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<document id="516C4C75B7F430BB9B643A2E1A2EC00A" ID-DOI="10.15553/c2012v672a12" ID-ISSN="2235-3658" ID-Zenodo-Dep="5790002" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637611591098" checkinUser="carolina" docAuthor="Callmander, Martin W., Lowry Ii, Porter P., Forest, Félix, Devey, Dion S., Beentje, Henk &amp; Buerki, Sven" docDate="2012" docId="03F20C491D142B4E5E06FADF76815225" docLanguage="en" docName="Candollea.67.2.323-345.pdf.imf" docOrigin="Candollea 67 (2)" docStyle="DocumentStyle:FB35ED2BCB231A5F446DBDA23BDBADFC.2:Candollea.2012-2014.journal_article" docStyleId="FB35ED2BCB231A5F446DBDA23BDBADFC" docStyleName="Candollea.2012-2014.journal_article" docStyleVersion="2" docTitle="Benstonea copelandii Callm. &amp; Buerki, comb. nova." docType="treatment" docVersion="10" lastPageNumber="333" masterDocId="FFCB74311D1D2B445D3AFFF3771C5134" masterDocTitle="Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species" masterLastPageNumber="345" masterPageNumber="323" pageNumber="332" updateTime="1699425018902" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="9B9DF8B4BECD56367AF402BE4E140DAF">Benstonea Callm. &amp; Buerki (Pandanaceae): characterization, circumscription, and distribution of a new genus of screw-pines, with a synopsis of accepted species</mods:title>
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<mods:affiliation id="0F510867FF328CAF35E45B5D36A43FB7">Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Conservatoire et Jardin botaniques de la Ville de Genève, CP 60, CH- 1292 Chambésy, Switzerland.</mods:affiliation>
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<mods:affiliation id="7D515716FE41A223CD27AA60D11DBCCA">Missouri Botanical Garden, P. O. Box 299, St. Louis, MO, 63166 - 0299, U. S. A. and Département Systématique et Evolution (UMR 7205), Muséum National dHistoire Naturelle, case postale 39, rue Cuvier 57, 75231 Paris (Cedex) 05, France.</mods:affiliation>
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<mods:affiliation id="5C6B1D5D2C8A1012D034E22ADEC02E34">JodrellLaboratory, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 DS, United Kingdom.</mods:affiliation>
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<mods:title id="9CF2C31EC815FC5C8CC282223652A539">Candollea</mods:title>
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13.
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<emphasis id="B92F614D1D142B4D5E50FADF73425470" bold="true" box="[874,1118,1324,1348]" italics="true" pageId="9" pageNumber="332">Benstonea copelandii</emphasis>
(Merr.) Callm. &amp; Buerki
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,
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<treatmentCitation id="0AFA9B4E1D142B4D5EB2FA807316549F" author="Merr." journal="Publ. Bur. Sci. Gov. Lab." page="7" pageId="9" pageNumber="332" score="0.039372851653953024" volume="17" year="1904">
<taxonomicName id="4C5BC6DC1D142B4D5EB2FA8073B854BF" ID-CoL="6TLBW" authority="Merr." authorityName="Merr." box="[904,1188,1395,1419]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="species" species="copelandii">
<emphasis id="B92F614D1D142B4D5EB2FA80737954BF" box="[904,1125,1395,1419]" italics="true" pageId="9" pageNumber="332">Pandanus copelandii</emphasis>
Merr.
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in Publ. Bur. Sci. Gov. Lab. 17: 7. 1904.
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.
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:
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140
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(
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destroyed
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;
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(
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):
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,
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, fr.,
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<collectorName id="26AED8891D142B4D5801F9D07260570F" box="[1339,1404,1571,1595]" pageId="9" pageNumber="332">Elmer</collectorName>
15267
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(
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<typeStatus id="54E003FD1D142B4D590BF9B0739F576F" box="[1073,1155,1603,1627]" pageId="9" pageNumber="332" type="isoneotype">isoneo-</typeStatus>
:
<collectionCode id="ED4A259A1D142B4D59B7F9B073B9576F" box="[1165,1189,1603,1627]" country="Italy" name="Natural History Museum" pageId="9" pageNumber="332" type="Herbarium">FI</collectionCode>
[
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,
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!,
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[
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[
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[
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</paragraph>
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=
<treatmentCitation id="0AFA9B4E1D142B4D5EB2F959737157D6" author="B. C. Stone" page="25" pageId="9" pageNumber="332" year="1978">
<taxonomicName id="4C5BC6DC1D142B4D5EB2F959726A57F6" ID-CoL="5QFRB" authority="B. C. Stone" authorityName="B. C. Stone" box="[904,1398,1706,1730]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="variety" species="copelandii" variety="panchoi">
<emphasis id="B92F614D1D142B4D5EB2F959737657F6" box="[904,1130,1706,1730]" italics="true" pageId="9" pageNumber="332">Pandanus copelandii</emphasis>
var.
<emphasis id="B92F614D1D142B4D59A4F95973E857F6" box="[1182,1268,1706,1730]" italics="true" pageId="9" pageNumber="332">panchoi</emphasis>
B. C. Stone
</taxonomicName>
in Fed. Mus. J. 23: 25. 1978.
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D142B4D594EF93973CA5616" pageId="9" pageNumber="332" type="materials_examined">
<paragraph id="8BE4BD5F1D142B4D594EF93973CA5616" blockId="9.[874,1481,1467,1826]" pageId="9" pageNumber="332">
<materialsCitation id="3B33B7021D142B4D594EF93973CA5616" collectingDate="1960-10-25" collectionCode="CAHP" collectorName="Pancho &amp; Bardenas" country="Philippines" location="Gubat" municipality="Gubat" pageId="9" pageNumber="332" specimenCount="1" stateProvince="Sorsogon Prov." typeStatus="holotype">
<emphasis id="B92F614D1D142B4D594EF939732F5636" bold="true" pageId="9" pageNumber="332">
Typus:
<collectingCountry id="F34CFDCF1D142B4D59F1F939725357D5" box="[1227,1359,1738,1762]" name="Philippines" pageId="9" pageNumber="332">PHILIPPINES</collectingCountry>
.
<collectingRegion id="499F73BD1D142B4D586EF93972DC57D6" box="[1364,1472,1738,1762]" pageId="9" pageNumber="332">Sorsogon</collectingRegion>
Prov. (
<collectingRegion id="499F73BD1D142B4D5EE8F91973395636" box="[978,1061,1770,1794]" pageId="9" pageNumber="332">Luzon</collectingRegion>
):
</emphasis>
<collectingMunicipality id="6B8027251D142B4D597DF91973915636" box="[1095,1165,1770,1794]" pageId="9" pageNumber="332">Gubat</collectingMunicipality>
,
<date id="FFE59B9F1D142B4D59A7F91972085636" box="[1181,1300,1770,1794]" pageId="9" pageNumber="332" value="1960-10-25">
<collectingDate id="EFA162771D142B4D59A7F91972085636" box="[1181,1300,1770,1794]" pageId="9" pageNumber="332" value="1960-10-25">25.X.1960</collectingDate>
</date>
, fr.,
<emphasis id="B92F614D1D142B4D586BF91973355616" italics="true" pageId="9" pageNumber="332">
<collectorName id="26AED8891D142B4D586BF91972B95636" box="[1361,1445,1770,1794]" pageId="9" pageNumber="332">Pancho</collectorName>
&amp;
<collectorName id="26AED8891D142B4D5EB2F8F974F15616" box="[904,1005,1802,1826]" pageId="9" pageNumber="332">Bardenas</collectorName>
4820
</emphasis>
(
<typeStatus id="54E003FD1D142B4D5902F8F973695616" box="[1080,1141,1802,1826]" pageId="9" pageNumber="332" type="holotype">holo-</typeStatus>
:
<collectionCode id="ED4A259A1D142B4D5945F8F973D65616" box="[1151,1226,1802,1826]" pageId="9" pageNumber="332">CAHP</collectionCode>
),
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D142B4D59E6F8F972225616" box="[1244,1342,1802,1826]" pageId="9" pageNumber="332" type="multiple">
<paragraph id="8BE4BD5F1D142B4D59E6F8F972225616" blockId="9.[874,1481,1467,1826]" box="[1244,1342,1802,1826]" pageId="9" pageNumber="332">
<emphasis id="B92F614D1D142B4D59E6F8F972225616" bold="true" box="[1244,1342,1802,1826]" pageId="9" pageNumber="332">
<taxonomicNameLabel id="A21CDC361D142B4D59E6F8F972225616" box="[1244,1342,1802,1826]" pageId="9" pageNumber="332">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D142B4D5E50F8C172D6565E" pageId="9" pageNumber="332" type="distribution">
<paragraph id="8BE4BD5F1D142B4D5E50F8C172D6565E" blockId="9.[828,1482,1842,1898]" pageId="9" pageNumber="332">
<emphasis id="B92F614D1D142B4D5E50F8C1739B567E" box="[874,1159,1842,1866]" italics="true" pageId="9" pageNumber="332">Distribution and ecology</emphasis>
.
<taxonomicName id="4C5BC6DC1D142B4D598EF8C172B4567E" box="[1204,1448,1842,1866]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="9" pageNumber="332" phylum="Tracheophyta" rank="species" species="copelandii">
<emphasis id="B92F614D1D142B4D598EF8C172B4567E" box="[1204,1448,1842,1866]" italics="true" pageId="9" pageNumber="332">Benstonea copelandii</emphasis>
</taxonomicName>
is endemic to the
<collectingCountry id="F34CFDCF1D142B4D5EE6F8A1734C565E" box="[988,1104,1874,1898]" name="Philippines" pageId="9" pageNumber="332">Philippines</collectingCountry>
(ranging from Luzon to Mindanao).
</paragraph>
</subSubSection>
<subSubSection id="C341EED41D172B4E5D99FEEA76815225" pageId="10" pageNumber="333" type="discussion">
<paragraph id="8BE4BD5F1D172B4E5D99FEEA76815225" blockId="10.[118,769,281,785]" pageId="10" pageNumber="333">
<emphasis id="B92F614D1D172B4E5D99FEEA764D5005" box="[163,337,281,305]" italics="true" pageId="10" pageNumber="333">Observations. </emphasis>
The
<typeStatus id="54E003FD1D172B4E5CB1FEEA76F25005" box="[395,494,281,305]" pageId="10" pageNumber="333" type="holotype">holotype</typeStatus>
, deposited at the herbarium in Manila (
<collectingCountry id="F34CFDCF1D172B4E5C1AFECA76405065" box="[288,348,313,337]" name="Papua New Guinea" pageId="10" pageNumber="333">PNG</collectingCountry>
), was destroyed during World War II, but a fragment has been located in FI. The original collection was rather incomplete, comprising only a few drupes, which were illustrated in
<bibRefCitation id="EFCAC0AE1D172B4E5CB4FE6A75565085" author="MARTELLI" box="[398,586,409,433]" firstAuthor="MARTELLI" journalOrPublisher="Webbia" pageId="10" pageNumber="333" pagination="399 - 435" part="4" refId="ref15813" refString="MARTELLI, U. (1914). Le specie e varieta nuove di &quot; Pandanus &quot; menzionate nella enumerazione delle Pandanaceae. Webbia 4: 399 - 435." title="Le specie e varieta nuove di &quot; Pandanus &quot; menzionate nella enumerazione delle Pandanaceae" type="journal article" year="1914">MARTELLI (1914</bibRefCitation>
: Tab. 31, Fig. 5- 7) and
<bibRefCitation id="EFCAC0AE1D172B4E5D85FE4A767850E5" author="ST. JOHN, H." box="[191,356,441,465]" journalOrPublisher="Pacific Sci." pageId="10" pageNumber="333" pagination="355 - 366" part="23" refId="ref15986" refString="ST. JOHN, H. (1969). Revision of the genus Pandanus Stickman, Part 34. Four species from the Philippines. Pacific Sci. 23: 355 - 366." title="Revision of the genus Pandanus Stickman, Part 34. Four species from the Philippines" type="journal article" year="1969">
ST.
<bibRefCitation id="EFCAC0AE1D172B4E5DDFFE4A767850E5" author="JOHN" box="[229,356,441,465]" firstAuthor="JOHN" journalOrPublisher="Pacific Sci." page="357" pageId="10" pageNumber="333" pagination="355 - 366" part="23" refId="ref15986" refString="ST. JOHN, H. (1969). Revision of the genus Pandanus Stickman, Part 34. Four species from the Philippines. Pacific Sci. 23: 355 - 366." title="Revision of the genus Pandanus Stickman, Part 34. Four species from the Philippines" type="journal article" year="1969">JOHN (1969</bibRefCitation>
</bibRefCitation>
: 357, Fig. 295). We have not been able to locate the duplicate material at FI despite an extensive search.
<typeStatus id="54E003FD1D172B4E5DFCFE0A762E5325" box="[198,306,505,529]" pageId="10" pageNumber="333">Aneotype</typeStatus>
<emphasis id="B92F614D1D172B4E5C03FE0A76CF5325" box="[313,467,505,529]" italics="true" pageId="10" pageNumber="333">(Elmer 15267)</emphasis>
is therefore designated here. This collection is complete and well represented in several herbaria.
<taxonomicName id="4C5BC6DC1D172B4E5DE6FDCA75525365" box="[220,590,569,593]" class="Liliopsida" family="Pandanaceae" genus="Pandanus" kingdom="Plantae" order="Pandanales" pageId="10" pageNumber="333" phylum="Tracheophyta" rank="variety" species="copelandii" variety="panchoi">
<emphasis id="B92F614D1D172B4E5DE6FDCA76DF5365" box="[220,451,569,593]" italics="true" pageId="10" pageNumber="333">Pandanus copelandii</emphasis>
var.
<emphasis id="B92F614D1D172B4E5CCDFDCA75525365" box="[503,590,569,593]" italics="true" pageId="10" pageNumber="333">panchoi</emphasis>
</taxonomicName>
is considered as a synonym. It was described on the basis of a collection bearing only a single syncarp (vs 3-8 crowded, spicate syncarp in the remaining collections of
<taxonomicName id="4C5BC6DC1D172B4E5CFFFD6A75B25385" box="[453,686,665,689]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="10" pageNumber="333" phylum="Tracheophyta" rank="species" species="copelandii">
<emphasis id="B92F614D1D172B4E5CFFFD6A75B25385" box="[453,686,665,689]" italics="true" pageId="10" pageNumber="333">Benstonea copelandii</emphasis>
</taxonomicName>
), but is considered to fall within the morphological variation of the species, even though it is rather atypical and is documented only by a single collection.
</paragraph>
</subSubSection>
</treatment>
</document>