194 lines
20 KiB
XML
194 lines
20 KiB
XML
<document id="33730E9A05068947D7BB46CED81EE959" ID-DOI="10.5281/zenodo.186542" ID-GBIF-Dataset="cc2e1d8e-63c7-4b6e-95de-e415a22ccfe9" ID-ISSN="1175-5326" ID-Zenodo-Dep="186542" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461052940062" checkinUser="plazi" docAuthor="Wagner, Philipp, Böhme, Wolfgang, Pauwels, Olivier S. G. & Schmitz, Andreas" docDate="2009" docId="03BC72486944B818FE93FE132C9C9751" docLanguage="en" docName="zt02050p030.pdf" docOrigin="Zootaxa 2050" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lepidothyris hinkeli subsp. joei Wagner, Böhme, Pauwels & Schmitz, 2009, ssp. n." docType="treatment" docVersion="10" lastPageNumber="16" masterDocId="FF850A306949B817FE04FFE72E649622" masterDocTitle="A review of the African red – flanked skinks of the Lygosoma fernandi (BURTON, 1836) species group (Squamata: Scincidae) and the role of climate change in their speciation." masterLastPageNumber="30" masterPageNumber="1" pageNumber="14" updateTime="1698580187426" updateUser="plazi">
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<mods:titleInfo id="410FB820AADA65EE57BE5879D7F0DF2E">
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<mods:title id="4972120E40707AADAB3EA450376641C7">A review of the African red – flanked skinks of the Lygosoma fernandi (BURTON, 1836) species group (Squamata: Scincidae) and the role of climate change in their speciation.</mods:title>
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<mods:name id="9C5754CE819D1160E121E59F98B202AF" type="personal">
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<mods:roleTerm id="CC7E883755D081A8AC2ABC33700DCA93">Author</mods:roleTerm>
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<mods:namePart id="10B1D67030D4CA4952CBBF29C0964D4D">Wagner, Philipp</mods:namePart>
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<mods:namePart id="7CEEFBE0D64FC7D4A2D095656E88F149">Böhme, Wolfgang</mods:namePart>
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</mods:name>
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<mods:namePart id="0A7E03D645A67D50B582B84BBA4BD24D">Pauwels, Olivier S. G.</mods:namePart>
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</mods:name>
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<mods:name id="72F00F109B8D15700812DBADFE717B08" type="personal">
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<mods:roleTerm id="DEB22D879F18DEE885CC986D570298F9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E6638FEA5E4CAA142578EDC14247F3D2">Schmitz, Andreas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="0493E7897ED5EC8DB8B1942772BEBE33">text</mods:typeOfResource>
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<mods:title id="8FB957EC22E10AC21BBB2692F0574D45">Zootaxa</mods:title>
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<mods:part id="109F6F606F6ECE73DD86D61CBEB3379D">
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<mods:date id="561560EBAB3A0A2971A3EB383E6905BD">2009</mods:date>
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<mods:detail id="82D5DFBC270CD6D3F0FDB623BD112567" type="volume">
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<mods:number id="761DD90ED6A582302ACFBDAC34E2501F">2050</mods:number>
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<mods:classification id="151A3E015FFAA36376B88064D6F8DEF4">journal article</mods:classification>
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<mods:identifier id="2D95ABA88045BED95AA6ADB81F3DC427" type="DOI">10.5281/zenodo.186542</mods:identifier>
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<mods:identifier id="C261F4AC7B5D20D2977D481E1D21105A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03BC72486944B818FE93FE132C9C9751" ID-DOI="http://doi.org/10.5281/zenodo.5678614" ID-GBIF-Taxon="119612316" ID-Zenodo-Dep="5678614" LSID="urn:lsid:plazi:treatment:03BC72486944B818FE93FE132C9C9751" httpUri="http://treatment.plazi.org/id/03BC72486944B818FE93FE132C9C9751" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
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<subSubSection id="C30F90D56944B81AFE93FE132C76942F" box="[151,530,499,526]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8BAAC35E6944B81AFE93FE132C76942F" blockId="13.[151,530,499,526]" box="[151,530,499,526]" pageId="13" pageNumber="14">
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<heading id="D0E274326944B81AFE93FE132C76942F" bold="true" box="[151,530,499,526]" fontSize="11" level="1" pageId="13" pageNumber="14" reason="1">
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<emphasis id="B9611F4C6944B81AFE93FE132C76942F" bold="true" box="[151,530,499,526]" pageId="13" pageNumber="14">
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<taxonomicName id="4C15B8DD6944B81AFE93FE132FD9942C" box="[151,445,500,526]" class="Reptilia" family="Scincidae" genus="Lepidothyris" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="hinkeli" status="subsp. nov." subSpecies="joei">
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<emphasis id="B9611F4C6944B81AFE93FE132FD9942C" bold="true" box="[151,445,500,526]" italics="true" pageId="13" pageNumber="14">Lepidothyris hinkeli joei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A252A2376944B81AFFC1FE142C76942F" box="[453,530,499,525]" pageId="13" pageNumber="14" rank="subSpecies">ssp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F90D56944B81AFE93FDDA2B229486" pageId="13" pageNumber="14" type="materials_examined">
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<paragraph id="8BAAC35E6944B81AFE93FDDA2A9A945C" blockId="13.[151,1436,573,1102]" pageId="13" pageNumber="14">
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<typeStatus id="54AE7DFC6944B81AFE93FDDA2F6C9475" box="[151,264,573,599]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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:
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<collectionCode id="ED045B9B6944B81AFF17FDDA2F019475" LSID="urn:lsid:biocol.org:col:34613" box="[275,357,573,599]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="13" pageNumber="14">ZFMK</collectionCode>
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64410 Oyo, Bokouelé, Peoples Rep.
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<collectingCountry id="F30283CE6944B81AFD19FDDA2D0A9475" box="[797,878,573,599]" name="Republic of the Congo" pageId="13" pageNumber="14">Congo</collectingCountry>
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, leg. E. Fischer & H.Hinkel, 1993.
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<typeStatus id="54AE7DFC6944B81AFE93FD832F6A945C" box="[151,270,612,638]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
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:
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<collectionCode id="ED045B9B6944B81AFF1CFD832F15945C" box="[280,369,612,638]" pageId="13" pageNumber="14">MNHG</collectionCode>
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2539.71 Kasai–Occidental, Dekese, Dem. Rep.
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<collectingCountry id="F30283CE6944B81AFDADFD832D9E945C" box="[937,1018,612,638]" name="Republic of the Congo" pageId="13" pageNumber="14">Congo</collectingCountry>
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, leg. S. Battoni, 1959.
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</paragraph>
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<paragraph id="8BAAC35E6944B81AFEC2FD6D2B229486" blockId="13.[151,1436,573,1102]" box="[198,1350,650,676]" pageId="13" pageNumber="14">
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<collectionCode id="ED045B9B6944B81AFEC2FD6D2F789486" box="[198,284,650,676]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="14">IRSNB</collectionCode>
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2644 Yangambi, Distr. Stanleyville,
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<collectingCountry id="F30283CE6944B81AFCC9FD6D2D039486" box="[717,871,650,676]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">Congo Belge</collectingCountry>
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(=
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<collectingCountry id="F30283CE6944B81AFD82FD6D2A689486" box="[902,1036,650,676]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">DR Congo</collectingCountry>
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), leg. G. Gilert,
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<date id="FFABE59E6944B81AFAC1FD6D2B229486" box="[1221,1350,650,676]" pageId="13" pageNumber="14" value="1946-04-18">18.IV.1946</date>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F90D56944B81AFEC2FD562B6794D0" pageId="13" pageNumber="14" type="description">
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<paragraph id="8BAAC35E6944B81AFEC2FD562B6794E9" blockId="13.[151,1436,573,1102]" box="[198,1283,689,715]" pageId="13" pageNumber="14">
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IRSNB 2645 Ibembo (forêt), Uele,
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<collectingCountry id="F30283CE6944B81AFC62FD562C9B94E9" box="[614,767,689,715]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">Congo Belge</collectingCountry>
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(=
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<collectingCountry id="F30283CE6944B81AFD1BFD562DC794E9" box="[799,931,689,715]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">DR Congo</collectingCountry>
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), leg. H. Siméons,
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<date id="FFABE59E6944B81AFA86FD562A9B94E9" box="[1154,1279,689,715]" pageId="13" pageNumber="14" value="1940-02-20">20.II.1940</date>
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.
