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<document id="B161F184B0993A9B5C7B3904600AB465" ID-DOI="10.15298/rusentj.32.1.04" ID-ISSN="0132-8069" ID-Zenodo-Dep="10920271" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1712198856878" checkinUser="felipe" docAuthor="Gildenkov, M. Yu." docDate="2023" docId="03F687CCE85AFF8CFEEEE8349991FCF7" docLanguage="en" docName="RussEntomolJ.32.1.40-43.pdf" docOrigin="Russian Entomological Journal 32 (1)" docSource="http://dx.doi.org/10.15298/rusentj.32.1.04" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.1:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="1" docTitle="Carpelimus (Trogophloeus) kwauensis Gildenkov 2023, sp.n." docType="treatment" docVersion="2" lastPageNumber="42" masterDocId="FFCFFFB4E85BFF8EFFA5E90F9B7BFFE0" masterDocTitle="Two new species of the genus Carpelimus Leach, 1819 from New Guinea, related to Carpelimus (Trogophloeus) notumus Gildenkov, 2019 (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="43" masterPageNumber="40" pageNumber="41" updateTime="1712735098337" updateUser="tatiana">
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<mods:title id="AE9DB220F515E613BD12401AE6DCC90D">Two new species of the genus Carpelimus Leach, 1819 from New Guinea, related to Carpelimus (Trogophloeus) notumus Gildenkov, 2019 (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:title id="543375A2C7683F930B58B2D2FE59B2A1">Russian Entomological Journal</mods:title>
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<mods:date id="4C97BEDFE97B6CE659879E2A275479CC">2023</mods:date>
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<treatment id="03F687CCE85AFF8CFEEEE8349991FCF7" LSID="urn:lsid:plazi:treatment:03F687CCE85AFF8CFEEEE8349991FCF7" httpUri="http://treatment.plazi.org/id/03F687CCE85AFF8CFEEEE8349991FCF7" lastPageId="2" lastPageNumber="42" pageId="1" pageNumber="41">
<subSubSection id="C3456551E85AFF8FFEEEE834990DFE91" pageId="1" pageNumber="41" type="nomenclature">
<paragraph id="8BE036DAE85AFF8FFEEEE834990DFE91" blockId="1.[189,776,315,1030]" pageId="1" pageNumber="41">
<heading id="D0A881B6E85AFF8FFEEEE834990DFE91" centered="true" fontSize="10" level="2" pageId="1" pageNumber="41" reason="8">
<taxonomicName id="4C5F4D59E85AFF8FFEEEE8349948FE91" ID-CoL="8VS9V" authority="Gildenkov, 2023" authorityName="Gildenkov" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="kwauensis" status="sp. nov." subGenus="Trogophloeus">
<emphasis id="B92BEAC8E85AFF8FFEEEE8349ABCFEB3" box="[331,455,315,339]" italics="true" pageId="1" pageNumber="41">Carpelimus</emphasis>
(
<emphasis id="B92BEAC8E85AFF8FFE7EE8349914FEB3" box="[475,623,315,339]" italics="true" pageId="1" pageNumber="41">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E85AFF8FFEE8E8569AC0FE91" box="[333,443,345,369]" italics="true" pageId="1" pageNumber="41">kwauensis</emphasis>
Gildenkov
</taxonomicName>
,
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<taxonomicNameLabel id="A21857B3E85AFF8FFDE4E856990DFE91" box="[577,630,345,369]" pageId="1" pageNumber="41" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3456551E85AFF8FFE07E8799959FE6E" box="[418,546,374,398]" pageId="1" pageNumber="41" type="description">
<paragraph id="8BE036DAE85AFF8FFE07E8799959FE6E" blockId="1.[189,776,315,1030]" box="[418,546,374,398]" pageId="1" pageNumber="41">
<figureCitation id="13642A5FE85AFF8FFE07E8799965FE6E" box="[418,542,374,398]" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="1" pageNumber="41">Figs 1, 34</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3456551E85AFF8FFF47E89499A6FDC6" pageId="1" pageNumber="41" type="materials_examined">
<paragraph id="8BE036DAE85AFF8FFF47E89499A6FDC6" blockId="1.[189,776,315,1030]" pageId="1" pageNumber="41">
MATERIAL.
<materialsCitation id="3B373C87E85AFF8FFEC4E8949A73FE3E" collectingDate="1993-04-17" collectionCode="SMNS" country="Indonesia" elevation="1475" location="West Papua" pageId="1" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="holotype">
<typeStatus id="54E48878E85AFF8FFEC4E8949AC8FE4E" box="[353,435,411,430]" pageId="1" pageNumber="41" type="holotype">Holotype</typeStatus>
,
<specimenCount id="9D59FD53E85AFF8FFE67E8979AADFE4E" box="[450,470,408,430]" count="1" pageId="1" pageNumber="41" type="male"></specimenCount>
,
<collectingCountry id="F348764AE85AFF8FFE43E8949947FE4E" box="[486,572,411,430]" name="Indonesia" pageId="1" pageNumber="41">Indonesia</collectingCountry>
,
<collectingRegion id="499BF838E85AFF8FFDEEE89499CCFE4E" box="[587,695,411,430]" country="Indonesia" name="Papua Barat" pageId="1" pageNumber="41">West Papua</collectingRegion>
“IRIAN JAYA: Mokwam Kwau
<quantity id="4CA79B3FE85AFF8FFE37E8BC997BFE26" box="[402,512,435,454]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.65" metricValueMin="1.3" pageId="1" pageNumber="41" unit="m" value="1475.0" valueMax="1650.0" valueMin="1300.0">
<elevation id="0072D1E9E85AFF8FFE37E8BC997BFE26" box="[402,512,435,454]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.65" metricValueMin="1.3" pageId="1" pageNumber="41" unit="m" value="1475.0" valueMax="1650.0" valueMin="1300.0">1300-1650m</elevation>
</quantity>
<date id="FFE1101AE85AFF8FFDA2E8BC9912FE26" box="[519,617,435,454]" pageId="1" pageNumber="41" value="1993-04-17">
<collectingDate id="EFA5E9F2E85AFF8FFDA2E8BC9912FE26" box="[519,617,435,454]" pageId="1" pageNumber="41" value="1993-04-17">17.IV.1993</collectingDate>
</date>
leg. A. RIEDEL” (
<collectionCode id="ED4EAE1FE85AFF8FFF66E8C49A79FE3E" box="[195,258,459,478]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="1" pageNumber="41">SMNS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B373C87E85AFF8FFEB1E8C49878FE16" collectingDate="1993-04-17" country="Indonesia" elevation="1475" location="Mokwam Kwau" pageId="1" pageNumber="41" specimenCount="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="paratype">
<typeStatus id="54E48878E85AFF8FFEB1E8C49A10FE3E" box="[276,363,459,478]" pageId="1" pageNumber="41" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D59FD53E85AFF8FFED2E8C49AEDFE3E" box="[375,406,456,478]" count="1" pageId="1" pageNumber="41" type="male">1♂</specimenCount>
,
<collectingCountry id="F348764AE85AFF8FFE00E8C49A80FE3E" box="[421,507,459,478]" name="Indonesia" pageId="1" pageNumber="41">Indonesia</collectingCountry>
,
<collectingRegion id="499BF838E85AFF8FFDADE8C49909FE3E" box="[520,626,459,478]" country="Indonesia" name="Papua Barat" pageId="1" pageNumber="41">West Papua</collectingRegion>
“IRIAN JAYA:
<location id="8E806001E85AFF8FFF18E8EC9A3CFE16" LSID="urn:lsid:plazi:treatment:03F687CCE85AFF8CFEEEE8349991FCF7:8E806001E85AFF8FFF18E8EC9A3CFE16" box="[189,327,483,502]" country="Indonesia" name="Mokwam Kwau" pageId="1" pageNumber="41" stateProvince="Papua Barat">Mokwam Kwau</location>
<quantity id="4CA79B3FE85AFF8FFEE8E8EC9AC5FE16" box="[333,446,483,502]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.65" metricValueMin="1.3" pageId="1" pageNumber="41" unit="m" value="1475.0" valueMax="1650.0" valueMin="1300.0">
<elevation id="0072D1E9E85AFF8FFEE8E8EC9AC5FE16" box="[333,446,483,502]" metricMagnitude="3" metricUnit="m" metricValue="1.475" metricValueMax="1.65" metricValueMin="1.3" pageId="1" pageNumber="41" unit="m" value="1475.0" valueMax="1650.0" valueMin="1300.0">13001650m</elevation>
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<date id="FFE1101AE85AFF8FFE61E8EC995DFE16" box="[452,550,483,502]" pageId="1" pageNumber="41" value="1993-04-17">
<collectingDate id="EFA5E9F2E85AFF8FFE61E8EC995DFE16" box="[452,550,483,502]" pageId="1" pageNumber="41" value="1993-04-17">17.IV.1993</collectingDate>
</date>
leg. A. RIEDEL” (cMG)
</materialsCitation>
;
<materialsCitation id="3B373C87E85AFF8FFF18E8F499A2FDC6" collectionCode="MNHG" collectorName="W. G. Ullrich" country="Papua New Guinea" location="Papua New Guinea" pageId="1" pageNumber="41" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D59FD53E85AFF8FFF18E8F49BA8FDEE" box="[189,211,504,526]" count="1" pageId="1" pageNumber="41" type="female">1♀</specimenCount>
,
<collectingCountry id="F348764AE85AFF8FFF44E8F49AF0FDEE" box="[225,395,507,526]" name="Papua New Guinea" pageId="1" pageNumber="41">Papua New Guinea</collectingCountry>
“18 VI 79 | PNG EN Prov. Umg. Igipinti Korindaiop. R.” “Papua Nlle Guinee
<collectorName id="26AA530CE85AFF8FFDA1EB1C990CFDC6" box="[516,631,531,550]" pageId="1" pageNumber="41">W.G. Ullrich</collectorName>
” (
<collectionCode id="ED4EAE1FE85AFF8FFD28EB1C99AFFDC6" box="[653,724,531,550]" pageId="1" pageNumber="41">MNHG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3456551E85AFF8CFF47EB249A72FD71" lastPageId="2" lastPageNumber="42" pageId="1" pageNumber="41" type="description">
<paragraph id="8BE036DAE85AFF8FFF47EB24996AFD71" blockId="1.[189,776,315,1030]" pageId="1" pageNumber="41">
DESCRIPTION (
<typeStatus id="54E48878E85AFF8FFE2EEB249A9EFDA1" box="[395,485,555,577]" pageId="1" pageNumber="41" type="holotype">holotype</typeStatus>
). Length
<quantity id="4CA79B3FE85AFF8FFD9AEB2499F0FDA0" box="[575,651,555,576]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="1" pageNumber="41" unit="mm" value="1.9">1.9 mm</quantity>
. Colouration brown. Legs and antennal bases yellow-brown, antennal apices darker, light brown. Integument slightly shining, body with short, light-coloured setation.
</paragraph>
<paragraph id="8BE036DAE85AFF8FFF47EB9A9A0CFBE6" blockId="1.[189,776,315,1030]" pageId="1" pageNumber="41">
Head transverse, with a wide base, ratio of its length (from posterior margin of head to anterior margin of clypeus) to maximum width about 18:25. Neck constriction prominent. Eyes large, convex. Temples well-developed, round, eye diameter in dorsal view barely exceeds temple length. Head width across eyes approximately equal to its width across temples (
<figureCitation id="13642A5FE85AFF8FFEFCEA3A9AE9FCAC" box="[345,402,821,844]" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="1" pageNumber="41">Fig. 1</figureCitation>
). Head surface with delicate, fine and dense punctation. Diameter of punctures about 3.0 times as small as eye facet. Distances between punctures approximately equal to their diameter, interspaces smooth, slightly shining. Antennae rather long, antennal segments 13 elongate; segments 410 slightly elongate; segment 11 elongate, conical. Last 3 segments more massive than others and form loose club (
<figureCitation id="13642A5FE85AFF8FFE8AEAFF9A13FBE6" box="[303,360,1008,1030]" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="1" pageNumber="41">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BE036DAE85AFF8FFE93EE579900F892" blockId="1.[310,635,1880,1906]" box="[310,635,1880,1906]" pageId="1" pageNumber="41">
<emphasis id="B92BEAC8E85AFF8FFE93EE579900F892" bold="true" box="[310,635,1880,1906]" pageId="1" pageNumber="41">1 2</emphasis>
</paragraph>
<paragraph id="8BE036DAE85AFF8FFCF7E83398E7FDBC" blockId="1.[812,1399,315,1031]" pageId="1" pageNumber="41">
Pronotum maximum broad after about 2/3 its length from base, then narrowed. Lateral margin smoothly rounded (
<figureCitation id="13642A5FE85AFF8FFAEAE859984CFE67" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="1" pageNumber="41">Fig. 1</figureCitation>
). Ratio of pronotum length to its maximum width about 20:26. Surface of pronotum with delicate, fine and dense punctation, like head surface; Pronotal disc with 2 pairs of prominent, symmetrical depressions and 1 unpaired oval depression along midline near apex. Base of pronotal disc with narrow, crescent-shaped depressions, weakly separated by medial ridge. Central part of disc with rather deep, oval depressions fused across midline to single butterfly-shaped depression.
</paragraph>
<paragraph id="8BE036DAE85AFF8FFCF7EB6E9F9AFD1C" blockId="1.[812,1399,315,1031]" pageId="1" pageNumber="41">
Length of elytra related to their combined width approximately as 31:36. Scutellum with shallow, round depressions (
<figureCitation id="13642A5FE85AFF8FFC96EB99980AFD4C" box="[819,881,662,684]" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="1" pageNumber="41">Fig. 1</figureCitation>
). Elytra with delicate, fine and dense punctation. Diameter of punctures about 2.5 times as small as eye facet. Distances between punctures slightly smaller than their diameter, interspaces smooth, slightly shining.
</paragraph>
<paragraph id="8BE036DAE85AFF8FFCF7EA0E9FE9FCF7" blockId="1.[812,1399,315,1031]" box="[850,1170,769,791]" pageId="1" pageNumber="41">Abdomen delicately shagreened.</paragraph>
<paragraph id="8BE036DAE85AFF8FFCF7EA149E5FFCD1" blockId="1.[812,1399,315,1031]" box="[850,1316,795,817]" pageId="1" pageNumber="41">
Aedeagus of characteristic structure (
<figureCitation id="13642A5FE85AFF8FFB1BEA149E63FCD1" box="[1214,1304,795,817]" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="1" pageNumber="41">Figs 34</figureCitation>
).
</paragraph>
<paragraph id="8BE036DAE85AFF8FFCF7EA399F7AFC61" blockId="1.[812,1399,315,1031]" pageId="1" pageNumber="41">
<emphasis id="B92BEAC8E85AFF8FFCF7EA3998E6FCAB" bold="true" box="[850,925,822,843]" pageId="1" pageNumber="41">Female</emphasis>
. Sexual dimorphism absent, female morphologically similar to male. Spermatheca failed to isolate and its structure is unknown.
</paragraph>
<paragraph id="8BE036DAE85AFF8CFCF7EA899A72FD71" blockId="1.[812,1399,315,1031]" lastBlockId="2.[189,775,315,791]" lastPageId="2" lastPageNumber="42" pageId="1" pageNumber="41">
COMPARATIVE REMARKS. The new species is very similar and closely related to
<taxonomicName id="4C5F4D59E85AFF8FFBC1EAAE9F50FC30" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="notumus" subGenus="Trogophloeus">
<emphasis id="B92BEAC8E85AFF8FFBC1EAAE9FAFFC56" box="[1124,1236,929,950]" italics="true" pageId="1" pageNumber="41">Carpelimus</emphasis>
(
<emphasis id="B92BEAC8E85AFF8FFB4DEAAE9E15FC56" box="[1256,1390,929,950]" italics="true" pageId="1" pageNumber="41">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E85AFF8FFC89EAB39805FC31" box="[812,894,956,977]" italics="true" pageId="1" pageNumber="41">notumus</emphasis>
Gildenkov, 2019
</taxonomicName>
from the
<collectingCountry id="F348764AE85AFF8FFB34EAB49F85FC31" box="[1169,1278,955,977]" name="Philippines" pageId="1" pageNumber="41">Philippines</collectingCountry>
and
<taxonomicName id="4C5F4D59E85AFF8FFA97EAB39F86FC0B" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="vilisus" subGenus="Trogophloeus">
<emphasis id="B92BEAC8E85AFF8FFA97EAB3981BFC0B" italics="true" pageId="1" pageNumber="41">Carpelimus</emphasis>
(
<emphasis id="B92BEAC8E85AFF8FFCD1EAD99881FC0B" box="[884,1018,982,1003]" italics="true" pageId="1" pageNumber="41">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E85AFF8FFBA8EAD99F30FC0B" box="[1037,1099,982,1003]" italics="true" pageId="1" pageNumber="41">vilisus</emphasis>
Gildenkov, 2019
</taxonomicName>
from
<collectingCountry id="F348764AE85AFF8FFAE7EAD99822FBE7" name="Thailand" pageId="1" pageNumber="41">Thailand</collectingCountry>
. It differs from them by slightly less developed eyes, slightly finer punctation of elytra and slightly longer antennae. Reliably differs only by the structure of aedeagus [
<figureCitation id="13642A5FE859FF8CFD7FE8599B98FE66" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="2" pageNumber="42">Figs 34</figureCitation>
;
<bibRefCitation id="EFCE4B2BE859FF8CFF57E87E9AD3FE66" author="Gildenkov M. Yu." box="[242,424,369,391]" pageId="2" pageNumber="42" refId="ref2388" refString="Gildenkov M. Yu. 2019 a. Five new species of the Genus Carpelimus Leach, 1819 from Thailand and the Philippines (Coleoptera: Staphylinidae: Oxytelinae) // Russian Entomological Journal. Vol. 28. No. 1. P. 30 - 35. DOI: 10.15298 / rusentj. 28.1.05." type="book" year="2019">Gildenkov, 2019a</bibRefCitation>
:
<figureCitation id="13642A5FE859FF8CFE13E87E996AFE66" box="[438,529,368,391]" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="2" pageNumber="42">Figs 56</figureCitation>
, 89]. The new species differs from
<taxonomicName id="4C5F4D59E859FF8CFE9EE8849B96FE5B" authority="Gildenkov, 2019" authorityName="Gildenkov" authorityYear="2019" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="plenus" subGenus="Trogophloeus">
<emphasis id="B92BEAC8E859FF8CFE9EE8849AD0FE40" box="[315,427,395,416]" italics="true" pageId="2" pageNumber="42">Carpelimus</emphasis>
(
<emphasis id="B92BEAC8E859FF8CFE19E8849939FE40" box="[444,578,395,416]" italics="true" pageId="2" pageNumber="42">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E859FF8CFDF6E88499E9FE40" box="[595,658,395,416]" italics="true" pageId="2" pageNumber="42">plenus</emphasis>
Gildenkov, 2019
</taxonomicName>
from the
<collectingCountry id="F348764AE859FF8CFEFDE8A99ABEFE5C" box="[344,453,422,444]" name="Philippines" pageId="2" pageNumber="42">Philippines</collectingCountry>
by smaller body size and finer punctation; reliably differs by the structure of aedeagus [
<figureCitation id="13642A5FE859FF8CFD7FE8CE9B98FE10" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="2" pageNumber="42">Figs 34</figureCitation>
;
<bibRefCitation id="EFCE4B2BE859FF8CFF51E8D49AD0FE10" author="Gildenkov M. Yu." box="[244,427,475,497]" pageId="2" pageNumber="42" refId="ref2388" refString="Gildenkov M. Yu. 2019 a. Five new species of the Genus Carpelimus Leach, 1819 from Thailand and the Philippines (Coleoptera: Staphylinidae: Oxytelinae) // Russian Entomological Journal. Vol. 28. No. 1. P. 30 - 35. DOI: 10.15298 / rusentj. 28.1.05." type="book" year="2019">Gildenkov, 2019a</bibRefCitation>
: Figs 910]. It well differs from
<taxonomicName id="4C5F4D59E859FF8CFF18E8F999A0FDEB" authority=", Gildenkov, 2020" authorityName=", Gildenkov" authorityYear="2020" box="[189,731,502,524]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="ibelensis" subGenus="Trogophloeus">
<emphasis id="B92BEAC8E859FF8CFF18E8F99A56FDEB" box="[189,301,502,523]" italics="true" pageId="2" pageNumber="42">Carpelimus</emphasis>
(
<emphasis id="B92BEAC8E859FF8CFE98E8F99AB8FDEB" box="[317,451,502,523]" italics="true" pageId="2" pageNumber="42">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E859FF8CFE71E8F99953FDEB" box="[468,552,502,523]" italics="true" pageId="2" pageNumber="42">ibelensis</emphasis>
, Gildenkov, 2020
</taxonomicName>
and
<taxonomicName id="4C5F4D59E859FF8CFF18EB1E99A6FDC6" authority="Gildenkov, 2020" authorityName="Gildenkov" authorityYear="2020" box="[189,733,529,551]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="irianensis" subGenus="Trogophloeus">
<emphasis id="B92BEAC8E859FF8CFF18EB1E9A56FDC6" box="[189,301,529,550]" italics="true" pageId="2" pageNumber="42">Carpelimus</emphasis>
(
<emphasis id="B92BEAC8E859FF8CFE98EB1E9AB8FDC6" box="[317,451,529,550]" italics="true" pageId="2" pageNumber="42">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E859FF8CFE76EB1E9949FDC6" box="[467,562,529,550]" italics="true" pageId="2" pageNumber="42">irianensis</emphasis>
Gildenkov, 2020
</taxonomicName>
living in New
<collectingCountry id="F348764AE859FF8CFE97EB249A03FDA1" box="[306,376,555,577]" name="Guinea-Bissau" pageId="2" pageNumber="42">Guinea</collectingCountry>
by lighter coloration, significantly more developed depressions on pronotal disc, reliably differs by the structure of aedeagus [
<figureCitation id="13642A5FE859FF8CFE18EB6E996DFD96" box="[445,534,609,631]" captionStart="Figs 16" captionStartId="1.[226,263,1967,1986]" captionTargetBox="[817,1374,1067,1951]" captionTargetId="figure-432@1.[812,1375,1039,1956]" captionTargetPageId="1" captionText="Figs 16. Carpelimus spp., holotypes, males: 1, 34 — C. (Trogophloeus) kwauensis, sp.n.; 2, 56 — C. (Trogophloeus) nabireensis, sp.n.; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm. Рис. 16. Carpelimus spp., голотипы, самцы,: 1, 34 — Carpelimus (Trogophloeus) kwauensis, sp.n.; 2, 56 — Carpelimus (Trogophloeus) nabireensis, sp.n.; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм." figureDoi="http://doi.org/10.5281/zenodo.10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="2" pageNumber="42">Figs 34</figureCitation>
;
<bibRefCitation id="EFCE4B2BE859FF8CFD84EB6E99A8FD96" author="Gildenkov M. Yu." box="[545,723,609,631]" pageId="2" pageNumber="42" refId="ref2770" refString="Gildenkov M. Yu. 2020 a. Eight new species of the genus Carpelimus Leach, 1819 from Indonesia (Coleoptera: Staphylinidae: Oxytelinae) // Russian Entomological Journal. Vol. 29. No. 1. P. 53 - 60. DOI: 10.15298 / rusentj. 29.1.07." type="book" year="2020">Gildenkov, 2020a</bibRefCitation>
: Figs 1013].
</paragraph>
</subSubSection>
<caption id="DF206652E85AFF8FFF47EEA09E7BF7EF" ID-DOI="http://doi.org/10.5281/zenodo.10920273" ID-Zenodo-Dep="10920273" httpUri="https://zenodo.org/record/10920273/files/figure.png" pageId="1" pageNumber="41" startId="1.[226,263,1967,1986]" targetBox="[817,1374,1067,1951]" targetPageId="1" targetType="figure">
<paragraph id="8BE036DAE85AFF8FFF47EEA09FFAF83B" blockId="1.[188,1398,1967,2063]" pageId="1" pageNumber="41">
Figs 16.
<taxonomicName id="4C5F4D59E85AFF8FFE9CEEA09AB1F823" box="[313,458,1967,1987]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92BEAC8E85AFF8FFE9CEEA09AE6F822" box="[313,413,1967,1986]" italics="true" pageId="1" pageNumber="41">Carpelimus</emphasis>
spp.
</taxonomicName>
, holotypes, males: 1, 34 —
<taxonomicName id="4C5F4D59E85AFF8FFD6DEEBF98BDF822" authority="Gildenkov, 2023" authorityName="Gildenkov" authorityYear="2023" box="[712,966,1967,1987]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="kwauensis" status="sp.n." subGenus="Trogophloeus">
<emphasis id="B92BEAC8E85AFF8FFD6DEEBF99ADF823" box="[712,726,1968,1987]" italics="true" pageId="1" pageNumber="41">C</emphasis>
. (
<emphasis id="B92BEAC8E85AFF8FFD4CEEA0981BF822" box="[745,864,1967,1986]" italics="true" pageId="1" pageNumber="41">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E85AFF8FFCCBEEA098BDF822" box="[878,966,1967,1986]" italics="true" pageId="1" pageNumber="41">kwauensis</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAC8E85AFF8FFC77EEBF9886F823" bold="true" box="[978,1021,1968,1987]" pageId="1" pageNumber="41">
<taxonomicNameLabel id="A21857B3E85AFF8FFC77EEBF9886F823" box="[978,1021,1968,1987]" pageId="1" pageNumber="41" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 2, 56 —
<taxonomicName id="4C5F4D59E85AFF8FFBC2EEBF9E0BF822" authority="Gildenkov, 2023" authorityName="Gildenkov" authorityYear="2023" box="[1127,1392,1967,1987]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="nabireensis" status="sp.n." subGenus="Trogophloeus">
<emphasis id="B92BEAC8E85AFF8FFBC2EEBF9F0EF823" box="[1127,1141,1968,1987]" italics="true" pageId="1" pageNumber="41">C</emphasis>
. (
<emphasis id="B92BEAC8E85AFF8FFB2CEEA09E7BF822" box="[1161,1280,1967,1986]" italics="true" pageId="1" pageNumber="41">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E85AFF8FFAA8EEA09E0BF822" box="[1293,1392,1967,1986]" italics="true" pageId="1" pageNumber="41">nabireensis</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAC8E85AFF8FFF19EEC79B9CF83B" bold="true" box="[188,231,1992,2011]" pageId="1" pageNumber="41">
<taxonomicNameLabel id="A21857B3E85AFF8FFF19EEC79B9CF83B" box="[188,231,1992,2011]" pageId="1" pageNumber="41" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 12 — dorsal view; 3, 5 — aedeagus, ventral view; 4, 6 — aedeagus, lateral view. Scale bar: 0.25 mm.
</paragraph>
<paragraph id="8BE036DAE85AFF8FFF47EEEB9E7BF7EF" blockId="1.[188,1398,1967,2063]" pageId="1" pageNumber="41">
Рис. 16.
<taxonomicName id="4C5F4D59E85AFF8FFEE6EEEC9AA3F817" box="[323,472,2019,2039]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B92BEAC8E85AFF8FFEE6EEEC9ADCF816" box="[323,423,2019,2038]" italics="true" pageId="1" pageNumber="41">Carpelimus</emphasis>
spp.
</taxonomicName>
, голотипы, самцы,: 1, 34 —
<taxonomicName id="4C5F4D59E85AFF8FFCAEEEEC9F19F816" authority="Gildenkov, 2023" authorityName="Gildenkov" authorityYear="2023" box="[779,1122,2019,2038]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="kwauensis" status="sp. nov." subGenus="Trogophloeus">
<emphasis id="B92BEAC8E85AFF8FFCAEEEEC9814F816" box="[779,879,2019,2038]" italics="true" pageId="1" pageNumber="41">Carpelimus</emphasis>
(
<emphasis id="B92BEAC8E85AFF8FFC24EEEC9883F816" box="[897,1016,2019,2038]" italics="true" pageId="1" pageNumber="41">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E85AFF8FFBAFEEEC9F19F816" box="[1034,1122,2019,2038]" italics="true" pageId="1" pageNumber="41">kwauensis</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAC8E85AFF8FFBD7EEEC9FE6F816" bold="true" box="[1138,1181,2019,2038]" pageId="1" pageNumber="41">
<taxonomicNameLabel id="A21857B3E85AFF8FFBD7EEEC9FE6F816" box="[1138,1181,2019,2038]" pageId="1" pageNumber="41" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 2, 56 —
<taxonomicName id="4C5F4D59E85AFF8FFAB7EEEC9AD7F7EE" authority="Gildenkov, 2023" authorityName="Gildenkov" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="nabireensis" status="sp. nov." subGenus="Trogophloeus">
<emphasis id="B92BEAC8E85AFF8FFAB7EEEC9E0DF816" box="[1298,1398,2019,2038]" italics="true" pageId="1" pageNumber="41">Carpelimus</emphasis>
(
<emphasis id="B92BEAC8E85AFF8FFF61EEF49A40F7EE" box="[196,315,2043,2062]" italics="true" pageId="1" pageNumber="41">Trogophloeus</emphasis>
)
<emphasis id="B92BEAC8E85AFF8FFEECEEF49AD7F7EE" box="[329,428,2043,2062]" italics="true" pageId="1" pageNumber="41">nabireensis</emphasis>
</taxonomicName>
,
<emphasis id="B92BEAC8E85AFF8FFE1CEEF49A98F7EE" bold="true" box="[441,483,2043,2062]" pageId="1" pageNumber="41">
<taxonomicNameLabel id="A21857B3E85AFF8FFE1CEEF49A98F7EE" box="[441,483,2043,2062]" pageId="1" pageNumber="41" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 12 — вид сверху; 3, 5 — Эдеагус, сниЗу; 4, 6 — Эдеагус, сбоку. МасШтаб: 0,25 мм.
</paragraph>
</caption>
<subSubSection id="C3456551E859FF8CFF47EB999A71FD27" pageId="2" pageNumber="42" type="distribution">
<paragraph id="8BE036DAE859FF8CFF47EB999A71FD27" blockId="2.[189,775,315,791]" pageId="2" pageNumber="42">
DISTRIBUTION.
<collectingCountry id="F348764AE859FF8CFE3FEB999A83FD4C" box="[410,504,662,684]" name="Indonesia" pageId="2" pageNumber="42">Indonesia</collectingCountry>
(
<collectingRegion id="499BF838E859FF8CFDA2EB9999F9FD4B" box="[519,642,662,683]" country="Indonesia" name="Papua Barat" pageId="2" pageNumber="42">West Papua</collectingRegion>
),
<collectingCountry id="F348764AE859FF8CFD30EB999A7DFD27" name="Papua New Guinea" pageId="2" pageNumber="42">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3456551E859FF8CFF47EBC49991FCF7" pageId="2" pageNumber="42" type="etymology">
<paragraph id="8BE036DAE859FF8CFF47EBC49991FCF7" blockId="2.[189,775,315,791]" pageId="2" pageNumber="42">
ETYMOLOGY. The species name is given according to the name of the village (Kwau Village) in West Papuas, in Manokwari Regencies where the
<typeStatus id="54E48878E859FF8CFDA6EA0E9923FCF7" box="[515,600,769,791]" pageId="2" pageNumber="42" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
</treatment>
</document>