314 lines
38 KiB
XML
314 lines
38 KiB
XML
<document id="FBB218BDD309C74A0E3F6456EC898C73" ID-DOI="10.5281/zenodo.170940" ID-GBIF-Dataset="60bedb06-0856-4fd2-b1ce-c8061fd663cf" ID-ISSN="11755326" ID-Zenodo-Dep="170940" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459717131927" checkinUser="plazi" docAuthor="Miralles, Aurélien, Rivas, Gilson & Schargel, Walter E." docDate="2005" docId="563DE827D835FFB6FEF7FEBBFE3EFDB1" docLanguage="en" docName="zt00895.pdf" docOrigin="Zootaxa 895" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Mabuya meridensis Miralles, Rivas & Schargel, 2005, new species" docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="AA04905FD837FFB1FFFFFFA7FFA5FFC7" masterDocTitle="A new species of Mabuya (Squamata, Scincidae) from the Venezuelan Andes" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1698208181939" updateUser="plazi">
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<mods:mods id="C46BA459BC3145F4A4B6374BF4169D22" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1F83185D024F59B2F8E94F6C31A8CF35">
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<mods:title id="497E6F00AAB10A206FA51E6282B49B11">A new species of Mabuya (Squamata, Scincidae) from the Venezuelan Andes</mods:title>
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</mods:titleInfo>
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<mods:name id="AC301C037C99A61BE7847B24C984C388" type="personal">
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<mods:role id="819F67AA379F69932C2800070BFB09D2">
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<mods:roleTerm id="CBBCE7D8AC47F32FB9F4EA2C1D24F0F4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8D8030DFC5E8BAA8AE665B0A6965ADB4">Miralles, Aurélien</mods:namePart>
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<mods:name id="6B49B7DE904B32FA4EA28103EC5B3E45" type="personal">
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<mods:roleTerm id="0AA8E205C5295FC502A5E8E7E930616D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3BEC0155A865D77AB33F658D52C48B1D">Rivas, Gilson</mods:namePart>
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</mods:name>
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<mods:name id="DF4A6B0603671364E6934DCD2BEC4C0D" type="personal">
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<mods:role id="1B32C04B95C67500BE66494CD46A0BAA">
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<mods:roleTerm id="C77AC6FF586AB9D7F4D7C05214E2059D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1D6D9EB2C9E3354EB059D5194E636FF9">Schargel, Walter E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="CAF33EDB9ED4005BD6A444BD70F3A437">text</mods:typeOfResource>
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<mods:titleInfo id="25C7B374FB4F61DA0478D7FAF5B25510">
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<mods:title id="4C020866B6C3298F5343848D97B09FB0">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1A98B77E62C6F1C14F31F39F6A55A587">
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<mods:date id="844FDDA6F4784C65BB728E3FE8BC4DC4">2005</mods:date>
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<mods:detail id="092639386F1EC6371EBF59F612457DE6" type="volume">
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<mods:number id="53D557D50D76AC7CD971E664C00046A5">895</mods:number>
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<mods:classification id="C9176A2878E9344DE9B039ACA59BA576">journal article</mods:classification>
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<mods:identifier id="488F44CB1A62EF22A7CF47704949BFF8" type="DOI">10.5281/zenodo.170940</mods:identifier>
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<mods:identifier id="97D8E04CAE815B26D34BD998B0C04CFA" type="GBIF-Dataset">60bedb06-0856-4fd2-b1ce-c8061fd663cf</mods:identifier>
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<mods:identifier id="2DFD8C33828821B7F4F7B72EEAE51C22" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="D2DADB1E27EEC950B2A41FF7B7C60543" type="Zenodo-Dep">170940</mods:identifier>
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<treatment id="563DE827D835FFB6FEF7FEBBFE3EFDB1" ID-DOI="http://doi.org/10.5281/zenodo.6268770" ID-GBIF-Taxon="119334838" ID-Zenodo-Dep="6268770" LSID="urn:lsid:plazi:treatment:563DE827D835FFB6FEF7FEBBFE3EFDB1" httpUri="http://treatment.plazi.org/id/563DE827D835FFB6FEF7FEBBFE3EFDB1" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3">
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<subSubSection id="968E0ABAD835FFB3FEF7FEBBFD36FEF2" box="[264,659,283,310]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="DE2B5931D835FFB3FEF7FEBBFD36FEF2" blockId="2.[264,659,283,310]" box="[264,659,283,310]" pageId="2" pageNumber="3">
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<heading id="8563EE5DD835FFB3FEF7FEBBFD36FEF2" bold="true" box="[264,659,283,310]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
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||
<emphasis id="ECE08523D835FFB3FEF7FEBBFD36FEF2" bold="true" box="[264,659,283,310]" pageId="2" pageNumber="3">
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||
<emphasis id="ECE08523D835FFB3FEF7FEBBFE59FEF1" bold="true" box="[264,508,284,310]" italics="true" pageId="2" pageNumber="3">
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<taxonomicName id="199422B2D835FFB3FEF7FEBBFE50FEF1" box="[264,501,284,310]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="meridensis" status="sp. nov.">Mabuya meridensis</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="F7D33858D835FFB3FDFBFEBCFD36FEF2" box="[516,659,283,309]" pageId="2" pageNumber="3" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="968E0ABAD835FFB3FEF7FEC1FD86FD5E" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="DE2B5931D835FFB3FEF7FEC1FD03FE16" blockId="2.[264,1324,358,1905]" pageId="2" pageNumber="3">
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<emphasis id="ECE08523D835FFB3FEF7FEC1FE26FE47" bold="true" box="[264,387,358,384]" pageId="2" pageNumber="3">
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<typeStatus id="012FE793D835FFB3FEF7FEC1FEDBFE47" box="[264,382,358,384]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectionCode id="B885C1F4D835FFB3FE74FEC0FE55FE46" box="[395,496,359,385]" httpUri="http://grbio.org/cool/fxnk-ibav" name="Universidad de los Andes, Laboratorio de Biogeografia" pageId="2" pageNumber="3">ULABG</collectionCode>
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1570, an adult male. Typelocality: City of Mérida, Mérida state,
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<collectingCountry id="A68319A1D835FFB3FB0CFEC0FEF7FE6E" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
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(Ciudad de Mérida, Estado Mérida,
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<collectingCountry id="A68319A1D835FFB3FCFBFE28FC24FE6E" box="[772,897,399,425]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
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), collected
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<date id="AA2A7FF1D835FFB3FBF9FE28FB3AFE6E" box="[1030,1183,399,425]" pageId="2" pageNumber="3" value="1985-07-10">10 July 1985</date>
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by Enrique La Marca, at an altitude of
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<quantity id="196CF4D4D835FFB3FDB9FE10FD3AFE16" box="[582,671,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="2" pageNumber="3" unit="m" value="1750.0">1750 m</quantity>
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.
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</paragraph>
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<paragraph id="DE2B5931D835FFB3FEC7FE79FD86FD5E" blockId="2.[264,1324,358,1905]" pageId="2" pageNumber="3">
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<emphasis id="ECE08523D835FFB3FEC7FE79FE1BFE3F" bold="true" box="[312,446,478,504]" pageId="2" pageNumber="3">
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<typeStatus id="012FE793D835FFB3FEC7FE79FE1FFE3F" box="[312,442,478,504]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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Nine specimens.
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<collectingCountry id="A68319A1D835FFB3FD6BFE78FCAAFE3E" box="[660,783,479,505]" name="Venezuela" pageId="2" pageNumber="3">Venezuela</collectingCountry>
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, Mérida state:
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<collectionCode id="B885C1F4D835FFB3FC3EFE78FBBEFE3E" LSID="urn:lsid:biocol.org:col:34925" box="[961,1051,479,505]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="3">AMNH</collectionCode>
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R–13405 to R–13408, Ríos Albarregas y Milla, all adults, collected by W. F. Rosenberg;
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<collectionCode id="B885C1F4D835FFB3FBAFFDA0FB0FFDE6" LSID="urn:lsid:biocol.org:col:34925" box="[1104,1194,519,545]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="2" pageNumber="3">AMNH</collectionCode>
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R–13526;
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<collectionCode id="B885C1F4D835FFB3FEF7FD88FEC8FD8E" box="[264,365,559,585]" httpUri="http://grbio.org/cool/fxnk-ibav" name="Universidad de los Andes, Laboratorio de Biogeografia" pageId="2" pageNumber="3">ULABG</collectionCode>
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4153, 4281, Urbanization of Santa Maria, City of Mérida,
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<quantity id="196CF4D4D835FFB3FBDBFD88FBD8FD8E" box="[1060,1149,559,585]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="2" pageNumber="3" unit="m" value="1750.0">1750 m</quantity>
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alt., all adults, collected in 1996 by E. La Marca;
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<collectionCode id="B885C1F4D835FFB3FD50FDF0FCAFFDB6" box="[687,778,599,625]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="2" pageNumber="3">UMMZ</collectionCode>
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57435, City of Mérida, juvenile, collected in 1922 by M. A. Carriker.
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</paragraph>
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</subSubSection>
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<subSubSection id="968E0ABAD835FFB3FEC7FD01FED0FCA6" pageId="2" pageNumber="3" type="diagnosis">
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||
<paragraph id="DE2B5931D835FFB3FEC7FD01FED0FCA6" blockId="2.[264,1324,358,1905]" pageId="2" pageNumber="3">
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<emphasis id="ECE08523D835FFB3FEC7FD01FE1FFD07" bold="true" box="[312,442,678,704]" pageId="2" pageNumber="3">Diagnosis:</emphasis>
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A small to medium sized
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<taxonomicName id="199422B2D835FFB3FCF3FD00FCC9FD07" box="[780,876,679,704]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
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<emphasis id="ECE08523D835FFB3FCF3FD00FCC9FD07" box="[780,876,679,704]" italics="true" pageId="2" pageNumber="3">Mabuya</emphasis>
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</taxonomicName>
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with an undivided lower eyelid, all scales smooth, auricular lobes absent, and having: a short and acute snout, paired prefrontals and frontoparietals, a single pair of nuchals, four supraoculars, four supraciliaries with the second longest, seven dark and four white/pale stripes along body, and light cream colored lips.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="968E0ABAD835FFB3FEC7FCC8FB7AFAAE" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="DE2B5931D835FFB3FEC7FCC8FB7AFAAE" blockId="2.[264,1324,358,1905]" pageId="2" pageNumber="3">
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Within
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<taxonomicName id="199422B2D835FFB3FE6EFCC8FE54FC4F" box="[401,497,879,904]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
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<emphasis id="ECE08523D835FFB3FE6EFCC8FE54FC4F" box="[401,497,879,904]" italics="true" pageId="2" pageNumber="3">Mabuya</emphasis>
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</taxonomicName>
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, only three species have seven longitudinal dark stripes on the body:
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<emphasis id="ECE08523D835FFB3FEF7FC30FD32FC77" box="[264,663,919,944]" italics="true" pageId="2" pageNumber="3">
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<taxonomicName id="199422B2D835FFB3FEF7FC30FE7DFC77" box="[264,472,919,944]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="cochabambae">M. cochabambae</taxonomicName>
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,
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<taxonomicName id="199422B2D835FFB3FE19FC30FD32FC77" box="[486,663,919,944]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="dorsivittata">M. dorsivittata</taxonomicName>
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</emphasis>
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and
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<taxonomicName id="199422B2D835FFB3FD2AFC30FC2BFC77" box="[725,910,919,944]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="guaporicola">
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<emphasis id="ECE08523D835FFB3FD2AFC30FC2BFC77" box="[725,910,919,944]" italics="true" pageId="2" pageNumber="3">M. guaporicola</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="199422B2D835FFB3FC60FC30FBEDFC77" box="[927,1096,919,944]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="meridensis">
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<emphasis id="ECE08523D835FFB3FC60FC30FBEDFC77" box="[927,1096,919,944]" italics="true" pageId="2" pageNumber="3">M. meridensis</emphasis>
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</taxonomicName>
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differs from all of them [data for the latter species in parentheses, after
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<bibRefCitation id="BA0524C0D835FFB3FC8DFC18FC52FC1E" author="Dunn" box="[882,1015,959,985]" pageId="2" pageNumber="10" refString="Dunn, E. R. (1936) Notes on American mabuyas. Proceedings of the Academy of Natural Sciences, Philadelphia, 87, 533 - 557." type="journal article" year="1936">Dunn 1936</bibRefCitation>
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,
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<bibRefCitation id="BA0524C0D835FFB3FBFCFC18FBCAFC1E" author="Cei" box="[1027,1135,959,985]" pageId="2" pageNumber="10" refString="Cei, J. M. (1993) Reptiles del noroeste, nordeste y este de la Argentina. Herpetofauna de las selvas subtropicales, Puna y Pampas. Monograpfia XIV. Museo Regionale di Scienze Naturali, Torino, 949 pp." type="book" year="1993">Cei 1993</bibRefCitation>
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and
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<bibRefCitation id="BA0524C0D835FFB3FB58FC18FEE0FBC6" author="Avila-Pires" pageId="2" pageNumber="9" refString="Avila-Pires, T. C. S. (1995) Lizards of Brazilian Amazonia (Reptilia: Squamata). Zoologische Verhandelingen (Leiden), 599, 1 - 706." type="journal article" year="1995">AvilaPires 1995</bibRefCitation>
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] in having reticulated and not well defined dorsolateral stripes, that widen posteriorly and fuse with each other and to the vertebral thin stripe on the posterior part of the body (vertebral and dorsolateral stripes sharp and well defined throughout all the length of the body). Additionally,
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<taxonomicName id="199422B2D835FFB3FE03FBF8FD06FBBF" box="[508,675,1119,1144]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="meridensis">
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<emphasis id="ECE08523D835FFB3FE03FBF8FD06FBBF" box="[508,675,1119,1144]" italics="true" pageId="2" pageNumber="3">M. meridensis</emphasis>
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</taxonomicName>
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differs from both
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<taxonomicName id="199422B2D835FFB3FC86FBF8FBE6FBBF" box="[889,1091,1119,1144]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="cochabambae">
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<emphasis id="ECE08523D835FFB3FC86FBF8FBE6FBBF" box="[889,1091,1119,1144]" italics="true" pageId="2" pageNumber="3">M. cochabambae</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="199422B2D835FFB3FB83FBF8FA8EFBBF" box="[1148,1323,1119,1144]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="dorsivittata">
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<emphasis id="ECE08523D835FFB3FB83FBF8FA8EFBBF" box="[1148,1323,1119,1144]" italics="true" pageId="2" pageNumber="3">M. dorsivittata</emphasis>
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</taxonomicName>
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in having four supraoculars (three); from both
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<taxonomicName id="199422B2D835FFB3FCD0FB20FC7BFB67" box="[815,990,1159,1184]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="dorsivittata">
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<emphasis id="ECE08523D835FFB3FCD0FB20FC7BFB67" box="[815,990,1159,1184]" italics="true" pageId="2" pageNumber="3">M. dorsivittata</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="199422B2D835FFB3FBE6FB20FB74FB67" box="[1049,1233,1159,1184]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="guaporicola">
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<emphasis id="ECE08523D835FFB3FBE6FB20FB74FB67" box="[1049,1233,1159,1184]" italics="true" pageId="2" pageNumber="3">M. guaporicola</emphasis>
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</taxonomicName>
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in having a relatively shorter body, fore and hindlimbs easily touching each other when adpressed against body (fore and hindlimbs never touch when adpressed to body); from
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<taxonomicName id="199422B2D835FFB3FEF7FB58FE77FADF" box="[264,466,1279,1304]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="cochabambae">
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<emphasis id="ECE08523D835FFB3FEF7FB58FE77FADF" box="[264,466,1279,1304]" italics="true" pageId="2" pageNumber="3">M. cochabambae</emphasis>
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</taxonomicName>
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in having two frontoparietals (a single one); and from
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<taxonomicName id="199422B2D835FFB3FBA9FB58FAA9FADF" box="[1110,1292,1279,1304]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="guaporicola">
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<emphasis id="ECE08523D835FFB3FBA9FB58FAA9FADF" box="[1110,1292,1279,1304]" italics="true" pageId="2" pageNumber="3">M. guaporicola</emphasis>
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</taxonomicName>
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in having 52 to 59 dorsal scales rows (65 to 68) and a whitish stripe on each side separating the dorsolateral from the dark upper lateral stripe (white intermediate stripe absent).
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</paragraph>
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</subSubSection>
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<subSubSection id="968E0ABAD835FFB4FEC7FAD1FA88FA79" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="DE2B5931D835FFB3FEC7FAD1FD8EFA26" blockId="2.[264,1324,358,1905]" pageId="2" pageNumber="3">
|
||
<emphasis id="ECE08523D835FFB3FEC7FAD1FD04FA57" bold="true" box="[312,673,1398,1424]" pageId="2" pageNumber="3">
|
||
Description of the
|
||
<typeStatus id="012FE793D835FFB3FDD3FAD1FD39FA57" box="[556,668,1398,1424]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Specimen (Plate 1a–b) in good condition, with long medioabdominal slit; hemipenes not extruded, testes white, enlarged, apparently sexually active at time of capture.
|
||
</paragraph>
|
||
<paragraph id="DE2B5931D835FFB3FEC7FA48FED8F99E" blockId="2.[264,1324,358,1905]" pageId="2" pageNumber="3">
|
||
Snoutvent length
|
||
<quantity id="196CF4D4D835FFB3FDEDFA48FD28F9CE" box="[530,653,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="6.648" pageId="2" pageNumber="3" unit="mm" value="66.48">66.48 mm</quantity>
|
||
; tail length
|
||
<quantity id="196CF4D4D835FFB3FCE5FA48FC30F9CE" box="[794,917,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="8.66" pageId="2" pageNumber="3" unit="mm" value="86.6">86.60 mm</quantity>
|
||
; head length
|
||
<quantity id="196CF4D4D835FFB3FBCCFA48FB0BF9CE" box="[1075,1198,1519,1545]" metricMagnitude="-2" metricUnit="m" metricValue="1.108" pageId="2" pageNumber="3" unit="mm" value="11.08">11.08 mm</quantity>
|
||
. Fore and hindlimbs easily touching each other when adpressed against body. Twentyeight presacral vertebrae.
|
||
</paragraph>
|
||
<paragraph id="DE2B5931D835FFB2FEC7F9C0FBD4FD89" blockId="2.[264,1324,358,1905]" lastBlockId="3.[264,1323,284,590]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
Rostral wider than high, contacting first supralabials, nasals and supranasals. Paired supranasals in median contact, contacting anteriormost loreal. Frontonasal approximately hexagonal, wider than long, laterally contacting anterior loreal. Paired prefrontals roughly pentagonal, in median contact, wider than long, contacting frontonasal, both anterior and posterior loreals, first supraciliaries, first and second supraoculars, and frontal. Frontal lanceolate, approximately twice as long as wide, wider anteriorly, in contact with prefrontal, second supraoculars and frontoparietals. Four supraoculars; the first the smallest, the second the longest and widest. Posteriormost supraocular in contact with the frontal is the second (
|
||
<bibRefCitation id="BA0524C0D834FFB2FEBBFEE3FDC0FE99" author="Greer" box="[324,613,324,350]" pageId="3" pageNumber="10" refString="Greer, A. E. & Broadley, D. (2000) Six characters of systematic importance in the scincid lizard genus Mabuya. Hamadryad, 25, 1 - 12." type="journal article" year="2000">Greer & Broadley 2000</bibRefCitation>
|
||
). Four supraciliaries, second longest. Paired frontoparietals, longer than wide, in broad contact at midline, in contact with frontal, all supraoculars except the first, parietal and interparietal. Interparietal rhomboid, longer than wide, wider anteriorly, posterior tip barely in pointcontact with nuchals, and preventing the parietals from touching each other. Parietal eye hardly distinct. Parietals larger than interparietal, wider than long, overlapping the upper temporal scale (
|
||
<bibRefCitation id="BA0524C0D834FFB2FC65FDABFB6EFDE1" author="Greer" box="[922,1227,524,550]" pageId="3" pageNumber="10" refString="Greer, A. E. & Nussbaum, R. A. (2000) New character useful in the Systematics of the Scincid Lizard Genus Mabuya. Copeia, 2000, 615 - 618." type="journal article" year="2000">Greer & Nussbaum 2000</bibRefCitation>
|
||
). Single pair of transversely enlarged nuchals, each as wide as two rows of dorsals.
|
||
</paragraph>
|
||
<paragraph id="DE2B5931D834FFB2FEF7F8A6FB0AF8FC" blockId="3.[264,1323,1793,1851]" pageId="3" pageNumber="4">
|
||
<emphasis id="ECE08523D834FFB2FEF7F8A6FED7F8DE" bold="true" box="[264,370,1793,1817]" pageId="3" pageNumber="4">PLATE 1</emphasis>
|
||
. Drawings of the head of the
|
||
<typeStatus id="012FE793D834FFB2FD51F8A6FCAEF8DF" box="[686,779,1793,1816]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="199422B2D834FFB2FCD1F8A5FBA5F8DE" box="[814,1024,1794,1817]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="4" phylum="Chordata" rank="species" species="meridensis">
|
||
<emphasis id="ECE08523D834FFB2FCD1F8A5FBA5F8DE" box="[814,1024,1794,1817]" italics="true" pageId="3" pageNumber="4">Mabuya meridensis</emphasis>
|
||
</taxonomicName>
|
||
(ULABG 1570): (A) lateral view of the head and (B) dorsal view of the anterior part of the body. Scale bar =
|
||
<quantity id="196CF4D4D834FFB2FB94F883FB09F8FC" box="[1131,1196,1828,1851]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="4" unit="mm" value="2.0">2 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DE2B5931D833FFB5FEC7FEBBFCF8FCD1" blockId="4.[264,1324,284,1873]" pageId="4" pageNumber="5">
|
||
Nasal subrectangular. Nostril located posteriorly. Postnasal small, in contact with supranasal, anterior loreal and first supralabial. Two subrectangular loreals behind nasal, subequal in size, the second slightly higher. First loreal in contact with first and second supralabials, second loreal in contact with second and third supralabials. One presubocular in contact with fourth, fifth and sixth supralabials. One preocular, in front of presubocular and behind second loreal, in contact with third and fourth supralabials. Lower eyelid undivided with a transparent disk, one row of small scales across its dorsal edge (
|
||
<bibRefCitation id="BA0524C0D833FFB5FB3BFDABFE19FD89" author="Greer" pageId="4" pageNumber="10" refString="Greer, A. E. & Broadley, D. (2000) Six characters of systematic importance in the scincid lizard genus Mabuya. Hamadryad, 25, 1 - 12." type="journal article" year="2000">Greer & Broadley 2000</bibRefCitation>
|
||
). Eight supralabials, the sixth being the widest and forming the lower bor der of the eyelid. Seven infralabials. Temporals imbricate, smooth, cycloid, not distinctly delimited from scales on the nape or sides of the neck. Two pretemporals. One primary temporal, two secondary temporals in contact and three tertiary temporals (
|
||
<bibRefCitation id="BA0524C0D833FFB5FB82FD0BFE2FFD29" author="Greer" pageId="4" pageNumber="10" refString="Greer, A. E. & Broadley, D. (2000) Six characters of systematic importance in the scincid lizard genus Mabuya. Hamadryad, 25, 1 - 12." type="journal article" year="2000">Greer & Broadley 2000</bibRefCitation>
|
||
). Earopening relatively small, oval, with undulating anterior margin and smooth posterior margin. Auricular lobules absent.
|
||
</paragraph>
|
||
<paragraph id="DE2B5931D833FFB5FEC7FC83FCF9FC71" blockId="4.[264,1324,284,1873]" pageId="4" pageNumber="5">Mental wider than long, posterior margin straight. Postmental wider than long adjacent to first and twothirds of second infralabials. Two pairs of chin shields, first in contact with postmental, posterior onethird of second and anterior threequarters of third infralabials. Gulars similar in size and outline to ventrals.</paragraph>
|
||
<paragraph id="DE2B5931D833FFB5FEC7FC63FB7FFB91" blockId="4.[264,1324,284,1873]" pageId="4" pageNumber="5">Palms and soles covered with small tubercles, subequal in size. Both regions delimited by a row of larger and flatter scales. Subdigital lamellae smooth, single, 12 under fourth finger (on each sides), 15 under left fourth toe (right toe partly amputated). Fingers and toes clawed. Relative length of the toes in the following order: I <II <III = V <IV.</paragraph>
|
||
<paragraph id="DE2B5931D833FFB5FEC7FBC3FB24FB31" blockId="4.[264,1324,284,1873]" pageId="4" pageNumber="5">All scales, except head shields and scales on sole and digits, cycloid, smooth and imbricate. Thirty scale rows around midbody, 52 transverse rows of dorsal scales, 31 transverse rows of ventral scales. Four preanals larger than adjacent ventral scales. Median subcaudal series of scales twice as wide as long on the posterior half of the tail.</paragraph>
|
||
<paragraph id="DE2B5931D833FFB5FEBEFAA3FDDCFAA9" blockId="4.[264,1324,284,1873]" pageId="4" pageNumber="5">Coloration in preservative: background colour of flanks and upper side of the head, neck, back, limbs and tail dark bronze. Venter, lower side of head, throat, lower side of limbs and tail immaculate grey.</paragraph>
|
||
<paragraph id="DE2B5931D833FFB5FEC7FADBFCC8F9F1" blockId="4.[264,1324,284,1873]" pageId="4" pageNumber="5">Lower half of rostral and each supralabial, and upper third part of each infralabial and mental white, immaculate, resulting in distinct and contrasting “lip marks”. A dark spot on posterior edge of each fourth supraocular. Lateral and upper sides of limbs spotted with small fused dark dots. Palms and soles slightly darker than venter. Preanals pale cream with four little chocolate brown longitudinal dashes.</paragraph>
|
||
<paragraph id="DE2B5931D833FFB4FEC7F9E3FB0AFE11" blockId="4.[264,1324,284,1873]" lastBlockId="5.[264,1325,284,1830]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">Seven dark brown stripes run along body, some of them continuing along tail. One very thin vertebral stripe, from 8th dorsal until base of the tail. Two dorsolateral stripes not well defined, formed by a succession of aligned dots; widen posteriorly and fused with each other and to the vertebral thin stripe in the posterior half part of the body; from nape (first dorsal scales) until first third of tail where the two stripes merge into a single middorsal stripe with several interruptions; about one scale wide at midbody and astride two rows of scales. Two upper lateral stripes on each side; margins darker and strongly contrasted; about three scales wide at midbody; from nostrils, loreals, upper half part of supralabials, around eyes and temporals, along upper half part of earopenings, on neck, above arms, on sides until insertion of hindlimbs. Two lower lateral stripes; from corner of mouth, below earopening, above forelimb until insertion of hindlimb; dorsal margins darker and very well contrasted, whereas limits between ventral margins and venter not distinct.</paragraph>
|
||
<paragraph id="DE2B5931D832FFB4FEC7FE43FD66FD89" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">Four whitish stripes run along body; two dorsolateral stripes separating dark dorsolateral from the upper lateral dark stripes; and two lateral stripes separating the dark upper lateral from lower lateral dark stripes.</paragraph>
|
||
<paragraph id="DE2B5931D832FFB4FEC7FDFBFE59FD59" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">Although the coloration in life was not observed, it seems that the specimen has darkened in preservative.</paragraph>
|
||
<paragraph id="DE2B5931D832FFB4FEC7FD0BFBC0FBC1" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">
|
||
<emphasis id="ECE08523D832FFB4FEC7FD0BFE11FD01" bold="true" box="[312,436,684,710]" pageId="5" pageNumber="6">Variation:</emphasis>
|
||
The following summary of meristic and mensural variation gives the range for each character, followed by the mean, ± the standard deviation, and sample size in parentheses. For some bilateral characters, the sample size has been noted as the number of sides rather than specimens, and this is then indicated after the sample size. Dorsal scale rows: 52–59 (55.66 ± 2.39, 9); midbody scale rows: 28–32 (30.66 ± 1.32, 9); ventral scale rows: 31–39 (34.88 ± 2.71, 9); lamellae under fourth finger: 12–14 (12.77 ± 0.87, 18 sides); lamellae under fourth toe: 15–18 (16.37 ± 1.14, 16 sides); head length: 10.49–14.46 (12.50 ± 1.37, 8); snout–vent length: 55.02–76.73 (69.63 ± 7.40, 8); tail length: 79.67– 133.26 (103.45 ± 21.43, 7); presacral vertebrae: 28–30 (29.00 ± 0.92, 8).
|
||
</paragraph>
|
||
<paragraph id="DE2B5931D832FFB4FEC7FBB3FE5AFB09" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">
|
||
All specimens studied have separated prefrontals and parietals in contact, except the
|
||
<typeStatus id="012FE793D832FFB4FEF7FB9BFECBFB91" box="[264,366,1084,1110]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
which has prefrontals in contact and parietals barely in pointcontact. Seven specimens have four supraoculars on both sides, two specimens have five supraoculars on the right and four on the left side, and one specimen has five supraoculars on the right and three on the left side.
|
||
</paragraph>
|
||
<paragraph id="DE2B5931D832FFB4FEC7FB7BFA88FA79" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">All specimens from AMNH and UMMZ are lighter in coloration than those from ULABG, with cream clear venter. This difference seems to result from a difference in fixation conditions. The single juvenile (UMMZ 57435) has a more contrasted and vivid coloration than the adults. Of the eight adult specimens, four have a dark dot on each fourth supraocular. Position of the dorsal scale where the thin dark vertebral stripe begins: 1–6 (2.75 ± 1.83, 8), and where the two dark dorsolateral stripes begin: 8–17 (11.62 ± 3.24, 8).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="968E0ABAD832FFB4FEC7FA6BFEF0F9F1" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="DE2B5931D832FFB4FEC7FA6BFEF0F9F1" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">
|
||
<emphasis id="ECE08523D832FFB4FEC7FA6BFE6DFA21" bold="true" box="[312,456,1484,1510]" pageId="5" pageNumber="6">Etymology:</emphasis>
|
||
The specific name was given in reference to Mérida, capital city of the state of Mérida,
|
||
<collectingCountry id="A68319A1D832FFB4FE39FA53FDE4F9C9" box="[454,577,1524,1550]" name="Venezuela" pageId="5" pageNumber="6">Venezuela</collectingCountry>
|
||
, the locality where the
|
||
<typeStatus id="012FE793D832FFB4FCB1FA53FC10F9C9" box="[846,949,1524,1550]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="199422B2D832FFB4FC23FA53FB67F9CA" box="[988,1218,1524,1549]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="meridensis">
|
||
<emphasis id="ECE08523D832FFB4FC23FA53FB67F9CA" box="[988,1218,1524,1549]" italics="true" pageId="5" pageNumber="6">Mabuya meridensis</emphasis>
|
||
</taxonomicName>
|
||
was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="968E0ABAD832FFB6FEC7F9E3FE3EFDB1" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="distribution">
|
||
<paragraph id="DE2B5931D832FFB4FEC7F9E3FD43F8E1" blockId="5.[264,1325,284,1830]" pageId="5" pageNumber="6">
|
||
<emphasis id="ECE08523D832FFB4FEC7F9E3FD78F999" bold="true" box="[312,733,1604,1630]" pageId="5" pageNumber="6">Distribution, habitat and ecology:</emphasis>
|
||
According to
|
||
<bibRefCitation id="BA0524C0D832FFB4FC79F9E3FB1BF999" author="Huber" box="[902,1214,1604,1630]" pageId="5" pageNumber="10" refString="Huber, O. & Alarcon, C. (1988) Mapa de Vegetacion de Venezuela. 1: 2,000,000. Republica de Venezuela, Ministerio del Ambiente y de los Recursos Naturales Renovables, Caracas, Venezuela." type="book chapter" year="1988">Huber and Alarcón (1988)</bibRefCitation>
|
||
, the
|
||
<typeStatus id="012FE793D832FFB4FB07F9E3FA8FF999" box="[1272,1322,1604,1630]" pageId="5" pageNumber="6">type</typeStatus>
|
||
locality is situated among farmlands and areas for cattle raising in the Venezuelan Andes. This area is bordered by cloud forests that are found at an altitudinal range from
|
||
<quantity id="196CF4D4D832FFB4FB2FF933FEC4F911" metricMagnitude="3" metricUnit="m" metricValue="2.6" metricValueMax="3.2" metricValueMin="2.0" pageId="5" pageNumber="6" unit="m" value="2600.0" valueMax="3200.0" valueMin="2000.0">2000 to 3200 m</quantity>
|
||
above sea level. These forests are dense, with medium to large sized trees and also provide a welldeveloped understory with numerous epiphytic plants as well as endemic species of ferns, bromeliads and orchids.
|
||
</paragraph>
|
||
<caption id="8AEB09B9D831FFB7FEF7FC3DFCABFC75" box="[264,782,922,946]" httpUri="https://zenodo.org/record/170941/files/figure.png" pageId="6" pageNumber="7" targetBox="[264,1323,326,898]" targetPageId="6">
|
||
<paragraph id="DE2B5931D831FFB7FEF7FC3DFCABFC75" blockId="6.[264,782,922,946]" box="[264,782,922,946]" pageId="6" pageNumber="7">
|
||
<emphasis id="ECE08523D831FFB7FEF7FC3DFE2FFC75" bold="true" box="[264,394,922,946]" pageId="6" pageNumber="7">FIGURE 1.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="199422B2D831FFB7FDCAFC3CFCA2FC75" box="[565,775,923,946]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="meridensis">
|
||
<emphasis id="ECE08523D831FFB7FDCAFC3CFCA2FC75" box="[565,775,923,946]" italics="true" pageId="6" pageNumber="7">Mabuya meridensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8AEB09B9D831FFB7FEF7F9F1FDE1F957" httpUri="https://zenodo.org/record/170942/files/figure.png" pageId="6" pageNumber="7" targetBox="[340,1246,1000,1597]" targetPageId="6">
|
||
<paragraph id="DE2B5931D831FFB7FEF7F9F1FDE1F957" blockId="6.[264,1322,1622,1680]" pageId="6" pageNumber="7">
|
||
<emphasis id="ECE08523D831FFB7FEF7F9F1FE2CF9A9" bold="true" box="[264,393,1622,1646]" pageId="6" pageNumber="7">FIGURE 2.</emphasis>
|
||
Living specimen of
|
||
<taxonomicName id="199422B2D831FFB7FD9BF9F0FC91F9A9" box="[612,820,1623,1646]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="meridensis">
|
||
<emphasis id="ECE08523D831FFB7FD9BF9F0FC91F9A9" box="[612,820,1623,1646]" italics="true" pageId="6" pageNumber="7">Mabuya meridensis</emphasis>
|
||
</taxonomicName>
|
||
. Photograph by César L. BarrioAmorós, courtesy of Fundación AndígenA.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DE2B5931D831FFB6FEC7F94BFCFAFE39" blockId="6.[264,1323,1772,1878]" lastBlockId="7.[264,1323,284,630]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
|
||
The
|
||
<typeStatus id="012FE793D831FFB7FE91F94BFE05F8C1" box="[366,416,1772,1798]" pageId="6" pageNumber="7">type</typeStatus>
|
||
specimen, along with the
|
||
<typeStatus id="012FE793D831FFB7FD26F94BFCECF8C1" box="[729,841,1772,1798]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
|
||
ULABG 4153 and 4281, were collected in a garden of a house in Mérida city.
|
||
<taxonomicName id="199422B2D831FFB7FD3FF8B3FC00F8EA" box="[704,933,1812,1837]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="meridensis">
|
||
<emphasis id="ECE08523D831FFB7FD3FF8B3FC00F8EA" box="[704,933,1812,1837]" italics="true" pageId="6" pageNumber="7">Mabuya meridensis</emphasis>
|
||
</taxonomicName>
|
||
was observed by us in secondary forests, farmlands, rocky areas, margins of rivers or creeks, as well as disturbed areas such as dirt roads, stonewalls and gardens. We saw
|
||
<taxonomicName id="199422B2D830FFB6FCD6FEBBFBB5FEF2" box="[809,1040,284,309]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="meridensis">
|
||
<emphasis id="ECE08523D830FFB6FCD6FEBBFBB5FEF2" box="[809,1040,284,309]" italics="true" pageId="7" pageNumber="8">Mabuya meridensis</emphasis>
|
||
</taxonomicName>
|
||
in Mérida from
|
||
<quantity id="196CF4D4D830FFB6FB2DFEBBFA8EFEF1" box="[1234,1323,284,310]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="8" unit="m" value="1300.0">1300 m</quantity>
|
||
(in the city of Ejido) to ca.
|
||
<quantity id="196CF4D4D830FFB6FDBEFEE3FD3CFE99" box="[577,665,324,350]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="8" unit="m" value="2200.0">2200 m</quantity>
|
||
above sea level in Mucuruba. However, the species was never found in the cloud forest near Mérida city (Cesar Barrio, pers. comm.). At the least sign of danger,
|
||
<taxonomicName id="199422B2D830FFB6FE37FE33FDD4FE6A" box="[456,625,404,429]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="meridensis">
|
||
<emphasis id="ECE08523D830FFB6FE37FE33FDD4FE6A" box="[456,625,404,429]" italics="true" pageId="7" pageNumber="8">M. meridensis</emphasis>
|
||
</taxonomicName>
|
||
rapidly hides amidst clumps of herbaceous vegetation or into cracks on the walls of houses. At present,
|
||
<taxonomicName id="199422B2D830FFB6FCD1FE1BFC70FE12" box="[814,981,444,469]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="meridensis">
|
||
<emphasis id="ECE08523D830FFB6FCD1FE1BFC70FE12" box="[814,981,444,469]" italics="true" pageId="7" pageNumber="8">M. meridensis</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="012FE793D830FFB6FB07FE1BFA8FFE11" box="[1272,1322,444,470]" pageId="7" pageNumber="8">type</typeStatus>
|
||
locality, and a few other places around Mérida city.
|
||
</paragraph>
|
||
<paragraph id="DE2B5931D830FFB6FEC7FDABFE3EFDB1" blockId="7.[264,1323,284,630]" pageId="7" pageNumber="8">
|
||
Like all other species belonging to the genus
|
||
<taxonomicName id="199422B2D830FFB6FC9BFDABFC61FDE2" box="[868,964,524,549]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="genus">
|
||
<emphasis id="ECE08523D830FFB6FC9BFDABFC61FDE2" box="[868,964,524,549]" italics="true" pageId="7" pageNumber="8">Mabuya</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="199422B2D830FFB6FC29FDABFBDAFDE2" box="[982,1151,524,549]" class="Reptilia" family="Scincidae" genus="Mabuya" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="species" species="meridensis">
|
||
<emphasis id="ECE08523D830FFB6FC29FDABFBDAFDE2" box="[982,1151,524,549]" italics="true" pageId="7" pageNumber="8">M. meridensis</emphasis>
|
||
</taxonomicName>
|
||
is viviparous, four and five embryos having been found in the uterus of females ULABG 4153 and 4281, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |