treatments-xml/data/03/F6/87/03F687F7FF9F6C1ADAFD9BA96EF1B40B.xml

<document id="AB1FCBDE8E1C09EA1F57DDFE82BE0482" ID-DOI="10.5281/zenodo.207732" ID-GBIF-Dataset="f012a7d5-c4ce-4d77-86e0-1b9215fd5211" ID-ISSN="1175-5326" ID-Zenodo-Dep="207732" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460327140569" checkinUser="plazi" docAuthor="Pabijan, Maciej, Gehring, Philip-Sebastian, Köhler, Jörn, Glaw, Frank &amp; Vences, Miguel" docDate="2011" docId="03F687F7FF9F6C1ADAFD9BA96EF1B40B" docLanguage="en" docName="zt02978p050.pdf" docOrigin="Zootaxa 2978" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Blommersia variabilis Pabijan, Gehring, Köhler, Glaw &amp; Vences, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="45" masterDocId="FFCFFF8FFF986C11DA6A9B3E6A74B371" masterDocTitle="A new microendemic frog species of the genus Blommersia (Anura: Mantellidae) from the east coast of Madagascar" masterLastPageNumber="50" masterPageNumber="34" pageNumber="41" updateTime="1698257913655" updateUser="plazi">
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<mods:title id="4677706DA933D546A4BC6FF906604D90">A new microendemic frog species of the genus Blommersia (Anura: Mantellidae) from the east coast of Madagascar</mods:title>
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<mods:namePart id="CCE9849F9AC8D2FD9D750D2C9BA63778">Pabijan, Maciej</mods:namePart>
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<mods:namePart id="0ADAD70806FFB3ACE1BDE03DD0CB9B8B">Gehring, Philip-Sebastian</mods:namePart>
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<mods:namePart id="2551B32B669F9B691CAF1D7558F589C3">Köhler, Jörn</mods:namePart>
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<mods:namePart id="A313DEC82692F1EBE1DBA6083043D8E5">Glaw, Frank</mods:namePart>
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<mods:namePart id="0A7BABBAEA4AE5B20EBB250F5AD679C2">Vences, Miguel</mods:namePart>
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<mods:date id="712FCD3F35E68113CD2ACDA64F9AC05C">2011</mods:date>
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<treatment id="03F687F7FF9F6C1ADAFD9BA96EF1B40B" ID-DOI="http://doi.org/10.5281/zenodo.6189887" ID-GBIF-Taxon="119403947" ID-Zenodo-Dep="6189887" LSID="urn:lsid:plazi:treatment:03F687F7FF9F6C1ADAFD9BA96EF1B40B" httpUri="http://treatment.plazi.org/id/03F687F7FF9F6C1ADAFD9BA96EF1B40B" lastPageId="11" lastPageNumber="45" pageId="7" pageNumber="41">
<subSubSection id="C345656AFF9F6C16DAFD9BA96B91B3C1" box="[151,485,151,176]" pageId="7" pageNumber="41" type="nomenclature">
<paragraph id="8BE036E1FF9F6C16DAFD9BA96B91B3C1" blockId="7.[151,485,151,176]" box="[151,485,151,176]" pageId="7" pageNumber="41">
<heading id="D0A8818DFF9F6C16DAFD9BA96B91B3C1" bold="true" box="[151,485,151,176]" fontSize="10" level="2" pageId="7" pageNumber="41" reason="2">
<emphasis id="B92BEAF3FF9F6C16DAFD9BA96B91B3C1" bold="true" box="[151,485,151,176]" pageId="7" pageNumber="41">
<taxonomicName id="4C5F4D62FF9F6C16DAFD9BA96BFEB3C1" ID-CoL="M74R" box="[151,394,151,176]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B92BEAF3FF9F6C16DAFD9BA96BFEB3C1" bold="true" box="[151,394,151,176]" italics="true" pageId="7" pageNumber="41">Blommersia variabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2185788FF9F6C16DBFA9BA66B91B3C1" box="[400,485,152,176]" pageId="7" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C345656AFF9F6C16DAFC9BDD696BB154" pageId="7" pageNumber="41" type="materials_examined">
<paragraph id="8BE036E1FF9F6C16DAFC9BDD694BB234" blockId="7.[150,1437,226,1930]" pageId="7" pageNumber="41">
<emphasis id="B92BEAF3FF9F6C16DAFC9BDD6B71B38A" bold="true" box="[150,261,227,251]" pageId="7" pageNumber="41">
<typeStatus id="54E48843FF9F6C16DAFC9BDD6B74B38A" box="[150,256,227,251]" pageId="7" pageNumber="41" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED4EAE24FF9F6C16DB789BDC6B3EB38B" box="[274,330,226,250]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="41">ZSM</collectionCode>
237/2010 (field number 
<collectionCode id="ED4EAE24FF9F6C16D8079BDC68C6B38B" box="[621,690,226,250]" pageId="7" pageNumber="41">FGZC</collectionCode>
4305), adult male (
<figureCitation id="13642A64FF9F6C16D9F19BDC6996B38B" box="[923,994,226,250]" captionStart="FIGURE 5" captionStartId="9.[151,250,1972,1995]" captionTargetBox="[214,1382,181,1925]" captionTargetId="figure@9.[214,1383,181,1925]" captionTargetPageId="9" captionText="FIGURE 5. Dorsolateral (a) and ventral (b) views of the male holotype of Blommersia variabilis sp. nov. (ZSM 237 / 2010) in life." httpUri="https://zenodo.org/record/207737/files/figure.png" pageId="7" pageNumber="41">Fig. 5</figureCitation>
), from Ambodivoahangy, northwest of Maroantsetra, northeastern 
<collectingCountry id="F3487671FF9F6C16DBD59A396835B26E" box="[447,577,263,287]" name="Madagascar" pageId="7" pageNumber="41">Madagascar</collectingCountry>
(coordinates: 
<geoCoordinate id="EE6B5026FF9F6C16D8B79A396903B26E" box="[733,887,263,287]" direction="south" orientation="latitude" pageId="7" pageNumber="41" precision="1" value="-15.289945">15°17’23.8'' S</geoCoordinate>
, 
<geoCoordinate id="EE6B5026FF9F6C16D9EF9A396E55B26E" box="[901,1057,263,287]" direction="east" orientation="longitude" pageId="7" pageNumber="41" precision="1" value="43.620277">43°37’13.0'' E</geoCoordinate>
, below 
<quantity id="4CA79B04FF9F6C16DE139A396EC5B26E" box="[1145,1201,263,287]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="41" unit="m" value="50.0">50 m</quantity>
a.s.l.), collected on 
<date id="FFE11021FF9F6C16DFE49A396B7FB234" pageId="7" pageNumber="41" value="2010-04-03">3 April 2010</date>
by P.-S. Gehring, F. Glaw, J. Köhler and M. Pabijan.
</paragraph>
<paragraph id="8BE036E1FF9F6C16DAAD9A6D696BB154" blockId="7.[150,1437,226,1930]" pageId="7" pageNumber="41">
<emphasis id="B92BEAF3FF9F6C16DAAD9A6D6B35B21A" bold="true" box="[199,321,339,363]" pageId="7" pageNumber="41">
<typeStatus id="54E48843FF9F6C16DAAD9A6D6B48B21A" box="[199,316,339,363]" pageId="7" pageNumber="41" type="paratype">Paratypes</typeStatus>
.
</emphasis>
Eleven adult males: 
<collectionCode id="ED4EAE24FF9F6C16D8499A6C682FB21B" box="[547,603,338,362]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="41">ZSM</collectionCode>
236/2010 (field number 
<collectionCode id="ED4EAE24FF9F6C16D9009A6C69DBB21B" box="[874,943,338,362]" pageId="7" pageNumber="41">FGZC</collectionCode>
4289), 
<collectionCode id="ED4EAE24FF9F6C16DE699A6D6E10B21A" box="[1027,1124,339,363]" httpUri="http://grbio.org/cool/q3sh-smdb" name="University dAntananarivo, Department de Biologie Animale" pageId="7" pageNumber="41">UADBA</collectionCode>
uncatalogued (
<collectionCode id="ED4EAE24FF9F6C16DF629A6C6F3BB21B" box="[1288,1359,338,362]" pageId="7" pageNumber="41">FGZC</collectionCode>
4290), 
<collectionCode id="ED4EAE24FF9F6C16DAFD9A466A8CB2E1" box="[151,248,376,400]" httpUri="http://grbio.org/cool/q3sh-smdb" name="University dAntananarivo, Department de Biologie Animale" pageId="7" pageNumber="41">UADBA</collectionCode>
uncatalogued (
<collectionCode id="ED4EAE24FF9F6C16DBF79A496B90B2FE" box="[413,484,375,399]" pageId="7" pageNumber="41">FGZC</collectionCode>
4306), with same data as 
<typeStatus id="54E48843FF9F6C16D9699A496910B2FE" box="[771,868,375,399]" pageId="7" pageNumber="41" type="holotype">holotype</typeStatus>
; 
<collectionCode id="ED4EAE24FF9F6C16D91A9A4669CFB2FE" LSID="urn:lsid:biocol.org:col:34613" box="[880,955,376,399]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="7" pageNumber="41">ZFMK</collectionCode>
52608–52610, collected by F. Glaw and M. Vences on 
<date id="FFE11021FF9F6C16DB609AA36BD9B2C4" box="[266,429,413,437]" pageId="7" pageNumber="41" value="1991-03-18">
18 March 
<quantity id="4CA79B04FF9F6C16DB1D9AA36BD9B2C4" box="[375,429,413,437]" metricMagnitude="1" metricUnit="m" metricValue="5.05714" pageId="7" pageNumber="41" unit="in" value="1991.0">1991</quantity>
</date>
in the coastal town of Maroantsetra (
<geoCoordinate id="EE6B5026FF9F6C16D9539AA369E0B2C4" box="[825,916,413,437]" direction="south" orientation="latitude" pageId="7" pageNumber="41" precision="925" value="-15.433333">15°26' S</geoCoordinate>
, 
<geoCoordinate id="EE6B5026FF9F6C16D9CA9AA3698AB2C4" box="[928,1022,413,437]" direction="east" orientation="longitude" pageId="7" pageNumber="41" precision="925" value="49.733334">49°44' E</geoCoordinate>
, ca. 
<quantity id="4CA79B04FF9F6C16DE479AA36E17B2C4" box="[1069,1123,413,437]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="7" pageNumber="41" unit="m" value="20.0">20 m</quantity>
a.s.l.); 
<collectionCode id="ED4EAE24FF9F6C16DEDA9AA06E8FB2C4" LSID="urn:lsid:biocol.org:col:34613" box="[1200,1275,414,437]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="7" pageNumber="41">ZFMK</collectionCode>
52612–52615, collected by F. Glaw and M. Vences on 
<date id="FFE11021FF9F6C16D8569AFC6894B2AB" box="[572,736,450,474]" pageId="7" pageNumber="41" value="1991-03-19">19 March 1991</date>
at a site slightly inland from the coastal village of Voloina (
<geoCoordinate id="EE6B5026FF9F6C16DF3D9AFC6AD3B28E" direction="south" orientation="latitude" pageId="7" pageNumber="41" precision="925" value="-15.566667">15°34' S</geoCoordinate>
, 
<geoCoordinate id="EE6B5026FF9F6C16DADF9AD96B61B28E" box="[181,277,487,511]" direction="east" orientation="longitude" pageId="7" pageNumber="41" precision="925" value="49.616665">49°37' E</geoCoordinate>
) which is located south of Maroantsetra; 
<collectionCode id="ED4EAE24FF9F6C16D8899AD6696BB171" box="[739,799,488,512]" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="7" pageNumber="41">ZMA</collectionCode>
19509 (field number 
<collectionCode id="ED4EAE24FF9F6C16DE799AD96E16B28E" box="[1043,1122,487,511]" pageId="7" pageNumber="41">FGMV</collectionCode>
2002.2214) collected by M. Vences and A. Sarovy on 
<date id="FFE11021FF9F6C16DBC19933681DB154" box="[427,617,525,549]" pageId="7" pageNumber="41" value="2003-02-12">
12 February 
<quantity id="4CA79B04FF9F6C16D8599933681DB154" box="[563,617,525,549]" metricMagnitude="1" metricUnit="m" metricValue="5.087619999999999" pageId="7" pageNumber="41" unit="in" value="2003.0">2003</quantity>
</date>
in Maroantsetra.
</paragraph>
</subSubSection>
<subSubSection id="C345656AFF9F6C16DAAD990C6EDBB644" pageId="7" pageNumber="41" type="diagnosis">
<paragraph id="8BE036E1FF9F6C16DAAD990C6F50B0CE" blockId="7.[150,1437,226,1930]" pageId="7" pageNumber="41">
<emphasis id="B92BEAF3FF9F6C16DAAD990C6B48B13A" bold="true" box="[199,316,562,587]" pageId="7" pageNumber="41">Diagnosis.</emphasis>
Assigned to the genus 
<taxonomicName id="4C5F4D62FF9F6C16D85A990D68C6B13A" box="[560,690,563,587]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="B92BEAF3FF9F6C16D85A990D68C6B13A" box="[560,690,563,587]" italics="true" pageId="7" pageNumber="41">Blommersia</emphasis>
</taxonomicName>
in the 
<taxonomicName id="4C5F4D62FF9F6C16D893990C690DB13B" box="[761,889,562,586]" class="Amphibia" family="Mantellidae" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="family">Mantellidae</taxonomicName>
by a combination of (1) presence of intercalary elements between ultimate and penultimate phalanges of fingers and toes (verified by external examination), (2) presence of femoral glands and absence of nuptial pads in males, (3) presence of a moderately distensible, inconspicuously coloured single subgular vocal sac in males, (4) small size (adult SVL &lt;
<quantity id="4CA79B04FF9F6C16D926999C69E9B1CB" box="[844,925,674,698]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="41" unit="mm" value="25.0">25 mm</quantity>
), (5) semiarboreal habits and calling behaviour from vegetation above stagnant water. Among species of 
<taxonomicName id="4C5F4D62FF9F6C16D89699F6690AB191" box="[764,894,712,736]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="B92BEAF3FF9F6C16D89699F6690AB191" box="[764,894,712,736]" italics="true" pageId="7" pageNumber="41">Blommersia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C5F4D62FF9F6C16D9E099F76E7AB191" box="[906,1038,712,736]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B92BEAF3FF9F6C16D9E099F76E7AB191" box="[906,1038,712,736]" italics="true" pageId="7" pageNumber="41">B. variabilis</emphasis>
</taxonomicName>
is distinguished from 
<taxonomicName id="4C5F4D62FF9F6C16DE9199F76FE8B191" box="[1275,1436,712,736]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="blommersae">
<emphasis id="B92BEAF3FF9F6C16DE9199F76FE8B191" box="[1275,1436,712,736]" italics="true" pageId="7" pageNumber="41">B. blommersae</emphasis>
</taxonomicName>
by vomerine teeth present (vs. absent); from 
<taxonomicName id="4C5F4D62FF9F6C16D81D99D06972B074" box="[631,774,749,773]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="domerguei">
<emphasis id="B92BEAF3FF9F6C16D81D99D06972B074" box="[631,774,749,773]" italics="true" pageId="7" pageNumber="41">B. domerguei</emphasis>
</taxonomicName>
by different colouration (absence of light brown dorsum with dark longitudinal markings) and vomerine teeth present (vs. absent); from 
<taxonomicName id="4C5F4D62FF9F6C16D988982A6EFEB05A" box="[994,1162,787,811]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="grandisonae">
<emphasis id="B92BEAF3FF9F6C16D988982A6EFEB05A" box="[994,1162,787,811]" italics="true" pageId="7" pageNumber="41">B. grandisonae</emphasis>
</taxonomicName>
by different colouration (absence of distinct black lateral line ventrally bordered by white, vs. presence) and vomerine teeth present (vs. absent); from 
<taxonomicName id="4C5F4D62FF9F6C16DABB98606B6CB004" box="[209,280,861,885]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="kely">
<emphasis id="B92BEAF3FF9F6C16DABB98606B6CB004" box="[209,280,861,885]" italics="true" pageId="7" pageNumber="41">B. kely</emphasis>
</taxonomicName>
by larger size (male SVL 
<quantity id="4CA79B04FF9F6C16D847986368D4B004" box="[557,672,861,885]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="7" pageNumber="41" unit="mm" value="22.0" valueMax="25.0" valueMin="19.0">19–25 mm</quantity>
vs. 
<quantity id="4CA79B04FF9F6C16D8A09863694BB004" box="[714,831,861,885]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="7" pageNumber="41" unit="mm" value="15.0" valueMax="16.0" valueMin="14.0">14–16 mm</quantity>
) and vomerine teeth present (vs. absent); from 
<taxonomicName id="4C5F4D62FF9F6C16DF5898606FE8B004" box="[1330,1436,862,885]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="sarotra">
<emphasis id="B92BEAF3FF9F6C16DF5898606FE8B004" box="[1330,1436,862,885]" italics="true" pageId="7" pageNumber="41">B. sarotra</emphasis>
</taxonomicName>
by larger size (male SVL 
<quantity id="4CA79B04FF9F6C16DBC298BC6868B0EB" box="[424,540,898,922]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="7" pageNumber="41" unit="mm" value="22.0" valueMax="25.0" valueMin="19.0">19–25 mm</quantity>
vs. 
<quantity id="4CA79B04FF9F6C16D82298BC68C9B0EB" box="[584,701,898,922]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="7" pageNumber="41" unit="mm" value="16.0" valueMax="17.0" valueMin="15.0">15–17 mm</quantity>
) and vomerine teeth present (vs. absent); from 
<taxonomicName id="4C5F4D62FF9F6C16DEDE98BA6F44B0EA" box="[1204,1328,899,923]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="angolafa">
<emphasis id="B92BEAF3FF9F6C16DEDE98BA6F44B0EA" box="[1204,1328,899,923]" italics="true" pageId="7" pageNumber="41">B. angolafa</emphasis>
</taxonomicName>
by colouration (absence of uniformly brownish dorsal colour with white dots) and vomerine teeth present (vs. absent).
</paragraph>
<paragraph id="8BE036E1FF9F6C16DAAF98F36EDBB644" blockId="7.[150,1437,226,1930]" pageId="7" pageNumber="41">
The new species is morphologically most similar to 
<taxonomicName id="4C5F4D62FF9F6C16D96398F06910B094" box="[777,868,974,997]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="wittei">
<emphasis id="B92BEAF3FF9F6C16D96398F06910B094" box="[777,868,974,997]" italics="true" pageId="7" pageNumber="41">B. wittei</emphasis>
</taxonomicName>
from which it mainly differs by the wider distance between inner margins of femoral glands (FGD 
<quantity id="4CA79B04FF9F6C16D8F598CC6956B77B" box="[671,802,1010,1034]" metricMagnitude="-3" metricUnit="m" metricValue="3.65" metricValueMax="4.5" metricValueMin="2.8" pageId="7" pageNumber="41" unit="mm" value="3.65" valueMax="4.5" valueMin="2.8">2.8–4.5 mm</quantity>
vs. 1.6–2.0 in the 
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series of 
<taxonomicName id="4C5F4D62FF9F6C16DEEB98CA6EAEB77A" box="[1153,1242,1012,1035]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="wittei">
<emphasis id="B92BEAF3FF9F6C16DEEB98CA6EAEB77A" box="[1153,1242,1012,1035]" italics="true" pageId="7" pageNumber="41">B. wittei</emphasis>
</taxonomicName>
; see Vences 
<emphasis id="B92BEAF3FF9F6C16DF0C98CA6FE8B77A" box="[1382,1436,1011,1035]" italics="true" pageId="7" pageNumber="41">et al.</emphasis>
2010; FGD 12–20% of SVL vs. 7–9%), connected metatarsalia in most specimens (vs. always separated by webbing), and advertisement calls consisting of a short series of 3-6 notes of a mean duration of 107 ms and a mean inter-note interval of 27 ms (vs. longer series of up to 25 notes with a mean duration of 23-46 ms and a mean interval duration of 30–72 ms). The new species forms a monophyletic group with the recently described 
<taxonomicName id="4C5F4D62FF9F6C16D9B19FB76E3CB7D1" box="[987,1096,1160,1184]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="dejongi">
<emphasis id="B92BEAF3FF9F6C16D9B19FB76E3CB7D1" box="[987,1096,1160,1184]" italics="true" pageId="7" pageNumber="41">B. dejongi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5F4D62FF9F6C16DE169FB76EA9B7D1" box="[1148,1245,1160,1184]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="galani">
<emphasis id="B92BEAF3FF9F6C16DE169FB76EA9B7D1" box="[1148,1245,1160,1184]" italics="true" pageId="7" pageNumber="41">B. galani</emphasis>
</taxonomicName>
. It differs from 
<taxonomicName id="4C5F4D62FF9F6C16DFEF9FB76AAFB7B4" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="galani">
<emphasis id="B92BEAF3FF9F6C16DFEF9FB76AAFB7B4" italics="true" pageId="7" pageNumber="41">B. galani</emphasis>
</taxonomicName>
by the presence of vomerine teeth (vs. absence) and smaller femoral glands (see 
<figureCitation id="13642A64FF9F6C16DE569F936E09B7B4" box="[1084,1149,1197,1221]" captionStart="FIGURE 4" captionStartId="6.[151,250,1860,1883]" captionTargetBox="[258,1329,987,1817]" captionTargetId="figure@6.[258,1329,987,1817]" captionTargetPageId="6" captionText="FIGURE 4. Scatterplot of relative femoral gland length (FGL / SVL ratio) and relative distance between inner edges of femoral glands (FGD / SVL) in the three endemic Blommersia species of Madagascar's northern central east coast, and B. wittei. Based on measurements provided in Table 1 (B. variabilis sp. nov. and some B. wittei) and Vences et al. (2010) for B. dejongi and B. galani, as well as type specimens of B. wittei (only male specimens with complete measurement data were used). Arrow points to a putative hybrid between a female B. galani and male B. dejongi (specimen ZSM 452 / 2006)." httpUri="https://zenodo.org/record/207736/files/figure.png" pageId="7" pageNumber="41">Fig. 4</figureCitation>
; femoral gland length 3.5– 5.2 vs. 
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). It differs from 
<taxonomicName id="4C5F4D62FF9F6C16D84F9FEA68EFB79A" box="[549,667,1235,1259]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="species" species="dejongi">
<emphasis id="B92BEAF3FF9F6C16D84F9FEA68EFB79A" box="[549,667,1235,1259]" italics="true" pageId="7" pageNumber="41">B. dejongi</emphasis>
</taxonomicName>
by a shorter distance between the inner margins of femoral glands (
<figureCitation id="13642A64FF9F6C16DAF59FC96A90B67E" box="[159,228,1271,1295]" captionStart="FIGURE 4" captionStartId="6.[151,250,1860,1883]" captionTargetBox="[258,1329,987,1817]" captionTargetId="figure@6.[258,1329,987,1817]" captionTargetPageId="6" captionText="FIGURE 4. Scatterplot of relative femoral gland length (FGL / SVL ratio) and relative distance between inner edges of femoral glands (FGD / SVL) in the three endemic Blommersia species of Madagascar's northern central east coast, and B. wittei. Based on measurements provided in Table 1 (B. variabilis sp. nov. and some B. wittei) and Vences et al. (2010) for B. dejongi and B. galani, as well as type specimens of B. wittei (only male specimens with complete measurement data were used). Arrow points to a putative hybrid between a female B. galani and male B. dejongi (specimen ZSM 452 / 2006)." httpUri="https://zenodo.org/record/207736/files/figure.png" pageId="7" pageNumber="41">Fig. 4</figureCitation>
; FGD 
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vs. 
<quantity id="4CA79B04FF9F6C16DB889FC96819B67E" box="[482,621,1271,1295]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" metricValueMax="9.8" metricValueMin="7.8" pageId="7" pageNumber="41" unit="mm" value="8.8" valueMax="9.8" valueMin="7.8">7.8–9.8 mm</quantity>
). Furthermore, the new species differs from all nominal species of 
<taxonomicName id="4C5F4D62FF9F6C16DF3D9FC66A97B644" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="7" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="B92BEAF3FF9F6C16DF3D9FC66A97B644" italics="true" pageId="7" pageNumber="41">Blommersia</emphasis>
</taxonomicName>
by a substantial genetic divergence and by its advertisement call (as described below).
</paragraph>
</subSubSection>
<subSubSection id="C345656AFF9F6C1BDAAD9E7C6966B05B" lastPageId="10" lastPageNumber="44" pageId="7" pageNumber="41" type="description">
<paragraph id="8BE036E1FF9F6C16DAAD9E7C6BD6B4FB" blockId="7.[150,1437,226,1930]" pageId="7" pageNumber="41">
<emphasis id="B92BEAF3FF9F6C16DAAD9E7C6861B62A" bold="true" box="[199,533,1346,1371]" pageId="7" pageNumber="41">
Description of the 
<typeStatus id="54E48843FF9F6C16DBCD9E7D6864B62A" box="[423,528,1347,1371]" pageId="7" pageNumber="41" type="holotype">holotype</typeStatus>
.
</emphasis>
Specimen in a good state of preservation, tongue removed as tissue sample for molecular analysis. SVL 
<quantity id="4CA79B04FF9F6C16DBD39E596856B60E" box="[441,546,1383,1407]" metricMagnitude="-2" metricUnit="m" metricValue="2.37" pageId="7" pageNumber="41" unit="mm" value="23.7">23.7 mm</quantity>
, for further measurements see Table 1. Body slender; head longer than wide, wider than body; snout rounded in dorsal and lateral views, nostrils directed laterally, slightly protuberant, nearer to tip of snout than to eye; canthus rostralis distinct, straight; loreal region slightly concave; tympanum distinct, rounded, its diameter 54% of eye diameter; supratympanic fold distinct posterior to tympanum, indistinct between eye and tympanum, curved; tongue removed, its shape therefore not verifiable; vomerine teeth present as two distinct protuberances immediately posterior-medially of choanae, maxillary teeth present; choanae small, rounded. Arms slender, subarticular tubercles single; fingers without webbing; relative length of fingers 1&lt;2&lt;4&lt;3 on the right hand, 1&lt;2=4&lt;3 on the left hand; finger discs distinctly enlarged; nuptial pads absent. 
<collectingCountry id="F3487671FF9F6C16DEE09D536EB6B5F4" box="[1162,1218,1645,1669]" name="India" pageId="7" pageNumber="41">Hind</collectingCountry>
limbs slender; tibiotarsal articulation reaches nostril when the hind limb is adpressed along the body; lateral metatarsalia largely separated; inner and outer metatarsal tubercles distinct; webbing formula (according to 
<bibRefCitation id="EFCE4B10FF9F6C16DE0D9D896FE0B5BE" author="Blommers-Schlosser" box="[1127,1428,1719,1743]" pageId="7" pageNumber="41" refString="Blommers-Schlosser, R. M. A. (1979) Biosystematics of the Malagasy frogs. I. Mantellinae (Ranidae). Beaufortia, 352, 1 - 77." type="journal article" year="1979">Blommers-Schlösser 1979</bibRefCitation>
) between toes 1(0.25), 
<date id="FFE11021FF9F6C16DBFA9DE36BD3B584" box="[400,423,1757,1781]" pageId="7" pageNumber="41">2i</date>
(1.25), 2e(0.5), 
<date id="FFE11021FF9F6C16D83D9DE36818B584" box="[599,620,1757,1781]" pageId="7" pageNumber="41">3i</date>
(1.5), 3e(1), 
<date id="FFE11021FF9F6C16D8909DE36966B584" box="[762,786,1757,1781]" pageId="7" pageNumber="41">4i</date>
(2.5), 4e(2.5), 5(0.75); relative length of toes 1&lt;2&lt;5&lt;3&lt;4. Skin on the upper surface smooth, without folds or ridges. No distinct enlarged tubercles in the cloacal region; ventral skin smooth. Femoral glands distinct, measuring 5.2 x 
<quantity id="4CA79B04FF9F6C16D94E9C19690FB44E" box="[804,891,1831,1855]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="41" unit="mm" value="1.5">1.5 mm</quantity>
, of 
<typeStatus id="54E48843FF9F6C16D9CE9C1669A0B431" box="[932,980,1832,1856]" pageId="7" pageNumber="41">type</typeStatus>
2 sensu 
<bibRefCitation id="EFCE4B10FF9F6C16DE5E9C196F76B44E" box="[1076,1282,1831,1856]" pageId="7" pageNumber="41" refString="Glaw, F., Vences, M., &amp; Gossmann, V. (2000) A new species of Mantidactylus from Madagascar, with a comparative survey of internal femoral gland structure in the genus (Amphibia: Ranidae: Mantellinae). Journal of Natural History, 34, 1135 - 1154." type="journal article">
Glaw 
<emphasis id="B92BEAF3FF9F6C16DE1D9C176ED3B431" box="[1143,1191,1832,1856]" italics="true" pageId="7" pageNumber="41">et al</emphasis>
. (2000)
</bibRefCitation>
, consisting of 17 distinct granules as verified from internal view (after reflexing of the ventral skin on thigh), distance between femoral glands 
<quantity id="4CA79B04FF9F6C16DB2F9C4C6BEAB4FB" box="[325,414,1906,1930]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="41" unit="mm" value="2.8">2.8 mm</quantity>
.
</paragraph>
<paragraph id="8BE036E1FF906C19DA1799086D7AB13B" blockId="8.[125,1866,565,639]" box="[125,1806,565,587]" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DA1799086AB9B13A" bold="true" box="[125,205,566,587]" pageId="8" pageNumber="42">Table 1.</emphasis>
Measurements of the 
<typeStatus id="54E48843FF906C19DBF0990B6BB6B13B" box="[410,450,565,586]" pageId="8" pageNumber="42">type</typeStatus>
series of 
<taxonomicName id="4C5F4D62FF906C19D870990B689FB13B" box="[538,747,565,586]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="42" phylum="Chordata" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B92BEAF3FF906C19D870990B689FB13B" box="[538,747,565,586]" italics="true" pageId="8" pageNumber="42">Blommersia variabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2185788FF906C19D89A99086948B13A" box="[752,828,566,587]" pageId="8" pageNumber="42" rank="species">
<emphasis id="B92BEAF3FF906C19D89A99086948B13A" bold="true" box="[752,828,566,587]" pageId="8" pageNumber="42">sp. nov.</emphasis>
</taxonomicNameLabel>
and comparative specimens of 
<taxonomicName id="4C5F4D62FF906C19DE0A990B6ED9B13B" box="[1120,1197,565,586]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="42" phylum="Chordata" rank="species" species="wittei">
<emphasis id="B92BEAF3FF906C19DE0A990B6ED9B13B" box="[1120,1197,565,586]" italics="true" pageId="8" pageNumber="42">B. wittei</emphasis>
</taxonomicName>
(all males). ZSM 237/2010 is the 
<typeStatus id="54E48843FF906C19DF86990B6C34B13B" box="[1516,1600,565,586]" pageId="8" pageNumber="42" type="holotype">holotype</typeStatus>
, all other 
<taxonomicName id="4C5F4D62FF906C19DCF0990B6D7AB13B" box="[1690,1806,565,586]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="42" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B92BEAF3FF906C19DCF0990B6D7AB13B" box="[1690,1806,565,586]" italics="true" pageId="8" pageNumber="42">B. variabilis</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BE036E1FF906C19DA17996E6D3EB114" blockId="8.[125,1866,565,639]" box="[125,1866,592,613]" pageId="8" pageNumber="42">
specimens are 
<typeStatus id="54E48843FF906C19DB6E996E6B14B114" box="[260,352,592,613]" pageId="8" pageNumber="42" type="paratype">paratypes</typeStatus>
. For abbreviations, see Material and Methods. Relative hindlimb length (RHL) is coded as follows: when hindlimb is adpressed along body, tibiotarsal articulation
</paragraph>
<paragraph id="8BE036E1FF906C19DA1799546E60B10E" blockId="8.[125,1866,565,639]" box="[125,1044,618,639]" pageId="8" pageNumber="42">reaches (1) anterior eye corner, (2) eye center, (3) between eye and nostril, (4) nostril, (5) snout tip.</paragraph>
<paragraph id="8BE036E1FF906C19DAED99B96D36B50C" pageId="8" pageNumber="42">
<table id="F95FC441FF9093EEDA1799B96D27B50C" box="[125,1875,647,1661]" gridcols="21" gridrows="13" pageId="8" pageNumber="42">
<tr id="356F34A3FF9093EEDA1799B96D27B193" box="[125,1875,647,738]" gridrow="0" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA1799B96B52B193" box="[125,294,647,738]" gridcol="0" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DAED99B96A9BB1BA" bold="true" box="[135,239,647,715]" pageId="8" pageNumber="42">
Locality 
<taxonomicName id="4C5F4D62FF906C19DAED99866A9BB1BA" authority="Maroantsetra" authorityName="Maroantsetra" box="[135,239,696,715]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="42" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B92BEAF3FF906C19DAED99866A9BB1BA" bold="true" box="[135,239,696,715]" italics="true" pageId="8" pageNumber="42">B. variabilis</emphasis>
</taxonomicName>
</emphasis>
Maroantsetra
</th>
<th id="76BE5DDFFF9093EEDB3199B96BAFB193" box="[347,475,647,738]" gridcol="1" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DB3199B96BD9B1C0" bold="true" box="[347,429,647,689]" pageId="8" pageNumber="42">Voucher specimen</emphasis>
ZFMK 52608
</th>
<th id="76BE5DDFFF9093EED86A99B9682EB193" box="[512,602,647,738]" gridcol="2" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19D86A99B96832B1C0" bold="true" box="[512,582,647,689]" pageId="8" pageNumber="42">Field number</emphasis>
n.a.
</th>
<th id="76BE5DDFFF9093EED81D99B968EBB193" box="[631,671,647,738]" gridcol="3" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19D81D99B968EBB1E8" bold="true" box="[631,671,647,665]" pageId="8" pageNumber="42">SVL</emphasis>
20.6
</th>
<th id="76BE5DDFFF9093EED8D699B96895B193" box="[700,737,647,738]" gridcol="4" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19D8D699B96895B1E8" bold="true" box="[700,737,647,665]" pageId="8" pageNumber="42">HW</emphasis>
6.6
</th>
<th id="76BE5DDFFF9093EED89799B9696FB193" box="[765,795,647,738]" gridcol="5" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19D89799B9696FB1E8" bold="true" box="[765,795,647,665]" pageId="8" pageNumber="42">HL</emphasis>
8.3
</th>
<th id="76BE5DDFFF9093EED95D99B96920B193" box="[823,852,647,738]" gridcol="6" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19D95D99B96920B1E8" bold="true" box="[823,852,647,665]" pageId="8" pageNumber="42">TD</emphasis>
1.5
</th>
<th id="76BE5DDFFF9093EED91A99B969F9B193" box="[880,909,647,738]" gridcol="7" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19D91A99B969F9B1E8" bold="true" box="[880,909,647,665]" pageId="8" pageNumber="42">ED</emphasis>
2.6
</th>
<th id="76BE5DDFFF9093EED9C399B969A1B193" box="[937,981,647,738]" gridcol="8" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19D9C399B969A1B1E8" bold="true" box="[937,981,647,665]" pageId="8" pageNumber="42">END</emphasis>
2.2
</th>
<th id="76BE5DDFFF9093EED99B99B96E6FB193" box="[1009,1051,647,738]" gridcol="9" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19D99B99B96E6FB1E8" bold="true" box="[1009,1051,647,665]" pageId="8" pageNumber="42">NSD</emphasis>
1.4
</th>
<th id="76BE5DDFFF9093EEDE5D99B96E10B193" box="[1079,1124,647,738]" gridcol="10" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DE5D99B96E10B1E8" bold="true" box="[1079,1124,647,665]" pageId="8" pageNumber="42">NND</emphasis>
2.3
</th>
<th id="76BE5DDFFF9093EEDEEA99B96ECEB193" box="[1152,1210,647,738]" gridcol="11" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DEEA99B96ECEB1E8" bold="true" box="[1152,1210,647,665]" pageId="8" pageNumber="42">FORL</emphasis>
12.3
</th>
<th id="76BE5DDFFF9093EEDEBC99B96F77B193" box="[1238,1283,647,738]" gridcol="12" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DEBC99B96F77B1E8" bold="true" box="[1238,1283,647,665]" pageId="8" pageNumber="42">HAL</emphasis>
5.9
</th>
<th id="76BE5DDFFF9093EEDF7599B96F31B193" box="[1311,1349,647,738]" gridcol="13" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DF7599B96F31B1E8" bold="true" box="[1311,1349,647,665]" pageId="8" pageNumber="42">HIL</emphasis>
34
</th>
<th id="76BE5DDFFF9093EEDF0B99B96FEDB193" box="[1377,1433,647,738]" gridcol="14" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DF0B99B96FEDB1E8" bold="true" box="[1377,1433,647,665]" pageId="8" pageNumber="42">FOTL</emphasis>
15.3
</th>
<th id="76BE5DDFFF9093EEDFDF99B96F94B193" box="[1461,1504,647,738]" gridcol="15" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DFDF99B96F94B1E8" bold="true" box="[1461,1504,647,665]" pageId="8" pageNumber="42">FOL</emphasis>
9.4
</th>
<th id="76BE5DDFFF9093EEDF9699B96C5AB193" box="[1532,1582,647,738]" gridcol="16" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DF9699B96C5AB1E8" bold="true" box="[1532,1582,647,665]" pageId="8" pageNumber="42">TIBL</emphasis>
10.6
</th>
<th id="76BE5DDFFF9093EEDC2099B96C01B193" box="[1610,1653,647,738]" gridcol="17" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DC2099B96C01B1E8" bold="true" box="[1610,1653,647,665]" pageId="8" pageNumber="42">FGL</emphasis>
3.8
</th>
<th id="76BE5DDFFF9093EEDCFB99B96CB7B193" box="[1681,1731,647,738]" gridcol="18" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DCFB99B96CB7B1E8" bold="true" box="[1681,1731,647,665]" pageId="8" pageNumber="42">FGW</emphasis>
1.5
</th>
<th id="76BE5DDFFF9093EEDCB499B96D7EB193" box="[1758,1802,647,738]" gridcol="19" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DCB499B96D7EB1E8" bold="true" box="[1758,1802,647,665]" pageId="8" pageNumber="42">FGD</emphasis>
3.8
</th>
<th id="76BE5DDFFF9093EEDD4C99B96D27B193" box="[1830,1875,647,738]" gridcol="20" gridrow="0" pageId="8" pageNumber="42">
<emphasis id="B92BEAF3FF906C19DD4C99B96D27B1E8" bold="true" box="[1830,1875,647,665]" pageId="8" pageNumber="42">RHL</emphasis>
4
</th>
</tr>
<tr id="356F34A3FF9093EEDA1799D16D27B073" box="[125,1875,751,770]" gridrow="1" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA1799D16B52B073" box="[125,294,751,770]" gridcol="0" gridrow="1" pageId="8" pageNumber="42">Maroantsetra</th>
<td id="76BE5DDFFF9093EEDB3199D16BAFB073" box="[347,475,751,770]" gridcol="1" gridrow="1" pageId="8" pageNumber="42">ZFMK 52609</td>
<td id="76BE5DDFFF9093EED86A99D1682EB073" box="[512,602,751,770]" gridcol="2" gridrow="1" pageId="8" pageNumber="42">n.a.</td>
<td id="76BE5DDFFF9093EED81D99D168EBB073" box="[631,671,751,770]" gridcol="3" gridrow="1" pageId="8" pageNumber="42">18.6</td>
<td id="76BE5DDFFF9093EED8D699D16895B073" box="[700,737,751,770]" gridcol="4" gridrow="1" pageId="8" pageNumber="42">6</td>
<td id="76BE5DDFFF9093EED89799D1696FB073" box="[765,795,751,770]" gridcol="5" gridrow="1" pageId="8" pageNumber="42">7.8</td>
<td id="76BE5DDFFF9093EED95D99D16920B073" box="[823,852,751,770]" gridcol="6" gridrow="1" pageId="8" pageNumber="42">1.4</td>
<td id="76BE5DDFFF9093EED91A99D169F9B073" box="[880,909,751,770]" gridcol="7" gridrow="1" pageId="8" pageNumber="42">2.4</td>
<td id="76BE5DDFFF9093EED9C399D169A1B073" box="[937,981,751,770]" gridcol="8" gridrow="1" pageId="8" pageNumber="42">2.1</td>
<td id="76BE5DDFFF9093EED99B99D16E6FB073" box="[1009,1051,751,770]" gridcol="9" gridrow="1" pageId="8" pageNumber="42">1.6</td>
<td id="76BE5DDFFF9093EEDE5D99D16E10B073" box="[1079,1124,751,770]" gridcol="10" gridrow="1" pageId="8" pageNumber="42">2.4</td>
<td id="76BE5DDFFF9093EEDEEA99D16ECEB073" box="[1152,1210,751,770]" gridcol="11" gridrow="1" pageId="8" pageNumber="42">12.4</td>
<td id="76BE5DDFFF9093EEDEBC99D16F77B073" box="[1238,1283,751,770]" gridcol="12" gridrow="1" pageId="8" pageNumber="42">5.5</td>
<td id="76BE5DDFFF9093EEDF7599D16F31B073" box="[1311,1349,751,770]" gridcol="13" gridrow="1" pageId="8" pageNumber="42">32</td>
<td id="76BE5DDFFF9093EEDF0B99D16FEDB073" box="[1377,1433,751,770]" gridcol="14" gridrow="1" pageId="8" pageNumber="42">14.8</td>
<td id="76BE5DDFFF9093EEDFDF99D16F94B073" box="[1461,1504,751,770]" gridcol="15" gridrow="1" pageId="8" pageNumber="42">9.4</td>
<td id="76BE5DDFFF9093EEDF9699D16C5AB073" box="[1532,1582,751,770]" gridcol="16" gridrow="1" pageId="8" pageNumber="42">10.7</td>
<td id="76BE5DDFFF9093EEDC2099D16C01B073" box="[1610,1653,751,770]" gridcol="17" gridrow="1" pageId="8" pageNumber="42">3.6</td>
<td id="76BE5DDFFF9093EEDCFB99D16CB7B073" box="[1681,1731,751,770]" gridcol="18" gridrow="1" pageId="8" pageNumber="42">1.4</td>
<td id="76BE5DDFFF9093EEDCB499D16D7EB073" box="[1758,1802,751,770]" gridcol="19" gridrow="1" pageId="8" pageNumber="42">3.8</td>
<td id="76BE5DDFFF9093EEDD4C99D16D27B073" box="[1830,1875,751,770]" gridcol="20" gridrow="1" pageId="8" pageNumber="42">5</td>
</tr>
<tr id="356F34A3FF9093EEDA1798316D27B053" box="[125,1875,783,802]" gridrow="2" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA1798316B52B053" box="[125,294,783,802]" gridcol="0" gridrow="2" pageId="8" pageNumber="42">Maroantsetra</th>
<td id="76BE5DDFFF9093EEDB3198316BAFB053" box="[347,475,783,802]" gridcol="1" gridrow="2" pageId="8" pageNumber="42">ZFMK 52610</td>
<td id="76BE5DDFFF9093EED86A9831682EB053" box="[512,602,783,802]" gridcol="2" gridrow="2" pageId="8" pageNumber="42">n.a.</td>
<td id="76BE5DDFFF9093EED81D983168EBB053" box="[631,671,783,802]" gridcol="3" gridrow="2" pageId="8" pageNumber="42">20.3</td>
<td id="76BE5DDFFF9093EED8D698316895B053" box="[700,737,783,802]" gridcol="4" gridrow="2" pageId="8" pageNumber="42">6</td>
<td id="76BE5DDFFF9093EED8979831696FB053" box="[765,795,783,802]" gridcol="5" gridrow="2" pageId="8" pageNumber="42">7.9</td>
<td id="76BE5DDFFF9093EED95D98316920B053" box="[823,852,783,802]" gridcol="6" gridrow="2" pageId="8" pageNumber="42">1.4</td>
<td id="76BE5DDFFF9093EED91A983169F9B053" box="[880,909,783,802]" gridcol="7" gridrow="2" pageId="8" pageNumber="42">2.3</td>
<td id="76BE5DDFFF9093EED9C3983169A1B053" box="[937,981,783,802]" gridcol="8" gridrow="2" pageId="8" pageNumber="42">2.5</td>
<td id="76BE5DDFFF9093EED99B98316E6FB053" box="[1009,1051,783,802]" gridcol="9" gridrow="2" pageId="8" pageNumber="42">1.4</td>
<td id="76BE5DDFFF9093EEDE5D98316E10B053" box="[1079,1124,783,802]" gridcol="10" gridrow="2" pageId="8" pageNumber="42">2.1</td>
<td id="76BE5DDFFF9093EEDEEA98316ECEB053" box="[1152,1210,783,802]" gridcol="11" gridrow="2" pageId="8" pageNumber="42">12.2</td>
<td id="76BE5DDFFF9093EEDEBC98316F77B053" box="[1238,1283,783,802]" gridcol="12" gridrow="2" pageId="8" pageNumber="42">5.7</td>
<td id="76BE5DDFFF9093EEDF7598316F31B053" box="[1311,1349,783,802]" gridcol="13" gridrow="2" pageId="8" pageNumber="42">32.6</td>
<td id="76BE5DDFFF9093EEDF0B98316FEDB053" box="[1377,1433,783,802]" gridcol="14" gridrow="2" pageId="8" pageNumber="42">14.7</td>
<td id="76BE5DDFFF9093EEDFDF98316F94B053" box="[1461,1504,783,802]" gridcol="15" gridrow="2" pageId="8" pageNumber="42">9.8</td>
<td id="76BE5DDFFF9093EEDF9698316C5AB053" box="[1532,1582,783,802]" gridcol="16" gridrow="2" pageId="8" pageNumber="42">10.6</td>
<td id="76BE5DDFFF9093EEDC2098316C01B053" box="[1610,1653,783,802]" gridcol="17" gridrow="2" pageId="8" pageNumber="42">3.6</td>
<td id="76BE5DDFFF9093EEDCFB98316CB7B053" box="[1681,1731,783,802]" gridcol="18" gridrow="2" pageId="8" pageNumber="42">1.5</td>
<td id="76BE5DDFFF9093EEDCB498316D7EB053" box="[1758,1802,783,802]" gridcol="19" gridrow="2" pageId="8" pageNumber="42">3.5</td>
<td id="76BE5DDFFF9093EEDD4C98316D27B053" box="[1830,1875,783,802]" gridcol="20" gridrow="2" pageId="8" pageNumber="42">4</td>
</tr>
<tr id="356F34A3FF9093EEDA1798116D27B033" box="[125,1875,815,834]" gridrow="3" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA1798116B52B033" box="[125,294,815,834]" gridcol="0" gridrow="3" pageId="8" pageNumber="42">Voloina</th>
<td id="76BE5DDFFF9093EEDB3198116BAFB033" box="[347,475,815,834]" gridcol="1" gridrow="3" pageId="8" pageNumber="42">ZFMK 52612</td>
<td id="76BE5DDFFF9093EED86A9811682EB033" box="[512,602,815,834]" gridcol="2" gridrow="3" pageId="8" pageNumber="42">n.a.</td>
<td id="76BE5DDFFF9093EED81D981168EBB033" box="[631,671,815,834]" gridcol="3" gridrow="3" pageId="8" pageNumber="42">21.6</td>
<td id="76BE5DDFFF9093EED8D698116895B033" box="[700,737,815,834]" gridcol="4" gridrow="3" pageId="8" pageNumber="42">6.5</td>
<td id="76BE5DDFFF9093EED8979811696FB033" box="[765,795,815,834]" gridcol="5" gridrow="3" pageId="8" pageNumber="42">8.3</td>
<td id="76BE5DDFFF9093EED95D98116920B033" box="[823,852,815,834]" gridcol="6" gridrow="3" pageId="8" pageNumber="42">1.3</td>
<td id="76BE5DDFFF9093EED91A981169F9B033" box="[880,909,815,834]" gridcol="7" gridrow="3" pageId="8" pageNumber="42">2.4</td>
<td id="76BE5DDFFF9093EED9C3981169A1B033" box="[937,981,815,834]" gridcol="8" gridrow="3" pageId="8" pageNumber="42">2.2</td>
<td id="76BE5DDFFF9093EED99B98116E6FB033" box="[1009,1051,815,834]" gridcol="9" gridrow="3" pageId="8" pageNumber="42">1.6</td>
<td id="76BE5DDFFF9093EEDE5D98116E10B033" box="[1079,1124,815,834]" gridcol="10" gridrow="3" pageId="8" pageNumber="42">2.7</td>
<td id="76BE5DDFFF9093EEDEEA98116ECEB033" box="[1152,1210,815,834]" gridcol="11" gridrow="3" pageId="8" pageNumber="42">13.4</td>
<td id="76BE5DDFFF9093EEDEBC98116F77B033" box="[1238,1283,815,834]" gridcol="12" gridrow="3" pageId="8" pageNumber="42">5.8</td>
<td id="76BE5DDFFF9093EEDF7598116F31B033" box="[1311,1349,815,834]" gridcol="13" gridrow="3" pageId="8" pageNumber="42">35.8</td>
<td id="76BE5DDFFF9093EEDF0B98116FEDB033" box="[1377,1433,815,834]" gridcol="14" gridrow="3" pageId="8" pageNumber="42">16.3</td>
<td id="76BE5DDFFF9093EEDFDF98116F94B033" box="[1461,1504,815,834]" gridcol="15" gridrow="3" pageId="8" pageNumber="42">10</td>
<td id="76BE5DDFFF9093EEDF9698116C5AB033" box="[1532,1582,815,834]" gridcol="16" gridrow="3" pageId="8" pageNumber="42">11.8</td>
<td id="76BE5DDFFF9093EEDC2098116C01B033" box="[1610,1653,815,834]" gridcol="17" gridrow="3" pageId="8" pageNumber="42">3.2</td>
<td id="76BE5DDFFF9093EEDCFB98116CB7B033" box="[1681,1731,815,834]" gridcol="18" gridrow="3" pageId="8" pageNumber="42">1.5</td>
<td id="76BE5DDFFF9093EEDCB498116D7EB033" box="[1758,1802,815,834]" gridcol="19" gridrow="3" pageId="8" pageNumber="42">4</td>
<td id="76BE5DDFFF9093EEDD4C98116D27B033" box="[1830,1875,815,834]" gridcol="20" gridrow="3" pageId="8" pageNumber="42">4</td>
</tr>
<tr id="356F34A3FF9093EEDA1798716D27B013" box="[125,1875,847,866]" gridrow="4" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA1798716B52B013" box="[125,294,847,866]" gridcol="0" gridrow="4" pageId="8" pageNumber="42">Voloina</th>
<td id="76BE5DDFFF9093EEDB3198716BAFB013" box="[347,475,847,866]" gridcol="1" gridrow="4" pageId="8" pageNumber="42">ZFMK 52613</td>
<td id="76BE5DDFFF9093EED86A9871682EB013" box="[512,602,847,866]" gridcol="2" gridrow="4" pageId="8" pageNumber="42">n.a.</td>
<td id="76BE5DDFFF9093EED81D987168EBB013" box="[631,671,847,866]" gridcol="3" gridrow="4" pageId="8" pageNumber="42">24.7</td>
<td id="76BE5DDFFF9093EED8D698716895B013" box="[700,737,847,866]" gridcol="4" gridrow="4" pageId="8" pageNumber="42">6.4</td>
<td id="76BE5DDFFF9093EED8979871696FB013" box="[765,795,847,866]" gridcol="5" gridrow="4" pageId="8" pageNumber="42">9.4</td>
<td id="76BE5DDFFF9093EED95D98716920B013" box="[823,852,847,866]" gridcol="6" gridrow="4" pageId="8" pageNumber="42">1.3</td>
<td id="76BE5DDFFF9093EED91A987169F9B013" box="[880,909,847,866]" gridcol="7" gridrow="4" pageId="8" pageNumber="42">2.4</td>
<td id="76BE5DDFFF9093EED9C3987169A1B013" box="[937,981,847,866]" gridcol="8" gridrow="4" pageId="8" pageNumber="42">2.6</td>
<td id="76BE5DDFFF9093EED99B98716E6FB013" box="[1009,1051,847,866]" gridcol="9" gridrow="4" pageId="8" pageNumber="42">1.9</td>
<td id="76BE5DDFFF9093EEDE5D98716E10B013" box="[1079,1124,847,866]" gridcol="10" gridrow="4" pageId="8" pageNumber="42">2.6</td>
<td id="76BE5DDFFF9093EEDEEA98716ECEB013" box="[1152,1210,847,866]" gridcol="11" gridrow="4" pageId="8" pageNumber="42">14.7</td>
<td id="76BE5DDFFF9093EEDEBC98716F77B013" box="[1238,1283,847,866]" gridcol="12" gridrow="4" pageId="8" pageNumber="42">7</td>
<td id="76BE5DDFFF9093EEDF7598716F31B013" box="[1311,1349,847,866]" gridcol="13" gridrow="4" pageId="8" pageNumber="42">38.3</td>
<td id="76BE5DDFFF9093EEDF0B98716FEDB013" box="[1377,1433,847,866]" gridcol="14" gridrow="4" pageId="8" pageNumber="42">17.3</td>
<td id="76BE5DDFFF9093EEDFDF98716F94B013" box="[1461,1504,847,866]" gridcol="15" gridrow="4" pageId="8" pageNumber="42">11</td>
<td id="76BE5DDFFF9093EEDF9698716C5AB013" box="[1532,1582,847,866]" gridcol="16" gridrow="4" pageId="8" pageNumber="42">12</td>
<td id="76BE5DDFFF9093EEDC2098716C01B013" box="[1610,1653,847,866]" gridcol="17" gridrow="4" pageId="8" pageNumber="42">4.2</td>
<td id="76BE5DDFFF9093EEDCFB98716CB7B013" box="[1681,1731,847,866]" gridcol="18" gridrow="4" pageId="8" pageNumber="42">1.6</td>
<td id="76BE5DDFFF9093EEDCB498716D7EB013" box="[1758,1802,847,866]" gridcol="19" gridrow="4" pageId="8" pageNumber="42">4.5</td>
<td id="76BE5DDFFF9093EEDD4C98716D27B013" box="[1830,1875,847,866]" gridcol="20" gridrow="4" pageId="8" pageNumber="42">3</td>
</tr>
<tr id="356F34A3FF9093EEDA1798516D27B0F3" box="[125,1875,879,898]" gridrow="5" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA1798516B52B0F3" box="[125,294,879,898]" gridcol="0" gridrow="5" pageId="8" pageNumber="42">Voloina</th>
<td id="76BE5DDFFF9093EEDB3198516BAFB0F3" box="[347,475,879,898]" gridcol="1" gridrow="5" pageId="8" pageNumber="42">ZFMK 52614</td>
<td id="76BE5DDFFF9093EED86A9851682EB0F3" box="[512,602,879,898]" gridcol="2" gridrow="5" pageId="8" pageNumber="42">n.a.</td>
<td id="76BE5DDFFF9093EED81D985168EBB0F3" box="[631,671,879,898]" gridcol="3" gridrow="5" pageId="8" pageNumber="42">23.7</td>
<td id="76BE5DDFFF9093EED8D698516895B0F3" box="[700,737,879,898]" gridcol="4" gridrow="5" pageId="8" pageNumber="42">6.7</td>
<td id="76BE5DDFFF9093EED8979851696FB0F3" box="[765,795,879,898]" gridcol="5" gridrow="5" pageId="8" pageNumber="42">10</td>
<td id="76BE5DDFFF9093EED95D98516920B0F3" box="[823,852,879,898]" gridcol="6" gridrow="5" pageId="8" pageNumber="42">1.6</td>
<td id="76BE5DDFFF9093EED91A985169F9B0F3" box="[880,909,879,898]" gridcol="7" gridrow="5" pageId="8" pageNumber="42">2.5</td>
<td id="76BE5DDFFF9093EED9C3985169A1B0F3" box="[937,981,879,898]" gridcol="8" gridrow="5" pageId="8" pageNumber="42">2.4</td>
<td id="76BE5DDFFF9093EED99B98516E6FB0F3" box="[1009,1051,879,898]" gridcol="9" gridrow="5" pageId="8" pageNumber="42">1.7</td>
<td id="76BE5DDFFF9093EEDE5D98516E10B0F3" box="[1079,1124,879,898]" gridcol="10" gridrow="5" pageId="8" pageNumber="42">2.5</td>
<td id="76BE5DDFFF9093EEDEEA98516ECEB0F3" box="[1152,1210,879,898]" gridcol="11" gridrow="5" pageId="8" pageNumber="42">14.8</td>
<td id="76BE5DDFFF9093EEDEBC98516F77B0F3" box="[1238,1283,879,898]" gridcol="12" gridrow="5" pageId="8" pageNumber="42">6.4</td>
<td id="76BE5DDFFF9093EEDF7598516F31B0F3" box="[1311,1349,879,898]" gridcol="13" gridrow="5" pageId="8" pageNumber="42">37</td>
<td id="76BE5DDFFF9093EEDF0B98516FEDB0F3" box="[1377,1433,879,898]" gridcol="14" gridrow="5" pageId="8" pageNumber="42">17</td>
<td id="76BE5DDFFF9093EEDFDF98516F94B0F3" box="[1461,1504,879,898]" gridcol="15" gridrow="5" pageId="8" pageNumber="42">11</td>
<td id="76BE5DDFFF9093EEDF9698516C5AB0F3" box="[1532,1582,879,898]" gridcol="16" gridrow="5" pageId="8" pageNumber="42">11.7</td>
<td id="76BE5DDFFF9093EEDC2098516C01B0F3" box="[1610,1653,879,898]" gridcol="17" gridrow="5" pageId="8" pageNumber="42">3.8</td>
<td id="76BE5DDFFF9093EEDCFB98516CB7B0F3" box="[1681,1731,879,898]" gridcol="18" gridrow="5" pageId="8" pageNumber="42">1.8</td>
<td id="76BE5DDFFF9093EEDCB498516D7EB0F3" box="[1758,1802,879,898]" gridcol="19" gridrow="5" pageId="8" pageNumber="42">3.9</td>
<td id="76BE5DDFFF9093EEDD4C98516D27B0F3" box="[1830,1875,879,898]" gridcol="20" gridrow="5" pageId="8" pageNumber="42">4</td>
</tr>
<tr id="356F34A3FF9093EEDA1798B16D27B0D3" box="[125,1875,911,930]" gridrow="6" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA1798B16B52B0D3" box="[125,294,911,930]" gridcol="0" gridrow="6" pageId="8" pageNumber="42">Voloina</th>
<td id="76BE5DDFFF9093EEDB3198B16BAFB0D3" box="[347,475,911,930]" gridcol="1" gridrow="6" pageId="8" pageNumber="42">ZFMK 52615</td>
<td id="76BE5DDFFF9093EED86A98B1682EB0D3" box="[512,602,911,930]" gridcol="2" gridrow="6" pageId="8" pageNumber="42">n.a.</td>
<td id="76BE5DDFFF9093EED81D98B168EBB0D3" box="[631,671,911,930]" gridcol="3" gridrow="6" pageId="8" pageNumber="42">21.3</td>
<td id="76BE5DDFFF9093EED8D698B16895B0D3" box="[700,737,911,930]" gridcol="4" gridrow="6" pageId="8" pageNumber="42">6.7</td>
<td id="76BE5DDFFF9093EED89798B1696FB0D3" box="[765,795,911,930]" gridcol="5" gridrow="6" pageId="8" pageNumber="42">8.5</td>
<td id="76BE5DDFFF9093EED95D98B16920B0D3" box="[823,852,911,930]" gridcol="6" gridrow="6" pageId="8" pageNumber="42">1.4</td>
<td id="76BE5DDFFF9093EED91A98B169F9B0D3" box="[880,909,911,930]" gridcol="7" gridrow="6" pageId="8" pageNumber="42">2.5</td>
<td id="76BE5DDFFF9093EED9C398B169A1B0D3" box="[937,981,911,930]" gridcol="8" gridrow="6" pageId="8" pageNumber="42">2.4</td>
<td id="76BE5DDFFF9093EED99B98B16E6FB0D3" box="[1009,1051,911,930]" gridcol="9" gridrow="6" pageId="8" pageNumber="42">1.6</td>
<td id="76BE5DDFFF9093EEDE5D98B16E10B0D3" box="[1079,1124,911,930]" gridcol="10" gridrow="6" pageId="8" pageNumber="42">2.8</td>
<td id="76BE5DDFFF9093EEDEEA98B16ECEB0D3" box="[1152,1210,911,930]" gridcol="11" gridrow="6" pageId="8" pageNumber="42">12.7</td>
<td id="76BE5DDFFF9093EEDEBC98B16F77B0D3" box="[1238,1283,911,930]" gridcol="12" gridrow="6" pageId="8" pageNumber="42">6.5</td>
<td id="76BE5DDFFF9093EEDF7598B16F31B0D3" box="[1311,1349,911,930]" gridcol="13" gridrow="6" pageId="8" pageNumber="42">37.7</td>
<td id="76BE5DDFFF9093EEDF0B98B16FEDB0D3" box="[1377,1433,911,930]" gridcol="14" gridrow="6" pageId="8" pageNumber="42">16.5</td>
<td id="76BE5DDFFF9093EEDFDF98B16F94B0D3" box="[1461,1504,911,930]" gridcol="15" gridrow="6" pageId="8" pageNumber="42">10.5</td>
<td id="76BE5DDFFF9093EEDF9698B16C5AB0D3" box="[1532,1582,911,930]" gridcol="16" gridrow="6" pageId="8" pageNumber="42">11.4</td>
<td id="76BE5DDFFF9093EEDC2098B16C01B0D3" box="[1610,1653,911,930]" gridcol="17" gridrow="6" pageId="8" pageNumber="42">3.7</td>
<td id="76BE5DDFFF9093EEDCFB98B16CB7B0D3" box="[1681,1731,911,930]" gridcol="18" gridrow="6" pageId="8" pageNumber="42">1.7</td>
<td id="76BE5DDFFF9093EEDCB498B16D7EB0D3" box="[1758,1802,911,930]" gridcol="19" gridrow="6" pageId="8" pageNumber="42">3.6</td>
<td id="76BE5DDFFF9093EEDD4C98B16D27B0D3" box="[1830,1875,911,930]" gridcol="20" gridrow="6" pageId="8" pageNumber="42">4</td>
</tr>
<tr id="356F34A3FF9093EEDA1798916D27B753" box="[125,1875,943,1058]" gridrow="7" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA1798916B52B753" box="[125,294,943,1058]" gridcol="0" gridrow="7" pageId="8" pageNumber="42">Ambodivoahangy Ambodivoahangy Maroantsetra</th>
<td id="76BE5DDFFF9093EEDB3198916BAFB753" box="[347,475,943,1058]" gridcol="1" gridrow="7" pageId="8" pageNumber="42">ZSM 236/2010 ZSM 237/2010 ZMA 19509</td>
<td id="76BE5DDFFF9093EED86A9891682EB753" box="[512,602,943,1058]" gridcol="2" gridrow="7" pageId="8" pageNumber="42">FGZC 4289 FGZC 4305 n.a.</td>
<td id="76BE5DDFFF9093EED81D989168EBB753" box="[631,671,943,1058]" gridcol="3" gridrow="7" pageId="8" pageNumber="42">21.4 23.7 22.3</td>
<td id="76BE5DDFFF9093EED8D698916895B753" box="[700,737,943,1058]" gridcol="4" gridrow="7" pageId="8" pageNumber="42">6.3 6.8 6.4</td>
<td id="76BE5DDFFF9093EED8979891696FB753" box="[765,795,943,1058]" gridcol="5" gridrow="7" pageId="8" pageNumber="42">8.2 8.9 8.1</td>
<td id="76BE5DDFFF9093EED95D98916920B753" box="[823,852,943,1058]" gridcol="6" gridrow="7" pageId="8" pageNumber="42">1.5 1.5 1.3</td>
<td id="76BE5DDFFF9093EED91A989169F9B753" box="[880,909,943,1058]" gridcol="7" gridrow="7" pageId="8" pageNumber="42">2.5 2.8 2.5</td>
<td id="76BE5DDFFF9093EED9C3989169A1B753" box="[937,981,943,1058]" gridcol="8" gridrow="7" pageId="8" pageNumber="42">2.1 2.5 2.4</td>
<td id="76BE5DDFFF9093EED99B98916E6FB753" box="[1009,1051,943,1058]" gridcol="9" gridrow="7" pageId="8" pageNumber="42">1.5 1.5 1.2</td>
<td id="76BE5DDFFF9093EEDE5D98916E10B753" box="[1079,1124,943,1058]" gridcol="10" gridrow="7" pageId="8" pageNumber="42">2.1 2.8 2.4</td>
<td id="76BE5DDFFF9093EEDEEA98916ECEB753" box="[1152,1210,943,1058]" gridcol="11" gridrow="7" pageId="8" pageNumber="42">14.2 16 13.8</td>
<td id="76BE5DDFFF9093EEDEBC98916F77B753" box="[1238,1283,943,1058]" gridcol="12" gridrow="7" pageId="8" pageNumber="42">6.6 7.2 6.3</td>
<td id="76BE5DDFFF9093EEDF7598916F31B753" box="[1311,1349,943,1058]" gridcol="13" gridrow="7" pageId="8" pageNumber="42">n.a. 40.9 34</td>
<td id="76BE5DDFFF9093EEDF0B98916FEDB753" box="[1377,1433,943,1058]" gridcol="14" gridrow="7" pageId="8" pageNumber="42">n.a. 18.5 15.8</td>
<td id="76BE5DDFFF9093EEDFDF98916F94B753" box="[1461,1504,943,1058]" gridcol="15" gridrow="7" pageId="8" pageNumber="42">n.a. 12.3 10.3</td>
<td id="76BE5DDFFF9093EEDF9698916C5AB753" box="[1532,1582,943,1058]" gridcol="16" gridrow="7" pageId="8" pageNumber="42">12.2 12.9 n.a.</td>
<td id="76BE5DDFFF9093EEDC2098916C01B753" box="[1610,1653,943,1058]" gridcol="17" gridrow="7" pageId="8" pageNumber="42">4.8 5.2 3.5</td>
<td id="76BE5DDFFF9093EEDCFB98916CB7B753" box="[1681,1731,943,1058]" gridcol="18" gridrow="7" pageId="8" pageNumber="42">1.5 1.5 1.1</td>
<td id="76BE5DDFFF9093EEDCB498916D7EB753" box="[1758,1802,943,1058]" gridcol="19" gridrow="7" pageId="8" pageNumber="42">3.1 2.8 n.a.</td>
<td id="76BE5DDFFF9093EEDD4C98916D27B753" box="[1830,1875,943,1058]" gridcol="20" gridrow="7" pageId="8" pageNumber="42">5 4 1</td>
</tr>
<tr id="356F34A3FF9093EEDA179F0C6D27B6AC" box="[125,1875,1074,1501]" gridrow="8" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA179F0C6B52B6AC" box="[125,294,1074,1501]" gridcol="0" gridrow="8" pageId="8" pageNumber="42">
<taxonomicName id="4C5F4D62FF906C19DAED9F0C6A83B6DF" authority="Nosy Be Nosy Be Nosy Be Montagne d'Ambre Montagne d'Ambre Montagne d'Ambre Montagne d'Ambre Maevatanana" authorityName="Nosy Be Nosy Be Nosy Be Montagne d'Ambre Montagne d'Ambre Montagne d'Ambre Montagne d'Ambre Maevatanana" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="8" pageNumber="42" phylum="Chordata" rank="species" species="wittei">
<emphasis id="B92BEAF3FF906C19DAED9F0C6AB8B734" bold="true" box="[135,204,1074,1093]" italics="true" pageId="8" pageNumber="42">B. wittei</emphasis>
Nosy Be Nosy Be Nosy Be Montagne d'Ambre Montagne d'Ambre Montagne d'Ambre Montagne d'Ambre Maevatanana
</taxonomicName>
/ Manongarivo Nosy Be
</th>
<td id="76BE5DDFFF9093EEDB319F0C6BAFB6AC" box="[347,475,1074,1501]" gridcol="1" gridrow="8" pageId="8" pageNumber="42">ZSM 55/2002 ZSM 56/2002 ZSM 57/2002 ZSM 879/2003 ZSM 880/2003 ZSM 881/2003 ZSM 882/2003 ZMA 19503 ZMA 7160</td>
<td id="76BE5DDFFF9093EED86A9F0C682EB6AC" box="[512,602,1074,1501]" gridcol="2" gridrow="8" pageId="8" pageNumber="42">MV 2001.1253 MV 2001.1255 MV 2001.1265 FGMV 2002.876 FGMV 2002.878 FGMV 2002.879 FGMV 2002.880 n.a. 594</td>
<td id="76BE5DDFFF9093EED81D9F0C68EBB6AC" box="[631,671,1074,1501]" gridcol="3" gridrow="8" pageId="8" pageNumber="42">24.3 23.7 23.0 22.7 21.8 21.0 22.0 20.3 24.0</td>
<td id="76BE5DDFFF9093EED8D69F0C6895B6AC" box="[700,737,1074,1501]" gridcol="4" gridrow="8" pageId="8" pageNumber="42">7.7 7.5 7.2 7.2 7.0 6.7 7.0 6.1 7.0</td>
<td id="76BE5DDFFF9093EED8979F0C696FB6AC" box="[765,795,1074,1501]" gridcol="5" gridrow="8" pageId="8" pageNumber="42">9.5 9.3 9.0 8.7 8.3 8.0 8.3 8.0 9.1</td>
<td id="76BE5DDFFF9093EED95D9F0C6920B6AC" box="[823,852,1074,1501]" gridcol="6" gridrow="8" pageId="8" pageNumber="42">2.2 1.6 1.6 1.7 1.5 1.6 1.5 1.4 1.6</td>
<td id="76BE5DDFFF9093EED91A9F0C69F9B6AC" box="[880,909,1074,1501]" gridcol="7" gridrow="8" pageId="8" pageNumber="42">2.6 2.8 3.0 2.7 2.3 2.5 2.5 2.4 2.7</td>
<td id="76BE5DDFFF9093EED9C39F0C69A1B6AC" box="[937,981,1074,1501]" gridcol="8" gridrow="8" pageId="8" pageNumber="42">2.5 2.1 2.2 2.2 1.9 2.0 2.0 2.2 2.0</td>
<td id="76BE5DDFFF9093EED99B9F0C6E6FB6AC" box="[1009,1051,1074,1501]" gridcol="9" gridrow="8" pageId="8" pageNumber="42">1.7 1.7 1.5 2.0 1.6 1.6 1.6 1.5 1.5</td>
<td id="76BE5DDFFF9093EEDE5D9F0C6E10B6AC" box="[1079,1124,1074,1501]" gridcol="10" gridrow="8" pageId="8" pageNumber="42">2.5 2.6 2.4 2.8 2.6 2.6 2.3 2.5 2.5</td>
<td id="76BE5DDFFF9093EEDEEA9F0C6ECEB6AC" box="[1152,1210,1074,1501]" gridcol="11" gridrow="8" pageId="8" pageNumber="42">15.9 15.7 15.3 15.0 14.2 13.6 14.1 13.6 16.3</td>
<td id="76BE5DDFFF9093EEDEBC9F0C6F77B6AC" box="[1238,1283,1074,1501]" gridcol="12" gridrow="8" pageId="8" pageNumber="42">7.5 7.5 7.2 7.0 6.7 6.3 6.6 6.0 6.9</td>
<td id="76BE5DDFFF9093EEDF759F0C6F31B6AC" box="[1311,1349,1074,1501]" gridcol="13" gridrow="8" pageId="8" pageNumber="42">41.0 41.0 38.4 35.3 36.3 34.6 35.3 36.0 40.0</td>
<td id="76BE5DDFFF9093EEDF0B9F0C6FEDB6AC" box="[1377,1433,1074,1501]" gridcol="14" gridrow="8" pageId="8" pageNumber="42">19.4 19.0 17.7 16.8 17.2 15.7 16.5 16.2 19.0</td>
<td id="76BE5DDFFF9093EEDFDF9F0C6F94B6AC" box="[1461,1504,1074,1501]" gridcol="15" gridrow="8" pageId="8" pageNumber="42">13.0 12.5 11.9 11.0 11.3 10.4 10.7 10.5 12.4</td>
<td id="76BE5DDFFF9093EEDF969F0C6C5AB6AC" box="[1532,1582,1074,1501]" gridcol="16" gridrow="8" pageId="8" pageNumber="42">n.a. n.a. n.a. n.a. n.a. n.a. n.a. n.a. n.a.</td>
<td id="76BE5DDFFF9093EEDC209F0C6C01B6AC" box="[1610,1653,1074,1501]" gridcol="17" gridrow="8" pageId="8" pageNumber="42">4.1 4.2 4.0 3.8 3.6 4.2 4.7 4.4 3.4</td>
<td id="76BE5DDFFF9093EEDCFB9F0C6CB7B6AC" box="[1681,1731,1074,1501]" gridcol="18" gridrow="8" pageId="8" pageNumber="42">2.3 2.0 1.6 1.9 1.9 1.7 1.7 2.0 1.7</td>
<td id="76BE5DDFFF9093EEDCB49F0C6D7EB6AC" box="[1758,1802,1074,1501]" gridcol="19" gridrow="8" pageId="8" pageNumber="42">1.1 1.3 1.9 2.5 2.4 2.3 1.8 8.0 2.0</td>
<td id="76BE5DDFFF9093EEDD4C9F0C6D27B6AC" box="[1830,1875,1074,1501]" gridcol="20" gridrow="8" pageId="8" pageNumber="42">3 3 3 1 4 3 1 4 3</td>
</tr>
<tr id="356F34A3FF9093EEDA179ED36D27B571" box="[125,1875,1517,1536]" gridrow="9" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA179ED36B52B571" box="[125,294,1517,1536]" gridcol="0" gridrow="9" pageId="8" pageNumber="42">Nosy Be</th>
<td id="76BE5DDFFF9093EEDB319ED36BAFB571" box="[347,475,1517,1536]" gridcol="1" gridrow="9" pageId="8" pageNumber="42">ZMA 7160</td>
<td id="76BE5DDFFF9093EED86A9ED3682EB571" box="[512,602,1517,1536]" gridcol="2" gridrow="9" pageId="8" pageNumber="42">595</td>
<td id="76BE5DDFFF9093EED81D9ED368EBB571" box="[631,671,1517,1536]" gridcol="3" gridrow="9" pageId="8" pageNumber="42">25.0</td>
<td id="76BE5DDFFF9093EED8D69ED36895B571" box="[700,737,1517,1536]" gridcol="4" gridrow="9" pageId="8" pageNumber="42">7.7</td>
<td id="76BE5DDFFF9093EED8979ED3696FB571" box="[765,795,1517,1536]" gridcol="5" gridrow="9" pageId="8" pageNumber="42">9.2</td>
<td id="76BE5DDFFF9093EED95D9ED36920B571" box="[823,852,1517,1536]" gridcol="6" gridrow="9" pageId="8" pageNumber="42">1.7</td>
<td id="76BE5DDFFF9093EED91A9ED369F9B571" box="[880,909,1517,1536]" gridcol="7" gridrow="9" pageId="8" pageNumber="42">2.8</td>
<td id="76BE5DDFFF9093EED9C39ED369A1B571" box="[937,981,1517,1536]" gridcol="8" gridrow="9" pageId="8" pageNumber="42">2.3</td>
<td id="76BE5DDFFF9093EED99B9ED36E6FB571" box="[1009,1051,1517,1536]" gridcol="9" gridrow="9" pageId="8" pageNumber="42">1.6</td>
<td id="76BE5DDFFF9093EEDE5D9ED36E10B571" box="[1079,1124,1517,1536]" gridcol="10" gridrow="9" pageId="8" pageNumber="42">2.4</td>
<td id="76BE5DDFFF9093EEDEEA9ED36ECEB571" box="[1152,1210,1517,1536]" gridcol="11" gridrow="9" pageId="8" pageNumber="42">16.1</td>
<td id="76BE5DDFFF9093EEDEBC9ED36F77B571" box="[1238,1283,1517,1536]" gridcol="12" gridrow="9" pageId="8" pageNumber="42">7.6</td>
<td id="76BE5DDFFF9093EEDF759ED36F31B571" box="[1311,1349,1517,1536]" gridcol="13" gridrow="9" pageId="8" pageNumber="42">41.1</td>
<td id="76BE5DDFFF9093EEDF0B9ED36FEDB571" box="[1377,1433,1517,1536]" gridcol="14" gridrow="9" pageId="8" pageNumber="42">19.7</td>
<td id="76BE5DDFFF9093EEDFDF9ED36F94B571" box="[1461,1504,1517,1536]" gridcol="15" gridrow="9" pageId="8" pageNumber="42">13.2</td>
<td id="76BE5DDFFF9093EEDF969ED36C5AB571" box="[1532,1582,1517,1536]" gridcol="16" gridrow="9" pageId="8" pageNumber="42">n.a.</td>
<td id="76BE5DDFFF9093EEDC209ED36C01B571" box="[1610,1653,1517,1536]" gridcol="17" gridrow="9" pageId="8" pageNumber="42">3.6</td>
<td id="76BE5DDFFF9093EEDCFB9ED36CB7B571" box="[1681,1731,1517,1536]" gridcol="18" gridrow="9" pageId="8" pageNumber="42">1.6</td>
<td id="76BE5DDFFF9093EEDCB49ED36D7EB571" box="[1758,1802,1517,1536]" gridcol="19" gridrow="9" pageId="8" pageNumber="42">2.1</td>
<td id="76BE5DDFFF9093EEDD4C9ED36D27B571" box="[1830,1875,1517,1536]" gridcol="20" gridrow="9" pageId="8" pageNumber="42">3</td>
</tr>
<tr id="356F34A3FF9093EEDA179D316D27B553" box="[125,1875,1551,1570]" gridrow="10" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA179D316B52B553" box="[125,294,1551,1570]" gridcol="0" gridrow="10" pageId="8" pageNumber="42">Ankarafantsika</th>
<td id="76BE5DDFFF9093EEDB319D316BAFB553" box="[347,475,1551,1570]" gridcol="1" gridrow="10" pageId="8" pageNumber="42">ZMA 6870</td>
<td id="76BE5DDFFF9093EED86A9D31682EB553" box="[512,602,1551,1570]" gridcol="2" gridrow="10" pageId="8" pageNumber="42">1110</td>
<td id="76BE5DDFFF9093EED81D9D3168EBB553" box="[631,671,1551,1570]" gridcol="3" gridrow="10" pageId="8" pageNumber="42">24.0</td>
<td id="76BE5DDFFF9093EED8D69D316895B553" box="[700,737,1551,1570]" gridcol="4" gridrow="10" pageId="8" pageNumber="42">7.0</td>
<td id="76BE5DDFFF9093EED8979D31696FB553" box="[765,795,1551,1570]" gridcol="5" gridrow="10" pageId="8" pageNumber="42">9.1</td>
<td id="76BE5DDFFF9093EED95D9D316920B553" box="[823,852,1551,1570]" gridcol="6" gridrow="10" pageId="8" pageNumber="42">1.8</td>
<td id="76BE5DDFFF9093EED91A9D3169F9B553" box="[880,909,1551,1570]" gridcol="7" gridrow="10" pageId="8" pageNumber="42">2.5</td>
<td id="76BE5DDFFF9093EED9C39D3169A1B553" box="[937,981,1551,1570]" gridcol="8" gridrow="10" pageId="8" pageNumber="42">2.3</td>
<td id="76BE5DDFFF9093EED99B9D316E6FB553" box="[1009,1051,1551,1570]" gridcol="9" gridrow="10" pageId="8" pageNumber="42">1.5</td>
<td id="76BE5DDFFF9093EEDE5D9D316E10B553" box="[1079,1124,1551,1570]" gridcol="10" gridrow="10" pageId="8" pageNumber="42">2.5</td>
<td id="76BE5DDFFF9093EEDEEA9D316ECEB553" box="[1152,1210,1551,1570]" gridcol="11" gridrow="10" pageId="8" pageNumber="42">14.6</td>
<td id="76BE5DDFFF9093EEDEBC9D316F77B553" box="[1238,1283,1551,1570]" gridcol="12" gridrow="10" pageId="8" pageNumber="42">8.5</td>
<td id="76BE5DDFFF9093EEDF759D316F31B553" box="[1311,1349,1551,1570]" gridcol="13" gridrow="10" pageId="8" pageNumber="42">37.9</td>
<td id="76BE5DDFFF9093EEDF0B9D316FEDB553" box="[1377,1433,1551,1570]" gridcol="14" gridrow="10" pageId="8" pageNumber="42">18.0</td>
<td id="76BE5DDFFF9093EEDFDF9D316F94B553" box="[1461,1504,1551,1570]" gridcol="15" gridrow="10" pageId="8" pageNumber="42">12.2</td>
<td id="76BE5DDFFF9093EEDF969D316C5AB553" box="[1532,1582,1551,1570]" gridcol="16" gridrow="10" pageId="8" pageNumber="42">n.a.</td>
<td id="76BE5DDFFF9093EEDC209D316C01B553" box="[1610,1653,1551,1570]" gridcol="17" gridrow="10" pageId="8" pageNumber="42">4.0</td>
<td id="76BE5DDFFF9093EEDCFB9D316CB7B553" box="[1681,1731,1551,1570]" gridcol="18" gridrow="10" pageId="8" pageNumber="42">1.7</td>
<td id="76BE5DDFFF9093EEDCB49D316D7EB553" box="[1758,1802,1551,1570]" gridcol="19" gridrow="10" pageId="8" pageNumber="42">1.4</td>
<td id="76BE5DDFFF9093EEDD4C9D316D27B553" box="[1830,1875,1551,1570]" gridcol="20" gridrow="10" pageId="8" pageNumber="42">2</td>
</tr>
<tr id="356F34A3FF9093EEDA179D0F6D27B535" box="[125,1875,1585,1604]" gridrow="11" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA179D0F6B52B535" box="[125,294,1585,1604]" gridcol="0" gridrow="11" pageId="8" pageNumber="42">Ankarafantsika</th>
<td id="76BE5DDFFF9093EEDB319D0F6BAFB535" box="[347,475,1585,1604]" gridcol="1" gridrow="11" pageId="8" pageNumber="42">ZMA 6870</td>
<td id="76BE5DDFFF9093EED86A9D0F682EB535" box="[512,602,1585,1604]" gridcol="2" gridrow="11" pageId="8" pageNumber="42">1108</td>
<td id="76BE5DDFFF9093EED81D9D0F68EBB535" box="[631,671,1585,1604]" gridcol="3" gridrow="11" pageId="8" pageNumber="42">23.1</td>
<td id="76BE5DDFFF9093EED8D69D0F6895B535" box="[700,737,1585,1604]" gridcol="4" gridrow="11" pageId="8" pageNumber="42">6.9</td>
<td id="76BE5DDFFF9093EED8979D0F696FB535" box="[765,795,1585,1604]" gridcol="5" gridrow="11" pageId="8" pageNumber="42">8.6</td>
<td id="76BE5DDFFF9093EED95D9D0F6920B535" box="[823,852,1585,1604]" gridcol="6" gridrow="11" pageId="8" pageNumber="42">1.5</td>
<td id="76BE5DDFFF9093EED91A9D0F69F9B535" box="[880,909,1585,1604]" gridcol="7" gridrow="11" pageId="8" pageNumber="42">2.8</td>
<td id="76BE5DDFFF9093EED9C39D0F69A1B535" box="[937,981,1585,1604]" gridcol="8" gridrow="11" pageId="8" pageNumber="42">1.9</td>
<td id="76BE5DDFFF9093EED99B9D0F6E6FB535" box="[1009,1051,1585,1604]" gridcol="9" gridrow="11" pageId="8" pageNumber="42">1.4</td>
<td id="76BE5DDFFF9093EEDE5D9D0F6E10B535" box="[1079,1124,1585,1604]" gridcol="10" gridrow="11" pageId="8" pageNumber="42">2.1</td>
<td id="76BE5DDFFF9093EEDEEA9D0F6ECEB535" box="[1152,1210,1585,1604]" gridcol="11" gridrow="11" pageId="8" pageNumber="42">15.1</td>
<td id="76BE5DDFFF9093EEDEBC9D0F6F77B535" box="[1238,1283,1585,1604]" gridcol="12" gridrow="11" pageId="8" pageNumber="42">7.2</td>
<td id="76BE5DDFFF9093EEDF759D0F6F31B535" box="[1311,1349,1585,1604]" gridcol="13" gridrow="11" pageId="8" pageNumber="42">40.4</td>
<td id="76BE5DDFFF9093EEDF0B9D0F6FEDB535" box="[1377,1433,1585,1604]" gridcol="14" gridrow="11" pageId="8" pageNumber="42">18.4</td>
<td id="76BE5DDFFF9093EEDFDF9D0F6F94B535" box="[1461,1504,1585,1604]" gridcol="15" gridrow="11" pageId="8" pageNumber="42">12.7</td>
<td id="76BE5DDFFF9093EEDF969D0F6C5AB535" box="[1532,1582,1585,1604]" gridcol="16" gridrow="11" pageId="8" pageNumber="42">n.a.</td>
<td id="76BE5DDFFF9093EEDC209D0F6C01B535" box="[1610,1653,1585,1604]" gridcol="17" gridrow="11" pageId="8" pageNumber="42">4.0</td>
<td id="76BE5DDFFF9093EEDCFB9D0F6CB7B535" box="[1681,1731,1585,1604]" gridcol="18" gridrow="11" pageId="8" pageNumber="42">1.9</td>
<td id="76BE5DDFFF9093EEDCB49D0F6D7EB535" box="[1758,1802,1585,1604]" gridcol="19" gridrow="11" pageId="8" pageNumber="42">2.2</td>
<td id="76BE5DDFFF9093EEDD4C9D0F6D27B535" box="[1830,1875,1585,1604]" gridcol="20" gridrow="11" pageId="8" pageNumber="42">1</td>
</tr>
<tr id="356F34A3FF9093EEDA179D6D6D27B50C" box="[125,1875,1619,1661]" gridrow="12" pageId="8" pageNumber="42">
<th id="76BE5DDFFF9093EEDA179D6D6B52B50C" box="[125,294,1619,1661]" gridcol="0" gridrow="12" pageId="8" pageNumber="42">Ankarafantsika Ankarafantsika</th>
<td id="76BE5DDFFF9093EEDB319D6D6BAFB50C" box="[347,475,1619,1661]" gridcol="1" gridrow="12" pageId="8" pageNumber="42">ZMA 6870 ZMA 6870</td>
<td id="76BE5DDFFF9093EED86A9D6D682EB50C" box="[512,602,1619,1661]" gridcol="2" gridrow="12" pageId="8" pageNumber="42">1109 1111</td>
<td id="76BE5DDFFF9093EED81D9D6D68EBB50C" box="[631,671,1619,1661]" gridcol="3" gridrow="12" pageId="8" pageNumber="42">23.0 23.0</td>
<td id="76BE5DDFFF9093EED8D69D6D6895B50C" box="[700,737,1619,1661]" gridcol="4" gridrow="12" pageId="8" pageNumber="42">6.6 6.4</td>
<td id="76BE5DDFFF9093EED8979D6D696FB50C" box="[765,795,1619,1661]" gridcol="5" gridrow="12" pageId="8" pageNumber="42">8.5 8.5</td>
<td id="76BE5DDFFF9093EED95D9D6D6920B50C" box="[823,852,1619,1661]" gridcol="6" gridrow="12" pageId="8" pageNumber="42">1.6 1.4</td>
<td id="76BE5DDFFF9093EED91A9D6D69F9B50C" box="[880,909,1619,1661]" gridcol="7" gridrow="12" pageId="8" pageNumber="42">2.5 2.8</td>
<td id="76BE5DDFFF9093EED9C39D6D69A1B50C" box="[937,981,1619,1661]" gridcol="8" gridrow="12" pageId="8" pageNumber="42">1.7 2.2</td>
<td id="76BE5DDFFF9093EED99B9D6D6E6FB50C" box="[1009,1051,1619,1661]" gridcol="9" gridrow="12" pageId="8" pageNumber="42">1.6 1.6</td>
<td id="76BE5DDFFF9093EEDE5D9D6D6E10B50C" box="[1079,1124,1619,1661]" gridcol="10" gridrow="12" pageId="8" pageNumber="42">2.4 2.2</td>
<td id="76BE5DDFFF9093EEDEEA9D6D6ECEB50C" box="[1152,1210,1619,1661]" gridcol="11" gridrow="12" pageId="8" pageNumber="42">14.7 15.6</td>
<td id="76BE5DDFFF9093EEDEBC9D6D6F77B50C" box="[1238,1283,1619,1661]" gridcol="12" gridrow="12" pageId="8" pageNumber="42">6.6 6.6</td>
<td id="76BE5DDFFF9093EEDF759D6D6F31B50C" box="[1311,1349,1619,1661]" gridcol="13" gridrow="12" pageId="8" pageNumber="42">38.8 38.2</td>
<td id="76BE5DDFFF9093EEDF0B9D6D6FEDB50C" box="[1377,1433,1619,1661]" gridcol="14" gridrow="12" pageId="8" pageNumber="42">17.7 17.3</td>
<td id="76BE5DDFFF9093EEDFDF9D6D6F94B50C" box="[1461,1504,1619,1661]" gridcol="15" gridrow="12" pageId="8" pageNumber="42">11.8 11.4</td>
<td id="76BE5DDFFF9093EEDF969D6D6C5AB50C" box="[1532,1582,1619,1661]" gridcol="16" gridrow="12" pageId="8" pageNumber="42">n.a. n.a.</td>
<td id="76BE5DDFFF9093EEDC209D6D6C01B50C" box="[1610,1653,1619,1661]" gridcol="17" gridrow="12" pageId="8" pageNumber="42">4.4 3.6</td>
<td id="76BE5DDFFF9093EEDCFB9D6D6CB7B50C" box="[1681,1731,1619,1661]" gridcol="18" gridrow="12" pageId="8" pageNumber="42">1.5 1.8</td>
<td id="76BE5DDFFF9093EEDCB49D6D6D7EB50C" box="[1758,1802,1619,1661]" gridcol="19" gridrow="12" pageId="8" pageNumber="42">2.0 1.4</td>
<td id="76BE5DDFFF9093EEDD4C9D6D6D27B50C" box="[1830,1875,1619,1661]" gridcol="20" gridrow="12" pageId="8" pageNumber="42">1 1</td>
</tr>
</table>
</paragraph>
<caption id="DF206669FF916C18DAFD9C8A6AB4B498" httpUri="https://zenodo.org/record/207737/files/figure.png" pageId="9" pageNumber="43" targetBox="[214,1382,181,1925]" targetPageId="9">
<paragraph id="8BE036E1FF916C18DAFD9C8A6AB4B498" blockId="9.[151,1436,1972,2025]" pageId="9" pageNumber="43">
<emphasis id="B92BEAF3FF916C18DAFD9C8A6B67B4BB" bold="true" box="[151,275,1972,1995]" pageId="9" pageNumber="43">FIGURE 5.</emphasis>
Dorsolateral (a) and ventral (b) views of the male holotype of 
<taxonomicName id="4C5F4D62FF916C18D9FE9C8B6E03B4BA" box="[916,1143,1973,1995]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="43" phylum="Chordata" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B92BEAF3FF916C18D9FE9C8B6E03B4BA" box="[916,1143,1973,1995]" italics="true" pageId="9" pageNumber="43">Blommersia variabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2185788FF916C18DE149C8B6EBBB4BB" box="[1150,1231,1973,1994]" pageId="9" pageNumber="43" rank="species">
<emphasis id="B92BEAF3FF916C18DE149C8B6EBBB4BB" bold="true" box="[1150,1231,1973,1994]" pageId="9" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
(ZSM 237/2010) in life.
</paragraph>
</caption>
<paragraph id="8BE036E1FF926C1BDAAF9BA9692BB2AB" blockId="10.[151,1437,151,1968]" pageId="10" pageNumber="44">
After nine months in preservative, the dorsum appears greyish-brown because of many densely spaced, single dark melanophores. A translucent pink colour covers the entire dorsum, but is more prominent between and around the eye sockets and the tympanum. A thin, single medio-dorsal whitish line stretches from the neck in a posterior direction until the cloaca. There is a moderately distinct colour border between the flanks and the lighter dorsum. The hind limbs are light brown with four distinct dark brown crossbands on the thighs, three indistinct crossbands on tibia; tarsus light brown. The arms show indistinct, irregular dark patterns. Posterior to the tympanum a lateral dark streak runs along the supratympanic fold and ends at the forelimb insertion. Ventrally, the 
<typeStatus id="54E48843FF926C1BDEAF9A496F53B2FE" box="[1221,1319,375,399]" pageId="10" pageNumber="44" type="holotype">holotype</typeStatus>
was whitish with single, small and irregular dark melanophores. Throat almost uniformly whitish. Femoral glands largely white, contrasting with the yellowish colour of the inner thighs.
</paragraph>
<paragraph id="8BE036E1FF926C1BDAAF9AD66966B05B" blockId="10.[151,1437,151,1968]" pageId="10" pageNumber="44">
In life the 
<typeStatus id="54E48843FF926C1BDB509AD96BE8B28E" box="[314,412,487,511]" pageId="10" pageNumber="44" type="holotype">holotype</typeStatus>
(
<figureCitation id="13642A64FF926C1BDBC69AD96B9BB28E" box="[428,495,487,511]" captionStart="FIGURE 5" captionStartId="9.[151,250,1972,1995]" captionTargetBox="[214,1382,181,1925]" captionTargetId="figure@9.[214,1383,181,1925]" captionTargetPageId="9" captionText="FIGURE 5. Dorsolateral (a) and ventral (b) views of the male holotype of Blommersia variabilis sp. nov. (ZSM 237 / 2010) in life." httpUri="https://zenodo.org/record/207737/files/figure.png" pageId="10" pageNumber="44">Fig. 5</figureCitation>
) had a brown dorsum with a reddish hue on the anterior half of the body, including the head and neck, whereas the posterior is grayish-brown. Indistinct, gray crossbands line the limbs and a distinct white vertebral stripe started in the neck region and continued down the length of the body. Iris golden dorsally, ventrally poorly recognizable due to the widely opened pupil. Ventrally, the throat is yellow, but the rest of the body is white with some yellow mottling along the sternum and along the sides. The undersides of the front limbs are pinkish-blue. The ventral side of the hindlimbs is a darker blue, especially on the digits and joints. The ventral side of the thighs is pinkish-blue with yellow, clearly discernible femoral glands. The skin on the venter was translucent such that larger blood vessels could be easily discerned. The ventral colouration extends slightly onto the flanks, merging rather indistinctly with the dorsal brown colour.
</paragraph>
</subSubSection>
<subSubSection id="C345656AFF926C1BDAAD980668E7B0EB" pageId="10" pageNumber="44" type="discussion">
<paragraph id="8BE036E1FF926C1BDAAD980668E7B0EB" blockId="10.[151,1437,151,1968]" pageId="10" pageNumber="44">
<emphasis id="B92BEAF3FF926C1BDAAD98066B44B021" bold="true" box="[199,304,824,848]" pageId="10" pageNumber="44">Remark.</emphasis>
This species was referred to as 
<taxonomicName id="4C5F4D62FF926C1BD8F698066957B021" box="[668,803,824,848]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B92BEAF3FF926C1BD8F698066957B021" box="[668,803,824,848]" italics="true" pageId="10" pageNumber="44">Blommersia</emphasis>
</taxonomicName>
sp. 
<taxonomicName id="4C5F4D62FF926C1BD93998096E7EB021" authority="Maroantsetra" authorityName="Maroantsetra" box="[851,1034,823,848]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="species" species="blommersae">
aff. 
<emphasis id="B92BEAF3FF926C1BD9EB98066E7EB021" box="[897,1034,824,848]" italics="true" pageId="10" pageNumber="44">blommersae</emphasis>
</taxonomicName>
“Maroantsetra” by 
<bibRefCitation id="EFCE4B10FF926C1BDE8098096A90B004" author="Glaw" pageId="10" pageNumber="44" refString="Glaw, F., &amp; Vences, M. (2007) A Field Guide to the Amphibians and Reptiles of Madagascar. Third edition. Vences &amp; Glaw Verlag, Cologne, 496 pp." type="book" year="2007">Glaw &amp; Vences (2007)</bibRefCitation>
, 
<taxonomicName id="4C5F4D62FF926C1BDA9B98636B0CB004" box="[241,376,861,885]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B92BEAF3FF926C1BDA9B98636B0CB004" box="[241,376,861,885]" italics="true" pageId="10" pageNumber="44">Blommersia</emphasis>
</taxonomicName>
sp. 
<taxonomicName id="4C5F4D62FF926C1BDBC3986368B7B004" authority="Maroantsetra" authorityName="Maroantsetra" box="[425,707,861,885]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="species" species="wittei">
aff. 
<emphasis id="B92BEAF3FF926C1BDBBD98606861B004" box="[471,533,862,885]" italics="true" pageId="10" pageNumber="44">wittei</emphasis>
(Maroantsetra)
</taxonomicName>
by Vences 
<emphasis id="B92BEAF3FF926C1BD922986069F6B004" box="[840,898,861,885]" italics="true" pageId="10" pageNumber="44">et al.</emphasis>
(2006), and 
<taxonomicName id="4C5F4D62FF926C1BDE7F98636EE9B004" box="[1045,1181,861,885]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B92BEAF3FF926C1BDE7F98636EE9B004" box="[1045,1181,861,885]" italics="true" pageId="10" pageNumber="44">Blommersia</emphasis>
</taxonomicName>
sp. 2 Maroantsetra by Vieites 
<emphasis id="B92BEAF3FF926C1BDA8698BA6B69B0EA" box="[236,285,899,923]" italics="true" pageId="10" pageNumber="44">et al</emphasis>
. (2009) and Vences 
<emphasis id="B92BEAF3FF926C1BD86898BA6846B0EA" box="[514,562,899,923]" italics="true" pageId="10" pageNumber="44">et al</emphasis>
. (2010).
</paragraph>
</subSubSection>
<subSubSection id="C345656AFF926C1BDAAD98996E43B62B" pageId="10" pageNumber="44" type="description">
<paragraph id="8BE036E1FF926C1BDAAD98996E43B62B" blockId="10.[151,1437,151,1968]" pageId="10" pageNumber="44">
<emphasis id="B92BEAF3FF926C1BDAAD98996B4FB0B1" bold="true" box="[199,315,935,960]" pageId="10" pageNumber="44">Variation.</emphasis>
Morphological variation of nine 
<typeStatus id="54E48843FF926C1BD8DE98966954B0B1" box="[692,800,936,960]" pageId="10" pageNumber="44" type="paratype">paratypes</typeStatus>
is provided in Table 1. In general the morphology of the 
<typeStatus id="54E48843FF926C1BDAFD98F06B76B097" box="[151,258,974,998]" pageId="10" pageNumber="44" type="paratype">paratypes</typeStatus>
was similar to the 
<typeStatus id="54E48843FF926C1BDBBD98F36849B094" box="[471,573,973,997]" pageId="10" pageNumber="44" type="holotype">holotype</typeStatus>
. However, there is some variability in the separation of the lateral metatarsalia. For instance, in ZFMK 52608 they are separated, whereas in other specimens they are partly connected, and ZFMK 52612 shows connected metatarsalia on the left foot and nearly completely separated metatarsalia on the right foot. Dorsal colouration of the 
<typeStatus id="54E48843FF926C1BD8449F0068EDB727" box="[558,665,1086,1110]" pageId="10" pageNumber="44" type="paratype">paratypes</typeStatus>
is rather variable. The reddish hue characterizing the 
<typeStatus id="54E48843FF926C1BDE9E9F036F22B724" box="[1268,1366,1085,1109]" pageId="10" pageNumber="44" type="holotype">holotype</typeStatus>
is not typically observed in other specimens which are usually light brown with dark brown markings. A narrow white median line can be present on the dorsum as well as a broader band that becomes triangular at the anterior part of the dorsum, covering the entire surface of the head. Brown or grey markings and spots can be present on the venter. In 2003 we also collected one specimen of very large body size in Maroantsetra, mixed with normal-sized specimens. Because of a labeling error we cannot ascertain with full reliability the current identity of this specimen and therefore did not include it in Table 1, almost certainly it is the male ZMA 19511 of 29.0 mm SVL with largely separated outer metatarsalia (intermediate between the connected and separated state).
</paragraph>
</subSubSection>
<subSubSection id="C345656AFF926C1ADAAD9E566824B504" lastPageId="11" lastPageNumber="45" pageId="10" pageNumber="44" type="etymology">
<paragraph id="8BE036E1FF926C1BDAAD9E566E76B6BB" blockId="10.[151,1437,151,1968]" pageId="10" pageNumber="44">
<emphasis id="B92BEAF3FF926C1BDAAD9E566B3AB6F1" bold="true" box="[199,334,1384,1408]" pageId="10" pageNumber="44">Etymology.</emphasis>
The species name 
<taxonomicName id="4C5F4D62FF926C1BD8499E5668FAB6F1" box="[547,654,1384,1408]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B92BEAF3FF926C1BD8499E5668FAB6F1" box="[547,654,1384,1408]" italics="true" pageId="10" pageNumber="44">variabilis</emphasis>
</taxonomicName>
is an adjective referring to the remarkable variation observed in various characters of this new species, especially in the separation or connection of the lateral metatarsalia which otherwise is considered to be a very stable character within species of mantellids.
</paragraph>
<paragraph id="8BE036E1FF926C1BDAAD9EE66F6FB4C1" blockId="10.[151,1437,151,1968]" pageId="10" pageNumber="44">
<emphasis id="B92BEAF3FF926C1BDAAD9EE66BF2B681" bold="true" box="[199,390,1495,1520]" pageId="10" pageNumber="44">Natural history.</emphasis>
At the 
<typeStatus id="54E48843FF926C1BDBB69EE66878B681" box="[476,524,1496,1520]" pageId="10" pageNumber="44">type</typeStatus>
locality in the vicinities of Ambodivoahangy village, 
<taxonomicName id="4C5F4D62FF926C1BDE199EE76E8AB681" box="[1139,1278,1496,1520]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B92BEAF3FF926C1BDE199EE76E8AB681" box="[1139,1278,1496,1520]" italics="true" pageId="10" pageNumber="44">B. variabilis</emphasis>
</taxonomicName>
was common outside of the forest in cultivated landscape. It was not seen or heard within primary or selectively logged forest. Calling males were observed in dense secondary vegetation on the borders of partially inundated ricefields, about 
<quantity id="4CA79B04FF926C1BDAFD9D796AA8B52E" box="[151,220,1607,1631]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="44" unit="cm" value="30.0">30 cm</quantity>
to 
<quantity id="4CA79B04FF926C1BDB6B9D796B58B52E" box="[257,300,1607,1631]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="44" unit="m" value="1.0">1 m</quantity>
off the ground. The male 
<typeStatus id="54E48843FF926C1BD8399D7968C1B52E" box="[595,693,1607,1631]" pageId="10" pageNumber="44" type="holotype">holotype</typeStatus>
was discovered while calling within a dense bush at the edge of a rice field. During the day specimens were difficult to find, hiding underneath the leaves of plants. Calling was most frequent in the evening and at night, but was also occasionally heard during the day, especially after rains. Calls were often intermingled with choruses of 
<taxonomicName id="4C5F4D62FF926C1BD8149D866947B5A1" box="[638,819,1720,1744]" class="Amphibia" family="Mantellidae" genus="Aglyptodactylus" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B92BEAF3FF926C1BD8149D866947B5A1" box="[638,819,1720,1744]" italics="true" pageId="10" pageNumber="44">Aglyptodactylus</emphasis>
</taxonomicName>
sp., 
<taxonomicName id="4C5F4D62FF926C1BD9049D866EEEB5A1" box="[878,1178,1719,1744]" class="Amphibia" family="Mantellidae" genus="Guibemantis" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="species" species="kathrinae">
<emphasis id="B92BEAF3FF926C1BD9049D866989B5A1" box="[878,1021,1720,1744]" italics="true" pageId="10" pageNumber="44">Guibemantis</emphasis>
cf. 
<emphasis id="B92BEAF3FF926C1BDE459D866EEEB5A1" box="[1071,1178,1720,1744]" italics="true" pageId="10" pageNumber="44">kathrinae</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C5F4D62FF926C1BDEBD9D866B61B584" class="Amphibia" family="Ptychadenidae" genus="Ptychadena" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="species" species="mascareniensis">
<emphasis id="B92BEAF3FF926C1BDEBD9D866B61B584" italics="true" pageId="10" pageNumber="44">Ptychadena mascareniensis</emphasis>
</taxonomicName>
. At Maroantsetra it is a common species living in ditches covered by dense vegetation within the town itself. Near Voloina we heard and collected the species in flooded areas in a mosaic of rice paddies, shrubs, and remains of primary rainforest. A single clutch of eggs deposited on the surface of a leaf was discovered within the village of Ambinanitelo on 
<date id="FFE11021FF926C1BDBBE9C73681EB414" box="[468,618,1869,1893]" pageId="10" pageNumber="44" value="2010-04-01">1 April 2010</date>
. Mating in this species takes place on a leaf usually overhanging lentic water and has been observed by us in Voloina. 
<collectingCountry id="F3487671FF926C1BD8C89C4D68B5B4FA" box="[674,705,1907,1931]" name="American Samoa" pageId="10" pageNumber="44">As</collectingCountry>
in other 
<taxonomicName id="4C5F4D62FF926C1BD94F9C4D69D8B4FA" box="[805,940,1907,1931]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="10" pageNumber="44" phylum="Chordata" rank="genus">
<emphasis id="B92BEAF3FF926C1BD94F9C4D69D8B4FA" box="[805,940,1907,1931]" italics="true" pageId="10" pageNumber="44">Blommersia</emphasis>
</taxonomicName>
, the male and female sit vertically on a leaf, the male on top of the female with his legs extending over the female’s dorsum (
<bibRefCitation id="EFCE4B10FF926C1BDE709CA66F7AB4C1" author="Glaw" box="[1050,1294,1944,1968]" pageId="10" pageNumber="44" refString="Glaw, F., &amp; Vences, M. (1994) A Fieldguide to the Amphibians and Reptiles of Madagascar. Second edition. Vences &amp; Glaw Verlag, Cologne, 480 pp." type="book" year="1994">Glaw &amp; Vences 1994</bibRefCitation>
).
</paragraph>
<caption id="DF206669FF936C1ADAFD98F56934B771" httpUri="https://zenodo.org/record/207738/files/figure.png" pageId="11" pageNumber="45" targetBox="[343,1243,193,926]" targetPageId="11">
<paragraph id="8BE036E1FF936C1ADAFD98F56934B771" blockId="11.[151,1435,971,1024]" pageId="11" pageNumber="45">
<emphasis id="B92BEAF3FF936C1ADAFD98F56B61B090" bold="true" box="[151,277,971,994]" pageId="11" pageNumber="45">FIGURE 6.</emphasis>
Audiospectrogram and corresponding oscillogram of the advertisement call of 
<taxonomicName id="4C5F4D62FF936C1ADE3398F26F4AB093" box="[1113,1342,972,994]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B92BEAF3FF936C1ADE3398F26F4AB093" box="[1113,1342,972,994]" italics="true" pageId="11" pageNumber="45">Blommersia variabilis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2185788FF936C1ADF2298F26FEFB090" box="[1352,1435,972,993]" pageId="11" pageNumber="45" rank="species">
<emphasis id="B92BEAF3FF936C1ADF2298F26FEFB090" bold="true" box="[1352,1435,972,993]" pageId="11" pageNumber="45">sp. nov.</emphasis>
</taxonomicNameLabel>
from the type locality Ambodivoahangy, recorded on 3 April 2010.
</paragraph>
</caption>
<paragraph id="8BE036E1FF936C1ADAAD9F136ECCB6CB" blockId="11.[151,1437,1069,1914]" pageId="11" pageNumber="45">
<emphasis id="B92BEAF3FF936C1ADAAD9F136B15B737" bold="true" box="[199,353,1069,1094]" pageId="11" pageNumber="45">Vocalization.</emphasis>
The advertisement call of 
<taxonomicName id="4C5F4D62FF936C1AD8FA9F10696FB734" box="[656,795,1069,1093]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B92BEAF3FF936C1AD8FA9F10696FB734" box="[656,795,1069,1093]" italics="true" pageId="11" pageNumber="45">B. variabilis</emphasis>
</taxonomicName>
(
<figureCitation id="13642A64FF936C1AD9419F13691BB734" box="[811,879,1069,1093]" captionStart="FIGURE 6" captionStartId="11.[151,250,971,994]" captionTargetBox="[343,1243,193,926]" captionTargetId="figure@11.[343,1243,193,926]" captionTargetPageId="11" captionText="FIGURE 6. Audiospectrogram and corresponding oscillogram of the advertisement call of Blommersia variabilis sp. nov. from the type locality Ambodivoahangy, recorded on 3 April 2010." httpUri="https://zenodo.org/record/207738/files/figure.png" pageId="11" pageNumber="45">Fig. 6</figureCitation>
) was recorded at the 
<typeStatus id="54E48843FF936C1ADE099F106EE7B737" box="[1123,1171,1070,1094]" pageId="11" pageNumber="45">type</typeStatus>
locality (Ambodivoahangy) on 
<date id="FFE11021FF936C1ADB689F6C6BE6B71B" box="[258,402,1106,1130]" pageId="11" pageNumber="45" value="2010-04-03">3 April 2010</date>
at an estimated air temperature of 26°C. It consists of a series of short inharmonious notes, repeated at regular intervals in fast succession with a repetition rate of approximately 7.4 notes/second. Each call is composed of 3–6 notes, whereas in most cases a long call is followed by a shorter call containing fewer notes, in combination resulting in a call series with two calls. Within these call series, the interval between first and second call ranges between 500 and 720 ms. Within calls, the initial note is longer and contains more pulses than subsequent secondary notes. Numerical call parameters are as follows: call duration 344–887 ms (532 ± 219; n = 9); note duration 54–223 ms (107 ± 48; n = 23); pulses/note 3–14 (6.4 ± 3.0; n = 23); inter-note interval 21–33 ms (26.7 ± 3.2 ms; n = 19). Overall frequency is distributed in a broad band from app. 2000–6500 Hz, with a dominant frequency at around 5100 Hz. Calls were repeated at irregular intervals. Similar calls were recorded in 
<quantity id="4CA79B04FF936C1ADF299E436FE8B6E4" box="[1347,1436,1405,1429]" metricMagnitude="1" metricUnit="m" metricValue="5.05714" pageId="11" pageNumber="45" unit="in" value="1991.0">1991 in</quantity>
Maroantsetra, but due to the poor quality of the recordings we refrain from a detailed analysis.
</paragraph>
<paragraph id="8BE036E1FF936C1ADAAD9EF96824B504" blockId="11.[151,1437,1069,1914]" pageId="11" pageNumber="45">
Detailed call data for various populations of 
<emphasis id="B92BEAF3FF936C1AD8AB9EF769C6B691" box="[705,946,1480,1504]" italics="true" pageId="11" pageNumber="45">
<taxonomicName id="4C5F4D62FF936C1AD8AB9EF76958B691" box="[705,812,1480,1504]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="galani">B. galani</taxonomicName>
, 
<taxonomicName id="4C5F4D62FF936C1AD95D9EF769DAB691" box="[823,942,1480,1504]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="dejongi">B. dejongi</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="4C5F4D62FF936C1AD9819EF76E3DB691" box="[1003,1097,1481,1504]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="wittei">
<emphasis id="B92BEAF3FF936C1AD9819EF76E3DB691" box="[1003,1097,1481,1504]" italics="true" pageId="11" pageNumber="45">B. wittei</emphasis>
</taxonomicName>
are presented in Vences 
<emphasis id="B92BEAF3FF936C1ADF099EF76FE1B691" box="[1379,1429,1480,1504]" italics="true" pageId="11" pageNumber="45">et al</emphasis>
. (2010). In comparison, the call of 
<taxonomicName id="4C5F4D62FF936C1AD8709ED068D1B574" box="[538,677,1517,1541]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B92BEAF3FF936C1AD8709ED068D1B574" box="[538,677,1517,1541]" italics="true" pageId="11" pageNumber="45">B. variabilis</emphasis>
</taxonomicName>
differs from that of 
<taxonomicName id="4C5F4D62FF936C1AD9E79ED0699FB574" box="[909,1003,1518,1541]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="wittei">
<emphasis id="B92BEAF3FF936C1AD9E79ED0699FB574" box="[909,1003,1518,1541]" italics="true" pageId="11" pageNumber="45">B. wittei</emphasis>
</taxonomicName>
by longer note duration, shorter internote intervals and a lower note repetition rate. Calls of 
<taxonomicName id="4C5F4D62FF936C1AD9699D2A691FB55A" box="[771,875,1555,1579]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="galani">
<emphasis id="B92BEAF3FF936C1AD9699D2A691FB55A" box="[771,875,1555,1579]" italics="true" pageId="11" pageNumber="45">B. galani</emphasis>
</taxonomicName>
differ from those of 
<taxonomicName id="4C5F4D62FF936C1ADE3C9D2A6E94B55A" box="[1110,1248,1555,1579]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B92BEAF3FF936C1ADE3C9D2A6E94B55A" box="[1110,1248,1555,1579]" italics="true" pageId="11" pageNumber="45">B. variabilis</emphasis>
</taxonomicName>
by shorter notes and much longer inter-note intervals. In calls of 
<taxonomicName id="4C5F4D62FF936C1AD8D69D076945B521" box="[700,817,1592,1616]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="dejongi">
<emphasis id="B92BEAF3FF936C1AD8D69D076945B521" box="[700,817,1592,1616]" italics="true" pageId="11" pageNumber="45">B. dejongi</emphasis>
</taxonomicName>
, notes are barely spaced and partly fused and separate pulses within notes are unrecognizable.
</paragraph>
</subSubSection>
<subSubSection id="C345656AFF936C1ADAAD9DBC6EF1B40B" pageId="11" pageNumber="45" type="distribution">
<paragraph id="8BE036E1FF936C1ADAAD9DBC6EF1B40B" blockId="11.[151,1437,1069,1914]" pageId="11" pageNumber="45">
<emphasis id="B92BEAF3FF936C1ADAAD9DBC6B14B5EA" bold="true" box="[199,352,1666,1691]" pageId="11" pageNumber="45">Distribution.</emphasis>
<taxonomicName id="4C5F4D62FF936C1ADB0D9DBA6B86B5EA" box="[359,498,1667,1691]" class="Amphibia" family="Mantellidae" genus="Blommersia" kingdom="Animalia" order="Anura" pageId="11" pageNumber="45" phylum="Chordata" rank="species" species="variabilis">
<emphasis id="B92BEAF3FF936C1ADB0D9DBA6B86B5EA" box="[359,498,1667,1691]" italics="true" pageId="11" pageNumber="45">B. variabilis</emphasis>
</taxonomicName>
is known from (1) Maroantsetra, (2) Voloina, (3) Ambodivoahangy and (4) Ambohinantely (
<geoCoordinate id="EE6B5026FF936C1ADB649D996BCDB5CE" box="[270,441,1703,1727]" direction="south" orientation="latitude" pageId="11" pageNumber="45" precision="1" value="-15.347419">15°20'50.71'' S</geoCoordinate>
, 
<geoCoordinate id="EE6B5026FF936C1ADBAF9D996805B5CE" box="[453,625,1703,1727]" direction="east" orientation="longitude" pageId="11" pageNumber="45" precision="1" value="49.58412">49°35'02.84'' E</geoCoordinate>
). The largest straight line distance between any of these sites is approximately 
<quantity id="4CA79B04FF936C1ADA809DF36B45B594" box="[234,305,1741,1765]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="45" unit="km" value="30.0">30 km</quantity>
(Ambodivoahangy-Voloina). The altitude was not precisely measured at the collecting localities, but all except Voloina are situated below 
<quantity id="4CA79B04FF936C1AD8239DCC68F0B47B" box="[585,644,1778,1802]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="45" unit="m" value="50.0">50 m</quantity>
a.s.l. We estimate an elevation of 
<quantity id="4CA79B04FF936C1ADE7E9DCC6EE5B47B" box="[1044,1169,1778,1802]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="45" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">100-300 m</quantity>
a.s.l. for the collecting locality near Voloina. The Voloina site lies close to a medium-sized river, but due to the lack of reliable coordinates for our precise collecting locality in 1991, we cannot ascertain if our samples originate from north or south of the river. The remaining three sites are in the immediate vicinity of the Antainambalana river.
</paragraph>
</subSubSection>
</treatment>
</document>