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<document id="8170E3B46F8A333CF4182B32F8106834" ID-DOI="10.5281/zenodo.275645" ID-GBIF-Dataset="c9d212c2-bfed-4043-971c-b80ab6d7ff9c" ID-ISSN="1175-5326" ID-Zenodo-Dep="275645" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460153538636" checkinUser="plazi" docAuthor="Geiger, Daniel L. &amp; Mclean, James H." docDate="2010" docId="03DCBC11C24CFFA40ED21CBDFBF967C6" docLanguage="en" docName="zt02356p035.pdf" docOrigin="Zootaxa 2356" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thieleella kelseyi Dall 1905" docType="treatment" docVersion="13" lastPageNumber="28" masterDocId="FFE5C469C259FFBF0E451F42FFF1673A" masterDocTitle="New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="22" updateTime="1698235584404" updateUser="plazi">
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<mods:title id="1DA7FC29D59EE5D293129EC5B82F8E88">New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda)</mods:title>
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<mods:namePart id="0111E8284DE699329B7E72A6F37DCDED">Geiger, Daniel L.</mods:namePart>
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<mods:namePart id="39714371E3892089A7135E0F6BFA57CE">Mclean, James H.</mods:namePart>
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<mods:date id="1A5AB02D1804A8CACD5FDDDC3EC7A602">2010</mods:date>
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<treatment id="03DCBC11C24CFFA40ED21CBDFBF967C6" ID-DOI="http://doi.org/10.5281/zenodo.6210883" ID-GBIF-Taxon="127654801" ID-Zenodo-Dep="6210883" LSID="urn:lsid:plazi:treatment:03DCBC11C24CFFA40ED21CBDFBF967C6" httpUri="http://treatment.plazi.org/id/03DCBC11C24CFFA40ED21CBDFBF967C6" lastPageId="27" lastPageNumber="28" pageId="21" pageNumber="22">
<subSubSection id="C36F5E8CC24CFFAA0ED21CBDFECE6306" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BCA0D07C24CFFAA0ED21CBDFDF16323" blockId="21.[151,512,1023,1084]" box="[151,512,1023,1049]" pageId="21" pageNumber="22">
<heading id="D082BA6BC24CFFAA0ED21CBDFDF16323" bold="true" box="[151,512,1023,1049]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="4C757684C24CFFAA0ED21CBDFDF16323" ID-CoL="56GHX" authority="Dall, 1905" authorityName="Dall" authorityYear="1905" box="[151,512,1023,1049]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21CBDFDF16323" bold="true" box="[151,512,1023,1049]" pageId="21" pageNumber="22">
<emphasis id="B901D115C24CFFAA0ED21CBDFE996323" bold="true" box="[151,360,1023,1049]" italics="true" pageId="21" pageNumber="22">Thieleella kelseyi</emphasis>
(
<bibRefCitation id="EFE470F6C24CFFAA0F3D1CBDFE096323" author="Dall" box="[376,504,1023,1049]" pageId="21" pageNumber="34" refString="Dall, W. H. (1905). Some new species of mollusks from California. The Nautilus, 18, 123 - 125." type="journal article" year="1905">Dall, 1905</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21B60FECE6306" blockId="21.[151,512,1023,1084]" box="[151,319,1058,1084]" pageId="21" pageNumber="22">
<figureCitation id="134E1182C24CFFAA0ED21B60FECE6306" box="[151,319,1058,1084]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStart-3="FIGURE 19" captionStartId-0="23.[151,260,1869,1893]" captionStartId-1="24.[151,256,1104,1128]" captionStartId-2="25.[151,254,1321,1345]" captionStartId-3="26.[151,254,1674,1698]" captionTargetBox-0="[151,1436,335,1856]" captionTargetBox-1="[151,1435,194,1081]" captionTargetBox-2="[151,1435,717,1300]" captionTargetBox-3="[151,1435,493,1652]" captionTargetId-0="figure@23.[151,1436,335,1857]" captionTargetId-1="figure@24.[151,1436,194,1082]" captionTargetId-2="figure@25.[151,1436,717,1300]" captionTargetId-3="figure@26.[151,1436,493,1653]" captionTargetPageId-0="23" captionTargetPageId-1="24" captionTargetPageId-2="25" captionTargetPageId-3="26" captionText-0="FIGURE 16. Thieleella kelseyi. A. Neotype, here designated. 183 m, off La Jolla, California, USA, 32.843 ˚ N, 117.392 ˚ W (SBMNH 3215). Radula shown in Figure 13 A – C. B. Between Tanner Bank and Cortez Bank, California, USA, 32.450 ˚ N, 119.250 ˚ W (SBMNH 33350). Radula shown in Figure 13 D – E. Scale bar shell = 1 mm. Scale bar protoconch = 100 µm." captionText-1="FIGURE 17. Radula of Thieleella kelseyi. A – C. From here designated neotype, 183 m, off La Jolla, California, USA, 32.843 ˚ N, 117.392 ˚ W (SBMNH 3215). For shell see Figure 12 A. D – E. Between Tanner Bank and Cortez Bank, California, USA, 32.450 ˚ N, 119.250 ˚ W (SBMNH 33350). For shell see Figure 12 B. Scale bars A, D = 500 µm. Scale bars B, C, E. = 50 µm." captionText-2="FIGURE 18. Preserved body of Thieleella kelseyi. 421 m, 2.3 miles, 48 ˚ T, to S Pt. Light, Santa Rosa Island, California, 33.850 ˚ N, 120.183 ˚ W (LACM 23129). Stereomicroscope images, z-stacked with HeliconFocus. Scale bar = 1 mm. A. Ventral view. B. Lateral view. ct: cephalic tentacle; es: eye stalk; et: epipodial tentacle; j: jaws; m: mantle; mp: metapodium; op: operculum; pp: propodium; sn: snout; st: suboptic tentacle." captionText-3="FIGURE 19. Preserved body of Thieleella kelseyi. 421 m, 2.3 miles 48 ˚ T to S Pt. Light, Santa Rosa Island, California, 33. 850 ˚ N, 120. 183 ˚ W (LACM 23129). A. Operculum. Scale bar = 1 mm. B. Entire preserved body, hexamethyldisilizane treated. Scale bar = 1 mm. C. Ventral view of anterior portion. Scale bar = 1 mm. D. Ventral view of anterior epipodial tentacles plus ESO. Scale bar = 200 µm. E. Dorsal view of epipodial tentacles and ESO; note biserial papillation of epipodial tentacles. Scale bar = 200 µm. F. Mantle tentacle at posterior end of mantle enlarged. Scale bar = 100 µm. G. Overview of gill and mantle. Fat arrow: direction of view of H. Scale bar = 1 mm. H. Bursicles with openings. Scale bar = 100 µm. Abbreviations: act: accessory cephalic tentacle. b: bursicle. bo: bursicle opening. ct: cephalic tentacle. es: eyestalk. ESO: epipodial sense organ. et: epipodial tentacles. gf: gill filaments. j: jaw. mp: metapodium. mt: mantle tentacle. oaf: operculum attachment flap. op: operculum. opg: opening of propodial gland. pp: propodium. rp: rectal papilla. sn: snout. st. suboptic tentacle." figureDoi-0="http://doi.org/10.5281/zenodo.3558592" figureDoi-1="http://doi.org/10.5281/zenodo.3558594" figureDoi-2="http://doi.org/10.5281/zenodo.3558505" figureDoi-3="http://doi.org/10.5281/zenodo.3558596" httpUri-0="https://zenodo.org/record/3558592/files/figure.png" httpUri-1="https://zenodo.org/record/3558594/files/figure.png" httpUri-2="https://zenodo.org/record/3558505/files/figure.png" httpUri-3="https://zenodo.org/record/3558596/files/figure.png" pageId="21" pageNumber="22">Figures 16–19</figureCitation>
</paragraph>
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<paragraph id="8BCA0D07C24CFFAA0ED21B2FFC6363BE" blockId="21.[151,1436,1133,1636]" box="[151,914,1133,1156]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED21B2FFC6363BE" box="[151,914,1133,1156]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21B2FFD4463BE" authority="Dall, 1905: 124" authorityName="Dall" authorityPageNumber="124" authorityYear="1905" box="[151,693,1133,1156]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi" subGenus="Schizotrochus">
<emphasis id="B901D115C24CFFAA0ED21B2FFEFA63BE" box="[151,267,1133,1156]" italics="true" pageId="21" pageNumber="22">Scissurella</emphasis>
(
<emphasis id="B901D115C24CFFAA0F5F1B2FFE5F63BE" box="[282,430,1133,1156]" italics="true" pageId="21" pageNumber="22">Schizotrochus</emphasis>
)
<emphasis id="B901D115C24CFFAA0FFB1B2FFDF663BE" box="[446,519,1133,1156]" italics="true" pageId="21" pageNumber="22">kelseyi</emphasis>
<treatmentCitation id="0AD42B16C24CFFAA0C4B1B2FFD4463BE" author="Dall" box="[526,693,1133,1156]" page="124" pageId="21" pageNumber="22" year="1905">
<bibRefCitation id="EFE470F6C24CFFAA0C4B1B2FFD7363BE" author="Dall" box="[526,642,1133,1156]" pageId="21" pageNumber="34" refString="Dall, W. H. (1905). Some new species of mollusks from California. The Nautilus, 18, 123 - 125." type="journal article" year="1905">Dall, 1905</bibRefCitation>
: 124
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–125, not illustrated.
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<paragraph id="8BCA0D07C24CFFAA0ED21BCFFDEF639E" blockId="21.[151,1436,1133,1636]" box="[151,542,1165,1188]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED21BCFFDEF639E" box="[151,542,1165,1188]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21BCFFDEB639E" authority="Thiele, 1912: 18" authorityName="Thiele" authorityPageNumber="18" authorityYear="1912" box="[151,538,1165,1188]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21BCFFEAB639E" box="[151,346,1165,1188]" italics="true" pageId="21" pageNumber="22">Scissurella kelseyi</emphasis>
:
<treatmentCitation id="0AD42B16C24CFFAA0F2D1BCFFDEB639E" author="Thiele" box="[360,538,1165,1188]" page="18" pageId="21" pageNumber="22" year="1912">
<bibRefCitation id="EFE470F6C24CFFAA0F2D1BCFFE02639E" author="Thiele" box="[360,499,1165,1188]" pageId="21" pageNumber="35" refString="Thiele, J. (1912) Scissurelliden und Fissurelliden. In: Kobelt, H. C. &amp; Kuster, W. (eds), Systematisches Conchylien- Cabinet von Martini und Chemnitz, Bauer &amp; Raspe, Nurnberg, pp. 1 - 34, pls. 1 - 4." type="book chapter" year="1912">Thiele, 1912</bibRefCitation>
: 18
</treatmentCitation>
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.
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</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21BEFFDD363FE" blockId="21.[151,1436,1133,1636]" box="[151,546,1197,1220]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21BEFFDEF63FE" authority="Dall, 1921a: 183" authorityName="Dall" authorityPageNumber="183" authorityYear="1921" box="[151,542,1197,1220]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21BEFFEAB63FE" box="[151,346,1197,1220]" italics="true" pageId="21" pageNumber="22">Scissurella kelseyi</emphasis>
: Dall, 1921a: 183
</taxonomicName>
.
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21B8FFDB063DE" blockId="21.[151,1436,1133,1636]" box="[151,577,1229,1252]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED21B8FFDB063DE" box="[151,577,1229,1252]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21B8FFDCC63DE" authority="Oldroyd, 1924: 180" authorityName="Oldroyd" authorityPageNumber="180" authorityYear="1924" box="[151,573,1229,1252]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21B8FFEAB63DE" box="[151,346,1229,1252]" italics="true" pageId="21" pageNumber="22">Scissurella kelseyi</emphasis>
:
<treatmentCitation id="0AD42B16C24CFFAA0F2D1B8FFDCC63DE" author="Oldroyd" box="[360,573,1229,1252]" page="180" pageId="21" pageNumber="22" year="1924">
<bibRefCitation id="EFE470F6C24CFFAA0F2D1B8FFDF663DE" author="Oldroyd" box="[360,519,1229,1252]" pageId="21" pageNumber="35" refString="Oldroyd, I. S. (1924) Marine shells of Puget Sound and vicinity. Publications Puget Sound Biological Station of the University of Washington, 4, 1 - 272." type="journal article" year="1924">Oldroyd, 1924</bibRefCitation>
: 180
</treatmentCitation>
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.
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</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21BAFFDB0623E" blockId="21.[151,1436,1133,1636]" box="[151,577,1261,1284]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED21BAFFDB0623E" box="[151,577,1261,1284]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21BAFFDCC623E" authority="Oldroyd, 1927: 228" authorityName="Oldroyd" authorityPageNumber="228" authorityYear="1927" box="[151,573,1261,1284]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21BAFFEAB623E" box="[151,346,1261,1284]" italics="true" pageId="21" pageNumber="22">Scissurella kelseyi</emphasis>
:
<treatmentCitation id="0AD42B16C24CFFAA0F2D1BAFFDCC623E" author="Oldroyd" box="[360,573,1261,1284]" page="228" pageId="21" pageNumber="22" year="1927">
<bibRefCitation id="EFE470F6C24CFFAA0F2D1BAFFDF6623E" author="Oldroyd" box="[360,519,1261,1284]" pageId="21" pageNumber="35" refString="Oldroyd, I. S. (1927) The Marine Shells of the West Coast of North America vol. 2, part 3. Stanford University Press, Stanford. 339 pp., pls. 73 - 108." type="book chapter" year="1927">Oldroyd, 1927</bibRefCitation>
: 228
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21A4FFDE3621E" blockId="21.[151,1436,1133,1636]" box="[151,530,1293,1316]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED21A4FFDE3621E" box="[151,530,1293,1316]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21A4FFDFE621E" authority="Keen, 1937: 45" authorityName="Keen" authorityPageNumber="45" authorityYear="1937" box="[151,527,1293,1316]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21A4FFEAB621E" box="[151,346,1293,1316]" italics="true" pageId="21" pageNumber="22">Scissurella kelseyi</emphasis>
:
<treatmentCitation id="0AD42B16C24CFFAA0F2D1A4FFDFE621E" author="Keen" box="[360,527,1293,1316]" page="45" pageId="21" pageNumber="22" year="1937">
<bibRefCitation id="EFE470F6C24CFFAA0F2D1A4FFE16621E" author="Keen" box="[360,487,1293,1316]" pageId="21" pageNumber="35" refString="Keen, A. M. (1937) An Abridged Check List and Bibilography of West North American Marine Mollusca. Stanford University Press, Stanford. 87 pp." type="book" year="1937">Keen, 1937</bibRefCitation>
: 45
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21A6FFDD9627E" blockId="21.[151,1436,1133,1636]" box="[151,552,1325,1348]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED21A6FFDD9627E" box="[151,552,1325,1348]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21A6FFDD5627E" authority="Burch, 1946a: 27" authorityName="Burch" authorityPageNumber="27" authorityYear="1946" box="[151,548,1325,1348]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21A6FFEAB627E" box="[151,346,1325,1348]" italics="true" pageId="21" pageNumber="22">Scissurella kelseyi</emphasis>
:
<treatmentCitation id="0AD42B16C24CFFAA0F2D1A6FFDD5627E" author="Burch" box="[360,548,1325,1348]" page="27" pageId="21" pageNumber="22" year="1946">
<bibRefCitation id="EFE470F6C24CFFAA0F2D1A6FFE0A627E" author="Burch" box="[360,507,1325,1348]" pageId="21" pageNumber="34" refString="Burch, J. Q. (1946 a) Check List of West North American Marine Mollusks. Burch, Los Angeles. 30 pp." type="book" year="1946" yearSuffix="a">Burch, 1946a</bibRefCitation>
: 27
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21A0FFDD8625E" blockId="21.[151,1436,1133,1636]" box="[151,553,1357,1380]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED21A0FFDD8625E" box="[151,553,1357,1380]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21A0FFDD4625E" authority="Burch, 1946b: 23" authorityName="Burch" authorityPageNumber="23" authorityYear="1946" box="[151,549,1357,1380]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21A0FFEAB625E" box="[151,346,1357,1380]" italics="true" pageId="21" pageNumber="22">Scissurella kelseyi</emphasis>
:
<treatmentCitation id="0AD42B16C24CFFAA0F2D1A0FFDD4625E" author="Burch" box="[360,549,1357,1380]" page="23" pageId="21" pageNumber="22" year="1946">
<bibRefCitation id="EFE470F6C24CFFAA0F2D1A0FFE0C625E" author="Burch" box="[360,509,1357,1380]" pageId="21" pageNumber="34" refString="Burch, J. Q. (1946 b) [No title]. Minutes of the Conchological Club of Southern California, 58, 1 - 27." type="journal article" year="1946" yearSuffix="b">Burch, 1946b</bibRefCitation>
: 23
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21A2FFDE462BE" blockId="21.[151,1436,1133,1636]" box="[151,533,1389,1412]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED21A2FFDE462BE" box="[151,533,1389,1412]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21A2FFDE362BE" authority="Draper, 1973: 8" authorityName="Draper" authorityPageNumber="8" authorityYear="1973" box="[151,530,1389,1412]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21A2FFEAB62BE" box="[151,346,1389,1412]" italics="true" pageId="21" pageNumber="22">Scissurella kelseyi</emphasis>
:
<treatmentCitation id="0AD42B16C24CFFAA0F2D1A2FFDE362BE" author="Draper" box="[360,530,1389,1412]" page="8" pageId="21" pageNumber="22" year="1973">
<bibRefCitation id="EFE470F6C24CFFAA0F2D1A2FFE0962BE" author="Draper" box="[360,504,1389,1412]" pageId="21" pageNumber="34" refString="Draper, B. (1973) Minute shells, part 3. Tabulata, 6 (2), 8 - 12." type="journal article" year="1973">Draper, 1973</bibRefCitation>
: 8
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21ACEFE92629E" blockId="21.[151,1436,1133,1636]" box="[151,355,1420,1444]" pageId="21" pageNumber="22">
<emphasis id="B901D115C24CFFAA0ED21ACEFE92629E" bold="true" box="[151,355,1420,1444]" pageId="21" pageNumber="22">Misidentifications</emphasis>
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED21AEFFE7A613E" blockId="21.[151,1436,1133,1636]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED21AEFFEAC62FE" box="[151,349,1453,1476]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0ED21AEFFEAC62FE" box="[151,349,1453,1476]" italics="true" pageId="21" pageNumber="22">Scissurella kelseyi</emphasis>
</taxonomicName>
non
<bibRefCitation id="EFE470F6C24CFFAA0FDE1AEFFDE262FE" author="Dall" box="[411,531,1453,1476]" pageId="21" pageNumber="34" refString="Dall, W. H. (1905). Some new species of mollusks from California. The Nautilus, 18, 123 - 125." type="journal article" year="1905">Dall, 1905</bibRefCitation>
:
<bibRefCitation id="EFE470F6C24CFFAA0C641AEFFD2462FE" author="Talmadge" box="[545,725,1453,1476]" pageId="21" pageNumber="35" refString="Talmadge, R. R. (1966) Notes on the Mollusca of Prince William Sound, Alaska. The Veliger, 9, 82 - 86." type="journal article" year="1966">Talmadge, 1966</bibRefCitation>
: 83 [Identification
<emphasis id="B901D115C24CFFAA0DE91AEFFC2562FE" box="[940,980,1453,1476]" italics="true" pageId="21" pageNumber="22">fide</emphasis>
<bibRefCitation id="EFE470F6C24CFFAA0D9B1AEFFB7062FE" author="McLean" box="[990,1153,1453,1476]" pageId="21" pageNumber="35" refString="McLean, J. H. (1967) West American Scissurellidae. The Veliger, 9, 404 - 410." type="journal article" year="1967">McLean, 1967</bibRefCitation>
, is
<taxonomicName id="4C757684C24CFFAA0AE91AEFFAC362FE" ID-CoL="66MMC" box="[1196,1330,1453,1476]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="lamellata">
<emphasis id="B901D115C24CFFAA0AE91AEFFAC362FE" box="[1196,1330,1453,1476]" italics="true" pageId="21" pageNumber="22">A. lamellata</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFE470F6C24CFFAA0B061AEFFEFF62DE" author="McLean" pageId="21" pageNumber="35" refString="McLean, J. H. (1984) New species of northeast Pacific archaeogastropods. The Veliger, 26, 233 - 239." type="journal article" year="1984">McLean (1984)</bibRefCitation>
identified “
<taxonomicName id="4C757684C24CFFAA0FD61A8FFCEA62DE" ID-CoL="66MMC" authority="McLean (1967)" authorityName="McLean" authorityYear="1967" box="[403,795,1485,1508]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="lamellata">
<emphasis id="B901D115C24CFFAA0FD61A8FFDE962DE" box="[403,536,1485,1508]" italics="true" pageId="21" pageNumber="22">A. lamellata</emphasis>
” sensu
<bibRefCitation id="EFE470F6C24CFFAA0C371A8FFCEA62DE" author="McLean" box="[626,795,1485,1508]" pageId="21" pageNumber="35" refString="McLean, J. H. (1967) West American Scissurellidae. The Veliger, 9, 404 - 410." type="journal article" year="1967">McLean (1967)</bibRefCitation>
</taxonomicName>
from the eastern Pacific as
<taxonomicName id="4C757684C24CFFAA0A161A8FFB4E62DE" ID-CoL="7C92W" box="[1107,1215,1485,1508]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="baxteri">
<emphasis id="B901D115C24CFFAA0A161A8FFB4E62DE" box="[1107,1215,1485,1508]" italics="true" pageId="21" pageNumber="22">A. baxteri</emphasis>
</taxonomicName>
, but did not list the Talmadge record].
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED2194FFC0C611E" blockId="21.[151,1436,1133,1636]" box="[151,1021,1549,1572]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED2194FFC0C611E" box="[151,1021,1549,1572]" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED2194FFEA4611E" ID-CoL="DJBG" box="[151,341,1549,1572]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="crispata">
<emphasis id="B901D115C24CFFAA0ED2194FFEA4611E" box="[151,341,1549,1572]" italics="true" pageId="21" pageNumber="22">Anatoma crispata</emphasis>
</taxonomicName>
(non Fleming, 1828):
<treatmentCitation id="0AD42B16C24CFFAA0C00194FFCFD611E" author="McLean" box="[581,780,1549,1572]" page="17" pageId="21" pageNumber="22" year="1996">
<bibRefCitation id="EFE470F6C24CFFAA0C00194FFD14611E" author="McLean" box="[581,741,1549,1572]" pageId="21" pageNumber="35" refString="McLean, J. H. (1996) Gastropoda, the Prosobranchia. In: Scott, P. H., Blake, J. A. &amp; Lissner, A. L. (Eds), Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel, vol. 9. Santa Barbara Museum of Natural History, Santa Barbara, pp. i - v, 1 - 160." type="journal article" year="1996">McLean, 1996</bibRefCitation>
: 17
</treatmentCitation>
, fig. 1.2 [is
<taxonomicName id="4C757684C24CFFAA0DC9194FFC1F611E" box="[908,1006,1549,1572]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0DC9194FFC1F611E" box="[908,1006,1549,1572]" italics="true" pageId="21" pageNumber="22">T. kelseyi</emphasis>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0ED2196FFE4D615E" blockId="21.[151,1436,1133,1636]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="AB652A29C24CFFAA0ED2196FFE4D615E" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0ED2196FFD3B617E" authority="McLean, 1967: 405" authorityName="McLean" authorityPageNumber="405" authorityYear="1967" box="[151,714,1581,1604]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="crispata" subGenus="Anatoma">
<emphasis id="B901D115C24CFFAA0ED2196FFEFA617E" box="[151,267,1581,1604]" italics="true" pageId="21" pageNumber="22">Scissurella</emphasis>
(
<emphasis id="B901D115C24CFFAA0F59196FFE8D617E" box="[284,380,1581,1604]" italics="true" pageId="21" pageNumber="22">Anatoma</emphasis>
)
<emphasis id="B901D115C24CFFAA0FC9196FFE12617E" box="[396,483,1581,1604]" italics="true" pageId="21" pageNumber="22">crispata</emphasis>
:
<treatmentCitation id="0AD42B16C24CFFAA0FB6196FFD3B617E" author="McLean" box="[499,714,1581,1604]" page="405" pageId="21" pageNumber="22" year="1967">
<bibRefCitation id="EFE470F6C24CFFAA0FB6196FFD65617E" author="McLean" box="[499,660,1581,1604]" pageId="21" pageNumber="35" refString="McLean, J. H. (1967) West American Scissurellidae. The Veliger, 9, 404 - 410." type="journal article" year="1967">McLean, 1967</bibRefCitation>
: 405
</treatmentCitation>
</taxonomicName>
, pl. 56, figs 1–4 [includes
<taxonomicName id="4C757684C24CFFAA0DAE196FFB9E617E" ID-CoL="DJBB" box="[1003,1135,1581,1604]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="chiricova">
<emphasis id="B901D115C24CFFAA0DAE196FFC0A617E" box="[1003,1019,1581,1604]" italics="true" pageId="21" pageNumber="22">A</emphasis>
.
<emphasis id="B901D115C24CFFAA0A4F196FFB9E617E" box="[1034,1135,1581,1604]" italics="true" pageId="21" pageNumber="22">chiricova</emphasis>
</taxonomicName>
and
<emphasis id="B901D115C24CFFAA0AE0196FFAD8617E" box="[1189,1321,1581,1604]" italics="true" pageId="21" pageNumber="22">
<taxonomicName id="4C757684C24CFFAA0AE0196FFAD4617E" ID-CoL="DJC9" box="[1189,1317,1581,1604]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="japonica">A. japonica</taxonomicName>
.
</emphasis>
Illustrated specimen is
<taxonomicName id="4C757684C24CFFAA0F0E190FFE5C615E" box="[331,429,1613,1636]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24CFFAA0F0E190FFE5C615E" box="[331,429,1613,1636]" italics="true" pageId="21" pageNumber="22">T. kelseyi</emphasis>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36F5E8CC24CFFAA0ED219C8FD166000" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BCA0D07C24CFFAA0ED219C8FC6361F0" blockId="21.[151,1437,1674,2036]" pageId="21" pageNumber="22">
<emphasis id="B901D115C24CFFAA0ED219C8FEB5619E" bold="true" box="[151,324,1674,1700]" pageId="21" pageNumber="22">
<typeStatus id="54CEB3A5C24CFFAA0ED219C8FF25619E" box="[151,212,1674,1700]" pageId="21" pageNumber="22">Type</typeStatus>
material
</emphasis>
.
<typeStatus id="54CEB3A5C24CFFAA0F1319C8FE32619E" box="[342,451,1674,1700]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED6495C2C24CFFAA0F9619C8FDDB619E" LSID="urn:lsid:biocol.org:col:34871" box="[467,554,1674,1700]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22">USNM</collectionCode>
181820. Lost:
<bibRefCitation id="EFE470F6C24CFFAA0C9B19C8FC76619E" author="McLean" box="[734,903,1674,1700]" pageId="21" pageNumber="35" refString="McLean, J. H. (1967) West American Scissurellidae. The Veliger, 9, 404 - 410." type="journal article" year="1967">McLean 1967</bibRefCitation>
: 405, Geiger pers. obs. 2. 2004), 5.5 ×
<quantity id="4C8DA0E2C24CFFAA0B1319C8FA6A619E" box="[1366,1435,1674,1700]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="mm" value="6.0">6 mm</quantity>
(H × W).
<typeStatus id="54CEB3A5C24CFFAA0F4C19F2FE9D61F0" box="[265,364,1712,1738]" pageId="21" pageNumber="22" type="neotype">Neotype</typeStatus>
here designated (
<collectionCode id="ED6495C2C24CFFAA0C7F19F2FD5561F0" LSID="urn:lsid:biocol.org:col:34834" box="[570,676,1712,1738]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34834" name="Santa Barbara Museum of Natural History" pageId="21" pageNumber="22">SBMNH</collectionCode>
3215: see remarks).
</paragraph>
<paragraph id="8BCA0D07C24CFFAA0E811997FD166000" blockId="21.[151,1437,1674,2036]" pageId="21" pageNumber="22">
<emphasis id="B901D115C24CFFAA0E811997FE9861D5" bold="true" box="[196,361,1749,1775]" pageId="21" pageNumber="22">
<typeStatus id="54CEB3A5C24CFFAA0E811997FEF261D5" box="[196,259,1749,1775]" pageId="21" pageNumber="22">Type</typeStatus>
locality
</emphasis>
. California at
<collectingCountry id="F3624D97C24CFFAA0C5E1997FDA161D5" box="[539,592,1749,1775]" name="United States of America" pageId="21" pageNumber="22">U.S.</collectingCountry>
Fish Commission Station 4353 [32.700˚N, 117.233˚W], also off San Diego (OD), 639 fms [=
<quantity id="4C8DA0E2C24CFFAA0FFA19B8FDEC602E" box="[447,541,1786,1812]" metricMagnitude="3" metricUnit="m" metricValue="1.169" pageId="21" pageNumber="22" unit="m" value="1169.0">1169 m</quantity>
], 15 mls from Pt. Loma, 47˚W (
<collectionCode id="ED6495C2C24CFFAA0DDF19B8FC00602E" LSID="urn:lsid:biocol.org:col:34871" box="[922,1009,1786,1812]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="21" pageNumber="22">USNM</collectionCode>
label).
<typeStatus id="54CEB3A5C24CFFAA0A0E19B8FB5F602E" box="[1099,1198,1786,1812]" pageId="21" pageNumber="22" type="neotype">Neotype</typeStatus>
locality:
<quantity id="4C8DA0E2C24CFFAA0B5A19B8FA9C602E" box="[1311,1389,1786,1812]" metricMagnitude="2" metricUnit="m" metricValue="1.83" pageId="21" pageNumber="22" unit="m" value="183.0">183 m</quantity>
, off La Jolla, California,
<collectingCountry id="F3624D97C24CFFAA0FCD1862FE326000" box="[392,451,1824,1850]" name="United States of America" pageId="21" pageNumber="22">USA</collectingCountry>
[32.843˚N, 117.392˚W].
</paragraph>
</subSubSection>
<subSubSection id="C36F5E8CC24CFFAA0E811807FBF96065" box="[196,1032,1861,1887]" pageId="21" pageNumber="22" type="etymology">
<paragraph id="8BCA0D07C24CFFAA0E811807FBF96065" blockId="21.[151,1437,1674,2036]" box="[196,1032,1861,1887]" pageId="21" pageNumber="22">
<emphasis id="B901D115C24CFFAA0E811807FEB86065" bold="true" box="[196,329,1861,1887]" pageId="21" pageNumber="22">Etymology</emphasis>
. Named in honor of Prof. F. W. Kelsey, of San Diego (OD).
</paragraph>
</subSubSection>
<subSubSection id="C36F5E8CC24CFFA90E811828FDA3620D" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="8BCA0D07C24CFFA90E811828FD37668D" blockId="21.[151,1437,1674,2036]" lastBlockId="22.[151,1437,152,2007]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="B901D115C24CFFAA0E811828FEAB60BE" bold="true" box="[196,346,1898,1924]" pageId="21" pageNumber="22">Description</emphasis>
. Shell large (to
<quantity id="4C8DA0E2C24CFFAA0C641828FD9D60BE" box="[545,620,1898,1924]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="21" pageNumber="22" unit="mm" value="6.0">6 mm</quantity>
: fide
<bibRefCitation id="EFE470F6C24CFFAA0CF41828FCC760BE" author="Dall" box="[689,822,1898,1924]" pageId="21" pageNumber="34" refString="Dall, W. H. (1905). Some new species of mollusks from California. The Nautilus, 18, 123 - 125." type="journal article" year="1905">Dall, 1905</bibRefCitation>
), trochiform, biconical, white. Protoconch of 0.75 whorls, with reticulate sculpture, apertural varix barely connected to embryonic cap, apertural margin straight. TI of 0.75 whorls, with approximately 38 axial cordlets, one spiral cordlet in position of selenizone, interstices with flocculent sculpture. TII of 3.125 whorls in
<quantity id="4C8DA0E2C24CFFAA0CA41898FCCC60CE" box="[737,829,2010,2036]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="21" pageNumber="22" unit="mm" value="3.2">3.2 mm</quantity>
specimen, space between suture and selenizone as wide as selenizone showing approximately the uppermost three spirals. Shoulder somewhat convex, suture lightly impressed, axials approximately of same density and strength as on TI, approximately 22 equispaced fine spiral lines running over cordlets. Base with slight constriction below selenizone, axials of same strength and density as on shoulder, crossed with approximately 32 equispaced spiral lines, interstices with fine irregular growth lines. Umbilicus very narrow, evenly curving with base, funiculus present. Selenizone slightly above periphery, keels of moderate strength and moderate elevation, growth marks rather weak; slit open with parallel margins. Aperture rounded, adumbilical portion flared, roof overhanging. Operculum corneous, thin, multispiral with central nucleus.
</paragraph>
<paragraph id="8BCA0D07C24FFFA90E811E80FEFD65D8" blockId="22.[151,1437,152,2007]" pageId="22" pageNumber="23">Radula. Rachidian tooth trapezoid, central denticle largest, approximately 6 denticles on each side. Lateral teeth 1–3 similar, apical denticle largest; 3–4 denticles on outer cutting edge; 1–2 denticles on along inner edge, half size of outer denticles. Lateral tooth 4 reduced, hookshaped, apical denticle largest, 3–4 minute denticles on inner edge. Lateral tooth 5 enlarged by elongation, apical denticle largest, approximately 12 denticles along inner edge, approximately 4 denticles along distal half of cusp’s outer edge. Inner marginal teeth with triangular cusp on long shafts, apical denticle largest, approximately 6 denticles on each side. Outer marginal teeth thin, broadened, with many small bristles along edge of cusp. Radular interlock of central field moderate.</paragraph>
<paragraph id="8BCA0D07C24FFFA90E811DAFFAF464A6" blockId="22.[151,1437,152,2007]" pageId="22" pageNumber="23">Body. Tentacles in anterior-posterior order: cephalic tentacles papillate; ocular stalk without pigmented eye; two distinct, non-papillate tentacles in neck region in anterior-posterior alignment with cephalic tentacle and first epipodial tentacle; bifid papillate epipodial tentacles with single posterior blunt, stubby non-papillate structure about half as long as second epipodial tentacle, most likely elongated epipodial sense organ (ESO); two single papillate epipodial tentacles. Operculum attached to discrete fine flap of metapodium.</paragraph>
<paragraph id="8BCA0D07C24FFFA90E811CEAFDD76336" blockId="22.[151,1437,152,2007]" pageId="22" pageNumber="23">Mantle with papillate tentacle at posterior junction of slit near anal papilla. Two monopectinate gills, approximately 25–30 gill leaflets each, distinct bottle-shaped bursicles in distal half of each filament, opening of bursicles in middle of filament.</paragraph>
<paragraph id="8BCA0D07C24FFFA90E811B55FDA3620D" blockId="22.[151,1437,152,2007]" pageId="22" pageNumber="23">
<emphasis id="B901D115C24FFFA90E811B55FE13630B" bold="true" box="[196,482,1047,1073]" pageId="22" pageNumber="23">Differential Diagnosis</emphasis>
.
<taxonomicName id="4C757684C24FFFA90FB71B5AFC7F6308" authority="McLean, 1966" authorityName="McLean" authorityYear="1966" box="[498,910,1048,1074]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="keenae">
<emphasis id="B901D115C24FFFA90FB71B5AFD4F630B" box="[498,702,1048,1073]" italics="true" pageId="22" pageNumber="23">Anatoma keenae</emphasis>
(
<bibRefCitation id="EFE470F6C24FFFA90C8A1B5AFC746308" author="McLean" box="[719,901,1048,1074]" pageId="22" pageNumber="35" refString="McLean, J. H. (1966) West American Prosobranch Gastropoda: Superfamilies Patellacea, Pleurotomariacea and Fissurellacea. Dissertation Stanford University, Stanford. 245 pp., 7 pls." type="book" year="1966">McLean, 1966</bibRefCitation>
)
</taxonomicName>
[=
<taxonomicName id="4C757684C24FFFA90DFE1B5AFAD26308" authority="McLean, 1966" authorityName="McLean" authorityYear="1966" box="[955,1315,1048,1074]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="epicharis">
<emphasis id="B901D115C24FFFA90DFE1B5AFBBE630B" box="[955,1103,1048,1073]" italics="true" pageId="22" pageNumber="23">A. epicharis</emphasis>
(
<bibRefCitation id="EFE470F6C24FFFA90A241B5AFAE86308" author="McLean" box="[1121,1305,1048,1074]" pageId="22" pageNumber="35" refString="McLean, J. H. (1966) West American Prosobranch Gastropoda: Superfamilies Patellacea, Pleurotomariacea and Fissurellacea. Dissertation Stanford University, Stanford. 245 pp., 7 pls." type="book" year="1966">McLean, 1966</bibRefCitation>
)
</taxonomicName>
] from the Panamic province has a protoconch with flocculent sculpture (reticulate in
<taxonomicName id="4C757684C24FFFA90DBA1B7FFB9D636C" box="[1023,1132,1085,1110]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24FFFA90DBA1B7FFB9D636C" box="[1023,1132,1085,1110]" italics="true" pageId="22" pageNumber="23">T. kelseyi</emphasis>
</taxonomicName>
), on TI lacks a spiral cord in the position of the selenizone (present in
<taxonomicName id="4C757684C24FFFA90CDA1B21FCFC6346" box="[671,781,1123,1148]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24FFFA90CDA1B21FCFC6346" box="[671,781,1123,1148]" italics="true" pageId="22" pageNumber="23">T. kelseyi</emphasis>
</taxonomicName>
), and forms small but distinct points at the intersection of spiral and axial cords on TII (no points in
<taxonomicName id="4C757684C24FFFA90CE31BCAFCE0639B" box="[678,785,1160,1185]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24FFFA90CE31BCAFCE0639B" box="[678,785,1160,1185]" italics="true" pageId="22" pageNumber="23">T. kelseyi</emphasis>
</taxonomicName>
).
<taxonomicName id="4C757684C24FFFA90D691BCAFC0D639B" box="[812,1020,1160,1185]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="baxteri">
<emphasis id="B901D115C24FFFA90D691BCAFC0D639B" box="[812,1020,1160,1185]" italics="true" pageId="22" pageNumber="23">Thieleella baxteri</emphasis>
</taxonomicName>
from the northeastern Pacific has a TI of approximately 0.5 whorls (
<quantity id="4C8DA0E2C24FFFA90C5B1BEFFD8263FD" box="[542,627,1197,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" pageId="22" pageNumber="23" unit="in" value="0.75">0.75 in</quantity>
<taxonomicName id="4C757684C24FFFA90C3E1BEFFD1B63FC" box="[635,746,1197,1222]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24FFFA90C3E1BEFFD1B63FC" box="[635,746,1197,1222]" italics="true" pageId="22" pageNumber="23">T. kelseyi</emphasis>
</taxonomicName>
) and axial sculpture consisting of raised lamellae (simple cords in
<taxonomicName id="4C757684C24FFFA90F401B91FE8D63D6" box="[261,380,1235,1260]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24FFFA90F401B91FE8D63D6" box="[261,380,1235,1260]" italics="true" pageId="22" pageNumber="23">T. kelseyi</emphasis>
</taxonomicName>
).
<taxonomicName id="4C757684C24FFFA90FDD1B91FD0A63D6" authority="Berry, 1947" authorityName="Berry" authorityYear="1947" box="[408,763,1234,1260]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="lyra">
<emphasis id="B901D115C24FFFA90FDD1B91FDB463D6" box="[408,581,1235,1260]" italics="true" pageId="22" pageNumber="23">Anatoma lyra</emphasis>
(Berry, 1947)
</taxonomicName>
, from the northeastern Pacific has a protoconch with flocculent sculpture (reticulate in
<taxonomicName id="4C757684C24FFFA90C641BBAFD7B622B" box="[545,650,1272,1297]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24FFFA90C641BBAFD7B622B" box="[545,650,1272,1297]" italics="true" pageId="22" pageNumber="23">T. kelseyi</emphasis>
</taxonomicName>
), overall stronger and less dense sculpture, and a distinct funiculus in the umbilicus (absent in
<taxonomicName id="4C757684C24FFFA90F901A5FFDB3620C" box="[469,578,1309,1334]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24FFFA90F901A5FFDB3620C" box="[469,578,1309,1334]" italics="true" pageId="22" pageNumber="23">T. kelseyi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36F5E8CC24FFFA60E811A00FB7E67C6" lastPageId="25" lastPageNumber="26" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8BCA0D07C24FFFA90E811A00FDB862B8" blockId="22.[151,1437,152,2007]" pageId="22" pageNumber="23">
<emphasis id="B901D115C24FFFA90E811A00FE906266" bold="true" box="[196,353,1346,1372]" pageId="22" pageNumber="23">Distribution</emphasis>
. Queen Charlotte Islands, British
<collectingCountry id="F3624D97C24FFFA90D451A00FC8B6266" box="[768,890,1346,1372]" name="United States of America" pageId="22" pageNumber="23">Columbia</collectingCountry>
,
<collectingCountry id="F3624D97C24FFFA90DC31A00FC126266" box="[902,995,1346,1372]" name="Canada" pageId="22" pageNumber="23">Canada</collectingCountry>
, to South Coronado Island, northern Baja California,
<collectingCountry id="F3624D97C24FFFA90F1C1A2AFE4262B8" box="[345,435,1384,1410]" name="Mexico" pageId="22" pageNumber="23">Mexico</collectingCountry>
(Dall 1921).
</paragraph>
<paragraph id="8BCA0D07C24FFFA90E811ACFFB3062F6" blockId="22.[151,1437,152,2007]" pageId="22" pageNumber="23">
<emphasis id="B901D115C24FFFA90E811ACFFE52629D" bold="true" box="[196,419,1421,1447]" pageId="22" pageNumber="23">Specimen records</emphasis>
.
<collectingCountry id="F3624D97C24FFFA90FF61ACFFE03629D" box="[435,498,1421,1447]" name="United States of America" pageId="22" pageNumber="23">USA</collectingCountry>
: California.
<quantity id="4C8DA0E2C24FFFA90CC01ACFFD10629D" box="[645,737,1421,1447]" metricMagnitude="3" metricUnit="m" metricValue="1.503" pageId="22" pageNumber="23" unit="m" value="1503.0">1503 m</quantity>
,
<collectingCountry id="F3624D97C24FFFA90CAB1ACFFCE2629D" box="[750,787,1421,1447]" name="United States of America" pageId="22" pageNumber="23">US</collectingCountry>
Fish Comm. St. 2923, off San Diego (USNM 209412, 4).
<quantity id="4C8DA0E2C24FFFA90EFB1AF0FE9262F6" box="[190,355,1458,1484]" metricMagnitude="3" metricUnit="m" metricValue="1.1385" metricValueMax="1.244" metricValueMin="1.033" pageId="22" pageNumber="23" unit="m" value="1138.5" valueMax="1244.0" valueMin="1033.0">1033–1244 m</quantity>
,
<quantity id="4C8DA0E2C24FFFA90F2B1AF0FE2362F6" box="[366,466,1458,1484]" metricMagnitude="4" metricUnit="m" metricValue="2.5749504" pageId="22" pageNumber="23" unit="mi" value="16.0">16 miles</quantity>
SW off Point Loma, 32.700˚N, 117.246˚W (USNM
<date id="FFCB2BC7C24FFFA90A781AF0FB6762F6" box="[1085,1174,1458,1484]" pageId="22" pageNumber="23" value="66-11-21">211166</date>
, 4).
</paragraph>
<paragraph id="8BCA0D07C24FFFA90E811A9AFEE561BD" blockId="22.[151,1437,152,2007]" pageId="22" pageNumber="23">
San Diego Trough:
<quantity id="4C8DA0E2C24FFFA90FFF1A9AFD9962C8" box="[442,616,1496,1522]" metricMagnitude="3" metricUnit="m" metricValue="1.206" metricValueMax="1.241" metricValueMin="1.1709999999999998" pageId="22" pageNumber="23" unit="m" value="1206.0" valueMax="1241.0" valueMin="1171.0">1171–1241 m</quantity>
, 32.417˚N, 117.467˚W (SIO M1974, 1: complete).
<quantity id="4C8DA0E2C24FFFA90AAF1A9AFA6962C8" box="[1258,1432,1496,1522]" metricMagnitude="3" metricUnit="m" metricValue="1.194" metricValueMax="1.222" metricValueMin="1.166" pageId="22" pageNumber="23" unit="m" value="1194.0" valueMax="1222.0" valueMin="1166.0">1166–1222 m</quantity>
, 32.450˚N, 117.467˚W (SIO M1991, 1, 5: complete).
<quantity id="4C8DA0E2C24FFFA90D4B1ABFFC47612D" box="[782,950,1533,1559]" metricMagnitude="3" metricUnit="m" metricValue="1.2205" metricValueMax="1.234" metricValueMin="1.207" pageId="22" pageNumber="23" unit="m" value="1220.5" valueMax="1234.0" valueMin="1207.0">1207–1234 m</quantity>
, SIO69-487, 32.450˚N, 117.525˚W (SIO M2070, 3: complete).
<quantity id="4C8DA0E2C24FFFA90FE01960FDBC6106" box="[421,589,1570,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.1995" metricValueMax="1.216" metricValueMin="1.183" pageId="22" pageNumber="23" unit="m" value="1199.5" valueMax="1216.0" valueMin="1183.0">1183–1216 m</quantity>
, R-28, 32.583˚N, 117.558˚W (SIO 2171, 1: complete).
<quantity id="4C8DA0E2C24FFFA90AA81960FABD6106" box="[1261,1356,1570,1596]" metricMagnitude="3" metricUnit="m" metricValue="1.241" pageId="22" pageNumber="23" unit="m" value="1241.0">1241 m</quantity>
, R-42, 32.440˚N, 117.483˚W (SIO M1778, 1: complete).
<quantity id="4C8DA0E2C24FFFA90CA6190AFC776158" box="[739,902,1608,1634]" metricMagnitude="3" metricUnit="m" metricValue="1.22" metricValueMax="1.241" metricValueMin="1.199" pageId="22" pageNumber="23" unit="m" value="1220.0" valueMax="1241.0" valueMin="1199.0">1199–1241 m</quantity>
, R-17, 32.447˚N, 117.467˚W (SIO M1834, 2: complete).
</paragraph>
<paragraph id="8BCA0D07C24FFFA90E8119D0FB736026" blockId="22.[151,1437,152,2007]" pageId="22" pageNumber="23">
San Clemente Island:
<quantity id="4C8DA0E2C24FFFA90F9F19D0FDDC6196" box="[474,557,1682,1708]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="22" pageNumber="23" unit="m" value="390.0">390 m</quantity>
, off Tanner Bank, 32.683˚N, 119.234˚W (LACM 76-313, 6).
<quantity id="4C8DA0E2C24FFFA90B7319D0FA696196" box="[1334,1432,1682,1708]" metricMagnitude="3" metricUnit="m" metricValue="1.047" pageId="22" pageNumber="23" unit="m" value="1047.0">1047 m</quantity>
, 32.585˚N, 118.965˚W (LACM 76-378, 6).
<quantity id="4C8DA0E2C24FFFA90CEA19FAFCF061E8" box="[687,769,1720,1746]" metricMagnitude="2" metricUnit="m" metricValue="9.33" pageId="22" pageNumber="23" unit="m" value="933.0">933 m</quantity>
,
<quantity id="4C8DA0E2C24FFFA90D4B19FAFC8061E8" box="[782,881,1720,1746]" metricMagnitude="4" metricUnit="m" metricValue="4.95677952" pageId="22" pageNumber="23" unit="mi" value="30.8">30.8 mi</quantity>
, 65˚T of
<collectingCountry id="F3624D97C24FFFA90DA319FAFBC161E8" box="[998,1072,1720,1746]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
Point, 32.583˚N, 118.982˚W (LACM 76-377, 2).
<quantity id="4C8DA0E2C24FFFA90FC8199FFE2D61CD" box="[397,476,1757,1783]" metricMagnitude="2" metricUnit="m" metricValue="9.73" pageId="22" pageNumber="23" unit="m" value="973.0">973 m</quantity>
, California,
<collectingCountry id="F3624D97C24FFFA90C36199FFD4361CD" box="[627,690,1757,1783]" name="United States of America" pageId="22" pageNumber="23">USA</collectingCountry>
, 32.569˚N, 118.984˚W (LACM 76-381, 5).
<quantity id="4C8DA0E2C24FFFA90A83199FFAE761CD" box="[1222,1302,1757,1783]" metricMagnitude="2" metricUnit="m" metricValue="2.92" pageId="22" pageNumber="23" unit="m" value="292.0">292 m</quantity>
, 32.566˚N, 119.131˚W (LACM 76-387, 4).
<quantity id="4C8DA0E2C24FFFA90C551840FDAC6026" box="[528,605,1794,1820]" metricMagnitude="2" metricUnit="m" metricValue="2.01" pageId="22" pageNumber="23" unit="m" value="201.0">201 m</quantity>
, 32.772˚N, 118.379˚W (LACM AHF 1082, 1).
</paragraph>
<paragraph id="8BCA0D07C24FFFA90E81186AFD6160B6" blockId="22.[151,1437,152,2007]" pageId="22" pageNumber="23">
Between Tanner Bank and Cortez Bank, 32.450˚N, 119.250˚W (SBMNH 33350, 1).
<quantity id="4C8DA0E2C24FFFA90A9B186AFACE6078" box="[1246,1343,1832,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="22" pageNumber="23" unit="m" value="1200.0">1200 m</quantity>
, Off N. Coronado Island, 32.750˚N, 117.500˚W (USNM 208901, 3; USNM 209054, 3).
<quantity id="4C8DA0E2C24FFFA90AD8180FFB01605D" box="[1181,1264,1869,1895]" metricMagnitude="2" metricUnit="m" metricValue="1.83" pageId="22" pageNumber="23" unit="m" value="183.0">183 m</quantity>
, off La Jolla, 32.8430˚N, 117.392˚W (SBMNH 3215, 1).
</paragraph>
<paragraph id="8BCA0D07C24FFFA80E8118DAFEEC67C6" blockId="22.[151,1437,152,2007]" lastBlockId="23.[151,1437,152,290]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
Santa Rosa Island:
<quantity id="4C8DA0E2C24FFFA90FE618DAFE106088" box="[419,481,1944,1970]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="22" pageNumber="23" unit="m" value="70.0">70 m</quantity>
, 33.531˚N, 119.982˚W (LACM 77-303, 1).
<quantity id="4C8DA0E2C24FFFA90D9A18DAFBDA6088" box="[991,1067,1944,1970]" metricMagnitude="2" metricUnit="m" metricValue="1.15" pageId="22" pageNumber="23" unit="m" value="115.0">115 m</quantity>
, 33.865˚N, 119.917˚W (LACM 75-300, 6).
<quantity id="4C8DA0E2C24FFFA90F6718FFFE9260ED" box="[290,355,1981,2007]" metricMagnitude="1" metricUnit="m" metricValue="6.1" pageId="22" pageNumber="23" unit="m" value="61.0">61 m</quantity>
, 33.865˚N, 120.030˚W (LACM 75-306, 1).
<quantity id="4C8DA0E2C24FFFA90D3518FFFC4E60ED" box="[880,959,1981,2007]" metricMagnitude="2" metricUnit="m" metricValue="1.05" pageId="22" pageNumber="23" unit="m" value="105.0">105 m</quantity>
, 33.934˚N, 119.901˚W (LACM 75-480, 1).
<quantity id="4C8DA0E2C24EFFA80E871FDAFEE56788" box="[194,276,152,178]" metricMagnitude="2" metricUnit="m" metricValue="4.42" pageId="23" pageNumber="24" unit="m" value="442.0">442 m</quantity>
, 33.884˚N, 120.316˚W (LACM 76-213, 8).
<quantity id="4C8DA0E2C24EFFA80D7B1FDAFC616788" box="[830,912,152,178]" metricMagnitude="2" metricUnit="m" metricValue="4.21" pageId="23" pageNumber="24" unit="m" value="421.0">421 m</quantity>
, 33.850˚N, 120.183˚W (LACM 23129, 2: complete).
<quantity id="4C8DA0E2C24EFFA80F671FFFFE8267ED" box="[290,371,189,215]" metricMagnitude="2" metricUnit="m" metricValue="4.92" pageId="23" pageNumber="24" unit="m" value="492.0">492 m</quantity>
, Off Point Sal, California,
<collectingCountry id="F3624D97C24EFFA80C841FFFFCF067ED" box="[705,769,189,215]" name="United States of America" pageId="23" pageNumber="24">USA</collectingCountry>
, 34.970˚N, 121.298˚W (SBMNH 350392, 1; SBMNH 350392, 2).
</paragraph>
<paragraph id="8BCA0D07C24EFFA80E811E4AFC3D6618" blockId="23.[151,1437,152,290]" box="[196,972,264,290]" pageId="23" pageNumber="24">
<collectingCountry id="F3624D97C24EFFA80E811E4AFEF26618" box="[196,259,264,290]" name="United States of America" pageId="23" pageNumber="24">USA</collectingCountry>
: Oregon.
<quantity id="4C8DA0E2C24EFFA80F301E4AFE216618" box="[373,464,264,290]" metricMagnitude="3" metricUnit="m" metricValue="1.58" pageId="23" pageNumber="24" unit="m" value="1580.0">1580 m</quantity>
, 45.828˚N, 125.233˚W (LACM 67-167, 1).
</paragraph>
<caption id="DF0A5D8FC24EFFA80ED2180FFE7460FE" ID-DOI="http://doi.org/10.5281/zenodo.3558592" ID-Zenodo-Dep="3558592" httpUri="https://zenodo.org/record/3558592/files/figure.png" pageId="23" pageNumber="24" targetBox="[151,1436,335,1856]" targetPageId="23">
<paragraph id="8BCA0D07C24EFFA80ED2180FFE7460FE" blockId="23.[151,1437,1869,1988]" pageId="23" pageNumber="24">
<emphasis id="B901D115C24EFFA80ED2180FFEC0605F" bold="true" box="[151,305,1869,1893]" pageId="23" pageNumber="24">FIGURE 16.</emphasis>
<taxonomicName id="4C757684C24EFFA80F7D180FFDF2605E" box="[312,515,1869,1892]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C24EFFA80F7D180FFDF2605E" box="[312,515,1869,1892]" italics="true" pageId="23" pageNumber="24">Thieleella kelseyi</emphasis>
</taxonomicName>
. A. Neotype, here designated. 183 m, off La Jolla, California, USA, 32.843˚N, 117.392˚W (SBMNH 3215). Radula shown in Figure 13A–C. B. Between Tanner Bank and Cortez Bank, California, USA, 32.450˚N, 119.250˚W (SBMNH 33350). Radula shown in Figure 13D–E. Scale bar shell = 1 mm. Scale bar protoconch = 100 µm.
</paragraph>
</caption>
<caption id="DF0A5D8FC241FFA70ED21B12FE7863FD" ID-DOI="http://doi.org/10.5281/zenodo.3558594" ID-Zenodo-Dep="3558594" httpUri="https://zenodo.org/record/3558594/files/figure.png" pageId="24" pageNumber="25" targetBox="[151,1435,194,1081]" targetPageId="24">
<paragraph id="8BCA0D07C241FFA70ED21B12FE7863FD" blockId="24.[151,1436,1104,1223]" pageId="24" pageNumber="25">
<emphasis id="B901D115C241FFA70ED21B12FED96352" bold="true" box="[151,296,1104,1128]" pageId="24" pageNumber="25">FIGURE 17.</emphasis>
Radula of
<taxonomicName id="4C757684C241FFA70FE71B12FDAE635D" box="[418,607,1104,1127]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="24" pageNumber="25" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C241FFA70FE71B12FDAE635D" box="[418,607,1104,1127]" italics="true" pageId="24" pageNumber="25">Thieleella kelseyi</emphasis>
</taxonomicName>
. A–C. From here designated neotype, 183 m, off La Jolla, California, USA, 32.843˚N, 117.392˚W (SBMNH 3215). For shell see Figure 12 A. D–E. Between Tanner Bank and Cortez Bank, California, USA, 32.450˚N, 119.250˚W (SBMNH 33350). For shell see Figure 12B. Scale bars A, D = 500 µm. Scale bars B, C, E. = 50 µm.
</paragraph>
</caption>
<paragraph id="8BCA0D07C241FFA70E811BB7FE9E629F" blockId="24.[151,1437,1269,2005]" pageId="24" pageNumber="25">
<collectingCountry id="F3624D97C241FFA70E811BB7FED36235" box="[196,290,1269,1295]" name="Canada" pageId="24" pageNumber="25">Canada</collectingCountry>
: British
<collectingCountry id="F3624D97C241FFA70FC31BB7FE0A6235" box="[390,507,1269,1295]" name="United States of America" pageId="24" pageNumber="25">Columbia</collectingCountry>
.
<quantity id="4C8DA0E2C241FFA70C4F1BB7FD996235" box="[522,616,1269,1295]" metricMagnitude="3" metricUnit="m" metricValue="2.1759999999999997" pageId="24" pageNumber="25" unit="m" value="2176.0">2176 m</quantity>
, Estevan Point Light, Hesquiat Peninsula, Noetka Sound, Vancouver Island, 48.635˚N, 126.967˚W (LACM 71-374.7, 3: complete).
<quantity id="4C8DA0E2C241FFA70D301A59FC21620F" box="[885,976,1307,1333]" metricMagnitude="3" metricUnit="m" metricValue="1.6019999999999999" pageId="24" pageNumber="25" unit="m" value="1602.0">1602 m</quantity>
, off Queen Charlotte Sound, 51.500˚N, 129.000˚W (USNM 206690, 1).
<quantity id="4C8DA0E2C241FFA70C6C1A02FD9A6260" box="[553,619,1344,1370]" metricMagnitude="1" metricUnit="m" metricValue="5.5" pageId="24" pageNumber="25" unit="m" value="55.0">55 m</quantity>
, Naden Harbor, N side of Graham Island, Queen Charlotte Island, 54.000˚N, 132.667˚W (LACM 66-66, 0).
<quantity id="4C8DA0E2C241FFA70CC41A27FD3F6245" box="[641,718,1381,1407]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="24" pageNumber="25" unit="m" value="260.0">260 m</quantity>
, Dixon Entrance, N of Graham Island, 54.483˚N, 133.200˚W (LACM 67-28, 2).
</paragraph>
<paragraph id="8BCA0D07C241FFA70E811AF2FEE9612F" blockId="24.[151,1437,1269,2005]" pageId="24" pageNumber="25">
<collectingCountry id="F3624D97C241FFA70E811AF2FEF562F0" box="[196,260,1456,1482]" name="United States of America" pageId="24" pageNumber="25">USA</collectingCountry>
: Alaska.
<quantity id="4C8DA0E2C241FFA70F341AF2FE4162F0" box="[369,432,1456,1482]" metricMagnitude="1" metricUnit="m" metricValue="8.8" pageId="24" pageNumber="25" unit="m" value="88.0">88 m</quantity>
, NW of Gibson Island, off NE side Attu Island, Near Islands, 52.958˚N, 173.237˚W (LACM 86-312, 5).
<quantity id="4C8DA0E2C241FFA70FCF1A97FE2862D5" box="[394,473,1493,1519]" metricMagnitude="2" metricUnit="m" metricValue="1.7899999999999998" pageId="24" pageNumber="25" unit="m" value="179.0">179 m</quantity>
, S of Coronation Island, W of N end Dall
<collectingCountry id="F3624D97C241FFA70D931A97FBD062D5" box="[982,1057,1493,1519]" name="Iceland" pageId="24" pageNumber="25">Island</collectingCountry>
, 55.143˚N, 134.210˚W (LACM 87-377, 2).
</paragraph>
<paragraph id="8BCA0D07C241FFA70E811962FE7C6005" blockId="24.[151,1437,1269,2005]" pageId="24" pageNumber="25">
Baranof Island:
<quantity id="4C8DA0E2C241FFA70FC21962FE276100" box="[391,470,1568,1594]" metricMagnitude="2" metricUnit="m" metricValue="2.78" pageId="24" pageNumber="25" unit="m" value="278.0">278 m</quantity>
, W of gulf of Esquibel, S of Byron Bay, 55.537˚N, 134.891˚W (LACM 87-369, 27).
<quantity id="4C8DA0E2C241FFA70E891907FEE86165" box="[204,281,1605,1631]" metricMagnitude="2" metricUnit="m" metricValue="2.05" pageId="24" pageNumber="25" unit="m" value="205.0">205 m</quantity>
, W of N end
<collectingCountry id="F3624D97C241FFA70FF71907FDBB6165" box="[434,586,1605,1631]" name="United States Pacific Island Wildlife Refuges" pageId="24" pageNumber="25">Baker Island</collectingCountry>
, S of Cape Ommany, 55.394˚N, 134.675˚W (LACM 87-370, 20).
<quantity id="4C8DA0E2C241FFA70B0E1907FA696165" box="[1355,1432,1605,1631]" metricMagnitude="2" metricUnit="m" metricValue="4.43" pageId="24" pageNumber="25" unit="m" value="443.0">443 m</quantity>
, W of Nation Point, Coronation Island, SW of Cape Ommaney, 55.925˚N, 135.438˚W (LACM 87-365, 8).
<quantity id="4C8DA0E2C241FFA70B351929FF416190" metricMagnitude="2" metricUnit="m" metricValue="2.52" pageId="24" pageNumber="25" unit="m" value="252.0">252 m</quantity>
, SW of Cape Ommaney, S end, 56.000˚N, 135.000˚W (LACM 87-357, 2).
<quantity id="4C8DA0E2C241FFA70A6819D2FB8A6190" box="[1069,1147,1680,1706]" metricMagnitude="2" metricUnit="m" metricValue="3.6399999999999997" pageId="24" pageNumber="25" unit="m" value="364.0">364 m</quantity>
, W of Pt. Howard, Kulu Island, S of Sitka, 56.037˚N, 135.307˚W (LACM 87-364, 13).
<quantity id="4C8DA0E2C241FFA70DCC19F7FC2961F5" box="[905,984,1717,1743]" metricMagnitude="2" metricUnit="m" metricValue="3.62" pageId="24" pageNumber="25" unit="m" value="362.0">362 m</quantity>
, W of Pt. Crowley, Kulu Island, S. of Necker Bay, 56.090˚N, 135.150˚W (LACM 87-363, 9).
<quantity id="4C8DA0E2C241FFA70D1E1999FC5C61CF" box="[859,941,1755,1781]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="24" pageNumber="25" unit="m" value="240.0">240 m</quantity>
, S of Kruzof Island, W of Necker Bay, 56.598˚N, 135.782˚W (LACM 87-358, 100). W of Whale Bay, SSW of Biorka Island, 56.514˚N, 135.661˚W (LACM 87-395, 29).
</paragraph>
<paragraph id="8BCA0D07C241FFA60E811809FB7E67C6" blockId="24.[151,1437,1269,2005]" lastBlockId="25.[151,1437,152,663]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<quantity id="4C8DA0E2C241FFA70E811809FEE7605F" box="[196,278,1867,1893]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="24" pageNumber="25" unit="m" value="390.0">390 m</quantity>
, SW of mouth of Sitka Sound, SSW of Cape Edgecumbe, S end of Kruzof Island, 56.778˚N, 135.970˚W (LACM 87-356, 25).
<quantity id="4C8DA0E2C241FFA70C771832FD7260B0" box="[562,643,1904,1930]" metricMagnitude="2" metricUnit="m" metricValue="2.7399999999999998" pageId="24" pageNumber="25" unit="m" value="274.0">274 m</quantity>
, S of Cape Edward, W of Gilmer Bay, Kruzof Island, 57.219˚N, 136.234˚W (LACM 87-354, 17).
<quantity id="4C8DA0E2C241FFA70C7118D7FD746095" box="[564,645,1941,1967]" metricMagnitude="2" metricUnit="m" metricValue="4.08" pageId="24" pageNumber="25" unit="m" value="408.0">408 m</quantity>
, S of Icy Point, W of Point Theodore, Yakobi Island, 57.853˚N, 137.111˚W (LACM 87-346, 1).
<quantity id="4C8DA0E2C241FFA70C5C18F9FD9960EF" box="[537,616,1979,2005]" metricMagnitude="2" metricUnit="m" metricValue="2.1399999999999997" pageId="24" pageNumber="25" unit="m" value="214.0">214 m</quantity>
, SSW of Cape Fairweather, Chichagof Island, W of Cape Bingham, 58.073˚N, 138.476˚W (LACM 87-330, 3).
<quantity id="4C8DA0E2C240FFA60CFA1FDAFCE26788" box="[703,787,152,178]" metricMagnitude="2" metricUnit="m" metricValue="3.84" pageId="25" pageNumber="26" unit="m" value="384.0">384 m</quantity>
, W of Yakobi Island, S of Yakutat Bay, 58.000˚N, 140.000˚W (LACM 87-335, 1).
<quantity id="4C8DA0E2C240FFA60C551FFFFDAC67ED" box="[528,605,189,215]" metricMagnitude="2" metricUnit="m" metricValue="1.78" pageId="25" pageNumber="26" unit="m" value="178.0">178 m</quantity>
, SE of Icy Bay, SW of Yakutat Bay, 59.453˚N, 141.150˚W (LACM 87- 336, 1).
<quantity id="4C8DA0E2C240FFA60EBD1FA0FEB767C6" box="[248,326,226,252]" metricMagnitude="2" metricUnit="m" metricValue="3.53" pageId="25" pageNumber="26" unit="m" value="353.0">353 m</quantity>
, S of Icy Bay, E of Dry Bay, 59.118˚N, 141.495˚E (LACM 87-333, 10).
</paragraph>
</subSubSection>
<subSubSection id="C36F5E8CC240FFA60E811E45FED265AD" pageId="25" pageNumber="26" type="discussion">
<paragraph id="8BCA0D07C240FFA60E811E45FED265AD" blockId="25.[151,1437,152,663]" pageId="25" pageNumber="26">
<emphasis id="B901D115C240FFA60E811E45FEC9661B" bold="true" box="[196,312,263,289]" pageId="25" pageNumber="26">Remarks</emphasis>
. Although well figured by
<bibRefCitation id="EFE470F6C240FFA60CC41E4AFCCE6618" author="McLean" box="[641,831,264,290]" pageId="25" pageNumber="35" refString="McLean, J. H. (1996) Gastropoda, the Prosobranchia. In: Scott, P. H., Blake, J. A. &amp; Lissner, A. L. (Eds), Taxonomic Atlas of the Benthic Fauna of the Santa Maria Basin and Western Santa Barbara Channel, vol. 9. Santa Barbara Museum of Natural History, Santa Barbara, pp. i - v, 1 - 160." type="journal article" year="1996">McLean (1996)</bibRefCitation>
, the identification of this familiar species must be changed, as it is clearly not conspecific with
<taxonomicName id="4C757684C240FFA60CEE1E6FFC02667D" authority="Fleming, 1828" authorityName="Fleming" authorityYear="1828" box="[683,1011,301,327]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="crispata">
<emphasis id="B901D115C240FFA60CEE1E6FFCDD667C" box="[683,812,301,326]" italics="true" pageId="25" pageNumber="26">A. crispata</emphasis>
(Fleming, 1828)
</taxonomicName>
with
<typeStatus id="54CEB3A5C240FFA60A721E6FFB9B667D" box="[1079,1130,301,327]" pageId="25" pageNumber="26">type</typeStatus>
locality in Europe. Given the controversial nature of the name, it is prudent for taxon stabilization to designate a
<typeStatus id="54CEB3A5C240FFA60AE71E10FAF26656" box="[1186,1283,338,364]" pageId="25" pageNumber="26" type="neotype">neotype</typeStatus>
. A specimen collected near to the
<typeStatus id="54CEB3A5C240FFA60FD51E3AFE3266A8" box="[400,451,376,402]" pageId="25" pageNumber="26">type</typeStatus>
locality is here so designated. The specimen agrees in all aspects with the original description, though the latter lacks much specific detail. Specimens in the original
<typeStatus id="54CEB3A5C240FFA60A321EDFFB5B668D" box="[1143,1194,413,439]" pageId="25" pageNumber="26">type</typeStatus>
repository (USNM) originally identified as
<taxonomicName id="4C757684C240FFA60FEC1E80FE0866E1" box="[425,505,450,475]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C240FFA60FEC1E80FE0866E1" box="[425,505,450,475]" italics="true" pageId="25" pageNumber="26">kelseyi</emphasis>
</taxonomicName>
were examined, and the best specimen was imaged by SEM (USNM 209054). Unfortunately, its protoconch is eroded, which makes generic assignment impossible (see
<bibRefCitation id="EFE470F6C240FFA60A9E1EAAFA806538" author="Geiger" box="[1243,1393,488,514]" pageId="25" pageNumber="34" refString="Geiger, D. L. (2003) Phylogenetic assessment of characters proposed for the generic classification of Recent Scissurellidae (Gastropoda: Vetigastropoda) with a description of one new genus and six new species from Easter Island and Australia. Molluscan Research, 23, 21 - 83." type="journal article" year="2003">Geiger 2003</bibRefCitation>
for discussion of genera in
<taxonomicName id="4C757684C240FFA60FEF1D4FFDCE651D" box="[426,575,525,551]" class="Gastropoda" family="Anatomidae" kingdom="Animalia" pageId="25" pageNumber="26" phylum="Mollusca" rank="family">Anatomidae</taxonomicName>
). A specimen with preserved protoconch sculpture, agreeing in all aspects of teleoconch morphology with the USNM
<taxonomicName id="4C757684C240FFA60CFC1D70FCFF6571" box="[697,782,562,587]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C240FFA60CFC1D70FCFF6571" box="[697,782,562,587]" italics="true" pageId="25" pageNumber="26">kelseyi</emphasis>
</taxonomicName>
specimens and obtained from near the original
<typeStatus id="54CEB3A5C240FFA60B221D70FA6D6576" box="[1383,1436,562,588]" pageId="25" pageNumber="26">type</typeStatus>
locality, was chosen instead. The generic placement of
<taxonomicName id="4C757684C240FFA60D721D1AFC7B654B" box="[823,906,600,625]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C240FFA60D721D1AFC7B654B" box="[823,906,600,625]" italics="true" pageId="25" pageNumber="26">kelseyi</emphasis>
</taxonomicName>
follows from the reticulate sculpture of the protoconch.
</paragraph>
</subSubSection>
<caption id="DF0A5D8FC240FFA60ED21A6BFC39629A" ID-DOI="http://doi.org/10.5281/zenodo.3558505" ID-Zenodo-Dep="3558505" httpUri="https://zenodo.org/record/3558505/files/figure.png" pageId="25" pageNumber="26" targetBox="[151,1435,717,1300]" targetPageId="25">
<paragraph id="8BCA0D07C240FFA60ED21A6BFC39629A" blockId="25.[151,1437,1321,1440]" pageId="25" pageNumber="26">
<emphasis id="B901D115C240FFA60ED21A6BFED7627B" bold="true" box="[151,294,1321,1345]" pageId="25" pageNumber="26">FIGURE 18.</emphasis>
Preserved body of
<taxonomicName id="4C757684C240FFA60FB21A68FD40627B" box="[503,689,1322,1345]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C240FFA60FB21A68FD40627B" box="[503,689,1322,1345]" italics="true" pageId="25" pageNumber="26">Thieleella kelseyi</emphasis>
</taxonomicName>
. 421 m, 2.3 miles, 48˚T, to S Pt. Light, Santa Rosa Island, California, 33.850˚N, 120.183˚W (LACM 23129). Stereomicroscope images, z-stacked with
<emphasis id="B901D115C240FFA60A481A08FB55625B" box="[1037,1188,1354,1377]" italics="true" pageId="25" pageNumber="26">HeliconFocus</emphasis>
. Scale bar = 1 mm. A. Ventral view. B. Lateral view. ct: cephalic tentacle; es: eye stalk; et: epipodial tentacle; j: jaws; m: mantle; mp: metapodium; op: operculum; pp: propodium; sn: snout; st: suboptic tentacle.
</paragraph>
</caption>
<subSubSection id="C36F5E8CC240FFA60E811A8DFD9C60BE" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="8BCA0D07C240FFA60E811A8DFAD5619E" blockId="25.[151,1437,1487,2036]" pageId="25" pageNumber="26">
<taxonomicName id="4C757684C240FFA60E811A8DFDC862D3" ID-CoL="DJBB" authority="Dall, 1919" authorityName="Dall" authorityYear="1919" box="[196,569,1487,1513]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="chiricova">
<emphasis id="B901D115C240FFA60E811A8DFE5262D2" box="[196,419,1487,1512]" italics="true" pageId="25" pageNumber="26">Anatoma chiricova</emphasis>
(Dall, 1919)
</taxonomicName>
had been placed in synonymy with the Pacific
<emphasis id="B901D115C240FFA60A271A8DFB8A62D2" box="[1122,1147,1487,1512]" italics="true" pageId="25" pageNumber="26">A.</emphasis>
“
<emphasis id="B901D115C240FFA60ACA1A8DFB0662D2" box="[1167,1271,1487,1512]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="4C757684C240FFA60ACA1A8DFB0362D2" box="[1167,1266,1487,1512]" class="Gastropoda" family="Scissurellidae" genus="Scissurella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="crispata">crispata</taxonomicName>
,
</emphasis>
” under which also
<taxonomicName id="4C757684C240FFA60E951AB6FED26137" ID-CoL="56GHX" box="[208,291,1524,1549]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C240FFA60E951AB6FED26137" box="[208,291,1524,1549]" italics="true" pageId="25" pageNumber="26">kelseyi</emphasis>
</taxonomicName>
was synonymized (
<bibRefCitation id="EFE470F6C240FFA60C501AB6FD346134" author="McLean" box="[533,709,1524,1550]" pageId="25" pageNumber="35" refString="McLean, J. H. (1967) West American Scissurellidae. The Veliger, 9, 404 - 410." type="journal article" year="1967">McLean 1967</bibRefCitation>
). We do not consider
<taxonomicName id="4C757684C240FFA60D941AB6FBB56137" ID-CoL="DJBB" box="[977,1092,1524,1549]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="chiricova">
<emphasis id="B901D115C240FFA60D941AB6FBB56137" box="[977,1092,1524,1549]" italics="true" pageId="25" pageNumber="26">chiricova</emphasis>
</taxonomicName>
a synonym of
<taxonomicName id="4C757684C240FFA60ABE1AB6FABF6137" ID-CoL="56GHX" box="[1275,1358,1524,1549]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C240FFA60ABE1AB6FABF6137" box="[1275,1358,1524,1549]" italics="true" pageId="25" pageNumber="26">kelseyi</emphasis>
</taxonomicName>
either given the more biconical overall shape of the shell of
<taxonomicName id="4C757684C240FFA60D021958FC4F6109" ID-CoL="DJBB" box="[839,958,1562,1587]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="chiricova">
<emphasis id="B901D115C240FFA60D021958FC4F6109" box="[839,958,1562,1587]" italics="true" pageId="25" pageNumber="26">chiricova</emphasis>
</taxonomicName>
. The taxon
<taxonomicName id="4C757684C240FFA60A1D1958FB3E6109" ID-CoL="DJBB" box="[1112,1231,1562,1587]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="chiricova">
<emphasis id="B901D115C240FFA60A1D1958FB3E6109" box="[1112,1231,1562,1587]" italics="true" pageId="25" pageNumber="26">chiricova</emphasis>
</taxonomicName>
requires further investigation. The
<typeStatus id="54CEB3A5C240FFA60F37197DFE546163" box="[370,421,1599,1625]" pageId="25" pageNumber="26">type</typeStatus>
specimen (USNM 206509) was shown by
<bibRefCitation id="EFE470F6C240FFA60DDB197DFB056163" author="McLean" box="[926,1268,1599,1625]" pageId="25" pageNumber="35" refString="McLean, J. H. (1967) West American Scissurellidae. The Veliger, 9, 404 - 410." type="journal article" year="1967">McLean (1967: pl. 56, fig. 2)</bibRefCitation>
as a complete specimen, but has since fragmented most likely due to glass disease (cf. Geiger
<emphasis id="B901D115C240FFA60A081926FB736147" box="[1101,1154,1636,1661]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
. 2007) as evidenced by crystalline precipitations particularly on the largest fragment of the base (Geiger pers. obs. 3/2007).
</paragraph>
<paragraph id="8BCA0D07C240FFA60E8119EDFD9C60BE" blockId="25.[151,1437,1487,2036]" pageId="25" pageNumber="26">
A fully extended preserved body was available for examination of external anatomy by light and scanning electron microscopy. The operculum is not attached to the main portion of the metapodium, but to a rather small tissue flap. The attachment area only occupies approximately a wedge of 60˚, or 1/6 of the total area of the operculum. A similar attachment area is also evident in
<taxonomicName id="4C757684C240FFA60D08185DFC2E6002" ID-CoL="56GHY" box="[845,991,1823,1848]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="peruviana" status="sp. nov.">
<emphasis id="B901D115C240FFA60D08185DFC2E6002" box="[845,991,1823,1848]" italics="true" pageId="25" pageNumber="26">T. peruviana</emphasis>
</taxonomicName>
<emphasis id="B901D115C240FFA60DA2185CFBD86002" bold="true" box="[999,1065,1822,1848]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A2326C6EC240FFA60DA2185CFBD86002" box="[999,1065,1822,1848]" pageId="25" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
and
<emphasis id="B901D115C240FFA60A24185DFB826002" box="[1121,1139,1823,1848]" italics="true" pageId="25" pageNumber="26">A</emphasis>
.
<emphasis id="B901D115C240FFA60AC4185CFB326002" bold="true" box="[1153,1219,1822,1848]" pageId="25" pageNumber="26">n. sp.</emphasis>
of Sasaki
<emphasis id="B901D115C240FFA60B79185DFA816002" box="[1340,1392,1823,1848]" italics="true" pageId="25" pageNumber="26">et al</emphasis>
., in press, from which the operculum had been removed. We are unaware of any other gastropod in which the operculum is attached in such a fashion.
</paragraph>
</subSubSection>
<subSubSection id="C36F5E8CC240FFA40E8118CDFBF967C6" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="description">
<paragraph id="8BCA0D07C240FFA50E8118CDFEF167C6" blockId="25.[151,1437,1487,2036]" lastBlockId="26.[151,1437,152,439]" lastPageId="26" lastPageNumber="27" pageId="25" pageNumber="26">
The two smooth tentacles located between the cephalic tentacle and the first epipodial tentacles are most likely homologous to what
<bibRefCitation id="EFE470F6C240FFA60FA918F6FD6760F4" author="Quinn" box="[492,662,1972,1998]" pageId="25" pageNumber="35" refString="Quinn, J. F. Jr. (1983) A revision of the Seguenziacea Verrill, 1884 (Gastropoda: Prosobranchia). I. Summary and evaluation of the superfamily. Proceedings of the Biological Society of Washington, 96, 725 - 757." type="journal article" year="1983">Quinn (1983)</bibRefCitation>
,
<bibRefCitation id="EFE470F6C240FFA60CE118F6FCA160F4" author="Sasaki" box="[676,848,1972,1998]" pageId="25" pageNumber="35" refString="Sasaki, T. 1998. Comparative anatomy and phylogeny of the Recent Archaeogastropoda (Mollusca: Gastropoda). The University Museum, University of Tokyo, Bulletin, 38, 1 - 224." type="journal article" year="1998">Sasaki (1998)</bibRefCitation>
and
<bibRefCitation id="EFE470F6C240FFA60DCB18F6FBDA60F4" author="Kano" box="[910,1067,1972,1998]" pageId="25" pageNumber="35" refString="Kano, Y, Chikyu E. &amp; A. Waren (2009) Morphological, ecological and molecular characterization of the enigmatic planispiral snail genus Adeuomphalus (Vetigastropoda: Seguenzoidea). Journal of Molluscan Studies, 75, 397 - 418." type="journal article" year="2009">Kano (2009)</bibRefCitation>
called the accessory cephalic tentacle. A single suboptic tentacle can be found in some species, but it is more closely associated with the eye, and usually significantly smaller. As these accessory cephalic tentacles are paired, and as fertilization is most likely external and no ciliary tracks are associated with them, it is unlikely that they have any penial function.
</paragraph>
<paragraph id="8BCA0D07C243FFA50E811E4AFC2E668D" blockId="26.[151,1437,152,439]" pageId="26" pageNumber="27">
The existence of an epipodial sense organ (ESO) in
<taxonomicName id="4C757684C243FFA50D151E4AFC176618" box="[848,998,264,290]" class="Gastropoda" family="Anatomidae" kingdom="Animalia" pageId="26" pageNumber="27" phylum="Mollusca" rank="family">Anatomidae</taxonomicName>
has been uncertain. Here, the nonpapillate stubby projection associated with the first two epipodial tentacles is interpreted as the ESO. Its nonpapillate, blunt nature agrees with typical ESO, which usually, however, are located ventrally to epipodial tentacles and are only about has high as wide (here 3–4 times as high as wide). Sasaki
<emphasis id="B901D115C243FFA50A981E3AFAE466AB" box="[1245,1301,376,401]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
. (in press) interpreted the homologous structure in
<taxonomicName id="4C757684C243FFA50C291EDFFD27668C" box="[620,726,413,438]" class="Gastropoda" family="Anatomidae" genus="Anatoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="26" pageNumber="27" phylum="Mollusca" rank="genus" status="sp. nov.">
<emphasis id="B901D115C243FFA50C291EDFFD27668C" box="[620,726,413,438]" italics="true" pageId="26" pageNumber="27">Anatoma</emphasis>
</taxonomicName>
<emphasis id="B901D115C243FFA50C981EDFFCD1668D" bold="true" box="[733,800,413,439]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A2326C6EC243FFA50C981EDFFCD1668D" box="[733,800,413,439]" pageId="26" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
also as an ESO.
</paragraph>
<caption id="DF0A5D8FC243FFA50ED219C8FCED60FB" ID-DOI="http://doi.org/10.5281/zenodo.3558596" ID-Zenodo-Dep="3558596" httpUri="https://zenodo.org/record/3558596/files/figure.png" pageId="26" pageNumber="27" targetBox="[151,1435,493,1652]" targetPageId="26">
<paragraph id="8BCA0D07C243FFA50ED219C8FCED60FB" blockId="26.[151,1437,1674,1985]" pageId="26" pageNumber="27">
<emphasis id="B901D115C243FFA50ED219C8FED66198" bold="true" box="[151,295,1674,1698]" pageId="26" pageNumber="27">FIGURE 19.</emphasis>
Preserved body of
<taxonomicName id="4C757684C243FFA50FBF19C8FD44619B" box="[506,693,1674,1697]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="kelseyi">
<emphasis id="B901D115C243FFA50FBF19C8FD44619B" box="[506,693,1674,1697]" italics="true" pageId="26" pageNumber="27">Thieleella kelseyi</emphasis>
</taxonomicName>
. 421 m, 2.3 miles 48˚T to S Pt. Light, Santa Rosa Island, California, 33.850˚ N, 120.183˚ W (LACM 23129). A. Operculum. Scale bar = 1 mm. B. Entire preserved body, hexamethyldisilizane treated. Scale bar = 1 mm. C. Ventral view of anterior portion. Scale bar = 1 mm. D. Ventral view of anterior epipodial tentacles plus ESO. Scale bar = 200 µm. E. Dorsal view of epipodial tentacles and ESO; note biserial papillation of epipodial tentacles. Scale bar = 200 µm. F. Mantle tentacle at posterior end of mantle enlarged. Scale bar = 100 µm. G. Overview of gill and mantle. Fat arrow: direction of view of H. Scale bar = 1 mm. H. Bursicles with openings. Scale bar = 100 µm. Abbreviations: act: accessory cephalic tentacle. b: bursicle. bo: bursicle opening. ct: cephalic tentacle. es: eyestalk. ESO: epipodial sense organ. et: epipodial tentacles. gf: gill filaments. j: jaw. mp: metapodium. mt: mantle tentacle. oaf: operculum attachment flap. op: operculum. opg: opening of propodial gland. pp: propodium. rp: rectal papilla. sn: snout. st. suboptic tentacle.
</paragraph>
</caption>
<paragraph id="8BCA0D07C242FFA40E831FDAFBF967C6" blockId="27.[151,1437,152,252]" pageId="27" pageNumber="28">
The presumably chemosensory bursicles (see Geiger
<emphasis id="B901D115C242FFA40D7C1FDAFC9D678B" box="[825,876,152,177]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
. 2008 for review) were found by
<bibRefCitation id="EFE470F6C242FFA40ABF1FDAFA6C6788" author="Sasaki" box="[1274,1437,152,178]" pageId="27" pageNumber="35" refString="Sasaki, T. 1998. Comparative anatomy and phylogeny of the Recent Archaeogastropoda (Mollusca: Gastropoda). The University Museum, University of Tokyo, Bulletin, 38, 1 - 224." type="journal article" year="1998">Sasaki (1998)</bibRefCitation>
in
<taxonomicName id="4C757684C242FFA40EF01FFFFEB567ED" box="[181,324,189,215]" class="Gastropoda" family="Anatomidae" kingdom="Animalia" pageId="27" pageNumber="28" phylum="Mollusca" rank="family">Anatomidae</taxonomicName>
and their presence in the family is here confirmed. The bursicles are distinct, with their typical bottle shape and the opening to the chamber could be seen in several cases.
</paragraph>
</subSubSection>
</treatment>
</document>
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