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<document id="A9EA4AC97FB8031E75302E6031A5CB4E" ID-DOI="10.11646/zootaxa.3888.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10086415" ID-ZooBank="1B437AE1-CF28-4C1B-95B6-C31A295905A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623730057475" checkinUser="felipe" docAuthor="Schwarzhans, Werner" docDate="2014" docId="463A8F36FF94FFDD12979D03E3C8FD08" docLanguage="en" docName="zt03888p073.pdf.imf" docOrigin="Zootaxa 3888 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hymenocephalus papyraceus : Okamura 1970" docType="treatment" docVersion="2" lastPageNumber="38" masterDocId="BA03F74EFFB0FFF812009814E16BFFF8" masterDocTitle="Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species" masterLastPageNumber="73" masterPageNumber="1" pageNumber="37" updateTime="1699488604623" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="9C1FB5978352CC52CCDBE5087D12398C">Head and otolith morphology of the genera Hymenocephalus, Hymenogadus and Spicomacrurus (Macrouridae), with the description of three new species</mods:title>
</mods:titleInfo>
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<mods:namePart id="49EA82551F8AD27B2AA2451612D7BF51">Schwarzhans, Werner</mods:namePart>
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<mods:typeOfResource id="0BA05C899A146A77FE72297798D4E96D">text</mods:typeOfResource>
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<mods:title id="DE6357E000433BFAA64C5C59B21FA12D">Zootaxa</mods:title>
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<mods:date id="C080CF7EA63C3EF7031E4F84ACFFEDBD">2014</mods:date>
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<mods:number id="FCBD72F296E5D4121CF205CBD3017BBD">2014-11-28</mods:number>
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<mods:number id="603E7DEEE1DAB7D477A6AA9ED54AC44C">3888</mods:number>
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<mods:classification id="2DE27A89918E6D64C88E8B5BDBB1C16B">journal article</mods:classification>
<mods:identifier id="79828346C660FDAE7F540EB90BD7D076" type="DOI">10.11646/zootaxa.3888.1.1</mods:identifier>
<mods:identifier id="FB37B713E4FEF9E181603045313C0F5C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D6ABF2C25DF5F27E09F2CAA3642AEFEB" type="Zenodo-Dep">10086415</mods:identifier>
<mods:identifier id="696428456F2776F3A74F764F5B843520" type="ZooBank">1B437AE1-CF28-4C1B-95B6-C31A295905A0</mods:identifier>
</mods:mods>
<treatment id="463A8F36FF94FFDD12979D03E3C8FD08" LSID="urn:lsid:plazi:treatment:463A8F36FF94FFDD12979D03E3C8FD08" httpUri="http://treatment.plazi.org/id/463A8F36FF94FFDD12979D03E3C8FD08" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<subSubSection id="86896DABFF94FFDC12979D03E24DFACA" box="[151,806,1303,1330]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="CE2C3E20FF94FFDC12979D03E24DFACA" blockId="36.[151,806,1303,1365]" box="[151,806,1303,1330]" pageId="36" pageNumber="37">
<heading id="9564894CFF94FFDC12979D03E24DFACA" bold="true" box="[151,806,1303,1330]" fontSize="11" level="2" pageId="36" pageNumber="37" reason="4">
<emphasis id="FCE7E232FF94FFDC12979D03E24DFACA" bold="true" box="[151,806,1303,1330]" pageId="36" pageNumber="37">
<taxonomicName id="099345A3FF94FFDC12979D03E092FAC9" ID-CoL="3NJ9J" authorityName=": Okamura" authorityYear="1970" box="[151,505,1303,1329]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="papyraceus">
<emphasis id="FCE7E232FF94FFDC12979D03E092FAC9" bold="true" box="[151,505,1303,1329]" italics="true" pageId="36" pageNumber="37">Hymenocephalus papyraceus</emphasis>
</taxonomicName>
<bibRefCitation id="AA0243D1FF94FFDC10009D0CE24DFACA" author="Jordan, D. S. &amp; Gilbert, C. H." box="[512,806,1304,1330]" pageId="36" pageNumber="37" pagination="577 - 630" refId="ref51264" refString="Jordan, D. S. &amp; Gilbert, C. H. (1904) Macrouridae. In: Jordan, D. S. &amp; Starks, E. C. List of fishes dredged by the streamer Albatross off the coasts of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae. Bulletin of the United States Fishery Commission, 22, 577 - 630." type="journal article" year="1904">
<collectingCountry id="B6847EB0FF94FFDC10009D0CE331FACA" box="[512,602,1304,1330]" name="Jordan" pageId="36" pageNumber="37">Jordan</collectingCountry>
&amp; Gilbert, 1904
</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF94FFDC12979D28E028FAAD" box="[151,323,1340,1365]" pageId="36" pageNumber="37" type="description">
<paragraph id="CE2C3E20FF94FFDC12979D28E028FAAD" blockId="36.[151,806,1303,1365]" box="[151,323,1340,1365]" pageId="36" pageNumber="37">
<figureCitation id="56A822A5FF94FFDC12979D28E070FAAD" box="[151,283,1340,1365]" captionStart="FIGURE 14" captionStartId="35.[151,250,1641,1663]" captionTargetBox="[202,1394,197,1613]" captionTargetId="figure-112@35.[187,1402,189,1622]" captionTargetPageId="35" captionText="FIGURE 14. A–B: Hymenocephalus barbatulus: A—Figure from Gilbert &amp; Hubbs (1920). B—Figure from Cohen et al. (1990). C–E: Hymenocephalus nesaeae: Holotype, MNHN 1995-0869, C—Lateral view of head, D—Dorsal view of head, E—inner face of otolith. F–J: Hymenocephalus papyraceus: F—BSKU 099483, lateral view of head, G—Holotype, USNM 50935, lateral view of head, H—BSKU 099483, dorsal view of head, I—BSKU 44761, inner face of otolith, J—BSKU 44761, ventral view of otolith." figureDoi="http://doi.org/10.5281/zenodo.10086445" httpUri="https://zenodo.org/record/10086445/files/figure.png" pageId="36" pageNumber="37">Figs. 14F–J</figureCitation>
,
<figureCitation id="56A822A5FF94FFDC13279D28E028FAAD" box="[295,323,1340,1365]" captionStart="FIGURE 16" captionStartId="39.[151,250,1970,1992]" captionTargetBox="[165,1418,640,1938]" captionTargetId="figure-204@39.[151,1436,628,1946]" captionTargetPageId="39" captionText="FIGURE 16. Geographical distribution of Hymenocephalus aterrimus, H. barbatulus, H. nesaeae, H. papyraceus and H. sazonovi." figureDoi="http://doi.org/10.5281/zenodo.10086451" httpUri="https://zenodo.org/record/10086451/files/figure.png" pageId="36" pageNumber="37">16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF94FFDC12979D90E3F2FA46" pageId="36" pageNumber="37" type="reference_group">
<paragraph id="CE2C3E20FF94FFDC12979D90E525FA61" blockId="36.[151,1102,1412,1470]" box="[151,1102,1412,1434]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="EE83190EFF94FFDC12979D90E525FA61" box="[151,1102,1412,1434]" pageId="36" pageNumber="37">
<taxonomicName id="099345A3FF94FFDC12979D90E0ABFA61" ID-CoL="3NJ9J" authorityName=": Okamura" authorityYear="1970" box="[151,448,1412,1434]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="papyraceus">
<emphasis id="FCE7E232FF94FFDC12979D90E0ABFA61" box="[151,448,1412,1434]" italics="true" pageId="36" pageNumber="37">Hymenocephalus papyraceus</emphasis>
</taxonomicName>
<materialsCitation id="7EFB347DFF94FFDC13C69D90E295FA61" box="[454,1022,1412,1434]" country="Jordan" location="Sagami Bay" pageId="36" pageNumber="37" specimenCount="1">
<treatmentCitation id="4F321831FF94FFDC13C69D90E38EFA62" author="Jordan, D. S. &amp; Gilbert, C. H." box="[454,741,1412,1434]" page="614" pageId="36" pageNumber="37" year="1904">
<bibRefCitation id="AA0243D1FF94FFDC13C69D90E38EFA62" author="Jordan, D. S. &amp; Gilbert, C. H." box="[454,741,1412,1434]" pageId="36" pageNumber="37" pagination="577 - 630" refId="ref51264" refString="Jordan, D. S. &amp; Gilbert, C. H. (1904) Macrouridae. In: Jordan, D. S. &amp; Starks, E. C. List of fishes dredged by the streamer Albatross off the coasts of Japan in the summer of 1900, with descriptions of new species and a review of the Japanese Macrouridae. Bulletin of the United States Fishery Commission, 22, 577 - 630." type="journal article" year="1904">
<collectingCountry id="B6847EB0FF94FFDC13C69D90E361FA62" box="[454,522,1412,1434]" name="Jordan" pageId="36" pageNumber="37">Jordan</collectingCountry>
&amp; Gilbert, 1904: 614
</bibRefCitation>
</treatmentCitation>
(
<typeStatus id="11288082FF94FFDC10F29D90E274FA62" box="[754,799,1412,1434]" pageId="36" pageNumber="37">type</typeStatus>
locality:
<location id="CB4C68FBFF94FFDC117E9D90E290FA61" LSID="urn:lsid:plazi:treatment:463A8F36FF94FFDD12979D03E3C8FD08:CB4C68FBFF94FFDC117E9D90E290FA61" box="[894,1019,1412,1434]" country="Jordan" name="Sagami Bay" pageId="36" pageNumber="37">Sagami Bay</location>
,
</materialsCitation>
<collectingCountry id="B6847EB0FF94FFDC16049D90E529FA61" box="[1028,1090,1412,1433]" name="Japan" pageId="36" pageNumber="37">Japan</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="CE2C3E20FF94FFDC12979DBCE3F2FA46" blockId="36.[151,1102,1412,1470]" box="[151,665,1448,1470]" pageId="36" pageNumber="37">
<treatmentCitationGroup id="EE83190EFF94FFDC12979DBCE3F2FA46" box="[151,665,1448,1470]" pageId="36" pageNumber="37">
<treatmentCitation id="4F321831FF94FFDC12979DBCE3FFFA46" author="Okamura, O." box="[151,660,1448,1470]" page="56" pageId="36" pageNumber="37" year="1970">
<taxonomicName id="099345A3FF94FFDC12979DBCE3FFFA46" ID-CoL="3NJ9J" authority=": Okamura, 1970: 56" authorityName=": Okamura" authorityPageNumber="56" authorityYear="1970" box="[151,660,1448,1470]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="papyraceus">
<emphasis id="FCE7E232FF94FFDC12979DBCE0ABFA46" box="[151,448,1448,1470]" italics="true" pageId="36" pageNumber="37">Hymenocephalus papyraceus</emphasis>
:
<bibRefCitation id="AA0243D1FF94FFDC13CD9DBCE3FFFA46" author="Okamura, O." box="[461,660,1448,1470]" pageId="36" pageNumber="37" pagination="1 - 216" refId="ref51620" refString="Okamura, O. (1970) Fauna Japonica. Macrourina (Pisces). Academic Press of Japan, Tokyo, pp. 1 - 216." type="book chapter" year="1970">Okamura, 1970: 56</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF94FFDC12979DFBE47EF960" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="CE2C3E20FF94FFDC12979DFBE47EF960" blockId="36.[151,1437,1519,2013]" pageId="36" pageNumber="37">
<emphasis id="FCE7E232FF94FFDC12979DFBE0EAF9F0" bold="true" box="[151,385,1519,1544]" pageId="36" pageNumber="37">Material examined.</emphasis>
<specimenCount id="D895F5A9FF94FFDC138A9DFBE34DF9F0" box="[394,550,1519,1544]" pageId="36" pageNumber="37" type="generic">12 specimens</specimenCount>
;
<materialsCitation id="7EFB347DFF94FFDC10339DFBE051F9D3" collectionCode="BSKU" country="India" elevation="600" location="central Tosa Bay" pageId="36" pageNumber="37" specimenCode="BSKU 44292-44297, 120" specimenCount="6">
<specimenCount id="D895F5A9FF94FFDC10339DFBE3D5F9F0" box="[563,702,1519,1544]" pageId="36" pageNumber="37" type="generic">6 specimens</specimenCount>
<specimenCode id="9E35965BFF94FFDC10C69DFBE2DAF9F0" box="[710,945,1519,1544]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="36" pageNumber="37">BSKU 44292-44297</specimenCode>
,
<specimenCode id="9E35965BFF94FFDC11BE9DFBE283F9F0" box="[958,1000,1519,1544]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="36" pageNumber="37">120</specimenCode>
+ –132+ mm TL,
<location id="CB4C68FBFF94FFDC16B59DFBE41FF9F0" LSID="urn:lsid:plazi:treatment:463A8F36FF94FFDD12979D03E3C8FD08:CB4C68FBFF94FFDC16B59DFBE41FF9F0" box="[1205,1396,1519,1544]" country="India" name="central Tosa Bay" pageId="36" pageNumber="37">central Tosa Bay</location>
off
<collectingCountry id="B6847EB0FF94FFDC12979E00E1B5F9D5" box="[151,222,1556,1581]" name="India" pageId="36" pageNumber="37">Kochi</collectingCountry>
,
<quantity id="096B93C5FF94FFDC12ED9E00E051F9D3" box="[237,314,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="37" unit="m" value="600.0">
<elevation id="45BED913FF94FFDC12ED9E00E051F9D3" box="[237,314,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="37" unit="m" value="600.0">600 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="7EFB347DFF94FFDC13489E00E5DCF9D3" box="[328,1207,1556,1581]" collectionCode="BSKU" country="India" elevation="600" location="central Tosa Bay" pageId="36" pageNumber="37" specimenCount="1">
<specimenCount id="D895F5A9FF94FFDC13489E00E0A1F9D5" box="[328,458,1556,1581]" pageId="36" pageNumber="37" type="generic">1 specimen</specimenCount>
<collectionCode id="A882A6E5FF94FFDC13D49E00E374F9D5" box="[468,543,1556,1581]" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="36" pageNumber="37">BSKU</collectionCode>
<quantity id="096B93C5FF94FFDC10299E00E38BF9D3" box="[553,736,1556,1581]" metricMagnitude="4" metricUnit="m" metricValue="4.476117" pageId="36" pageNumber="37" unit="mm" value="4.476117E7">44761, 170 mm</quantity>
TL,
<location id="CB4C68FBFF94FFDC111E9E00E2B4F9D4" LSID="urn:lsid:plazi:treatment:463A8F36FF94FFDD12979D03E3C8FD08:CB4C68FBFF94FFDC111E9E00E2B4F9D4" box="[798,991,1556,1581]" country="India" name="central Tosa Bay" pageId="36" pageNumber="37">central Tosa Bay</location>
off
<collectingCountry id="B6847EB0FF94FFDC16149E00E530F9D5" box="[1044,1115,1556,1581]" name="India" pageId="36" pageNumber="37">Kochi</collectingCountry>
,
<quantity id="096B93C5FF94FFDC166A9E00E5DCF9D3" box="[1130,1207,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="37" unit="m" value="600.0">
<elevation id="45BED913FF94FFDC166A9E00E5DCF9D3" box="[1130,1207,1556,1581]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="37" unit="m" value="600.0">600 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="7EFB347DFF94FFDC16C59E00E27BF9A8" collectionCode="BSKU" country="Japan" location="Japan" pageId="36" pageNumber="37" specimenCount="1">
