286 lines
30 KiB
XML
286 lines
30 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2011" docId="9A6E243C61EB46267E19D35B46B56C17" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha rotundicollis Attems 1937" docType="treatment" docVersion="6" lastPageNumber="54" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="49" updateTime="1668152297771" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>131</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>161</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152031111" LSID="urn:lsid:plazi:treatment:9A6E243C61EB46267E19D35B46B56C17" httpUri="http://treatment.plazi.org/id/9A6E243C61EB46267E19D35B46B56C17" lastPageId="53" lastPageNumber="54" pageId="48" pageNumber="49">
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<subSubSection pageId="48" pageNumber="49" type="nomenclature">
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName LSID="http://species-id.net/wiki/Orthomorpha_rotundicollis" authority="Attems, 1937" authorityName="Attems" authorityYear="1937" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis (Attems, 1937)</taxonomicName>
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Figs 4346
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="reference_group">
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Attems 1937</bibRefCitation>
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: 118 (D).
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
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-
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<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">Attems 1938</bibRefCitation>
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: 217 (D);
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<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">1953</bibRefCitation>
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: 170 (R).
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
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<bibRefCitation pageId="48" pageNumber="49">Attems 1937</bibRefCitation>
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: 119 (D), syn. n.
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
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-
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<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">Attems 1938</bibRefCitation>
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: 219 (D).
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
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-
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<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
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: 265 (M);
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<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">1964</bibRefCitation>
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: 361 (M, D);
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<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">1968</bibRefCitation>
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: 56 (M);
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<bibRefCitation author="Hoffman, RL" journalOrPublisher="Journal of Natural History" pageId="114" pageNumber="115" pagination="361 - 389" title="Descriptions and allocation of new or poorly known genera and species of Paradoxosomatidae from south-eastern Asia (Diplopoda: Polydesmida)." url="doi: 10.1080/00222937300770281" volume="7" year="1973">Hoffman 1973</bibRefCitation>
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: 363 (M);
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<bibRefCitation author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">1977</bibRefCitation>
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: 700 (M);
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<bibRefCitation pageId="48" pageNumber="49">Golovatch 1998</bibRefCitation>
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: 42 (M, D);
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<bibRefCitation author="Enghoff, H" journalOrPublisher="Arthropoda Selecta" pageId="114" pageNumber="115" pagination="29 - 43" title="A review of the millipede fauna of Vietnam (Diplopoda)." volume="13" year="2004">Enghoff et al. 2004</bibRefCitation>
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: 38 (M, R).
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</paragraph>
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<paragraph pageId="48" pageNumber="49">
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
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-
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<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
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: 265 (M);
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<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">1964</bibRefCitation>
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||
: 361 (M, D);
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<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">1968</bibRefCitation>
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: 56 (M);
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<bibRefCitation author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">Hoffman 1977</bibRefCitation>
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: 700 (M);
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<bibRefCitation pageId="48" pageNumber="49">Golovatch 1998</bibRefCitation>
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: 42 (M, D);
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<bibRefCitation author="Enghoff, H" journalOrPublisher="Arthropoda Selecta" pageId="114" pageNumber="115" pagination="29 - 43" title="A review of the millipede fauna of Vietnam (Diplopoda)." volume="13" year="2004">Enghoff et al. 2004</bibRefCitation>
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: 39 (M, R).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="lectotype">
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<paragraph pageId="48" pageNumber="49">Lectotype.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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♂ of
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
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(NHMW-3515), Vietnam, Lamdong Prov., Lang Biang,
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<normalizedToken originalValue="Arbre-Broyé">Arbre-Broye</normalizedToken>
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, 1500 m, 01.02.1931, leg. C. Dawydoff.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="paralectotype">
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<paragraph pageId="48" pageNumber="49">Paralectotype.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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1 ♂ of
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
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(NHMW-3515), Vietnam, Lamdong Prov., Dalat, 1500 m, 02.1933, leg. C. Dawydoff.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="48" pageNumber="49" type="non-type material">
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<paragraph pageId="48" pageNumber="49">Non-type material.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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1 ♀ of
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
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(NHMW-8000), Laos, Xiangkhoang Prov., Xiangkhoang Plateau, Xiang Kuang, no date, leg. C. Dawydoff, det. Attems.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="syntypes">
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<paragraph pageId="48" pageNumber="49">Syntypes.</paragraph>
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<paragraph pageId="48" pageNumber="49">
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2 ♂, 1 ♀, 1 juv. of
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
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(NHMW-3519), Vietnam, Lamdong Prov., Peak Lang Biang,
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<normalizedToken originalValue="Arbre-Broyé">Arbre-Broye</normalizedToken>
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, 1500 m, 01.02.1931, leg. C. Dawydoff.
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</paragraph>
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<paragraph lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
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Lectotype designation proposed herewith is necessary to ensure the species is based on a complete male coming from a certain locality, because (1) (
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<bibRefCitation pageId="48" pageNumber="49">Attems (1937</bibRefCitation>
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,
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<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
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)
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<pageBreakToken pageId="49" pageNumber="50" start="start">had</pageBreakToken>
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provided no information on the number of syntypes, and (2) he stated their provenance to have been both from
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<normalizedToken originalValue="Arbre-Broyé">Arbre-Broye</normalizedToken>
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and Dalat, Vietnam.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="53" lastPageNumber="54" pageId="49" pageNumber="50" type="redescription">
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<paragraph pageId="49" pageNumber="50">Redescription.</paragraph>
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<paragraph lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
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Length 29-31 mm (♂), ca 31-35 mm (♀), width of midbody pro- and metazona 2.5-3.0 and 3.7-4.1 mm (♂), 3.4 and 4.5 mm (♀), respectively (vs 2.5 and 4.0 mm in width, as given in the available descriptions (
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<bibRefCitation pageId="49" pageNumber="50">Attems 1937</bibRefCitation>
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,
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<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
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)). Lectotype ca 31 mm long, 2.9 and 4.1 mm wide on midbody pro- and metazona, respectively. Coloration of alcohol material upon long-term preservation rather uniformly light grey-brown (Figs 43 & 45) (vs with a cingulate pattern of castaneous brown head and collum, as well as regions of pro- and metazona adjacent to stricture
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<pageBreakToken pageId="50" pageNumber="51" start="start">alternating</pageBreakToken>
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with light brownish remaining parts, including antennae and legs, as given in the descriptions (
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<bibRefCitation pageId="50" pageNumber="51">Attems 1937</bibRefCitation>
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,
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<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
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)).
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</paragraph>
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<paragraph lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51">
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Head usual, clypeolabral region densely setose, surface of vertex smooth, epicranial suture distinct. Antennae rather long and slender (Figs 43J, 45B & J), extending behind body segment 4 (♂) or 3 (♀) dorsally. Head in width <collum <segments 3 and 4 <segment 2 <segments 5-17 (♂, ♀), gently and gradually tapering thereafter. Collum smooth, with three transverse rows of setae, 4+4 anterior, 2+2 intermediate, and 3+3 posterior setae; caudal corner of paraterga subrectangular, narrowly rounded (Figs 43A, B, J, 45A, B & J). Tegument smooth and shining; metaterga very faintly rugulose, each postcollum one with one caudal row of 3(4)+3(4) very small tubercles growing a little larger laterally, as well as with 2+2 either minute or fully obliterated knobs with insertion points of abraded setae in front row; prozona very finely shagreened, surface below paraterga finely microgranulate. Axial line visible, starting from collum. Paraterga very strongly developed (Figs 43A-G, J-L, 45A-G & J-L), all subhorizontal
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<pageBreakToken pageId="51" pageNumber="52" start="start">and</pageBreakToken>
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lying below dorsum, thin in lateral view, like blunt blades, a little thicker only on pore-bearing segments, on postcollum segments extending increasingly beyond rear tergal margin, nearly pointed to pointed, caudal tip on paraterga 16-19 evidently curved mesad. Calluses on delimited by a sulcus only dorsally, rather narrow. Paraterga 2 broad, anterior edge angulate, lateral edge with three small incisions in anterior half; posterior edge evidently concave (Figs 43A, B, J, 45A, B & J). Paraterga 3 and 4 subequal, like subsequent paraterga, anterior edge broadly rounded, bordered and fused to callus, lateral edge with two small incisions in anterior half on poreless segments, with
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<pageBreakToken pageId="52" pageNumber="53" start="start">only</pageBreakToken>
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one incision near front 1/3 on pore-bearing ones. Ozopores evident, lateral, lying inside an ovoid groove at about 1/4 metazonital length. Transverse sulcus complete on metaterga 5-18, incomplete on metatergum 4, shallow, not reaching bases of paraterga, beaded at bottom, slightly sinuate anteromedially (Figs 43A- D, F, J-L, 45A-F & J-L). Stricture between pro- and metazona rather narrow, deep, ribbed at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segments 2-4 (Figs 43B, D, E, J-K, 45A-F & J-L), each with an evident sharp denticle caudally, thereafter increasingly strongly reduced until segment 10 (♂, ♀). Epiproct (Figs 43E-G, 45E-G) conical, flattened dorsoventrally, apical papillae small; tip subtruncate; pre-apical papillae small, but visible, lying close to tip. Hypoproct (Figs 43G, 45G) roundly subtriangular, setiferous knobs at caudal margin small and well-separated.
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</paragraph>
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<paragraph lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
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Sterna sparsely setose, without modifications, but with two rather large, rounded, fully separated, but nearly contiguous, setose cones between ♂ coxae 4 (Figs 43H, I, 45H & I). A paramedian pair of small tubercles in front of gonopod aperture. Legs
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<pageBreakToken pageId="53" pageNumber="54" start="start">long</pageBreakToken>
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and slender, midbody ones ca 1.2-1.3 (♂) or 0.8-0.9 (♀) as long as midbody height, prefemora without modifications, tarsal brushes present until ♂ legs 4.
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</paragraph>
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<paragraph pageId="53" pageNumber="54">
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Gonopods (Figs 44 & 46) simple. Coxa long and slender, with several setae distodorsally. Prefemur densely setose, nearly 3 times shorter than femorite +
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<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part. Femorite very slender, evidently curved, very slightly enlarged distad,
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<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part demarcated by an oblique lateral sulcus; tip of solenophore small, trifid, with two subequal denticles (one terminal, the other middle) and a broader subterminal lobule.
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</paragraph>
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<caption pageId="53" pageNumber="54">
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<paragraph pageId="53" pageNumber="54">
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Figure 43.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
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Attems, 1937, ♂ lectotype (
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<normalizedToken originalValue="A–I">A-I</normalizedToken>
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), ♀ (
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<normalizedToken originalValue="J–L">J-L</normalizedToken>
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) of
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
|
||
Attems, 1937. A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, D, K segments 10 and 11, dorsal, lateral and dorsal views, respectively
|
||
<normalizedToken originalValue="E–G">E-G</normalizedToken>
|
||
, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="53" pageNumber="54">
|
||
<paragraph pageId="53" pageNumber="54">
|
||
Figure 44.
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
|
||
Attems, 1937, ♂ lectotype of
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Pratinus rotundicollis</taxonomicName>
|
||
Attems, 1937. A, B right gonopod, lateral and mesal views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="53" pageNumber="54">
|
||
<paragraph pageId="53" pageNumber="54">
|
||
Figure 45.
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
|
||
Attems, 1937, ♂ syntype (
|
||
<normalizedToken originalValue="A–I">A-I</normalizedToken>
|
||
), ♀ syntype (
|
||
<normalizedToken originalValue="J–L">J-L</normalizedToken>
|
||
) of
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
|
||
<bibRefCitation pageId="53" pageNumber="54">Attems 1937</bibRefCitation>
|
||
. A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, D segments 11 and 12, dorsal and lateral veiws, respectively K segments 10 and 11, dorsal view
|
||
<normalizedToken originalValue="E–F">E-F</normalizedToken>
|
||
, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="53" pageNumber="54">
|
||
<paragraph pageId="53" pageNumber="54">
|
||
Figure 46.
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
|
||
Attems, 1937, ♂ syntype of
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus tuberculatus" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculatus">Pratinus tuberculatus</taxonomicName>
|
||
<bibRefCitation pageId="53" pageNumber="54">Attems 1937</bibRefCitation>
|
||
. A, B right gonopod, lateral and mesal views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="53" pageNumber="54" type="remark">
|
||
<paragraph pageId="53" pageNumber="54">Remark.</paragraph>
|
||
<paragraph pageId="53" pageNumber="54">
|
||
This species is known from Lamdong Province, southern Vietnam and Luang Prabang Province, Laos (
|
||
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">Attems 1938</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">1953</bibRefCitation>
|
||
). The type locality of both
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
|
||
and
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
|
||
being exactly the same (
|
||
<normalizedToken originalValue="Arbre-Broyé">Arbre-Broye</normalizedToken>
|
||
at Peak Lang Biang), and the differences between them so minor, rather to be treated as infraspecific variations, we are inclined to formally synonymize these species. Indeed, distinctions are observed only in
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
|
||
showing a slightly more rounded caudal corner of the paraterga on the collum (vs slightly more angular in
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
|
||
), in the front row of metatergal setae (= insertion points) borne on virtually fully obliterated knobs (vs on minute knobs in
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
|
||
), and in the apical papillae on the epiproct directed more caudally than ventrally (vs more ventrally than caudally in
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
|
||
) (
|
||
<bibRefCitation pageId="53" pageNumber="54">Attems 1937</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Serie A, Zoologie" pageId="114" pageNumber="115" pagination="133 - 230" title="Myriopoden von Indochina. Expedition von Dr. C. Dawydoff (1938 - 1939)." volume="5" year="1953">1953</bibRefCitation>
|
||
). We can also add another two differences: paraterga 18 and 19 seem to be a little shorter in
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
|
||
than in
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
|
||
, and the caudal row of tubercles on a few caudalmost metaterga demonstrating a pattern of 3+4 or 4+3 in
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha tuberculata" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tuberculata">Orthomorpha tuberculata</taxonomicName>
|
||
(vs invariably 3+3 in
|
||
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha rotundicollis" order="Polydesmida" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="rotundicollis">Orthomorpha rotundicollis</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="53" pageNumber="54">The above is a combined redescription, based on material of both nominate species.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |