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<document id="8FC40E61FC5B169E7AE4E481A69A5625" ID-DOI="10.5281/zenodo.5170203" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170203" ID-ZooBank="AB1A57F0-7CB4-4830-920B-DF219740A596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628281492031" checkinUser="felipe" docAuthor="Blahnik, Roger J. &amp; Holzenthal, Ralph W." docDate="2017" docId="03F687A7FF9DF861FF01BBE6412BFECF" docLanguage="en" docName="InsectaMundi.0602.1-251.pdf" docOrigin="Insecta Mundi 2017 (602)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Mortoniella (Nanotrichia) quadridactyla Blahnik &amp; Holzenthal 2017, new species" docType="treatment" docVersion="4" lastPageNumber="101" masterDocId="FFCFFFDFFFF8F807FFC1B865400BFFB4" masterDocTitle="Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *" masterLastPageNumber="251" masterPageNumber="1" pageNumber="100" updateTime="1699039969677" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="F117BE71CBE3C9E1AA1A57E4AFD258BA">Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *</mods:title>
</mods:titleInfo>
<mods:name id="5B7047AB729370710C2A8B308CAACE53" type="personal">
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<mods:namePart id="E8C1D5196F6E8BCD11CBBBB929141AFB">Blahnik, Roger J.</mods:namePart>
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<mods:namePart id="C7180A9C8600730537FFBB12AC829A27">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="6142D020016E0243B892F682F7CEC660">text</mods:typeOfResource>
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<mods:title id="A6536E398B6B35CC479D7EB344D97462">Insecta Mundi</mods:title>
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<mods:part id="65FB139A2665E49A97BDE4D4666D05EB">
<mods:date id="76A7AD903507CD980B585B93E51FEBEE">2017</mods:date>
<mods:detail id="F0E6112401E40E6E695181D91758D3A1" type="pubDate">
<mods:number id="C68E0806E3249EAB182AC17377C37B7D">2017-12-29</mods:number>
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<mods:detail id="5A1A1BF0285E295E81CA30383CF9C4E1" type="volume">
<mods:number id="C1CFB13BEDD28BD46E1BC9A8C770BA56">2017</mods:number>
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<mods:number id="1E545FB584D36605FCF4C120E20F55A4">602</mods:number>
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<mods:classification id="F8238C59FDF63CA6713A486078EB5F15">journal article</mods:classification>
<mods:identifier id="3DACCE808A3068F8E8DA81A3066B0364" type="DOI">10.5281/zenodo.5170203</mods:identifier>
<mods:identifier id="0010E4077123E8D60F642010C52E3127" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="CD16A06690888C12FAC67A91E240437E" type="Zenodo-Dep">5170203</mods:identifier>
<mods:identifier id="1CFE4D214875875621575193F01F4CCA" type="ZooBank">AB1A57F0-7CB4-4830-920B-DF219740A596</mods:identifier>
</mods:mods>
<treatment id="03F687A7FF9DF861FF01BBE6412BFECF" ID-DOI="http://doi.org/10.5281/zenodo.5186339" ID-Zenodo-Dep="5186339" LSID="urn:lsid:plazi:treatment:03F687A7FF9DF861FF01BBE6412BFECF" httpUri="http://treatment.plazi.org/id/03F687A7FF9DF861FF01BBE6412BFECF" lastPageId="102" lastPageNumber="101" pageId="101" pageNumber="100">
<subSubSection id="C345653AFF9DF862FF01BBE643BAFC2F" box="[192,945,899,923]" pageId="101" pageNumber="100" type="nomenclature">
<paragraph id="8BE036B1FF9DF862FF01BBE643BAFC2F" blockId="101.[192,945,899,955]" box="[192,945,899,923]" pageId="101" pageNumber="100">
<heading id="D0A881DDFF9DF862FF01BBE643BAFC2F" bold="true" box="[192,945,899,923]" fontSize="10" level="2" pageId="101" pageNumber="100" reason="3">
<emphasis id="B92BEAA3FF9DF862FF01BBE643BAFC2F" bold="true" box="[192,945,899,923]" pageId="101" pageNumber="100">
<taxonomicName id="4C5F4D32FF9DF862FF01BBE642F0FC2F" authority="Blahnik &amp; Holzenthal, 2017" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[192,763,899,923]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="101" pageNumber="100" phylum="Tracheophyta" rank="species" species="quadridactyla" status="sp. nov." subGenus="Nanotrichia">
<emphasis id="B92BEAA3FF9DF862FF01BBE64168FC2F" bold="true" box="[192,355,899,923]" italics="true" pageId="101" pageNumber="100">Mortoniella</emphasis>
(
<emphasis id="B92BEAA3FF9DF862FEB7BBE6422AFC2F" bold="true" box="[374,545,899,923]" italics="true" pageId="101" pageNumber="100">Nanotrichia</emphasis>
)
<emphasis id="B92BEAA3FF9DF862FDF5BBE642F0FC2F" bold="true" box="[564,763,899,923]" italics="true" pageId="101" pageNumber="100">quadridactyla</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21857D8FF9DF862FCCBBBE643BAFC2F" box="[778,945,899,923]" pageId="101" pageNumber="100" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C345653AFF9DF862FF01BBC642BCF8CF" pageId="101" pageNumber="100" type="description">
<paragraph id="8BE036B1FF9DF862FF01BBC6411DFC0F" blockId="101.[192,945,899,955]" box="[192,278,931,955]" pageId="101" pageNumber="100">
<figureCitation id="13642A34FF9DF862FF01BBC6411DFC0F" box="[192,278,931,955]" captionStart="Figure 85" captionStartId="225.[192,276,1836,1858]" captionTargetBox="[354,1278,256,1704]" captionTargetId="figure-54@225.[354,1279,256,1704]" captionTargetPageId="225" captionText="Figure 85. Mortoniella (Nanotrichia) quadridactyla, new species, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5170381" httpUri="https://zenodo.org/record/5170381/files/figure.png" pageId="101" pageNumber="100">Fig. 85</figureCitation>
</paragraph>
<paragraph id="8BE036B1FF9DF862FF31BB864554FBCF" blockId="101.[192,1440,995,1148]" pageId="101" pageNumber="100">
This species is probably most closely related to
<taxonomicName id="4C5F4D32FF9DF862FCAABB81443FFC4F" authority="Blahnik &amp; Holzenthal, 2017" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[875,1076,995,1019]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="101" pageNumber="100" phylum="Tracheophyta" rank="species" species="triangularis" status="sp. nov.">
<emphasis id="B92BEAA3FF9DF862FCAABB81443FFC4F" box="[875,1076,995,1019]" italics="true" pageId="101" pageNumber="100">M. triangularis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21857D8FF9DF862FB86BB814484FC4F" box="[1095,1167,996,1019]" pageId="101" pageNumber="100" rank="species">n. sp.</taxonomicNameLabel>
Both species have a deep mesal incision at the apex of the dorsal phallic spine.
<taxonomicName id="4C5F4D32FF9DF862FC27BC664535FBAF" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[998,1342,1027,1051]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="101" pageNumber="100" phylum="Tracheophyta" rank="species" species="quadridactyla">
<emphasis id="B92BEAA3FF9DF862FC27BC664535FBAF" box="[998,1342,1027,1051]" italics="true" pageId="101" pageNumber="100">Mortoniella quadridactyla</emphasis>
</taxonomicName>
can be easily distinguished by having the dorsal phallic spine much narrower in lateral view, with the apex upturned, and with each half of the divided apex subdivided into a pair of acute spines. Tergum X is also simpler in construction, with the apicolateral lobes very broadly rounded.
</paragraph>
<paragraph id="8BE036B1FF9DF862FF31BCC644A4FAAF" blockId="101.[192,1441,1187,1916]" pageId="101" pageNumber="100">
<emphasis id="B92BEAA3FF9DF862FF31BCC6413DFB0F" box="[240,310,1187,1211]" italics="true" pageId="101" pageNumber="100">Adult</emphasis>
—Length of forewing: male
<quantity id="4CA79B54FF9DF862FD54BCC6432EFB0F" box="[661,805,1187,1211]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="101" pageNumber="100" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.7-1.9 mm</quantity>
; female 2.0 mm. Forewing with forks I, II, and III present, hind wing with fork II only; both wings narrow, acute apically. Spur formula 0:3:4. Overall color (in alcohol) yellowish-brown (specimen faded and partially rubbed, wing bar not evident). Forewings of male with scale-like setae paralleling major veins (except apically).
</paragraph>
<paragraph id="8BE036B1FF9DF862FF31BD4642BCF8CF" blockId="101.[192,1441,1187,1916]" pageId="101" pageNumber="100">
<emphasis id="B92BEAA3FF9DF862FF31BD4641A0FA8F" box="[240,427,1315,1339]" italics="true" pageId="101" pageNumber="100">Male genitalia</emphasis>
—Ventral process of segment VI posteriorly projecting, narrow basally, length about 3 times width at base. Segment IX with anterolateral margin rounded and distinctly produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by more than ½ width of segment. Tergum X moderately elongate, lateral margins subparallel, apicomesally with deep V-shaped emargination, extending more than ½ length of tergum; apicolateral lobes simple, rounded apically, formed by mesal invagination; tergum, in lateral view, with rounded apicolateral invagination, demarcating prominent and apically rounded ventrolateral lobe. Inferior appendages with small setose lateral projections, apparently fused ventrally to spine-like projections of mesal pockets. Mesal pockets of inferior appendage small, with very elongate, narrow, posteriorly-directed, spinelike, apicoventral projections. Paramere appendage elongate, strap-like, apex acute, extending about same length as dorsal phallic spine. Phallobase with very short ventral rod-like projections, each distinctly flared apically. Dorsal phallic spine, as viewed laterally, relatively narrow, strongly and sinuously upturned in about middle, apex distinctly forked into pair of acute projections; as viewed dorsally, distinctly widened in apical ½, apicomesally with deep U-shaped invagination, extending about ½ length of spine, each half of divided apex forked preapically to produce 4 apical spine-like projections. Phallicata relatively lightly sclerotized and difficult to discern. Endophallic membrane elongate, with large basodorsal, balloon-like lobe; phallotremal spines very small.
</paragraph>
</subSubSection>
<subSubSection id="C345653AFF9EF861FF31B8A6412BFECF" pageId="102" pageNumber="101" type="materials_examined">
<paragraph id="8BE036B1FF9EF861FF31B8A642F7FF4F" blockId="102.[192,1440,195,379]" pageId="102" pageNumber="101">
<materialsCitation id="3B373CECFF9EF861FF31B8A642F7FF4F" collectingDate="1976-02-17" collectionCode="CM, OS, NMNH" country="Venezuela" county="Barinas" location="Flint" municipality="Rio Santo Domingo" pageId="102" pageNumber="101" specimenCode="UMSP000097089" specimenCount="1" stateProvince="Barinas" typeStatus="holotype">
<emphasis id="B92BEAA3FF9EF861FF31B8A641BFFF6F" bold="true" box="[240,436,195,219]" italics="true" pageId="102" pageNumber="101">
<typeStatus id="54E48813FF9EF861FF31B8A64162FF6F" box="[240,361,195,219]" pageId="102" pageNumber="101" type="holotype">Holotype</typeStatus>
male
</emphasis>
(alcohol)—
<emphasis id="B92BEAA3FF9EF861FDFFB8A64372FF6F" bold="true" box="[574,889,195,219]" pageId="102" pageNumber="101">
<collectingCountry id="F3487621FF9EF861FDFFB8A642F3FF6F" box="[574,760,195,219]" name="Venezuela" pageId="102" pageNumber="101">VENEZUELA</collectingCountry>
:
<collectingRegion id="499BF853FF9EF861FCC5B8A64378FF6F" box="[772,883,195,219]" country="Venezuela" name="Barinas" pageId="102" pageNumber="101">Barinas</collectingRegion>
:
</emphasis>
<collectingCounty id="62814E3DFF9EF861FC41B8A643EEFF6F" box="[896,997,195,219]" pageId="102" pageNumber="101">Barinas</collectingCounty>
,
<collectingMunicipality id="6B84ACCBFF9EF861FC31B8A644EAFF6F" box="[1008,1249,195,219]" pageId="102" pageNumber="101">Río Santo Domingo</collectingMunicipality>
,
<date id="FFE11071FF9EF861FB2DB8A64563FF6F" box="[1260,1384,195,219]" pageId="102" pageNumber="101" value="1976-02-17">
<collectingDate id="EFA5E999FF9EF861FB2DB8A64563FF6F" box="[1260,1384,195,219]" pageId="102" pageNumber="101" value="1976-02-17">17.ii.1976</collectingDate>
</date>
,
<collectionCode id="ED4EAE74FF9EF861FAB2B8A645ABFF6F" box="[1395,1440,195,219]" country="China" name="Chongqing Museum" pageId="102" pageNumber="101">CM</collectionCode>
and
<collectionCode id="ED4EAE74FF9EF861FF36B8864116FF4F" box="[247,285,227,251]" country="USA" name="Oregon State University" pageId="102" pageNumber="101">OS</collectionCode>
<location id="8E80606AFF9EF861FEE5B886416EFF4F" LSID="urn:lsid:plazi:treatment:03F687A7FF9DF861FF01BBE6412BFECF:8E80606AFF9EF861FEE5B886416EFF4F" box="[292,357,227,251]" country="Venezuela" county="Barinas" municipality="Rio Santo Domingo" name="Flint" pageId="102" pageNumber="101" stateProvince="Barinas">Flint</location>
,
<location id="8E80606AFF9EF861FEB0B8814187FF4F" LSID="urn:lsid:plazi:treatment:03F687A7FF9DF861FF01BBE6412BFECF:8E80606AFF9EF861FEB0B8814187FF4F" box="[369,396,228,251]" country="Venezuela" county="Barinas" municipality="Rio Santo Domingo" name="Jr" pageId="102" pageNumber="101" stateProvince="Barinas">Jr</location>
(
<specimenCode id="DBF99ECAFF9EF861FE5DB886427CFF4F" box="[412,631,227,251]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="102" pageNumber="101" type="University or college">UMSP000097089</specimenCode>
) (
<collectionCode id="ED4EAE74FF9EF861FD4EB88642FBFF4F" box="[655,752,227,251]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="102" pageNumber="101" type="Museum">NMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE036B1FF9EF861FF31B96645ABFEAF" blockId="102.[192,1440,195,379]" box="[240,1440,259,283]" pageId="102" pageNumber="101">
<emphasis id="B92BEAA3FF9EF861FF31B96642CCFEAF" bold="true" box="[240,711,259,283]" pageId="102" pageNumber="101">
<emphasis id="B92BEAA3FF9EF861FF31B9664173FEAE" bold="true" box="[240,376,259,282]" italics="true" pageId="102" pageNumber="101">
<typeStatus id="54E48813FF9EF861FF31B9664173FEAE" box="[240,376,259,282]" pageId="102" pageNumber="101" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F3487621FF9EF861FE53B9664242FEAF" box="[402,585,259,283]" name="Venezuela" pageId="102" pageNumber="101">VENEZUELA</collectingCountry>
:
<collectingRegion id="499BF853FF9EF861FD95B96642CAFEAF" box="[596,705,259,283]" country="Venezuela" name="Barinas" pageId="102" pageNumber="101">Barinas</collectingRegion>
:
</emphasis>
same data as
<typeStatus id="54E48813FF9EF861FCACB96643DEFEAF" box="[877,981,259,283]" pageId="102" pageNumber="101" type="holotype">holotype</typeStatus>
<specimenCount id="9D59FD38FF9EF861FC25B966444BFEAF" box="[996,1088,259,283]" pageId="102" pageNumber="101" type="male">2 males</specimenCount>
,
<specimenCount id="9D59FD38FF9EF861FB8BB96644B7FEAF" box="[1098,1212,259,283]" pageId="102" pageNumber="101" type="female">2 females</specimenCount>
(alcohol) (
<collectionCode id="ED4EAE74FF9EF861FAF4B966459FFEAF" box="[1333,1428,259,283]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="102" pageNumber="101" type="Museum">NMNH</collectionCode>
).
</paragraph>
<paragraph id="8BE036B1FF9EF861FF31B946412BFECF" blockId="102.[192,1440,195,379]" pageId="102" pageNumber="101">
<emphasis id="B92BEAA3FF9EF861FF31B9464179FE8F" box="[240,370,291,315]" italics="true" pageId="102" pageNumber="101">Etymology</emphasis>
—This species is named
<taxonomicName id="4C5F4D32FF9EF861FD60B94643E3FE8F" authorityName="Blahnik &amp; Holzenthal" authorityYear="2017" box="[673,1000,291,315]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="102" pageNumber="101" phylum="Tracheophyta" rank="species" species="quadridactyla">
<emphasis id="B92BEAA3FF9EF861FD60B94643E3FE8F" box="[673,1000,291,315]" italics="true" pageId="102" pageNumber="101">Mortoniella quadridactyla</emphasis>
</taxonomicName>
, derived from Latin for four fingers or appendages, and referring to the four acute projections at the apex of the dorsal phallic spine of the species.
</paragraph>
</subSubSection>
</treatment>
</document>