<specimenCount id="D895F5A9FF94FFDC16C59E00E423F9D5" box="[1221,1352,1556,1581]" pageId="36" pageNumber="37" type="generic">1 specimen</specimenCount>
<collectionCode id="A882A6E5FF94FFDC17529E00E4F6F9D5" box="[1362,1437,1556,1581]" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="36" pageNumber="37">BSKU</collectionCode>
<quantity id="096B93C5FF94FFDC12979E23E023F9B7" box="[151,328,1591,1616]" metricMagnitude="4" metricUnit="m" metricValue="9.9483171" pageId="36" pageNumber="37" unit="mm" value="9.9483171E7">99483, 171 mm</quantity>
TL, off southern
<collectingCountry id="B6847EB0FF94FFDC100E9E2CE325F9B7" box="[526,590,1592,1615]" name="Japan" pageId="36" pageNumber="37">Japan</collectingCountry>
(no further data)
</materialsCitation>
;
<materialsCitation id="7EFB347DFF94FFDC111B9E23E537F98D" collectionCode="R, V, BSKU" collectorName="Hakuro-maru &amp; St." country="Japan" location="Japan" pageId="36" pageNumber="37" specimenCode="BSKU 101959, NMIC-ZF-03092" specimenCount="1">
<specimenCount id="D895F5A9FF94FFDC111B9E23E2F1F9A8" box="[795,922,1591,1616]" pageId="36" pageNumber="37" type="generic">1 specimen</specimenCount>
<specimenCode id="9E35965BFF94FFDC11A19E23E521F9A8" box="[929,1098,1591,1616]" collectionCode="BSKU" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="36" pageNumber="37">BSKU 101959</specimenCode>
,
<collectionCode id="A882A6E5FF94FFDC16559E23E50CF9A8" box="[1109,1127,1591,1616]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="36" pageNumber="37">R</collectionCode>
/
<collectionCode id="A882A6E5FF94FFDC16709E23E5EFF9A8" box="[1136,1156,1591,1616]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="36" pageNumber="37" type="Museum">V</collectionCode>
<collectorName id="63665BF6FF94FFDC168B9E23E449F9A8" box="[1163,1314,1591,1616]" pageId="36" pageNumber="37">Hakuro-maru</collectorName>
(KH 73-2,
<collectorName id="63665BF6FF94FFDC12979E48E1DEF98D" box="[151,181,1628,1653]" pageId="36" pageNumber="37">St.</collectorName>
4);
<collectionCode id="A882A6E5FF94FFDC12EA9E48E05EF98D" box="[234,309,1628,1653]" country="Japan" httpUri="http://grbio.org/cool/enbt-yj9t" name="Kochi University" pageId="36" pageNumber="37">BSKU</collectionCode>
<specimenCode id="9E35965BFF94FFDC13409E48E36DF98D" box="[320,518,1628,1653]" collectionCode="NMIC-ZF-" pageId="36" pageNumber="37">NMIC-ZF-03092</specimenCode>
,
<quantity id="096B93C5FF94FFDC10159E48E31CF98B" box="[533,631,1628,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.47" pageId="36" pageNumber="37" unit="mm" value="147.0">147 mm</quantity>
TL, off southern
<collectingCountry id="B6847EB0FF94FFDC114D9E48E2E6F98B" box="[845,909,1628,1651]" name="Japan" pageId="36" pageNumber="37">Japan</collectingCountry>
(no further data)
</materialsCitation>
;
<materialsCitation id="7EFB347DFF94FFDC166B9E48E3A9F96F" collectionCode="USNM" country="Japan" elevation="353" location="Sagami Bay" pageId="36" pageNumber="37" specimenCode="USNM 50935" specimenCount="1" typeStatus="holotype">
<specimenCount id="D895F5A9FF94FFDC166B9E48E584F98D" box="[1131,1263,1628,1653]" pageId="36" pageNumber="37" type="generic">1 specimen</specimenCount>
<specimenCode id="9E35965BFF94FFDC16FA9E48E4F7F98D" box="[1274,1436,1628,1653]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="36" pageNumber="37" type="Museum">USNM 50935</specimenCode>
(
<typeStatus id="11288082FF94FFDC129F9E6BE06CF960" box="[159,263,1663,1688]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
),
<quantity id="096B93C5FF94FFDC13199E6BE01EF96F" box="[281,373,1663,1688]" metricMagnitude="-1" metricUnit="m" metricValue="1.1199999999999999" pageId="36" pageNumber="37" unit="mm" value="112.0">112 mm</quantity>
TL,
<location id="CB4C68FBFF94FFDC13AC9E6BE35EF960" LSID="urn:lsid:plazi:treatment:463A8F36FF94FFDD12979D03E3C8FD08:CB4C68FBFF94FFDC13AC9E6BE35EF960" box="[428,565,1663,1688]" country="Japan" name="Sagami Bay" pageId="36" pageNumber="37">Sagami Bay</location>
,
<quantity id="096B93C5FF94FFDC10409E6BE3A9F96F" box="[576,706,1663,1688]" metricMagnitude="2" metricUnit="m" metricValue="3.525" metricValueMax="4.85" metricValueMin="2.2" pageId="36" pageNumber="37" unit="m" value="352.5" valueMax="485.0" valueMin="220.0">
<elevation id="45BED913FF94FFDC10409E6BE3A9F96F" box="[576,706,1663,1688]" metricMagnitude="2" metricUnit="m" metricValue="3.525" metricValueMax="4.85" metricValueMin="2.2" pageId="36" pageNumber="37" unit="m" value="352.5" valueMax="485.0" valueMin="220.0">220–485 m</elevation>
</quantity>
</materialsCitation>
;
<materialsCitation id="7EFB347DFF94FFDC10CE9E6BE47BF960" box="[718,1296,1663,1688]" collectionCode="ZMMGU" country="Japan" latitude="30.5" longLatPrecision="1221" longitude="129.31667" pageId="36" pageNumber="37" specimenCount="1" typeStatus="holotype">
<specimenCount id="D895F5A9FF94FFDC10CE9E6BE226F960" box="[718,845,1663,1688]" pageId="36" pageNumber="37" type="generic">1 specimen</specimenCount>
<collectionCode id="A882A6E5FF94FFDC11549E6BE2D4F960" box="[852,959,1663,1688]" country="Russia" httpUri="http://grbio.org/cool/dvrf-d2q9" name="Zoological Museum" pageId="36" pageNumber="37">ZMMGU</collectionCode>
P-18258,
<geoCoordinate id="ABA758E7FF94FFDC16339E6BE5FDF960" box="[1075,1174,1663,1688]" degrees="30" direction="north" minutes="30" orientation="latitude" pageId="36" pageNumber="37" precision="925" value="30.5">30°30’N</geoCoordinate>
,
<geoCoordinate id="ABA758E7FF94FFDC16A29E6BE47BF960" box="[1186,1296,1663,1688]" degrees="129" direction="east" minutes="19" orientation="longitude" pageId="36" pageNumber="37" precision="925" value="129.31667">129°19’E</geoCoordinate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF94FFDD12C79EB0E3C7FF2D" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="CE2C3E20FF94FFDC12C79EB0E2C2F888" blockId="36.[151,1437,1519,2013]" pageId="36" pageNumber="37">
<emphasis id="FCE7E232FF94FFDC12C79EB0E02AF945" bold="true" box="[199,321,1700,1725]" pageId="36" pageNumber="37">Diagnosis.</emphasis>
Pelvic fin rays 10–11; pectoral fin rays 16–18; snout projecting, with thin, pointed tip, 20–25% HL; barbel short, 4–5% HL, not reaching vertical through anterior margin of orbit; orbit diameter moderate, 25–30% HL; interorbital width 70–80% HW; infraorbital width 16–20% HL; preopercular supporter very long, straight, 10–11% HL; gill rakers 18–22; ventral striae reaching to about ⅓ distance from pelvic fin bases to periproct; otolith with moderately high predorsal lobe, colliculi separated, closely placed across narrow collum, terminating far from anterior and posterior rims of otolith; OL:OH = 1.05; TCL:PCL = 2.1.
</paragraph>
<paragraph id="CE2C3E20FF94FFDD12C79F68E3C7FF2D" blockId="36.[151,1437,1519,2013]" lastBlockId="37.[151,1437,151,752]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
<emphasis id="FCE7E232FF94FFDC12C79F68E009F86D" bold="true" box="[199,354,1916,1941]" pageId="36" pageNumber="37">Comparison.</emphasis>
<taxonomicName id="099345A3FF94FFDC136C9F68E3D3F86C" authorityName=": Okamura" authorityYear="1970" box="[364,696,1916,1941]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="papyraceus">
<emphasis id="FCE7E232FF94FFDC136C9F68E3D3F86C" box="[364,696,1916,1941]" italics="true" pageId="36" pageNumber="37">Hymenocephalus papyraceus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="099345A3FF94FFDC113C9F69E2A5F86C" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[828,974,1917,1940]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="aterrimus">
<emphasis id="FCE7E232FF94FFDC113C9F69E2A5F86C" box="[828,974,1917,1940]" italics="true" pageId="36" pageNumber="37">H. aterrimus</emphasis>
</taxonomicName>
and
<taxonomicName id="099345A3FF94FFDC160B9F69E5F3F86C" box="[1035,1176,1917,1940]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="neasaeae">
<emphasis id="FCE7E232FF94FFDC160B9F69E5F3F86C" box="[1035,1176,1917,1940]" italics="true" pageId="36" pageNumber="37">H. neasaeae</emphasis>
</taxonomicName>
. From
<taxonomicName id="099345A3FF94FFDC16F09F69E4E9F86C" authorityName="Gilbert &amp; Hubbs" authorityYear="1920" box="[1264,1410,1917,1940]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="aterrimus">
<emphasis id="FCE7E232FF94FFDC16F09F69E4E9F86C" box="[1264,1410,1917,1940]" italics="true" pageId="36" pageNumber="37">H. aterrimus</emphasis>
</taxonomicName>
it differs in the number of pelvic and pectoral fin rays (10–11 vs 13–14 and 16–18 vs 13–16, respectively), the smaller preopercular canal width (14–17 vs 19–23% HL), the smaller postorbital-preopercular interspace (4–6 vs 10–14% HL), the long and slender preopercular supporter (11 vs 3–4% HL) and the otolith with a distinct predorsal lobe. For difference with
<taxonomicName id="099345A3FF95FFDD13B498A9E35BFF2C" authorityName="Merrett &amp; Iwamoto" authorityYear="2000" box="[436,560,189,212]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="nesaeae">
<emphasis id="FCE7E232FF95FFDD13B498A9E35BFF2C" box="[436,560,189,212]" italics="true" pageId="37" pageNumber="38">H. nesaeae</emphasis>
</taxonomicName>
see above.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF95FFDD12C798CBE5DCFD50" pageId="37" pageNumber="38" type="description">
<paragraph id="CE2C3E20FF95FFDD12C798CBE5C7FE28" blockId="37.[151,1437,151,752]" pageId="37" pageNumber="38">
<emphasis id="FCE7E232FF95FFDD12C798CBE033FF00" bold="true" box="[199,344,223,248]" pageId="37" pageNumber="38">Description.</emphasis>
Head morphology (n = 3) (
<figureCitation id="56A822A5FF95FFDD108D98CBE27FFF00" box="[653,788,223,248]" captionStart="FIGURE 14" captionStartId="35.[151,250,1641,1663]" captionTargetBox="[202,1394,197,1613]" captionTargetId="figure-112@35.[187,1402,189,1622]" captionTargetPageId="35" captionText="FIGURE 14. A–B: Hymenocephalus barbatulus: A—Figure from Gilbert &amp; Hubbs (1920). B—Figure from Cohen et al. (1990). C–E: Hymenocephalus nesaeae: Holotype, MNHN 1995-0869, C—Lateral view of head, D—Dorsal view of head, E—inner face of otolith. F–J: Hymenocephalus papyraceus: F—BSKU 099483, lateral view of head, G—Holotype, USNM 50935, lateral view of head, H—BSKU 099483, dorsal view of head, I—BSKU 44761, inner face of otolith, J—BSKU 44761, ventral view of otolith." figureDoi="http://doi.org/10.5281/zenodo.10086445" httpUri="https://zenodo.org/record/10086445/files/figure.png" pageId="37" pageNumber="38">Fig. 14F–H</figureCitation>
): Head large, snout projecting, long, with thin pointed tip, which is easily damaged, snout length 20–25% HL, orbit diameter 25–30% HL, interorbital width 70–80% HW. Barbel short, 4–5% HL, not reaching vertical through anterior margin of orbit. Head canals well developed, large, infraorbital width 16–20% HL, supraorbital canal with 4 segments, width 17–20% HL, supratemporal canal rather distinct and wide, above segment 3 of supraorbital canal, preopercular canal width 14–17% HL, postorbitalpreopercular interspace 4–6% HL. Infranasal supporter small; infraorbital supporter short, expanding only beyond rear part of orbit, 40–60% OD; preopercular supporter very long, thin and straight (11% HL).
</paragraph>
<paragraph id="CE2C3E20FF95FFDD12C799C8E5DCFD50" blockId="37.[151,1437,151,752]" pageId="37" pageNumber="38">
Otolith morphology (n = 1) (
<figureCitation id="56A822A5FF95FFDD101B99C8E3FDFE0D" box="[539,662,476,501]" captionStart="FIGURE 14" captionStartId="35.[151,250,1641,1663]" captionTargetBox="[202,1394,197,1613]" captionTargetId="figure-112@35.[187,1402,189,1622]" captionTargetPageId="35" captionText="FIGURE 14. A–B: Hymenocephalus barbatulus: A—Figure from Gilbert &amp; Hubbs (1920). B—Figure from Cohen et al. (1990). C–E: Hymenocephalus nesaeae: Holotype, MNHN 1995-0869, C—Lateral view of head, D—Dorsal view of head, E—inner face of otolith. F–J: Hymenocephalus papyraceus: F—BSKU 099483, lateral view of head, G—Holotype, USNM 50935, lateral view of head, H—BSKU 099483, dorsal view of head, I—BSKU 44761, inner face of otolith, J—BSKU 44761, ventral view of otolith." figureDoi="http://doi.org/10.5281/zenodo.10086445" httpUri="https://zenodo.org/record/10086445/files/figure.png" pageId="37" pageNumber="38">Fig. 14I–J</figureCitation>
): Otolith moderate in size, ventrally well rounded, dorsally with a distinct predorsal lobe, postdorsally somewhat undulating and regularly inclined towards rounded posterior tip; OL:OH = 1.05; OH:OT = 3.5. Inner face slightly convex, with median sulcus. Ostial and caudal colliculi small, narrowly placed across collum, terminating at considerable distance from anterior and posterior tips of otolith; pseudocolliculum moderately short. CCL:OCL = 0.85; TCL:PCL = 2.1. Dorsal depression small; ventral furrow distinct, moderately close to ventral rim, anteriorly and posteriorly turning away and upwards.
</paragraph>
</subSubSection>
<subSubSection id="86896DABFF95FFDD12C79AA0E3C8FD08" pageId="37" pageNumber="38" type="distribution">
<paragraph id="CE2C3E20FF95FFDD12C79AA0E3C8FD08" blockId="37.[151,1437,151,752]" pageId="37" pageNumber="38">
<emphasis id="FCE7E232FF95FFDD12C79AA0E032FD35" bold="true" box="[199,345,692,717]" pageId="37" pageNumber="38">Distribution</emphasis>
(
<figureCitation id="56A822A5FF95FFDD13689AA0E0D0FD35" box="[360,443,692,717]" captionStart="FIGURE 16" captionStartId="39.[151,250,1970,1992]" captionTargetBox="[165,1418,640,1938]" captionTargetId="figure-204@39.[151,1436,628,1946]" captionTargetPageId="39" captionText="FIGURE 16. Geographical distribution of Hymenocephalus aterrimus, H. barbatulus, H. nesaeae, H. papyraceus and H. sazonovi." figureDoi="http://doi.org/10.5281/zenodo.10086451" httpUri="https://zenodo.org/record/10086451/files/figure.png" pageId="37" pageNumber="38">Fig. 16</figureCitation>
).
<taxonomicName id="099345A3FF95FFDD13CE9AA0E27CFD34" authorityName=": Okamura" authorityYear="1970" box="[462,791,692,717]" class="Actinopterygii" family="Macrouridae" genus="Hymenocephalus" kingdom="Animalia" order="Gadiformes" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="papyraceus">
<emphasis id="FCE7E232FF95FFDD13CE9AA0E27CFD34" box="[462,791,692,717]" italics="true" pageId="37" pageNumber="38">Hymenocephalus papyraceus</emphasis>
</taxonomicName>
is only known off the southeastern coast of
<collectingCountry id="B6847EB0FF95FFDD17019AA0E42FFD33" box="[1281,1348,692,715]" name="Japan" pageId="37" pageNumber="38">Japan</collectingCountry>
, chiefly the Sagami and Tosa Bays and off
<collectingRegion id="0C57F0C2FF95FFDD101C9AC3E3F4FD08" box="[540,671,727,752]" country="Japan" name="Kagoshima" pageId="37" pageNumber="38">Kagoshima</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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