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</paragraph>
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<paragraph id="8BAAC35E6944B81AFEC2FD3F2B6794D0" blockId="13.[151,1436,573,1102]" box="[198,1283,728,754]" pageId="13" pageNumber="14">
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IRSNB 2646 Ibembo (forêt), Uele,
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<collectingCountry id="F30283CE6944B81AFC62FD3F2C9B94D0" box="[614,767,728,754]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">Congo Belge</collectingCountry>
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(=
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<collectingCountry id="F30283CE6944B81AFD1BFD3F2DC794D0" box="[799,931,728,754]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">DR Congo</collectingCountry>
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), leg. H. Siméons,
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<date id="FFABE59E6944B81AFA86FD3F2A9B94D0" box="[1154,1279,728,754]" pageId="13" pageNumber="14" value="1940-02-20">20.II.1940</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F90D56944B81AFEC2FD192A479205" pageId="13" pageNumber="14" type="diagnosis">
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<paragraph id="8BAAC35E6944B81AFEC2FD192EBC9591" blockId="13.[151,1436,573,1102]" pageId="13" pageNumber="14">
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<emphasis id="B9611F4C6944B81AFEC2FD192F2C953A" bold="true" box="[198,328,766,792]" pageId="13" pageNumber="14">Diagnosis:</emphasis>
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The new subspecies differs from the nominotypic form in having a monochromatic back, without lateral bars continuing on the back. Lateral bars in most cases smaller and lower in number than in the nominate form, not bordered with white dots; lower lip is lighter than in
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<taxonomicName id="4C15B8DD6944B81AFA02FCAB2AB59547" box="[1030,1233,844,869]" class="Reptilia" family="Scincidae" genus="Lepidothyris" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="hinkeli" subSpecies="hinkeli">
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<emphasis id="B9611F4C6944B81AFA02FCAB2AB59547" box="[1030,1233,844,869]" italics="true" pageId="13" pageNumber="14">L. hinkeli hinkeli</emphasis>
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</taxonomicName>
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and without any black markings on the first sublabial scales; body scales in most cases stronger keeled than in the nominate form.
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</paragraph>
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<paragraph id="8BAAC35E6944B81AFEC1FC582A479205" blockId="13.[151,1436,573,1102]" pageId="13" pageNumber="14">
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<emphasis id="B9611F4C6944B81AFEC1FC582F0795FB" bold="true" box="[197,355,959,985]" pageId="13" pageNumber="14">Colouration.</emphasis>
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Throat and belly white, without any markings; dark framed light patches on the sides of the head between ear opening and forelimb, dark framing is lacking towards ventrally; laterally between the limbs there are eight to eleven dark stripes, not reaching the dorsal side of the body.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F90D56944B818FEC1FBD42A069704" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
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<paragraph id="8BAAC35E6944B81AFEC1FBD42B02926C" blockId="13.[151,1436,573,1102]" box="[197,1382,1075,1102]" pageId="13" pageNumber="14">
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<emphasis id="B9611F4C6944B81AFEC1FBD42E9A926F" bold="true" box="[197,254,1075,1101]" pageId="13" pageNumber="14">Size.</emphasis>
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The maximum snout–vent length of the analysed specimens is 146.0 mm, average is
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<quantity id="4CED6EBB6944B81AFAEDFBD32B06926C" box="[1257,1378,1076,1102]" metricMagnitude="-1" metricUnit="m" metricValue="1.155" pageId="13" pageNumber="14" unit="mm" value="115.5">115.5 mm</quantity>
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.
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</paragraph>
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<caption id="DF6A93D66944B81AFE93F90D2DB59123" box="[151,977,1770,1794]" httpUri="https://zenodo.org/record/186548/files/figure.png" pageId="13" pageNumber="14" targetBox="[530,1057,1159,1744]" targetPageId="13">
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<paragraph id="8BAAC35E6944B81AFE93F90D2DB59123" blockId="13.[151,977,1770,1794]" box="[151,977,1770,1794]" pageId="13" pageNumber="14">
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<emphasis id="B9611F4C6944B81AFE93F90D2F7E9120" bold="true" box="[151,282,1770,1794]" pageId="13" pageNumber="14">FIGURE 8.</emphasis>
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<taxonomicName id="4C15B8DD6944B81AFF24F90D2C409123" box="[288,548,1770,1793]" class="Reptilia" family="Scincidae" genus="Lepidothyris" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="hinkeli" subSpecies="joei">
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<emphasis id="B9611F4C6944B81AFF24F90D2C409123" box="[288,548,1770,1793]" italics="true" pageId="13" pageNumber="14">Lepidothyris hinkeli joei</emphasis>
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</taxonomicName>
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, original published in LAURENT (1964).
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</paragraph>
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</caption>
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<paragraph id="8BAAC35E6944B819FEC2F8D42D429416" blockId="13.[151,1436,1843,2024]" lastBlockId="14.[151,1437,152,796]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<emphasis id="B9611F4C6944B81AFEC2F8D42F9E916F" bold="true" box="[198,506,1843,1869]" pageId="13" pageNumber="14">
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Description of
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<typeStatus id="54AE7DFC6944B81AFF7AF8D42F90916F" box="[382,500,1843,1869]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
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||
:
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</emphasis>
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Snout–vent length:
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<quantity id="4CED6EBB6944B81AFCE3F8D42D3B916F" box="[743,863,1843,1869]" metricMagnitude="-1" metricUnit="m" metricValue="1.119" pageId="13" pageNumber="14" unit="mm" value="111.9">111.9 mm</quantity>
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, tail is broken behind
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<quantity id="4CED6EBB6944B81AFA66F8D42AA9916F" box="[1122,1229,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="3.8899999999999997" pageId="13" pageNumber="14" unit="mm" value="38.9">38.9 mm</quantity>
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, head length
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<quantity id="4CED6EBB6944B81AFB6CF8D42EAC9156" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="13" pageNumber="14" unit="mm" value="23.4">23.4 mm</quantity>
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, head wide
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<quantity id="4CED6EBB6944B81AFF53F8BD2FA09156" box="[343,452,1882,1908]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="13" pageNumber="14" unit="mm" value="15.2">15.2 mm</quantity>
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, head height
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<quantity id="4CED6EBB6944B81AFC60F8BD2CB69156" box="[612,722,1882,1908]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="13" pageNumber="14" unit="mm" value="10.8">10.8 mm</quantity>
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. Body stout, sub–cylindrical; there are 36 scales around the body and 65 longitudinal ventral scales; limbs short and strong with five toes, dark above, light below; 4th fingers with eleven digits on both sides, 4th toes with 15 digits on the left and 16 digits on the right side; ear opening visible, oval to round, with two small ear lobules, little less in diameter than half of the length of the eye opening, little less broad than high, tympanum visible; lower eyelid with two scale rows; dorsal body scales moderately keeled with three keels; lateral body scales smooth to moderately keeled with three small keels; dorsal tail scales moderately keeled to smooth with three keels, lateral tail scales relatively smooth with three keels; head not distinctly set off from body; eye opening as long as the distance between the posterior edge of the eye to the ear opening; distance from tip of snout to anterior edge of eye little shorter than distance from posterior edge of eye to ear; occipital scale lacking; parietal organ set in the posterior edge of the interparietal scale; frontal scale a bit longer than broad; two frontoparietal and prefrontal scales at the anterior side of the frontal scale; one frontonasal scale; two internasal scales; rostral scale broader than high; two nasal scales equal in size, nostril in the middle of the scales; two loreal scales, equal in size; two preocular scales; four supraocular scales; eight supralabial and sublabial scales on each side; mental scale broader than high; three pairs of submaxillary scales, the first one is entire.
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</paragraph>
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<paragraph id="8BAAC35E6947B819FEC1FDA72B12953E" blockId="14.[151,1437,152,796]" pageId="14" pageNumber="15">
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<emphasis id="B9611F4C6947B819FEC1FDA72F099478" bold="true" box="[197,365,576,602]" pageId="14" pageNumber="15">Colouration.</emphasis>
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Head and back red to red–brown; on the back no darker bars visible; supralabial and sublabial scales the same colour as the head; lateral body colouration between ear opening and forelimb light, framed by a darker undulate bar on the upper side from ear opening to the forelimb; ground colour of the lateral body sides is red to red–brown, contrary to the
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<typeStatus id="54AE7DFC6947B819FD0BFD522D1094ED" box="[783,884,693,719]" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
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series only a few dark bars are visible between the limbs (artefact due to preservation); incomplete tail light red to red–brown, also an artefact due to preservation, without any markings or darker bars; belly and throat whitish to reddish, without any stripes.
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</paragraph>
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<caption id="DF6A93D66947B819FE93F85F2CD691F2" box="[151,690,1976,2000]" pageId="14" pageNumber="15">
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<paragraph id="8BAAC35E6947B819FE93F85F2CD691F2" blockId="14.[151,690,1976,2000]" box="[151,690,1976,2000]" pageId="14" pageNumber="15">
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<emphasis id="B9611F4C6947B819FE93F85F2F7D91F2" bold="true" box="[151,281,1976,2000]" pageId="14" pageNumber="15">FIGURE 9.</emphasis>
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Holotype of
|
||
<taxonomicName id="4C15B8DD6947B819FFA2F85E2CCF91F2" box="[422,683,1977,2000]" class="Reptilia" family="Scincidae" genus="Lepidothyris" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="subSpecies" species="hinkeli" subSpecies="joei">
|
||
<emphasis id="B9611F4C6947B819FFA2F85E2CCF91F2" box="[422,683,1977,2000]" italics="true" pageId="14" pageNumber="15">Lepidothyris hinkeli joei</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAAC35E6946B818FEC2FF702A069704" blockId="15.[151,1436,151,371]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9611F4C6946B818FEC2FF702F959693" bold="true" box="[198,497,151,177]" pageId="15" pageNumber="16">
|
||
Variation of
|
||
<typeStatus id="54AE7DFC6946B818FF68FF702F889693" box="[364,492,151,177]" pageId="15" pageNumber="16" type="paratype">paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="54AE7DFC6946B818FC32FF7F2CFF9690" box="[566,667,152,178]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
|
||
series agrees in morphology with the given description of the
|
||
<typeStatus id="54AE7DFC6946B818FE93FF592F6696FA" box="[151,258,190,216]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
. They differ slightly in aspects of the colouration: 8 to 11 small to medium sized black stripes on the lateral body sides, stripes not bordered with dots; the not autotomized tails have broader lateral stripes than the body, stripes cross the ventral part of the tail; incomplete tail diffuse black and red.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F90D56946B818FEC2FED52C9C9751" pageId="15" pageNumber="16" type="etymology">
|
||
<paragraph id="8BAAC35E6946B818FEC2FED52C9C9751" blockId="15.[151,1436,151,371]" pageId="15" pageNumber="16">
|
||
<emphasis id="B9611F4C6946B818FEC2FED52F33976E" bold="true" box="[198,343,306,332]" pageId="15" pageNumber="16">Etymology:</emphasis>
|
||
We dedicate the new subspecies to Dr. Rüdiger (Joe) Wagner, the father of the senior author, to acknowledge his valuable work over many years.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |