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<document id="66E28BE4DF9893C20D08BA1D61EBFF84" ID-DOI="10.11646/zootaxa.3925.1.9" ID-GBIF-Dataset="98786b6c-26a1-4204-9967-e2190ecce616" ID-ISSN="1175-5326" ID-Zenodo-Dep="243577" ID-ZooBank="52C2B852-7814-4313-B18A-717241C81E4F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461000984459" checkinUser="plazi" docAuthor="Marusik, Yuri M., Esyunin, Sergei L. &amp; Tuneva, Tatyana K." docDate="2015" docId="03F6879CB652FFC5FF01E1C12995B8D6" docLanguage="en" docName="zt03925p144.pdf" docOrigin="Zootaxa 3925 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dictynomorpha daemonis Marusik, Esyunin &amp; Tuneva, 2015, sp. n." docType="treatment" docVersion="12" lastPageNumber="131" masterDocId="FFCFFFE4B650FFC7FF96E2302C07BF25" masterDocTitle="A survey of Palaearctic Dictynidae (Araneae). 1. Taxonomic notes on Dictynomorpha Spassky, 1939, Brigittea Lehtinen, 1967 and Lathys Simon, 1884" masterLastPageNumber="144" masterPageNumber="129" pageNumber="131" updateTime="1698564595630" updateUser="plazi">
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<mods:title id="C2E4F5E7F9FB947DE144E5238D9D38D3">A survey of Palaearctic Dictynidae (Araneae). 1. Taxonomic notes on Dictynomorpha Spassky, 1939, Brigittea Lehtinen, 1967 and Lathys Simon, 1884</mods:title>
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<mods:namePart id="FC10218EFBFB7493BD35C231D27535BE">Marusik, Yuri M.</mods:namePart>
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<mods:namePart id="42306CFAC37166CA7A4E81D91744812F">Esyunin, Sergei L.</mods:namePart>
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<mods:namePart id="53487A88DFF283F88B35C94C4A69ED5D">Tuneva, Tatyana K.</mods:namePart>
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<treatment id="03F6879CB652FFC5FF01E1C12995B8D6" ID-DOI="http://doi.org/10.5281/zenodo.6094414" ID-GBIF-Taxon="119598501" ID-Zenodo-Dep="6094414" LSID="urn:lsid:plazi:treatment:03F6879CB652FFC5FF01E1C12995B8D6" httpUri="http://treatment.plazi.org/id/03F6879CB652FFC5FF01E1C12995B8D6" lastPageNumber="131" pageId="2" pageNumber="131">
<subSubSection id="C3456501B652FFC5FF01E1C12D13BB08" pageId="2" pageNumber="131" type="nomenclature">
<paragraph id="8BE0368AB652FFC5FF01E1C12E1FBB2E" blockId="2.[151,536,1009,1069]" box="[151,536,1009,1035]" pageId="2" pageNumber="131">
<heading id="D0A881E6B652FFC5FF01E1C12E1FBB2E" bold="true" box="[151,536,1009,1035]" fontSize="11" level="1" pageId="2" pageNumber="131" reason="1">
<emphasis id="B92BEA98B652FFC5FF01E1C12E1FBB2E" bold="true" box="[151,536,1009,1035]" pageId="2" pageNumber="131">
<taxonomicName id="4C5F4D09B652FFC5FF01E1C12DC8BB2E" ID-CoL="6D5KN" box="[151,463,1009,1035]" class="Arachnida" family="Dictynidae" genus="Dictynomorpha" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="daemonis" status="sp. nov.">
<emphasis id="B92BEA98B652FFC5FF01E1C12DC8BB2E" bold="true" box="[151,463,1009,1035]" italics="true" pageId="2" pageNumber="131">Dictynomorpha daemonis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21857E3B652FFC5FE40E1C12E1FBB2E" box="[470,536,1009,1035]" pageId="2" pageNumber="131" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE0368AB652FFC5FF01E6242D13BB08" blockId="2.[151,536,1009,1069]" box="[151,276,1044,1069]" pageId="2" pageNumber="131">
<figureCitation id="13642A0FB652FFC5FF01E6242D13BB08" box="[151,276,1044,1069]" captionStart-0="FIGURES 16 21" captionStart-1="FIGURES 22 25" captionStartId-0="5.[151,264,1788,1811]" captionStartId-1="6.[151,264,1125,1148]" captionTargetBox-0="[175,1418,209,1724]" captionTargetBox-1="[162,1424,193,1104]" captionTargetId-0="figure@5.[158,1428,193,1734]" captionTargetId-1="figure@6.[158,1428,193,1105]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 16 21. Dictynomorpha daemonis sp. n., copulatory organs and somatic features. 16 19 Male palp (16 ventral, 17 retrolateral, 18 dorso-retrolateral, 19 prolateral). 20 Habitus, dorsal. 21 Labium and gnathocoxae, ventral. Scale: 16 19 = 0.1 mm, 20 = 0.25 mm. Abbreviations: Cs—cymbial spur; Pa—prolateral apophysis; Ra—retrolateral apophysis; Pp—dorsal process of patella; Pr—retrolateral process of patella; Rp—retrolateral process of cymbium." captionText-1="FIGURES 22 25. Dictynomorpha daemonis sp. n. (22 23), Lathys spasskyi (24 25 from Kyrgyzstan), male palp (22, 24 ventral; 23 tip of conductor, ventral; 25 tibia, retrolateral). Abbreviations: Cf—cymbial fovea; Da—dorsal tibial apophysis; La—lower arm of conductor; Ra—retrolateral tibial apophysis; Va—ventral tibial apophysis; Ua—upper arm of conductor." httpUri-0="https://zenodo.org/record/243580/files/figure.png" httpUri-1="https://zenodo.org/record/243581/files/figure.png" pageId="2" pageNumber="131">Figs 1623</figureCitation>
</paragraph>
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<subSubSection id="C3456501B652FFC5FF01E66C2EAEBB57" box="[151,681,1116,1138]" pageId="2" pageNumber="131" type="reference_group">
<paragraph id="8BE0368AB652FFC5FF01E66C2EAEBB57" blockId="2.[151,681,1116,1138]" box="[151,681,1116,1138]" pageId="2" pageNumber="131">
<treatmentCitationGroup id="AB4F11A4B652FFC5FF01E66C2EAEBB57" box="[151,681,1116,1138]" pageId="2" pageNumber="131">
<taxonomicName id="4C5F4D09B652FFC5FF01E66C2E4EBB57" ID-CoL="6CTKP" authority="Spassky 1939: 138" authorityName="Spassky" authorityPageNumber="138" authorityYear="1939" box="[151,585,1116,1138]" class="Arachnida" family="Dictynidae" genus="Dictynomorpha" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="B92BEA98B652FFC5FF01E66C2D83BB57" box="[151,388,1116,1138]" italics="true" pageId="2" pageNumber="131">Dictynomorpha strandi</emphasis>
<treatmentCitation id="0AFE109BB652FFC5FE1CE66C2E4EBB57" author="Spassky" box="[394,585,1116,1138]" page="138" pageId="2" pageNumber="131" year="1939">
<bibRefCitation id="EFCE4B7BB652FFC5FE1CE66C2E1DBB57" author="Spassky" box="[394,538,1116,1138]" pageId="2" pageNumber="131" refString="Spassky, S. (1939) Araneae palaearcticae novae. III. Festschrift Embrik Strand, 5, 138 - 144." type="journal article" year="1939">Spassky 1939</bibRefCitation>
: 138
</treatmentCitation>
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(in part).
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</paragraph>
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<subSubSection id="C3456501B652FFC5FF01E6AE280EBBFE" pageId="2" pageNumber="131" type="materials_examined">
<paragraph id="8BE0368AB652FFC5FF01E6AE280EBBFE" blockId="2.[151,1437,1182,2035]" pageId="2" pageNumber="131">
<typeStatus id="54E48828B652FFC5FF01E6AE2CD7BB92" box="[151,208,1182,1207]" pageId="2" pageNumber="131">
<emphasis id="B92BEA98B652FFC5FF01E6AE2CD7BB92" bold="true" box="[151,208,1182,1207]" pageId="2" pageNumber="131">Type</emphasis>
</typeStatus>
.
<typeStatus id="54E48828B652FFC5FF74E6AE2D4DBB92" box="[226,330,1182,1207]" pageId="2" pageNumber="131" type="holotype">Holotype</typeStatus>
♂ (
<collectionCode id="ED4EAE4FB652FFC5FEEFE6AF2DD1BB92" box="[377,470,1183,1207]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="2" pageNumber="131">ZMMU</collectionCode>
),
<emphasis id="B92BEA98B652FFC5FE7AE6AE2FDEBB92" bold="true" box="[492,985,1182,1207]" pageId="2" pageNumber="131">
<collectingCountry id="F348761AB652FFC5FE7AE6AE2EB2BB92" box="[492,693,1182,1207]" name="Kazakhstan" pageId="2" pageNumber="131">KAZAKHSTAN</collectingCountry>
,
<emphasis id="B92BEA98B652FFC5FD55E6AE2FDEBB92" bold="true" box="[707,985,1182,1207]" italics="true" pageId="2" pageNumber="131">
South
<collectingCountry id="F348761AB652FFC5FC87E6AE2F9FBB92" box="[785,920,1182,1207]" name="Kazakhstan" pageId="2" pageNumber="131">Kazakhstan</collectingCountry>
Area
</emphasis>
</emphasis>
: ~
<quantity id="4CA79B6FB652FFC5FC6FE6AF2832BB92" box="[1017,1077,1182,1207]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="131" unit="km" value="7.0">7 km</quantity>
NE Terekty Village, Boralday- Tau Range, N 42°51.751', E 69°50.887',
<quantity id="4CA79B6FB652FFC5FDC9E6F22EAFBBFF" box="[607,680,1218,1243]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="2" pageNumber="131" unit="m" value="650.0">650 m</quantity>
, 11
<date id="FFE1104AB652FFC5FD4CE6F22F5ABBFE" box="[730,861,1218,1243]" pageId="2" pageNumber="131" value="2010-05-13">13.05.2010</date>
, T. Tuneva leg.
</paragraph>
</subSubSection>
<subSubSection id="C3456501B652FFC5FF51E6D62864BA07" pageId="2" pageNumber="131" type="etymology">
<paragraph id="8BE0368AB652FFC5FF51E6D62864BA07" blockId="2.[151,1437,1182,2035]" pageId="2" pageNumber="131">
<emphasis id="B92BEA98B652FFC5FF51E6D62D41BBDA" bold="true" box="[199,326,1254,1279]" pageId="2" pageNumber="131">Etymology</emphasis>
. The species name is derived from “daemon” a Latinized Greek word “δαίµων” meaning mythological creatures of mixed nature that can do good or evil; noun in genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3456501B652FFC5FF51E71E2F27BAAB" pageId="2" pageNumber="131" type="diagnosis">
<paragraph id="8BE0368AB652FFC5FF51E71E2F27BAAB" blockId="2.[151,1437,1182,2035]" pageId="2" pageNumber="131">
<emphasis id="B92BEA98B652FFC5FF51E71E2D3DBA62" bold="true" box="[199,314,1326,1351]" pageId="2" pageNumber="131">Diagnosis</emphasis>
. The species is most similar to
<taxonomicName id="4C5F4D09B652FFC5FD0BE71F2F08BA62" box="[669,783,1326,1351]" class="Arachnida" family="Dictynidae" genus="Dictynomorpha" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="B92BEA98B652FFC5FD0BE71F2F08BA62" box="[669,783,1326,1351]" italics="true" pageId="2" pageNumber="131">D. strandi</emphasis>
</taxonomicName>
from which it can be separated by the strongly bent tip of embolus and the presence of a cymbial spur. The two species differ also in the shape of cymbium, the lower arm of the conductor, the tibial apophyses and patellar processes.
</paragraph>
</subSubSection>
<subSubSection id="C3456501B652FFC5FF01E7AA2D97B88E" pageId="2" pageNumber="131" type="description">
<paragraph id="8BE0368AB652FFC5FF01E7AA2DA0B942" blockId="2.[151,1437,1182,2035]" pageId="2" pageNumber="131">
<emphasis id="B92BEA98B652FFC5FF01E7AA2D26BA96" bold="true" box="[151,289,1434,1459]" pageId="2" pageNumber="131">Description</emphasis>
. Male. Total length 3.2. Carapace 1.4 long, 1.2 wide; brown, lighter median area. Sternum yellow with brown margin, covered with white setae. Chelicerae brown, with 3 prolateral teeth and 1 retrolateral tooth. Mouth parts (
<figureCitation id="13642A0FB652FFC5FF4BE7D22D34BADE" box="[221,307,1506,1531]" captionStart="FIGURES 16 21" captionStartId="5.[151,264,1788,1811]" captionTargetBox="[175,1418,209,1724]" captionTargetId="figure@5.[158,1428,193,1734]" captionTargetPageId="5" captionText="FIGURES 16 21. Dictynomorpha daemonis sp. n., copulatory organs and somatic features. 16 19 Male palp (16 ventral, 17 retrolateral, 18 dorso-retrolateral, 19 prolateral). 20 Habitus, dorsal. 21 Labium and gnathocoxae, ventral. Scale: 16 19 = 0.1 mm, 20 = 0.25 mm. Abbreviations: Cs—cymbial spur; Pa—prolateral apophysis; Ra—retrolateral apophysis; Pp—dorsal process of patella; Pr—retrolateral process of patella; Rp—retrolateral process of cymbium." httpUri="https://zenodo.org/record/243580/files/figure.png" pageId="2" pageNumber="131">Fig. 21</figureCitation>
): labium about 1.6 times longer than wide; endites as wide as labium, two times longer than labium width, converging. Legs uniform yellow. Leg formula 1243. Femora and tibiae without spines; metatarsi IIIV with 3 ventro-distal spines. Abdomen light grеy with white spots and pattern composed of grey setae; dorsum with 3 pairs of sigilla (
<figureCitation id="13642A0FB652FFC5FED0E47E2D9CB942" box="[326,411,1614,1639]" captionStart="FIGURES 16 21" captionStartId="5.[151,264,1788,1811]" captionTargetBox="[175,1418,209,1724]" captionTargetId="figure@5.[158,1428,193,1734]" captionTargetPageId="5" captionText="FIGURES 16 21. Dictynomorpha daemonis sp. n., copulatory organs and somatic features. 16 19 Male palp (16 ventral, 17 retrolateral, 18 dorso-retrolateral, 19 prolateral). 20 Habitus, dorsal. 21 Labium and gnathocoxae, ventral. Scale: 16 19 = 0.1 mm, 20 = 0.25 mm. Abbreviations: Cs—cymbial spur; Pa—prolateral apophysis; Ra—retrolateral apophysis; Pp—dorsal process of patella; Pr—retrolateral process of patella; Rp—retrolateral process of cymbium." httpUri="https://zenodo.org/record/243580/files/figure.png" pageId="2" pageNumber="131">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="8BE0368AB652FFC5FF51E4432F12B98A" blockId="2.[151,1437,1182,2035]" pageId="2" pageNumber="131">Leg measurements: I 4.33 (1.30+1.55+0.95+0.53), II 3.73 (1.13+1.28+0.83+0.50), III 3.08 (0.90+1.13+0.68+0.38), IV 3.58 (1.08+1.30+0.88+0.33).</paragraph>
<paragraph id="8BE0368AB652FFC5FF51E48A2CBBB8A2" blockId="2.[151,1437,1182,2035]" pageId="2" pageNumber="131">
Palp as in
<figureCitation id="13642A0FB652FFC5FEABE48A2DBAB9F6" box="[317,445,1722,1747]" captionStart="FIGURES 16 21" captionStartId="5.[151,264,1788,1811]" captionTargetBox="[175,1418,209,1724]" captionTargetId="figure@5.[158,1428,193,1734]" captionTargetPageId="5" captionText="FIGURES 16 21. Dictynomorpha daemonis sp. n., copulatory organs and somatic features. 16 19 Male palp (16 ventral, 17 retrolateral, 18 dorso-retrolateral, 19 prolateral). 20 Habitus, dorsal. 21 Labium and gnathocoxae, ventral. Scale: 16 19 = 0.1 mm, 20 = 0.25 mm. Abbreviations: Cs—cymbial spur; Pa—prolateral apophysis; Ra—retrolateral apophysis; Pp—dorsal process of patella; Pr—retrolateral process of patella; Rp—retrolateral process of cymbium." httpUri="https://zenodo.org/record/243580/files/figure.png" pageId="2" pageNumber="131">Figs 1619</figureCitation>
,
<figureCitation id="13642A0FB652FFC5FE5FE48B2E14B9F6" box="[457,531,1723,1747]" captionStart="FIGURES 22 25" captionStartId="6.[151,264,1125,1148]" captionTargetBox="[162,1424,193,1104]" captionTargetId="figure@6.[158,1428,193,1105]" captionTargetPageId="6" captionText="FIGURES 22 25. Dictynomorpha daemonis sp. n. (22 23), Lathys spasskyi (24 25 from Kyrgyzstan), male palp (22, 24 ventral; 23 tip of conductor, ventral; 25 tibia, retrolateral). Abbreviations: Cf—cymbial fovea; Da—dorsal tibial apophysis; La—lower arm of conductor; Ra—retrolateral tibial apophysis; Va—ventral tibial apophysis; Ua—upper arm of conductor." httpUri="https://zenodo.org/record/243581/files/figure.png" pageId="2" pageNumber="131">2223</figureCitation>
. Femur modified, with small prolateral knob on anterior third and impression of knob on patella. Patella modified with two processes: retrolateral (
<emphasis id="B92BEA98B652FFC5FCEAE4EF2F9FB9D3" box="[892,920,1759,1782]" italics="true" pageId="2" pageNumber="131">Pr</emphasis>
) and a dorsal (
<emphasis id="B92BEA98B652FFC5FBA8E4EF285AB9D3" box="[1086,1117,1759,1782]" italics="true" pageId="2" pageNumber="131">Pp</emphasis>
). Tibia cylindrical, with two apophyses, prolateral (
<emphasis id="B92BEA98B652FFC5FE0DE5342DBDB83E" box="[411,442,1796,1819]" italics="true" pageId="2" pageNumber="131">Pa</emphasis>
) and retrolateral (
<emphasis id="B92BEA98B652FFC5FD1BE5342EABB83E" box="[653,684,1796,1819]" italics="true" pageId="2" pageNumber="131">Ra</emphasis>
). Cymbium with massive extension and retrolateral process (
<emphasis id="B92BEA98B652FFC5FAE2E5342994B83E" box="[1396,1427,1796,1819]" italics="true" pageId="2" pageNumber="131">Rp</emphasis>
) and long cymbial spur (
<emphasis id="B92BEA98B652FFC5FE30E5172DC4B81B" box="[422,451,1831,1854]" italics="true" pageId="2" pageNumber="131">Cs</emphasis>
) subdivided on top (
<figureCitation id="13642A0FB652FFC5FD26E5162F02B81A" box="[688,773,1830,1855]" captionStart="FIGURES 16 21" captionStartId="5.[151,264,1788,1811]" captionTargetBox="[175,1418,209,1724]" captionTargetId="figure@5.[158,1428,193,1734]" captionTargetPageId="5" captionText="FIGURES 16 21. Dictynomorpha daemonis sp. n., copulatory organs and somatic features. 16 19 Male palp (16 ventral, 17 retrolateral, 18 dorso-retrolateral, 19 prolateral). 20 Habitus, dorsal. 21 Labium and gnathocoxae, ventral. Scale: 16 19 = 0.1 mm, 20 = 0.25 mm. Abbreviations: Cs—cymbial spur; Pa—prolateral apophysis; Ra—retrolateral apophysis; Pp—dorsal process of patella; Pr—retrolateral process of patella; Rp—retrolateral process of cymbium." httpUri="https://zenodo.org/record/243580/files/figure.png" pageId="2" pageNumber="131">Fig. 19</figureCitation>
). Bulbus round (as high as wide); conductor with strongly reduced upper arm (
<emphasis id="B92BEA98B652FFC5FEEEE57C2D9DB846" box="[376,410,1868,1891]" italics="true" pageId="2" pageNumber="131">Ua</emphasis>
) and simple lower arm (
<emphasis id="B92BEA98B652FFC5FD38E57C2ECCB846" box="[686,715,1868,1891]" italics="true" pageId="2" pageNumber="131">La</emphasis>
); embolus relatively short, smoothly arching, with strongly bent tip.
</paragraph>
<paragraph id="8BE0368AB652FFC5FF51E5A22D97B88E" blockId="2.[151,1437,1182,2035]" box="[199,400,1938,1963]" pageId="2" pageNumber="131">Female unknown.</paragraph>
</subSubSection>
<subSubSection id="C3456501B652FFC5FF51E5862995B8D6" pageId="2" pageNumber="131" type="distribution">
<paragraph id="8BE0368AB652FFC5FF51E5862995B8D6" blockId="2.[151,1437,1182,2035]" pageId="2" pageNumber="131">
<emphasis id="B92BEA98B652FFC5FF51E5862D5EB8EA" bold="true" box="[199,345,1974,1999]" pageId="2" pageNumber="131">Distribution</emphasis>
. So far, the species is known only from the
<typeStatus id="54E48828B652FFC5FCC6E5862FB5B8EA" box="[848,946,1974,1999]" pageId="2" pageNumber="131" type="holotype">holotype</typeStatus>
male. It is very likely that the record of
<taxonomicName id="4C5F4D09B652FFC5FA17E5872CE1B8D6" class="Arachnida" family="Dictynidae" genus="Dictynomorpha" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="strandi">
<emphasis id="B92BEA98B652FFC5FA17E5872CE1B8D6" italics="true" pageId="2" pageNumber="131">D. strandi</emphasis>
</taxonomicName>
from the environs of Almaty (
<bibRefCitation id="EFCE4B7BB652FFC5FDAAE5EA2EDAB8D6" author="Spassky" box="[572,733,2010,2035]" pageId="2" pageNumber="131" refString="Spassky, S. (1939) Araneae palaearcticae novae. III. Festschrift Embrik Strand, 5, 138 - 144." type="journal article" year="1939">Spassky 1939</bibRefCitation>
), the same locality as
<taxonomicName id="4C5F4D09B652FFC5FC45E5EC2867B8D6" box="[979,1120,2010,2035]" class="Arachnida" family="Dictynidae" genus="Dictynomorpha" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="131" phylum="Arthropoda" rank="species" species="daemonis" status="sp. nov.">
<emphasis id="B92BEA98B652FFC5FC45E5EC2867B8D6" box="[979,1120,2010,2035]" italics="true" pageId="2" pageNumber="131">D. daemonis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21857E3B652FFC5FBF1E5EB28A2B8D7" box="[1127,1189,2011,2034]" pageId="2" pageNumber="131" rank="species">sp. n.</taxonomicNameLabel>
, refer to this species.
</paragraph>
</subSubSection>
</treatment>
</document>