249 lines
31 KiB
XML
249 lines
31 KiB
XML
<document id="E8204459C1CB59055DAB94F1FCB811CA" ID-DOI="10.5281/zenodo.275645" ID-GBIF-Dataset="c9d212c2-bfed-4043-971c-b80ab6d7ff9c" ID-ISSN="1175-5326" ID-Zenodo-Dep="275645" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460153538636" checkinUser="plazi" docAuthor="Geiger, Daniel L. & Mclean, James H." docDate="2010" docId="03DCBC11C242FFA10ED21E10FCAD66A8" docLanguage="en" docName="zt02356p035.pdf" docOrigin="Zootaxa 2356" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Thieleella peruviana Geiger & Mclean, 2010, n. sp." docType="treatment" docVersion="11" lastPageNumber="31" masterDocId="FFE5C469C259FFBF0E451F42FFF1673A" masterDocTitle="New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda)" masterLastPageNumber="35" masterPageNumber="1" pageNumber="28" updateTime="1698235584404" updateUser="plazi">
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<mods:mods id="DC6474F41528ADBF685E529B6ACE4076" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0EAEE5D08FFC56F3DC3301D8CAA924D7">
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<mods:title id="71483BEF2C90A660DB422F819BCEDDB0">New species and records of Scissurellidae and Anatomidae from the Americas (Mollusca: Gastropoda: Vetigastropoda)</mods:title>
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<mods:name id="D4C4C97851CB6377A355E3711D8FC6E5" type="personal">
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<mods:roleTerm id="CA859C1CD238B050E4703C8BFDE22DC7">Author</mods:roleTerm>
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<mods:namePart id="0D8EF3F08994E1278FEEF4BF63EB7A08">Geiger, Daniel L.</mods:namePart>
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<mods:name id="06C02C3F345F710A329FACA7BECB7745" type="personal">
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<mods:roleTerm id="56841F56479F8E79A4C6B270A98882EF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="91E8E548B8692E70286AD7083A7D1C9F">Mclean, James H.</mods:namePart>
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<mods:typeOfResource id="BB86AF4678B7660CDFBAB6B6A1183221">text</mods:typeOfResource>
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<mods:title id="682AE42D6B326EB26D80D5FCDBDD58DB">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="19A0266E53AF17E446843619CA8A1C54">
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<mods:date id="C166B41835AB6CA6143B301E1A334E10">2010</mods:date>
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<mods:detail id="E2B138EC9D5228D4D2256383FAA10CC6" type="volume">
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<mods:number id="44EB01203C16CFC0108DC56DAA1C4699">2356</mods:number>
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<mods:start id="11F58FFC9CDF2E1E51C007EEB78D0DB1">1</mods:start>
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<mods:end id="9854B8B473232818C49D0C707FDB70BD">35</mods:end>
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<mods:classification id="88240686506C01AD3BB9D21218C8E337">journal article</mods:classification>
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<mods:identifier id="4B9F34F822D53B036EE71F1A87608CCD" type="DOI">10.5281/zenodo.275645</mods:identifier>
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<mods:identifier id="3D29BC5C984E41311A775C62FF2A45F4" type="GBIF-Dataset">c9d212c2-bfed-4043-971c-b80ab6d7ff9c</mods:identifier>
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<mods:identifier id="5CC0B3E543FDB08D55003DF193261DFF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F2C62735D8E584AAC736473BC5BA1B8C" type="Zenodo-Dep">275645</mods:identifier>
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<treatment id="03DCBC11C242FFA10ED21E10FCAD66A8" ID-DOI="http://doi.org/10.5281/zenodo.6210885" ID-GBIF-Taxon="127654804" ID-Zenodo-Dep="6210885" LSID="urn:lsid:plazi:treatment:03DCBC11C242FFA10ED21E10FCAD66A8" httpUri="http://treatment.plazi.org/id/03DCBC11C242FFA10ED21E10FCAD66A8" lastPageId="30" lastPageNumber="31" pageId="27" pageNumber="28">
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<subSubSection id="C36F5E8CC242FFA40ED21E10FECE66B5" pageId="27" pageNumber="28" type="nomenclature">
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<paragraph id="8BCA0D07C242FFA40ED21E10FE2B6656" blockId="27.[151,474,338,399]" box="[151,474,338,364]" pageId="27" pageNumber="28">
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<heading id="D082BA6BC242FFA40ED21E10FE2B6656" bold="true" box="[151,474,338,364]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
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<emphasis id="B901D115C242FFA40ED21E10FE2B6656" bold="true" box="[151,474,338,364]" pageId="27" pageNumber="28">
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<taxonomicName id="4C757684C242FFA40ED21E10FE616656" ID-CoL="56GHY" box="[151,400,338,364]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="peruviana" status="sp. nov.">
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||
<emphasis id="B901D115C242FFA40ED21E10FE616656" bold="true" box="[151,400,338,364]" italics="true" pageId="27" pageNumber="28">Thieleella peruviana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2326C6EC242FFA40FDD1E10FE2B6656" box="[408,474,338,364]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BCA0D07C242FFA40ED21E37FECE66B5" blockId="27.[151,474,338,399]" box="[151,319,373,399]" pageId="27" pageNumber="28">
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<figureCitation id="134E1182C242FFA40ED21E37FECE66B5" box="[151,319,373,399]" captionStart-0="FIGURE 20" captionStart-1="FIGURE 21" captionStart-2="FIGURE 22" captionStartId-0="27.[151,255,1926,1950]" captionStartId-1="28.[151,259,1798,1822]" captionStartId-2="29.[151,258,597,621]" captionTargetBox-0="[151,1435,679,1901]" captionTargetBox-1="[151,1436,194,1776]" captionTargetBox-2="[151,1435,194,575]" captionTargetId-0="figure@27.[151,1436,679,1901]" captionTargetId-1="figure@28.[151,1436,194,1777]" captionTargetId-2="figure@29.[151,1436,194,576]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="29" captionText-0="FIGURE 20. Holotype of Thieleella peruviana n. sp. Arrows indicate remnant of protoconch sculpture; most of apical protoconch surface eroded. Scale bar shell = 1 mm. Scale bar protoconch = 100 µm." captionText-1="FIGURE 21. Anatomy of Thieleella peruviana n. sp. A. Head-foot complex from left side. Scale bar = 1 mm. B. Cephalic tentacle and suboptic tentacle enlarged; note absence of eye or eyestalk. Scale bar = 100 µm. C. Epipodial tentacles with bi-serial papillae on anterior and posterior margin. Box shows enlarged area shown in D. Scale bar = 100 µm. D. Tip of epipodial tentacle with papillae Scale bar = 20 µm. E. Ventral view of epipodial tentacle and potential epipodial sense organ. Scale bar = 100 µm. F. Mantle tentacles at posterior junction of slit. Scale bar = 100 µm. act: accessory cephalic tentacle. ct: cephalic tentacle. ESO: epipodial sense organ. et: epipodial tentacle. oa: opercular attachment site. sn: snout. st: suboptic tentacle. vm: visceral mass." captionText-2="FIGURE 22. Radula and operculum from holotype of Thieleella peruviana n. sp. A. Overview of entire radula. B. Central field enlarged. D. Series of lateral teeth. D. Operculum. Scale bar A = 100 µm. Scale bars B, C = 10 µm. Scale bar D = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3558598" figureDoi-1="http://doi.org/10.5281/zenodo.3558600" figureDoi-2="http://doi.org/10.5281/zenodo.3558602" httpUri-0="https://zenodo.org/record/3558598/files/figure.png" httpUri-1="https://zenodo.org/record/3558600/files/figure.png" httpUri-2="https://zenodo.org/record/3558602/files/figure.png" pageId="27" pageNumber="28">Figures 20–22</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C36F5E8CC242FFA40ED21EFDFDD06570" pageId="27" pageNumber="28" type="materials_examined">
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<paragraph id="8BCA0D07C242FFA40ED21EFDFDD06570" blockId="27.[151,1436,447,623]" pageId="27" pageNumber="28">
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<emphasis id="B901D115C242FFA40ED21EFDFEBC66E3" bold="true" box="[151,333,447,473]" pageId="27" pageNumber="28">
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<typeStatus id="54CEB3A5C242FFA40ED21EFDFF2366E3" box="[151,210,447,473]" pageId="27" pageNumber="28">Type</typeStatus>
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material.
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</emphasis>
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<materialsCitation id="3B1D075AC242FFA40F101E82FAA866E0" ID-GBIF-Occurrence="2460525619" box="[341,1369,448,474]" collectionCode="UF" pageId="27" pageNumber="28" specimenCode="UF 383672" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54CEB3A5C242FFA40F101E82FE3366E0" box="[341,450,448,474]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
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(
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<specimenCode id="DBD3A57CC242FFA40F971E82FDAD66E0" box="[466,604,448,474]" pageId="27" pageNumber="28">UF 383672</specimenCode>
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, shell, mounted soft parts with radula removed, mounted radula)
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</materialsCitation>
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. Four
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<typeStatus id="54CEB3A5C242FFA40ED21EA7FEF666C5" box="[151,263,485,511]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
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(UF 424420, fluid preserved shells with preserved bodies), from
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<typeStatus id="54CEB3A5C242FFA40A421EA7FBC866C5" box="[1031,1081,485,511]" pageId="27" pageNumber="28">type</typeStatus>
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locality.
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<materialsCitation id="3B1D075AC242FFA40E811D48FDD06570" ID-GBIF-Occurrence="2460525621" country="Peru" elevation="1003" latitude="-7.983" location="approximately 175 km WNW of Trujillo" longitude="-80.617" pageId="27" pageNumber="28" specimenCount="1" typeStatus="holotype">
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<emphasis id="B901D115C242FFA40E811D48FE98651E" bold="true" box="[196,361,522,548]" pageId="27" pageNumber="28">
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<typeStatus id="54CEB3A5C242FFA40E811D48FEF2651E" box="[196,259,522,548]" pageId="27" pageNumber="28">Type</typeStatus>
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locality
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</emphasis>
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.
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<quantity id="4C8DA0E2C242FFA40F3C1D48FDE5651E" box="[377,532,522,548]" metricMagnitude="3" metricUnit="m" metricValue="1.003" metricValueMax="1.015" metricValueMin="0.991" pageId="27" pageNumber="28" unit="m" value="1003.0" valueMax="1015.0" valueMin="991.0">
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<elevation id="0058EA34C242FFA40F3C1D48FDE5651E" box="[377,532,522,548]" metricMagnitude="3" metricUnit="m" metricValue="1.003" metricValueMax="1.015" metricValueMin="0.991" pageId="27" pageNumber="28" unit="m" value="1003.0" valueMax="1015.0" valueMin="991.0">991–1015 m</elevation>
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</quantity>
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, off Northern
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<collectingCountry id="F3624D97C242FFA40C861D48FD0A651E" box="[707,763,522,548]" name="Peru" pageId="27" pageNumber="28">Peru</collectingCountry>
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[
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<location id="8EAA5BDCC242FFA40D4F1D48FAF4651E" LSID="urn:lsid:plazi:treatment:03DCBC11C242FFA10ED21E10FCAD66A8:8EAA5BDCC242FFA40D4F1D48FAF4651E" box="[778,1285,522,548]" country="Peru" name="approximately 175 km WNW of Trujillo" pageId="27" pageNumber="28">
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approximately
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||
<quantity id="4C8DA0E2C242FFA40D801D48FBD0651E" box="[965,1057,522,548]" metricMagnitude="5" metricUnit="m" metricValue="1.75" pageId="27" pageNumber="28" unit="km" value="175.0">175 km</quantity>
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WNW of Trujillo
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</location>
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],
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<location id="8EAA5BDCC242FFA40B5C1D48FF1D6570" LSID="urn:lsid:plazi:treatment:03DCBC11C242FFA10ED21E10FCAD66A8:8EAA5BDCC242FFA40B5C1D48FF1D6570" country="Peru" name="Peru-Chile Trench" pageId="27" pageNumber="28">Peru-Chile Trench</location>
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,
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<collectingCountry id="F3624D97C242FFA40EB21D72FEC16570" box="[247,304,560,586]" name="Peru" pageId="27" pageNumber="28">Peru</collectingCountry>
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,
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<geoCoordinate id="EE416BC0C242FFA40F791D72FE6B6570" box="[316,410,560,586]" pageId="27" pageNumber="28">7.983˚S</geoCoordinate>
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,
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<geoCoordinate id="EE416BC0C242FFA40FE31D72FDEC6570" box="[422,541,560,586]" pageId="27" pageNumber="28">80.617˚W</geoCoordinate>
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.
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C36F5E8CC242FFA40E811D17FCB86555" box="[196,841,597,623]" pageId="27" pageNumber="28" type="etymology">
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<paragraph id="8BCA0D07C242FFA40E811D17FCB86555" blockId="27.[151,1436,447,623]" box="[196,841,597,623]" pageId="27" pageNumber="28">
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<emphasis id="B901D115C242FFA40E811D17FEB86555" bold="true" box="[196,329,597,623]" pageId="27" pageNumber="28">Etymology</emphasis>
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. Named after its provenance from off
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<collectingCountry id="F3624D97C242FFA40D4E1D17FCB46555" box="[779,837,597,623]" name="Peru" pageId="27" pageNumber="28">Peru</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<caption id="DF0A5D8FC242FFA40ED218C4FBE86087" ID-DOI="http://doi.org/10.5281/zenodo.3558598" ID-Zenodo-Dep="3558598" httpUri="https://zenodo.org/record/3558598/files/figure.png" pageId="27" pageNumber="28" targetBox="[151,1435,679,1901]" targetPageId="27">
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<paragraph id="8BCA0D07C242FFA40ED218C4FBE86087" blockId="27.[151,1436,1926,1981]" pageId="27" pageNumber="28">
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<emphasis id="B901D115C242FFA40ED218C4FED660A4" bold="true" box="[151,295,1926,1950]" pageId="27" pageNumber="28">FIGURE 20.</emphasis>
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Holotype of
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<taxonomicName id="4C757684C242FFA40FF218C4FD6760A7" box="[439,662,1926,1949]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="peruviana" status="sp. nov.">
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<emphasis id="B901D115C242FFA40FF218C4FD6760A7" box="[439,662,1926,1949]" italics="true" pageId="27" pageNumber="28">Thieleella peruviana</emphasis>
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</taxonomicName>
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<emphasis id="B901D115C242FFA40CDB18C4FD2B60A4" bold="true" box="[670,730,1926,1950]" pageId="27" pageNumber="28">
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<taxonomicNameLabel id="A2326C6EC242FFA40CDB18C4FD2B60A4" box="[670,730,1926,1950]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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Arrows indicate remnant of protoconch sculpture; most of apical protoconch surface eroded. Scale bar shell = 1 mm. Scale bar protoconch = 100 µm.
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</paragraph>
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</caption>
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<caption id="DF0A5D8FC245FFA30ED21844FCA860E7" ID-DOI="http://doi.org/10.5281/zenodo.3558600" ID-Zenodo-Dep="3558600" httpUri="https://zenodo.org/record/3558600/files/figure.png" pageId="28" pageNumber="29" targetBox="[151,1436,194,1776]" targetPageId="28">
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||
<paragraph id="8BCA0D07C245FFA30ED21844FCA860E7" blockId="28.[151,1437,1798,2013]" pageId="28" pageNumber="29">
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||
<emphasis id="B901D115C245FFA30ED21844FEDC6024" bold="true" box="[151,301,1798,1822]" pageId="28" pageNumber="29">FIGURE 21.</emphasis>
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Anatomy of
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<taxonomicName id="4C757684C245FFA30F811845FD5C6024" box="[452,685,1799,1822]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="peruviana" status="sp. nov.">
|
||
<emphasis id="B901D115C245FFA30F811845FD5C6024" box="[452,685,1799,1822]" italics="true" pageId="28" pageNumber="29">Thieleella peruviana</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B901D115C245FFA30CF31844FD076024" bold="true" box="[694,758,1798,1822]" pageId="28" pageNumber="29">
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||
<taxonomicNameLabel id="A2326C6EC245FFA30CF31844FD076024" box="[694,758,1798,1822]" pageId="28" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
A. Head-foot complex from left side. Scale bar = 1 mm. B. Cephalic tentacle and suboptic tentacle enlarged; note absence of eye or eyestalk. Scale bar = 100 µm. C. Epipodial tentacles with bi-serial papillae on anterior and posterior margin. Box shows enlarged area shown in D. Scale bar = 100 µm. D. Tip of epipodial tentacle with papillae Scale bar = 20 µm. E. Ventral view of epipodial tentacle and potential epipodial sense organ. Scale bar = 100 µm. F. Mantle tentacles at posterior junction of slit. Scale bar = 100 µm. act: accessory cephalic tentacle. ct: cephalic tentacle. ESO: epipodial sense organ. et: epipodial tentacle. oa: opercular attachment site. sn: snout. st: suboptic tentacle. vm: visceral mass.
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</paragraph>
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</caption>
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||
<caption id="DF0A5D8FC244FFA20ED21D17FEC56597" ID-DOI="http://doi.org/10.5281/zenodo.3558602" ID-Zenodo-Dep="3558602" httpUri="https://zenodo.org/record/3558602/files/figure.png" pageId="29" pageNumber="30" targetBox="[151,1435,194,575]" targetPageId="29">
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||
<paragraph id="8BCA0D07C244FFA20ED21D17FEC56597" blockId="29.[151,1436,597,685]" pageId="29" pageNumber="30">
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||
<emphasis id="B901D115C244FFA20ED21D17FEDA6557" bold="true" box="[151,299,597,621]" pageId="29" pageNumber="30">FIGURE 22.</emphasis>
|
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Radula and operculum from holotype of
|
||
<taxonomicName id="4C757684C244FFA20CB91D14FC126557" box="[764,995,598,621]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="peruviana" status="sp. nov.">
|
||
<emphasis id="B901D115C244FFA20CB91D14FC126557" box="[764,995,598,621]" italics="true" pageId="29" pageNumber="30">Thieleella peruviana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901D115C244FFA20DAE1D17FBDB6557" bold="true" box="[1003,1066,597,621]" pageId="29" pageNumber="30">
|
||
<taxonomicNameLabel id="A2326C6EC244FFA20DAE1D17FBDB6557" box="[1003,1066,597,621]" pageId="29" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Overview of entire radula. B. Central field enlarged. D. Series of lateral teeth. D. Operculum. Scale bar A = 100 µm. Scale bars B, C = 10 µm. Scale bar D = 1 mm.
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||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36F5E8CC244FFA10E811D99FCAD66A8" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
|
||
<paragraph id="8BCA0D07C244FFA20E811D99FC03637F" blockId="29.[151,1437,731,2027]" pageId="29" pageNumber="30">
|
||
<emphasis id="B901D115C244FFA20E811D99FEA465CF" bold="true" box="[196,341,731,757]" pageId="29" pageNumber="30">Description</emphasis>
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. Shell medium size for genus (up to 3.0 mm), trochoid globular. Protoconch of 0.75 whorls, reticulate sculpture, (see arrows on
|
||
<figureCitation id="134E1182C244FFA20C1D1C42FD426420" box="[600,691,768,794]" captionStart="FIGURE 20" captionStartId="27.[151,255,1926,1950]" captionTargetBox="[151,1435,679,1901]" captionTargetId="figure@27.[151,1436,679,1901]" captionTargetPageId="27" captionText="FIGURE 20. Holotype of Thieleella peruviana n. sp. Arrows indicate remnant of protoconch sculpture; most of apical protoconch surface eroded. Scale bar shell = 1 mm. Scale bar protoconch = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.3558598" httpUri="https://zenodo.org/record/3558598/files/figure.png" pageId="29" pageNumber="30">Fig. 20</figureCitation>
|
||
) condition of apertural varix and shape of apertural margin unknown. TI of 1 whorl, approximately 42 fine axial cords, spiral cord in position of selenizone. TII of 2.5 whorls, suture impressed, below periphery of previous whorl. Shoulder convex, approximately 43 axial cords on first whorl, spiral lines starting at onset of selenizone, approximately 10 after 1 whorl, 12 at apertural margin of
|
||
<typeStatus id="54CEB3A5C244FFA20F561CD4FE8F648A" box="[275,382,918,944]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
, slightly irregularly spaced between suture and selenizone. Base with slight constriction below selenizone, 1.5 times as many axials cords as on shoulder, approximately 28 spiral lines stronger than on shoulder, forming points at intersection with axials. Umbilicus narrow, weak funiculus. Aperture D-shaped with columellar portion almost straight. Selenizone slightly above periphery; keels moderately strong, moderately elevated; lunules irregular, distinct; slit with parallel margins.
|
||
</paragraph>
|
||
<paragraph id="8BCA0D07C244FFA20E811B12FB886350" blockId="29.[151,1437,731,2027]" box="[196,1145,1104,1130]" pageId="29" pageNumber="30">Operculum covering aperture, corneous, thin, round, multispiral, nucleus central.</paragraph>
|
||
<paragraph id="8BCA0D07C244FFA20E811B34FEFD6271" blockId="29.[151,1437,731,2027]" pageId="29" pageNumber="30">Radula: rachidian tooth triangular, cusp with central denticle larges, 4–5 denticles on each side, decreasing in size laterally. Lateral teeth 1–3 similar, decreasing in size two fold, 4,4,3 denticles on outer edge of cusp, respectively. Lateral tooth 4 reduced, hook-shaped. Lateral tooth 5 enlarged by broadening, cusp with penultimate denticle largest, one smaller one on outer edge, five on inner edge, decreasing in size towards base. Marginal teeth hook-shaped with many small denticles on each side. Radular interlock of central field moderate.</paragraph>
|
||
<paragraph id="8BCA0D07C244FFA20E811A14FB04613F" blockId="29.[151,1437,731,2027]" pageId="29" pageNumber="30">Body pale cream in color. Head with non-papillate cephalic tentacles (ct), no eyestalk or eye, suboptic tentacle (ct) present, accessory cephalic tentacle (act). Four epipodial tentacles (et), biserially papillate on anterior and posterior edge, apical papillary tuft. Possible epipodial sense organ (ESO) on ventral base of second epipodial tentacle [or base of broken secondary ramus of epipodial tentacle?]. Opercular attachment (oa) with scale-like muscle fibres. Two mantle tentacles at posterior convergence of mantle slit.</paragraph>
|
||
<paragraph id="8BCA0D07C244FFA20E811952FEF461FA" blockId="29.[151,1437,731,2027]" pageId="29" pageNumber="30">
|
||
<emphasis id="B901D115C244FFA20E811952FE276110" bold="true" box="[196,470,1552,1578]" pageId="29" pageNumber="30">Differential diagnosis.</emphasis>
|
||
This is the first record of any anatomid from the Peruvian coastline. It differs from
|
||
<taxonomicName id="4C757684C244FFA20ED21974FEF46175" box="[151,261,1590,1615]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="baxteri">
|
||
<emphasis id="B901D115C244FFA20ED21974FEF46175" box="[151,261,1590,1615]" italics="true" pageId="29" pageNumber="30">T. baxteri</emphasis>
|
||
</taxonomicName>
|
||
from the Pacific Northwest by the turreted shape (lenticular in
|
||
<taxonomicName id="4C757684C244FFA20DB11974FB956175" box="[1012,1124,1590,1615]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="baxteri">
|
||
<emphasis id="B901D115C244FFA20DB11974FB956175" box="[1012,1124,1590,1615]" italics="true" pageId="29" pageNumber="30">T. baxteri</emphasis>
|
||
</taxonomicName>
|
||
), TI of 1 whorl (
|
||
<quantity id="4C8DA0E2C244FFA20B6E1974FA8F616A" box="[1323,1406,1590,1616]" metricMagnitude="-2" metricUnit="m" metricValue="1.6764" pageId="29" pageNumber="30" unit="in" value="0.66">0.66 in</quantity>
|
||
<taxonomicName id="4C757684C244FFA20BC31974FF00614E" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="baxteri">
|
||
<emphasis id="B901D115C244FFA20BC31974FF00614E" italics="true" pageId="29" pageNumber="30">T. baxteri</emphasis>
|
||
</taxonomicName>
|
||
), and the less distinct axial cords (raised lamellae in
|
||
<taxonomicName id="4C757684C244FFA20DE41919FBEA614E" box="[929,1051,1627,1652]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="baxteri">
|
||
<emphasis id="B901D115C244FFA20DE41919FBEA614E" box="[929,1051,1627,1652]" italics="true" pageId="29" pageNumber="30">T. baxteri</emphasis>
|
||
</taxonomicName>
|
||
). The comparison is based on examination of the
|
||
<typeStatus id="54CEB3A5C244FFA20F3A19C3FE1661A1" box="[383,487,1665,1691]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C757684C244FFA20C5419C3FD7261A0" box="[529,643,1665,1690]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="baxteri">
|
||
<emphasis id="B901D115C244FFA20C5419C3FD7261A0" box="[529,643,1665,1690]" italics="true" pageId="29" pageNumber="30">T. baxteri</emphasis>
|
||
</taxonomicName>
|
||
by SEM, plus conspecific material as identified by comparison to
|
||
<typeStatus id="54CEB3A5C244FFA20ED219E4FEF361FA" box="[151,258,1702,1728]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCA0D07C244FFA10E811989FE3C67C6" blockId="29.[151,1437,731,2027]" lastBlockId="30.[151,1436,152,402]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C757684C244FFA20E811989FE6361DE" box="[196,402,1739,1764]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="kelseyi">
|
||
<emphasis id="B901D115C244FFA20E811989FE6361DE" box="[196,402,1739,1764]" italics="true" pageId="29" pageNumber="30">Thieleella kelseyi</emphasis>
|
||
</taxonomicName>
|
||
from the Pacific North-West shares the overall trochoid-turreted shape, and the overall sculptural pattern. The difference in TI whorls (
|
||
<quantity id="4C8DA0E2C244FFA20C9119B3FCD86031" box="[724,809,1777,1803]" metricMagnitude="-2" metricUnit="m" metricValue="1.6764" pageId="29" pageNumber="30" unit="in" value="0.66">0.66 in</quantity>
|
||
<taxonomicName id="4C757684C244FFA20D7719B3FC386030" box="[818,969,1777,1802]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="peruviana">
|
||
<emphasis id="B901D115C244FFA20D7719B3FC386030" box="[818,969,1777,1802]" italics="true" pageId="29" pageNumber="30">T. peruviana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4C8DA0E2C244FFA20D9D19B3FBDD6031" box="[984,1068,1777,1803]" metricMagnitude="-2" metricUnit="m" metricValue="1.905" pageId="29" pageNumber="30" unit="in" value="0.75">0.75 in</quantity>
|
||
<taxonomicName id="4C757684C244FFA20A7119B3FB526030" box="[1076,1187,1777,1802]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="kelseyi">
|
||
<emphasis id="B901D115C244FFA20A7119B3FB526030" box="[1076,1187,1777,1802]" italics="true" pageId="29" pageNumber="30">T. kelseyi</emphasis>
|
||
</taxonomicName>
|
||
) is not diagnostic as intraspecific variability is known to be ±0.125 whorls (
|
||
<bibRefCitation id="EFE470F6C244FFA20D531854FC5E600A" author="Geiger" box="[790,943,1814,1840]" pageId="29" pageNumber="34" refString="Geiger, D. L. (2003) Phylogenetic assessment of characters proposed for the generic classification of Recent Scissurellidae (Gastropoda: Vetigastropoda) with a description of one new genus and six new species from Easter Island and Australia. Molluscan Research, 23, 21 - 83." type="journal article" year="2003">Geiger 2003</bibRefCitation>
|
||
). The overall shell shape is less turreted in
|
||
<taxonomicName id="4C757684C244FFA20ED21879FEFC606E" box="[151,269,1851,1876]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="kelseyi">
|
||
<emphasis id="B901D115C244FFA20ED21879FEFC606E" box="[151,269,1851,1876]" italics="true" pageId="29" pageNumber="30">T. kelseyi</emphasis>
|
||
</taxonomicName>
|
||
, with the suture being only one width of the selenizone below the selenizone, whereas in
|
||
<taxonomicName id="4C757684C244FFA20BC01879FEFE6040" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="peruviana">
|
||
<emphasis id="B901D115C244FFA20BC01879FEFE6040" italics="true" pageId="29" pageNumber="30">T. peruviana</emphasis>
|
||
</taxonomicName>
|
||
that space is up to three times as wide. Although the position of the suture descends with growth, the distinction holds particularly for larger specimens (
|
||
<quantity id="4C8DA0E2C244FFA20D0618C4FC61609A" box="[835,912,1926,1952]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="mm" value="3.0">3 mm</quantity>
|
||
). The axials are about twice as dense in
|
||
<taxonomicName id="4C757684C244FFA20BC318C4FF1660FE" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="kelseyi">
|
||
<emphasis id="B901D115C244FFA20BC318C4FF1660FE" italics="true" pageId="29" pageNumber="30">T. kelseyi</emphasis>
|
||
</taxonomicName>
|
||
, particularly towards the apertural margin of large specimens (
|
||
<quantity id="4C8DA0E2C244FFA20D9118E9FBEC60FF" box="[980,1053,1963,1989]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="mm" value="3.0">3 mm</quantity>
|
||
). Furthermore, the radula shows some striking dissimilarities, particularly in the shape of lateral tooth 5. While it is broad with five denticles on inner margin in
|
||
<taxonomicName id="4C757684C247FFA10F311FDAFDF6678B" box="[372,519,152,177]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="peruviana">
|
||
<emphasis id="B901D115C247FFA10F311FDAFDF6678B" box="[372,519,152,177]" italics="true" pageId="30" pageNumber="31">T. peruviana</emphasis>
|
||
</taxonomicName>
|
||
, it is more elongated with approximately 12 denticles on the inner margin in
|
||
<taxonomicName id="4C757684C247FFA10BC31FDAFF1B67EC" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="kelseyi">
|
||
<emphasis id="B901D115C247FFA10BC31FDAFF1B67EC" italics="true" pageId="30" pageNumber="31">T. kelseyi</emphasis>
|
||
</taxonomicName>
|
||
. Although the individual distinctions may each appear slight, in conjunction they allow for a clear separation of the two taxa.
|
||
</paragraph>
|
||
<paragraph id="8BCA0D07C247FFA10E811E4AFCAD66A8" blockId="30.[151,1436,152,402]" pageId="30" pageNumber="31">
|
||
<taxonomicName id="4C757684C247FFA10E811E4AFE2E661B" box="[196,479,264,289]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="bathypacifica" status="sp. nov.">
|
||
<emphasis id="B901D115C247FFA10E811E4AFE2E661B" box="[196,479,264,289]" italics="true" pageId="30" pageNumber="31">Thieleella bathypacifica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B901D115C247FFA10FA21E45FDD8661B" bold="true" box="[487,553,263,289]" pageId="30" pageNumber="31">
|
||
<taxonomicNameLabel id="A2326C6EC247FFA10FA21E45FDD8661B" box="[487,553,263,289]" pageId="30" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
from deep water west of Central
|
||
<collectingCountry id="F3624D97C247FFA10DF71E4AFBEB6618" box="[946,1050,264,290]" name="United States of America" pageId="30" pageNumber="31">America</collectingCountry>
|
||
, has on TI a strong spiral cord in the position of the selenizone where the shell profile shows a distinct angle and has fewer (15 vs. 42) and stronger axial cords on TI, while
|
||
<taxonomicName id="4C757684C247FFA10C581E10FD416651" box="[541,688,338,363]" class="Gastropoda" family="Anatomidae" genus="Thieleella" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepetellida" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="peruviana">
|
||
<emphasis id="B901D115C247FFA10C581E10FD416651" box="[541,688,338,363]" italics="true" pageId="30" pageNumber="31">T. peruviana</emphasis>
|
||
</taxonomicName>
|
||
has only a fine spiral line, does not show an angle in the shell’s profile, and has more (42 vs. 15) and finer axial cords on TI.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |