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<document id="A5AF53BBA4F0003635059C4A71204C01" ID-CLB-Dataset="38998" ID-DOI="10.11646/zootaxa.4114.4.5" ID-GBIF-Dataset="e23a4c9f-0c59-4962-bd1d-dae0e1f0f22a" ID-ISSN="1175-5326" ID-Zenodo-Dep="258082" ID-ZooBank="13E78B18-DDD8-47EF-B4E4-2DE9608E1F8D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1464159392924" checkinUser="plazi" docAuthor="Bayarsaikhan, Ulziijargal &amp; Bae, Yang-Seop" docDate="2016" docId="806905426E6BFFD865CEC8C28E6BFC27" docLanguage="en" docName="zootaxa.4114.4.5.pdf.imf" docOrigin="Zootaxa 4114 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyana perornata Walker 1854" docType="treatment" docVersion="9" lastPageNumber="452" masterDocId="7C507D3A6E6EFFDD6559C8558A6CFFC8" masterDocTitle="A review of the genus Cyana Walker, 1854 (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with description of new species" masterLastPageNumber="463" masterPageNumber="447" pageNumber="452" updateTime="1698662618307" updateUser="ExternalLinkService">
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<mods:title id="A83CF26D2780033E3B9CBD79F4BA0A1C">A review of the genus Cyana Walker, 1854 (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with description of new species</mods:title>
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<mods:namePart id="BFA88AD915DBF83A43DC27959221EA68">Bayarsaikhan, Ulziijargal</mods:namePart>
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<mods:namePart id="862375401D5F5EA4C4A9159EC8D1082F">Bae, Yang-Seop</mods:namePart>
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<mods:date id="7A7583F59A5C60A7331F09673713FC07">2016</mods:date>
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<treatment id="806905426E6BFFD865CEC8C28E6BFC27" ID-DOI="http://doi.org/10.5281/zenodo.6069418" ID-GBIF-Taxon="120701455" ID-Zenodo-Dep="6069418" LSID="urn:lsid:plazi:treatment:806905426E6BFFD865CEC8C28E6BFC27" httpUri="http://treatment.plazi.org/id/806905426E6BFFD865CEC8C28E6BFC27" lastPageNumber="452" pageId="5" pageNumber="452">
<subSubSection id="40DAE7DF6E6BFFD865CEC8C28BA7FF1B" pageId="5" pageNumber="452" type="nomenclature">
<paragraph id="087FB4546E6BFFD865CEC8C2884DFF7A" blockId="5.[151,545,151,211]" box="[151,545,151,178]" pageId="5" pageNumber="452">
<heading id="533703386E6BFFD865CEC8C2884DFF7A" bold="true" box="[151,545,151,178]" fontSize="11" level="1" pageId="5" pageNumber="452" reason="1">
<taxonomicName id="CFC0CFD76E6BFFD865CEC8C2884DFF7A" authority="Walker, 1854" authorityName="Walker" authorityYear="1854" box="[151,545,151,178]" class="Insecta" family="Arctiidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="perornata">
<emphasis id="3AB468466E6BFFD865CEC8C2884DFF7A" bold="true" box="[151,545,151,178]" pageId="5" pageNumber="452">
<emphasis id="3AB468466E6BFFD865CEC8C28B0EFF79" bold="true" box="[151,354,151,177]" italics="true" pageId="5" pageNumber="452">Cyana perornata</emphasis>
(Walker, 1854)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="087FB4546E6BFFD865CEC8EF8BA7FF1B" blockId="5.[151,545,151,211]" box="[151,459,186,211]" pageId="5" pageNumber="452">(Pl. 1: 4a, 4b, 4c; 3: 4; 5: 4)</paragraph>
</subSubSection>
<subSubSection id="40DAE7DF6E6BFFD865CEC9568B98FE7A" pageId="5" pageNumber="452" type="reference_group">
<paragraph id="087FB4546E6BFFD865CEC9568B57FEBC" blockId="5.[151,1436,258,434]" pageId="5" pageNumber="452">
<taxonomicName id="CFC0CFD76E6BFFD865CEC9568BB9FED0" authority="Walker, 1854" authorityName="Walker" authorityYear="1854" box="[151,469,258,280]" class="Insecta" family="Erebidae" genus="Bizone" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="perornata">
<emphasis id="3AB468466E6BFFD865CEC9568B2BFED0" box="[151,327,259,280]" italics="true" pageId="5" pageNumber="452">Bizone perornata</emphasis>
Walker, 1854
</taxonomicName>
,
<emphasis id="3AB468466E6BFFD864B9C9568911FED0" box="[480,893,258,280]" italics="true" pageId="5" pageNumber="452">List. Spec. Lepid. Insects Colln. Br. Mus.</emphasis>
2: 548. TL: NE
<collectingCountry id="70D7F4C46E6BFFD8617AC9578EF5FED0" box="[1059,1177,258,280]" name="Bangladesh" pageId="5" pageNumber="452">Bangladesh</collectingCountry>
(Silhet).
<taxonomicName id="CFC0CFD76E6BFFD865CEC9778B9FFEFF" authority="Hampson, 1894" authorityName="Hampson" authorityYear="1894" box="[151,499,289,311]" class="Insecta" family="Arctiidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="perornata">
<emphasis id="3AB468466E6BFFD865CEC9778B2FFEFF" box="[151,323,290,311]" italics="true" pageId="5" pageNumber="452">Cyana perornata</emphasis>
: Hampson, 1894
</taxonomicName>
,
<emphasis id="3AB468466E6BFFD864A7C974881FFEFF" box="[510,627,289,311]" italics="true" pageId="5" pageNumber="452">
Moths.
<collectingCountry id="70D7F4C46E6BFFD86713C9748803FEFF" box="[586,623,289,311]" name="India" pageId="5" pageNumber="452">Ind</collectingCountry>
.
</emphasis>
2: 57; Fang, 1992, 9: 258; Kishida, 1994, 14 (1): 70, pl. 79: 18, 19;
<bibRefCitation id="6C51C9A56E6BFFD8607AC9778FFBFEFF" author="Fang" box="[1315,1431,289,311]" pageId="5" pageNumber="452" refString="Fang, C. (2000) Fauna Sinica Insecta, Vol. 19 (Lepidoptera: Arctiidae). Science Press, Beijing, China, 589 pp." type="book" year="2000">Fang, 2000</bibRefCitation>
, 19: 163, pl. 4: 18;
<bibRefCitation id="6C51C9A56E6BFFD864DBC915884AFE9E" author="Holloway" box="[386,550,320,342]" pageId="5" pageNumber="452" refString="Holloway, J. D. (2001) The Moths of Borneo: Family Arctiidae, Subfamily Lithosiinae. Malayan Nature Journal, 7, 279 - 486." type="journal article" year="2001">Holloway, 2001</bibRefCitation>
, 7: 332, pl. 1: 135, 140;
<bibRefCitation id="6C51C9A56E6BFFD86646C9158E73FE9E" author="Cerny" box="[799,1055,320,342]" pageId="5" pageNumber="452" refString="Cerny, K. &amp; Pinratana, A. (2009) Moths of Thailand, Vol. 6, Arctiidae. Brothers of Saint Gabriel in Thailand, Bangkok, Thailand, 283 pp." type="book" year="2009">Černy &amp; Pinratana, 2009</bibRefCitation>
, 6: 51, pl. 10: 103;
<bibRefCitation id="6C51C9A56E6BFFD861BCC9158F1FFE9E" author="Bucsek" box="[1253,1395,320,342]" pageId="5" pageNumber="452" refString="Bucsek, K. (2012) Erebidae, Arctiinae (Lithosiini, Arctiini) of Malaya Peninsula-Malaysia. Institute of Zoology SAS, Bratislava, 170 pp." type="book" year="2012">Bucsek, 2012</bibRefCitation>
: 98, pl. 17: 244.
</paragraph>
<paragraph id="087FB4546E6BFFD865CEC9288B98FE7A" blockId="5.[151,1436,258,434]" pageId="5" pageNumber="452">
<taxonomicName id="CFC0CFD76E6BFFD865CEC9288846FE5B" authority="Hampson, 1900" authorityName="Hampson" authorityYear="1900" box="[151,554,381,403]" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="perornata">
<emphasis id="3AB468466E6BFFD865CEC9288B1BFE5B" box="[151,375,381,403]" italics="true" pageId="5" pageNumber="452">Chionaema perornata</emphasis>
:
<bibRefCitation id="6C51C9A56E6BFFD864DFC92B8846FE5B" author="Hampson" box="[390,554,381,403]" pageId="5" pageNumber="452" refString="Hampson, G. F. (1900) Catalogue of the Arctiadae (Nolinae, Lithosinae) in the collection of the British Museum. Catalogue of the Lepidoptera Phalaenae in the British Museum, 2, 590 pp." type="book" year="1900">Hampson, 1900</bibRefCitation>
</taxonomicName>
,
<emphasis id="3AB468466E6BFFD8676CC92B895DFE5B" box="[565,817,381,403]" italics="true" pageId="5" pageNumber="452">Cat. Lep. Phal. Br. Mus.</emphasis>
2: 308; Draudt, 1914, 10: 171, pl. 16: i; Holloway, 1976: 3; Fang, 1982, 2: 203, fig. 1491.
</paragraph>
</subSubSection>
<subSubSection id="40DAE7DF6E6BFFD865CEC98B88F8FD07" pageId="5" pageNumber="452" type="diagnosis">
<paragraph id="087FB4546E6BFFD865CEC98B88F8FD07" blockId="5.[151,1436,478,1007]" pageId="5" pageNumber="452">
<emphasis id="3AB468466E6BFFD865CEC98B8B7DFE3F" bold="true" box="[151,273,478,503]" pageId="5" pageNumber="452">Diagnosis.</emphasis>
Wingspan ♂
<quantity id="CF3819B16E6BFFD864F7C98A8847FE3E" box="[430,555,479,503]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" metricValueMax="4.4" metricValueMin="3.7" pageId="5" pageNumber="452" unit="mm" value="40.5" valueMax="44.0" valueMin="37.0">3744 mm</quantity>
, ♀
<quantity id="CF3819B16E6BFFD8670DC98B88BDFE3E" box="[596,721,478,503]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.3" metricValueMin="4.8" pageId="5" pageNumber="452" unit="mm" value="50.5" valueMax="53.0" valueMin="48.0">4853 mm</quantity>
. This species is the largest of all
<taxonomicName id="CFC0CFD76E6BFFD86113C98A8EFFFE3E" box="[1098,1171,479,502]" class="Insecta" family="Arctiidae" genus="Cyana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="genus">
<emphasis id="3AB468466E6BFFD86113C98A8EFFFE3E" box="[1098,1171,479,502]" italics="true" pageId="5" pageNumber="452">Cyana</emphasis>
</taxonomicName>
species which occur in
<collectingCountry id="70D7F4C46E6BFFD865CECA578B62FDD3" box="[151,270,514,539]" name="Cambodia" pageId="5" pageNumber="452">Cambodia</collectingCountry>
.
<taxonomicName id="CFC0CFD76E6BFFD86442CA568BC1FDD2" box="[283,429,515,538]" class="Insecta" family="Arctiidae" genus="Chionaema" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="452" phylum="Arthropoda" rank="species" species="perornata">
<emphasis id="3AB468466E6BFFD86442CA568BC1FDD2" box="[283,429,515,538]" italics="true" pageId="5" pageNumber="452">C. perornata</emphasis>
</taxonomicName>
differs from congeners by having the forewing of both sexes with dark edged crimson lines; one dark, elliptical discoidal spot; terminal line slightly paler red, zig-zag shaped; forewing of some males with dark or pale red band before apex; in male genitalia, valvae bowling pin-shaped, with more rectangular apex, upper angle of apex rounded, lower angle of apex angled; saccular process tapered, and curved to acute apex, almost the same length of valvae; vesica with scobination; in female genitalia, corpus bursae globular, with one large, elliptical and one small, rounded signa.
</paragraph>
</subSubSection>
<subSubSection id="40DAE7DF6E6BFFD8659ECA8F8F55FC4B" pageId="5" pageNumber="452" type="materials_examined">
<paragraph id="087FB4546E6BFFD8659ECA8F8F55FC4B" blockId="5.[151,1436,478,1007]" pageId="5" pageNumber="452">
<emphasis id="3AB468466E6BFFD8659ECA8F8BDDFD3B" bold="true" box="[199,433,730,755]" pageId="5" pageNumber="452">Material examined.</emphasis>
<specimenCount id="1EC67FDD6E6BFFD864E0CA8E8BB1FD3B" box="[441,477,731,755]" pageId="5" pageNumber="452" type="generic">1♂</specimenCount>
, 3♀,
<collectingCountry id="70D7F4C46E6BFFD86744CA8F88FFFD3B" box="[541,659,730,755]" name="Cambodia" pageId="5" pageNumber="452">Cambodia</collectingCountry>
, Cardamom (N11˚5847″, E103˚2222″),
<date id="7C7E92946E6BFFD8613DCA8E8F61FD3B" box="[1124,1293,731,755]" pageId="5" pageNumber="452" value="2011-06-28" valueMax="2011-06-29" valueMin="2011-06-28">2829.VI.2011</date>
(Y.S. Bae
<emphasis id="3AB468466E6BFFD860D1CA8E8AD8FCDE" italics="true" pageId="5" pageNumber="452">et al.</emphasis>
), Gen. Slide No. INU-1067, INU-1069;
<specimenCount id="1EC67FDD6E6BFFD86722CAAA88CCFCDF" box="[635,672,767,791]" pageId="5" pageNumber="452" type="generic">1♂</specimenCount>
,
<collectingCountry id="70D7F4C46E6BFFD867F5CAAB894EFCDF" box="[684,802,766,791]" name="Cambodia" pageId="5" pageNumber="452">Cambodia</collectingCountry>
, Koh Kong (N11˚3504″, E103˚1321″),
<date id="7C7E92946E6BFFD861B0CAAA8F32FCDF" box="[1257,1374,767,791]" pageId="5" pageNumber="452" value="2012-02-22">22.II.2012</date>
(Y.S. Bae
<emphasis id="3AB468466E6BFFD86594CB768B65FCF3" box="[205,265,803,827]" italics="true" pageId="5" pageNumber="452">et al.</emphasis>
), Gen. Slide No. INU-1068;
<specimenCount id="1EC67FDD6E6BFFD8673DCB7688E5FCF3" box="[612,649,803,827]" pageId="5" pageNumber="452" type="generic">3♂</specimenCount>
,
<collectingCountry id="70D7F4C46E6BFFD867C0CB778963FCF3" box="[665,783,802,827]" name="Cambodia" pageId="5" pageNumber="452">Cambodia</collectingCountry>
, Koh Kong (N11˚4144.9″, E103˚0651.8″),
<date id="7C7E92946E6BFFD8604ACB778FF0FCF3" box="[1299,1436,802,827]" pageId="5" pageNumber="452" value="2014-12-25">25.XII.2014</date>
(Y.S. Bae
<emphasis id="3AB468466E6BFFD86454CB128B24FC96" box="[269,328,838,862]" italics="true" pageId="5" pageNumber="452">et al.</emphasis>
);
<specimenCount id="1EC67FDD6E6BFFD8643ACB128BEBFC97" box="[355,391,839,863]" pageId="5" pageNumber="452" type="generic">1♂</specimenCount>
,
<collectingCountry id="70D7F4C46E6BFFD864CFCB138860FC97" box="[406,524,838,863]" name="Cambodia" pageId="5" pageNumber="452">Cambodia</collectingCountry>
, Samkos (N12˚1240.43″, E102˚5342.38″),
<date id="7C7E92946E6BFFD86151CB128EFDFC97" box="[1032,1169,839,863]" pageId="5" pageNumber="452" value="2012-07-20">20.VII.2012</date>
(Y.S. Bae
<emphasis id="3AB468466E6BFFD86048CB128F20FC96" box="[1297,1356,838,862]" italics="true" pageId="5" pageNumber="452">et al.</emphasis>
), Gen. Slide No. INU-1066;
<specimenCount id="1EC67FDD6E6BFFD864D0CB3E8BC2FC4B" box="[393,430,875,899]" pageId="5" pageNumber="452" type="generic">1♂</specimenCount>
,
<collectingCountry id="70D7F4C46E6BFFD864E3CB3F885CFC4B" box="[442,560,874,899]" name="Cambodia" pageId="5" pageNumber="452">Cambodia</collectingCountry>
, Seima (N12˚1829″, E107˚0454.80″),
<date id="7C7E92946E6BFFD866B2CB3E8E3AFC4B" box="[1003,1110,875,899]" pageId="5" pageNumber="452" value="2013-01-11">11.I.2013</date>
(I) (Y.S. Bae
<emphasis id="3AB468466E6BFFD861A8CB3E8F45FC4B" box="[1265,1321,875,899]" italics="true" pageId="5" pageNumber="452">et al.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="40DAE7DF6E6BFFD8659ECBDB894FFC03" pageId="5" pageNumber="452" type="distribution">
<paragraph id="087FB4546E6BFFD8659ECBDB894FFC03" blockId="5.[151,1436,478,1007]" pageId="5" pageNumber="452">
<emphasis id="3AB468466E6BFFD8659ECBDB8B0CFC6F" bold="true" box="[199,352,910,935]" pageId="5" pageNumber="452">Distribution.</emphasis>
<collectingCountry id="70D7F4C46E6BFFD86428CBDB8B88FC6F" box="[369,484,910,935]" name="Cambodia" pageId="5" pageNumber="452">Cambodia</collectingCountry>
(Cardamom, Koh Kong, Samkos, Seima), N.E. Himalaya,
<collectingCountry id="70D7F4C46E6BFFD86192CBDB8F7DFC6F" box="[1227,1297,910,935]" name="Nepal" pageId="5" pageNumber="452">Nepal</collectingCountry>
, Sundaland (Borneo),
<collectingCountry id="70D7F4C46E6BFFD86453CBE78B1CFC03" box="[266,368,946,971]" name="Thailand" pageId="5" pageNumber="452">Thailand</collectingCountry>
, peninsular
<collectingCountry id="70D7F4C46E6BFFD864AECBE7880CFC03" box="[503,608,946,971]" name="Malaysia" pageId="5" pageNumber="452">Malaysia</collectingCountry>
,
<collectingCountry id="70D7F4C46E6BFFD86735CBE788A2FC03" box="[620,718,946,971]" name="Vietnam" pageId="5" pageNumber="452">Vietnam</collectingCountry>
,
<collectingCountry id="70D7F4C46E6BFFD86781CBE78973FC03" box="[728,799,946,971]" name="China" pageId="5" pageNumber="452">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="40DAE7DF6E6BFFD8659ECB838E6BFC27" box="[199,1031,982,1007]" pageId="5" pageNumber="452" type="discussion">
<paragraph id="087FB4546E6BFFD8659ECB838E6BFC27" blockId="5.[151,1436,478,1007]" box="[199,1031,982,1007]" pageId="5" pageNumber="452">
<emphasis id="3AB468466E6BFFD8659ECB838B57FC27" bold="true" box="[199,315,982,1007]" pageId="5" pageNumber="452">Remarks.</emphasis>
This species was reported from
<collectingCountry id="70D7F4C46E6BFFD867FFCB838975FC27" box="[678,793,982,1007]" name="Cambodia" pageId="5" pageNumber="452">Cambodia</collectingCountry>
by
<bibRefCitation id="6C51C9A56E6BFFD8661DCB828E6FFC27" box="[836,1027,982,1007]" pageId="5" pageNumber="452" refString="Bae, Y. S., Ahn, N. H., Chu, Y. D. &amp; Lee, H. (2012) Lepidoptera. In: Lee, W. S., Rhim, S. J., Choe, J. I. &amp; Park, S. (Eds), Biodiversity of Cambodia - Cardamom Protected Forest &amp; Seima Biodiversity Conservation Area. National Institute of Biological Resources and Incheon. pp. 87 - 92." type="book chapter">
Bae
<emphasis id="3AB468466E6BFFD8662FCB8289C2FC26" box="[886,942,982,1006]" italics="true" pageId="5" pageNumber="452">et al.</emphasis>
(2012)
</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>