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<document id="B554D09424195DB5398E6E4CD38BFA37" ID-CLB-Dataset="20233" ID-DOI="10.5852/ejt.2022.797.1667" ID-GBIF-Dataset="e4b9a92e-23ef-4d55-85c9-ccabcbd8316e" ID-ISSN="2118-9773" ID-Zenodo-Dep="6299487" ID-ZooBank="8DF57743-9C53-4265-BCB5-743276A3A16C" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645807222198" checkinUser="felipe" docAuthor="Telnov, Dmitry" docDate="2022" docId="6D52212B21292F70FD9A489669F7F89D" docLanguage="en" docName="ejt-797_1-100.pdf" docOrigin="European Journal of Taxonomy 797" docSource="http://dx.doi.org/10.5852/ejt.2022.797.1667" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Macrotomoderus angelinii Telnov 2022, sp. nov." docType="treatment" docUuid="8E264897-1526-4DB8-A6CA-BF9888A270EA" docUuidSource="ZooBank" docVersion="10" lastPageNumber="4" masterDocId="916B5953212B2F73FFD24C506A5AFFF0" masterDocTitle="Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species" masterLastPageNumber="100" masterPageNumber="1" pageNumber="3" updateTime="1699316540125" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="ED68AC728F04CF23815ADE4A71B7487F">Revision of the Tomoderinae (Coleoptera: Anthicidae). Part III. New species and records of Macrotomoderus Pic, 1901 from China and a key to the Palaearctic species</mods:title>
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<mods:namePart id="AA76D60F53CF3CDB097AF3F9FB11571C">Telnov, Dmitry</mods:namePart>
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<mods:affiliation id="70657E4D273124CFE216BED2C8050E59">Department of Life Sciences, Natural History Museum, Cromwell Road, SW 7 5 BD, London, United Kingdom &amp; Institute of Biology, University of Latvia, O. Vācieša iela 4, LV – 1004, Rīga, Latvia. Coleopterological Research Center, Institute of Life Sciences and Technology, Daugavpils University, Vienības iela 13, LV – 5401, Daugavpils, Latvia.</mods:affiliation>
<mods:nameIdentifier id="431A32E8D4BEBCD79B4A4281A222D508" type="email">anthicus@gmail.com</mods:nameIdentifier>
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<mods:title id="97120D24250AA84700FD93B7B10EAF85">European Journal of Taxonomy</mods:title>
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<mods:date id="8379A7646F5C69F7178A4C0286813CDE">2022</mods:date>
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<treatment id="6D52212B21292F70FD9A489669F7F89D" ID-DOI="http://doi.org/10.5281/zenodo.6317906" ID-GBIF-Taxon="194038738" ID-Zenodo-Dep="6317906" LSID="urn:lsid:zoobank.org:act:8E264897-1526-4DB8-A6CA-BF9888A270EA" httpUri="http://treatment.plazi.org/id/6D52212B21292F70FD9A489669F7F89D" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="ADE1C3B621292F71FD9A489669B6FB11" box="[584,1004,1222,1249]" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="E544903D21292F71FD9A489669B6FB11" blockId="2.[366,1223,1222,1319]" box="[584,1004,1222,1249]" pageId="2" pageNumber="3">
<heading id="BE0C275121292F71FD9A489669B6FB11" box="[584,1004,1222,1249]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="22FBEBBE21292F71FD9A489669D0FB11" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[584,906,1222,1249]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="angelinii" status="sp. nov.">
<emphasis id="D78F4C2F21292F71FD9A489669D0FB11" bold="true" box="[584,906,1222,1249]" italics="true" pageId="2" pageNumber="3">Macrotomoderus angelinii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421292F71FC43489769B6FB11" box="[913,1004,1223,1249]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
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</paragraph>
<paragraph id="E544903D21292F71FD35495D6917FAD7" blockId="2.[366,1223,1222,1319]" box="[743,845,1293,1319]" pageId="2" pageNumber="3">
<figureCitation id="7DC08CB821292F71FD35495D6917FAD7" box="[743,845,1293,1319]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="4.[189,232,1963,1989]" captionStartId-1="5.[189,232,1928,1954]" captionTargetBox-0="[227,1338,273,1915]" captionTargetBox-1="[365,1211,281,1860]" captionTargetId-0="figure-21@4.[214,1374,265,1924]" captionTargetId-1="figure-12@5.[359,1229,265,1886]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="Fig. 1. Macrotomoderus angelinii sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." captionText-1="Fig. 2. Macrotomoderus angelinii sp. nov., holotype, ♂ (ZIN), aedeagus. A–B. General view, different positions. C. Gonopore armature in median portion of aedeagus. D. Ditto, basal portion of apicale. E. Apical portion. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6299489" figureDoi-1="http://doi.org/10.5281/zenodo.6299491" httpUri-0="https://zenodo.org/record/6299489/files/figure.png" httpUri-1="https://zenodo.org/record/6299491/files/figure.png" pageId="2" pageNumber="3">Figs 1–2</figureCitation>
</paragraph>
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<subSubSection id="ADE1C3B621292F71FF6F491E69D8F907" pageId="2" pageNumber="3" type="diagnosis">
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<heading id="BE0C275121292F71FF6F491E6B93FA98" bold="true" box="[189,457,1358,1384]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="D78F4C2F21292F71FF6F491E6B93FA98" bold="true" box="[189,457,1358,1384]" pageId="2" pageNumber="3">Differential diagnosis</emphasis>
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<paragraph id="E544903D21292F71FF6F492D69D8F907" blockId="2.[188,1400,1405,1784]" pageId="2" pageNumber="3">
This species falls in a group of species from continental
<collectingCountry id="9DECD0AD21292F71FCB3492D69FDFA67" box="[865,935,1405,1431]" name="China" pageId="2" pageNumber="3">China</collectingCountry>
with the lateral fovea of pronotum not or only poorly visible in dorsal view, not forming deep notches at both sides of the lateral constriction. Among these species,
<taxonomicName id="22FBEBBE21292F71FE10499369DDFA2E" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" box="[450,903,1475,1502]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="bukejsi">
<emphasis id="D78F4C2F21292F71FE10499368BFFA2D" box="[450,741,1475,1501]" italics="true" pageId="2" pageNumber="3">Macrotomoderus bukejsi</emphasis>
<bibRefCitation id="816AEDCC21292F71FD3E499369DDFA2E" author="Telnov D." box="[748,903,1475,1502]" pageId="2" pageNumber="3" pagination="463 - 492" refId="ref31178" refString="Telnov D. 2018. Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales Zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.3.008" type="journal article" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="22FBEBBE21292F71FC4049946EE3FA2E" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" box="[914,1209,1475,1502]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="chingpo">
<emphasis id="D78F4C2F21292F71FC4049946E42FA2D" box="[914,1048,1475,1501]" italics="true" pageId="2" pageNumber="3">M. chingpo</emphasis>
<bibRefCitation id="816AEDCC21292F71FBCC49936EE3FA2E" author="Telnov D." box="[1054,1209,1475,1502]" pageId="2" pageNumber="3" pagination="463 - 492" refId="ref31178" refString="Telnov D. 2018. Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales Zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.3.008" type="journal article" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="22FBEBBE21292F71FB2449946B02F9F1" authority="Telnov, 2018" authorityName="Telnov" authorityYear="2018" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="D78F4C2F21292F71FB2449946F2CFA2D" box="[1270,1398,1475,1501]" italics="true" pageId="2" pageNumber="3">M. gracilis</emphasis>
<bibRefCitation id="816AEDCC21292F71FF6F49B76B02F9F1" author="Telnov D." box="[189,344,1511,1537]" pageId="2" pageNumber="3" pagination="463 - 492" refId="ref31178" refString="Telnov D. 2018. Revision of the Tomoderinae (Coleoptera: Anthicidae). Part II. Macrotomoderus Pic, 1901 species from China and Taiwan. Annales Zoologici 68 (3): 463 - 492. https: // doi. org / 10.3161 / 00034541 ANZ 2018.68.3.008" type="journal article" year="2018">Telnov, 2018</bibRefCitation>
</taxonomicName>
have slender and elongate pronotum. The pronotum is medially longitudinally carinate at the constriction area in the two first species, non-carinate in
<taxonomicName id="22FBEBBE21292F71FC634A5B6E18F9D4" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[945,1090,1546,1572]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="angelinii" status="sp. nov.">
<emphasis id="D78F4C2F21292F71FC634A5B6E18F9D4" box="[945,1090,1546,1572]" italics="true" pageId="2" pageNumber="3">M. angelinii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421292F71FB9C4A5A6EF2F9D4" box="[1102,1192,1546,1572]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
The pronotum is peculiarly modified in
<taxonomicName id="22FBEBBE21292F71FE164A7E681EF9B7" authorityName="Telnov" authorityYear="2018" box="[452,580,1581,1607]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="D78F4C2F21292F71FE164A7E681EF9B7" box="[452,580,1581,1607]" italics="true" pageId="2" pageNumber="3">M. gracilis</emphasis>
</taxonomicName>
, with the median part of the anterior margin of it elevated and strongly projecting anteriad, provided with a group of ך-shaped (bent) setae directed anteriad (both modifications are not present in
<taxonomicName id="22FBEBBE21292F71FE594A246840F97D" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[395,538,1651,1677]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="angelinii" status="sp. nov.">
<emphasis id="D78F4C2F21292F71FE594A246840F97D" box="[395,538,1651,1677]" italics="true" pageId="2" pageNumber="3">M. angelinii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421292F71FDF24A246824F97E" box="[544,638,1652,1678]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
).
<taxonomicName id="22FBEBBE21292F71FD434A2369FDF97D" authority="Telnov, 2022" authorityName="Telnov" authorityYear="2022" box="[657,935,1651,1677]" class="Insecta" family="Anthicidae" genus="Macrotomoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tenuis" status="sp. nov.">
<emphasis id="D78F4C2F21292F71FD434A2369FDF97D" box="[657,935,1651,1677]" italics="true" pageId="2" pageNumber="3">Macrotomoderus tenuis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CCBCF15421292F71FC7C4A246E5DF97E" box="[942,1031,1652,1678]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
(see description below) has the lateral foveae of the pronotum short and narrow, but well-visible in dorsal view (this includes denticles of the fovea), the median setae on the anterior margin of the male pronotum either straight and anteriadpointed or slightly C-shaped (curved) in their apical portion.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621292F71FF6F4B4F6901F87A" pageId="2" pageNumber="3" type="etymology">
<paragraph id="E544903D21292F71FF6F4B4F6B1EF8C9" blockId="2.[189,324,1823,1849]" box="[189,324,1823,1849]" pageId="2" pageNumber="3">
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<emphasis id="D78F4C2F21292F71FF6F4B4F6B1EF8C9" bold="true" box="[189,324,1823,1849]" pageId="2" pageNumber="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21292F71FF6F4B1D6901F87A" blockId="2.[189,1399,1869,1931]" pageId="2" pageNumber="3">
Patronymic. This species named after the famous Italian coleopterist and my good colleague Fernando Angelini who has passed away on
<date id="9145B6FD21292F71FD814B2068AAF87A" box="[595,752,1904,1931]" pageId="2" pageNumber="3" value="2021-05-28">28 May 2021</date>
aged 72.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621282F70FF6F4D596E88FE0C" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="E544903D21282F70FF6F4D596B36FED2" blockId="3.[189,364,264,291]" box="[189,364,264,291]" pageId="3" pageNumber="4">
<heading id="BE0C275121282F70FF6F4D596B36FED2" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="D78F4C2F21282F70FF6F4D596B36FED2" bold="true" box="[189,364,264,291]" pageId="3" pageNumber="4">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21282F70FF384D676E49FE68" blockId="3.[189,1399,311,408]" pageId="3" pageNumber="4">
<materialsCitation id="55939A6021282F70FF384D676E49FE68" ID-GBIF-Occurrence="3705075327" collectingDate="2018-05-20" collectionCode="ZIN" collectorName="I. Belousov &amp; I. Kabak" country="China" elevation="3405" latitude="26.447222" location="Yunnan" longLatPrecision="20" longitude="99.30528" pageId="3" pageNumber="4" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="D78F4C2F21282F70FF384D676B06FEA1" bold="true" box="[234,348,311,337]" pageId="3" pageNumber="4">
<typeStatus id="3A402E9F21282F70FF384D676B06FEA1" box="[234,348,311,337]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="9DECD0AD21282F70FF6F4D0A6B42FE84" box="[189,280,346,372]" name="China" pageId="3" pageNumber="4">CHINA</collectingCountry>
•
<specimenCount id="F3FD5BB421282F70FEE94D0B6B0EFE85" box="[315,340,347,373]" pageId="3" pageNumber="4" type="male">♂</specimenCount>
; “CHINA,
<collectingRegion id="273F5EDF21282F70FE314D0B681AFE85" box="[483,576,347,373]" country="China" name="Yunnan" pageId="3" pageNumber="4">Yunnan</collectingRegion>
,
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W Laiping, SE Lajing,
<quantity id="22033DD821282F70FC344D0A6E75FE84" box="[998,1071,346,372]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="4" unit="km" value="3.4">3.4km</quantity>
SSE Jiaolieshan
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,
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<elevation id="6ED6770E21282F70FAC34D0A6F2BFE85" box="[1297,1393,346,373]" metricMagnitude="3" metricUnit="m" metricValue="3.405" pageId="3" pageNumber="4" unit="m" value="3405.0">3405 m</elevation>
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,
<date id="9145B6FD21282F70FF6F4D2E6B69FE68" box="[189,307,382,408]" pageId="3" pageNumber="4" value="2018-05-20">
<collectingDate id="81014F1521282F70FF6F4D2E6B69FE68" box="[189,307,382,408]" pageId="3" pageNumber="4" value="2018-05-20">20.v.2018</collectingDate>
</date>
,
<collectorName id="480EF5EB21282F70FEED4D2E6B9DFE67" box="[319,455,381,408]" pageId="3" pageNumber="4">I. Belousov</collectorName>
&amp;
<collectorName id="480EF5EB21282F70FE3E4D2E680BFE67" box="[492,593,381,408]" pageId="3" pageNumber="4">I. Kabak</collectorName>
leg. //
<geoCoordinate id="80CFF6FA21282F70FD704D2D6974FE68" box="[674,814,380,408]" degrees="26" direction="north" minutes="26" orientation="latitude" pageId="3" pageNumber="4" precision="15" seconds="50" value="26.447222">26°26′50″N</geoCoordinate>
<geoCoordinate id="80CFF6FA21282F70FCE74D2E6999FE68" box="[821,963,380,408]" degrees="99" direction="east" minutes="18" orientation="longitude" pageId="3" pageNumber="4" precision="15" seconds="19" value="99.30528">99°18′19″E</geoCoordinate>
”;
<collectionCode id="83EA08F821282F70FC084D2E6E55FE68" box="[986,1039,382,408]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="4" type="Museum">ZIN</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="E544903D21282F70FF384DEF6E88FE0C" blockId="3.[189,1234,447,509]" pageId="3" pageNumber="4">
<materialsCitation id="55939A6021282F70FF384DEF68B7FE0D" ID-GBIF-Occurrence="3705075302" collectingDate="2018-05-20" collectionCode="ZIN" collectorName="I. Belousov &amp; I. Kabak" country="China" elevation="3405" latitude="26.447222" location="CHINA" longLatPrecision="20" longitude="99.30528" pageId="3" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="D78F4C2F21282F70FF384DEF6B3DFE29" bold="true" box="[234,359,447,473]" pageId="3" pageNumber="4">
<typeStatus id="3A402E9F21282F70FF384DEF6B3DFE29" box="[234,359,447,473]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
</emphasis>
(2 specimens)
<collectingCountry id="9DECD0AD21282F70FF6F4DB26B42FE0C" box="[189,280,482,508]" name="China" pageId="3" pageNumber="4">CHINA</collectingCountry>
•
<specimenCount id="F3FD5BB421282F70FEE34DB26B04FE0D" box="[305,350,482,509]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
; same labels as for holotype;
<collectionCode id="83EA08F821282F70FD6E4DB368B7FE0D" box="[700,749,483,509]" collectionName="Russia, St. Petersburg, Russian Academy of Sciences, Zoological Institute" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="3" pageNumber="4" type="Museum">ZIN</collectionCode>
</materialsCitation>
•
<materialsCitation id="55939A6021282F70FCD74DB26E94FE0C" ID-GBIF-Occurrence="3705075313" box="[773,1230,482,509]" collectingDate="2018-05-20" collectionCode="DTC" collectorName="I. Belousov &amp; I. Kabak" country="China" elevation="3405" latitude="26.447222" location="Yunnan" longLatPrecision="20" longitude="99.30528" pageId="3" pageNumber="4" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<specimenCount id="F3FD5BB421282F70FCD74DB2696EFE0D" box="[773,820,482,509]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
; same labels as for holotype;
<collectionCode id="83EA08F821282F70FB424DB26E94FE0C" box="[1168,1230,482,508]" collectionName="Dmitrijs Telnovs" pageId="3" pageNumber="4">DTC</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621282F70FF6F4E736923F97D" pageId="3" pageNumber="4" type="description">
<paragraph id="E544903D21282F70FF6F4E736B14FDCD" blockId="3.[189,334,547,573]" box="[189,334,547,573]" pageId="3" pageNumber="4">
<heading id="BE0C275121282F70FF6F4E736B14FDCD" bold="true" box="[189,334,547,573]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="D78F4C2F21282F70FF6F4E736B14FDCD" bold="true" box="[189,334,547,573]" pageId="3" pageNumber="4">Description</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21282F70FF6F4E036B22FD42" blockId="3.[189,1400,594,690]" pageId="3" pageNumber="4">
MEASUREMENTS.
<typeStatus id="3A402E9F21282F70FEAC4E026BB4FD9C" box="[382,494,594,620]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
, total body length
<quantity id="22033DD821282F70FD1E4E026963FD9D" box="[716,825,594,621]" metricMagnitude="-3" metricUnit="m" metricValue="3.48" pageId="3" pageNumber="4" unit="mm" value="3.48">3.48 mm</quantity>
; head
<quantity id="22033DD821282F70FC544E0269B5FD9D" box="[902,1007,594,621]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="3" pageNumber="4" unit="mm" value="0.88">0.88 mm</quantity>
long, across eyes
<quantity id="22033DD821282F70FB1B4E026F7EFD9D" box="[1225,1316,594,621]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="4" unit="mm" value="0.6">0.6 mm</quantity>
broad, pronotum
<quantity id="22033DD821282F70FEE54E256BCAFD60" box="[311,400,629,656]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="4" unit="mm" value="0.8">0.8 mm</quantity>
long, maximum width
<quantity id="22033DD821282F70FD734E2568A7FD60" box="[673,765,629,656]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="0.5">0.5 mm</quantity>
, minimum width
<quantity id="22033DD821282F70FC194E256E7DFD60" box="[971,1063,629,656]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="4" unit="mm" value="0.2">0.2 mm</quantity>
, elytra
<quantity id="22033DD821282F70FBAE4E256E8FFD60" box="[1148,1237,629,656]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="4" unit="mm" value="1.8">1.8 mm</quantity>
long,
<quantity id="22033DD821282F70FACC4E256F22FD60" box="[1310,1400,629,656]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="4" unit="mm" value="1.2">1.2 mm</quantity>
combined wide.
</paragraph>
<paragraph id="E544903D21282F70FF6F4E8A6EDAF9EE" blockId="3.[187,1401,730,1566]" pageId="3" pageNumber="4">
Dorsum and venter uniformly pale brown. Mouthparts, antennae, palps and legs yellow. Head ovoid with moderately large, ovoid compound eyes, which are not protruding beyond head outline laterally. Head rounded in broad arc posterior to eyes. Head dorsal punctures minute and inconspicuous. Head dorsal setae inconspicuous, sparse. Antennae nearly extending towards base of pronotum. Antennomere three about as long as antennomere two, antennomeres 8–10 transverse, of which penultimate antennomere strongly transverse. Terminal antennomere asymmetrically triangular with rounded apex, nearly twice as long as penultimate antennomere. Terminal maxillary palpomere slightly securiform. Pronotum elongate and slender, with broad, medially slightly notched (in dorsal view) postmedian lateral constriction. Front margin of anterior lobe very broadly rounded, dorsally without anterior rim, medially with a bunch of yellowish, moderately long, Ɔ-shaped (curved), posterodorsad-pointed setae (
<figureCitation id="7DC08CB821282F70FBAF48476EA4FBC1" box="[1149,1278,1047,1073]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[227,1338,273,1915]" captionTargetId="figure-21@4.[214,1374,265,1924]" captionTargetPageId="4" captionText="Fig. 1. Macrotomoderus angelinii sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299489" httpUri="https://zenodo.org/record/6299489/files/figure.png" pageId="3" pageNumber="4">Fig. 1B–C</figureCitation>
). Anterior lobe slightly convex in lateral view (
<figureCitation id="7DC08CB821282F70FDA6486A688BFBA4" box="[628,721,1082,1108]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[227,1338,273,1915]" captionTargetId="figure-21@4.[214,1374,265,1924]" captionTargetPageId="4" captionText="Fig. 1. Macrotomoderus angelinii sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299489" httpUri="https://zenodo.org/record/6299489/files/figure.png" pageId="3" pageNumber="4">Fig. 1C</figureCitation>
). Lateral constriction continues onto disc in lateral view, shallow (
<figureCitation id="7DC08CB821282F70FEFB480D6BDEFB87" box="[297,388,1117,1143]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[227,1338,273,1915]" captionTargetId="figure-21@4.[214,1374,265,1924]" captionTargetPageId="4" captionText="Fig. 1. Macrotomoderus angelinii sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299489" httpUri="https://zenodo.org/record/6299489/files/figure.png" pageId="3" pageNumber="4">Fig. 1C</figureCitation>
). Lateral pronotal fovea narrow at lower external margin of pronotum, strongly widens upwards towards pronotal disc in lateral view, external margins protruding into a pair of inconspicuous, obtuse (in lateral view), glabrous, narrowly separated denticles (in lateral view). Cavity in lateral wall of pronotum between lateral denticles small, narrowly rectangular. In dorsal view, lateral pronotal fovea narrow, anterior and posterior pair of denticles poorly visible, glabrous (
<figureCitation id="7DC08CB821282F70FBF348BA6E24FAF4" box="[1057,1150,1258,1284]" captionStart="Fig" captionStartId="4.[189,232,1963,1989]" captionTargetBox="[227,1338,273,1915]" captionTargetId="figure-21@4.[214,1374,265,1924]" captionTargetPageId="4" captionText="Fig. 1. Macrotomoderus angelinii sp. nov., holotype, ♂ (ZIN). A. Habitus, dorsal view. B. Forebody, dorsal view. C. Ditto, lateral view. Scale bar: A = 1 mm; B–C = not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299489" httpUri="https://zenodo.org/record/6299489/files/figure.png" pageId="3" pageNumber="4">Fig. 1B</figureCitation>
). Pronotal punctures minute on disc; lateral constriction dorsally with dense, moderately large to large circular punctures separated by about their diameters. Dorsal pronotal setae inconspicuous, moderately long. Scutellar shield minute, triangular. Elytra dorsally elliptical, convex in lateral view, strongly widened laterally around midlength, lateral margins evenly broadly rounded, shoulders obsolete (apterous species). Elytral punctures much stronger and larger than those on dorsal forebody but somewhat less coarse than dorsal punctures on lateral pronotal constriction. Elytral setae long and sparse, suberect. Male and female tergite and morphological sternite VII broadly rounded at posterior margin. Aedeagus as in
<figureCitation id="7DC08CB821282F70FB3349B06F7DFA0A" box="[1249,1319,1504,1530]" captionStart="Fig" captionStartId="5.[189,232,1928,1954]" captionTargetBox="[365,1211,281,1860]" captionTargetId="figure-12@5.[359,1229,265,1886]" captionTargetPageId="5" captionText="Fig. 2. Macrotomoderus angelinii sp. nov., holotype, ♂ (ZIN), aedeagus. A–B. General view, different positions. C. Gonopore armature in median portion of aedeagus. D. Ditto, basal portion of apicale. E. Apical portion. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6299491" httpUri="https://zenodo.org/record/6299491/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
, rather narrow and slender, with very delicate, tube-like encapsulated gonopore armature.
</paragraph>
<paragraph id="E544903D21282F70FF6F4A156BEAF9AF" blockId="3.[189,432,1605,1631]" box="[189,432,1605,1631]" pageId="3" pageNumber="4">
<heading id="BE0C275121282F70FF6F4A156BEAF9AF" bold="true" box="[189,432,1605,1631]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="D78F4C2F21282F70FF6F4A156BEAF9AF" bold="true" box="[189,432,1605,1631]" pageId="3" pageNumber="4">Sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21282F70FF6F4A236923F97D" blockId="3.[189,889,1651,1678]" box="[189,889,1651,1678]" pageId="3" pageNumber="4">Female anterior margin of pronotum without modifications.</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621282F70FF6F4AE5687FF90D" box="[189,549,1717,1790]" pageId="3" pageNumber="4" type="biology_ecology">
<paragraph id="E544903D21282F70FF6F4AE56B7AF93F" blockId="3.[189,288,1717,1743]" box="[189,288,1717,1743]" pageId="3" pageNumber="4">
<heading id="BE0C275121282F70FF6F4AE56B7AF93F" bold="true" box="[189,288,1717,1743]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="D78F4C2F21282F70FF6F4AE56B7AF93F" bold="true" box="[189,288,1717,1743]" pageId="3" pageNumber="4">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21282F70FF6F4AB3687FF90D" blockId="3.[189,549,1763,1790]" box="[189,549,1763,1790]" pageId="3" pageNumber="4">
Collected at
<quantity id="22033DD821282F70FE834AB36BF0F90E" box="[337,426,1763,1790]" metricMagnitude="3" metricUnit="m" metricValue="3.405" pageId="3" pageNumber="4" unit="m" value="3405.0">3405 m</quantity>
elevation.
</paragraph>
</subSubSection>
<subSubSection id="ADE1C3B621282F70FF6F4B7569F7F89D" box="[189,941,1829,1902]" pageId="3" pageNumber="4" type="distribution">
<paragraph id="E544903D21282F70FF6F4B756B0CF8CF" blockId="3.[189,342,1829,1855]" box="[189,342,1829,1855]" pageId="3" pageNumber="4">
<heading id="BE0C275121282F70FF6F4B756B0CF8CF" bold="true" box="[189,342,1829,1855]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="D78F4C2F21282F70FF6F4B756B0CF8CF" bold="true" box="[189,342,1829,1855]" pageId="3" pageNumber="4">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="E544903D21282F70FF6F4B0469F7F89D" blockId="3.[189,941,1875,1902]" box="[189,941,1875,1902]" pageId="3" pageNumber="4">
Known only from northern part of
<collectingRegion id="273F5EDF21282F70FD874B04697BF89D" box="[597,801,1875,1902]" country="China" name="Yunnan" pageId="3" pageNumber="4">Yunnan Province</collectingRegion>
, SW
<collectingCountry id="9DECD0AD21282F70FCB24B0369F0F89D" box="[864,938,1875,1901]" name="China" pageId="3" pageNumber="4">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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