docOrigin="Zootaxa 4858 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Euepedanus vietnamicus Tchemeris 2020, sp. nov." docType="treatment" docUuid="39C412B2-02F2-4645-A6E5-33155CDC7819" docUuidSource="ZooBank" docVersion="6" lastPageNumber="436" masterDocId="6674FF867372676FFF87FF9D3D30FFCD" masterDocTitle="Remarkable new species of epedanid from Vietnam (Arachnida: Opiliones Laniatores: Epedanidae)" masterLastPageNumber="437" masterPageNumber="427" pageNumber="429" updateTime="1698851608036" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="275F38539F95DCF9677AAA516CF100C0">
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<mods:title id="8E62DAB12B0208F36E204763B3DAF2F5">Remarkable new species of epedanid from Vietnam (Arachnida: Opiliones Laniatores: Epedanidae)</mods:title>
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<mods:name id="6A3E7073955864B37C75DD1A4B46CD40" type="personal">
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<mods:roleTerm id="35F16979F6FF2E0B85564BD14CE3338F">Author</mods:roleTerm>
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<mods:namePart id="DF72F35D80D9C10AA5E75C7816C3EF4A">Tchemeris, Aleksey N.</mods:namePart>
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<mods:typeOfResource id="54EC0DE76BA925B6169AB5F18B254215">text</mods:typeOfResource>
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<mods:title id="58AA0B49BC3FCD9A4AEB7E66919D1084">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="94A155F9B213EA92498730ED27B8B303">
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<mods:date id="2B57CF49A371BE95D76B22DE3A57BD34">2020</mods:date>
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<mods:number id="D00ADCF2C927B2533F6A88BB7C4959CD">2020-10-01</mods:number>
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<mods:identifier id="226F61A01A9ADF0F61BBAF674FA29382" type="DOI">10.11646/zootaxa.4858.3.8</mods:identifier>
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<treatment id="9A4D87FE73706766FF10FD803E02FC04" ID-DOI="http://doi.org/10.5281/zenodo.4412514" ID-GBIF-Taxon="176419403" ID-Zenodo-Dep="4412514" LSID="urn:lsid:zoobank.org:act:39C412B2-02F2-4645-A6E5-33155CDC7819" httpUri="http://treatment.plazi.org/id/9A4D87FE73706766FF10FD803E02FC04" lastPageId="9" lastPageNumber="436" pageId="2" pageNumber="429">
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<subSubSection id="5AFE65637370676DFF10FD803F1AFDF5" box="[151,554,541,568]" pageId="2" pageNumber="429" type="nomenclature">
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<paragraph id="125B36E87370676DFF10FD803F1AFDF5" blockId="2.[151,554,541,604]" box="[151,554,541,568]" pageId="2" pageNumber="429">
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<heading id="491381847370676DFF10FD803F1AFDF5" bold="true" box="[151,554,541,568]" fontSize="11" level="1" pageId="2" pageNumber="429" reason="1">
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<emphasis id="2090EAFA7370676DFF10FD803F1AFDF5" bold="true" box="[151,554,541,568]" pageId="2" pageNumber="429">
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<taxonomicName id="D5E44D6B7370676DFF10FD803CF5FDF5" authority="Tchemeris, 2020" authorityName="Tchemeris" authorityYear="2020" box="[151,453,541,568]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="vietnamicus" status="sp. nov.">
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<emphasis id="2090EAFA7370676DFF10FD803CF5FDF5" bold="true" box="[151,453,541,568]" italics="true" pageId="2" pageNumber="429">Euepedanus vietnamicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="3BA357817370676DFE4AFD833F1AFDF5" box="[461,554,542,568]" pageId="2" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5AFE65637370676DFF10FDDF3FCAFD2B" pageId="2" pageNumber="429" type="description">
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<paragraph id="125B36E87370676DFF10FDDF3C07FD91" blockId="2.[151,554,541,604]" box="[151,311,578,604]" pageId="2" pageNumber="429">
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<figureCitation id="8ADF2A6D7370676DFF10FDDF3C20FD91" box="[151,272,578,604]" captionStart-0="FIGURES 1–5" captionStart-1="FIGURES 6–10" captionStart-2="FIGURES 11–18" captionStartId-0="1.[151,264,1937,1961]" captionStartId-1="3.[151,264,1943,1967]" captionStartId-2="4.[151,264,1909,1933]" captionTargetBox-0="[167,1415,294,1887]" captionTargetBox-1="[300,1282,210,1897]" captionTargetBox-2="[253,1340,196,1859]" captionTargetId-0="figure-18@1.[151,1436,271,1912]" captionTargetId-1="figure-15@3.[276,1312,181,1920]" captionTargetId-2="figure-24@4.[230,1357,181,1885]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–5. Euepedanus vietnamicus sp. nov., male holotype: 1—body dorsal view, 2—body, left side frontolateral view, 3—left chelicera ectal view, 4—chelicerae dorsofrontal view, 5—distal segment of chelicerae frontal view. Scale bar 1.0 mm." captionText-1="FIGURES 6–10. Euepedanus vietnamicus sp. nov., male holotype: 6—palp mesal view, 7—palp dorsal view, 8—palp ectal view, 9—distal segments of chelicerae dorsofrontal view, 10—habitus left side lateral view. Scale bar 1.0 mm." captionText-2="FIGURES 11–18. Euepedanus vietnamicus sp. nov., 11–14 penis of male holotype: 11—distal area dorsal view, 12—distal area lateral view, 13—distal area ventral view. Scale bar 0.125 mm, 14—corpus dorsally. Scale bar 0.5 mm. 15–18 penis of male paratype: 15—distal area dorsal view, 16—distal area dorsolateral view, 17—distal area ventral view. Scale bar 0.125 mm, 18—corpus dorsally. Scale bar 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4412539" figureDoi-1="http://doi.org/10.5281/zenodo.4412541" figureDoi-2="http://doi.org/10.5281/zenodo.4412543" httpUri-0="https://zenodo.org/record/4412539/files/figure.png" httpUri-1="https://zenodo.org/record/4412541/files/figure.png" httpUri-2="https://zenodo.org/record/4412543/files/figure.png" pageId="2" pageNumber="429">Figs. 1–18</figureCitation>
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,
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<figureCitation id="8ADF2A6D7370676DFE9CFDDF3C07FD91" box="[283,311,578,604]" captionStart="FIGURE 28" captionStartId="7.[151,255,1818,1842]" captionTargetBox="[386,1206,188,1792]" captionTargetId="figure-15@7.[380,1207,181,1794]" captionTargetPageId="7" captionText="FIGURE 28. The type and paratype locality of Euepedanus vietnamicus sp. nov. in Vietnam. Scale bar 200 km." figureDoi="http://doi.org/10.5281/zenodo.4412551" httpUri="https://zenodo.org/record/4412551/files/figure.png" pageId="2" pageNumber="429">28</figureCitation>
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</paragraph>
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<paragraph id="125B36E87370676DFF10FD183FCAFD2B" blockId="2.[151,1437,644,2039]" pageId="2" pageNumber="429">
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<emphasis id="2090EAFA7370676DFF10FD183DD4FD52" bold="true" box="[151,228,645,671]" pageId="2" pageNumber="429">Types.</emphasis>
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<materialsCitation id="A28C3CB57370676DFF6AFD193F2BFD0F" ID-GBIF-Occurrence="3008789303" collectingDate="2011-12" collectionCode="MMUE" collectorName="A. A. Goncharov" country="Vietnam" location="Dong Noi Province" municipality="Cat Tien National-Park" pageId="2" pageNumber="429" specimenCode="G7641.1" specimenCount="1" typeStatus="holotype">
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<typeStatus id="CD5F884A7370676DFF6AFD193C65FD53" box="[237,341,644,670]" pageId="2" pageNumber="429" type="holotype">Holotype</typeStatus>
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male (
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<collectionCode id="74F5AE2D7370676DFE22FD183F31FD52" box="[421,513,645,671]" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="2" pageNumber="429">MMUE</collectionCode>
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,
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<specimenCode id="42429E937370676DFD89FD193F41FD52" box="[526,625,644,671]" collectionCode="MMUE" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="2" pageNumber="429">G7641.1</specimenCode>
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) from
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<collectingCountry id="6AF376787370676DFD47FD193E12FD53" box="[704,802,644,670]" name="Vietnam" pageId="2" pageNumber="429">Vietnam</collectingCountry>
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,
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<location id="173B60337370676DFCA8FD18393FFD53" LSID="urn:lsid:plazi:treatment:9A4D87FE73706766FF10FD803E02FC04:173B60337370676DFCA8FD18393FFD53" box="[815,1039,644,671]" country="Vietnam" municipality="Cat Tien National-Park" name="Dong Noi Province" pageId="2" pageNumber="429">Dong Noi Province</location>
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,
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<collectingMunicipality id="F23FAC927370676DFB9BFD183817FD53" box="[1052,1319,644,671]" pageId="2" pageNumber="429">Cat Tien National-Park</collectingMunicipality>
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, October–
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<date id="665A10287370676DFF10FD353C7FFD0E" box="[151,335,680,707]" pageId="2" pageNumber="429" value="2011-12">
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||
<collectingDate id="761EE9C07370676DFF10FD353C7FFD0E" box="[151,335,680,707]" pageId="2" pageNumber="429" value="2011-12">December 2011</collectingDate>
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</date>
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,
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<collectorName id="BF11533E7370676DFEDBFD343F2BFD0F" box="[348,539,680,707]" pageId="2" pageNumber="429">A.A. Goncharov</collectorName>
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</materialsCitation>
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||
.
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<typeStatus id="CD5F884A7370676DFDAEFD343FAAFD0E" box="[553,666,681,707]" pageId="2" pageNumber="429" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="04E2FD617370676DFD2FFD343FC7FD0F" box="[680,759,680,707]" pageId="2" pageNumber="429" type="male">1 male</specimenCount>
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||
(damaged),
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||
<materialsCitation id="A28C3CB57370676DFC0CFD343C35FD2A" ID-GBIF-Occurrence="3008789304" collectionCode="MMUE" country="Vietnam" pageId="2" pageNumber="429" specimenCode="G7641.2" specimenCount="6" specimenCount-female="6" typeStatus="paratype">
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||
<specimenCount id="04E2FD617370676DFC0CFD343ECBFD0F" box="[907,1019,680,707]" pageId="2" pageNumber="429" type="female">3 females</specimenCount>
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and
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||
<specimenCount id="04E2FD617370676DFBBFFD343998FD0F" box="[1080,1192,680,707]" pageId="2" pageNumber="429" type="female">3 females</specimenCount>
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||
(damaged) (
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||
<collectionCode id="74F5AE2D7370676DFABBFD3438A8FD0E" box="[1340,1432,681,707]" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="2" pageNumber="429">MMUE</collectionCode>
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,
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<specimenCode id="42429E937370676DFF10FD513DCDFD2A" box="[151,253,716,743]" collectionCode="MMUE" country="United Kingdom" httpUri="http://grbio.org/cool/he6q-xsyw" name="Museum of Manchester University" pageId="2" pageNumber="429">G7641.2</specimenCode>
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)
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</materialsCitation>
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,
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<materialsCitation id="A28C3CB57370676DFE97FD503FC7FD2B" ID-GBIF-Occurrence="3008789302" box="[272,759,716,743]" collectionCode="ISEA" country="Vietnam" pageId="2" pageNumber="429" specimenCode="ISEA 001.8549" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
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<specimenCount id="04E2FD617370676DFE97FD503C6EFD2B" box="[272,350,716,743]" pageId="2" pageNumber="429" type="male">1 male</specimenCount>
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,
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||
<specimenCount id="04E2FD617370676DFEEEFD503CFAFD2B" box="[361,458,716,743]" pageId="2" pageNumber="429" type="female">1 female</specimenCount>
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(
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<specimenCode id="42429E937370676DFE5EFD503FB9FD2A" box="[473,649,716,743]" collectionCode="ISEA" pageId="2" pageNumber="429">ISEA 001.8549</specimenCode>
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) — idem
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</materialsCitation>
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||
.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="5AFE65637370676DFF40FD6D3949FCE2" pageId="2" pageNumber="429" type="materials_examined">
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<paragraph id="125B36E87370676DFF40FD6D3949FCE2" blockId="2.[151,1437,644,2039]" pageId="2" pageNumber="429">
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<emphasis id="2090EAFA7370676DFF40FD6D3CE9FCC7" bold="true" box="[199,473,752,778]" pageId="2" pageNumber="429">Comparative material.</emphasis>
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<taxonomicName id="D5E44D6B7370676DFE66FD6D3EA7FCC6" authority="Roewer, 1915" authorityName="Roewer" authorityYear="1915" box="[481,919,752,779]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="trispinosus">
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<emphasis id="2090EAFA7370676DFE66FD6D3FDDFCC7" box="[481,749,752,778]" italics="true" pageId="2" pageNumber="429">Euepedanus trispinosus</emphasis>
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||
<materialsCitation id="A28C3CB57370676DFD71FD6C3EA7FCC6" ID-GBIF-Occurrence="3008789301" box="[758,919,752,779]" collectionCode="SMF" country="Singapore" location="Malakka" municipality="Roewer" pageId="2" pageNumber="429" specimenCode="SMF9901124" specimenCount="1" specimenCount-male="1" typeStatus="holotype">Roewer, 1915</materialsCitation>
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||
</taxonomicName>
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||
:
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<typeStatus id="CD5F884A7370676DFC24FD6D393BFCC7" box="[931,1035,752,778]" pageId="2" pageNumber="429" type="holotype">Holotype</typeStatus>
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||
<specimenCount id="04E2FD617370676DFB93FD6C3919FCC6" box="[1044,1065,753,779]" pageId="2" pageNumber="429" type="male">♂</specimenCount>
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||
(
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<specimenCode id="42429E937370676DFBBFFD6C39E5FCC6" box="[1080,1237,753,779]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="2" pageNumber="429">SMF9901124</specimenCode>
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||
) from
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<collectingCountry id="6AF376787370676DFAA3FD6D38A8FCC7" box="[1316,1432,752,778]" name="Singapore" pageId="2" pageNumber="429">Singapore</collectingCountry>
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,
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<location id="173B60337370676DFF10FC893DCDFCE3" LSID="urn:lsid:plazi:treatment:9A4D87FE73706766FF10FD803E02FC04:173B60337370676DFF10FC893DCDFCE3" box="[151,253,788,814]" country="Singapore" municipality="Roewer" name="Malakka" pageId="2" pageNumber="429">Malakka</location>
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||
; see
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<collectingMunicipality id="F23FAC927370676DFEB4FC883CBAFCE2" box="[307,394,789,815]" pageId="2" pageNumber="429">Roewer</collectingMunicipality>
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(1915: 63, fig. 31 a–b), for its general appearance see
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<figureCitation id="8ADF2A6D7370676DFC6AFC893945FCE2" box="[1005,1141,788,815]" captionStart-0="FIGURES 19–21" captionStart-1="FIGURES 22–27" captionStartId-0="5.[151,264,1944,1968]" captionStartId-1="6.[151,264,1943,1967]" captionTargetBox-0="[413,1176,187,1910]" captionTargetBox-1="[157,1431,181,1920]" captionTargetId-0="figure-15@5.[403,1184,181,1920]" captionTargetId-1="figure-24@6.[157,1431,181,1920]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 19–21. Habitus of the male holotype Euepedanus trispinosus: 19—lateral view, 20–21 dorsal view. Scale bar 1.0 mm." captionText-1="FIGURES 22–27. Male holotype of Euepedanus trispinosus Roewer, 1915: 22–23—lateral view of anterior part of male (22— focused of chelicerae, 23—focused of palp), 24–25—habitusdorsolateral view, 26—habitus frontal view (focused of chelicerae). Scale bar 1.0 mm, 27—the original label." figureDoi-0="http://doi.org/10.5281/zenodo.4412545" figureDoi-1="http://doi.org/10.5281/zenodo.4412547" httpUri-0="https://zenodo.org/record/4412545/files/figure.png" httpUri-1="https://zenodo.org/record/4412547/files/figure.png" pageId="2" pageNumber="429">Figs. 19–27</figureCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5AFE65637370676DFF40FCA43DD2FCBB" pageId="2" pageNumber="429" type="etymology">
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||
<paragraph id="125B36E87370676DFF40FCA43DD2FCBB" blockId="2.[151,1437,644,2039]" pageId="2" pageNumber="429">
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||
<emphasis id="2090EAFA7370676DFF40FCA43C78FC9E" bold="true" box="[199,328,825,851]" pageId="2" pageNumber="429">Etymology</emphasis>
|
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. The species epithet originates from
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<collectingCountry id="6AF376787370676DFD7CFCA53E6DFC9F" box="[763,861,824,850]" name="Vietnam" pageId="2" pageNumber="429">Vietnam</collectingCountry>
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||
, the country from where the
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||
<typeStatus id="CD5F884A7370676DFB32FCA439D5FC9E" box="[1205,1253,825,851]" pageId="2" pageNumber="429">type</typeStatus>
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||
series was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5AFE65637370676DFF40FC1D39EBFB5A" pageId="2" pageNumber="429" type="diagnosis">
|
||
<paragraph id="125B36E87370676DFF40FC1D39EBFB5A" blockId="2.[151,1437,644,2039]" pageId="2" pageNumber="429">
|
||
<emphasis id="2090EAFA7370676DFF40FC1D3C72FC57" bold="true" box="[199,322,896,922]" pageId="2" pageNumber="429">Diagnosis.</emphasis>
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By the structure of body and appendages, this species is closest to
|
||
<taxonomicName id="D5E44D6B7370676DFBB4FC1D3C1EFC73" authority="Zhu & Lian, 2006" authorityName="Zhu & Lian" authorityYear="2006" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="flavimaculatus">
|
||
<emphasis id="2090EAFA7370676DFBB4FC1D3858FC57" box="[1075,1384,896,922]" italics="true" pageId="2" pageNumber="429">Euepedanus flavimaculatus</emphasis>
|
||
<bibRefCitation id="76754B197370676DFAE8FC1D3C1EFC73" author="Zhu, M. S. & Lian, W. G." pageId="2" pageNumber="429" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D5E44D6B7370676DFEE2FC383FB2FC73" authority="Suzuki, 1969" authorityName="Suzuki" authorityYear="1969" box="[357,642,932,958]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="dividuus">
|
||
<emphasis id="2090EAFA7370676DFEE2FC383CD4FC73" box="[357,484,932,958]" italics="true" pageId="2" pageNumber="429">E. dividuus</emphasis>
|
||
<bibRefCitation id="76754B197370676DFE6CFC393FB2FC73" author="Suzuki, S." box="[491,642,932,958]" pageId="2" pageNumber="429" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
. The main diagnostic characters are as follows: 1) penis apex truncate, not trapeziform (
|
||
<figureCitation id="8ADF2A6D7370676DFED5FC553CEAFC2E" box="[338,474,968,995]" captionStart="FIGURES 11–18" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[253,1340,196,1859]" captionTargetId="figure-24@4.[230,1357,181,1885]" captionTargetPageId="4" captionText="FIGURES 11–18. Euepedanus vietnamicus sp. nov., 11–14 penis of male holotype: 11—distal area dorsal view, 12—distal area lateral view, 13—distal area ventral view. Scale bar 0.125 mm, 14—corpus dorsally. Scale bar 0.5 mm. 15–18 penis of male paratype: 15—distal area dorsal view, 16—distal area dorsolateral view, 17—distal area ventral view. Scale bar 0.125 mm, 18—corpus dorsally. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4412543" httpUri="https://zenodo.org/record/4412543/files/figure.png" pageId="2" pageNumber="429">Figs. 11–18</figureCitation>
|
||
); 2) cheliceral basal segment relatively long, somewhat curved and armed with numerous large and robust setae-tipped tubercles (but not strongly drawn), except for ectal and ventral surfaces (
|
||
<figureCitation id="8ADF2A6D7370676DFAC0FC713DF4FBE6" captionStart="FIGURES 1–5" captionStartId="1.[151,264,1937,1961]" captionTargetBox="[167,1415,294,1887]" captionTargetId="figure-18@1.[151,1436,271,1912]" captionTargetPageId="1" captionText="FIGURES 1–5. Euepedanus vietnamicus sp. nov., male holotype: 1—body dorsal view, 2—body, left side frontolateral view, 3—left chelicera ectal view, 4—chelicerae dorsofrontal view, 5—distal segment of chelicerae frontal view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412539" httpUri="https://zenodo.org/record/4412539/files/figure.png" pageId="2" pageNumber="429">Figs. 1, 3–5</figureCitation>
|
||
); 3) distal cheliceral segment markedly swollen, with an apophysis-like structure near chelae formed by several long spines, protruding forward beyond the segment limit (
|
||
<figureCitation id="8ADF2A6D7370676DFCAAFBA93EA9FB82" box="[813,921,1076,1103]" captionStart="FIGURES 1–5" captionStartId="1.[151,264,1937,1961]" captionTargetBox="[167,1415,294,1887]" captionTargetId="figure-18@1.[151,1436,271,1912]" captionTargetPageId="1" captionText="FIGURES 1–5. Euepedanus vietnamicus sp. nov., male holotype: 1—body dorsal view, 2—body, left side frontolateral view, 3—left chelicera ectal view, 4—chelicerae dorsofrontal view, 5—distal segment of chelicerae frontal view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412539" httpUri="https://zenodo.org/record/4412539/files/figure.png" pageId="2" pageNumber="429">Figs. 2–4</figureCitation>
|
||
,
|
||
<figureCitation id="8ADF2A6D7370676DFC23FBA83ED0FB82" box="[932,992,1077,1103]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[300,1282,210,1897]" captionTargetId="figure-15@3.[276,1312,181,1920]" captionTargetPageId="3" captionText="FIGURES 6–10. Euepedanus vietnamicus sp. nov., male holotype: 6—palp mesal view, 7—palp dorsal view, 8—palp ectal view, 9—distal segments of chelicerae dorsofrontal view, 10—habitus left side lateral view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412541" httpUri="https://zenodo.org/record/4412541/files/figure.png" pageId="2" pageNumber="429">9–10</figureCitation>
|
||
); 4) fingers of chelae widened and long (
|
||
<figureCitation id="8ADF2A6D7370676DFF18FBC53C3BFBBE" box="[159,267,1112,1139]" captionStart="FIGURES 1–5" captionStartId="1.[151,264,1937,1961]" captionTargetBox="[167,1415,294,1887]" captionTargetId="figure-18@1.[151,1436,271,1912]" captionTargetPageId="1" captionText="FIGURES 1–5. Euepedanus vietnamicus sp. nov., male holotype: 1—body dorsal view, 2—body, left side frontolateral view, 3—left chelicera ectal view, 4—chelicerae dorsofrontal view, 5—distal segment of chelicerae frontal view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412539" httpUri="https://zenodo.org/record/4412539/files/figure.png" pageId="2" pageNumber="429">Figs. 2–5</figureCitation>
|
||
,
|
||
<figureCitation id="8ADF2A6D7370676DFE91FBC43C62FBBE" box="[278,338,1113,1139]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[300,1282,210,1897]" captionTargetId="figure-15@3.[276,1312,181,1920]" captionTargetPageId="3" captionText="FIGURES 6–10. Euepedanus vietnamicus sp. nov., male holotype: 6—palp mesal view, 7—palp dorsal view, 8—palp ectal view, 9—distal segments of chelicerae dorsofrontal view, 10—habitus left side lateral view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412541" httpUri="https://zenodo.org/record/4412541/files/figure.png" pageId="2" pageNumber="429">9–10</figureCitation>
|
||
); 5) palpal armament (
|
||
<tableCitation id="5F6603537370676DFDCAFBC53F93FBBE" box="[589,675,1112,1139]" captionStart="TABLE 1" captionStartId="8.[151,239,152,176]" captionText="TABLE 1. Main diagnostic characters of four Euepedanus species" pageId="2" pageNumber="429">Table 1</tableCitation>
|
||
;
|
||
<figureCitation id="8ADF2A6D7370676DFD37FBC53E75FBBE" box="[688,837,1112,1139]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[300,1282,210,1897]" captionTargetId="figure-15@3.[276,1312,181,1920]" captionTargetPageId="3" captionText="FIGURES 6–10. Euepedanus vietnamicus sp. nov., male holotype: 6—palp mesal view, 7—palp dorsal view, 8—palp ectal view, 9—distal segments of chelicerae dorsofrontal view, 10—habitus left side lateral view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412541" httpUri="https://zenodo.org/record/4412541/files/figure.png" pageId="2" pageNumber="429">Figs. 6–8, 10</figureCitation>
|
||
). Morphological differences between the new species and three other
|
||
<taxonomicName id="D5E44D6B7370676DFEC1FBE13CFFFB5B" authorityName="Roewer" authorityYear="1915" box="[326,463,1148,1174]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="429" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2090EAFA7370676DFEC1FBE13CFFFB5B" box="[326,463,1148,1174]" italics="true" pageId="2" pageNumber="429">Euepedanus</emphasis>
|
||
</taxonomicName>
|
||
species, including
|
||
<taxonomicName id="D5E44D6B7370676DFD21FBE03E70FB5B" authorityName="Roewer" authorityYear="1915" box="[678,832,1149,1174]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="429" phylum="Arthropoda" rank="species" species="trispinosus">
|
||
<emphasis id="2090EAFA7370676DFD21FBE03E70FB5B" box="[678,832,1149,1174]" italics="true" pageId="2" pageNumber="429">E. trispinosus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="8ADF2A6D7370676DFCC8FBE13EE8FB5A" box="[847,984,1148,1175]" captionStart-0="FIGURES 19–21" captionStart-1="FIGURES 22–27" captionStartId-0="5.[151,264,1944,1968]" captionStartId-1="6.[151,264,1943,1967]" captionTargetBox-0="[413,1176,187,1910]" captionTargetBox-1="[157,1431,181,1920]" captionTargetId-0="figure-15@5.[403,1184,181,1920]" captionTargetId-1="figure-24@6.[157,1431,181,1920]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 19–21. Habitus of the male holotype Euepedanus trispinosus: 19—lateral view, 20–21 dorsal view. Scale bar 1.0 mm." captionText-1="FIGURES 22–27. Male holotype of Euepedanus trispinosus Roewer, 1915: 22–23—lateral view of anterior part of male (22— focused of chelicerae, 23—focused of palp), 24–25—habitusdorsolateral view, 26—habitus frontal view (focused of chelicerae). Scale bar 1.0 mm, 27—the original label." figureDoi-0="http://doi.org/10.5281/zenodo.4412545" figureDoi-1="http://doi.org/10.5281/zenodo.4412547" httpUri-0="https://zenodo.org/record/4412545/files/figure.png" httpUri-1="https://zenodo.org/record/4412547/files/figure.png" pageId="2" pageNumber="429">Figs. 19–27</figureCitation>
|
||
), are shown in
|
||
<tableCitation id="5F6603537370676DFB06FBE139E5FB5A" box="[1153,1237,1148,1175]" captionStart="TABLE 1" captionStartId="8.[151,239,152,176]" captionText="TABLE 1. Main diagnostic characters of four Euepedanus species" pageId="2" pageNumber="429">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5AFE656373706766FF40FB3D3CFEFC68" lastPageId="9" lastPageNumber="436" pageId="2" pageNumber="429" type="description">
|
||
<paragraph id="125B36E873706766FF40FB3D3FB7FE1C" blockId="2.[151,1437,644,2039]" lastBlockId="9.[151,1437,151,969]" lastPageId="9" lastPageNumber="436" pageId="2" pageNumber="429">
|
||
<emphasis id="2090EAFA7370676DFF40FB3D3C69FB77" bold="true" box="[199,345,1184,1210]" pageId="2" pageNumber="429">Description.</emphasis>
|
||
MALE.
|
||
<emphasis id="2090EAFA7370676DFE46FB3C3F53FB77" box="[449,611,1185,1210]" italics="true" pageId="2" pageNumber="429">Measurements</emphasis>
|
||
. Body: length 5.92; maximal width 4.03. Scutum length: 4.90; width in the middle 3.58. ‘Clypeus’ length: 0.68. Eye tubercle width: 1.58; height (from the base of eye tubercle to top) 2.0. Chelicera: basal segment 2.57 long; distal segment 2.81 long; chela 2.35 long. Length of palp segments: 1.08 + 3.53 + 1.62 + 2.05 + 1.80 + claw 1.93 = 12.01. Length of leg segments: I: 4.92 + 1.13 + 3.40 + 6.02 + 1.32 = 16.79; II: 8.82 + 1.63 + 7.58 + 11.12 + 5.09 = 34.24; III: 6.76 + 1.53 + 3.63 + 8.16 + 3.28 = 23.36. IV: 9.28 + 1.78 + 4.65 + 10.95 + 4.12 = 30.78. Penis 2.58 long. Body. Almost rectangular, slightly trapezoid-like as shown in
|
||
<figureCitation id="8ADF2A6D7370676DFB7EFAC9380AFAA2" box="[1273,1338,1364,1391]" captionStart="FIGURES 1–5" captionStartId="1.[151,264,1937,1961]" captionTargetBox="[167,1415,294,1887]" captionTargetId="figure-18@1.[151,1436,271,1912]" captionTargetPageId="1" captionText="FIGURES 1–5. Euepedanus vietnamicus sp. nov., male holotype: 1—body dorsal view, 2—body, left side frontolateral view, 3—left chelicera ectal view, 4—chelicerae dorsofrontal view, 5—distal segment of chelicerae frontal view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412539" httpUri="https://zenodo.org/record/4412539/files/figure.png" pageId="2" pageNumber="429">Fig. 1</figureCitation>
|
||
. Scutum with a wide portion consisting of four sclerites, of which the most narrow one is situated near the rear end. Abdomen widely rounded behind. Anterior carapace margin with 6–8 granules. Eye tubercle separated from the anterior carapace margin and armed with a long median spine on its top. Carapace region with small dome-shaped hump situated around odoriferous glands and a big dome-shaped hump on both sides beloweye tubercle. Opisthosomal region of scutum with four scuta; scutal area II with a pair of median sharp relatively long spines, others areas smooth; fourth area in corners with short spine; each lateral margin of the scutum with a longitudinal row of conical and distinctly transparent tiny tubercles. Each of the free tergites with a row of granules spread out across its entire width. Last granule from a longitudinal row modified to short spine. Venter. Palpal coxae distally armed with 2–3 blunt tubercles. Leg coxae I–II considerably narrower than the others. I—armed with a row of irregular blunt tubercles, IIwith a row of setae-tipped granules and 2–3 distal blunt tubercles, III—on its sides with a row of pointed tubercles, IV—with 1–2 distal blunt tubercles situated at its corners. Tracheal stigma clearly visible; it is equal to about half length of coxa IV. Each of the free sternites with a row of tiny translucentsetae. Chelicerae (
|
||
<figureCitation id="8ADF2A6D7370676DFB10F8993832F8D2" box="[1175,1282,1796,1823]" captionStart="FIGURES 1–5" captionStartId="1.[151,264,1937,1961]" captionTargetBox="[167,1415,294,1887]" captionTargetId="figure-18@1.[151,1436,271,1912]" captionTargetPageId="1" captionText="FIGURES 1–5. Euepedanus vietnamicus sp. nov., male holotype: 1—body dorsal view, 2—body, left side frontolateral view, 3—left chelicera ectal view, 4—chelicerae dorsofrontal view, 5—distal segment of chelicerae frontal view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412539" httpUri="https://zenodo.org/record/4412539/files/figure.png" pageId="2" pageNumber="429">Figs. 1–5</figureCitation>
|
||
,
|
||
<figureCitation id="8ADF2A6D7370676DFA89F8983878F8D2" box="[1294,1352,1797,1823]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[300,1282,210,1897]" captionTargetId="figure-15@3.[276,1312,181,1920]" captionTargetPageId="3" captionText="FIGURES 6–10. Euepedanus vietnamicus sp. nov., male holotype: 6—palp mesal view, 7—palp dorsal view, 8—palp ectal view, 9—distal segments of chelicerae dorsofrontal view, 10—habitus left side lateral view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412541" httpUri="https://zenodo.org/record/4412541/files/figure.png" pageId="2" pageNumber="429">9–10</figureCitation>
|
||
) strong and robust. Basal segment relatively long, not swollen, somewhat curved and armed with numerous large and robust setae-tipped tubercles, except for ectal and ventral surfaces. Distal segment large and swollen, frontally armed with two irregular rows of setae-tipped tubercles, the three lowermost long setae-tipped tubercles at the base of chelae are fused, forming a prominent apophysis resembling a crown. Chela fingers relatively long, widened and strong, movable finger curved with strong and robust cutting teeth, their cutting edges toothed as in
|
||
<figureCitation id="8ADF2A6D7370676DFB24F825383EF81E" box="[1187,1294,1976,2003]" captionStart="FIGURES 1–5" captionStartId="1.[151,264,1937,1961]" captionTargetBox="[167,1415,294,1887]" captionTargetId="figure-18@1.[151,1436,271,1912]" captionTargetPageId="1" captionText="FIGURES 1–5. Euepedanus vietnamicus sp. nov., male holotype: 1—body dorsal view, 2—body, left side frontolateral view, 3—left chelicera ectal view, 4—chelicerae dorsofrontal view, 5—distal segment of chelicerae frontal view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412539" httpUri="https://zenodo.org/record/4412539/files/figure.png" pageId="2" pageNumber="429">Figs. 2–5</figureCitation>
|
||
,
|
||
<figureCitation id="8ADF2A6D7370676DFA9DF8243865F81E" box="[1306,1365,1977,2003]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[300,1282,210,1897]" captionTargetId="figure-15@3.[276,1312,181,1920]" captionTargetPageId="3" captionText="FIGURES 6–10. Euepedanus vietnamicus sp. nov., male holotype: 6—palp mesal view, 7—palp dorsal view, 8—palp ectal view, 9—distal segments of chelicerae dorsofrontal view, 10—habitus left side lateral view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412541" httpUri="https://zenodo.org/record/4412541/files/figure.png" pageId="2" pageNumber="429">9–10</figureCitation>
|
||
. Palps (
|
||
<figureCitation id="8ADF2A6D7370676DFF18F8413C3DF83A" box="[159,269,2012,2039]" captionStart="FIGURES 1–5" captionStartId="1.[151,264,1937,1961]" captionTargetBox="[167,1415,294,1887]" captionTargetId="figure-18@1.[151,1436,271,1912]" captionTargetPageId="1" captionText="FIGURES 1–5. Euepedanus vietnamicus sp. nov., male holotype: 1—body dorsal view, 2—body, left side frontolateral view, 3—left chelicera ectal view, 4—chelicerae dorsofrontal view, 5—distal segment of chelicerae frontal view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412539" httpUri="https://zenodo.org/record/4412539/files/figure.png" pageId="2" pageNumber="429">Figs. 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="8ADF2A6D7370676DFE9BF8413C43F83A" box="[284,371,2012,2039]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[300,1282,210,1897]" captionTargetId="figure-15@3.[276,1312,181,1920]" captionTargetPageId="3" captionText="FIGURES 6–10. Euepedanus vietnamicus sp. nov., male holotype: 6—palp mesal view, 7—palp dorsal view, 8—palp ectal view, 9—distal segments of chelicerae dorsofrontal view, 10—habitus left side lateral view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412541" httpUri="https://zenodo.org/record/4412541/files/figure.png" pageId="2" pageNumber="429">6–8, 10</figureCitation>
|
||
) relatively long, slightly swollen, armed with strong spines. Coxa DE—with strong crescent -shaped tubercle, D and M with rare LCT. Palpal spinulation formula: Tr: D—2 LCT, V—1 SLT; Fm: D—7–8 LCT, V—5 SLT + 2 SMT, E—2 distal SLT, DE and VE—5–6 TCT; Pt: VE—2 SLT, VM—1 SLT; Tb: VE—3 SLT, VM—4 SLT; Tr: VE and VM—3 SLT. Tarsal claw almost equal to tarsus long, slightly curved. Legs (
|
||
<figureCitation id="8ADF2A6D737B6766FA86FF42385BFF34" box="[1281,1387,223,249]" captionStart="FIGURES 1–5" captionStartId="1.[151,264,1937,1961]" captionTargetBox="[167,1415,294,1887]" captionTargetId="figure-18@1.[151,1436,271,1912]" captionTargetPageId="1" captionText="FIGURES 1–5. Euepedanus vietnamicus sp. nov., male holotype: 1—body dorsal view, 2—body, left side frontolateral view, 3—left chelicera ectal view, 4—chelicerae dorsofrontal view, 5—distal segment of chelicerae frontal view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412539" httpUri="https://zenodo.org/record/4412539/files/figure.png" pageId="9" pageNumber="436">Figs. 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="8ADF2A6D737B6766FAF0FF4238A4FF34" box="[1399,1428,223,249]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[300,1282,210,1897]" captionTargetId="figure-15@3.[276,1312,181,1920]" captionTargetPageId="3" captionText="FIGURES 6–10. Euepedanus vietnamicus sp. nov., male holotype: 6—palp mesal view, 7—palp dorsal view, 8—palp ectal view, 9—distal segments of chelicerae dorsofrontal view, 10—habitus left side lateral view. Scale bar 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4412541" httpUri="https://zenodo.org/record/4412541/files/figure.png" pageId="9" pageNumber="436">10</figureCitation>
|
||
) slender and relatively long, III–IV thicker than others. Femora I slightly curved, II–IV straight, femoral bases of all legs with 1–2 pseudosegments. All metatarsi with numerous pseudosegments. Tarsi III–IV with a pair of claws, no scopulae. Distitarsus I leg with two tarsomeres. Tarsal articulations: I: 9, II: 25–28, III: 8, IV: 9. Penis (
|
||
<figureCitation id="8ADF2A6D737B6766FA88FED638A4FEA8" box="[1295,1428,331,357]" captionStart="FIGURES 11–18" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[253,1340,196,1859]" captionTargetId="figure-24@4.[230,1357,181,1885]" captionTargetPageId="4" captionText="FIGURES 11–18. Euepedanus vietnamicus sp. nov., 11–14 penis of male holotype: 11—distal area dorsal view, 12—distal area lateral view, 13—distal area ventral view. Scale bar 0.125 mm, 14—corpus dorsally. Scale bar 0.5 mm. 15–18 penis of male paratype: 15—distal area dorsal view, 16—distal area dorsolateral view, 17—distal area ventral view. Scale bar 0.125 mm, 18—corpus dorsally. Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4412543" httpUri="https://zenodo.org/record/4412543/files/figure.png" pageId="9" pageNumber="436">Figs. 11–18</figureCitation>
|
||
) long and slender, its truncus is gradually widened distad. Distal area with broad deepening along the middle line and rows of spicules, glans (capsules externa and interna) are located inside the depression. Apex of the ventral plate emarginate in the middle. Base very narrow.
|
||
</paragraph>
|
||
<caption id="469B66607371676CFF10F80A39C6F81E" ID-DOI="http://doi.org/10.5281/zenodo.4412541" ID-Zenodo-Dep="4412541" httpUri="https://zenodo.org/record/4412541/files/figure.png" pageId="3" pageNumber="430" startId="3.[151,264,1943,1967]" targetBox="[300,1282,210,1897]" targetPageId="3">
|
||
<paragraph id="125B36E87371676CFF10F80A39C6F81E" blockId="3.[151,1436,1943,2004]" pageId="3" pageNumber="430">
|
||
<emphasis id="2090EAFA7371676CFF10F80A3C79F862" bold="true" box="[151,329,1943,1967]" pageId="3" pageNumber="430">FIGURES 6–10.</emphasis>
|
||
<taxonomicName id="D5E44D6B7371676CFED5F8053F7FF862" authority="Tchemeris, 2020" authorityName="Tchemeris" authorityYear="2020" box="[338,591,1944,1968]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="430" phylum="Arthropoda" rank="species" species="vietnamicus" status="sp. nov.">
|
||
<emphasis id="2090EAFA7371676CFED5F8053F7FF862" box="[338,591,1944,1968]" italics="true" pageId="3" pageNumber="430">Euepedanus vietnamicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3BA357817371676CFDD0F8053F9AF862" box="[599,682,1944,1967]" pageId="3" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male holotype: 6—palp mesal view, 7—palp dorsal view, 8—palp ectal view, 9—distal segments of chelicerae dorsofrontal view, 10—habitus left side lateral view. Scale bar 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="469B66607376676BFF10F8E83F19F835" ID-DOI="http://doi.org/10.5281/zenodo.4412543" ID-Zenodo-Dep="4412543" httpUri="https://zenodo.org/record/4412543/files/figure.png" pageId="4" pageNumber="431" startId="4.[151,264,1909,1933]" targetBox="[253,1340,196,1859]" targetPageId="4">
|
||
<paragraph id="125B36E87376676BFF10F8E83F19F835" blockId="4.[151,1437,1909,2041]" pageId="4" pageNumber="431">
|
||
<emphasis id="2090EAFA7376676BFF10F8E83C65F840" bold="true" box="[151,341,1909,1933]" pageId="4" pageNumber="431">FIGURES 11–18.</emphasis>
|
||
<taxonomicName id="D5E44D6B7376676BFED9F8E83F6BF840" authority="Tchemeris, 2020" authorityName="Tchemeris" authorityYear="2020" box="[350,603,1909,1933]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="431" phylum="Arthropoda" rank="species" species="vietnamicus" status="sp. nov.">
|
||
<emphasis id="2090EAFA7376676BFED9F8E83F6BF840" box="[350,603,1909,1933]" italics="true" pageId="4" pageNumber="431">Euepedanus vietnamicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3BA357817376676BFDE3F8E83F87F841" box="[612,695,1909,1932]" pageId="4" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, 11–14 penis of male holotype: 11—distal area dorsal view, 12—distal area lateral view, 13—distal area ventral view. Scale bar 0.125 mm, 14—corpus dorsally. Scale bar 0.5 mm. 15–18 penis of male paratype: 15—distal area dorsal view, 16—distal area dorsolateral view, 17—distal area ventral view. Scale bar 0.125 mm, 18—corpus dorsally. Scale bar 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="469B66607377676AFF10F8053DF6F819" ID-DOI="http://doi.org/10.5281/zenodo.4412545" ID-Zenodo-Dep="4412545" httpUri="https://zenodo.org/record/4412545/files/figure.png" pageId="5" pageNumber="432" startId="5.[151,264,1944,1968]" targetBox="[413,1176,187,1910]" targetPageId="5">
|
||
<paragraph id="125B36E87377676AFF10F8053DF6F819" blockId="5.[151,1437,1944,2004]" pageId="5" pageNumber="432">
|
||
<emphasis id="2090EAFA7377676AFF10F8053C66F87D" bold="true" box="[151,342,1944,1968]" pageId="5" pageNumber="432">FIGURES 19–21.</emphasis>
|
||
Habitus of the male holotype
|
||
<taxonomicName id="D5E44D6B7377676AFD16F8053EB4F87D" authorityName="Roewer" authorityYear="1915" box="[657,900,1944,1968]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="432" phylum="Arthropoda" rank="species" species="trispinosus">
|
||
<emphasis id="2090EAFA7377676AFD16F8053EB4F87D" box="[657,900,1944,1968]" italics="true" pageId="5" pageNumber="432">Euepedanus trispinosus</emphasis>
|
||
</taxonomicName>
|
||
: 19—lateral view, 20–21 dorsal view. Scale bar 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="469B666073746769FF10F80A3F0BF835" ID-DOI="http://doi.org/10.5281/zenodo.4412547" ID-Zenodo-Dep="4412547" httpUri="https://zenodo.org/record/4412547/files/figure.png" pageId="6" pageNumber="433" startId="6.[151,264,1943,1967]" targetBox="[157,1431,181,1920]" targetPageId="6">
|
||
<paragraph id="125B36E873746769FF10F80A3F0BF835" blockId="6.[151,1437,1943,2040]" pageId="6" pageNumber="433">
|
||
<emphasis id="2090EAFA73746769FF10F80A3C64F862" bold="true" box="[151,340,1943,1967]" pageId="6" pageNumber="433">FIGURES 22–27.</emphasis>
|
||
Male holotype of
|
||
<taxonomicName id="D5E44D6B73746769FD8AF8053EEBF87D" authority="Roewer, 1915: 22 - 23" authorityName="Roewer" authorityPageNumber="22 - 23" authorityYear="1915" box="[525,987,1943,1968]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="433" phylum="Arthropoda" rank="species" species="trispinosus">
|
||
<emphasis id="2090EAFA73746769FD8AF8053FCDF862" box="[525,765,1944,1968]" italics="true" pageId="6" pageNumber="433">Euepedanus trispinosus</emphasis>
|
||
Roewer, 1915: 22–23
|
||
</taxonomicName>
|
||
—lateral view of anterior part of male (22— focused of chelicerae, 23—focused of palp), 24–25—habitusdorsolateral view, 26—habitus frontal view (focused of chelicerae). Scale bar 1.0 mm, 27—the original label.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="469B666073756768FF10F8873870F8FF" ID-DOI="http://doi.org/10.5281/zenodo.4412551" ID-Zenodo-Dep="4412551" box="[151,1344,1818,1842]" httpUri="https://zenodo.org/record/4412551/files/figure.png" pageId="7" pageNumber="434" startId="7.[151,255,1818,1842]" targetBox="[386,1206,188,1792]" targetPageId="7">
|
||
<paragraph id="125B36E873756768FF10F8873870F8FF" blockId="7.[151,1344,1818,1842]" box="[151,1344,1818,1842]" pageId="7" pageNumber="434">
|
||
<emphasis id="2090EAFA73756768FF10F8873C11F8FF" bold="true" box="[151,289,1818,1842]" pageId="7" pageNumber="434">FIGURE 28</emphasis>
|
||
. The type and paratype locality of
|
||
<taxonomicName id="D5E44D6B73756768FD12F8873EADF8FF" authority="Tchemeris, 2020" authorityName="Tchemeris" authorityYear="2020" box="[661,925,1818,1842]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="434" phylum="Arthropoda" rank="species" species="vietnamicus" status="sp. nov.">
|
||
<emphasis id="2090EAFA73756768FD12F8873EADF8FF" box="[661,925,1818,1842]" italics="true" pageId="7" pageNumber="434">Euepedanus vietnamicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3BA3578173756768FC23F8873EC7F8FC" box="[932,1015,1818,1841]" pageId="7" pageNumber="434" rank="species">sp. nov.</taxonomicNameLabel>
|
||
in Vietnam. Scale bar 200 km.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="469B6660737A6767FF10FF053E6DFF7D" box="[151,861,152,176]" pageId="8" pageNumber="435">
|
||
<paragraph id="125B36E8737A6767FF10FF053E6DFF7D" blockId="8.[151,861,152,176]" box="[151,861,152,176]" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF10FF053C3AFF7D" bold="true" box="[151,266,152,176]" pageId="8" pageNumber="435">TABLE 1.</emphasis>
|
||
Main diagnostic characters of four
|
||
<taxonomicName id="D5E44D6B737A6767FD00FF053E39FF7D" authorityName="Roewer" authorityYear="1915" box="[647,777,152,176]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="435" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2090EAFA737A6767FD00FF053E39FF7D" box="[647,777,152,176]" italics="true" pageId="8" pageNumber="435">Euepedanus</emphasis>
|
||
</taxonomicName>
|
||
species
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="125B36E8737A6767FF35FF5238BDF876" pageId="8" pageNumber="435">
|
||
<table id="60E4C448737A9890FF24FF5D38A0F876" box="[163,1424,192,1979]" gridcols="5" gridrows="13" pageId="8" pageNumber="435">
|
||
<tr id="ACD434AA737A9890FF24FF5D38A0FF3B" box="[163,1424,192,246]" gridrow="0" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24FF5D3C08FF3B" box="[163,312,192,246]" gridcol="0" gridrow="0" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF35FF523C1BFF2A" bold="true" box="[178,299,207,231]" pageId="8" pageNumber="435">Characters</emphasis>
|
||
</th>
|
||
<th id="EF055DD6737A9890FED4FF5D3F7EFF3B" box="[339,590,192,246]" gridcol="1" gridrow="0" pageId="8" pageNumber="435">
|
||
<taxonomicName id="D5E44D6B737A6767FEDAFF523CC7FF2A" authority="Tchemeris, 2020" authorityName="Tchemeris" authorityYear="2020" box="[349,503,207,231]" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="vietnamicus" status="sp. nov.">
|
||
<emphasis id="2090EAFA737A6767FEDAFF523CC7FF2A" bold="true" box="[349,503,207,231]" italics="true" pageId="8" pageNumber="435">E. vietnamicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="3BA35781737A6767FE79FF4D3F76FF2A" box="[510,582,208,231]" pageId="8" pageNumber="435" rank="species">sp.nov.</taxonomicNameLabel>
|
||
</th>
|
||
<th id="EF055DD6737A9890FDE0FF5D3E56FF3B" box="[615,870,192,246]" gridcol="2" gridrow="0" pageId="8" pageNumber="435">
|
||
<taxonomicName id="D5E44D6B737A6767FD0CFF5C3E72FF3B" authority="Zhu et Lian, 2006" authorityName="Zhu et Lian" authorityYear="2006" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="flavimaculatus">
|
||
<emphasis id="2090EAFA737A6767FD0CFF5C3E73FF15" bold="true" box="[651,835,192,216]" italics="true" pageId="8" pageNumber="435">E. flavimaculatus</emphasis>
|
||
<bibRefCitation id="76754B19737A6767FD0BFF433E72FF3B" author="Zhu, M. S. & Lian, W. G." box="[652,834,222,246]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu et Lian, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="EF055DD6737A9890FCF8FF5D394AFF3B" box="[895,1146,192,246]" gridcol="3" gridrow="0" pageId="8" pageNumber="435">
|
||
<taxonomicName id="D5E44D6B737A6767FC46FF5C3970FF3B" authority="Suzuki, 1969" authorityName="Suzuki" authorityYear="1969" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="dividuus">
|
||
<emphasis id="2090EAFA737A6767FC46FF5C3908FF15" bold="true" box="[961,1080,192,216]" italics="true" pageId="8" pageNumber="435">E. dividuus</emphasis>
|
||
<bibRefCitation id="76754B19737A6767FC3DFF433970FF3B" author="Suzuki, S." box="[954,1088,222,246]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki, 1969</bibRefCitation>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="EF055DD6737A9890FB12FF5D38A0FF3B" box="[1173,1424,192,246]" gridcol="4" gridrow="0" pageId="8" pageNumber="435">
|
||
<taxonomicName id="D5E44D6B737A6767FB4CFF5C3869FF38" authority="Roewer, 1915" authorityName="Roewer" authorityYear="1915" class="Arachnida" family="Epedanidae" genus="Euepedanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="435" phylum="Arthropoda" rank="species" species="trispinosus">
|
||
<emphasis id="2090EAFA737A6767FB4CFF5C386AFF15" bold="true" box="[1227,1370,192,216]" italics="true" pageId="8" pageNumber="435">E. trispinosus</emphasis>
|
||
Roewer, 1915
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24FE9B38A0FE5E" box="[163,1424,262,403]" gridrow="1" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24FE9B3C08FE5E" box="[163,312,262,403]" gridcol="0" gridrow="1" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24FEBE3DCBFEBB" bold="true" pageId="8" pageNumber="435">Median spine of eye tubercle</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4FE9B3F7EFE5E" box="[339,590,262,403]" gridcol="1" gridrow="1" pageId="8" pageNumber="435">Long (Figs. 2, 10)</td>
|
||
<td id="EF055DD6737A9890FDE0FE9B3E56FE5E" box="[615,870,262,403]" gridcol="2" gridrow="1" pageId="8" pageNumber="435">
|
||
Long (
|
||
<bibRefCitation id="76754B19737A6767FDFEFEDD3E00FE95" author="Zhu, M. S. & Lian, W. G." box="[633,816,320,345]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian 2006</bibRefCitation>
|
||
: fig. 1)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8FE9B394AFE5E" box="[895,1146,262,403]" gridcol="3" gridrow="1" pageId="8" pageNumber="435">
|
||
Long (
|
||
<bibRefCitation id="76754B19737A6767FC1DFED2392EFEAA" author="Suzuki, S." box="[922,1054,335,359]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: fig. 6)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12FE9B38A0FE5E" box="[1173,1424,262,403]" gridcol="4" gridrow="1" pageId="8" pageNumber="435">Short (Figs. 19–25) Remark: Roewer 1915, fig. 31a is inaccurate)</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24FE3E38A0FDD9" box="[163,1424,419,532]" gridrow="2" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24FE3E3C08FDD9" box="[163,312,419,532]" gridcol="0" gridrow="2" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24FE2F3D87FDC8" bold="true" pageId="8" pageNumber="435">Pair of spines on scutal area II</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4FE3E3F7EFDD9" box="[339,590,419,532]" gridcol="1" gridrow="2" pageId="8" pageNumber="435">Long (Figs. 1–2, 10)</td>
|
||
<td id="EF055DD6737A9890FDE0FE3E3E56FDD9" box="[615,870,419,532]" gridcol="2" gridrow="2" pageId="8" pageNumber="435">
|
||
Long (
|
||
<bibRefCitation id="76754B19737A6767FDFEFE523E00FE2A" author="Zhu, M. S. & Lian, W. G." box="[633,816,463,488]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian 2006</bibRefCitation>
|
||
: fig. 1)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8FE3E394AFDD9" box="[895,1146,419,532]" gridcol="3" gridrow="2" pageId="8" pageNumber="435">
|
||
Short (
|
||
<bibRefCitation id="76754B19737A6767FC1DFE43392EFE3B" author="Suzuki, S." box="[922,1054,478,502]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: fig. 6)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12FE3E38A0FDD9" box="[1173,1424,419,532]" gridcol="4" gridrow="2" pageId="8" pageNumber="435">Short (Figs. 19–21) Remark: Roewer 1915, fig. 31a is inaccurate)</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24FDB938A0FC89" box="[163,1424,548,836]" gridrow="3" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24FDB93C08FC89" box="[163,312,548,836]" gridcol="0" gridrow="3" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24FD173C34FD10" bold="true" pageId="8" pageNumber="435">Basal segment of chelicera</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4FDB93F7EFC89" box="[339,590,548,836]" gridcol="1" gridrow="3" pageId="8" pageNumber="435">Relatively long somewhat curved and armed with numerous large and robust setae- tipped tubercles (but not strongly drawn), except for ectal and ventral surface (Figs. 1, 3–5)</td>
|
||
<td id="EF055DD6737A9890FDE0FDB93E56FC89" box="[615,870,548,836]" gridcol="2" gridrow="3" pageId="8" pageNumber="435">
|
||
Relatively long and curved, swollen distal- dorsally. Dorsally and mesally almost not armed, with sparse tiny setae-tipped tubercles (
|
||
<bibRefCitation id="76754B19737A6767FDF3FD6C3E1BFCC4" author="Zhu, M. S. & Lian, W. G." box="[628,811,753,777]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian 2006</bibRefCitation>
|
||
: figs. 1–3)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8FDB9394AFC89" box="[895,1146,548,836]" gridcol="3" gridrow="3" pageId="8" pageNumber="435">
|
||
Considerably elongated and swollen distal- dorsally. Dorsally with strong prominent setae-tipped tubercles of different forms. One long and conical others short and robust (
|
||
<bibRefCitation id="76754B19737A6767FC0AFC933921FCEB" author="Suzuki, S." box="[909,1041,782,806]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: figs. 6, 7 B–D)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12FDB938A0FC89" box="[1173,1424,548,836]" gridcol="4" gridrow="3" pageId="8" pageNumber="435">Some elongated and swollen distal-dorsally. Dorsally with small prominent setae-tipped tubercles (Figs. 20–21, 23–25 and Roewer 1915: fig. 31a)</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24FCC938A0FC44" box="[163,1424,852,905]" gridrow="4" pageId="8" pageNumber="435" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<td id="EF055DD6737A9890FED4FCC93F7EFC44" box="[339,590,852,905]" gridcol="1" gridrow="4" pageId="8" pageNumber="435">Markedly swollen, with an apophysis-like</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24FC1238A0FB9A" box="[163,1424,911,1111]" gridrow="5" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24FC123C08FB9A" box="[163,312,911,1111]" gridcol="0" gridrow="5" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24FC273C34FBC0" bold="true" pageId="8" pageNumber="435">Distal segment of chelicera</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4FC123F7EFB9A" box="[339,590,911,1111]" gridcol="1" gridrow="5" pageId="8" pageNumber="435">structure near chelae, which is formed by several long spines, protruding forward beyond the segment limit (Figs. 2–4, 9–10)</td>
|
||
<td id="EF055DD6737A9890FDE0FC123E56FB9A" box="[615,870,911,1111]" gridcol="2" gridrow="5" pageId="8" pageNumber="435">
|
||
Slightly swollen, near chelae with a structure made of low setae,
|
||
<emphasis id="2090EAFA737A6767FCB7FC573E6BFC2C" box="[816,859,970,993]" italics="true" pageId="8" pageNumber="435">e.g.,</emphasis>
|
||
tipped tubercles, slightly protruding forward. (
|
||
<bibRefCitation id="76754B19737A6767FCBBFB993FC5FBF4" author="Zhu, M. S. & Lian, W. G." pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian 2006</bibRefCitation>
|
||
: figs. 2, 4)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8FC12394AFB9A" box="[895,1146,911,1111]" gridcol="3" gridrow="5" pageId="8" pageNumber="435">
|
||
Markedly swollen. No complicated structure near chelae (
|
||
<bibRefCitation id="76754B19737A6767FC0AFC683921FBC0" author="Suzuki, S." box="[909,1041,1013,1037]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: figs. 6, 7 C and E)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12FC1238A0FB9A" box="[1173,1424,911,1111]" gridcol="4" gridrow="5" pageId="8" pageNumber="435">Slightly swollen. No complicated structure near chelae (Figs. 19, 21–26) Remark: Roewer 1915: fig. 31b is innacurate)</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24FB1938A0FAE2" box="[163,1424,1156,1327]" gridrow="6" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24FB193C08FAE2" box="[163,312,1156,1327]" gridcol="0" gridrow="6" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24FB233DD7FB39" bold="true" pageId="8" pageNumber="435">Fingers of chelae</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4FB193F7EFAE2" box="[339,590,1156,1327]" gridcol="1" gridrow="6" pageId="8" pageNumber="435">Widened and long. Movable finger curved with strong and robust cutting teeth (Figs. 2–5, 9–10)</td>
|
||
<td id="EF055DD6737A9890FDE0FB193E56FAE2" box="[615,870,1156,1327]" gridcol="2" gridrow="6" pageId="8" pageNumber="435">
|
||
Usual for the epedanids (
|
||
<bibRefCitation id="76754B19737A6767FDF3FB503E1BFB28" author="Zhu, M. S. & Lian, W. G." box="[628,811,1229,1254]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian 2006</bibRefCitation>
|
||
: figs. 2, 4)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8FB19394AFAE2" box="[895,1146,1156,1327]" gridcol="3" gridrow="6" pageId="8" pageNumber="435">
|
||
Usual for the epedanids (
|
||
<bibRefCitation id="76754B19737A6767FC0AFB503921FB28" author="Suzuki, S." box="[909,1041,1229,1253]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: figs. 6–7 B–C, E)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12FB1938A0FAE2" box="[1173,1424,1156,1327]" gridcol="4" gridrow="6" pageId="8" pageNumber="435">Thin and long. Movable finger markedly curved, with strong and cutting teeth (Figs. 19, 22, 26 and Roewer 1915: fig. 31b)</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24FAA238A0FA5F" box="[163,1424,1343,1426]" gridrow="7" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24FAA23C08FA5F" box="[163,312,1343,1426]" gridcol="0" gridrow="7" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24FAD03C2BFA4E" bold="true" pageId="8" pageNumber="435">Armament of palp femur</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4FAA23F7EFA5F" box="[339,590,1343,1426]" gridcol="1" gridrow="7" pageId="8" pageNumber="435">Fm: V—5 SLT + 2 SMT (Figs. 1, 6–7, 10)</td>
|
||
<td id="EF055DD6737A9890FDE0FAA23E56FA5F" box="[615,870,1343,1426]" gridcol="2" gridrow="7" pageId="8" pageNumber="435">
|
||
Fm: V—6 SLT + 1 SMT (
|
||
<bibRefCitation id="76754B19737A6767FDF3FAC13E1BFAB9" author="Zhu, M. S. & Lian, W. G." box="[628,811,1372,1397]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian 2006</bibRefCitation>
|
||
: figs. 1, 5–6)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8FAA2394AFA5F" box="[895,1146,1343,1426]" gridcol="3" gridrow="7" pageId="8" pageNumber="435">
|
||
Fm: V—6 SLT (
|
||
<bibRefCitation id="76754B19737A6767FC11FAC1392AFAB9" author="Suzuki, S." box="[918,1050,1372,1396]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: figs. 6, 7 F)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12FAA238A0FA5F" box="[1173,1424,1343,1426]" gridcol="4" gridrow="7" pageId="8" pageNumber="435">Fm: V—5 SLT (Figs. 19–25 and Roewer 1915: fig. 31a)</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24FA3F38A0F9DF" box="[163,1424,1442,1554]" gridrow="8" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24FA3F3C08F9DF" box="[163,312,1442,1554]" gridcol="0" gridrow="8" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24FA223C12FA38" bold="true" pageId="8" pageNumber="435">Armament of palp patella</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4FA3F3F7EF9DF" box="[339,590,1442,1554]" gridcol="1" gridrow="8" pageId="8" pageNumber="435">Pt: VE—2 SLT, VM—1 SLT (Fig. 7)</td>
|
||
<td id="EF055DD6737A9890FDE0FA3F3E56F9DF" box="[615,870,1442,1554]" gridcol="2" gridrow="8" pageId="8" pageNumber="435">
|
||
Pt: VE—2 SLT, VM—2 SLT (
|
||
<bibRefCitation id="76754B19737A6767FDF6FA403E1EFA38" author="Zhu, M. S. & Lian, W. G." box="[625,814,1501,1525]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian, 2006</bibRefCitation>
|
||
: figs. 1, 5–6)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8FA3F394AF9DF" box="[895,1146,1442,1554]" gridcol="3" gridrow="8" pageId="8" pageNumber="435">
|
||
Pt: VE—2 SLT, VM—1 SLT (
|
||
<bibRefCitation id="76754B19737A6767FC11FA40392AFA38" author="Suzuki, S." box="[918,1050,1501,1525]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: figs. 6, 7 F)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12FA3F38A0F9DF" box="[1173,1424,1442,1554]" gridcol="4" gridrow="8" pageId="8" pageNumber="435">Pt: VE—2 SLT, VM—1 SLT (Figs. 20–24 and Roewer 1915: fig. 31a)</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24F9BF38A0F95F" box="[163,1424,1570,1682]" gridrow="9" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24F9BF3C08F95F" box="[163,312,1570,1682]" gridcol="0" gridrow="9" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24F9A23C3AF9B8" bold="true" pageId="8" pageNumber="435">Armament of palp tibia</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4F9BF3F7EF95F" box="[339,590,1570,1682]" gridcol="1" gridrow="9" pageId="8" pageNumber="435">Tb: VE—3 SLT, VM—4 SLT (Fig. 7)</td>
|
||
<td id="EF055DD6737A9890FDE0F9BF3E56F95F" box="[615,870,1570,1682]" gridcol="2" gridrow="9" pageId="8" pageNumber="435">
|
||
Tb: VE—3 SLT, VM—4 SLT + 1 SMT (
|
||
<bibRefCitation id="76754B19737A6767FDF3F9C03E1BF9B8" author="Zhu, M. S. & Lian, W. G." box="[628,811,1629,1653]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian 2006</bibRefCitation>
|
||
: figs. 1, 5–6)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8F9BF394AF95F" box="[895,1146,1570,1682]" gridcol="3" gridrow="9" pageId="8" pageNumber="435">
|
||
Tb: VE—3 SLT, VM—4 SLT (
|
||
<bibRefCitation id="76754B19737A6767FC11F9C0392AF9B8" author="Suzuki, S." box="[918,1050,1629,1653]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: figs. 6, 7 F)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12F9BF38A0F95F" box="[1173,1424,1570,1682]" gridcol="4" gridrow="9" pageId="8" pageNumber="435">Tb: VE—3 SLT, VM—3 SLT (Figs. 22–25 and Roewer 1915: fig. 31a)</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24F93F38A0F8DE" box="[163,1424,1698,1811]" gridrow="10" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24F93F3C08F8DE" box="[163,312,1698,1811]" gridcol="0" gridrow="10" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24F9223DEBF938" bold="true" pageId="8" pageNumber="435">Corpus of penis</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4F93F3F7EF8DE" box="[339,590,1698,1811]" gridcol="1" gridrow="10" pageId="8" pageNumber="435">Gradually expanded from its base to apex (Figs. 14, 18)</td>
|
||
<td id="EF055DD6737A9890FDE0F93F3E56F8DE" box="[615,870,1698,1811]" gridcol="2" gridrow="10" pageId="8" pageNumber="435">
|
||
Gradually expanded from its base to apex (
|
||
<bibRefCitation id="76754B19737A6767FDF3F9403E1BF938" author="Zhu, M. S. & Lian, W. G." box="[628,811,1757,1781]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian 2006</bibRefCitation>
|
||
: figs. 10)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8F93F394AF8DE" box="[895,1146,1698,1811]" gridcol="3" gridrow="10" pageId="8" pageNumber="435">
|
||
Sharply expanded in its distal portion (
|
||
<bibRefCitation id="76754B19737A6767FBA5F9533EE4F8C9" author="Suzuki, S." pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: figs. 8 A–B)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12F93F38A0F8DE" box="[1173,1424,1698,1811]" gridcol="4" gridrow="10" pageId="8" pageNumber="435">–</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24F8BE38A0F85E" box="[163,1424,1827,1939]" gridrow="11" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24F8BE3C08F85E" box="[163,312,1827,1939]" gridcol="0" gridrow="11" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24F8DD3DEBF8B8" bold="true" pageId="8" pageNumber="435">Distal part of penis</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4F8BE3F7EF85E" box="[339,590,1827,1939]" gridcol="1" gridrow="11" pageId="8" pageNumber="435">Not trapeziform, with truncate apex (Figs. 11, 15, 13, 17)</td>
|
||
<td id="EF055DD6737A9890FDE0F8BE3E56F85E" box="[615,870,1827,1939]" gridcol="2" gridrow="11" pageId="8" pageNumber="435">
|
||
Clearly trapeziform, with tapering prominent apex (
|
||
<bibRefCitation id="76754B19737A6767FD2EF8C03E6FF8B8" author="Zhu, M. S. & Lian, W. G." box="[681,863,1885,1909]" pageId="8" pageNumber="435" pagination="63 - 68" refId="ref4050" refString="Zhu, M. S. & Lian, W. G. (2006) First record of the genus Euepedanus from China, with the description of a new species (Opiliones: Laniatores: Epedanidae). Zootaxa, 1367 (1), 63 - 68. https: // doi. org / 10.11646 / zootaxa. 1367.1.4" type="journal article" year="2006">Zhu & Lian 2006</bibRefCitation>
|
||
: figs. 7–10)
|
||
</td>
|
||
<td id="EF055DD6737A9890FCF8F8BE394AF85E" box="[895,1146,1827,1939]" gridcol="3" gridrow="11" pageId="8" pageNumber="435">
|
||
Clearly trapeziform, with truncate apex (
|
||
<bibRefCitation id="76754B19737A6767FC1EF8C0392DF8B8" author="Suzuki, S." box="[921,1053,1885,1909]" pageId="8" pageNumber="435" pagination="79 - 101" refId="ref3876" refString="Suzuki, S. (1969) Some phalangids from Thailand. Journal of Science of the Hiroshima University, Series B, Division 1 (Zoology), 22 (3), 79 - 101." type="journal article" year="1969">Suzuki 1969</bibRefCitation>
|
||
: figs. 8 A–C)
|
||
</td>
|
||
<td id="EF055DD6737A9890FB12F8BE38A0F85E" box="[1173,1424,1827,1939]" gridcol="4" gridrow="11" pageId="8" pageNumber="435">–</td>
|
||
</tr>
|
||
<tr id="ACD434AA737A9890FF24F83F38A0F876" box="[163,1424,1954,1979]" gridrow="12" pageId="8" pageNumber="435">
|
||
<th id="EF055DD6737A9890FF24F83F3C08F876" box="[163,312,1954,1979]" gridcol="0" gridrow="12" pageId="8" pageNumber="435">
|
||
<emphasis id="2090EAFA737A6767FF24F83F3C17F877" bold="true" box="[163,295,1954,1978]" pageId="8" pageNumber="435">Distribution</emphasis>
|
||
</th>
|
||
<td id="EF055DD6737A9890FED4F83F3F7EF876" box="[339,590,1954,1979]" gridcol="1" gridrow="12" pageId="8" pageNumber="435">Vietnam</td>
|
||
<td id="EF055DD6737A9890FDE0F83F3E56F876" box="[615,870,1954,1979]" gridcol="2" gridrow="12" pageId="8" pageNumber="435">China: Hainan Island</td>
|
||
<td id="EF055DD6737A9890FCF8F83F394AF876" box="[895,1146,1954,1979]" gridcol="3" gridrow="12" pageId="8" pageNumber="435">Thailand</td>
|
||
<td id="EF055DD6737A9890FB12F83F38A0F876" box="[1173,1424,1954,1979]" gridcol="4" gridrow="12" pageId="8" pageNumber="435">Singapore and Malaysia</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="125B36E8737B6766FF40FE463CFEFC68" blockId="9.[151,1437,151,969]" pageId="9" pageNumber="436">
|
||
FEMALE.
|
||
<emphasis id="2090EAFA737B6766FEC1FE463CD8FE39" box="[326,488,475,500]" italics="true" pageId="9" pageNumber="436">Measurements</emphasis>
|
||
. Body: length 5.50; maximal width 4.11. Scutum length 4.42; width in middle 4.12. ‘Clypeus’ length: 0.60. Eye tubercle width 0.97, height (from the base of eye tubercle to its top) 1.47. Chelicera: basal segment 1.42 long; distal segment 1.48 long; chela 1.22 long. Length of palp segments: 0.60 + 2.91 + 1.48 + 1.70 + 1.58 + claw 1.68 = 9.95. Length of leg segments: I: 4.36 + 0.91 + 2.98 + 5.47 + 1.98 = 15.70; II: 7.44 + 1.12 + 6.35 + 9.82 + 3.85 = 28.58; III: 5.68 + 1.28 + 3.10 + 6.82 + 2.73 = 19.61; IV: 7.95 + 1.27 + 4.07 + 9.36 + 3.43 = 26.08. Female general appearance as in the male, but body trapezoid and smaller. Chelicerae smaller, not swollen, basal segment without any apophysis-like structure as in the male. Females darker than the males, their prevailing colour—dirty yellowish, on the dorsal surface with a pattern resembling the Latin letter X. Carapace region, in front and eye tubercle, with a dark brown reticulation. Opistosomal region and each lateral margin of the scutum with the similar reticulation and dark brown spots and patches. Dome-shaped humps yellowish, situated on both sides below eye tubercle. Chelicerae and palps yellowish, with the same reticulation. Distal cheliceral segment with a zebra-pattern. Legs yellowish, with an ochre-brown reticulation, distal segments lighter, grey-milky. Venter yellowish, darkest at its distal end.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5AFE6563737B6766FF40FC333E02FC04" box="[199,818,942,969]" pageId="9" pageNumber="436" type="distribution">
|
||
<paragraph id="125B36E8737B6766FF40FC333E02FC04" blockId="9.[151,1437,151,969]" box="[199,818,942,969]" pageId="9" pageNumber="436">
|
||
<emphasis id="2090EAFA737B6766FF40FC333C69FC05" bold="true" box="[199,345,942,968]" pageId="9" pageNumber="436">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="6AF37678737B6766FEE0FC323CFAFC04" box="[359,458,943,969]" name="Vietnam" pageId="9" pageNumber="436">Vietnam</collectingCountry>
|
||
: the
|
||
<typeStatus id="CD5F884A737B6766FE79FC323F1EFC04" box="[510,558,943,969]" pageId="9" pageNumber="436">type</typeStatus>
|
||
locality only (
|
||
<figureCitation id="8ADF2A6D737B6766FD55FC323E16FC04" box="[722,806,943,969]" captionStart="FIGURE 28" captionStartId="7.[151,255,1818,1842]" captionTargetBox="[386,1206,188,1792]" captionTargetId="figure-15@7.[380,1207,181,1794]" captionTargetPageId="7" captionText="FIGURE 28. The type and paratype locality of Euepedanus vietnamicus sp. nov. in Vietnam. Scale bar 200 km." figureDoi="http://doi.org/10.5281/zenodo.4412551" httpUri="https://zenodo.org/record/4412551/files/figure.png" pageId="9" pageNumber="436">Fig. 28</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |