146 lines
18 KiB
XML
146 lines
18 KiB
XML
<document id="DDE4B46F94958340C7AF0BB00FF798D4" ID-DOI="10.5281/zenodo.5170203" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170203" ID-ZooBank="AB1A57F0-7CB4-4830-920B-DF219740A596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628281492031" checkinUser="felipe" docAuthor="Blahnik, Roger J. & Holzenthal, Ralph W." docDate="2017" docId="03F687A7FFE6F818FF01BB46426BFD0F" docLanguage="en" docName="InsectaMundi.0602.1-251.pdf" docOrigin="Insecta Mundi 2017 (602)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Mortoniella (Mortoniella) langleyae Blahnik & Holzenthal 2017, new species" docType="treatment" docVersion="4" lastPageNumber="30" masterDocId="FFCFFFDFFFF8F807FFC1B865400BFFB4" masterDocTitle="Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *" masterLastPageNumber="251" masterPageNumber="1" pageNumber="29" updateTime="1699039969677" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="3FC2A60E62F5263352F1BF3E4AFD7260">
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<mods:title id="74FAD6BAD20FF783FAC6054CD66862A5">Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *</mods:title>
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<mods:name id="CEA0C114E0CC6F084DA429729E34EF3E" type="personal">
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<mods:namePart id="2A07C1281F826A34658C666729A62B7D">Blahnik, Roger J.</mods:namePart>
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<mods:roleTerm id="0A94192A394F881B36DCA871F105E986">Author</mods:roleTerm>
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<mods:namePart id="ADFC2FB55C5917466C815438B4A74D97">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="E18B96E6E7BD363F484F286141218452">text</mods:typeOfResource>
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<mods:title id="5E52983F5807AD39EB31C1B3988EE98F">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="CEB532E234C2572C4F3482C978EDC8CA">
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<mods:date id="14F685823661BFDDE3AED2ED1AFF4E86">2017</mods:date>
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<mods:number id="293C1DAEC88DDAF2403347CE81ACB244">2017-12-29</mods:number>
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<mods:detail id="937BD55B761E1C509476F2BCF27A5F3D" type="volume">
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<mods:number id="1EF8401A1C8B33A892FEAA99EF603D86">2017</mods:number>
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<mods:detail id="17682CD59AF40706BEBF0E3199E626A9" type="issue">
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<mods:number id="64E8B67E7BD22F0E0F0DF220547F2F4A">602</mods:number>
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<mods:classification id="13CDBC393DF908343F220482DFAA205C">journal article</mods:classification>
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<mods:identifier id="B3641B9AD1EB757B82CBF5322B990601" type="DOI">10.5281/zenodo.5170203</mods:identifier>
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<mods:identifier id="238C09934E4891EC724D5D6FCFF7CC68" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="356108004CEC010FF64D56B45AC02D5B" type="Zenodo-Dep">5170203</mods:identifier>
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<mods:identifier id="15F66400F4C21667B79839B512EE81CA" type="ZooBank">AB1A57F0-7CB4-4830-920B-DF219740A596</mods:identifier>
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<treatment id="03F687A7FFE6F818FF01BB46426BFD0F" ID-DOI="http://doi.org/10.5281/zenodo.5186282" ID-Zenodo-Dep="5186282" LSID="urn:lsid:plazi:treatment:03F687A7FFE6F818FF01BB46426BFD0F" httpUri="http://treatment.plazi.org/id/03F687A7FFE6F818FF01BB46426BFD0F" lastPageId="31" lastPageNumber="30" pageId="30" pageNumber="29">
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<subSubSection id="C345653AFFE6F819FF01BB46436DFC8F" box="[192,870,803,827]" pageId="30" pageNumber="29" type="nomenclature">
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<paragraph id="8BE036B1FFE6F819FF01BB46436DFC8F" blockId="30.[192,870,803,859]" box="[192,870,803,827]" pageId="30" pageNumber="29">
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<heading id="D0A881DDFFE6F819FF01BB46436DFC8F" bold="true" box="[192,870,803,827]" fontSize="10" level="2" pageId="30" pageNumber="29" reason="3">
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<emphasis id="B92BEAA3FFE6F819FF01BB46436DFC8F" bold="true" box="[192,870,803,827]" pageId="30" pageNumber="29">
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<taxonomicName id="4C5F4D32FFE6F819FF01BB4642BBFC8F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[192,688,803,827]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="langleyae" status="sp. nov." subGenus="Mortoniella">
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<emphasis id="B92BEAA3FFE6F819FF01BB464168FC8F" bold="true" box="[192,355,803,827]" italics="true" pageId="30" pageNumber="29">Mortoniella</emphasis>
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(
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<emphasis id="B92BEAA3FFE6F819FEB7BB464212FC8F" bold="true" box="[374,537,803,827]" italics="true" pageId="30" pageNumber="29">Mortoniella</emphasis>
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)
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<emphasis id="B92BEAA3FFE6F819FDEABB4642BBFC8F" bold="true" box="[555,688,803,827]" italics="true" pageId="30" pageNumber="29">langleyae</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFE6F819FD00BB46436DFC8F" box="[705,870,803,827]" pageId="30" pageNumber="29" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C345653AFFE6F818FF01BB26447DFDAF" lastPageId="31" lastPageNumber="30" pageId="30" pageNumber="29" type="description">
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<paragraph id="8BE036B1FFE6F819FF01BB26411DFCEF" blockId="30.[192,870,803,859]" box="[192,278,835,859]" pageId="30" pageNumber="29">
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<figureCitation id="13642A34FFE6F819FF01BB26411DFCEF" box="[192,278,835,859]" captionStart="Figure 14" captionStartId="155.[192,279,1836,1858]" captionTargetBox="[345,1281,217,1772]" captionTargetId="figure-54@155.[345,1282,217,1785]" captionTargetPageId="155" captionText="Figure 14. Mortoniella (Mortoniella) langleyae, new species, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5170235" httpUri="https://zenodo.org/record/5170235/files/figure.png" pageId="30" pageNumber="29">Fig. 14</figureCitation>
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</paragraph>
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<paragraph id="8BE036B1FFE6F819FF31BBE64135FB4F" blockId="30.[192,1441,899,1275]" pageId="30" pageNumber="29">
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This is a very distinctive species, probably most closely related to
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<taxonomicName id="4C5F4D32FFE6F819FBB0BBE14597FC2F" authority="Sykora" authorityName="Sykora" authorityYear="1999" box="[1137,1436,899,923]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="denticulata">
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<emphasis id="B92BEAA3FFE6F819FBB0BBE14539FC2F" box="[1137,1330,899,923]" italics="true" pageId="30" pageNumber="29">M. denticulata</emphasis>
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Sykora
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</taxonomicName>
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, though superficially very different. Both species have a dorsal phallic spine with small apical spines (also present in
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<taxonomicName id="4C5F4D32FFE6F819FDC6BBA1429AFC6F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[519,657,963,987]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="silacea" status="sp. nov.">
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<emphasis id="B92BEAA3FFE6F819FDC6BBA1429AFC6F" box="[519,657,963,987]" italics="true" pageId="30" pageNumber="29">M. silacea</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFE6F819FD66BBA142F9FC6F" box="[679,754,963,987]" pageId="30" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
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) and an endophallic membrane with conspicuous membranous lateral lobes with minute spines. Both species also lack paramere appendages. As compared to either
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<taxonomicName id="4C5F4D32FFE6F819FE07BC61428DFBAF" authorityName="Sykora" authorityYear="1999" box="[454,646,1027,1051]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="denticulata">
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<emphasis id="B92BEAA3FFE6F819FE07BC61428DFBAF" box="[454,646,1027,1051]" italics="true" pageId="30" pageNumber="29">M. denticulata</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4C5F4D32FFE6F819FD79BC614335FBAF" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[696,830,1027,1051]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="silacea">
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<emphasis id="B92BEAA3FFE6F819FD79BC614335FBAF" box="[696,830,1027,1051]" italics="true" pageId="30" pageNumber="29">M. silacea</emphasis>
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</taxonomicName>
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, the apex of the dorsal phallic spine is much narrower, as viewed laterally. Especially distinctive aspects of
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<taxonomicName id="4C5F4D32FFE6F819FBD2BC4144B0FB8F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[1043,1211,1059,1083]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="langleyae">
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<emphasis id="B92BEAA3FFE6F819FBD2BC4144B0FB8F" box="[1043,1211,1059,1083]" italics="true" pageId="30" pageNumber="29">M. langleyae</emphasis>
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</taxonomicName>
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include the very elongate and apically curved tergum X, with a very deep mesal invagination and very elongate, pencil-like, ventrolateral lobes. Additionally, the tergum has very conspicuously developed, enlarged and projecting, ventromesal lobes. Like
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<taxonomicName id="4C5F4D32FFE6F819FCFDBCE143F3FB2F" authorityName="Sykora" authorityYear="1999" box="[828,1016,1155,1179]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="30" pageNumber="29" phylum="Tracheophyta" rank="species" species="denticulata">
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<emphasis id="B92BEAA3FFE6F819FCFDBCE143F3FB2F" box="[828,1016,1155,1179]" italics="true" pageId="30" pageNumber="29">M. denticulata</emphasis>
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</taxonomicName>
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, the tergum lacks either obvious apicoventral sclerotization or a distinctly inflated and demarcated basal portion. As compared to other species in the bilineata group, the anteroventral margin of segment IX is only weakly produced.
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</paragraph>
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<paragraph id="8BE036B1FFE6F819FF31BD464123FA2F" blockId="30.[192,1440,1315,1916]" pageId="30" pageNumber="29">
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<emphasis id="B92BEAA3FFE6F819FF31BD464133FA8F" box="[240,312,1315,1339]" italics="true" pageId="30" pageNumber="29">Adult</emphasis>
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—Length of forewing: male
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<quantity id="4CA79B54FFE6F819FD5BBD4642F7FA8F" box="[666,764,1315,1339]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="30" pageNumber="29" unit="mm" value="3.5">3.5 mm</quantity>
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. Forewing with forks I, II, and III present, hind wing with forks II, III, and V. Spur formula 0:4:4. Color (in alcohol) yellowish-brown; setae of forewing (apparently) golden or golden-brown. Wing membranes not obviously infuscated. Wing bars not evident.
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</paragraph>
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<paragraph id="8BE036B1FFE6F818FF31BDC6447DFDAF" blockId="30.[192,1440,1315,1916]" lastBlockId="31.[192,1440,195,699]" lastPageId="31" lastPageNumber="30" pageId="30" pageNumber="29">
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<emphasis id="B92BEAA3FFE6F819FF31BDC641A1FA0F" box="[240,426,1443,1467]" italics="true" pageId="30" pageNumber="29">Male genitalia</emphasis>
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—Ventral process of segment VI posteriorly projecting, relatively short, narrow basally, length about 3 times width at base. Tergum VIII relatively narrow, subtending ventral margin of segment IX; membranous connection to tergum IX elongate. Segment IX with anterolateral margin rounded and weakly produced in ventral half, posterolateral margin with nearly rectilinearly angular projection in dorsal half; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X very elongate and curved apically, base of segment not inflated, lateral margins slightly constricted at base, subparallel laterally, apex of tergum with very deep V-shaped mesal incision, apex sclerotized, but not forming evident “cap,” tergum ventrolaterally with very elongate narrow, pencil-like, lateral lobes, each with elongate apical seta, lobe narrowly separated from and subparallel to apicolateral lobes; tergum ventromesally with very prominent, paired, sclerotized and projecting, ventromesal lobes in basal half, each with short setae. Inferior appendages with very short (apparently vestigial) dorsolateral lobes and short paired, curved, apicoventral lobes, each with a distinct apicoventral spine-like projection. Mesal pockets of inferior appendage with elongate, posteriorly-directed, spine-like, apicoventral projections. Fused basal segments of parameres articulating near middle of stalk-like basal part of dorsal phallic spine, paramere appendages absent. Phallobase with relatively small rounded, laterally compressed, dorsomesal apodeme. Dorsal phallic spine, as viewed laterally, with dorsal margin nearly straight, obtusely upturned in about apical 1/3, base of spine narrow, stalk-like and sinuously curved, distinctly widened on ventral margin in basal ½, forming acute ventral projection; upturned apex of spine very distinctly narrowed, with scattered small lateral spines, apex rounded; spine, as viewed dorsally, narrow basally and apically, slightly widened in middle, apex rounded. Phallicata with sclerotized basodorsal projection, articulating with angular ventral projection of dorsal phallic spine; phallicata ventrally with indistinct, paired, lightly sclerotized lobes, extending beyond lateral lobes of endophallic membrane. Endophallic membrane with distinct projecting membranous lateral lobes at midlength, each with minute spines; phallotremal spines absent.
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</paragraph>
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</subSubSection>
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<subSubSection id="C345653AFFE7F818FF31BA46426BFD0F" pageId="31" pageNumber="30" type="materials_examined">
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<paragraph id="8BE036B1FFE7F818FF31BA4642E4FDEF" blockId="31.[192,1440,195,699]" pageId="31" pageNumber="30">
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<materialsCitation id="3B373CECFFE7F818FF31BA4642E4FDEF" collectingDate="1976-06-12" collectionCode="A, NMNH" country="Ecuador" location="Langley" municipality="Cumbaratza" pageId="31" pageNumber="30" specimenCode="UMSP000097033" specimenCount="1" stateProvince="Zamora-Chinchipe" typeStatus="holotype">
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<emphasis id="B92BEAA3FFE7F818FF31BA4641C9FD8F" bold="true" box="[240,450,547,571]" italics="true" pageId="31" pageNumber="30">
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<typeStatus id="54E48813FFE7F818FF31BA464164FD8F" box="[240,367,547,571]" pageId="31" pageNumber="30" type="holotype">Holotype</typeStatus>
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male
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</emphasis>
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(alcohol)—
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<emphasis id="B92BEAA3FFE7F818FD9CBA464421FD8F" bold="true" box="[605,1066,547,571]" pageId="31" pageNumber="30">
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<collectingCountry id="F3487621FFE7F818FD9CBA4642F7FD8F" box="[605,764,547,571]" name="Ecuador" pageId="31" pageNumber="30">ECUADOR</collectingCountry>
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:
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<collectingRegion id="499BF853FFE7F818FCCEBA464428FD8F" box="[783,1059,547,571]" country="Ecuador" name="Zamora-Chinchipe" pageId="31" pageNumber="30">Zamora-Chinchipe</collectingRegion>
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:
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</emphasis>
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<collectingMunicipality id="6B84ACCBFFE7F818FBF6BA4644D6FD8F" box="[1079,1245,547,571]" pageId="31" pageNumber="30">Cumbaratza</collectingMunicipality>
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,
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<date id="FFE11071FFE7F818FB2EBA464571FD8F" box="[1263,1402,547,571]" pageId="31" pageNumber="30" value="1976-06-12">
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<collectingDate id="EFA5E999FFE7F818FB2EBA464571FD8F" box="[1263,1402,547,571]" pageId="31" pageNumber="30" value="1976-06-12">12.vi.1976</collectingDate>
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</date>
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,
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<collectionCode id="ED4EAE74FFE7F818FA4CBA4645ABFD8F" box="[1421,1440,547,571]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="31" pageNumber="30" type="Herbarium">A</collectionCode>
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<location id="8E80606AFFE7F818FF01BA26412CFDEF" LSID="urn:lsid:plazi:treatment:03F687A7FFE6F818FF01BB46426BFD0F:8E80606AFFE7F818FF01BA26412CFDEF" box="[192,295,579,603]" country="Ecuador" municipality="Cumbaratza" name="Langley" pageId="31" pageNumber="30" stateProvince="Zamora-Chinchipe">Langley</location>
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et al. (
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<collectionCode id="ED4EAE74FFE7F818FE42BA2641E8FDEF" box="[387,483,579,603]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="30" type="Museum">NMNH</collectionCode>
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) (
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<specimenCode id="DBF99ECAFFE7F818FE3CBA2642E9FDEF" box="[509,738,579,603]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="31" pageNumber="30" type="University or college">UMSP000097033</specimenCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BE036B1FFE7F818FF31BA06426BFD0F" blockId="31.[192,1440,195,699]" pageId="31" pageNumber="30">
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<emphasis id="B92BEAA3FFE7F818FF31BA06417EFDCF" box="[240,373,611,635]" italics="true" pageId="31" pageNumber="30">Etymology</emphasis>
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—This species is named in honor of Andrea Langley, who collected the
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<typeStatus id="54E48813FFE7F818FB2DBA014528FDC8" box="[1260,1315,612,636]" pageId="31" pageNumber="30">type</typeStatus>
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specimen as part of a Peace Corps project in the 1970’s, in recognition of the value of this endeavor to the study of Neotropical caddisflies.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |