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<document id="216AC74358E2134DCB9D90FBCDCAF63D" ID-CLB-Dataset="292132" ID-DOI="10.5852/ejt.2024.930.2481" ID-GBIF-Dataset="7dcacff1-4cea-4dc9-8606-064d4308c9c5" ID-ISSN="2118-9773" ID-Zenodo-Dep="10893800" ID-ZooBank="A09EB44B-286F-439A-A970-48F09416584A" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1711701720926" checkinUser="plazi" docAuthor="Stec, Daniel" docDate="2024" docId="03D9AF2DB01C0A6EB266FCFF5F66FE47" docLanguage="en" docName="EJT.2024.930.79-123.pdf" docOrigin="European Journal of Taxonomy 930" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2481/11065" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Macrobiotus mileri Stec 2024, sp. nov." docType="treatment" docUuid="99372E2F-595A-4AB2-8398-21198A2CBD7E" docUuidSource="ZooBank" docVersion="5" lastPageNumber="106" masterDocId="FFE0D755B0120A73B013FFC95A05FFBF" masterDocTitle="Integrative taxonomy supports two new species of Macrobiotus (Tardigrada: Eutardigrada: Macrobiotidae) allowing further discussion on the genus phylogeny" masterLastPageNumber="123" masterPageNumber="79" pageNumber="93" updateTime="1711995707204" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C8EC51E41392336A6D61C44D6ACDC4DC">Integrative taxonomy supports two new species of Macrobiotus (Tardigrada: Eutardigrada: Macrobiotidae) allowing further discussion on the genus phylogeny</mods:title>
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<mods:namePart id="D708AF776AFAA0932DBEAA2CB1E81C23">Stec, Daniel</mods:namePart>
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<mods:affiliation id="C7BEE2F7698C517E3CB64BBE2A0D7CBF">Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Sławkowska 17, 31 - 016 Kraków, Poland</mods:affiliation>
<mods:nameIdentifier id="E7C4372E2E27FF31FC832FD5A01BE937" type="email">daniel.stec@isez.pan.krakow.pl</mods:nameIdentifier>
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<mods:title id="9E5EC776A9DA7805FABDA7E87590C0F3">European Journal of Taxonomy</mods:title>
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<mods:date id="E97709D65D9BC8D3432BE128F79D0B9E">2024</mods:date>
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<subSubSection id="C36A4DB0B01C0A7DB266FCFF59BAFCEE" box="[629,959,822,849]" pageId="14" pageNumber="93" type="nomenclature">
<paragraph id="8BCF1E3BB01C0A7DB266FCFF59BAFCEE" blockId="14.[369,1219,822,914]" box="[629,959,822,849]" pageId="14" pageNumber="93">
<heading id="D087A957B01C0A7DB266FCFF59BAFCEE" box="[629,959,822,849]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="93" reason="2">
<taxonomicName id="4C7065B8B01C0A7DB266FCFF5958FCEE" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[629,861,822,849]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="93" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B01C0A7DB266FCFF5958FCEE" bold="true" box="[629,861,822,849]" italics="true" pageId="14" pageNumber="93">Macrobiotus mileri</emphasis>
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<taxonomicNameLabel id="A2377F52B01C0A7DB377FCFE59BAFCEE" box="[868,959,823,849]" pageId="14" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="134B02BEB01C0A7DB28AFCBE590BFC2E" box="[665,782,887,913]" captionStart-0="Fig" captionStart-1="Fig" captionStart-10="Fig" captionStart-11="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="14.[189,232,1892,1918]" captionStartId-1="16.[189,232,1857,1883]" captionStartId-10="25.[189,232,1857,1883]" captionStartId-11="26.[189,232,1889,1915]" captionStartId-2="17.[189,232,1963,1989]" captionStartId-3="18.[189,232,1293,1319]" captionStartId-4="18.[189,232,1928,1954]" captionStartId-5="19.[189,232,1822,1848]" captionStartId-6="20.[189,232,1348,1374]" captionStartId-7="21.[189,232,1787,1813]" captionStartId-8="22.[189,232,1456,1482]" captionStartId-9="24.[189,232,1822,1848]" captionTargetBox-0="[189,1398,1122,1850]" captionTargetBox-1="[197,1391,265,1813]" captionTargetBox-10="[189,1398,859,1819]" captionTargetBox-11="[189,1398,265,1833]" captionTargetBox-2="[189,1398,659,1922]" captionTargetBox-3="[227,1361,265,1240]" captionTargetBox-4="[194,1394,1471,1871]" captionTargetBox-5="[189,1398,716,1782]" captionTargetBox-6="[189,1398,265,1309]" captionTargetBox-7="[238,1349,265,1746]" captionTargetBox-8="[189,1398,265,1417]" captionTargetBox-9="[189,1398,738,1781]" captionTargetId-0="figure-176@14.[189,1398,1122,1850]" captionTargetId-1="figure-0@16.[197,1391,265,1813]" captionTargetId-10="figure-1@25.[189,1398,859,1819]" captionTargetId-11="figure-0@26.[189,1398,265,1833]" captionTargetId-2="figure-0@17.[189,1398,659,1922]" captionTargetId-3="figure-0@18.[227,1361,265,1240]" captionTargetId-4="figure-1@18.[189,1398,1459,1888]" captionTargetId-5="figure-0@19.[189,1398,716,1782]" captionTargetId-6="figure-1@20.[189,1398,265,1309]" captionTargetId-7="figure-107@21.[238,1349,265,1746]" captionTargetId-8="figure-1@22.[189,1398,265,1417]" captionTargetId-9="figure-0@24.[189,1398,738,1781]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionTargetPageId-10="25" captionTargetPageId-11="26" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="18" captionTargetPageId-5="19" captionTargetPageId-6="20" captionTargetPageId-7="21" captionTargetPageId-8="22" captionTargetPageId-9="24" captionText-0="Fig. 9. Macrobiotus mileri sp. nov., PCM images of habitus and body granulation. A. Dorso-ventral projection (holotype, IL.001.11, ISEA PAS, forma aporata). B. Granulation in the dorsal cuticle (paratype, ISEA PAS, forma aporata). C. Granulation in the dorsal body cuticle (paratype, ISEA PAS, forma porata). Scale bars in μm." captionText-1="Fig. 10. Macrobiotus mileri sp. nov., SEM images of body granulation and cuticular pores (paratype, ISEA PAS, forma aporata). A. Body granulation in the dorsal head region. B. Body granulation in the dorsal central body region. C. Body granulation in the dorsal caudal body region. D. General view of body granulation in the lateral caudal body region. EF. Singular pores in the body cuticle. Scale bars in μm." captionText-10="Fig. 19. Macrobiotus mileri sp. nov., PCM images of the eggs under ×1000 magnification (ISEA PAS). AC. Egg surface. DE. Midsections of egg processes. Empty indented arrowheads indicate pores/ light-refracting dots in the center of the terminal discs, filled flat arrowheads indicate small pores around bases of the egg processes, filled indented arrowheads indicate dark thickenings around bases of the egg processes. Scale bars in μm." captionText-11="Fig. 20. Macrobiotus mileri sp. nov., SEM images of eggs (ISEA PAS). AB. Entire egg. CF. Details of egg processes and the surface between them. Empty indented arrowheads indicate pores in center of terminal discs, filled flat arrowheads indicate small pores around bases of egg processes, filled indented arrowheads indicate thickenings around bases of egg processes. Scale bars in μm." captionText-2="Fig. 11. Macrobiotus mileri sp. nov., PCM images of cuticular pores patches in forma porata (paratype, ISEA PAS). A. Patches II, III and IV. B. Patches IV and V. Scale bars in μm." captionText-3="Fig. 12. Macrobiotus mileri sp. nov., SEM images of cuticular pores patches in two paratypes of forma porata (ISEA PAS). A, C. Patches II, III and IV. B, D. Dorsal view on the caudal body region with continuous patch V of pores. Scale bars in μm." captionText-4="Fig. 13. Macrobiotus mileri sp. nov., a schematic drawing of a specimen belonging to forma porata, showing the distribution of patches of cuticular pores, body and leg granulation as well as the gartershaped structures on legs IIII." captionText-5="Fig. 14. Macrobiotus mileri sp. nov., PCM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation on the external, proximal and internal surface of leg II. B. Garter-shaped structure and granulation on the external surface of leg II. C. Granulation on the internal surface of leg III. D. Granulation on the hind leg. The empty arrow indicates a cuticular pore, filled arrows indicate the garter-shaped structure. All photographs taken from specimens belonging to forma aporata; scale bars in μm." captionText-6="Fig. 15. Macrobiotus mileri sp. nov., SEM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation and garter-shaped structure on the external surface of leg I (forma porata). B. Granulation and garter-shaped structure on the external surface of leg II (forma aporata). C. Granulation on the internal surface of leg II (forma porata). D. Granulation on the hind leg (forma porata). Empty arrows indicate cuticular pores, filled arrows indicate the garter-shaped structure, empty flat arrowheads indicate the small cuticular bulge / fold, filled flat arrowheads indicate small pores in between granulation that are visible only under SEM. Scale bars in μm." captionText-7="Fig. 16. Macrobiotus mileri sp. nov., images of claws (paratypes, ISEA PAS). A. Claws II with smooth lunulae (PCM, forma aporata). B. Claws IV with dentate lunulae (PCM, forma aporata). C. Dentate lunulae (PCM, forma aporata). D. Claws II with smooth lunulae, respectively (SEM, forma porata). E. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm." captionText-8="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." captionText-9="Fig. 18. Macrobiotus mileri sp. nov., mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS). AB. The mouth opening of a single paratype visible from lateral and frontal view respectively. CD. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm." figureDoi-0="http://doi.org/10.5281/zenodo.10893818" figureDoi-1="http://doi.org/10.5281/zenodo.10893820" figureDoi-10="http://doi.org/10.5281/zenodo.10893838" figureDoi-11="http://doi.org/10.5281/zenodo.10893840" figureDoi-2="http://doi.org/10.5281/zenodo.10893822" figureDoi-3="http://doi.org/10.5281/zenodo.10893824" figureDoi-4="http://doi.org/10.5281/zenodo.10893826" figureDoi-5="http://doi.org/10.5281/zenodo.10893828" figureDoi-6="http://doi.org/10.5281/zenodo.10893830" figureDoi-7="http://doi.org/10.5281/zenodo.10893832" figureDoi-8="http://doi.org/10.5281/zenodo.10893834" figureDoi-9="http://doi.org/10.5281/zenodo.10893836" httpUri-0="https://zenodo.org/record/10893818/files/figure.png" httpUri-1="https://zenodo.org/record/10893820/files/figure.png" httpUri-10="https://zenodo.org/record/10893838/files/figure.png" httpUri-11="https://zenodo.org/record/10893840/files/figure.png" httpUri-2="https://zenodo.org/record/10893822/files/figure.png" httpUri-3="https://zenodo.org/record/10893824/files/figure.png" httpUri-4="https://zenodo.org/record/10893826/files/figure.png" httpUri-5="https://zenodo.org/record/10893828/files/figure.png" httpUri-6="https://zenodo.org/record/10893830/files/figure.png" httpUri-7="https://zenodo.org/record/10893832/files/figure.png" httpUri-8="https://zenodo.org/record/10893834/files/figure.png" httpUri-9="https://zenodo.org/record/10893836/files/figure.png" pageId="14" pageNumber="93">Figs 920</figureCitation>
,
<tableCitation id="C6F22B80B01C0A7DB308FCBE599EFC2D" box="[795,923,887,914]" captionStart-0="Table 4" captionStart-1="Table 5" captionStartId-0="23.[189,258,265,291]" captionStartId-1="25.[189,258,265,291]" captionTargetPageId-0="23" captionTargetPageId-1="25" captionText-0="Table 4. Measurements [in μm] and pt values of selected morphological structures of animals of Macrobiotus mileri sp. nov.; specimens mounted in Hoyers medium; N: number of specimen/structures measured; range: refers to the smallest and the largest structure among all measured specimens; SD: standard deviation." captionText-1="Table 5. Measurements [in μm] of the eggs of Macrobiotus mileri sp. nov.; eggs mounted in Hoyers medium; process base/height ratio is expressed as percentage; N: number of eggs/structures measured; range: refers to the smallest and the largest structure among all measured specimens; SD: standard deviation." httpUri-0="http://table.plazi.org/id/DF0F4EB3B0050A64B0AEFEC05B98FE33" httpUri-1="http://table.plazi.org/id/DF0F4EB3B00B0A6AB0AEFEC05B37FE33" pageId="14" pageNumber="93" tableUuid-0="DF0F4EB3B0050A64B0AEFEC05B98FE33" tableUuid-1="DF0F4EB3B00B0A6AB0AEFEC05B37FE33">Tables 45</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A4DB0B01C0A7DB0AEFC7358C8FB99" pageId="14" pageNumber="93" type="etymology">
<paragraph id="8BCF1E3BB01C0A7DB0AEFC735B41FC6B" blockId="14.[189,324,954,980]" box="[189,324,954,980]" pageId="14" pageNumber="93">
<heading id="D087A957B01C0A7DB0AEFC735B41FC6B" bold="true" box="[189,324,954,980]" fontSize="11" level="3" pageId="14" pageNumber="93" reason="3">
<emphasis id="B904C229B01C0A7DB0AEFC735B41FC6B" bold="true" box="[189,324,954,980]" pageId="14" pageNumber="93">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1E3BB01C0A7DB0AEFC2058C8FB99" blockId="14.[189,1399,1001,1062]" pageId="14" pageNumber="93">The species is named in honour of Krzysztof Miler, who has developed an impressive tolerance for the daily tardigrade madness that surrounds him.</paragraph>
</subSubSection>
<caption id="DF0F4EB3B01C0A7DB0AEF8AD5832F856" ID-DOI="http://doi.org/10.5281/zenodo.10893818" ID-Zenodo-Dep="10893818" httpUri="https://zenodo.org/record/10893818/files/figure.png" pageId="14" pageNumber="93" startId="14.[189,232,1892,1918]" targetBox="[189,1398,1122,1850]" targetPageId="14">
<paragraph id="8BCF1E3BB01C0A7DB0AEF8AD5832F856" blockId="14.[189,1399,1892,2025]" pageId="14" pageNumber="93">
<emphasis id="B904C229B01C0A7DB0AEF8AD5B0BF8C0" bold="true" box="[189,270,1892,1919]" pageId="14" pageNumber="93">Fig. 9.</emphasis>
<taxonomicName id="4C7065B8B01C0A7DB10BF8AC5BF8F8C0" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[280,509,1893,1919]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="14" pageNumber="93" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B01C0A7DB10BF8AC5BF8F8C0" box="[280,509,1893,1919]" italics="true" pageId="14" pageNumber="93">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B01C0A7DB214F8AC5862F8C0" box="[519,615,1893,1919]" pageId="14" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
, PCM images of habitus and body granulation.
<emphasis id="B904C229B01C0A7DB4BEF8AD5EC7F8C1" bold="true" box="[1197,1218,1892,1918]" pageId="14" pageNumber="93">A</emphasis>
. Dorso-ventral projection (holotype, IL.001.11, ISEA PAS, forma aporata).
<emphasis id="B904C229B01C0A7DB3A6F84159CCF81D" bold="true" box="[949,969,1928,1954]" pageId="14" pageNumber="93">B</emphasis>
. Granulation in the dorsal cuticle (paratype, ISEA PAS, forma aporata).
<emphasis id="B904C229B01C0A7DB296F862589FF87A" bold="true" box="[645,666,1963,1989]" pageId="14" pageNumber="93">C</emphasis>
. Granulation in the dorsal body cuticle (paratype, ISEA PAS, forma porata). Scale bars in μm.
</paragraph>
</caption>
<subSubSection id="C36A4DB0B01D0A7CB0AEFEC1582BFD51" pageId="15" pageNumber="94" type="materials_examined">
<paragraph id="8BCF1E3BB01D0A7CB0AEFEC15BAFFE9D" blockId="15.[189,426,264,290]" box="[189,426,264,290]" pageId="15" pageNumber="94">
<heading id="D087A957B01D0A7CB0AEFEC15BAFFE9D" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="15" pageNumber="94" reason="3">
<emphasis id="B904C229B01D0A7CB0AEFEC15BAFFE9D" bold="true" box="[189,426,264,290]" pageId="15" pageNumber="94">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1E3BB01D0A7CB0AEFEF35977FEC6" blockId="15.[189,1398,313,377]" pageId="15" pageNumber="94">
39 animals,
<specimenCount id="9D76D5B2B01D0A7CB15EFEF35B9FFEEB" box="[333,410,314,340]" count="7" pageId="15" pageNumber="94" type="egg">7 eggs</specimenCount>
mounted on microscope slides in Hoyers medium, 7 animals and
<specimenCount id="9D76D5B2B01D0A7CB4A3FEF35EF8FEEB" box="[1200,1277,314,340]" count="3" pageId="15" pageNumber="94" type="egg">3 eggs</specimenCount>
examined under
<collectionCode id="ED6186FEB01D0A7CB114FE965B43FEC6" box="[263,326,351,377]" pageId="15" pageNumber="94">SEM</collectionCode>
, and 4 animals processed for DNA sequencing.
</paragraph>
<paragraph id="8BCF1E3BB01D0A7CB0AEFE6A5B69FE02" blockId="15.[189,364,419,445]" box="[189,364,419,445]" pageId="15" pageNumber="94">
<heading id="D087A957B01D0A7CB0AEFE6A5B69FE02" bold="true" box="[189,364,419,445]" fontSize="11" level="3" pageId="15" pageNumber="94" reason="3">
<emphasis id="B904C229B01D0A7CB0AEFE6A5B69FE02" bold="true" box="[189,364,419,445]" pageId="15" pageNumber="94">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1E3BB01D0A7CB0F9FE1D5B59FE51" blockId="15.[189,1399,468,569]" box="[234,348,468,494]" pageId="15" pageNumber="94">
<emphasis id="B904C229B01D0A7CB0F9FE1D5B59FE51" bold="true" box="[234,348,468,494]" pageId="15" pageNumber="94">
<typeStatus id="54CBA099B01D0A7CB0F9FE1D5B59FE51" box="[234,348,468,494]" pageId="15" pageNumber="94" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCF1E3BB01D0A7CB0AEFE305966FD86" blockId="15.[189,1399,468,569]" pageId="15" pageNumber="94">
<materialsCitation id="3B181466B01D0A7CB0AEFE305966FD86" collectedFrom="moss growing on a stone wall in urban park" collectingDate="2019-11" collectionCode="ISEA" collectorName="K. Miler" country="Israel" elevation="19" latitude="32.045227" location="Tel-Aviv" longLatPrecision="1" longitude="34.7708" pageId="15" pageNumber="94" specimenCount="1" stateProvince="Tel-Aviv" typeStatus="holotype">
<collectingCountry id="F3675EABB01D0A7CB0AEFE305B26FDAC" box="[189,291,505,531]" name="Israel" pageId="15" pageNumber="94">ISRAEL</collectingCountry>
<collectingRegion id="49B4D0D9B01D0A7CB12BFE305BA6FDAC" box="[312,419,505,531]" country="Israel" name="Tel Aviv" pageId="15" pageNumber="94">Tel-Aviv</collectingRegion>
;
<geoCoordinate id="EE4478FCB01D0A7CB1BDFE31585DFDAB" box="[430,600,504,532]" degrees="32" direction="north" minutes="2" orientation="latitude" pageId="15" pageNumber="94" precision="1" seconds="42.82" value="32.045227">32°242.82″ N</geoCoordinate>
,
<geoCoordinate id="EE4478FCB01D0A7CB270FE31591CFDAB" box="[611,793,504,532]" degrees="34" direction="east" minutes="46" orientation="longitude" pageId="15" pageNumber="94" precision="1" seconds="14.88" value="34.7708">34°4614.88″ E</geoCoordinate>
;
<elevation id="005DF908B01D0A7CB337FE305999FDAC" box="[804,924,505,532]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="15" pageNumber="94" unit="m" value="19.0">
<quantity id="4C88B3DEB01D0A7CB337FE30595BFDAB" box="[804,862,505,532]" metricMagnitude="1" metricUnit="m" metricValue="1.9" pageId="15" pageNumber="94" unit="m" value="19.0">19 m</quantity>
a.s.l.
</elevation>
;
<date id="FFCE38FBB01D0A7CB3BAFE335E20FDAC" box="[937,1061,505,532]" pageId="15" pageNumber="94" value="2019-11">
<collectingDate id="EF8AC113B01D0A7CB3BAFE335E20FDAC" box="[937,1061,505,532]" pageId="15" pageNumber="94" value="2019-11">Nov. 2019</collectingDate>
</date>
;
<collectorName id="26857BEDB01D0A7CB43CFE335E96FDAC" box="[1071,1171,505,532]" pageId="15" pageNumber="94">K. Miler</collectorName>
leg.;
<collectedFrom id="386C5DC6B01D0A7CB4C1FE335810FD86" pageId="15" pageNumber="94">moss growing on a stone wall in urban park</collectedFrom>
;
<collectionCode id="ED6186FEB01D0A7CB232FDD65864FD86" box="[545,609,543,569]" pageId="15" pageNumber="94">ISEA</collectionCode>
PAS, slide IL.001.11.
</materialsCitation>
</paragraph>
<paragraph id="8BCF1E3BB01D0A7CB0F9FDAA5B62FDC2" blockId="15.[189,1399,611,750]" box="[234,359,611,637]" pageId="15" pageNumber="94">
<emphasis id="B904C229B01D0A7CB0F9FDAA5B62FDC2" bold="true" box="[234,359,611,637]" pageId="15" pageNumber="94">
<typeStatus id="54CBA099B01D0A7CB0F9FDAA5B62FDC2" box="[234,359,611,637]" pageId="15" pageNumber="94" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BCF1E3BB01D0A7CB0AEFD41582BFD51" blockId="15.[189,1399,611,750]" pageId="15" pageNumber="94">
<materialsCitation id="3B181466B01D0A7CB0AEFD415BA0FD77" collectedFrom="moss growing on a stone wall in urban park" collectingDate="2019-11" collectionCode="ISEA, PAS" collectorName="K. Miler" country="Israel" elevation="19" latitude="32.045227" location="ISRAEL" longLatPrecision="1" longitude="34.7708" pageId="15" pageNumber="94" specimenCount="45" stateProvince="Tel Aviv" typeStatus="paratype">
<collectingCountry id="F3675EABB01D0A7CB0AEFD415B26FD1D" box="[189,291,648,674]" name="Israel" pageId="15" pageNumber="94">ISRAEL</collectingCountry>
<specimenCount id="9D76D5B2B01D0A7CB12BFD415B50FD1C" box="[312,341,648,675]" pageId="15" pageNumber="94">45</specimenCount>
animals; same collection data as for the holotype;
<collectionCode id="ED6186FEB01D0A7CB38BFD4159DDFD1D" box="[920,984,648,674]" collectionName="Poland, Krakow, Polish Academy of Sciences, Institute of Systematic Zoology" pageId="15" pageNumber="94">ISEA</collectionCode>
<collectionCode id="ED6186FEB01D0A7CB3CDFD415E11FD1D" box="[990,1044,648,674]" collectionName="PAS" pageId="15" pageNumber="94">PAS</collectionCode>
, slides IL.001.08 to IL.001.12, SEM stub TAR.014
</materialsCitation>
<materialsCitation id="3B181466B01D0A7CB1ACFD67582FFD51" collectedFrom="moss growing on a stone wall in urban park" collectingDate="2019-11" collectionCode="ISEA, PAS" collectorName="K. Miler" country="Israel" elevation="19" latitude="32.045227" location="Tel-Aviv" longLatPrecision="1" longitude="34.7708" pageId="15" pageNumber="94" specimenCount="10" specimenCount-egg="10" stateProvince="Tel Aviv" typeStatus="paratype">
<specimenCount id="9D76D5B2B01D0A7CB1ACFD675818FD77" box="[447,541,686,712]" count="10" pageId="15" pageNumber="94" type="egg">10 eggs</specimenCount>
; same collection data as for the holotype;
<collectionCode id="ED6186FEB01D0A7CB403FD675E55FD77" box="[1040,1104,686,712]" collectionName="Poland, Krakow, Polish Academy of Sciences, Institute of Systematic Zoology" pageId="15" pageNumber="94">ISEA</collectionCode>
<collectionCode id="ED6186FEB01D0A7CB444FD675E88FD77" box="[1111,1165,686,712]" collectionName="PAS" pageId="15" pageNumber="94">PAS</collectionCode>
, slides IL.001.06 to IL.001.07, SEM stub TAR.014
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A4DB0B01D0A68B0AEFCDE5E44FE26" lastPageId="27" lastPageNumber="106" pageId="15" pageNumber="94" type="description">
<paragraph id="8BCF1E3BB01D0A7CB0AEFCDE5B4BFC8E" blockId="15.[189,334,791,817]" box="[189,334,791,817]" pageId="15" pageNumber="94">
<heading id="D087A957B01D0A7CB0AEFCDE5B4BFC8E" bold="true" box="[189,334,791,817]" fontSize="11" level="3" pageId="15" pageNumber="94" reason="3">
<emphasis id="B904C229B01D0A7CB0AEFCDE5B4BFC8E" bold="true" box="[189,334,791,817]" pageId="15" pageNumber="94">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1E3BB01D0A7CB0F9FC815B57FCDD" blockId="15.[189,1399,840,1732]" box="[234,338,840,866]" pageId="15" pageNumber="94">
<emphasis id="B904C229B01D0A7CB0F9FC815B57FCDD" bold="true" box="[234,338,840,866]" pageId="15" pageNumber="94">Animals</emphasis>
</paragraph>
<paragraph id="8BCF1E3BB01D0A7CB0AEFCA75BEAF97B" blockId="15.[189,1399,840,1732]" pageId="15" pageNumber="94">
Body transparent in juveniles and white in adults, after fixation in Hoyers medium transparent (
<figureCitation id="134B02BEB01D0A7CB51FFCA75F6FFC37" box="[1292,1386,878,904]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[189,1398,1122,1850]" captionTargetId="figure-176@14.[189,1398,1122,1850]" captionTargetPageId="14" captionText="Fig. 9. Macrobiotus mileri sp. nov., PCM images of habitus and body granulation. A. Dorso-ventral projection (holotype, IL.001.11, ISEA PAS, forma aporata). B. Granulation in the dorsal cuticle (paratype, ISEA PAS, forma aporata). C. Granulation in the dorsal body cuticle (paratype, ISEA PAS, forma porata). Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893818" httpUri="https://zenodo.org/record/10893818/files/figure.png" pageId="15" pageNumber="94">Fig. 9A</figureCitation>
). Eyes present. Granulation is present on the entire body cuticle and is visible under PCM and SEM, but granulation on the ventral side of the body is less dense (
<figureCitation id="134B02BEB01D0A7CB35AFC7059CFFC6C" box="[841,970,953,979]" captionStart="Fig" captionStartId="14.[189,232,1892,1918]" captionTargetBox="[189,1398,1122,1850]" captionTargetId="figure-176@14.[189,1398,1122,1850]" captionTargetPageId="14" captionText="Fig. 9. Macrobiotus mileri sp. nov., PCM images of habitus and body granulation. A. Dorso-ventral projection (holotype, IL.001.11, ISEA PAS, forma aporata). B. Granulation in the dorsal cuticle (paratype, ISEA PAS, forma aporata). C. Granulation in the dorsal body cuticle (paratype, ISEA PAS, forma porata). Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893818" httpUri="https://zenodo.org/record/10893818/files/figure.png" pageId="15" pageNumber="94">Figs 9BC</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB3C5FC705E29FC6C" box="[982,1068,953,979]" captionStart="Fig" captionStartId="16.[189,232,1857,1883]" captionTargetBox="[197,1391,265,1813]" captionTargetId="figure-0@16.[197,1391,265,1813]" captionTargetPageId="16" captionText="Fig. 10. Macrobiotus mileri sp. nov., SEM images of body granulation and cuticular pores (paratype, ISEA PAS, forma aporata). A. Body granulation in the dorsal head region. B. Body granulation in the dorsal central body region. C. Body granulation in the dorsal caudal body region. D. General view of body granulation in the lateral caudal body region. EF. Singular pores in the body cuticle. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893820" httpUri="https://zenodo.org/record/10893820/files/figure.png" pageId="15" pageNumber="94">10AF</figureCitation>
). In terms of cuticular pores, two morphological forms of animals are present in this species. One form (forma porata) with large, evident pores arranged specifically in five patches (
<figureCitation id="134B02BEB01D0A7CB333FBCD59B6FBA1" box="[800,947,1028,1054]" captionStart="Fig" captionStartId="17.[189,232,1963,1989]" captionTargetBox="[189,1398,659,1922]" captionTargetId="figure-0@17.[189,1398,659,1922]" captionTargetPageId="17" captionText="Fig. 11. Macrobiotus mileri sp. nov., PCM images of cuticular pores patches in forma porata (paratype, ISEA PAS). A. Patches II, III and IV. B. Patches IV and V. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893822" httpUri="https://zenodo.org/record/10893822/files/figure.png" pageId="15" pageNumber="94">Figs 11AB</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB3D2FBCD5E1FFBA1" box="[961,1050,1028,1054]" captionStart="Fig" captionStartId="18.[189,232,1293,1319]" captionTargetBox="[227,1361,265,1240]" captionTargetId="figure-0@18.[227,1361,265,1240]" captionTargetPageId="18" captionText="Fig. 12. Macrobiotus mileri sp. nov., SEM images of cuticular pores patches in two paratypes of forma porata (ISEA PAS). A, C. Patches II, III and IV. B, D. Dorsal view on the caudal body region with continuous patch V of pores. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893824" httpUri="https://zenodo.org/record/10893824/files/figure.png" pageId="15" pageNumber="94">12AD</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB434FBCD5E40FBA1" box="[1063,1093,1028,1054]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[194,1394,1471,1871]" captionTargetId="figure-1@18.[189,1398,1459,1888]" captionTargetPageId="18" captionText="Fig. 13. Macrobiotus mileri sp. nov., a schematic drawing of a specimen belonging to forma porata, showing the distribution of patches of cuticular pores, body and leg granulation as well as the gartershaped structures on legs IIII." figureDoi="http://doi.org/10.5281/zenodo.10893826" httpUri="https://zenodo.org/record/10893826/files/figure.png" pageId="15" pageNumber="94">13</figureCitation>
) and second form (forma aporata) with only small, single pores randomly distributed on the body (almost indetectable under PCM and hardly detectable also under SEM; 0.20.4 μm in diameter;
<figureCitation id="134B02BEB01D0A7CB3DFFB865E5DFBD5" box="[972,1112,1103,1130]" captionStart="Fig" captionStartId="16.[189,232,1857,1883]" captionTargetBox="[197,1391,265,1813]" captionTargetId="figure-0@16.[197,1391,265,1813]" captionTargetPageId="16" captionText="Fig. 10. Macrobiotus mileri sp. nov., SEM images of body granulation and cuticular pores (paratype, ISEA PAS, forma aporata). A. Body granulation in the dorsal head region. B. Body granulation in the dorsal central body region. C. Body granulation in the dorsal caudal body region. D. General view of body granulation in the lateral caudal body region. EF. Singular pores in the body cuticle. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893820" httpUri="https://zenodo.org/record/10893820/files/figure.png" pageId="15" pageNumber="94">Figs 10EF</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB474FB995E99FBD5" box="[1127,1180,1104,1130]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[189,1398,716,1782]" captionTargetId="figure-0@19.[189,1398,716,1782]" captionTargetPageId="19" captionText="Fig. 14. Macrobiotus mileri sp. nov., PCM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation on the external, proximal and internal surface of leg II. B. Garter-shaped structure and granulation on the external surface of leg II. C. Granulation on the internal surface of leg III. D. Granulation on the hind leg. The empty arrow indicates a cuticular pore, filled arrows indicate the garter-shaped structure. All photographs taken from specimens belonging to forma aporata; scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893828" httpUri="https://zenodo.org/record/10893828/files/figure.png" pageId="15" pageNumber="94">14A</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB4B8FB865EDBFBD6" box="[1195,1246,1103,1129]" captionStart="Fig" captionStartId="21.[189,232,1787,1813]" captionTargetBox="[238,1349,265,1746]" captionTargetId="figure-107@21.[238,1349,265,1746]" captionTargetPageId="21" captionText="Fig. 16. Macrobiotus mileri sp. nov., images of claws (paratypes, ISEA PAS). A. Claws II with smooth lunulae (PCM, forma aporata). B. Claws IV with dentate lunulae (PCM, forma aporata). C. Dentate lunulae (PCM, forma aporata). D. Claws II with smooth lunulae, respectively (SEM, forma porata). E. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893832" httpUri="https://zenodo.org/record/10893832/files/figure.png" pageId="15" pageNumber="94">16C</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB4FDFB995F42FBD5" box="[1262,1351,1104,1130]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1398,738,1781]" captionTargetId="figure-0@24.[189,1398,738,1781]" captionTargetPageId="24" captionText="Fig. 18. Macrobiotus mileri sp. nov., mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS). AB. The mouth opening of a single paratype visible from lateral and frontal view respectively. CD. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893836" httpUri="https://zenodo.org/record/10893836/files/figure.png" pageId="15" pageNumber="94">18AB</figureCitation>
). In forma porata, the round and oval pores (0.40.7 μm in diameter) are arranged into five distinct patches: (I) a sparse patch of pores on the external surface of the distal portion of leg I (
<figureCitation id="134B02BEB01D0A7CB46CFB525EDEFB0A" box="[1151,1243,1179,1205]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[194,1394,1471,1871]" captionTargetId="figure-1@18.[189,1398,1459,1888]" captionTargetPageId="18" captionText="Fig. 13. Macrobiotus mileri sp. nov., a schematic drawing of a specimen belonging to forma porata, showing the distribution of patches of cuticular pores, body and leg granulation as well as the gartershaped structures on legs IIII." figureDoi="http://doi.org/10.5281/zenodo.10893826" httpUri="https://zenodo.org/record/10893826/files/figure.png" pageId="15" pageNumber="94">Figs 13</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB4F8FB535F24FB0A" box="[1259,1313,1178,1205]" captionStart="Fig" captionStartId="20.[189,232,1348,1374]" captionTargetBox="[189,1398,265,1309]" captionTargetId="figure-1@20.[189,1398,265,1309]" captionTargetPageId="20" captionText="Fig. 15. Macrobiotus mileri sp. nov., SEM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation and garter-shaped structure on the external surface of leg I (forma porata). B. Granulation and garter-shaped structure on the external surface of leg II (forma aporata). C. Granulation on the internal surface of leg II (forma porata). D. Granulation on the hind leg (forma porata). Empty arrows indicate cuticular pores, filled arrows indicate the garter-shaped structure, empty flat arrowheads indicate the small cuticular bulge / fold, filled flat arrowheads indicate small pores in between granulation that are visible only under SEM. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893830" httpUri="https://zenodo.org/record/10893830/files/figure.png" pageId="15" pageNumber="94">15A</figureCitation>
); (II) a dense patch of pores on the external surface of the proximal portion of leg II extending also towards the lateral body cuticle (
<figureCitation id="134B02BEB01D0A7CB1BEFB2F581EFABF" box="[429,539,1254,1280]" captionStart="Fig" captionStartId="17.[189,232,1963,1989]" captionTargetBox="[189,1398,659,1922]" captionTargetId="figure-0@17.[189,1398,659,1922]" captionTargetPageId="17" captionText="Fig. 11. Macrobiotus mileri sp. nov., PCM images of cuticular pores patches in forma porata (paratype, ISEA PAS). A. Patches II, III and IV. B. Patches IV and V. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893822" httpUri="https://zenodo.org/record/10893822/files/figure.png" pageId="15" pageNumber="94">Figs 11A</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB23BFB2F584DFABF" box="[552,584,1254,1280]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[194,1394,1471,1871]" captionTargetId="figure-1@18.[189,1398,1459,1888]" captionTargetPageId="18" captionText="Fig. 13. Macrobiotus mileri sp. nov., a schematic drawing of a specimen belonging to forma porata, showing the distribution of patches of cuticular pores, body and leg granulation as well as the gartershaped structures on legs IIII." figureDoi="http://doi.org/10.5281/zenodo.10893826" httpUri="https://zenodo.org/record/10893826/files/figure.png" pageId="15" pageNumber="94">13</figureCitation>
); (III) a dense patch of pores on the lateral body surface between legs II and II (
<figureCitation id="134B02BEB01D0A7CB13DFAC25B9BFA99" box="[302,414,1291,1318]" captionStart="Fig" captionStartId="17.[189,232,1963,1989]" captionTargetBox="[189,1398,659,1922]" captionTargetId="figure-0@17.[189,1398,659,1922]" captionTargetPageId="17" captionText="Fig. 11. Macrobiotus mileri sp. nov., PCM images of cuticular pores patches in forma porata (paratype, ISEA PAS). A. Patches II, III and IV. B. Patches IV and V. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893822" httpUri="https://zenodo.org/record/10893822/files/figure.png" pageId="15" pageNumber="94">Figs 11A</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB1B8FAC55806FA99" box="[427,515,1292,1318]" captionStart="Fig" captionStartId="18.[189,232,1293,1319]" captionTargetBox="[227,1361,265,1240]" captionTargetId="figure-0@18.[227,1361,265,1240]" captionTargetPageId="18" captionText="Fig. 12. Macrobiotus mileri sp. nov., SEM images of cuticular pores patches in two paratypes of forma porata (ISEA PAS). A, C. Patches II, III and IV. B, D. Dorsal view on the caudal body region with continuous patch V of pores. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893824" httpUri="https://zenodo.org/record/10893824/files/figure.png" pageId="15" pageNumber="94">12A, C</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB21CFAC5582AFA99" box="[527,559,1292,1318]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[194,1394,1471,1871]" captionTargetId="figure-1@18.[189,1398,1459,1888]" captionTargetPageId="18" captionText="Fig. 13. Macrobiotus mileri sp. nov., a schematic drawing of a specimen belonging to forma porata, showing the distribution of patches of cuticular pores, body and leg granulation as well as the gartershaped structures on legs IIII." figureDoi="http://doi.org/10.5281/zenodo.10893826" httpUri="https://zenodo.org/record/10893826/files/figure.png" pageId="15" pageNumber="94">13</figureCitation>
); (IV) a dense large patch of pores covering the whole external surface of leg III, extending also towards the lateral body cuticle (
<figureCitation id="134B02BEB01D0A7CB377FAF859F2FAF3" box="[868,1015,1329,1357]" captionStart="Fig" captionStartId="17.[189,232,1963,1989]" captionTargetBox="[189,1398,659,1922]" captionTargetId="figure-0@17.[189,1398,659,1922]" captionTargetPageId="17" captionText="Fig. 11. Macrobiotus mileri sp. nov., PCM images of cuticular pores patches in forma porata (paratype, ISEA PAS). A. Patches II, III and IV. B. Patches IV and V. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893822" httpUri="https://zenodo.org/record/10893822/files/figure.png" pageId="15" pageNumber="94">
Figs 11A
<emphasis id="B904C229B01D0A7CB3C2FAFB59E4FAF2" box="[977,993,1330,1357]" italics="true" pageId="15" pageNumber="94"></emphasis>
B
</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB410FAF85E5FFAF4" box="[1027,1114,1329,1355]" captionStart="Fig" captionStartId="18.[189,232,1293,1319]" captionTargetBox="[227,1361,265,1240]" captionTargetId="figure-0@18.[227,1361,265,1240]" captionTargetPageId="18" captionText="Fig. 12. Macrobiotus mileri sp. nov., SEM images of cuticular pores patches in two paratypes of forma porata (ISEA PAS). A, C. Patches II, III and IV. B, D. Dorsal view on the caudal body region with continuous patch V of pores. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893824" httpUri="https://zenodo.org/record/10893824/files/figure.png" pageId="15" pageNumber="94">12A, C</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB475FAF85E83FAF4" box="[1126,1158,1329,1355]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[194,1394,1471,1871]" captionTargetId="figure-1@18.[189,1398,1459,1888]" captionTargetPageId="18" captionText="Fig. 13. Macrobiotus mileri sp. nov., a schematic drawing of a specimen belonging to forma porata, showing the distribution of patches of cuticular pores, body and leg granulation as well as the gartershaped structures on legs IIII." figureDoi="http://doi.org/10.5281/zenodo.10893826" httpUri="https://zenodo.org/record/10893826/files/figure.png" pageId="15" pageNumber="94">13</figureCitation>
); and (V) the largest patch of pores that extends from the left caudo-lateral surface, through the dorsal caudal surface to the right caudo-lateral surface, extending also towards lateral and dorsal surfaces of legs IV (
<figureCitation id="134B02BEB01D0A7CB513FAB55F74FA29" box="[1280,1393,1404,1430]" captionStart="Fig" captionStartId="17.[189,232,1963,1989]" captionTargetBox="[189,1398,659,1922]" captionTargetId="figure-0@17.[189,1398,659,1922]" captionTargetPageId="17" captionText="Fig. 11. Macrobiotus mileri sp. nov., PCM images of cuticular pores patches in forma porata (paratype, ISEA PAS). A. Patches II, III and IV. B. Patches IV and V. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893822" httpUri="https://zenodo.org/record/10893822/files/figure.png" pageId="15" pageNumber="94">Figs 11B</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB0AEFA6B5B13FA03" box="[189,278,1442,1468]" captionStart="Fig" captionStartId="18.[189,232,1293,1319]" captionTargetBox="[227,1361,265,1240]" captionTargetId="figure-0@18.[227,1361,265,1240]" captionTargetPageId="18" captionText="Fig. 12. Macrobiotus mileri sp. nov., SEM images of cuticular pores patches in two paratypes of forma porata (ISEA PAS). A, C. Patches II, III and IV. B, D. Dorsal view on the caudal body region with continuous patch V of pores. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893824" httpUri="https://zenodo.org/record/10893824/files/figure.png" pageId="15" pageNumber="94">12AD</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB132FA6B5B45FA03" box="[289,320,1442,1468]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[194,1394,1471,1871]" captionTargetId="figure-1@18.[189,1398,1459,1888]" captionTargetPageId="18" captionText="Fig. 13. Macrobiotus mileri sp. nov., a schematic drawing of a specimen belonging to forma porata, showing the distribution of patches of cuticular pores, body and leg granulation as well as the gartershaped structures on legs IIII." figureDoi="http://doi.org/10.5281/zenodo.10893826" httpUri="https://zenodo.org/record/10893826/files/figure.png" pageId="15" pageNumber="94">13</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB158FA6B5B84FA02" box="[331,385,1442,1469]" captionStart="Fig" captionStartId="20.[189,232,1348,1374]" captionTargetBox="[189,1398,265,1309]" captionTargetId="figure-1@20.[189,1398,265,1309]" captionTargetPageId="20" captionText="Fig. 15. Macrobiotus mileri sp. nov., SEM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation and garter-shaped structure on the external surface of leg I (forma porata). B. Granulation and garter-shaped structure on the external surface of leg II (forma aporata). C. Granulation on the internal surface of leg II (forma porata). D. Granulation on the hind leg (forma porata). Empty arrows indicate cuticular pores, filled arrows indicate the garter-shaped structure, empty flat arrowheads indicate the small cuticular bulge / fold, filled flat arrowheads indicate small pores in between granulation that are visible only under SEM. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893830" httpUri="https://zenodo.org/record/10893830/files/figure.png" pageId="15" pageNumber="94">15D</figureCitation>
). Only the V patch is single and continuous, while patches IIV are doubled and present symmetrically on each side of the body. In both forms, some evident dense granulation patches are visible on the external and internal surfaces of all legs IIII, as well as on the lateral and dorsal surfaces of legs IV under PCM and SEM (
<figureCitation id="134B02BEB01D0A7CB25CF9DB58E0F992" box="[591,741,1554,1581]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[189,1398,716,1782]" captionTargetId="figure-0@19.[189,1398,716,1782]" captionTargetPageId="19" captionText="Fig. 14. Macrobiotus mileri sp. nov., PCM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation on the external, proximal and internal surface of leg II. B. Garter-shaped structure and granulation on the external surface of leg II. C. Granulation on the internal surface of leg III. D. Granulation on the hind leg. The empty arrow indicates a cuticular pore, filled arrows indicate the garter-shaped structure. All photographs taken from specimens belonging to forma aporata; scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893828" httpUri="https://zenodo.org/record/10893828/files/figure.png" pageId="15" pageNumber="94">Figs 14AD</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB2E1F9DB5948F992" box="[754,845,1554,1581]" captionStart="Fig" captionStartId="20.[189,232,1348,1374]" captionTargetBox="[189,1398,265,1309]" captionTargetId="figure-1@20.[189,1398,265,1309]" captionTargetPageId="20" captionText="Fig. 15. Macrobiotus mileri sp. nov., SEM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation and garter-shaped structure on the external surface of leg I (forma porata). B. Granulation and garter-shaped structure on the external surface of leg II (forma aporata). C. Granulation on the internal surface of leg II (forma porata). D. Granulation on the hind leg (forma porata). Empty arrows indicate cuticular pores, filled arrows indicate the garter-shaped structure, empty flat arrowheads indicate the small cuticular bulge / fold, filled flat arrowheads indicate small pores in between granulation that are visible only under SEM. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893830" httpUri="https://zenodo.org/record/10893830/files/figure.png" pageId="15" pageNumber="94">15AD</figureCitation>
). Small pores, visible only under SEM, can be seen in between the granulation on the hind legs (
<figureCitation id="134B02BEB01D0A7CB302F9F15985F9EC" box="[785,896,1592,1619]" captionStart="Fig" captionStartId="20.[189,232,1348,1374]" captionTargetBox="[189,1398,265,1309]" captionTargetId="figure-1@20.[189,1398,265,1309]" captionTargetPageId="20" captionText="Fig. 15. Macrobiotus mileri sp. nov., SEM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation and garter-shaped structure on the external surface of leg I (forma porata). B. Granulation and garter-shaped structure on the external surface of leg II (forma aporata). C. Granulation on the internal surface of leg II (forma porata). D. Granulation on the hind leg (forma porata). Empty arrows indicate cuticular pores, filled arrows indicate the garter-shaped structure, empty flat arrowheads indicate the small cuticular bulge / fold, filled flat arrowheads indicate small pores in between granulation that are visible only under SEM. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893830" httpUri="https://zenodo.org/record/10893830/files/figure.png" pageId="15" pageNumber="94">Fig. 15D</figureCitation>
). A pulvinus-shaped cuticular bulge is not visible on the internal surface of legs IIII, but there is a garter-shaped structure on the external surface of all legs IIII (
<figureCitation id="134B02BEB01D0A7CB16BF94A5BD7F922" box="[376,466,1667,1693]" captionStart="Fig" captionStartId="18.[189,232,1928,1954]" captionTargetBox="[194,1394,1471,1871]" captionTargetId="figure-1@18.[189,1398,1459,1888]" captionTargetPageId="18" captionText="Fig. 13. Macrobiotus mileri sp. nov., a schematic drawing of a specimen belonging to forma porata, showing the distribution of patches of cuticular pores, body and leg granulation as well as the gartershaped structures on legs IIII." figureDoi="http://doi.org/10.5281/zenodo.10893826" httpUri="https://zenodo.org/record/10893826/files/figure.png" pageId="15" pageNumber="94">Figs 13</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB1CEF94A5830F922" box="[477,565,1667,1693]" captionStart="Fig" captionStartId="19.[189,232,1822,1848]" captionTargetBox="[189,1398,716,1782]" captionTargetId="figure-0@19.[189,1398,716,1782]" captionTargetPageId="19" captionText="Fig. 14. Macrobiotus mileri sp. nov., PCM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation on the external, proximal and internal surface of leg II. B. Garter-shaped structure and granulation on the external surface of leg II. C. Granulation on the internal surface of leg III. D. Granulation on the hind leg. The empty arrow indicates a cuticular pore, filled arrows indicate the garter-shaped structure. All photographs taken from specimens belonging to forma aporata; scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893828" httpUri="https://zenodo.org/record/10893828/files/figure.png" pageId="15" pageNumber="94">14AB</figureCitation>
,
<figureCitation id="134B02BEB01D0A7CB253F94A5893F921" box="[576,662,1667,1694]" captionStart="Fig" captionStartId="20.[189,232,1348,1374]" captionTargetBox="[189,1398,265,1309]" captionTargetId="figure-1@20.[189,1398,265,1309]" captionTargetPageId="20" captionText="Fig. 15. Macrobiotus mileri sp. nov., SEM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation and garter-shaped structure on the external surface of leg I (forma porata). B. Granulation and garter-shaped structure on the external surface of leg II (forma aporata). C. Granulation on the internal surface of leg II (forma porata). D. Granulation on the hind leg (forma porata). Empty arrows indicate cuticular pores, filled arrows indicate the garter-shaped structure, empty flat arrowheads indicate the small cuticular bulge / fold, filled flat arrowheads indicate small pores in between granulation that are visible only under SEM. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893830" httpUri="https://zenodo.org/record/10893830/files/figure.png" pageId="15" pageNumber="94">15AB</figureCitation>
) above which there is a small cuticular bulge / fold (visible only under SEM;
<figureCitation id="134B02BEB01D0A7CB140F9605BE7F97B" box="[339,482,1705,1732]" captionStart="Fig" captionStartId="20.[189,232,1348,1374]" captionTargetBox="[189,1398,265,1309]" captionTargetId="figure-1@20.[189,1398,265,1309]" captionTargetPageId="20" captionText="Fig. 15. Macrobiotus mileri sp. nov., SEM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS). A. Granulation and garter-shaped structure on the external surface of leg I (forma porata). B. Granulation and garter-shaped structure on the external surface of leg II (forma aporata). C. Granulation on the internal surface of leg II (forma porata). D. Granulation on the hind leg (forma porata). Empty arrows indicate cuticular pores, filled arrows indicate the garter-shaped structure, empty flat arrowheads indicate the small cuticular bulge / fold, filled flat arrowheads indicate small pores in between granulation that are visible only under SEM. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893830" httpUri="https://zenodo.org/record/10893830/files/figure.png" pageId="15" pageNumber="94">Fig. 15AB</figureCitation>
).
</paragraph>
<paragraph id="8BCF1E3BB01D0A7CB0AEF9245885F857" blockId="15.[189,1399,1773,2025]" pageId="15" pageNumber="94">
Small and robust
<taxonomicName id="4C7065B8B01D0A7CB19DF9245BF8F8B8" authorityName="Schultze" authorityYear="1834" box="[398,509,1773,1799]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="15" pageNumber="94" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B904C229B01D0A7CB19DF9245BF8F8B8" box="[398,509,1773,1799]" italics="true" pageId="15" pageNumber="94">hufelandi</emphasis>
</taxonomicName>
-
<typeStatus id="54CBA099B01D0A7CB216F924583CF8B8" box="[517,569,1773,1799]" pageId="15" pageNumber="94">type</typeStatus>
claws (
<figureCitation id="134B02BEB01D0A7CB285F9245923F8B8" box="[662,806,1773,1799]" captionStart="Fig" captionStartId="21.[189,232,1787,1813]" captionTargetBox="[238,1349,265,1746]" captionTargetId="figure-107@21.[238,1349,265,1746]" captionTargetPageId="21" captionText="Fig. 16. Macrobiotus mileri sp. nov., images of claws (paratypes, ISEA PAS). A. Claws II with smooth lunulae (PCM, forma aporata). B. Claws IV with dentate lunulae (PCM, forma aporata). C. Dentate lunulae (PCM, forma aporata). D. Claws II with smooth lunulae, respectively (SEM, forma porata). E. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893832" httpUri="https://zenodo.org/record/10893832/files/figure.png" pageId="15" pageNumber="94">Fig. 16AE</figureCitation>
). Primary branches with distinct accessory points, a moderately long common tract, and an evident stalk connecting the claw to the lunula (
<figureCitation id="134B02BEB01D0A7CB4CFF8DB5F6FF893" box="[1244,1386,1810,1836]" captionStart="Fig" captionStartId="21.[189,232,1787,1813]" captionTargetBox="[238,1349,265,1746]" captionTargetId="figure-107@21.[238,1349,265,1746]" captionTargetPageId="21" captionText="Fig. 16. Macrobiotus mileri sp. nov., images of claws (paratypes, ISEA PAS). A. Claws II with smooth lunulae (PCM, forma aporata). B. Claws IV with dentate lunulae (PCM, forma aporata). C. Dentate lunulae (PCM, forma aporata). D. Claws II with smooth lunulae, respectively (SEM, forma porata). E. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893832" httpUri="https://zenodo.org/record/10893832/files/figure.png" pageId="15" pageNumber="94">Fig. 16AF</figureCitation>
). The lunulae on legs IIII are smooth (
<figureCitation id="134B02BEB01D0A7CB297F8F15912F8ED" box="[644,791,1848,1874]" captionStart="Fig" captionStartId="21.[189,232,1787,1813]" captionTargetBox="[238,1349,265,1746]" captionTargetId="figure-107@21.[238,1349,265,1746]" captionTargetPageId="21" captionText="Fig. 16. Macrobiotus mileri sp. nov., images of claws (paratypes, ISEA PAS). A. Claws II with smooth lunulae (PCM, forma aporata). B. Claws IV with dentate lunulae (PCM, forma aporata). C. Dentate lunulae (PCM, forma aporata). D. Claws II with smooth lunulae, respectively (SEM, forma porata). E. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893832" httpUri="https://zenodo.org/record/10893832/files/figure.png" pageId="15" pageNumber="94">Fig. 16A, D</figureCitation>
), while there is dentation in the lunulae on legs IV (
<figureCitation id="134B02BEB01D0A7CB0D5F8945B7EF8C7" box="[198,379,1885,1912]" captionStart="Fig" captionStartId="21.[189,232,1787,1813]" captionTargetBox="[238,1349,265,1746]" captionTargetId="figure-107@21.[238,1349,265,1746]" captionTargetPageId="21" captionText="Fig. 16. Macrobiotus mileri sp. nov., images of claws (paratypes, ISEA PAS). A. Claws II with smooth lunulae (PCM, forma aporata). B. Claws IV with dentate lunulae (PCM, forma aporata). C. Dentate lunulae (PCM, forma aporata). D. Claws II with smooth lunulae, respectively (SEM, forma porata). E. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893832" httpUri="https://zenodo.org/record/10893832/files/figure.png" pageId="15" pageNumber="94">Fig. 16B, C, E</figureCitation>
). The cuticular bars are absent, but double muscle attachments are present above the claws IIII (
<figureCitation id="134B02BEB01D0A7CB142F84A5BE0F822" box="[337,485,1923,1949]" captionStart="Fig" captionStartId="21.[189,232,1787,1813]" captionTargetBox="[238,1349,265,1746]" captionTargetId="figure-107@21.[238,1349,265,1746]" captionTargetPageId="21" captionText="Fig. 16. Macrobiotus mileri sp. nov., images of claws (paratypes, ISEA PAS). A. Claws II with smooth lunulae (PCM, forma aporata). B. Claws IV with dentate lunulae (PCM, forma aporata). C. Dentate lunulae (PCM, forma aporata). D. Claws II with smooth lunulae, respectively (SEM, forma porata). E. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893832" httpUri="https://zenodo.org/record/10893832/files/figure.png" pageId="15" pageNumber="94">Fig. 16A, D</figureCitation>
). Shadowed extensions extending from lunulae on legs IIII are present and faintly visible only under PCM (
<figureCitation id="134B02BEB01D0A7CB25CF86058C5F87C" box="[591,704,1961,1987]" captionStart="Fig" captionStartId="21.[189,232,1787,1813]" captionTargetBox="[238,1349,265,1746]" captionTargetId="figure-107@21.[238,1349,265,1746]" captionTargetPageId="21" captionText="Fig. 16. Macrobiotus mileri sp. nov., images of claws (paratypes, ISEA PAS). A. Claws II with smooth lunulae (PCM, forma aporata). B. Claws IV with dentate lunulae (PCM, forma aporata). C. Dentate lunulae (PCM, forma aporata). D. Claws II with smooth lunulae, respectively (SEM, forma porata). E. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893832" httpUri="https://zenodo.org/record/10893832/files/figure.png" pageId="15" pageNumber="94">Fig. 16A</figureCitation>
). A horseshoe-shaped structure connects the anterior and posterior lunules on leg IV (
<figureCitation id="134B02BEB01D0A7CB214F8075876F857" box="[519,627,1998,2024]" captionStart="Fig" captionStartId="21.[189,232,1787,1813]" captionTargetBox="[238,1349,265,1746]" captionTargetId="figure-107@21.[238,1349,265,1746]" captionTargetPageId="21" captionText="Fig. 16. Macrobiotus mileri sp. nov., images of claws (paratypes, ISEA PAS). A. Claws II with smooth lunulae (PCM, forma aporata). B. Claws IV with dentate lunulae (PCM, forma aporata). C. Dentate lunulae (PCM, forma aporata). D. Claws II with smooth lunulae, respectively (SEM, forma porata). E. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893832" httpUri="https://zenodo.org/record/10893832/files/figure.png" pageId="15" pageNumber="94">Fig. 16C</figureCitation>
).
</paragraph>
<caption id="DF0F4EB3B0020A63B0AEF8885B0CF856" ID-DOI="http://doi.org/10.5281/zenodo.10893820" ID-Zenodo-Dep="10893820" httpUri="https://zenodo.org/record/10893820/files/figure.png" pageId="16" pageNumber="95" startId="16.[189,232,1857,1883]" targetBox="[197,1391,265,1813]" targetPageId="16">
<paragraph id="8BCF1E3BB0020A63B0AEF8885B0CF856" blockId="16.[189,1399,1857,2025]" pageId="16" pageNumber="95">
<emphasis id="B904C229B0020A63B0AEF8885B19F8E4" bold="true" box="[189,284,1857,1883]" pageId="16" pageNumber="95">Fig. 10.</emphasis>
<taxonomicName id="4C7065B8B0020A63B136F888580CF8E4" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[293,521,1857,1883]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="16" pageNumber="95" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0020A63B136F888580CF8E4" box="[293,521,1857,1883]" italics="true" pageId="16" pageNumber="95">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0020A63B201F88B5874F8E3" box="[530,625,1858,1884]" pageId="16" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
, SEM images of body granulation and cuticular pores (paratype, ISEA PAS, forma aporata).
<emphasis id="B904C229B0020A63B214F8AD5819F8C1" bold="true" box="[519,540,1892,1918]" pageId="16" pageNumber="95">A</emphasis>
. Body granulation in the dorsal head region.
<emphasis id="B904C229B0020A63B42AF8AC5E48F8C0" bold="true" box="[1081,1101,1893,1919]" pageId="16" pageNumber="95">B</emphasis>
. Body granulation in the dorsal central body region.
<emphasis id="B904C229B0020A63B216F841581FF81D" bold="true" box="[517,538,1928,1954]" pageId="16" pageNumber="95">C</emphasis>
. Body granulation in the dorsal caudal body region.
<emphasis id="B904C229B0020A63B482F8415EA3F81D" bold="true" box="[1169,1190,1928,1954]" pageId="16" pageNumber="95">D</emphasis>
. General view of body granulation in the lateral caudal body region.
<emphasis id="B904C229B0020A63B30FF862594AF87A" bold="true" box="[796,847,1963,1989]" pageId="16" pageNumber="95">EF</emphasis>
. Singular pores in the body cuticle. Scale bars in μm.
</paragraph>
</caption>
<paragraph id="8BCF1E3BB0030A60B0AEFEC05EDDFDDF" blockId="17.[189,1399,265,573]" lastBlockId="19.[189,1399,265,608]" lastPageId="19" lastPageNumber="98" pageId="17" pageNumber="96">
Mouth antero-ventral. Bucco-pharyngeal apparatus of
<taxonomicName id="4C7065B8B0030A62B35AFEC059DBFE9C" authorityName="Schultze" authorityYear="1834" box="[841,990,265,291]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="17" pageNumber="96" phylum="Tardigrada" rank="genus">
<emphasis id="B904C229B0030A62B35AFEC059DBFE9C" box="[841,990,265,291]" italics="true" pageId="17" pageNumber="96">Macrobiotus</emphasis>
</taxonomicName>
<typeStatus id="54CBA099B0030A62B3F4FEC35E19FE9B" box="[999,1052,266,292]" pageId="17" pageNumber="96">type</typeStatus>
, with ventral lamina and ten small peribuccal lamellae followed by six buccal sensory lobes (
<figureCitation id="134B02BEB0030A62B3D2FEE55E36FEF9" box="[961,1075,300,326]" captionStart="Fig" captionStartId="22.[189,232,1456,1482]" captionTargetBox="[189,1398,265,1417]" captionTargetId="figure-1@22.[189,1398,265,1417]" captionTargetPageId="22" captionText="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="17" pageNumber="96">Figs 17A</figureCitation>
,
<figureCitation id="134B02BEB0030A62B452FEE55E99FEF9" box="[1089,1180,300,326]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1398,738,1781]" captionTargetId="figure-0@24.[189,1398,738,1781]" captionTargetPageId="24" captionText="Fig. 18. Macrobiotus mileri sp. nov., mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS). AB. The mouth opening of a single paratype visible from lateral and frontal view respectively. CD. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893836" httpUri="https://zenodo.org/record/10893836/files/figure.png" pageId="17" pageNumber="96">18AD</figureCitation>
). Under PCM, the oral cavity armature is of
<taxonomicName id="4C7065B8B0030A62B1E5FE865860FED6" authorityName="Schultze" authorityYear="1834" box="[502,613,335,361]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="17" pageNumber="96" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B904C229B0030A62B1E5FE865860FED6" box="[502,613,335,361]" italics="true" pageId="17" pageNumber="96">hufelandi</emphasis>
</taxonomicName>
<typeStatus id="54CBA099B0030A62B27CFE9958A7FED5" box="[623,674,336,362]" pageId="17" pageNumber="96">type</typeStatus>
three bands of teeth are always visible (
<figureCitation id="134B02BEB0030A62B4B1FE865F36FED5" box="[1186,1331,335,362]" captionStart="Fig" captionStartId="22.[189,232,1456,1482]" captionTargetBox="[189,1398,265,1417]" captionTargetId="figure-1@22.[189,1398,265,1417]" captionTargetPageId="22" captionText="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="17" pageNumber="96">Fig. 17BC</figureCitation>
). The first band of teeth is composed of numerous extremely small cones arranged in four to six rows located anteriorly in the oral cavity, just behind the bases of the peribuccal lamellae (
<figureCitation id="134B02BEB0030A62B47EFE5F5F07FE0F" box="[1133,1282,406,432]" captionStart="Fig" captionStartId="22.[189,232,1456,1482]" captionTargetBox="[189,1398,265,1417]" captionTargetId="figure-1@22.[189,1398,265,1417]" captionTargetPageId="22" captionText="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="17" pageNumber="96">Figs 17BC</figureCitation>
,
<figureCitation id="134B02BEB0030A62B502FE5F5F6FFE0F" box="[1297,1386,406,432]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1398,738,1781]" captionTargetId="figure-0@24.[189,1398,738,1781]" captionTargetPageId="24" captionText="Fig. 18. Macrobiotus mileri sp. nov., mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS). AB. The mouth opening of a single paratype visible from lateral and frontal view respectively. CD. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893836" httpUri="https://zenodo.org/record/10893836/files/figure.png" pageId="17" pageNumber="96">18CD</figureCitation>
). The second band of teeth is located between the ring fold and the third band of teeth and comprises about four rows of small cones, larger than those of the first band (
<figureCitation id="134B02BEB0030A62B3CAFE155E69FE49" box="[985,1132,476,502]" captionStart="Fig" captionStartId="22.[189,232,1456,1482]" captionTargetBox="[189,1398,265,1417]" captionTargetId="figure-1@22.[189,1398,265,1417]" captionTargetPageId="22" captionText="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="17" pageNumber="96">Figs 17BC</figureCitation>
,
<figureCitation id="134B02BEB0030A62B46BFE155ED7FE49" box="[1144,1234,476,502]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1398,738,1781]" captionTargetId="figure-0@24.[189,1398,738,1781]" captionTargetPageId="24" captionText="Fig. 18. Macrobiotus mileri sp. nov., mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS). AB. The mouth opening of a single paratype visible from lateral and frontal view respectively. CD. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893836" httpUri="https://zenodo.org/record/10893836/files/figure.png" pageId="17" pageNumber="96">18CD</figureCitation>
). The teeth of the third band are located within the posterior portion of the oral cavity, between the second band of teeth and the opening of the buccal tube (
<figureCitation id="134B02BEB0030A62B28DFDEB592AFD82" box="[670,815,546,573]" captionStart="Fig" captionStartId="22.[189,232,1456,1482]" captionTargetBox="[189,1398,265,1417]" captionTargetId="figure-1@22.[189,1398,265,1417]" captionTargetPageId="22" captionText="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="17" pageNumber="96">Figs 17BC</figureCitation>
,
<figureCitation id="134B02BEB0030A62B329FDEA5991FD82" box="[826,916,547,573]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1398,738,1781]" captionTargetId="figure-0@24.[189,1398,738,1781]" captionTargetPageId="24" captionText="Fig. 18. Macrobiotus mileri sp. nov., mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS). AB. The mouth opening of a single paratype visible from lateral and frontal view respectively. CD. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893836" httpUri="https://zenodo.org/record/10893836/files/figure.png" pageId="17" pageNumber="96">18CD</figureCitation>
). The third band of teeth is discontinuous and divided into the dorsal and ventral portions. Under PCM, dorsal teeth are seen as three distinct transverse ridges, and the medio-dorsal tooth is evidently longer than the latero-dorsal teeth (
<figureCitation id="134B02BEB0010A60B4EDFEE55F6DFEF9" box="[1278,1384,300,326]" captionStart="Fig" captionStartId="22.[189,232,1456,1482]" captionTargetBox="[189,1398,265,1417]" captionTargetId="figure-1@22.[189,1398,265,1417]" captionTargetPageId="22" captionText="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="19" pageNumber="98">Fig. 17B</figureCitation>
). The ventral teeth appear as two separate lateral transverse ridges between which a median tooth is visible and rarely divided into two teeth (
<figureCitation id="134B02BEB0010A60B2BFFEBB591CFE32" box="[684,793,370,397]" captionStart="Fig" captionStartId="22.[189,232,1456,1482]" captionTargetBox="[189,1398,265,1417]" captionTargetId="figure-1@22.[189,1398,265,1417]" captionTargetPageId="22" captionText="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="19" pageNumber="98">Fig. 17C</figureCitation>
). Under SEM, the dorsal and ventral teeth are also clearly distinct (
<figureCitation id="134B02BEB0010A60B16EFE5F580BFE0F" box="[381,526,406,432]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1398,738,1781]" captionTargetId="figure-0@24.[189,1398,738,1781]" captionTargetPageId="24" captionText="Fig. 18. Macrobiotus mileri sp. nov., mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS). AB. The mouth opening of a single paratype visible from lateral and frontal view respectively. CD. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893836" httpUri="https://zenodo.org/record/10893836/files/figure.png" pageId="19" pageNumber="98">Fig. 18CD</figureCitation>
). Under SEM, the margins of the dorsal teeth are serrated and the medio-dorsal tooth is clearly longer than latero-dorsal teeth (
<figureCitation id="134B02BEB0010A60B351FE7059A8FE6C" box="[834,941,441,467]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1398,738,1781]" captionTargetId="figure-0@24.[189,1398,738,1781]" captionTargetPageId="24" captionText="Fig. 18. Macrobiotus mileri sp. nov., mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS). AB. The mouth opening of a single paratype visible from lateral and frontal view respectively. CD. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893836" httpUri="https://zenodo.org/record/10893836/files/figure.png" pageId="19" pageNumber="98">Fig. 18C</figureCitation>
) whereas the ventral teeth are smaller and their margins are less serrated (
<figureCitation id="134B02BEB0010A60B27FFE1558D9FE49" box="[620,732,476,502]" captionStart="Fig" captionStartId="24.[189,232,1822,1848]" captionTargetBox="[189,1398,738,1781]" captionTargetId="figure-0@24.[189,1398,738,1781]" captionTargetPageId="24" captionText="Fig. 18. Macrobiotus mileri sp. nov., mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS). AB. The mouth opening of a single paratype visible from lateral and frontal view respectively. CD. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893836" httpUri="https://zenodo.org/record/10893836/files/figure.png" pageId="19" pageNumber="98">Fig. 18D</figureCitation>
). Pharyngeal bulb spherical, with triangular apophyses, two rod-shaped macroplacoids and a large triangular microplacoid (
<figureCitation id="134B02BEB0010A60B3C0FE365E45FDA6" box="[979,1088,511,537]" captionStart="Fig" captionStartId="22.[189,232,1456,1482]" captionTargetBox="[189,1398,265,1417]" captionTargetId="figure-1@22.[189,1398,265,1417]" captionTargetPageId="22" captionText="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="19" pageNumber="98">Fig. 17A</figureCitation>
). The macroplacoid length sequence being 2&lt;1. The first and the second macroplacoid are constricted centrally and subterminally, respectively (
<figureCitation id="134B02BEB0010A60B148FD8F5BEFFDDF" box="[347,490,582,608]" captionStart="Fig" captionStartId="22.[189,232,1456,1482]" captionTargetBox="[189,1398,265,1417]" captionTargetId="figure-1@22.[189,1398,265,1417]" captionTargetPageId="22" captionText="Fig. 17. Macrobiotus mileri sp. nov., PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata). A. An entire buccal apparatus. BC. The oral cavity armature, dorsal and ventral teeth respectively. DE. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="19" pageNumber="98">Fig. 17DE</figureCitation>
). The animals measurements and statistics are given in
<tableCitation id="C6F22B80B0010A60B469FD8F5ED1FDDF" box="[1146,1236,582,608]" captionStart="Table 4" captionStartId="23.[189,258,265,291]" captionTargetPageId="23" captionText="Table 4. Measurements [in μm] and pt values of selected morphological structures of animals of Macrobiotus mileri sp. nov.; specimens mounted in Hoyers medium; N: number of specimen/structures measured; range: refers to the smallest and the largest structure among all measured specimens; SD: standard deviation." httpUri="http://table.plazi.org/id/DF0F4EB3B0050A64B0AEFEC05B98FE33" pageId="19" pageNumber="98" tableUuid="DF0F4EB3B0050A64B0AEFEC05B98FE33">Table 4</tableCitation>
.
</paragraph>
<caption id="DF0F4EB3B0030A62B0AEF8625E39F856" ID-DOI="http://doi.org/10.5281/zenodo.10893822" ID-Zenodo-Dep="10893822" httpUri="https://zenodo.org/record/10893822/files/figure.png" pageId="17" pageNumber="96" startId="17.[189,232,1963,1989]" targetBox="[189,1398,659,1922]" targetPageId="17">
<paragraph id="8BCF1E3BB0030A62B0AEF8625E39F856" blockId="17.[189,1398,1963,2025]" pageId="17" pageNumber="96">
<emphasis id="B904C229B0030A62B0AEF8625B1CF87A" bold="true" box="[189,281,1963,1989]" pageId="17" pageNumber="96">Fig. 11.</emphasis>
<taxonomicName id="4C7065B8B0030A62B133F8625807F87A" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[288,514,1963,1989]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="17" pageNumber="96" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0030A62B133F8625807F87A" box="[288,514,1963,1989]" italics="true" pageId="17" pageNumber="96">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0030A62B21AF8655863F879" box="[521,614,1964,1990]" pageId="17" pageNumber="96" rank="species">sp. nov.</taxonomicNameLabel>
, PCM images of cuticular pores patches in forma porata (paratype, ISEA PAS).
<emphasis id="B904C229B0030A62B15CF8075B61F857" bold="true" box="[335,356,1998,2024]" pageId="17" pageNumber="96">A</emphasis>
. Patches II, III and IV.
<emphasis id="B904C229B0030A62B264F807588EF857" bold="true" box="[631,651,1998,2024]" pageId="17" pageNumber="96">B</emphasis>
. Patches IV and V. Scale bars in μm.
</paragraph>
</caption>
<caption id="DF0F4EB3B0000A61B0AEFAC458E4FAD1" ID-DOI="http://doi.org/10.5281/zenodo.10893824" ID-Zenodo-Dep="10893824" httpUri="https://zenodo.org/record/10893824/files/figure.png" pageId="18" pageNumber="97" startId="18.[189,232,1293,1319]" targetBox="[227,1361,265,1240]" targetPageId="18">
<paragraph id="8BCF1E3BB0000A61B0AEFAC458E4FAD1" blockId="18.[189,1398,1293,1390]" pageId="18" pageNumber="97">
<emphasis id="B904C229B0000A61B0AEFAC45B1FFA98" bold="true" box="[189,282,1293,1319]" pageId="18" pageNumber="97">Fig. 12.</emphasis>
<taxonomicName id="4C7065B8B0000A61B131FAC45801FA98" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[290,516,1293,1319]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="18" pageNumber="97" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0000A61B131FAC45801FA98" box="[290,516,1293,1319]" italics="true" pageId="18" pageNumber="97">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0000A61B218FAC7586CFA97" box="[523,617,1294,1320]" pageId="18" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
, SEM images of cuticular pores patches in two paratypes of forma porata (ISEA PAS).
<emphasis id="B904C229B0000A61B1A0FAF95BCDFAF5" bold="true" box="[435,456,1328,1354]" pageId="18" pageNumber="97">A</emphasis>
,
<emphasis id="B904C229B0000A61B1C9FAF95BEAFAF5" bold="true" box="[474,495,1328,1354]" pageId="18" pageNumber="97">C</emphasis>
. Patches II, III and IV.
<emphasis id="B904C229B0000A61B300FAF95922FAF5" bold="true" box="[787,807,1328,1354]" pageId="18" pageNumber="97">B</emphasis>
,
<emphasis id="B904C229B0000A61B32BFAF95948FAF5" bold="true" box="[824,845,1328,1354]" pageId="18" pageNumber="97">D</emphasis>
. Dorsal view on the caudal body region with continuous patch V of pores. Scale bars in μm.
</paragraph>
</caption>
<caption id="DF0F4EB3B0000A61B0AEF841582DF856" ID-DOI="http://doi.org/10.5281/zenodo.10893826" ID-Zenodo-Dep="10893826" httpUri="https://zenodo.org/record/10893826/files/figure.png" pageId="18" pageNumber="97" startId="18.[189,232,1928,1954]" targetBox="[194,1394,1471,1871]" targetPageId="18">
<paragraph id="8BCF1E3BB0000A61B0AEF841582DF856" blockId="18.[189,1399,1928,2025]" pageId="18" pageNumber="97">
<emphasis id="B904C229B0000A61B0AEF8415B18F81D" bold="true" box="[189,285,1928,1954]" pageId="18" pageNumber="97">Fig. 13.</emphasis>
<taxonomicName id="4C7065B8B0000A61B134F8415809F81D" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[295,524,1928,1954]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="18" pageNumber="97" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0000A61B134F8415809F81D" box="[295,524,1928,1954]" italics="true" pageId="18" pageNumber="97">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0000A61B205F8415870F81D" box="[534,629,1928,1954]" pageId="18" pageNumber="97" rank="species">sp. nov.</taxonomicNameLabel>
, a schematic drawing of a specimen belonging to forma porata, showing the distribution of patches of cuticular pores, body and leg granulation as well as the gartershaped structures on legs IIII.
</paragraph>
</caption>
<caption id="DF0F4EB3B0010A60B0AEF8D75832F856" ID-DOI="http://doi.org/10.5281/zenodo.10893828" ID-Zenodo-Dep="10893828" httpUri="https://zenodo.org/record/10893828/files/figure.png" pageId="19" pageNumber="98" startId="19.[189,232,1822,1848]" targetBox="[189,1398,716,1782]" targetPageId="19">
<paragraph id="8BCF1E3BB0010A60B0AEF8D75832F856" blockId="19.[189,1399,1822,2025]" pageId="19" pageNumber="98">
<emphasis id="B904C229B0010A60B0AEF8D75B1FF887" bold="true" box="[189,282,1822,1848]" pageId="19" pageNumber="98">Fig. 14.</emphasis>
<taxonomicName id="4C7065B8B0010A60B132F8D75806F887" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[289,515,1822,1848]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="19" pageNumber="98" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0010A60B132F8D75806F887" box="[289,515,1822,1848]" italics="true" pageId="19" pageNumber="98">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0010A60B219F8D65863F886" box="[522,614,1823,1849]" pageId="19" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
, PCM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS).
<emphasis id="B904C229B0010A60B251F8885852F8E4" bold="true" box="[578,599,1857,1883]" pageId="19" pageNumber="98">A</emphasis>
. Granulation on the external, proximal and internal surface of leg II.
<emphasis id="B904C229B0010A60B0F2F8AC5AF0F8C0" bold="true" box="[225,245,1893,1919]" pageId="19" pageNumber="98">B</emphasis>
. Garter-shaped structure and granulation on the external surface of leg II.
<emphasis id="B904C229B0010A60B462F8AD5E83F8C1" bold="true" box="[1137,1158,1892,1918]" pageId="19" pageNumber="98">C</emphasis>
. Granulation on the internal surface of leg III.
<emphasis id="B904C229B0010A60B1E5F841580EF81D" bold="true" box="[502,523,1928,1954]" pageId="19" pageNumber="98">D</emphasis>
. Granulation on the hind leg. The empty arrow indicates a cuticular pore, filled arrows indicate the garter-shaped structure. All photographs taken from specimens belonging to forma aporata; scale bars in μm.
</paragraph>
</caption>
<caption id="DF0F4EB3B0060A67B0AEFA8D5E24F98D" ID-DOI="http://doi.org/10.5281/zenodo.10893830" ID-Zenodo-Dep="10893830" httpUri="https://zenodo.org/record/10893830/files/figure.png" pageId="20" pageNumber="99" startId="20.[189,232,1348,1374]" targetBox="[189,1398,265,1309]" targetPageId="20">
<paragraph id="8BCF1E3BB0060A67B0AEFA8D5E24F98D" blockId="20.[189,1399,1348,1586]" pageId="20" pageNumber="99">
<emphasis id="B904C229B0060A67B0AEFA8D5B1FFAE1" bold="true" box="[189,282,1348,1374]" pageId="20" pageNumber="99">Fig. 15.</emphasis>
<taxonomicName id="4C7065B8B0060A67B132FA8D5806FAE1" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[289,515,1348,1374]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="20" pageNumber="99" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0060A67B132FA8D5806FAE1" box="[289,515,1348,1374]" italics="true" pageId="20" pageNumber="99">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0060A67B219FA8D5862FAE1" box="[522,615,1348,1374]" pageId="20" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
, SEM images of dense granulation patches and cuticular structures on legs (paratypes, ISEA PAS).
<emphasis id="B904C229B0060A67B22FFAAE5854FA3E" bold="true" box="[572,593,1383,1409]" pageId="20" pageNumber="99">A</emphasis>
. Granulation and garter-shaped structure on the external surface of leg I (forma porata).
<emphasis id="B904C229B0060A67B1A1FA435BC3FA1B" bold="true" box="[434,454,1418,1444]" pageId="20" pageNumber="99">B</emphasis>
. Granulation and garter-shaped structure on the external surface of leg II (forma aporata).
<emphasis id="B904C229B0060A67B13FFA645B44FA78" bold="true" box="[300,321,1453,1479]" pageId="20" pageNumber="99">C</emphasis>
. Granulation on the internal surface of leg II (forma porata).
<emphasis id="B904C229B0060A67B403FA675E20FA77" bold="true" box="[1040,1061,1454,1480]" pageId="20" pageNumber="99">D</emphasis>
. Granulation on the hind leg (forma porata). Empty arrows indicate cuticular pores, filled arrows indicate the garter-shaped structure, empty flat arrowheads indicate the small cuticular bulge / fold, filled flat arrowheads indicate small pores in between granulation that are visible only under SEM. Scale bars in μm.
</paragraph>
</caption>
<caption id="DF0F4EB3B0070A66B0AEF9325E9CF856" ID-DOI="http://doi.org/10.5281/zenodo.10893832" ID-Zenodo-Dep="10893832" httpUri="https://zenodo.org/record/10893832/files/figure.png" pageId="21" pageNumber="100" startId="21.[189,232,1787,1813]" targetBox="[238,1349,265,1746]" targetPageId="21">
<paragraph id="8BCF1E3BB0070A66B0AEF9325E9CF856" blockId="21.[189,1399,1787,2025]" pageId="21" pageNumber="100">
<emphasis id="B904C229B0070A66B0AEF9325B1FF8AA" bold="true" box="[189,282,1787,1813]" pageId="21" pageNumber="100">Fig. 16.</emphasis>
<taxonomicName id="4C7065B8B0070A66B131F9325801F8AA" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[290,516,1787,1813]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="21" pageNumber="100" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0070A66B131F9325801F8AA" box="[290,516,1787,1813]" italics="true" pageId="21" pageNumber="100">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0070A66B21FF935586CF8A9" box="[524,617,1788,1814]" pageId="21" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
, images of claws (paratypes, ISEA PAS).
<emphasis id="B904C229B0070A66B446F9325E6FF8AA" bold="true" box="[1109,1130,1787,1813]" pageId="21" pageNumber="100">A</emphasis>
. Claws II with smooth lunulae (PCM, forma aporata).
<emphasis id="B904C229B0070A66B22AF8D75848F887" bold="true" box="[569,589,1822,1848]" pageId="21" pageNumber="100">B</emphasis>
. Claws IV with dentate lunulae (PCM, forma aporata).
<emphasis id="B904C229B0070A66B4E7F8D75F0CF887" bold="true" box="[1268,1289,1822,1848]" pageId="21" pageNumber="100">C</emphasis>
. Dentate lunulae (PCM, forma aporata).
<emphasis id="B904C229B0070A66B229F88B584AF8E3" bold="true" box="[570,591,1858,1884]" pageId="21" pageNumber="100">D</emphasis>
. Claws II with smooth lunulae, respectively (SEM, forma porata).
<emphasis id="B904C229B0070A66B0AEF8AC5AD4F8C0" bold="true" box="[189,209,1893,1919]" pageId="21" pageNumber="100">E</emphasis>
. Claws IV with dentate lunulae (SEM, forma porata). The empty arrow indicates a singular cuticular pore, filled indented arrowheads indicate shadowed extensions extending from the lunulae (under PCM), filled flat arrowheads indicate paired muscles attachments, and empty indented arrowheads indicate the horseshoe structure connecting the anterior and the posterior claw. Scale bars in μm.
</paragraph>
</caption>
<caption id="DF0F4EB3B0040A65B0AEFA795B44F921" ID-DOI="http://doi.org/10.5281/zenodo.10893834" ID-Zenodo-Dep="10893834" httpUri="https://zenodo.org/record/10893834/files/figure.png" pageId="22" pageNumber="101" startId="22.[189,232,1456,1482]" targetBox="[189,1398,265,1417]" targetPageId="22">
<paragraph id="8BCF1E3BB0040A65B0AEFA795B44F921" blockId="22.[189,1399,1456,1694]" pageId="22" pageNumber="101">
<emphasis id="B904C229B0040A65B0AEFA795B1CFA75" bold="true" box="[189,281,1456,1482]" pageId="22" pageNumber="101">Fig. 17.</emphasis>
<taxonomicName id="4C7065B8B0040A65B10CFA795805FA75" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[287,512,1456,1482]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="22" pageNumber="101" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0040A65B10CFA795805FA75" box="[287,512,1456,1482]" italics="true" pageId="22" pageNumber="101">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0040A65B215FA785867FA74" box="[518,610,1457,1483]" pageId="22" pageNumber="101" rank="species">sp. nov.</taxonomicNameLabel>
, PCM images of the buccal apparatus (all from holotype, IL.001.11, ISEA PAS, forma aporata).
<emphasis id="B904C229B0040A65B21AFA1A581BFA52" bold="true" box="[521,542,1491,1517]" pageId="22" pageNumber="101">A</emphasis>
. An entire buccal apparatus.
<emphasis id="B904C229B0040A65B36EFA1A59B1FA52" bold="true" box="[893,948,1491,1518]" pageId="22" pageNumber="101">BC</emphasis>
. The oral cavity armature, dorsal and ventral teeth respectively.
<emphasis id="B904C229B0040A65B1FFFA3E5821F9AE" bold="true" box="[492,548,1527,1553]" pageId="22" pageNumber="101">DE</emphasis>
. Placoid morphology, dorsal and ventral placoids, respectively. The filled flat arrowheads indicate the first band of teeth, the empty flat arrowheads indicate the second band of teeth, the filled indented arrowheads indicate the third band of teeth, and the empty indented arrowheads indicate central and subterminal constrictions in the first and second macroplacoid, respectively. Scale bars in μm.
</paragraph>
</caption>
<caption id="DF0F4EB3B0050A64B0AEFEC05B98FE33" ID-Table-UUID="DF0F4EB3B0050A64B0AEFEC05B98FE33" httpUri="http://table.plazi.org/id/DF0F4EB3B0050A64B0AEFEC05B98FE33" pageId="23" pageNumber="102" startId="23.[189,258,265,291]" targetBox="[197,1385,427,1737]" targetIsTable="true" targetPageId="23">
<paragraph id="8BCF1E3BB0050A64B0AEFEC05B98FE33" blockId="23.[189,1399,265,396]" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B0AEFEC05B20FE9C" bold="true" box="[189,293,265,291]" pageId="23" pageNumber="102">Table 4.</emphasis>
Measurements [in μm] and
<emphasis id="B904C229B0050A64B29FFEC358A6FE9C" box="[652,675,266,291]" italics="true" pageId="23" pageNumber="102">pt</emphasis>
values of selected morphological structures of animals of
<taxonomicName id="4C7065B8B0050A64B0AEFEE55B9BFEF9" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[189,414,300,326]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="23" pageNumber="102" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0050A64B0AEFEE55B9BFEF9" box="[189,414,300,326]" italics="true" pageId="23" pageNumber="102">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0050A64B1B6FEE45807FEF8" box="[421,514,301,327]" pageId="23" pageNumber="102" rank="species">sp. nov.</taxonomicNameLabel>
; specimens mounted in Hoyers medium; N: number of specimen/structures measured; range: refers to the smallest and the largest structure among all measured specimens; SD: standard deviation.
</paragraph>
</caption>
<paragraph id="8BCF1E3BB0050A64B0D6FE625F50F977" pageId="23" pageNumber="102">
<table id="F970EC9BB005F58CB0D6FE625F6CF976" box="[197,1385,427,1737]" gridcols="10" gridrows="35" pageId="23" pageNumber="102">
<tr id="35401C79B005F58CB0D6FE625F6CFE7B" box="[197,1385,427,452]" gridrow="0" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FE625824FE7B" box="[197,545,427,452]" gridcol="0" gridrow="0" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B0D6FE625B3FFE7C" bold="true" box="[197,314,427,451]" pageId="23" pageNumber="102">Character</emphasis>
</th>
<th id="76917505B005F58CB25BFE625866FE7B" box="[584,611,427,452]" gridcol="1" gridrow="0" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B25FFE65585AFE7B" bold="true" box="[588,607,428,452]" pageId="23" pageNumber="102">N</emphasis>
</th>
<th id="76917505B005F58CB29EFE62598BFE7B" box="[653,910,427,452]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B2F5FE65592AFE7B" bold="true" box="[742,815,428,452]" pageId="23" pageNumber="102">Range</emphasis>
</th>
<th id="76917505B005F58CB3B8FE625E29FE7B" box="[939,1068,427,452]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3DAFE655E0FFE7B" bold="true" box="[969,1034,428,452]" pageId="23" pageNumber="102">Mean</emphasis>
</th>
<th id="76917505B005F58CB45DFE625EC5FE7B" box="[1102,1216,427,452]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B464FE625E9CFE7C" bold="true" box="[1143,1177,427,451]" pageId="23" pageNumber="102">SD</emphasis>
</th>
<th id="76917505B005F58CB4FBFE625F6CFE7B" box="[1256,1385,427,452]" colspan="2" colspanRight="1" gridcol="8" gridrow="0" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B4E0FE655F5EFE7B" bold="true" box="[1267,1371,428,452]" pageId="23" pageNumber="102">Holotype</emphasis>
</th>
</tr>
<tr id="35401C79B005F58CB0D6FE185F6CFE56" box="[197,1385,465,489]" gridrow="1" pageId="23" pageNumber="102" rowspan-0="1" rowspan-1="1">
<td id="76917505B005F58CB29EFE1858FDFE56" box="[653,760,465,489]" gridcol="2" gridrow="1" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B2A3FE1858D3FE56" bold="true" box="[688,726,465,489]" pageId="23" pageNumber="102">µm</emphasis>
</td>
<td id="76917505B005F58CB304FE18598BFE56" box="[791,910,465,489]" gridcol="3" gridrow="1" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B35BFE1B5958FE56" bold="true" box="[840,861,466,489]" italics="true" pageId="23" pageNumber="102">pt</emphasis>
</td>
<td id="76917505B005F58CB3B8FE1859DFFE56" box="[939,986,465,489]" gridcol="4" gridrow="1" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3BCFE1859D0FE56" bold="true" box="[943,981,465,489]" pageId="23" pageNumber="102">µm</emphasis>
</td>
<td id="76917505B005F58CB3E4FE185E29FE56" box="[1015,1068,465,489]" gridcol="5" gridrow="1" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B414FE1B5E19FE56" bold="true" box="[1031,1052,466,489]" italics="true" pageId="23" pageNumber="102">pt</emphasis>
</td>
<td id="76917505B005F58CB45DFE185E71FE56" box="[1102,1140,465,489]" gridcol="6" gridrow="1" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B45DFE185E71FE56" bold="true" box="[1102,1140,465,489]" pageId="23" pageNumber="102">µm</emphasis>
</td>
<td id="76917505B005F58CB48CFE185EC5FE56" box="[1183,1216,465,489]" gridcol="7" gridrow="1" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B4B5FE1B5EBEFE56" bold="true" box="[1190,1211,466,489]" italics="true" pageId="23" pageNumber="102">pt</emphasis>
</td>
<td id="76917505B005F58CB4FBFE185F12FE56" box="[1256,1303,465,489]" gridcol="8" gridrow="1" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B4FEFE185F16FE56" bold="true" box="[1261,1299,465,489]" pageId="23" pageNumber="102">µm</emphasis>
</td>
<td id="76917505B005F58CB527FE185F6CFE56" box="[1332,1385,465,489]" gridcol="9" gridrow="1" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B557FE1B5F5CFE56" bold="true" box="[1348,1369,466,489]" italics="true" pageId="23" pageNumber="102">pt</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FE3E5F6CFDB0" box="[197,1385,503,527]" gridrow="2" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FE3E5824FDB0" box="[197,545,503,527]" gridcol="0" gridrow="2" pageId="23" pageNumber="102">Body length</th>
<td id="76917505B005F58CB25BFE3E5866FDB0" box="[584,611,503,527]" gridcol="1" gridrow="2" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFE3E58FDFDB0" box="[653,760,503,527]" gridcol="2" gridrow="2" pageId="23" pageNumber="102">326523</td>
<td id="76917505B005F58CB304FE3E598BFDB0" box="[791,910,503,527]" gridcol="3" gridrow="2" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B304FE3E598BFDB0" box="[791,910,503,527]" italics="true" pageId="23" pageNumber="102">11011392</emphasis>
</td>
<td id="76917505B005F58CB3B8FE3E59DFFDB0" box="[939,986,503,527]" gridcol="4" gridrow="2" pageId="23" pageNumber="102">449</td>
<td id="76917505B005F58CB3E4FE3E5E29FDB0" box="[1015,1068,503,527]" gridcol="5" gridrow="2" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E4FE3E5E29FDB0" box="[1015,1068,503,527]" italics="true" pageId="23" pageNumber="102">1251</emphasis>
</td>
<td id="76917505B005F58CB45DFE3E5E71FDB0" box="[1102,1140,503,527]" gridcol="6" gridrow="2" pageId="23" pageNumber="102">54</td>
<td id="76917505B005F58CB48CFE3E5EC5FDB0" box="[1183,1216,503,527]" gridcol="7" gridrow="2" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B4B0FE3E5EBBFDB0" box="[1187,1214,503,527]" italics="true" pageId="23" pageNumber="102">71</emphasis>
</td>
<td id="76917505B005F58CB4FBFE3E5F12FDB0" box="[1256,1303,503,527]" gridcol="8" gridrow="2" pageId="23" pageNumber="102">488</td>
<td id="76917505B005F58CB527FE3E5F6CFDB0" box="[1332,1385,503,527]" gridcol="9" gridrow="2" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B527FE3E5F6CFDB0" box="[1332,1385,503,527]" italics="true" pageId="23" pageNumber="102">1217</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FDD45F6CFD8A" box="[197,1385,541,565]" gridrow="3" pageId="23" pageNumber="102" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76917505B005F58CB0D6FDD45824FD8A" box="[197,545,541,565]" gridcol="0" gridrow="3" pageId="23" pageNumber="102">Buccal tube</th>
</tr>
<tr id="35401C79B005F58CB0D6FD8A5F6CFDE4" box="[197,1385,579,603]" gridrow="4" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FD8A5824FDE4" box="[197,545,579,603]" gridcol="0" gridrow="4" pageId="23" pageNumber="102">Buccal tube length</th>
<td id="76917505B005F58CB25BFD8A5866FDE4" box="[584,611,579,603]" gridcol="1" gridrow="4" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFD8A58FDFDE4" box="[653,760,579,603]" gridcol="2" gridrow="4" pageId="23" pageNumber="102">29.540.8</td>
<td id="76917505B005F58CB304FD8A598BFDE4" box="[791,910,579,603]" gridcol="3" gridrow="4" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B35FFD8D595CFDE4" box="[844,857,580,603]" italics="true" pageId="23" pageNumber="102"></emphasis>
</td>
<td id="76917505B005F58CB3B8FD8A59DFFDE4" box="[939,986,579,603]" gridcol="4" gridrow="4" pageId="23" pageNumber="102">35.8</td>
<td id="76917505B005F58CB3E4FD8A5E29FDE4" box="[1015,1068,579,603]" gridcol="5" gridrow="4" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B418FD8D5E1DFDE4" box="[1035,1048,580,603]" italics="true" pageId="23" pageNumber="102"></emphasis>
</td>
<td id="76917505B005F58CB45DFD8A5E71FDE4" box="[1102,1140,579,603]" gridcol="6" gridrow="4" pageId="23" pageNumber="102">3.4</td>
<td id="76917505B005F58CB48CFD8A5EC5FDE4" box="[1183,1216,579,603]" gridcol="7" gridrow="4" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B4BAFD8D5EB3FDE4" box="[1193,1206,580,603]" italics="true" pageId="23" pageNumber="102"></emphasis>
</td>
<td id="76917505B005F58CB4FBFD8A5F12FDE4" box="[1256,1303,579,603]" gridcol="8" gridrow="4" pageId="23" pageNumber="102">40.1</td>
<td id="76917505B005F58CB527FD8A5F6CFDE4" box="[1332,1385,579,603]" gridcol="9" gridrow="4" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B55BFD8D5F50FDE4" box="[1352,1365,580,603]" italics="true" pageId="23" pageNumber="102"></emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FDA15F6CFD3E" box="[197,1385,616,641]" gridrow="5" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FDA15824FD3E" box="[197,545,616,641]" gridcol="0" gridrow="5" pageId="23" pageNumber="102">Stylet support insertion point</th>
<td id="76917505B005F58CB25BFDA15866FD3E" box="[584,611,616,641]" gridcol="1" gridrow="5" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFDA158FDFD3E" box="[653,760,616,641]" gridcol="2" gridrow="5" pageId="23" pageNumber="102">23.332.8</td>
<td id="76917505B005F58CB304FDA1598BFD3E" box="[791,910,616,641]" gridcol="3" gridrow="5" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFDA0598DFD3E" box="[797,904,617,641]" italics="true" pageId="23" pageNumber="102">78.981.3</emphasis>
</td>
<td id="76917505B005F58CB3B8FDA159DFFD3E" box="[939,986,616,641]" gridcol="4" gridrow="5" pageId="23" pageNumber="102">28.5</td>
<td id="76917505B005F58CB3E4FDA15E29FD3E" box="[1015,1068,616,641]" gridcol="5" gridrow="5" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FDA05E2CFD3E" box="[1018,1065,617,641]" italics="true" pageId="23" pageNumber="102">79.7</emphasis>
</td>
<td id="76917505B005F58CB45DFDA15E71FD3E" box="[1102,1140,616,641]" gridcol="6" gridrow="5" pageId="23" pageNumber="102">2.7</td>
<td id="76917505B005F58CB48CFDA15EC5FD3E" box="[1183,1216,616,641]" gridcol="7" gridrow="5" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFDA15EC5FD3F" box="[1183,1216,616,640]" italics="true" pageId="23" pageNumber="102">0.6</emphasis>
</td>
<td id="76917505B005F58CB4FBFDA15F12FD3E" box="[1256,1303,616,641]" gridcol="8" gridrow="5" pageId="23" pageNumber="102">32.0</td>
<td id="76917505B005F58CB527FDA15F6CFD3E" box="[1332,1385,616,641]" gridcol="9" gridrow="5" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FDA05F63FD3E" box="[1335,1382,617,641]" italics="true" pageId="23" pageNumber="102">79.8</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FD475F6CFD18" box="[197,1385,654,679]" gridrow="6" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FD475824FD18" box="[197,545,654,679]" gridcol="0" gridrow="6" pageId="23" pageNumber="102">Buccal tube external width</th>
<td id="76917505B005F58CB25BFD475866FD18" box="[584,611,654,679]" gridcol="1" gridrow="6" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFD4758FDFD18" box="[653,760,654,679]" gridcol="2" gridrow="6" pageId="23" pageNumber="102">4.36.7</td>
<td id="76917505B005F58CB304FD47598BFD18" box="[791,910,654,679]" gridcol="3" gridrow="6" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFD46598DFD18" box="[797,904,655,679]" italics="true" pageId="23" pageNumber="102">14.417.3</emphasis>
</td>
<td id="76917505B005F58CB3B8FD4759DFFD18" box="[939,986,654,679]" gridcol="4" gridrow="6" pageId="23" pageNumber="102">5.8</td>
<td id="76917505B005F58CB3E4FD475E29FD18" box="[1015,1068,654,679]" gridcol="5" gridrow="6" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FD475E2CFD19" box="[1018,1065,654,678]" italics="true" pageId="23" pageNumber="102">16.1</emphasis>
</td>
<td id="76917505B005F58CB45DFD475E71FD18" box="[1102,1140,654,679]" gridcol="6" gridrow="6" pageId="23" pageNumber="102">0.7</td>
<td id="76917505B005F58CB48CFD475EC5FD18" box="[1183,1216,654,679]" gridcol="7" gridrow="6" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFD465EC5FD18" box="[1183,1216,655,679]" italics="true" pageId="23" pageNumber="102">0.8</emphasis>
</td>
<td id="76917505B005F58CB4FBFD475F12FD18" box="[1256,1303,654,679]" gridcol="8" gridrow="6" pageId="23" pageNumber="102">6.4</td>
<td id="76917505B005F58CB527FD475F6CFD18" box="[1332,1385,654,679]" gridcol="9" gridrow="6" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FD475F63FD19" box="[1335,1382,654,678]" italics="true" pageId="23" pageNumber="102">16.0</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FD7D5F6CFD73" box="[197,1385,692,716]" gridrow="7" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FD7D5824FD73" box="[197,545,692,716]" gridcol="0" gridrow="7" pageId="23" pageNumber="102">Buccal tube internal width</th>
<td id="76917505B005F58CB25BFD7D5866FD73" box="[584,611,692,716]" gridcol="1" gridrow="7" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFD7D58FDFD73" box="[653,760,692,716]" gridcol="2" gridrow="7" pageId="23" pageNumber="102">3.05.2</td>
<td id="76917505B005F58CB304FD7D598BFD73" box="[791,910,692,716]" gridcol="3" gridrow="7" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B337FD7D5984FD73" box="[804,897,692,716]" italics="true" pageId="23" pageNumber="102">9.713.1</emphasis>
</td>
<td id="76917505B005F58CB3B8FD7D59DFFD73" box="[939,986,692,716]" gridcol="4" gridrow="7" pageId="23" pageNumber="102">4.2</td>
<td id="76917505B005F58CB3E4FD7D5E29FD73" box="[1015,1068,692,716]" gridcol="5" gridrow="7" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E8FD7D5E2DFD73" box="[1019,1064,692,716]" italics="true" pageId="23" pageNumber="102">11.7</emphasis>
</td>
<td id="76917505B005F58CB45DFD7D5E71FD73" box="[1102,1140,692,716]" gridcol="6" gridrow="7" pageId="23" pageNumber="102">0.6</td>
<td id="76917505B005F58CB48CFD7D5EC5FD73" box="[1183,1216,692,716]" gridcol="7" gridrow="7" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFD7D5EC5FD73" box="[1183,1216,692,716]" italics="true" pageId="23" pageNumber="102">0.9</emphasis>
</td>
<td id="76917505B005F58CB4FBFD7D5F12FD73" box="[1256,1303,692,716]" gridcol="8" gridrow="7" pageId="23" pageNumber="102">5.1</td>
<td id="76917505B005F58CB527FD7D5F6CFD73" box="[1332,1385,692,716]" gridcol="9" gridrow="7" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FD7D5F63FD73" box="[1335,1382,692,716]" italics="true" pageId="23" pageNumber="102">12.7</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FD135F6CFD4D" box="[197,1385,730,754]" gridrow="8" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FD135824FD4D" box="[197,545,730,754]" gridcol="0" gridrow="8" pageId="23" pageNumber="102">Ventral lamina length</th>
<td id="76917505B005F58CB25BFD135866FD4D" box="[584,611,730,754]" gridcol="1" gridrow="8" pageId="23" pageNumber="102">19</td>
<td id="76917505B005F58CB29EFD1358FDFD4D" box="[653,760,730,754]" gridcol="2" gridrow="8" pageId="23" pageNumber="102">17.224.2</td>
<td id="76917505B005F58CB304FD13598BFD4D" box="[791,910,730,754]" gridcol="3" gridrow="8" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFD13598DFD4D" box="[797,904,730,754]" italics="true" pageId="23" pageNumber="102">53.560.4</emphasis>
</td>
<td id="76917505B005F58CB3B8FD1359DFFD4D" box="[939,986,730,754]" gridcol="4" gridrow="8" pageId="23" pageNumber="102">20.8</td>
<td id="76917505B005F58CB3E4FD135E29FD4D" box="[1015,1068,730,754]" gridcol="5" gridrow="8" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FD135E2CFD4D" box="[1018,1065,730,754]" italics="true" pageId="23" pageNumber="102">57.6</emphasis>
</td>
<td id="76917505B005F58CB45DFD135E71FD4D" box="[1102,1140,730,754]" gridcol="6" gridrow="8" pageId="23" pageNumber="102">1.9</td>
<td id="76917505B005F58CB48CFD135EC5FD4D" box="[1183,1216,730,754]" gridcol="7" gridrow="8" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFD135EC5FD4D" box="[1183,1216,730,754]" italics="true" pageId="23" pageNumber="102">1.9</emphasis>
</td>
<td id="76917505B005F58CB4FBFD135F12FD4D" box="[1256,1303,730,754]" gridcol="8" gridrow="8" pageId="23" pageNumber="102">23.4</td>
<td id="76917505B005F58CB527FD135F6CFD4D" box="[1332,1385,730,754]" gridcol="9" gridrow="8" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FD135F63FD4D" box="[1335,1382,730,754]" italics="true" pageId="23" pageNumber="102">58.4</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FCC95F6CFCA7" box="[197,1385,768,792]" gridrow="9" pageId="23" pageNumber="102" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76917505B005F58CB0D6FCC95824FCA7" box="[197,545,768,792]" gridcol="0" gridrow="9" pageId="23" pageNumber="102">Placoid lengths</th>
</tr>
<tr id="35401C79B005F58CB0D6FCEC5F6CFC81" box="[197,1385,805,830]" gridrow="10" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FCEC5824FC81" box="[197,545,805,830]" gridcol="0" gridrow="10" pageId="23" pageNumber="102">Macroplacoid 1</th>
<td id="76917505B005F58CB25BFCEC5866FC81" box="[584,611,805,830]" gridcol="1" gridrow="10" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFCEC58FDFC81" box="[653,760,805,830]" gridcol="2" gridrow="10" pageId="23" pageNumber="102">7.012.2</td>
<td id="76917505B005F58CB304FCEC598BFC81" box="[791,910,805,830]" gridcol="3" gridrow="10" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFCEF598DFC81" box="[797,904,806,830]" italics="true" pageId="23" pageNumber="102">23.530.2</emphasis>
</td>
<td id="76917505B005F58CB3B8FCEC59DFFC81" box="[939,986,805,830]" gridcol="4" gridrow="10" pageId="23" pageNumber="102">9.7</td>
<td id="76917505B005F58CB3E4FCEC5E29FC81" box="[1015,1068,805,830]" gridcol="5" gridrow="10" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FCEC5E2CFC82" box="[1018,1065,805,829]" italics="true" pageId="23" pageNumber="102">26.9</emphasis>
</td>
<td id="76917505B005F58CB45DFCEC5E71FC81" box="[1102,1140,805,830]" gridcol="6" gridrow="10" pageId="23" pageNumber="102">1.6</td>
<td id="76917505B005F58CB48CFCEC5EC5FC81" box="[1183,1216,805,830]" gridcol="7" gridrow="10" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFCEF5EC5FC81" box="[1183,1216,806,830]" italics="true" pageId="23" pageNumber="102">2.2</emphasis>
</td>
<td id="76917505B005F58CB4FBFCEC5F12FC81" box="[1256,1303,805,830]" gridcol="8" gridrow="10" pageId="23" pageNumber="102">12.1</td>
<td id="76917505B005F58CB527FCEC5F6CFC81" box="[1332,1385,805,830]" gridcol="9" gridrow="10" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FCEF5F63FC81" box="[1335,1382,806,830]" italics="true" pageId="23" pageNumber="102">30.2</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FC825F6CFCDB" box="[197,1385,843,868]" gridrow="11" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FC825824FCDB" box="[197,545,843,868]" gridcol="0" gridrow="11" pageId="23" pageNumber="102">Macroplacoid 2</th>
<td id="76917505B005F58CB25BFC825866FCDB" box="[584,611,843,868]" gridcol="1" gridrow="11" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFC8258FDFCDB" box="[653,760,843,868]" gridcol="2" gridrow="11" pageId="23" pageNumber="102">4.67.5</td>
<td id="76917505B005F58CB304FC82598BFCDB" box="[791,910,843,868]" gridcol="3" gridrow="11" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFC855982FCDB" box="[797,903,844,868]" italics="true" pageId="23" pageNumber="102">12.320.8</emphasis>
</td>
<td id="76917505B005F58CB3B8FC8259DFFCDB" box="[939,986,843,868]" gridcol="4" gridrow="11" pageId="23" pageNumber="102">6.1</td>
<td id="76917505B005F58CB3E4FC825E29FCDB" box="[1015,1068,843,868]" gridcol="5" gridrow="11" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FC855E2CFCDB" box="[1018,1065,844,868]" italics="true" pageId="23" pageNumber="102">17.1</emphasis>
</td>
<td id="76917505B005F58CB45DFC825E71FCDB" box="[1102,1140,843,868]" gridcol="6" gridrow="11" pageId="23" pageNumber="102">0.9</td>
<td id="76917505B005F58CB48CFC825EC5FCDB" box="[1183,1216,843,868]" gridcol="7" gridrow="11" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFC855EC5FCDB" box="[1183,1216,844,868]" italics="true" pageId="23" pageNumber="102">1.8</emphasis>
</td>
<td id="76917505B005F58CB4FBFC825F12FCDB" box="[1256,1303,843,868]" gridcol="8" gridrow="11" pageId="23" pageNumber="102">7.3</td>
<td id="76917505B005F58CB527FC825F6CFCDB" box="[1332,1385,843,868]" gridcol="9" gridrow="11" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FC855F63FCDB" box="[1335,1382,844,868]" italics="true" pageId="23" pageNumber="102">18.2</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FCB85F6CFC36" box="[197,1385,881,905]" gridrow="12" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FCB85824FC36" box="[197,545,881,905]" gridcol="0" gridrow="12" pageId="23" pageNumber="102">Microplacoid</th>
<td id="76917505B005F58CB25BFCB85866FC36" box="[584,611,881,905]" gridcol="1" gridrow="12" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFCB858FDFC36" box="[653,760,881,905]" gridcol="2" gridrow="12" pageId="23" pageNumber="102">2.54.9</td>
<td id="76917505B005F58CB304FCB8598BFC36" box="[791,910,881,905]" gridcol="3" gridrow="12" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B337FCB85984FC36" box="[804,897,881,905]" italics="true" pageId="23" pageNumber="102">8.413.3</emphasis>
</td>
<td id="76917505B005F58CB3B8FCB859DFFC36" box="[939,986,881,905]" gridcol="4" gridrow="12" pageId="23" pageNumber="102">3.8</td>
<td id="76917505B005F58CB3E4FCB85E29FC36" box="[1015,1068,881,905]" gridcol="5" gridrow="12" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FCB85E2CFC36" box="[1018,1065,881,905]" italics="true" pageId="23" pageNumber="102">10.5</emphasis>
</td>
<td id="76917505B005F58CB45DFCB85E71FC36" box="[1102,1140,881,905]" gridcol="6" gridrow="12" pageId="23" pageNumber="102">0.6</td>
<td id="76917505B005F58CB48CFCB85EC5FC36" box="[1183,1216,881,905]" gridcol="7" gridrow="12" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFCB85EC5FC36" box="[1183,1216,881,905]" italics="true" pageId="23" pageNumber="102">1.2</emphasis>
</td>
<td id="76917505B005F58CB4FBFCB85F12FC36" box="[1256,1303,881,905]" gridcol="8" gridrow="12" pageId="23" pageNumber="102">4.1</td>
<td id="76917505B005F58CB527FCB85F6CFC36" box="[1332,1385,881,905]" gridcol="9" gridrow="12" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FCB85F63FC36" box="[1335,1382,881,905]" italics="true" pageId="23" pageNumber="102">10.2</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FC5E5F6CFC10" box="[197,1385,919,943]" gridrow="13" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FC5E5824FC10" box="[197,545,919,943]" gridcol="0" gridrow="13" pageId="23" pageNumber="102">Macroplacoid row</th>
<td id="76917505B005F58CB25BFC5E5866FC10" box="[584,611,919,943]" gridcol="1" gridrow="13" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFC5E58FDFC10" box="[653,760,919,943]" gridcol="2" gridrow="13" pageId="23" pageNumber="102">12.920.9</td>
<td id="76917505B005F58CB304FC5E598BFC10" box="[791,910,919,943]" gridcol="3" gridrow="13" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFC5E598DFC10" box="[797,904,919,943]" italics="true" pageId="23" pageNumber="102">42.752.2</emphasis>
</td>
<td id="76917505B005F58CB3B8FC5E59DFFC10" box="[939,986,919,943]" gridcol="4" gridrow="13" pageId="23" pageNumber="102">17.2</td>
<td id="76917505B005F58CB3E4FC5E5E29FC10" box="[1015,1068,919,943]" gridcol="5" gridrow="13" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FC5E5E2CFC10" box="[1018,1065,919,943]" italics="true" pageId="23" pageNumber="102">47.9</emphasis>
</td>
<td id="76917505B005F58CB45DFC5E5E71FC10" box="[1102,1140,919,943]" gridcol="6" gridrow="13" pageId="23" pageNumber="102">2.4</td>
<td id="76917505B005F58CB48CFC5E5EC5FC10" box="[1183,1216,919,943]" gridcol="7" gridrow="13" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFC5E5EC5FC10" box="[1183,1216,919,943]" italics="true" pageId="23" pageNumber="102">2.9</emphasis>
</td>
<td id="76917505B005F58CB4FBFC5E5F12FC10" box="[1256,1303,919,943]" gridcol="8" gridrow="13" pageId="23" pageNumber="102">20.9</td>
<td id="76917505B005F58CB527FC5E5F6CFC10" box="[1332,1385,919,943]" gridcol="9" gridrow="13" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FC5E5F63FC10" box="[1335,1382,919,943]" italics="true" pageId="23" pageNumber="102">52.1</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FC745F6CFC6A" box="[197,1385,957,981]" gridrow="14" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FC745824FC6A" box="[197,545,957,981]" gridcol="0" gridrow="14" pageId="23" pageNumber="102">Placoid row</th>
<td id="76917505B005F58CB25BFC745866FC6A" box="[584,611,957,981]" gridcol="1" gridrow="14" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EFC7458FDFC6A" box="[653,760,957,981]" gridcol="2" gridrow="14" pageId="23" pageNumber="102">16.326.6</td>
<td id="76917505B005F58CB304FC74598BFC6A" box="[791,910,957,981]" gridcol="3" gridrow="14" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFC745982FC6A" box="[797,903,957,981]" italics="true" pageId="23" pageNumber="102">54.367.7</emphasis>
</td>
<td id="76917505B005F58CB3B8FC7459DFFC6A" box="[939,986,957,981]" gridcol="4" gridrow="14" pageId="23" pageNumber="102">22.1</td>
<td id="76917505B005F58CB3E4FC745E29FC6A" box="[1015,1068,957,981]" gridcol="5" gridrow="14" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FC745E2CFC6A" box="[1018,1065,957,981]" italics="true" pageId="23" pageNumber="102">61.6</emphasis>
</td>
<td id="76917505B005F58CB45DFC745E71FC6A" box="[1102,1140,957,981]" gridcol="6" gridrow="14" pageId="23" pageNumber="102">3.0</td>
<td id="76917505B005F58CB48CFC745EC5FC6A" box="[1183,1216,957,981]" gridcol="7" gridrow="14" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFC745EC5FC6A" box="[1183,1216,957,981]" italics="true" pageId="23" pageNumber="102">3.8</emphasis>
</td>
<td id="76917505B005F58CB4FBFC745F12FC6A" box="[1256,1303,957,981]" gridcol="8" gridrow="14" pageId="23" pageNumber="102">26.6</td>
<td id="76917505B005F58CB527FC745F6CFC6A" box="[1332,1385,957,981]" gridcol="9" gridrow="14" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FC745F63FC6A" box="[1335,1382,957,981]" italics="true" pageId="23" pageNumber="102">66.3</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FC2A5F6CFC44" box="[197,1385,995,1019]" gridrow="15" pageId="23" pageNumber="102" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="76917505B005F58CB0D6FC2A5824FC44" box="[197,545,995,1019]" gridcol="0" gridrow="15" pageId="23" pageNumber="102">Claw I heights</th>
</tr>
<tr id="35401C79B005F58CB0D6FBC15F6CFB9F" box="[197,1385,1032,1056]" gridrow="16" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FBC15824FB9F" box="[197,545,1032,1056]" gridcol="0" gridrow="16" pageId="23" pageNumber="102">External primary branch</th>
<td id="76917505B005F58CB25BFBC15866FB9F" box="[584,611,1032,1056]" gridcol="1" gridrow="16" pageId="23" pageNumber="102">18</td>
<td id="76917505B005F58CB29EFBC158FDFB9F" box="[653,760,1032,1056]" gridcol="2" gridrow="16" pageId="23" pageNumber="102">7.212.9</td>
<td id="76917505B005F58CB304FBC1598BFB9F" box="[791,910,1032,1056]" gridcol="3" gridrow="16" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFBC15982FB9F" box="[797,903,1032,1056]" italics="true" pageId="23" pageNumber="102">24.434.1</emphasis>
</td>
<td id="76917505B005F58CB3B8FBC159DFFB9F" box="[939,986,1032,1056]" gridcol="4" gridrow="16" pageId="23" pageNumber="102">10.5</td>
<td id="76917505B005F58CB3E4FBC15E29FB9F" box="[1015,1068,1032,1056]" gridcol="5" gridrow="16" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FBC15E2CFB9F" box="[1018,1065,1032,1056]" italics="true" pageId="23" pageNumber="102">29.0</emphasis>
</td>
<td id="76917505B005F58CB45DFBC15E71FB9F" box="[1102,1140,1032,1056]" gridcol="6" gridrow="16" pageId="23" pageNumber="102">1.4</td>
<td id="76917505B005F58CB48CFBC15EC5FB9F" box="[1183,1216,1032,1056]" gridcol="7" gridrow="16" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFBC15EC5FB9F" box="[1183,1216,1032,1056]" italics="true" pageId="23" pageNumber="102">2.3</emphasis>
</td>
<td id="76917505B005F58CB4FBFBC15F12FB9F" box="[1256,1303,1032,1056]" gridcol="8" gridrow="16" pageId="23" pageNumber="102">11.0</td>
<td id="76917505B005F58CB527FBC15F6CFB9F" box="[1332,1385,1032,1056]" gridcol="9" gridrow="16" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FBC15F63FB9F" box="[1335,1382,1032,1056]" italics="true" pageId="23" pageNumber="102">27.4</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FBE75F6CFBF9" box="[197,1385,1070,1094]" gridrow="17" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FBE75824FBF9" box="[197,545,1070,1094]" gridcol="0" gridrow="17" pageId="23" pageNumber="102">External secondary branch</th>
<td id="76917505B005F58CB25BFBE75866FBF9" box="[584,611,1070,1094]" gridcol="1" gridrow="17" pageId="23" pageNumber="102">11</td>
<td id="76917505B005F58CB29EFBE758FDFBF9" box="[653,760,1070,1094]" gridcol="2" gridrow="17" pageId="23" pageNumber="102">7.110.0</td>
<td id="76917505B005F58CB304FBE7598BFBF9" box="[791,910,1070,1094]" gridcol="3" gridrow="17" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFBE7598DFBF9" box="[797,904,1070,1094]" italics="true" pageId="23" pageNumber="102">22.830.2</emphasis>
</td>
<td id="76917505B005F58CB3B8FBE759DFFBF9" box="[939,986,1070,1094]" gridcol="4" gridrow="17" pageId="23" pageNumber="102">9.1</td>
<td id="76917505B005F58CB3E4FBE75E29FBF9" box="[1015,1068,1070,1094]" gridcol="5" gridrow="17" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FBE75E2CFBF9" box="[1018,1065,1070,1094]" italics="true" pageId="23" pageNumber="102">25.2</emphasis>
</td>
<td id="76917505B005F58CB45DFBE75E71FBF9" box="[1102,1140,1070,1094]" gridcol="6" gridrow="17" pageId="23" pageNumber="102">0.9</td>
<td id="76917505B005F58CB48CFBE75EC5FBF9" box="[1183,1216,1070,1094]" gridcol="7" gridrow="17" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFBE75EC5FBF9" box="[1183,1216,1070,1094]" italics="true" pageId="23" pageNumber="102">2.1</emphasis>
</td>
<td id="76917505B005F58CB4FBFBE75F12FBF9" box="[1256,1303,1070,1094]" gridcol="8" gridrow="17" pageId="23" pageNumber="102">10.0</td>
<td id="76917505B005F58CB527FBE75F6CFBF9" box="[1332,1385,1070,1094]" gridcol="9" gridrow="17" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FBE75F63FBF9" box="[1335,1382,1070,1094]" italics="true" pageId="23" pageNumber="102">24.9</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FB9D5F6CFBD3" box="[197,1385,1108,1132]" gridrow="18" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FB9D5824FBD3" box="[197,545,1108,1132]" gridcol="0" gridrow="18" pageId="23" pageNumber="102">Internal primary branch</th>
<td id="76917505B005F58CB25BFB9D5866FBD3" box="[584,611,1108,1132]" gridcol="1" gridrow="18" pageId="23" pageNumber="102">18</td>
<td id="76917505B005F58CB29EFB9D58FDFBD3" box="[653,760,1108,1132]" gridcol="2" gridrow="18" pageId="23" pageNumber="102">7.111.5</td>
<td id="76917505B005F58CB304FB9D598BFBD3" box="[791,910,1108,1132]" gridcol="3" gridrow="18" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFB9D598DFBD3" box="[797,904,1108,1132]" italics="true" pageId="23" pageNumber="102">24.131.1</emphasis>
</td>
<td id="76917505B005F58CB3B8FB9D59DFFBD3" box="[939,986,1108,1132]" gridcol="4" gridrow="18" pageId="23" pageNumber="102">9.9</td>
<td id="76917505B005F58CB3E4FB9D5E29FBD3" box="[1015,1068,1108,1132]" gridcol="5" gridrow="18" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FB9D5E2CFBD3" box="[1018,1065,1108,1132]" italics="true" pageId="23" pageNumber="102">27.3</emphasis>
</td>
<td id="76917505B005F58CB45DFB9D5E71FBD3" box="[1102,1140,1108,1132]" gridcol="6" gridrow="18" pageId="23" pageNumber="102">1.1</td>
<td id="76917505B005F58CB48CFB9D5EC5FBD3" box="[1183,1216,1108,1132]" gridcol="7" gridrow="18" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFB9D5EC5FBD3" box="[1183,1216,1108,1132]" italics="true" pageId="23" pageNumber="102">1.6</emphasis>
</td>
<td id="76917505B005F58CB4FBFB9D5F12FBD3" box="[1256,1303,1108,1132]" gridcol="8" gridrow="18" pageId="23" pageNumber="102">10.7</td>
<td id="76917505B005F58CB527FB9D5F6CFBD3" box="[1332,1385,1108,1132]" gridcol="9" gridrow="18" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FB9D5F63FBD3" box="[1335,1382,1108,1132]" italics="true" pageId="23" pageNumber="102">26.7</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FBB35F6CFB2D" box="[197,1385,1146,1170]" gridrow="19" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FBB35824FB2D" box="[197,545,1146,1170]" gridcol="0" gridrow="19" pageId="23" pageNumber="102">Internal secondary branch</th>
<td id="76917505B005F58CB25BFBB35866FB2D" box="[584,611,1146,1170]" gridcol="1" gridrow="19" pageId="23" pageNumber="102">12</td>
<td id="76917505B005F58CB29EFBB358FDFB2D" box="[653,760,1146,1170]" gridcol="2" gridrow="19" pageId="23" pageNumber="102">7.010.1</td>
<td id="76917505B005F58CB304FBB3598BFB2D" box="[791,910,1146,1170]" gridcol="3" gridrow="19" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFBB35982FB2D" box="[797,903,1146,1170]" italics="true" pageId="23" pageNumber="102">21.427.1</emphasis>
</td>
<td id="76917505B005F58CB3B8FBB359DFFB2D" box="[939,986,1146,1170]" gridcol="4" gridrow="19" pageId="23" pageNumber="102">9.0</td>
<td id="76917505B005F58CB3E4FBB35E29FB2D" box="[1015,1068,1146,1170]" gridcol="5" gridrow="19" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FBB35E2CFB2D" box="[1018,1065,1146,1170]" italics="true" pageId="23" pageNumber="102">24.3</emphasis>
</td>
<td id="76917505B005F58CB45DFBB35E71FB2D" box="[1102,1140,1146,1170]" gridcol="6" gridrow="19" pageId="23" pageNumber="102">1.0</td>
<td id="76917505B005F58CB48CFBB35EC5FB2D" box="[1183,1216,1146,1170]" gridcol="7" gridrow="19" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFBB35EC5FB2D" box="[1183,1216,1146,1170]" italics="true" pageId="23" pageNumber="102">1.6</emphasis>
</td>
<td id="76917505B005F58CB4FBFBB35F12FB2D" box="[1256,1303,1146,1170]" gridcol="8" gridrow="19" pageId="23" pageNumber="102">9.1</td>
<td id="76917505B005F58CB527FBB35F6CFB2D" box="[1332,1385,1146,1170]" gridcol="9" gridrow="19" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FBB35F63FB2D" box="[1335,1382,1146,1170]" italics="true" pageId="23" pageNumber="102">22.7</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FB565F6CFB08" box="[197,1385,1183,1207]" gridrow="20" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FB565F6CFB08" box="[197,1385,1183,1207]" colspan="10" colspanRight="9" gridcol="0" gridrow="20" pageId="23" pageNumber="102">Claw II heights</th>
</tr>
<tr id="35401C79B005F58CB0D6FB0C5F6CFB62" box="[197,1385,1221,1245]" gridrow="21" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FB0C5824FB62" box="[197,545,1221,1245]" gridcol="0" gridrow="21" pageId="23" pageNumber="102">External primary branch</th>
<td id="76917505B005F58CB25BFB0C5866FB62" box="[584,611,1221,1245]" gridcol="1" gridrow="21" pageId="23" pageNumber="102">18</td>
<td id="76917505B005F58CB29EFB0C58FDFB62" box="[653,760,1221,1245]" gridcol="2" gridrow="21" pageId="23" pageNumber="102">8.313.0</td>
<td id="76917505B005F58CB304FB0C598BFB62" box="[791,910,1221,1245]" gridcol="3" gridrow="21" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFB0C598DFB62" box="[797,904,1221,1245]" italics="true" pageId="23" pageNumber="102">26.838.1</emphasis>
</td>
<td id="76917505B005F58CB3B8FB0C59DFFB62" box="[939,986,1221,1245]" gridcol="4" gridrow="21" pageId="23" pageNumber="102">11.3</td>
<td id="76917505B005F58CB3E4FB0C5E29FB62" box="[1015,1068,1221,1245]" gridcol="5" gridrow="21" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FB0C5E2CFB62" box="[1018,1065,1221,1245]" italics="true" pageId="23" pageNumber="102">31.3</emphasis>
</td>
<td id="76917505B005F58CB45DFB0C5E71FB62" box="[1102,1140,1221,1245]" gridcol="6" gridrow="21" pageId="23" pageNumber="102">1.5</td>
<td id="76917505B005F58CB48CFB0C5EC5FB62" box="[1183,1216,1221,1245]" gridcol="7" gridrow="21" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFB0C5EC5FB62" box="[1183,1216,1221,1245]" italics="true" pageId="23" pageNumber="102">2.8</emphasis>
</td>
<td id="76917505B005F58CB4FBFB0C5F12FB62" box="[1256,1303,1221,1245]" gridcol="8" gridrow="21" pageId="23" pageNumber="102">12.5</td>
<td id="76917505B005F58CB527FB0C5F6CFB62" box="[1332,1385,1221,1245]" gridcol="9" gridrow="21" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FB0C5F63FB62" box="[1335,1382,1221,1245]" italics="true" pageId="23" pageNumber="102">31.2</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FB225F6CFABC" box="[197,1385,1259,1283]" gridrow="22" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FB225824FABC" box="[197,545,1259,1283]" gridcol="0" gridrow="22" pageId="23" pageNumber="102">External secondary branch</th>
<td id="76917505B005F58CB25BFB225866FABC" box="[584,611,1259,1283]" gridcol="1" gridrow="22" pageId="23" pageNumber="102">10</td>
<td id="76917505B005F58CB29EFB2258FDFABC" box="[653,760,1259,1283]" gridcol="2" gridrow="22" pageId="23" pageNumber="102">7.712.0</td>
<td id="76917505B005F58CB304FB22598BFABC" box="[791,910,1259,1283]" gridcol="3" gridrow="22" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFB22598DFABC" box="[797,904,1259,1283]" italics="true" pageId="23" pageNumber="102">23.132.3</emphasis>
</td>
<td id="76917505B005F58CB3B8FB2259DFFABC" box="[939,986,1259,1283]" gridcol="4" gridrow="22" pageId="23" pageNumber="102">10.0</td>
<td id="76917505B005F58CB3E4FB225E29FABC" box="[1015,1068,1259,1283]" gridcol="5" gridrow="22" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FB225E2CFABC" box="[1018,1065,1259,1283]" italics="true" pageId="23" pageNumber="102">27.2</emphasis>
</td>
<td id="76917505B005F58CB45DFB225E71FABC" box="[1102,1140,1259,1283]" gridcol="6" gridrow="22" pageId="23" pageNumber="102">1.4</td>
<td id="76917505B005F58CB48CFB225EC5FABC" box="[1183,1216,1259,1283]" gridcol="7" gridrow="22" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFB225EC5FABC" box="[1183,1216,1259,1283]" italics="true" pageId="23" pageNumber="102">3.0</emphasis>
</td>
<td id="76917505B005F58CB4FBFB225F12FABC" box="[1256,1303,1259,1283]" gridcol="8" gridrow="22" pageId="23" pageNumber="102">10.9</td>
<td id="76917505B005F58CB527FB225F6CFABC" box="[1332,1385,1259,1283]" gridcol="9" gridrow="22" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FB225F63FABC" box="[1335,1382,1259,1283]" italics="true" pageId="23" pageNumber="102">27.2</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FAD85F6CFA96" box="[197,1385,1297,1321]" gridrow="23" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FAD85824FA96" box="[197,545,1297,1321]" gridcol="0" gridrow="23" pageId="23" pageNumber="102">Internal primary branch</th>
<td id="76917505B005F58CB25BFAD85866FA96" box="[584,611,1297,1321]" gridcol="1" gridrow="23" pageId="23" pageNumber="102">15</td>
<td id="76917505B005F58CB29EFAD858FDFA96" box="[653,760,1297,1321]" gridcol="2" gridrow="23" pageId="23" pageNumber="102">7.411.0</td>
<td id="76917505B005F58CB304FAD8598BFA96" box="[791,910,1297,1321]" gridcol="3" gridrow="23" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFAD85982FA96" box="[797,903,1297,1321]" italics="true" pageId="23" pageNumber="102">24.829.5</emphasis>
</td>
<td id="76917505B005F58CB3B8FAD859DFFA96" box="[939,986,1297,1321]" gridcol="4" gridrow="23" pageId="23" pageNumber="102">9.7</td>
<td id="76917505B005F58CB3E4FAD85E29FA96" box="[1015,1068,1297,1321]" gridcol="5" gridrow="23" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FAD85E2CFA96" box="[1018,1065,1297,1321]" italics="true" pageId="23" pageNumber="102">26.9</emphasis>
</td>
<td id="76917505B005F58CB45DFAD85E71FA96" box="[1102,1140,1297,1321]" gridcol="6" gridrow="23" pageId="23" pageNumber="102">1.1</td>
<td id="76917505B005F58CB48CFAD85EC5FA96" box="[1183,1216,1297,1321]" gridcol="7" gridrow="23" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFAD85EC5FA96" box="[1183,1216,1297,1321]" italics="true" pageId="23" pageNumber="102">1.4</emphasis>
</td>
<td id="76917505B005F58CB4FBFAD85F12FA96" box="[1256,1303,1297,1321]" gridcol="8" gridrow="23" pageId="23" pageNumber="102">10.1</td>
<td id="76917505B005F58CB527FAD85F6CFA96" box="[1332,1385,1297,1321]" gridcol="9" gridrow="23" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FAD85F63FA96" box="[1335,1382,1297,1321]" italics="true" pageId="23" pageNumber="102">25.2</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FAFE5F6CFAF0" box="[197,1385,1335,1359]" gridrow="24" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FAFE5824FAF0" box="[197,545,1335,1359]" gridcol="0" gridrow="24" pageId="23" pageNumber="102">Internal secondary branch</th>
<td id="76917505B005F58CB25BFAFE5866FAF0" box="[584,611,1335,1359]" gridcol="1" gridrow="24" pageId="23" pageNumber="102">10</td>
<td id="76917505B005F58CB29EFAFE58FDFAF0" box="[653,760,1335,1359]" gridcol="2" gridrow="24" pageId="23" pageNumber="102">6.210.2</td>
<td id="76917505B005F58CB304FAFE598BFAF0" box="[791,910,1335,1359]" gridcol="3" gridrow="24" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFAFE598DFAF0" box="[797,904,1335,1359]" italics="true" pageId="23" pageNumber="102">20.827.3</emphasis>
</td>
<td id="76917505B005F58CB3B8FAFE59DFFAF0" box="[939,986,1335,1359]" gridcol="4" gridrow="24" pageId="23" pageNumber="102">8.9</td>
<td id="76917505B005F58CB3E4FAFE5E29FAF0" box="[1015,1068,1335,1359]" gridcol="5" gridrow="24" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FAFE5E2CFAF0" box="[1018,1065,1335,1359]" italics="true" pageId="23" pageNumber="102">23.9</emphasis>
</td>
<td id="76917505B005F58CB45DFAFE5E71FAF0" box="[1102,1140,1335,1359]" gridcol="6" gridrow="24" pageId="23" pageNumber="102">1.2</td>
<td id="76917505B005F58CB48CFAFE5EC5FAF0" box="[1183,1216,1335,1359]" gridcol="7" gridrow="24" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFAFE5EC5FAF0" box="[1183,1216,1335,1359]" italics="true" pageId="23" pageNumber="102">2.1</emphasis>
</td>
<td id="76917505B005F58CB4FBFAFE5F12FAF0" box="[1256,1303,1335,1359]" gridcol="8" gridrow="24" pageId="23" pageNumber="102">9.8</td>
<td id="76917505B005F58CB527FAFE5F6CFAF0" box="[1332,1385,1335,1359]" gridcol="9" gridrow="24" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FAFE5F63FAF0" box="[1335,1382,1335,1359]" italics="true" pageId="23" pageNumber="102">24.4</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FA955F6CFACB" box="[197,1385,1372,1396]" gridrow="25" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FA955F6CFACB" box="[197,1385,1372,1396]" colspan="10" colspanRight="9" gridcol="0" gridrow="25" pageId="23" pageNumber="102">Claw III heights</th>
</tr>
<tr id="35401C79B005F58CB0D6FA4B5F6CFA25" box="[197,1385,1410,1434]" gridrow="26" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FA4B5824FA25" box="[197,545,1410,1434]" gridcol="0" gridrow="26" pageId="23" pageNumber="102">External primary branch</th>
<td id="76917505B005F58CB25BFA4B5866FA25" box="[584,611,1410,1434]" gridcol="1" gridrow="26" pageId="23" pageNumber="102">16</td>
<td id="76917505B005F58CB29EFA4B58FDFA25" box="[653,760,1410,1434]" gridcol="2" gridrow="26" pageId="23" pageNumber="102">8.112.9</td>
<td id="76917505B005F58CB304FA4B598BFA25" box="[791,910,1410,1434]" gridcol="3" gridrow="26" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFA4B5982FA25" box="[797,903,1410,1434]" italics="true" pageId="23" pageNumber="102">27.234.6</emphasis>
</td>
<td id="76917505B005F58CB3B8FA4B59DFFA25" box="[939,986,1410,1434]" gridcol="4" gridrow="26" pageId="23" pageNumber="102">11.2</td>
<td id="76917505B005F58CB3E4FA4B5E29FA25" box="[1015,1068,1410,1434]" gridcol="5" gridrow="26" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FA4B5E2CFA25" box="[1018,1065,1410,1434]" italics="true" pageId="23" pageNumber="102">30.9</emphasis>
</td>
<td id="76917505B005F58CB45DFA4B5E71FA25" box="[1102,1140,1410,1434]" gridcol="6" gridrow="26" pageId="23" pageNumber="102">1.2</td>
<td id="76917505B005F58CB48CFA4B5EC5FA25" box="[1183,1216,1410,1434]" gridcol="7" gridrow="26" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFA4B5EC5FA25" box="[1183,1216,1410,1434]" italics="true" pageId="23" pageNumber="102">1.9</emphasis>
</td>
<td id="76917505B005F58CB4FBFA4B5F12FA25" box="[1256,1303,1410,1434]" gridcol="8" gridrow="26" pageId="23" pageNumber="102">12.3</td>
<td id="76917505B005F58CB527FA4B5F6CFA25" box="[1332,1385,1410,1434]" gridcol="9" gridrow="26" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FA4B5F63FA25" box="[1335,1382,1410,1434]" italics="true" pageId="23" pageNumber="102">30.7</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FA615F6CFA7F" box="[197,1385,1448,1472]" gridrow="27" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FA615824FA7F" box="[197,545,1448,1472]" gridcol="0" gridrow="27" pageId="23" pageNumber="102">External secondary branch</th>
<td id="76917505B005F58CB25BFA615866FA7F" box="[584,611,1448,1472]" gridcol="1" gridrow="27" pageId="23" pageNumber="102">11</td>
<td id="76917505B005F58CB29EFA6158FDFA7F" box="[653,760,1448,1472]" gridcol="2" gridrow="27" pageId="23" pageNumber="102">7.511.9</td>
<td id="76917505B005F58CB304FA61598BFA7F" box="[791,910,1448,1472]" gridcol="3" gridrow="27" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFA615982FA7F" box="[797,903,1448,1472]" italics="true" pageId="23" pageNumber="102">22.831.9</emphasis>
</td>
<td id="76917505B005F58CB3B8FA6159DFFA7F" box="[939,986,1448,1472]" gridcol="4" gridrow="27" pageId="23" pageNumber="102">10.0</td>
<td id="76917505B005F58CB3E4FA615E29FA7F" box="[1015,1068,1448,1472]" gridcol="5" gridrow="27" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FA615E2CFA7F" box="[1018,1065,1448,1472]" italics="true" pageId="23" pageNumber="102">27.1</emphasis>
</td>
<td id="76917505B005F58CB45DFA615E71FA7F" box="[1102,1140,1448,1472]" gridcol="6" gridrow="27" pageId="23" pageNumber="102">1.3</td>
<td id="76917505B005F58CB48CFA615EC5FA7F" box="[1183,1216,1448,1472]" gridcol="7" gridrow="27" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFA615EC5FA7F" box="[1183,1216,1448,1472]" italics="true" pageId="23" pageNumber="102">2.6</emphasis>
</td>
<td id="76917505B005F58CB4FBFA615F12FA7F" box="[1256,1303,1448,1472]" gridcol="8" gridrow="27" pageId="23" pageNumber="102">10.6</td>
<td id="76917505B005F58CB527FA615F6CFA7F" box="[1332,1385,1448,1472]" gridcol="9" gridrow="27" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FA615F63FA7F" box="[1335,1382,1448,1472]" italics="true" pageId="23" pageNumber="102">26.4</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FA075F6CFA59" box="[197,1385,1486,1510]" gridrow="28" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FA075824FA59" box="[197,545,1486,1510]" gridcol="0" gridrow="28" pageId="23" pageNumber="102">Internal primary branch</th>
<td id="76917505B005F58CB25BFA075866FA59" box="[584,611,1486,1510]" gridcol="1" gridrow="28" pageId="23" pageNumber="102">13</td>
<td id="76917505B005F58CB29EFA0758FDFA59" box="[653,760,1486,1510]" gridcol="2" gridrow="28" pageId="23" pageNumber="102">7.311.2</td>
<td id="76917505B005F58CB304FA07598BFA59" box="[791,910,1486,1510]" gridcol="3" gridrow="28" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFA07598DFA59" box="[797,904,1486,1510]" italics="true" pageId="23" pageNumber="102">24.530.0</emphasis>
</td>
<td id="76917505B005F58CB3B8FA0759DFFA59" box="[939,986,1486,1510]" gridcol="4" gridrow="28" pageId="23" pageNumber="102">9.9</td>
<td id="76917505B005F58CB3E4FA075E29FA59" box="[1015,1068,1486,1510]" gridcol="5" gridrow="28" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FA075E2CFA59" box="[1018,1065,1486,1510]" italics="true" pageId="23" pageNumber="102">27.2</emphasis>
</td>
<td id="76917505B005F58CB45DFA075E71FA59" box="[1102,1140,1486,1510]" gridcol="6" gridrow="28" pageId="23" pageNumber="102">1.2</td>
<td id="76917505B005F58CB48CFA075EC5FA59" box="[1183,1216,1486,1510]" gridcol="7" gridrow="28" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFA075EC5FA59" box="[1183,1216,1486,1510]" italics="true" pageId="23" pageNumber="102">1.5</emphasis>
</td>
<td id="76917505B005F58CB4FBFA075F12FA59" box="[1256,1303,1486,1510]" gridcol="8" gridrow="28" pageId="23" pageNumber="102">10.8</td>
<td id="76917505B005F58CB527FA075F6CFA59" box="[1332,1385,1486,1510]" gridcol="9" gridrow="28" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FA075F63FA59" box="[1335,1382,1486,1510]" italics="true" pageId="23" pageNumber="102">26.9</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6FA3D5F6CF9B3" box="[197,1385,1524,1548]" gridrow="29" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6FA3D5824F9B3" box="[197,545,1524,1548]" gridcol="0" gridrow="29" pageId="23" pageNumber="102">Internal secondary branch</th>
<td id="76917505B005F58CB25BFA3D5866F9B3" box="[584,611,1524,1548]" gridcol="1" gridrow="29" pageId="23" pageNumber="102">10</td>
<td id="76917505B005F58CB29EFA3D58FDF9B3" box="[653,760,1524,1548]" gridcol="2" gridrow="29" pageId="23" pageNumber="102">6.010.4</td>
<td id="76917505B005F58CB304FA3D598BF9B3" box="[791,910,1524,1548]" gridcol="3" gridrow="29" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EFA3D5982F9B3" box="[797,903,1524,1548]" italics="true" pageId="23" pageNumber="102">20.127.9</emphasis>
</td>
<td id="76917505B005F58CB3B8FA3D59DFF9B3" box="[939,986,1524,1548]" gridcol="4" gridrow="29" pageId="23" pageNumber="102">9.0</td>
<td id="76917505B005F58CB3E4FA3D5E29F9B3" box="[1015,1068,1524,1548]" gridcol="5" gridrow="29" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9FA3D5E2CF9B3" box="[1018,1065,1524,1548]" italics="true" pageId="23" pageNumber="102">24.0</emphasis>
</td>
<td id="76917505B005F58CB45DFA3D5E71F9B3" box="[1102,1140,1524,1548]" gridcol="6" gridrow="29" pageId="23" pageNumber="102">1.5</td>
<td id="76917505B005F58CB48CFA3D5EC5F9B3" box="[1183,1216,1524,1548]" gridcol="7" gridrow="29" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CFA3D5EC5F9B3" box="[1183,1216,1524,1548]" italics="true" pageId="23" pageNumber="102">2.6</emphasis>
</td>
<td id="76917505B005F58CB4FBFA3D5F12F9B3" box="[1256,1303,1524,1548]" gridcol="8" gridrow="29" pageId="23" pageNumber="102">10.0</td>
<td id="76917505B005F58CB527FA3D5F6CF9B3" box="[1332,1385,1524,1548]" gridcol="9" gridrow="29" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524FA3D5F63F9B3" box="[1335,1382,1524,1548]" italics="true" pageId="23" pageNumber="102">24.9</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6F9D05F6CF98E" box="[197,1385,1561,1585]" gridrow="30" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6F9D05F6CF98E" box="[197,1385,1561,1585]" colspan="10" colspanRight="9" gridcol="0" gridrow="30" pageId="23" pageNumber="102">Claw IV heights</th>
</tr>
<tr id="35401C79B005F58CB0D6F9F65F6CF9E8" box="[197,1385,1599,1623]" gridrow="31" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6F9F65824F9E8" box="[197,545,1599,1623]" gridcol="0" gridrow="31" pageId="23" pageNumber="102">Anterior primary branch</th>
<td id="76917505B005F58CB25BF9F65866F9E8" box="[584,611,1599,1623]" gridcol="1" gridrow="31" pageId="23" pageNumber="102">18</td>
<td id="76917505B005F58CB29EF9F658FDF9E8" box="[653,760,1599,1623]" gridcol="2" gridrow="31" pageId="23" pageNumber="102">7.412.1</td>
<td id="76917505B005F58CB304F9F6598BF9E8" box="[791,910,1599,1623]" gridcol="3" gridrow="31" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EF9F65982F9E8" box="[797,903,1599,1623]" italics="true" pageId="23" pageNumber="102">24.831.4</emphasis>
</td>
<td id="76917505B005F58CB3B8F9F659DFF9E8" box="[939,986,1599,1623]" gridcol="4" gridrow="31" pageId="23" pageNumber="102">10.4</td>
<td id="76917505B005F58CB3E4F9F65E29F9E8" box="[1015,1068,1599,1623]" gridcol="5" gridrow="31" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9F9F65E2CF9E8" box="[1018,1065,1599,1623]" italics="true" pageId="23" pageNumber="102">28.8</emphasis>
</td>
<td id="76917505B005F58CB45DF9F65E71F9E8" box="[1102,1140,1599,1623]" gridcol="6" gridrow="31" pageId="23" pageNumber="102">1.4</td>
<td id="76917505B005F58CB48CF9F65EC5F9E8" box="[1183,1216,1599,1623]" gridcol="7" gridrow="31" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CF9F65EC5F9E8" box="[1183,1216,1599,1623]" italics="true" pageId="23" pageNumber="102">2.0</emphasis>
</td>
<td id="76917505B005F58CB4FBF9F65F12F9E8" box="[1256,1303,1599,1623]" gridcol="8" gridrow="31" pageId="23" pageNumber="102">12.1</td>
<td id="76917505B005F58CB527F9F65F6CF9E8" box="[1332,1385,1599,1623]" gridcol="9" gridrow="31" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524F9F65F63F9E8" box="[1335,1382,1599,1623]" italics="true" pageId="23" pageNumber="102">30.2</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6F9AC5F6CF9C2" box="[197,1385,1637,1661]" gridrow="32" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6F9AC5824F9C2" box="[197,545,1637,1661]" gridcol="0" gridrow="32" pageId="23" pageNumber="102">Anterior secondary branch</th>
<td id="76917505B005F58CB25BF9AC5866F9C2" box="[584,611,1637,1661]" gridcol="1" gridrow="32" pageId="23" pageNumber="102">13</td>
<td id="76917505B005F58CB29EF9AC58FDF9C2" box="[653,760,1637,1661]" gridcol="2" gridrow="32" pageId="23" pageNumber="102">8.210.7</td>
<td id="76917505B005F58CB304F9AC598BF9C2" box="[791,910,1637,1661]" gridcol="3" gridrow="32" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EF9AC598DF9C2" box="[797,904,1637,1661]" italics="true" pageId="23" pageNumber="102">20.127.8</emphasis>
</td>
<td id="76917505B005F58CB3B8F9AC59DFF9C2" box="[939,986,1637,1661]" gridcol="4" gridrow="32" pageId="23" pageNumber="102">9.1</td>
<td id="76917505B005F58CB3E4F9AC5E29F9C2" box="[1015,1068,1637,1661]" gridcol="5" gridrow="32" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9F9AC5E2CF9C2" box="[1018,1065,1637,1661]" italics="true" pageId="23" pageNumber="102">24.5</emphasis>
</td>
<td id="76917505B005F58CB45DF9AC5E71F9C2" box="[1102,1140,1637,1661]" gridcol="6" gridrow="32" pageId="23" pageNumber="102">0.7</td>
<td id="76917505B005F58CB48CF9AC5EC5F9C2" box="[1183,1216,1637,1661]" gridcol="7" gridrow="32" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CF9AC5EC5F9C2" box="[1183,1216,1637,1661]" italics="true" pageId="23" pageNumber="102">2.1</emphasis>
</td>
<td id="76917505B005F58CB4FBF9AC5F12F9C2" box="[1256,1303,1637,1661]" gridcol="8" gridrow="32" pageId="23" pageNumber="102">10.7</td>
<td id="76917505B005F58CB527F9AC5F6CF9C2" box="[1332,1385,1637,1661]" gridcol="9" gridrow="32" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524F9AC5F63F9C2" box="[1335,1382,1637,1661]" italics="true" pageId="23" pageNumber="102">26.7</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6F9425F6CF91C" box="[197,1385,1675,1699]" gridrow="33" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6F9425824F91C" box="[197,545,1675,1699]" gridcol="0" gridrow="33" pageId="23" pageNumber="102">Posterior primary branch</th>
<td id="76917505B005F58CB25BF9425866F91C" box="[584,611,1675,1699]" gridcol="1" gridrow="33" pageId="23" pageNumber="102">20</td>
<td id="76917505B005F58CB29EF94258FDF91C" box="[653,760,1675,1699]" gridcol="2" gridrow="33" pageId="23" pageNumber="102">8.213.1</td>
<td id="76917505B005F58CB304F942598BF91C" box="[791,910,1675,1699]" gridcol="3" gridrow="33" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EF9425982F91C" box="[797,903,1675,1699]" italics="true" pageId="23" pageNumber="102">23.838.4</emphasis>
</td>
<td id="76917505B005F58CB3B8F94259DFF91C" box="[939,986,1675,1699]" gridcol="4" gridrow="33" pageId="23" pageNumber="102">11.1</td>
<td id="76917505B005F58CB3E4F9425E29F91C" box="[1015,1068,1675,1699]" gridcol="5" gridrow="33" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9F9425E2CF91C" box="[1018,1065,1675,1699]" italics="true" pageId="23" pageNumber="102">31.1</emphasis>
</td>
<td id="76917505B005F58CB45DF9425E71F91C" box="[1102,1140,1675,1699]" gridcol="6" gridrow="33" pageId="23" pageNumber="102">1.5</td>
<td id="76917505B005F58CB48CF9425EC5F91C" box="[1183,1216,1675,1699]" gridcol="7" gridrow="33" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CF9425EC5F91C" box="[1183,1216,1675,1699]" italics="true" pageId="23" pageNumber="102">3.2</emphasis>
</td>
<td id="76917505B005F58CB4FBF9425F12F91C" box="[1256,1303,1675,1699]" gridcol="8" gridrow="33" pageId="23" pageNumber="102">12.2</td>
<td id="76917505B005F58CB527F9425F6CF91C" box="[1332,1385,1675,1699]" gridcol="9" gridrow="33" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B524F9425F63F91C" box="[1335,1382,1675,1699]" italics="true" pageId="23" pageNumber="102">30.4</emphasis>
</td>
</tr>
<tr id="35401C79B005F58CB0D6F9785F6CF976" box="[197,1385,1713,1737]" gridrow="34" pageId="23" pageNumber="102">
<th id="76917505B005F58CB0D6F9785824F976" box="[197,545,1713,1737]" gridcol="0" gridrow="34" pageId="23" pageNumber="102">Posterior secondary branch</th>
<td id="76917505B005F58CB25BF9785866F976" box="[584,611,1713,1737]" gridcol="1" gridrow="34" pageId="23" pageNumber="102">9</td>
<td id="76917505B005F58CB29EF97858FDF976" box="[653,760,1713,1737]" gridcol="2" gridrow="34" pageId="23" pageNumber="102">6.910.9</td>
<td id="76917505B005F58CB304F978598BF976" box="[791,910,1713,1737]" gridcol="3" gridrow="34" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B30EF9785982F976" box="[797,903,1713,1737]" italics="true" pageId="23" pageNumber="102">20.827.9</emphasis>
</td>
<td id="76917505B005F58CB3B8F97859DFF976" box="[939,986,1713,1737]" gridcol="4" gridrow="34" pageId="23" pageNumber="102">9.3</td>
<td id="76917505B005F58CB3E4F9785E29F976" box="[1015,1068,1713,1737]" gridcol="5" gridrow="34" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B3E9F9785E2CF976" box="[1018,1065,1713,1737]" italics="true" pageId="23" pageNumber="102">25.0</emphasis>
</td>
<td id="76917505B005F58CB45DF9785E71F976" box="[1102,1140,1713,1737]" gridcol="6" gridrow="34" pageId="23" pageNumber="102">1.3</td>
<td id="76917505B005F58CB48CF9785EC5F976" box="[1183,1216,1713,1737]" gridcol="7" gridrow="34" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B48CF9785EC5F976" box="[1183,1216,1713,1737]" italics="true" pageId="23" pageNumber="102">2.2</emphasis>
</td>
<td id="76917505B005F58CB4FBF9785F12F976" box="[1256,1303,1713,1737]" gridcol="8" gridrow="34" pageId="23" pageNumber="102">?</td>
<td id="76917505B005F58CB527F9785F6CF976" box="[1332,1385,1713,1737]" gridcol="9" gridrow="34" pageId="23" pageNumber="102">?</td>
</tr>
</table>
</paragraph>
<paragraph id="8BCF1E3BB0050A64B0F9F8D75B23F887" blockId="23.[189,1399,1822,2025]" box="[234,294,1822,1848]" pageId="23" pageNumber="102">
<emphasis id="B904C229B0050A64B0F9F8D75B23F887" bold="true" box="[234,294,1822,1848]" pageId="23" pageNumber="102">Eggs</emphasis>
</paragraph>
<paragraph id="8BCF1E3BB0050A6BB0AEF88859DDFDDE" blockId="23.[189,1399,1822,2025]" lastBlockId="24.[189,1399,265,609]" lastPageId="24" lastPageNumber="103" pageId="23" pageNumber="102">
Laid freely, white, spherical and ornamented (
<figureCitation id="134B02BEB0050A64B2F2F8885976F8E3" box="[737,883,1857,1884]" captionStart="Fig" captionStartId="25.[189,232,1857,1883]" captionTargetBox="[189,1398,859,1819]" captionTargetId="figure-1@25.[189,1398,859,1819]" captionTargetPageId="25" captionText="Fig. 19. Macrobiotus mileri sp. nov., PCM images of the eggs under ×1000 magnification (ISEA PAS). AC. Egg surface. DE. Midsections of egg processes. Empty indented arrowheads indicate pores/ light-refracting dots in the center of the terminal discs, filled flat arrowheads indicate small pores around bases of the egg processes, filled indented arrowheads indicate dark thickenings around bases of the egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893838" httpUri="https://zenodo.org/record/10893838/files/figure.png" pageId="23" pageNumber="102">Figs 19AE</figureCitation>
,
<figureCitation id="134B02BEB0050A64B392F88B59D2F8E3" box="[897,983,1858,1884]" captionStart="Fig" captionStartId="26.[189,232,1889,1915]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-0@26.[189,1398,265,1833]" captionTargetPageId="26" captionText="Fig. 20. Macrobiotus mileri sp. nov., SEM images of eggs (ISEA PAS). AB. Entire egg. CF. Details of egg processes and the surface between them. Empty indented arrowheads indicate pores in center of terminal discs, filled flat arrowheads indicate small pores around bases of egg processes, filled indented arrowheads indicate thickenings around bases of egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893840" httpUri="https://zenodo.org/record/10893840/files/figure.png" pageId="23" pageNumber="102">20AF</figureCitation>
). The surface between processes is of intermediate state between the
<emphasis id="B904C229B0050A64B247F8AC58C8F8C0" box="[596,717,1893,1919]" italics="true" pageId="23" pageNumber="102">maculatus</emphasis>
and the
<taxonomicName id="4C7065B8B0050A64B32AF8AC59A8F8C0" authorityName="Binda &amp; Pilato" authorityYear="1972" box="[825,941,1893,1919]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="23" pageNumber="102" phylum="Tardigrada" rank="species" species="persimilis">
<emphasis id="B904C229B0050A64B32AF8AC59A8F8C0" box="[825,941,1893,1919]" italics="true" pageId="23" pageNumber="102">persimilis</emphasis>
</taxonomicName>
<typeStatus id="54CBA099B0050A64B3A5F8AC59F2F8C0" box="[950,1015,1893,1919]" pageId="23" pageNumber="102">types</typeStatus>
, that is, the surface is solid and wrinkled with very small pores, which are present mainly around the bases of the egg processes, and only some are sparse and irregularly distributed in the egg surface between processes (
<figureCitation id="134B02BEB0050A64B464F8625F0FF87A" box="[1143,1290,1963,1989]" captionStart="Fig" captionStartId="25.[189,232,1857,1883]" captionTargetBox="[189,1398,859,1819]" captionTargetId="figure-1@25.[189,1398,859,1819]" captionTargetPageId="25" captionText="Fig. 19. Macrobiotus mileri sp. nov., PCM images of the eggs under ×1000 magnification (ISEA PAS). AC. Egg surface. DE. Midsections of egg processes. Empty indented arrowheads indicate pores/ light-refracting dots in the center of the terminal discs, filled flat arrowheads indicate small pores around bases of the egg processes, filled indented arrowheads indicate dark thickenings around bases of the egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893838" httpUri="https://zenodo.org/record/10893838/files/figure.png" pageId="23" pageNumber="102">Figs 19AC</figureCitation>
,
<figureCitation id="134B02BEB0050A64B506F8625F6EF87A" box="[1301,1387,1963,1989]" captionStart="Fig" captionStartId="26.[189,232,1889,1915]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-0@26.[189,1398,265,1833]" captionTargetPageId="26" captionText="Fig. 20. Macrobiotus mileri sp. nov., SEM images of eggs (ISEA PAS). AB. Entire egg. CF. Details of egg processes and the surface between them. Empty indented arrowheads indicate pores in center of terminal discs, filled flat arrowheads indicate small pores around bases of egg processes, filled indented arrowheads indicate thickenings around bases of egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893840" httpUri="https://zenodo.org/record/10893840/files/figure.png" pageId="23" pageNumber="102">20CF</figureCitation>
). These pores are visible under PCM, but better visible under SEM (0.10.3) μm in diameter;
<figureCitation id="134B02BEB0050A64B4FBF8075F77F857" box="[1256,1394,1998,2024]" captionStart="Fig" captionStartId="25.[189,232,1857,1883]" captionTargetBox="[189,1398,859,1819]" captionTargetId="figure-1@25.[189,1398,859,1819]" captionTargetPageId="25" captionText="Fig. 19. Macrobiotus mileri sp. nov., PCM images of the eggs under ×1000 magnification (ISEA PAS). AC. Egg surface. DE. Midsections of egg processes. Empty indented arrowheads indicate pores/ light-refracting dots in the center of the terminal discs, filled flat arrowheads indicate small pores around bases of the egg processes, filled indented arrowheads indicate dark thickenings around bases of the egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893838" httpUri="https://zenodo.org/record/10893838/files/figure.png" pageId="23" pageNumber="102">Figs19AC</figureCitation>
,
<figureCitation id="134B02BEB00A0A6BB0AEFEC05B17FE9C" box="[189,274,265,291]" captionStart="Fig" captionStartId="26.[189,232,1889,1915]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-0@26.[189,1398,265,1833]" captionTargetPageId="26" captionText="Fig. 20. Macrobiotus mileri sp. nov., SEM images of eggs (ISEA PAS). AB. Entire egg. CF. Details of egg processes and the surface between them. Empty indented arrowheads indicate pores in center of terminal discs, filled flat arrowheads indicate small pores around bases of egg processes, filled indented arrowheads indicate thickenings around bases of egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893840" httpUri="https://zenodo.org/record/10893840/files/figure.png" pageId="24" pageNumber="103">20CF</figureCitation>
). The processes are not in the shape of inverted goblets with mostly sigmoidal (sometimes concave) conical trunks and weakly defined convex terminal discs with smooth edges (
<figureCitation id="134B02BEB00A0A6BB4CFFEE55F77FEF9" box="[1244,1394,300,326]" captionStart="Fig" captionStartId="25.[189,232,1857,1883]" captionTargetBox="[189,1398,859,1819]" captionTargetId="figure-1@25.[189,1398,859,1819]" captionTargetPageId="25" captionText="Fig. 19. Macrobiotus mileri sp. nov., PCM images of the eggs under ×1000 magnification (ISEA PAS). AC. Egg surface. DE. Midsections of egg processes. Empty indented arrowheads indicate pores/ light-refracting dots in the center of the terminal discs, filled flat arrowheads indicate small pores around bases of the egg processes, filled indented arrowheads indicate dark thickenings around bases of the egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893838" httpUri="https://zenodo.org/record/10893838/files/figure.png" pageId="24" pageNumber="103">Figs 19AC</figureCitation>
,
<figureCitation id="134B02BEB00A0A6BB0AEFE865B17FED6" box="[189,274,335,361]" captionStart="Fig" captionStartId="26.[189,232,1889,1915]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-0@26.[189,1398,265,1833]" captionTargetPageId="26" captionText="Fig. 20. Macrobiotus mileri sp. nov., SEM images of eggs (ISEA PAS). AB. Entire egg. CF. Details of egg processes and the surface between them. Empty indented arrowheads indicate pores in center of terminal discs, filled flat arrowheads indicate small pores around bases of egg processes, filled indented arrowheads indicate thickenings around bases of egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893840" httpUri="https://zenodo.org/record/10893840/files/figure.png" pageId="24" pageNumber="103">20CF</figureCitation>
). Very faint annulations are visible on the process trunk, especially on the distal portion of the process (character visible only under SEM;
<figureCitation id="134B02BEB00A0A6BB2D1FEBB592AFE32" box="[706,815,370,397]" captionStart="Fig" captionStartId="26.[189,232,1889,1915]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-0@26.[189,1398,265,1833]" captionTargetPageId="26" captionText="Fig. 20. Macrobiotus mileri sp. nov., SEM images of eggs (ISEA PAS). AB. Entire egg. CF. Details of egg processes and the surface between them. Empty indented arrowheads indicate pores in center of terminal discs, filled flat arrowheads indicate small pores around bases of egg processes, filled indented arrowheads indicate thickenings around bases of egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893840" httpUri="https://zenodo.org/record/10893840/files/figure.png" pageId="24" pageNumber="103">Fig. 20D</figureCitation>
). A crown of gently marked thickenings is visible around the bases of the processes as darker dots under PCM (
<figureCitation id="134B02BEB00A0A6BB394FE5F5E16FE0F" box="[903,1043,406,432]" captionStart="Fig" captionStartId="25.[189,232,1857,1883]" captionTargetBox="[189,1398,859,1819]" captionTargetId="figure-1@25.[189,1398,859,1819]" captionTargetPageId="25" captionText="Fig. 19. Macrobiotus mileri sp. nov., PCM images of the eggs under ×1000 magnification (ISEA PAS). AC. Egg surface. DE. Midsections of egg processes. Empty indented arrowheads indicate pores/ light-refracting dots in the center of the terminal discs, filled flat arrowheads indicate small pores around bases of the egg processes, filled indented arrowheads indicate dark thickenings around bases of the egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893838" httpUri="https://zenodo.org/record/10893838/files/figure.png" pageId="24" pageNumber="103">Fig. 19AC</figureCitation>
) and as thicker wrinkles at the processes bases under SEM (
<figureCitation id="134B02BEB00A0A6BB200FE7058A4FE6C" box="[531,673,441,467]" captionStart="Fig" captionStartId="26.[189,232,1889,1915]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-0@26.[189,1398,265,1833]" captionTargetPageId="26" captionText="Fig. 20. Macrobiotus mileri sp. nov., SEM images of eggs (ISEA PAS). AB. Entire egg. CF. Details of egg processes and the surface between them. Empty indented arrowheads indicate pores in center of terminal discs, filled flat arrowheads indicate small pores around bases of egg processes, filled indented arrowheads indicate thickenings around bases of egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893840" httpUri="https://zenodo.org/record/10893840/files/figure.png" pageId="24" pageNumber="103">Fig. 20DF</figureCitation>
). In some processes under SEM the terminal discs have pores in the center (
<figureCitation id="134B02BEB00A0A6BB173FE155BE8FE49" box="[352,493,476,502]" captionStart="Fig" captionStartId="26.[189,232,1889,1915]" captionTargetBox="[189,1398,265,1833]" captionTargetId="figure-0@26.[189,1398,265,1833]" captionTargetPageId="26" captionText="Fig. 20. Macrobiotus mileri sp. nov., SEM images of eggs (ISEA PAS). AB. Entire egg. CF. Details of egg processes and the surface between them. Empty indented arrowheads indicate pores in center of terminal discs, filled flat arrowheads indicate small pores around bases of egg processes, filled indented arrowheads indicate thickenings around bases of egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893840" httpUri="https://zenodo.org/record/10893840/files/figure.png" pageId="24" pageNumber="103">Fig. 20DF</figureCitation>
), which under PCM are visible as large light-refracting dot in the disc center (
<figureCitation id="134B02BEB00A0A6BB0D5FE365B50FDA6" box="[198,341,511,537]" captionStart="Fig" captionStartId="25.[189,232,1857,1883]" captionTargetBox="[189,1398,859,1819]" captionTargetId="figure-1@25.[189,1398,859,1819]" captionTargetPageId="25" captionText="Fig. 19. Macrobiotus mileri sp. nov., PCM images of the eggs under ×1000 magnification (ISEA PAS). AC. Egg surface. DE. Midsections of egg processes. Empty indented arrowheads indicate pores/ light-refracting dots in the center of the terminal discs, filled flat arrowheads indicate small pores around bases of the egg processes, filled indented arrowheads indicate dark thickenings around bases of the egg processes. Scale bars in μm." figureDoi="http://doi.org/10.5281/zenodo.10893838" httpUri="https://zenodo.org/record/10893838/files/figure.png" pageId="24" pageNumber="103">Fig. 19AC</figureCitation>
). However, it cannot be excluded that the actual pores in the terminal discs are preparation artefacts, while light refracting dots visible under PCM are caused by thinner chorion layers in this place. The measurements and statistics of eggs are given in
<tableCitation id="C6F22B80B00A0A6BB36AFD8F59D7FDDE" box="[889,978,582,609]" captionStart="Table 5" captionStartId="25.[189,258,265,291]" captionTargetPageId="25" captionText="Table 5. Measurements [in μm] of the eggs of Macrobiotus mileri sp. nov.; eggs mounted in Hoyers medium; process base/height ratio is expressed as percentage; N: number of eggs/structures measured; range: refers to the smallest and the largest structure among all measured specimens; SD: standard deviation." httpUri="http://table.plazi.org/id/DF0F4EB3B00B0A6AB0AEFEC05B37FE33" pageId="24" pageNumber="103" tableUuid="DF0F4EB3B00B0A6AB0AEFEC05B37FE33">Table 5</tableCitation>
.
</paragraph>
<caption id="DF0F4EB3B00A0A6BB0AEF8D75E38F856" ID-DOI="http://doi.org/10.5281/zenodo.10893836" ID-Zenodo-Dep="10893836" httpUri="https://zenodo.org/record/10893836/files/figure.png" pageId="24" pageNumber="103" startId="24.[189,232,1822,1848]" targetBox="[189,1398,738,1781]" targetPageId="24">
<paragraph id="8BCF1E3BB00A0A6BB0AEF8D75E38F856" blockId="24.[189,1398,1822,2025]" pageId="24" pageNumber="103">
<emphasis id="B904C229B00A0A6BB0AEF8D75B1BF887" bold="true" box="[189,286,1822,1848]" pageId="24" pageNumber="103">Fig. 18.</emphasis>
<taxonomicName id="4C7065B8B00A0A6BB13AF8D7580AF887" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[297,527,1822,1848]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="24" pageNumber="103" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B00A0A6BB13AF8D7580AF887" box="[297,527,1822,1848]" italics="true" pageId="24" pageNumber="103">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B00A0A6BB209F8D6587EF886" box="[538,635,1823,1849]" pageId="24" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
, mouth opening and the oral cavity armature seen under SEM (paratypes, ISEA PAS).
<emphasis id="B904C229B00A0A6BB1CEF8885810F8E4" bold="true" box="[477,533,1857,1883]" pageId="24" pageNumber="103">AB</emphasis>
. The mouth opening of a single paratype visible from lateral and frontal view respectively.
<emphasis id="B904C229B00A0A6BB188F8AD5BD6F8C0" bold="true" box="[411,467,1892,1919]" pageId="24" pageNumber="103">CD</emphasis>
. The oral cavity armature of a single paratype seen under SEM from different angles, dorsal (C) and ventral (D) view, respectively. Empty arrows indicate cuticular pores, filled flat arrowheads indicate the first band of tenth, empty flat arrowheads indicate the second band of teeth, and filled indented arrowheads indicate the third band of teeth. Scale bars in μm.
</paragraph>
</caption>
<caption id="DF0F4EB3B00B0A6AB0AEFEC05B37FE33" ID-Table-UUID="DF0F4EB3B00B0A6AB0AEFEC05B37FE33" httpUri="http://table.plazi.org/id/DF0F4EB3B00B0A6AB0AEFEC05B37FE33" pageId="25" pageNumber="104" startId="25.[189,258,265,291]" targetBox="[197,1366,427,754]" targetIsTable="true" targetPageId="25">
<paragraph id="8BCF1E3BB00B0A6AB0AEFEC05B37FE33" blockId="25.[189,1398,265,396]" pageId="25" pageNumber="104">
<emphasis id="B904C229B00B0A6AB0AEFEC05B24FE9C" bold="true" box="[189,289,265,291]" pageId="25" pageNumber="104">Table 5.</emphasis>
Measurements [in μm] of the eggs of
<taxonomicName id="4C7065B8B00B0A6AB2E3FEC059D1FE9C" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[752,980,265,291]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="104" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B00B0A6AB2E3FEC059D1FE9C" box="[752,980,265,291]" italics="true" pageId="25" pageNumber="104">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B00B0A6AB3CEFEC35E39FE9B" box="[989,1084,266,292]" pageId="25" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
; eggs mounted in Hoyers medium; process base/height ratio is expressed as percentage; N: number of eggs/structures measured; range: refers to the smallest and the largest structure among all measured specimens; SD: standard deviation.
</paragraph>
</caption>
<paragraph id="8BCF1E3BB00B0A6AB0D6FE625F4BFD4D" pageId="25" pageNumber="104">
<table id="F970EC9BB00BF58CB0D6FE625F53FD4D" box="[197,1366,427,754]" gridcols="5" gridrows="9" pageId="25" pageNumber="104">
<tr id="35401C79B00BF58CB0D6FE625F53FE7B" box="[197,1366,427,452]" gridrow="0" pageId="25" pageNumber="104">
<th id="76917505B00BF58CB0D6FE6258BEFE7B" box="[197,699,427,452]" gridcol="0" gridrow="0" pageId="25" pageNumber="104">
<emphasis id="B904C229B00B0A6AB0D6FE625B3FFE7C" bold="true" box="[197,314,427,451]" pageId="25" pageNumber="104">Character</emphasis>
</th>
<th id="76917505B00BF58CB36DFE62599CFE7B" box="[894,921,427,452]" gridcol="1" gridrow="0" pageId="25" pageNumber="104">
<emphasis id="B904C229B00B0A6AB392FE655991FE7B" bold="true" box="[897,916,428,452]" pageId="25" pageNumber="104">N</emphasis>
</th>
<th id="76917505B00BF58CB3FBFE625E6EFE7B" box="[1000,1131,427,452]" gridcol="2" gridrow="0" pageId="25" pageNumber="104">
<emphasis id="B904C229B00B0A6AB415FE655E4AFE7B" bold="true" box="[1030,1103,428,452]" pageId="25" pageNumber="104">Range</emphasis>
</th>
<th id="76917505B00BF58CB4BFFE625EE8FE7B" box="[1196,1261,427,452]" gridcol="3" gridrow="0" pageId="25" pageNumber="104">
<emphasis id="B904C229B00B0A6AB4BFFE655EE8FE7B" bold="true" box="[1196,1261,428,452]" pageId="25" pageNumber="104">Mean</emphasis>
</th>
<th id="76917505B00BF58CB536FE625F53FE7B" box="[1317,1366,427,452]" gridcol="4" gridrow="0" pageId="25" pageNumber="104">
<emphasis id="B904C229B00B0A6AB53EFE625F4AFE7C" bold="true" box="[1325,1359,427,451]" pageId="25" pageNumber="104">SD</emphasis>
</th>
</tr>
<tr id="35401C79B00BF58CB0D6FE185F53FE55" box="[197,1366,465,490]" gridrow="1" pageId="25" pageNumber="104">
<th id="76917505B00BF58CB0D6FE1858BEFE55" box="[197,699,465,490]" gridcol="0" gridrow="1" pageId="25" pageNumber="104">Egg bare diameter</th>
<td id="76917505B00BF58CB36DFE18599CFE55" box="[894,921,465,490]" gridcol="1" gridrow="1" pageId="25" pageNumber="104">7</td>
<td id="76917505B00BF58CB3FBFE185E6EFE55" box="[1000,1131,465,490]" gridcol="2" gridrow="1" pageId="25" pageNumber="104">76.5 92.6</td>
<td id="76917505B00BF58CB4BFFE185EE8FE55" box="[1196,1261,465,490]" gridcol="3" gridrow="1" pageId="25" pageNumber="104">82.5</td>
<td id="76917505B00BF58CB536FE185F53FE55" box="[1317,1366,465,490]" gridcol="4" gridrow="1" pageId="25" pageNumber="104">6.2</td>
</tr>
<tr id="35401C79B00BF58CB0D6FE3E5F53FDAF" box="[197,1366,503,528]" gridrow="2" pageId="25" pageNumber="104">
<th id="76917505B00BF58CB0D6FE3E58BEFDAF" box="[197,699,503,528]" gridcol="0" gridrow="2" pageId="25" pageNumber="104">Egg full diameter</th>
<td id="76917505B00BF58CB36DFE3E599CFDAF" box="[894,921,503,528]" gridcol="1" gridrow="2" pageId="25" pageNumber="104">7</td>
<td id="76917505B00BF58CB3FBFE3E5E6EFDAF" box="[1000,1131,503,528]" gridcol="2" gridrow="2" pageId="25" pageNumber="104">93.0 107.6</td>
<td id="76917505B00BF58CB4BFFE3E5EE8FDAF" box="[1196,1261,503,528]" gridcol="3" gridrow="2" pageId="25" pageNumber="104">98.2</td>
<td id="76917505B00BF58CB536FE3E5F53FDAF" box="[1317,1366,503,528]" gridcol="4" gridrow="2" pageId="25" pageNumber="104">6.3</td>
</tr>
<tr id="35401C79B00BF58CB0D6FDD45F53FD8A" box="[197,1366,541,565]" gridrow="3" pageId="25" pageNumber="104">
<th id="76917505B00BF58CB0D6FDD458BEFD8A" box="[197,699,541,565]" gridcol="0" gridrow="3" pageId="25" pageNumber="104">Process height</th>
<td id="76917505B00BF58CB36DFDD4599CFD8A" box="[894,921,541,565]" gridcol="1" gridrow="3" pageId="25" pageNumber="104">21</td>
<td id="76917505B00BF58CB3FBFDD45E6EFD8A" box="[1000,1131,541,565]" gridcol="2" gridrow="3" pageId="25" pageNumber="104">6.2 8.7</td>
<td id="76917505B00BF58CB4BFFDD45EE8FD8A" box="[1196,1261,541,565]" gridcol="3" gridrow="3" pageId="25" pageNumber="104">7.6</td>
<td id="76917505B00BF58CB536FDD45F53FD8A" box="[1317,1366,541,565]" gridcol="4" gridrow="3" pageId="25" pageNumber="104">0.8</td>
</tr>
<tr id="35401C79B00BF58CB0D6FD8A5F53FDE4" box="[197,1366,579,603]" gridrow="4" pageId="25" pageNumber="104">
<th id="76917505B00BF58CB0D6FD8A58BEFDE4" box="[197,699,579,603]" gridcol="0" gridrow="4" pageId="25" pageNumber="104">Process base width</th>
<td id="76917505B00BF58CB36DFD8A599CFDE4" box="[894,921,579,603]" gridcol="1" gridrow="4" pageId="25" pageNumber="104">21</td>
<td id="76917505B00BF58CB3FBFD8A5E6EFDE4" box="[1000,1131,579,603]" gridcol="2" gridrow="4" pageId="25" pageNumber="104">7.3 10.3</td>
<td id="76917505B00BF58CB4BFFD8A5EE8FDE4" box="[1196,1261,579,603]" gridcol="3" gridrow="4" pageId="25" pageNumber="104">8.5</td>
<td id="76917505B00BF58CB536FD8A5F53FDE4" box="[1317,1366,579,603]" gridcol="4" gridrow="4" pageId="25" pageNumber="104">0.6</td>
</tr>
<tr id="35401C79B00BF58CB0D6FDA05F53FD3E" box="[197,1366,617,641]" gridrow="5" pageId="25" pageNumber="104">
<th id="76917505B00BF58CB0D6FDA058BEFD3E" box="[197,699,617,641]" gridcol="0" gridrow="5" pageId="25" pageNumber="104">Process base/height ratio</th>
<td id="76917505B00BF58CB36DFDA0599CFD3E" box="[894,921,617,641]" gridcol="1" gridrow="5" pageId="25" pageNumber="104">21</td>
<td id="76917505B00BF58CB3FBFDA05E6EFD3E" box="[1000,1131,617,641]" gridcol="2" gridrow="5" pageId="25" pageNumber="104">93% 140%</td>
<td id="76917505B00BF58CB4BFFDA05EE8FD3E" box="[1196,1261,617,641]" gridcol="3" gridrow="5" pageId="25" pageNumber="104">113%</td>
<td id="76917505B00BF58CB536FDA05F53FD3E" box="[1317,1366,617,641]" gridcol="4" gridrow="5" pageId="25" pageNumber="104">14%</td>
</tr>
<tr id="35401C79B00BF58CB0D6FD475F53FD18" box="[197,1366,654,679]" gridrow="6" pageId="25" pageNumber="104">
<th id="76917505B00BF58CB0D6FD4758BEFD18" box="[197,699,654,679]" gridcol="0" gridrow="6" pageId="25" pageNumber="104">Terminal disc width</th>
<td id="76917505B00BF58CB36DFD47599CFD18" box="[894,921,654,679]" gridcol="1" gridrow="6" pageId="25" pageNumber="104">21</td>
<td id="76917505B00BF58CB3FBFD475E6EFD18" box="[1000,1131,654,679]" gridcol="2" gridrow="6" pageId="25" pageNumber="104">2.2 4.2</td>
<td id="76917505B00BF58CB4BFFD475EE8FD18" box="[1196,1261,654,679]" gridcol="3" gridrow="6" pageId="25" pageNumber="104">3.2</td>
<td id="76917505B00BF58CB536FD475F53FD18" box="[1317,1366,654,679]" gridcol="4" gridrow="6" pageId="25" pageNumber="104">0.4</td>
</tr>
<tr id="35401C79B00BF58CB0D6FD7D5F53FD73" box="[197,1366,692,716]" gridrow="7" pageId="25" pageNumber="104">
<th id="76917505B00BF58CB0D6FD7D58BEFD73" box="[197,699,692,716]" gridcol="0" gridrow="7" pageId="25" pageNumber="104">Inter-process distance</th>
<td id="76917505B00BF58CB36DFD7D599CFD73" box="[894,921,692,716]" gridcol="1" gridrow="7" pageId="25" pageNumber="104">21</td>
<td id="76917505B00BF58CB3FBFD7D5E6EFD73" box="[1000,1131,692,716]" gridcol="2" gridrow="7" pageId="25" pageNumber="104">1.2 3.3</td>
<td id="76917505B00BF58CB4BFFD7D5EE8FD73" box="[1196,1261,692,716]" gridcol="3" gridrow="7" pageId="25" pageNumber="104">2.4</td>
<td id="76917505B00BF58CB536FD7D5F53FD73" box="[1317,1366,692,716]" gridcol="4" gridrow="7" pageId="25" pageNumber="104">0.5</td>
</tr>
<tr id="35401C79B00BF58CB0D6FD135F53FD4D" box="[197,1366,730,754]" gridrow="8" pageId="25" pageNumber="104">
<th id="76917505B00BF58CB0D6FD1358BEFD4D" box="[197,699,730,754]" gridcol="0" gridrow="8" pageId="25" pageNumber="104">Number of processes on the egg circumference</th>
<td id="76917505B00BF58CB36DFD13599CFD4D" box="[894,921,730,754]" gridcol="1" gridrow="8" pageId="25" pageNumber="104">7</td>
<td id="76917505B00BF58CB3FBFD135E6EFD4D" box="[1000,1131,730,754]" gridcol="2" gridrow="8" pageId="25" pageNumber="104">23 25</td>
<td id="76917505B00BF58CB4BFFD135EE8FD4D" box="[1196,1261,730,754]" gridcol="3" gridrow="8" pageId="25" pageNumber="104">24.0</td>
<td id="76917505B00BF58CB536FD135F53FD4D" box="[1317,1366,730,754]" gridcol="4" gridrow="8" pageId="25" pageNumber="104">0.8</td>
</tr>
</table>
</paragraph>
<caption id="DF0F4EB3B00B0A6AB0AEF8885861F856" ID-DOI="http://doi.org/10.5281/zenodo.10893838" ID-Zenodo-Dep="10893838" httpUri="https://zenodo.org/record/10893838/files/figure.png" pageId="25" pageNumber="104" startId="25.[189,232,1857,1883]" targetBox="[189,1398,859,1819]" targetPageId="25">
<paragraph id="8BCF1E3BB00B0A6AB0AEF8885861F856" blockId="25.[189,1399,1857,2025]" pageId="25" pageNumber="104">
<emphasis id="B904C229B00B0A6AB0AEF8885B25F8E4" bold="true" box="[189,288,1857,1883]" pageId="25" pageNumber="104">Fig. 19.</emphasis>
<taxonomicName id="4C7065B8B00B0A6AB13FF8885811F8E4" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[300,532,1857,1883]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="25" pageNumber="104" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B00B0A6AB13FF8885811F8E4" box="[300,532,1857,1883]" italics="true" pageId="25" pageNumber="104">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B00B0A6AB232F88B5886F8E3" box="[545,643,1858,1884]" pageId="25" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
, PCM images of the eggs under ×1000 magnification (ISEA PAS).
<emphasis id="B904C229B00B0A6AB11EF8AD5B43F8C1" bold="true" box="[269,326,1892,1918]" pageId="25" pageNumber="104">AC</emphasis>
. Egg surface.
<emphasis id="B904C229B00B0A6AB1E8F8AC5836F8C0" bold="true" box="[507,563,1893,1919]" pageId="25" pageNumber="104">DE</emphasis>
. Midsections of egg processes. Empty indented arrowheads indicate pores/ light-refracting dots in the center of the terminal discs, filled flat arrowheads indicate small pores around bases of the egg processes, filled indented arrowheads indicate dark thickenings around bases of the egg processes. Scale bars in μm.
</paragraph>
</caption>
<caption id="DF0F4EB3B0080A69B0AEF8A85E7FF85A" ID-DOI="http://doi.org/10.5281/zenodo.10893840" ID-Zenodo-Dep="10893840" httpUri="https://zenodo.org/record/10893840/files/figure.png" pageId="26" pageNumber="105" startId="26.[189,232,1889,1915]" targetBox="[189,1398,265,1833]" targetPageId="26">
<paragraph id="8BCF1E3BB0080A69B0AEF8A85E7FF85A" blockId="26.[189,1398,1889,2021]" pageId="26" pageNumber="105">
<emphasis id="B904C229B0080A69B0AEF8A85B1EF8C4" bold="true" box="[189,283,1889,1915]" pageId="26" pageNumber="105">Fig. 20.</emphasis>
<taxonomicName id="4C7065B8B0080A69B131F8A85800F8C4" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[290,517,1889,1915]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="26" pageNumber="105" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0080A69B131F8A85800F8C4" box="[290,517,1889,1915]" italics="true" pageId="26" pageNumber="105">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0080A69B21EF8AB586FF8C3" box="[525,618,1890,1916]" pageId="26" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
, SEM images of eggs (ISEA PAS).
<emphasis id="B904C229B0080A69B41CF8A85E42F8C4" bold="true" box="[1039,1095,1889,1916]" pageId="26" pageNumber="105">AB</emphasis>
. Entire egg.
<emphasis id="B904C229B0080A69B4CCF8A85F11F8C4" bold="true" box="[1247,1300,1889,1916]" pageId="26" pageNumber="105">CF</emphasis>
. Details of egg processes and the surface between them. Empty indented arrowheads indicate pores in center of terminal discs, filled flat arrowheads indicate small pores around bases of egg processes, filled indented arrowheads indicate thickenings around bases of egg processes. Scale bars in μm.
</paragraph>
</caption>
<paragraph id="8BCF1E3BB0090A68B0AEFEC15B6CFE9D" blockId="27.[189,361,264,290]" box="[189,361,264,290]" pageId="27" pageNumber="106">
<heading id="D087A957B0090A68B0AEFEC15B6CFE9D" bold="true" box="[189,361,264,290]" fontSize="11" level="3" pageId="27" pageNumber="106" reason="3">
<emphasis id="B904C229B0090A68B0AEFEC15B6CFE9D" bold="true" box="[189,361,264,290]" pageId="27" pageNumber="106">Reproduction</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1E3BB0090A68B0AEFEFE5E44FE26" blockId="27.[189,1398,311,409]" pageId="27" pageNumber="106">
The type population of
<taxonomicName id="4C7065B8B0090A68B1C8FEF1584EFEEE" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[475,587,311,337]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0090A68B1C8FEF1584EFEEE" box="[475,587,311,337]" italics="true" pageId="27" pageNumber="106">M. mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0090A68B246FEF158B7FEED" box="[597,690,312,338]" pageId="27" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
is dioecious. Both males with testes filled with sperm and females with ovaries containing oocytes were observed in specimens freshly mounted in Hoyers medium in both specimens ascribed to each of the two morphological forms.
</paragraph>
</subSubSection>
<subSubSection id="C36A4DB0B0090A68B0AEFE095F27FC67" pageId="27" pageNumber="106" type="diagnosis">
<paragraph id="8BCF1E3BB0090A68B0AEFE095BC3FE65" blockId="27.[189,454,448,474]" box="[189,454,448,474]" pageId="27" pageNumber="106">
<heading id="D087A957B0090A68B0AEFE095BC3FE65" bold="true" box="[189,454,448,474]" fontSize="11" level="3" pageId="27" pageNumber="106" reason="3">
<emphasis id="B904C229B0090A68B0AEFE095BC3FE65" bold="true" box="[189,454,448,474]" pageId="27" pageNumber="106">Differential diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1E3BB0090A68B0AEFE395F27FC67" blockId="27.[189,1399,495,985]" pageId="27" pageNumber="106">
By having (i) three bands of teeth in the oral cavity armature that are well visible under light microscope, (ii) the entire body cuticle covered by granulation (sometimes visible only under SEM), the new species is the most similar to five other taxa of
<taxonomicName id="4C7065B8B0090A68B2BBFDFE5938FDEE" authorityName="Schultze" authorityYear="1834" box="[680,829,567,593]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="genus">
<emphasis id="B904C229B0090A68B2BBFDFE5938FDEE" box="[680,829,567,593]" italics="true" pageId="27" pageNumber="106">Macrobiotus</emphasis>
</taxonomicName>
, namely
<taxonomicName id="4C7065B8B0090A68B3A3FDFE5AFDFDCB" authority="Pilato &amp; Binda, 1983" authorityName="Pilato &amp; Binda" authorityYear="1983" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="joannae">
<emphasis id="B904C229B0090A68B3A3FDFE5EABFDEF" box="[944,1198,567,593]" italics="true" pageId="27" pageNumber="106">Macrobiotus joannae</emphasis>
<bibRefCitation id="EFE163CAB0090A68B4ABFDFE5AFDFDCB" author="Pilato G. &amp; Binda M. G." pageId="27" pageNumber="106" pagination="262 - 272" refId="ref16025" refString="Pilato G. &amp; Binda M. G. 1983. Descrizione du una nuova specie di Eutardigrado d'Australia Macrobiotus joannae n. sp. Animalia 10: 262 - 272." type="journal article" year="1983">Pilato &amp; Binda, 1983</bibRefCitation>
</taxonomicName>
reported from its
<typeStatus id="54CBA099B0090A68B1DFFD925BFAFDCA" box="[460,511,603,629]" pageId="27" pageNumber="106">type</typeStatus>
locality in
<collectingCountry id="F3675EABB0090A68B290FD9358EAFDCB" box="[643,751,602,628]" name="Australia" pageId="27" pageNumber="106">Australia</collectingCountry>
(
<bibRefCitation id="EFE163CAB0090A68B2EDFD9359F3FDCB" author="Pilato G. &amp; Binda M. G." box="[766,1014,602,628]" pageId="27" pageNumber="106" pagination="262 - 272" refId="ref16025" refString="Pilato G. &amp; Binda M. G. 1983. Descrizione du una nuova specie di Eutardigrado d'Australia Macrobiotus joannae n. sp. Animalia 10: 262 - 272." type="journal article" year="1983">Pilato &amp; Binda 1983</bibRefCitation>
), and several uncertain localities in central, eastern, and south-eastern
<collectingCountry id="F3675EABB0090A68B295FDB758D1FD27" box="[646,724,638,664]" name="Russia" pageId="27" pageNumber="106">Russia</collectingCountry>
(
<bibRefCitation id="EFE163CAB0090A68B2FBFDB75988FD27" author="Biserov V. I." box="[744,909,638,664]" pageId="27" pageNumber="106" pagination="38 - 50" refId="ref13325" refString="Biserov V. I. 1990. On the revision of the genus Macrobiotus. The subgenus Macrobiotus s. st. is a new systematic status of the hufelandi group (Tardigrada, Macrobiotidae). Communication 2. Zoologicheskij Zhurnal 69 (12): 38 - 50." type="journal article" year="1990">Biserov 1990</bibRefCitation>
) and from
<collectingCountry id="F3675EABB0090A68B40FFDB75E57FD27" box="[1052,1106,638,664]" name="Italy" pageId="27" pageNumber="106">Italy</collectingCountry>
(
<bibRefCitation id="EFE163CAB0090A68B476FDB75F6FFD27" author="Bertolani R. &amp; Guidetti R. &amp; Marchioro T. &amp; Altiero T. &amp; Rebecchi L. &amp; Cesari M." box="[1125,1386,638,664]" pageId="27" pageNumber="106" pagination="110 - 126" refId="ref13175" refString="Bertolani R., Guidetti R., Marchioro T., Altiero T., Rebecchi L. &amp; Cesari M. 2014. Phylogeny of Eutardigrada: New molecular data and their morphological support lead to the identification of new evolutionary lineages. Molecular Phylogenetics and Evolution 76: 110 - 126. https: // doi. org / 10.1016 / j. ympev. 2014.03.006" type="journal article" year="2014">
Bertolani
<emphasis id="B904C229B0090A68B4F3FDB65F1AFD27" box="[1248,1311,638,664]" italics="true" pageId="27" pageNumber="106">et al.</emphasis>
2014
</bibRefCitation>
),
<taxonomicName id="4C7065B8B0090A68B0AEFD6858AAFD03" authority="Nowak &amp; Stec, 2018" authorityName="Nowak &amp; Stec" authorityYear="2018" box="[189,687,673,700]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="hannae">
<emphasis id="B904C229B0090A68B0AEFD685BAAFD04" box="[189,431,673,699]" italics="true" pageId="27" pageNumber="106">Macrobiotus hannae</emphasis>
<bibRefCitation id="EFE163CAB0090A68B1ABFD6858AAFD03" author="Nowak B. &amp; Stec D." box="[440,687,673,700]" pageId="27" pageNumber="106" pagination="269 - 286" refId="ref15949" refString="Nowak B. &amp; Stec D. 2018. An integrative description of Macrobiotus hannae sp. nov. (Tardigrada: Eutardigrada: Macrobiotidae: hufelandi group) from Poland. Turkish Journal of Zoology 42: 269 - 286. https: // doi. org / 10.3906 / zoo- 1712 - 31" type="journal article" year="2018">Nowak &amp; Stec, 2018</bibRefCitation>
</taxonomicName>
known only from its
<typeStatus id="54CBA099B0090A68B3BDFD6B59E4FD03" box="[942,993,674,700]" pageId="27" pageNumber="106">type</typeStatus>
locality in
<collectingCountry id="F3675EABB0090A68B475FD685EB2FD04" box="[1126,1207,673,699]" name="Poland" pageId="27" pageNumber="106">Poland</collectingCountry>
(
<bibRefCitation id="EFE163CAB0090A68B4D4FD685AF9FD60" author="Nowak B. &amp; Stec D." pageId="27" pageNumber="106" pagination="269 - 286" refId="ref15949" refString="Nowak B. &amp; Stec D. 2018. An integrative description of Macrobiotus hannae sp. nov. (Tardigrada: Eutardigrada: Macrobiotidae: hufelandi group) from Poland. Turkish Journal of Zoology 42: 269 - 286. https: // doi. org / 10.3906 / zoo- 1712 - 31" type="journal article" year="2018">Nowak &amp; Stec 2018</bibRefCitation>
),
<taxonomicName id="4C7065B8B0090A68B103FD0C5986FD60" authority="Pilato, Binda &amp; Azzaro, 1990" authorityName="Pilato, Binda &amp; Azzaro" authorityYear="1990" box="[272,899,709,735]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="punctillus">
<emphasis id="B904C229B0090A68B103FD0C5825FD60" box="[272,544,709,735]" italics="true" pageId="27" pageNumber="106">Macrobiotus punctillus</emphasis>
<bibRefCitation id="EFE163CAB0090A68B234FD0C5986FD60" author="Pilato G. &amp; Binda M. G. &amp; Azzaro M." box="[551,899,709,735]" pageId="27" pageNumber="106" pagination="123 - 129" refId="ref16106" refString="Pilato G., Binda M. G. &amp; Azzaro M. 1990. Tardigradi di Terra del Fuoco e Magallanes. III. Macrobiotus punctillus, nuova specie di Macrobiotidae del gruppo hufelandi. Animalia 17: 123 - 129." type="journal article" year="1990">Pilato, Binda &amp; Azzaro, 1990</bibRefCitation>
</taxonomicName>
known only from its
<typeStatus id="54CBA099B0090A68B493FD0F5EB6FD5F" box="[1152,1203,710,736]" pageId="27" pageNumber="106">type</typeStatus>
locality in
<collectingCountry id="F3675EABB0090A68B524FD0C5F72FD60" box="[1335,1399,709,735]" name="Chile" pageId="27" pageNumber="106">Chile</collectingCountry>
(
<bibRefCitation id="EFE163CAB0090A68B0D6FD205B90FCBC" author="Pilato G. &amp; Binda M. G. &amp; Azzaro M." box="[197,405,745,771]" pageId="27" pageNumber="106" pagination="123 - 129" refId="ref16106" refString="Pilato G., Binda M. G. &amp; Azzaro M. 1990. Tardigradi di Terra del Fuoco e Magallanes. III. Macrobiotus punctillus, nuova specie di Macrobiotidae del gruppo hufelandi. Animalia 17: 123 - 129." type="journal article" year="1990">
Pilato
<emphasis id="B904C229B0090A68B100FD205B4BFCBC" box="[275,334,745,771]" italics="true" pageId="27" pageNumber="106">et al.</emphasis>
1990
</bibRefCitation>
),
<taxonomicName id="4C7065B8B0090A68B1B9FD205932FCBC" authority="Stec, 2022" authorityName="Stec" authorityYear="2022" box="[426,823,745,771]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="rebecchii">
<emphasis id="B904C229B0090A68B1B9FD2058B7FCBC" box="[426,690,745,771]" italics="true" pageId="27" pageNumber="106">Macrobiotus rebecchii</emphasis>
Stec, 2022
</taxonomicName>
known only from its
<typeStatus id="54CBA099B0090A68B424FD205E6FFCBC" box="[1079,1130,745,771]" pageId="27" pageNumber="106">type</typeStatus>
locality in
<collectingCountry id="F3675EABB0090A68B4E3FD205F72FCBC" box="[1264,1399,745,771]" name="Kyrgyzstan" pageId="27" pageNumber="106">Kyrgyzstan</collectingCountry>
(
<bibRefCitation id="EFE163CAB0090A68B0D6FCC55B4EFC99" author="Stec D." box="[197,331,780,806]" pageId="27" pageNumber="106" pagination="2906" refId="ref16644" refString="Stec D. 2022 b. Macrobiotus rebecchii sp. nov.: a new limno-terrestrial and hermaphroditic tardigrade from Kyrgyzstan. Animals 12 (21): 2906. https: // doi. org / 10.3390 / ani 12212906" type="journal article" year="2022">Stec 2022b</bibRefCitation>
) and
<taxonomicName id="4C7065B8B0090A68B19EFCC45830FC99" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[397,565,781,806]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="ovovittatus" status="sp. nov.">
<emphasis id="B904C229B0090A68B19EFCC45830FC99" box="[397,565,781,806]" italics="true" pageId="27" pageNumber="106">M. ovovittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0090A68B22FFCC45892FC98" box="[572,663,781,807]" pageId="27" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
described above. However, it can be easily distinguished from all of them by having different pores arrangements on the body cuticles (two forms: porata with pores arranged in five distinct patches; and aporata with singular, small, almost undetectable pores vs typical, more or less evenly distributed cuticular pores in the other species) and a different morphology of the terminal discs (weakly defined convex terminal discs with smooth edges in the new species vs cog-shaped terminal discs, with a concave central area and 1018 distinct teeth in the other species).
</paragraph>
</subSubSection>
<subSubSection id="C36A4DB0B0090A6EB0AEFBC85F66FE47" lastPageId="29" lastPageNumber="108" pageId="27" pageNumber="106" type="discussion">
<paragraph id="8BCF1E3BB0090A68B0AEFBC858A2FBA1" blockId="27.[189,679,1025,1054]" box="[189,679,1025,1054]" pageId="27" pageNumber="106">
<emphasis id="B904C229B0090A68B0AEFBC858A2FBA1" bold="true" box="[189,679,1025,1054]" italics="true" pageId="27" pageNumber="106">Phylogenetic and delimitation results</emphasis>
</paragraph>
<paragraph id="8BCF1E3BB0090A68B0AEFBFF582DF94A" blockId="27.[189,1399,1078,1781]" pageId="27" pageNumber="106">
Both phylogenetic analyses resulted in trees of similar topology, and most of the nodes well and moderately supported, in which three distinct monophyletic
<taxonomicName id="4C7065B8B0090A68B390FB935E1DFBCB" authorityName="Schultze" authorityYear="1834" box="[899,1048,1114,1140]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="genus">
<emphasis id="B904C229B0090A68B390FB935E1DFBCB" box="[899,1048,1114,1140]" italics="true" pageId="27" pageNumber="106">Macrobiotus</emphasis>
</taxonomicName>
lineages (
<taxonomicName id="4C7065B8B0090A68B482FB935F23FBCB" authorityName="Schultze" authorityYear="1834" box="[1169,1318,1114,1140]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="genus">
<emphasis id="B904C229B0090A68B482FB935F23FBCB" box="[1169,1318,1114,1140]" italics="true" pageId="27" pageNumber="106">Macrobiotus</emphasis>
</taxonomicName>
clades A, B, and C) were confidently recovered (
<figureCitation id="134B02BEB0090A68B2B0FBB458FCFB28" box="[675,761,1149,1175]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[190,1399,265,1845]" captionTargetId="graphics-54@28.[190,1116,271,1836]" captionTargetPageId="28" captionText="Fig. 21. Bayesian phylogenetic reconstruction of Superclade I of the family Macrobiotidae. Values at nodes are BI posterior probability supports, nodes with support &lt;0.70 were collapsed whereas nodes with maximum support values (1.00) are not shown. Newly sequenced taxa are bolded and marked with red font. The scale bar represents substitutions per position." figureDoi="http://doi.org/10.5281/zenodo.10893842" httpUri="https://zenodo.org/record/10893842/files/figure.png" pageId="27" pageNumber="106">Fig. 21</figureCitation>
, Supp. file 3). The analyses confirmed also monophyly for the
<taxonomicName id="4C7065B8B0090A68B106FB6B5BE8FB04" authorityName="Weglarska" authorityYear="1965" box="[277,493,1185,1211]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="ariekammensis">
<emphasis id="B904C229B0090A68B106FB6B5BE8FB04" box="[277,493,1185,1211]" italics="true" pageId="27" pageNumber="106">M. ariekammensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7065B8B0090A68B1EEFB6B587BFB04" authorityName="Maucci" authorityYear="1954" box="[509,638,1185,1211]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="pallarii">
<emphasis id="B904C229B0090A68B1EEFB6B587BFB04" box="[509,638,1185,1211]" italics="true" pageId="27" pageNumber="106">M. pallarii</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7065B8B0090A68B2D0FB6B59A8FB04" box="[707,941,1185,1211]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="pseudohufelandi">
<emphasis id="B904C229B0090A68B2D0FB6B59A8FB04" box="[707,941,1185,1211]" italics="true" pageId="27" pageNumber="106">M. pseudohufelandi</emphasis>
</taxonomicName>
complexes (
<figureCitation id="134B02BEB0090A68B457FB685E98FB04" box="[1092,1181,1185,1211]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[190,1399,265,1845]" captionTargetId="graphics-54@28.[190,1116,271,1836]" captionTargetPageId="28" captionText="Fig. 21. Bayesian phylogenetic reconstruction of Superclade I of the family Macrobiotidae. Values at nodes are BI posterior probability supports, nodes with support &lt;0.70 were collapsed whereas nodes with maximum support values (1.00) are not shown. Newly sequenced taxa are bolded and marked with red font. The scale bar represents substitutions per position." figureDoi="http://doi.org/10.5281/zenodo.10893842" httpUri="https://zenodo.org/record/10893842/files/figure.png" pageId="27" pageNumber="106">Fig. 21</figureCitation>
). At first, it seems that the
<emphasis id="B904C229B0090A68B10CFB0D58AEFB61" box="[287,683,1220,1247]" italics="true" pageId="27" pageNumber="106">
<taxonomicName id="4C7065B8B0090A68B10CFB0D582BFB61" box="[287,558,1220,1246]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="polonicus">Macrobiotus polonicus</taxonomicName>
-
<taxonomicName id="4C7065B8B0090A68B224FB0D58AEFB61" authorityName="Binda &amp; Pilato" authorityYear="1972" box="[567,683,1220,1246]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="persimilis">persimilis</taxonomicName>
</emphasis>
complex as defined by
<bibRefCitation id="EFE163CAB0090A68B3D4FB0D5ECAFB60" author="Bertolani R. &amp; Cesari M. &amp; Giovannini I. &amp; Rebecchi L. &amp; Guidetti R. &amp; Kaczmarek L. &amp; Pilato G." box="[967,1231,1220,1247]" pageId="27" pageNumber="106" pagination="329 - 368" refId="ref13244" refString="Bertolani R., Cesari M., Giovannini I., Rebecchi L., Guidetti R., Kaczmarek L. &amp; Pilato G. 2023. The Macrobiotus persimilis-polonicus complex (Eutardigrada, Macrobiotidae), another example of problematic species identification, with the description of four new species. Organisms Diversity &amp; Evolution 23: 329 - 368. https: // doi. org / 10.1007 / s 13127 - 022 - 00599 - z" type="journal article" year="2023">
Bertolani
<emphasis id="B904C229B0090A68B42FFB0C5E7DFB61" box="[1084,1144,1220,1246]" italics="true" pageId="27" pageNumber="106">et al.</emphasis>
(2023)
</bibRefCitation>
has also been recovered to be monophyletic. However, the position of
<taxonomicName id="4C7065B8B0090A68B355FB205E04FABD" box="[838,1025,1256,1282]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="polonicus">
<emphasis id="B904C229B0090A68B355FB205963FABD" box="[838,870,1257,1282]" italics="true" pageId="27" pageNumber="106">M.</emphasis>
cf.
<emphasis id="B904C229B0090A68B39CFB215E04FABD" box="[911,1025,1256,1282]" italics="true" pageId="27" pageNumber="106">polonicus</emphasis>
</taxonomicName>
1 and 2 from
<collectingCountry id="F3675EABB0090A68B48DFB215EFEFABD" box="[1182,1275,1256,1282]" name="Sweden" pageId="27" pageNumber="106">Sweden</collectingCountry>
(
<bibRefCitation id="EFE163CAB0090A68B51BFB215B39FA99" author="Vecchi M. &amp; Stec D." pageId="27" pageNumber="106" pagination="281 - 306" refId="ref18257" refString="Vecchi M. &amp; Stec D. 2021. Integrative descriptions of two new Macrobiotus species (Tardigrada, Eutardigrada, Macrobiotidae) from Mississippi (USA) and Crete (Greece). Zoosystematics and Evolution 97 (1): 281 - 306. https: // doi. org / 10.3897 / zse. 97.65280" type="journal article" year="2021">Vecchi &amp; Stec 2021</bibRefCitation>
), whose morphology also fits this definition, makes this species complex paraphyletic. Also, the
<taxonomicName id="4C7065B8B0090A68B123FAF95BCFFAF6" box="[304,458,1327,1353]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="polonicus">
<emphasis id="B904C229B0090A68B123FAF95BCFFAF6" box="[304,458,1327,1353]" italics="true" pageId="27" pageNumber="106">M. polonicus</emphasis>
</taxonomicName>
species complex as defined by
<bibRefCitation id="EFE163CAB0090A68B353FAE65E1CFAF6" author="Stec D. &amp; Vecchi M. &amp; Calhim S. &amp; Michalczyk L." box="[832,1049,1327,1353]" pageId="27" pageNumber="106" pagination="106987" refId="ref17720" refString="Stec D., Vecchi M., Calhim S. &amp; Michalczyk L. 2021 a. New multilocus phylogeny reorganises the family Macrobiotidae (Eutardigrada) and unveils complex morphological evolution of the Macrobiotus hufelandi group. Molecular Phylogenetics &amp; Evolution 160: 106987. https: // doi. org / 10.1016 / j. ympev. 2020.106987" type="journal article" year="2021">
Stec
<emphasis id="B904C229B0090A68B368FAF959B3FAF6" box="[891,950,1327,1353]" italics="true" pageId="27" pageNumber="106">et al.</emphasis>
(2021a)
</bibRefCitation>
or the
<taxonomicName id="4C7065B8B0090A68B47DFAF95F0FFAF6" authorityName="Binda &amp; Pilato" authorityYear="1972" box="[1134,1290,1327,1353]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="persimilis">
<emphasis id="B904C229B0090A68B47DFAF95F0FFAF6" box="[1134,1290,1327,1353]" italics="true" pageId="27" pageNumber="106">M. persimilis</emphasis>
</taxonomicName>
morpho-group as defined by
<bibRefCitation id="EFE163CAB0090A68B1A5FA9A58C6FAD2" author="Bertolani R. &amp; Cesari M. &amp; Giovannini I. &amp; Rebecchi L. &amp; Guidetti R. &amp; Kaczmarek L. &amp; Pilato G." box="[438,707,1363,1389]" pageId="27" pageNumber="106" pagination="329 - 368" refId="ref13244" refString="Bertolani R., Cesari M., Giovannini I., Rebecchi L., Guidetti R., Kaczmarek L. &amp; Pilato G. 2023. The Macrobiotus persimilis-polonicus complex (Eutardigrada, Macrobiotidae), another example of problematic species identification, with the description of four new species. Organisms Diversity &amp; Evolution 23: 329 - 368. https: // doi. org / 10.1007 / s 13127 - 022 - 00599 - z" type="journal article" year="2023">
Bertolani
<emphasis id="B904C229B0090A68B23EFA9D586EFAD2" box="[557,619,1363,1389]" italics="true" pageId="27" pageNumber="106">et al.</emphasis>
(2023)
</bibRefCitation>
is paraphyletic, since
<taxonomicName id="4C7065B8B0090A68B3C7FA9D5E9DFAD2" box="[980,1176,1363,1389]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="polonicus">
<emphasis id="B904C229B0090A68B3C7FA9D59F1FAD2" box="[980,1012,1364,1389]" italics="true" pageId="27" pageNumber="106">M.</emphasis>
cf.
<emphasis id="B904C229B0090A68B435FA9A5E9DFAD2" box="[1062,1176,1363,1389]" italics="true" pageId="27" pageNumber="106">polonicus</emphasis>
</taxonomicName>
from
<collectingCountry id="F3675EABB0090A68B4F6FA9A5F47FAD2" box="[1253,1346,1363,1389]" name="Sweden" pageId="27" pageNumber="106">Sweden</collectingCountry>
and
<taxonomicName id="4C7065B8B0090A68B0AEFABF58F4FA2E" authority="Vecchi &amp; Stec, 2021" authorityName="Vecchi &amp; Stec" authorityYear="2021" box="[189,753,1398,1425]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="annewintersae">
<emphasis id="B904C229B0090A68B0AEFABF5807FA2F" box="[189,514,1398,1424]" italics="true" pageId="27" pageNumber="106">Macrobiotus annewintersae</emphasis>
<bibRefCitation id="EFE163CAB0090A68B214FABF58F4FA2E" author="Vecchi M. &amp; Stec D." box="[519,753,1398,1425]" pageId="27" pageNumber="106" pagination="281 - 306" refId="ref18257" refString="Vecchi M. &amp; Stec D. 2021. Integrative descriptions of two new Macrobiotus species (Tardigrada, Eutardigrada, Macrobiotidae) from Mississippi (USA) and Crete (Greece). Zoosystematics and Evolution 97 (1): 281 - 306. https: // doi. org / 10.3897 / zse. 97.65280" type="journal article" year="2021">Vecchi &amp; Stec, 2021</bibRefCitation>
</taxonomicName>
cluster together with species of the
<taxonomicName id="4C7065B8B0090A68B49BFABE5F00FA2F" authorityName="Maucci" authorityYear="1954" box="[1160,1285,1398,1424]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="pallarii">
<emphasis id="B904C229B0090A68B49BFABE5F00FA2F" box="[1160,1285,1398,1424]" italics="true" pageId="27" pageNumber="106">M. pallarii</emphasis>
</taxonomicName>
complex.
<taxonomicName id="4C7065B8B0090A68B0AEFA535BA4FA0B" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[189,417,1434,1460]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0090A68B0AEFA535BA4FA0B" box="[189,417,1434,1460]" italics="true" pageId="27" pageNumber="106">Macrobiotus mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0090A68B1BAFA525800FA0A" box="[425,517,1435,1461]" pageId="27" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the
<taxonomicName id="4C7065B8B0090A68B2ADFA535956FA0B" authorityName="Schultze" authorityYear="1834" box="[702,851,1434,1460]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="genus">
<emphasis id="B904C229B0090A68B2ADFA535956FA0B" box="[702,851,1434,1460]" italics="true" pageId="27" pageNumber="106">Macrobiotus</emphasis>
</taxonomicName>
clade B staying in sister relationship with the clade containing
<taxonomicName id="4C7065B8B0090A68B199FA775E4BFA67" authority="Coughlan, Michalczyk &amp; Stec, 2019" authorityName="Coughlan, Michalczyk &amp; Stec" authorityYear="2019" box="[394,1102,1470,1496]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="caelestis">
<emphasis id="B904C229B0090A68B199FA77588BFA67" box="[394,654,1470,1496]" italics="true" pageId="27" pageNumber="106">Macrobiotus caelestis</emphasis>
<bibRefCitation id="EFE163CAB0090A68B289FA775E4BFA67" author="Coughlan K. &amp; Michalczyk L. &amp; Stec D." box="[666,1102,1470,1496]" pageId="27" pageNumber="106" pagination="499 - 513" refId="ref13607" refString="Coughlan K., Michalczyk L. &amp; Stec D. 2019. Macrobiotus caelestis sp. nov., a new tardigrade species (Macrobiotidae: hufelandi group) from the Tien Shan mountains (Kyrgyzstan). Annales Zoologici 69 (3): 499 - 513. https: // doi. org / 10.3161 / 00034541 ANZ 2019.69.3.002" type="journal article" year="2019">Coughlan, Michalczyk &amp; Stec, 2019</bibRefCitation>
</taxonomicName>
and nominal taxa of the
<emphasis id="B904C229B0090A68B0AEFA2B5BD3FA44" box="[189,470,1505,1532]" italics="true" pageId="27" pageNumber="106">
<taxonomicName id="4C7065B8B0090A68B0AEFA2B5B5DFA44" box="[189,344,1505,1531]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="polonicus">M. polonicus</taxonomicName>
-
<taxonomicName id="4C7065B8B0090A68B171FA285BD3FA44" authorityName="Binda &amp; Pilato" authorityYear="1972" box="[354,470,1505,1531]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="persimilis">persimilis</taxonomicName>
</emphasis>
complex (
<figureCitation id="134B02BEB0090A68B244FA2858B4FA44" box="[599,689,1505,1531]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[190,1399,265,1845]" captionTargetId="graphics-54@28.[190,1116,271,1836]" captionTargetPageId="28" captionText="Fig. 21. Bayesian phylogenetic reconstruction of Superclade I of the family Macrobiotidae. Values at nodes are BI posterior probability supports, nodes with support &lt;0.70 were collapsed whereas nodes with maximum support values (1.00) are not shown. Newly sequenced taxa are bolded and marked with red font. The scale bar represents substitutions per position." figureDoi="http://doi.org/10.5281/zenodo.10893842" httpUri="https://zenodo.org/record/10893842/files/figure.png" pageId="27" pageNumber="106">Fig. 21</figureCitation>
). The second new species,
<taxonomicName id="4C7065B8B0090A68B3E9FA2B5EA1FA44" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[1018,1188,1506,1531]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="ovovittatus" status="sp. nov.">
<emphasis id="B904C229B0090A68B3E9FA2B5EA1FA44" box="[1018,1188,1506,1531]" italics="true" pageId="27" pageNumber="106">M. ovovittatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0090A68B4BDFA2B5F08FA43" box="[1198,1293,1506,1532]" pageId="27" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
, belongs to the
<taxonomicName id="4C7065B8B0090A68B114F9CC5B99F9A0" authorityName="Schultze" authorityYear="1834" box="[263,412,1541,1567]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="genus">
<emphasis id="B904C229B0090A68B114F9CC5B99F9A0" box="[263,412,1541,1567]" italics="true" pageId="27" pageNumber="106">Macrobiotus</emphasis>
</taxonomicName>
clade A, as the closest relative of
<taxonomicName id="4C7065B8B0090A68B33FF9CC5B24F9FC" authority="Yuan, Wang, Liu, Liu &amp; Li, 2022" authorityName="Yuan, Wang, Liu, Liu &amp; Li" authorityYear="2022" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="hupingensis">
<emphasis id="B904C229B0090A68B33FF9CC5E51F9A0" box="[812,1108,1541,1567]" italics="true" pageId="27" pageNumber="106">Macrobiotus hupingensis</emphasis>
<bibRefCitation id="EFE163CAB0090A68B448F9CC5B24F9FC" author="Yuan Z. &amp; Wang Y. &amp; Liu Q. &amp; Liu L. &amp; Li X. - C." pageId="27" pageNumber="106" pagination="86" refId="ref18630" refString="Yuan Z., Wang Y., Liu Q., Liu L. &amp; Li X. - C. 2022. Macrobiotus hupingensis, a new tardigrade species in the Macrobiotus pallarii complex from China. Zoological Studies 61: 86. https: // doi. org / 10.6620 / ZS. 2022.61 - 86" type="journal article" year="2022">Yuan, Wang, Liu, Liu &amp; Li, 2022</bibRefCitation>
</taxonomicName>
and together they cluster with
<taxonomicName id="4C7065B8B0090A68B298F9E15811F9D9" authority="Kayastha, Roszkowska, Mioduchowska, Gawlak &amp; Kaczmarek, 2021" authorityName="Kayastha, Roszkowska, Mioduchowska, Gawlak &amp; Kaczmarek" authorityYear="2021" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="birendrai">
<emphasis id="B904C229B0090A68B298F9E15990F9FD" box="[651,917,1576,1602]" italics="true" pageId="27" pageNumber="106">Macrobiotus birendrai</emphasis>
<bibRefCitation id="EFE163CAB0090A68B38FF9E15811F9D9" author="Kayastha P. &amp; Roszkowska M. &amp; Mioduchowska M. &amp; Gawlak M. &amp; Kaczmarek L." pageId="27" pageNumber="106" pagination="394" refId="ref14796" refString="Kayastha P., Roszkowska M., Mioduchowska M., Gawlak M. &amp; Kaczmarek L. 2021. Integrative descriptions of two new tardigrade species along with the new record of Mesobiotus skorackii Kaczmarek et al., 2018 from Canada. Diversity 13: 394. https: // doi. org / 10.3390 / d 13080394" type="journal article" year="2021">Kayastha, Roszkowska, Mioduchowska, Gawlak &amp; Kaczmarek, 2021</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7065B8B0090A68B231F98458A4F9D9" authorityName="Nowak &amp; Stec" authorityYear="2018" box="[546,673,1612,1638]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="hannae">
<emphasis id="B904C229B0090A68B231F98458A4F9D9" box="[546,673,1612,1638]" italics="true" pageId="27" pageNumber="106">M. hannae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7065B8B0090A68B2F3F9845970F9D9" authorityName="Stec" authorityYear="2022" box="[736,885,1612,1638]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="rebecchii">
<emphasis id="B904C229B0090A68B2F3F9845970F9D9" box="[736,885,1612,1638]" italics="true" pageId="27" pageNumber="106">M. rebecchii</emphasis>
</taxonomicName>
(
<figureCitation id="134B02BEB0090A68B394F98559DAF9D9" box="[903,991,1612,1638]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[190,1399,265,1845]" captionTargetId="graphics-54@28.[190,1116,271,1836]" captionTargetPageId="28" captionText="Fig. 21. Bayesian phylogenetic reconstruction of Superclade I of the family Macrobiotidae. Values at nodes are BI posterior probability supports, nodes with support &lt;0.70 were collapsed whereas nodes with maximum support values (1.00) are not shown. Newly sequenced taxa are bolded and marked with red font. The scale bar represents substitutions per position." figureDoi="http://doi.org/10.5281/zenodo.10893842" httpUri="https://zenodo.org/record/10893842/files/figure.png" pageId="27" pageNumber="106">Fig. 21</figureCitation>
). Importantly,
<taxonomicName id="4C7065B8B0090A68B49DF9845F46F9D9" authorityName="Yuan, Wang, Liu, Liu &amp; Li" authorityYear="2022" box="[1166,1347,1612,1638]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="hupingensis">
<emphasis id="B904C229B0090A68B49DF9845F46F9D9" box="[1166,1347,1612,1638]" italics="true" pageId="27" pageNumber="106">M. hupingensis</emphasis>
</taxonomicName>
is a species of the
<taxonomicName id="4C7065B8B0090A68B17AF9B95BECF935" authorityName="Maucci" authorityYear="1954" box="[361,489,1648,1674]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="pallarii">
<emphasis id="B904C229B0090A68B17AF9B95BECF935" box="[361,489,1648,1674]" italics="true" pageId="27" pageNumber="106">M. pallarii</emphasis>
</taxonomicName>
complex, but the DNA sequences associated with this description belong to different unspecified
<taxonomicName id="4C7065B8B0090A68B1AFF95A5854F912" authorityName="Schultze" authorityYear="1834" box="[444,593,1683,1709]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="genus">
<emphasis id="B904C229B0090A68B1AFF95A5854F912" box="[444,593,1683,1709]" italics="true" pageId="27" pageNumber="106">Macrobiotus</emphasis>
</taxonomicName>
and the authors are working on correcting this mistake (Z. Yuan, Shaanxi Normal University, pers. com.). Therefore, the species name is given within quotation marks in the phylogenetic tree (
<figureCitation id="134B02BEB0090A68B1D0F913581EF94A" box="[451,539,1754,1781]" captionStart="Fig" captionStartId="28.[189,232,1892,1918]" captionTargetBox="[190,1399,265,1845]" captionTargetId="graphics-54@28.[190,1116,271,1836]" captionTargetPageId="28" captionText="Fig. 21. Bayesian phylogenetic reconstruction of Superclade I of the family Macrobiotidae. Values at nodes are BI posterior probability supports, nodes with support &lt;0.70 were collapsed whereas nodes with maximum support values (1.00) are not shown. Newly sequenced taxa are bolded and marked with red font. The scale bar represents substitutions per position." figureDoi="http://doi.org/10.5281/zenodo.10893842" httpUri="https://zenodo.org/record/10893842/files/figure.png" pageId="27" pageNumber="106">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BCF1E3BB0090A6EB0AEF8D55F66FE47" blockId="27.[188,1399,1820,2025]" lastBlockId="29.[189,1399,265,504]" lastPageId="29" lastPageNumber="108" pageId="27" pageNumber="106">
The delimitation results of both ASAP analyses were congruent (Supp. file 4). The number of delimited species from the COI data set representing
<taxonomicName id="4C7065B8B0090A68B2AAF8895961F8E5" authorityName="Thulin" authorityYear="1928" box="[697,868,1856,1882]" class="Eutardigrada" family="Macrobiotidae" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="family">Macrobiotidae</taxonomicName>
superclade I, and for the data set comprising only taxa of
<taxonomicName id="4C7065B8B0090A68B15FF8AA5BE4F8C2" authorityName="Schultze" authorityYear="1834" box="[332,481,1891,1917]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="genus">
<emphasis id="B904C229B0090A68B15FF8AA5BE4F8C2" box="[332,481,1891,1917]" italics="true" pageId="27" pageNumber="106">Macrobiotus</emphasis>
</taxonomicName>
, were 73 and 47, respectively. The number of taxa of
<taxonomicName id="4C7065B8B0090A68B45CF8AA5EE1F8C2" authorityName="Schultze" authorityYear="1834" box="[1103,1252,1891,1917]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="genus">
<emphasis id="B904C229B0090A68B45CF8AA5EE1F8C2" box="[1103,1252,1891,1917]" italics="true" pageId="27" pageNumber="106">Macrobiotus</emphasis>
</taxonomicName>
delimited in the larger data set was the same as for the smaller data set (47; Supp. file 4). Two new species described in this study have always been distinguished as two distinct entities (Supp. file 4). Both morphological forms within
<taxonomicName id="4C7065B8B0090A68B14BF8065BC1F857" authority="Stec, 2024" authorityName="Stec" authorityYear="2024" box="[344,452,1998,2024]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="27" pageNumber="106" phylum="Tardigrada" rank="species" species="mileri" status="sp. nov.">
<emphasis id="B904C229B0090A68B14BF8065BC1F857" box="[344,452,1998,2024]" italics="true" pageId="27" pageNumber="106">M. mileri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377F52B0090A68B1D9F8065821F856" box="[458,548,1999,2025]" pageId="27" pageNumber="106" rank="species">sp. nov.</taxonomicNameLabel>
have also been recognized as a single species (Supp. file 4). Interestingly, there were several cases where taxa that were potentially thought as being distinct have been lumped together into singular putative species. These were: (i)
<taxonomicName id="4C7065B8B00F0A6EB34DFEE55AFDFED5" authority="Bertolani &amp; Rebecchi, 1993" authorityName="Bertolani &amp; Rebecchi" authorityYear="1993" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="29" pageNumber="108" phylum="Tardigrada" rank="species" species="sandrae">
<emphasis id="B904C229B00F0A6EB34DFEE55E58FEF9" box="[862,1117,300,326]" italics="true" pageId="29" pageNumber="108">Macrobiotus sandrae</emphasis>
Bertolani &amp; Rebecchi, 1993
</taxonomicName>
and
<taxonomicName id="4C7065B8B00F0A6EB127FE995E97FED5" authority="Ben Marnissi, Cesari, Rebecchi &amp; Bertolani, 2021" authorityName="Ben Marnissi, Cesari, Rebecchi &amp; Bertolani" authorityYear="2021" box="[308,1170,336,362]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="29" pageNumber="108" phylum="Tardigrada" rank="species" species="azzunae">
<emphasis id="B904C229B00F0A6EB127FE995835FED5" box="[308,560,336,362]" italics="true" pageId="29" pageNumber="108">Macrobiotus azzunae</emphasis>
<bibRefCitation id="EFE163CAB00F0A6EB22BFE995E97FED5" author="Ben Marnissi J. &amp; Cesari M. &amp; Rebecchi L. &amp; Bertolani R." box="[568,1170,336,362]" pageId="29" pageNumber="108" pagination="122 - 146" refId="ref13018" refString="Ben Marnissi J., Cesari M., Rebecchi L. &amp; Bertolani R. 2021. Integrative description of a new Tunisian tardigrade species, Macrobiotus azzunae sp. nov. (Eutardigrada, Macrobiotidae, hufelandi group). European Journal of Taxonomy 758: 122 - 146. https: // doi. org / 10.5852 / ejt. 2021.758.1429" type="journal article" year="2021">Ben Marnissi, Cesari, Rebecchi &amp; Bertolani, 2021</bibRefCitation>
</taxonomicName>
, (see
<bibRefCitation id="EFE163CAB00F0A6EB4CBFE995B3AFE32" author="Ben Marnissi J. &amp; Cesari M. &amp; Rebecchi L. &amp; Bertolani R." pageId="29" pageNumber="108" pagination="122 - 146" refId="ref13018" refString="Ben Marnissi J., Cesari M., Rebecchi L. &amp; Bertolani R. 2021. Integrative description of a new Tunisian tardigrade species, Macrobiotus azzunae sp. nov. (Eutardigrada, Macrobiotidae, hufelandi group). European Journal of Taxonomy 758: 122 - 146. https: // doi. org / 10.5852 / ejt. 2021.758.1429" type="journal article" year="2021">
Ben Marnissi
<emphasis id="B904C229B00F0A6EB0AEFEBD5AFDFE32" box="[189,248,371,397]" italics="true" pageId="29" pageNumber="108">et al.</emphasis>
2021
</bibRefCitation>
), (ii)
<taxonomicName id="4C7065B8B00F0A6EB16CFEBA5945FE32" authority="Schultze, 1834" authorityName="Schultze" authorityYear="1834" box="[383,832,371,397]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="29" pageNumber="108" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B904C229B00F0A6EB16CFEBA588FFE32" box="[383,650,371,397]" italics="true" pageId="29" pageNumber="108">Macrobiotus hufelandi</emphasis>
Schultze, 1834
</taxonomicName>
and
<taxonomicName id="4C7065B8B00F0A6EB369FEBD5E33FE32" authorityName="Schultze" authorityYear="1834" box="[890,1078,371,397]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="29" pageNumber="108" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B904C229B00F0A6EB369FEBD599FFE32" box="[890,922,372,397]" italics="true" pageId="29" pageNumber="108">M.</emphasis>
cf.
<emphasis id="B904C229B00F0A6EB3D4FEBA5E33FE32" box="[967,1078,371,397]" italics="true" pageId="29" pageNumber="108">hufelandi</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFE163CAB00F0A6EB467FEBA5F6FFE32" author="Bertolani R. &amp; Rebecchi L. &amp; Giovannini I. &amp; Cesari M." box="[1140,1386,371,397]" pageId="29" pageNumber="108" pagination="19 - 36" refId="ref13109" refString="Bertolani R., Rebecchi L., Giovannini I. &amp; Cesari M. 2011. DNA barcoding and integrative taxonomy of Macrobiotus hufelandi C. A. S. Schultze 1834, the first tardigrade species to be described, and some related species. Zootaxa 2997: 19 - 36. https: // doi. org / 10.11646 / zootaxa. 2997.1.2" type="journal article" year="2011">
Bertolani
<emphasis id="B904C229B00F0A6EB4FAFEBD5F21FE32" box="[1257,1316,371,397]" italics="true" pageId="29" pageNumber="108">et al.</emphasis>
2011
</bibRefCitation>
), (iii)
<taxonomicName id="4C7065B8B00F0A6EB0E6FE5F5AFDFE6B" authority="Bertolani, Cesari, Giovannini, Rebecchi, Guidetti, Kaczmarek &amp; Pilato, 2022" authorityName="Bertolani, Cesari, Giovannini, Rebecchi, Guidetti, Kaczmarek &amp; Pilato" authorityYear="2022" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="29" pageNumber="108" phylum="Tardigrada" rank="species" species="fontourai">
<emphasis id="B904C229B00F0A6EB0E6FE5F5807FE0E" box="[245,514,406,433]" italics="true" pageId="29" pageNumber="108">Macrobiotus fontourai</emphasis>
Bertolani, Cesari, Giovannini, Rebecchi, Guidetti, Kaczmarek &amp; Pilato, 2022
</taxonomicName>
and
<taxonomicName id="4C7065B8B00F0A6EB13DFE725BD6FE6B" authorityName="Bertolani, Cesari, Giovannini, Rebecchi, Guidetti, Kaczmarek &amp; Pilato" authorityYear="2022" box="[302,467,442,468]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="29" pageNumber="108" phylum="Tardigrada" rank="species" species="muralis">
<emphasis id="B904C229B00F0A6EB13DFE725B4AFE6B" box="[302,335,443,468]" italics="true" pageId="29" pageNumber="108">M.</emphasis>
cf.
<emphasis id="B904C229B00F0A6EB16AFE735BD6FE6B" box="[377,467,442,468]" italics="true" pageId="29" pageNumber="108">muralis</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFE163CAB00F0A6EB21DFE735905FE6B" author="Bertolani R. &amp; Cesari M. &amp; Giovannini I. &amp; Rebecchi L. &amp; Guidetti R. &amp; Kaczmarek L. &amp; Pilato G." box="[526,768,442,468]" pageId="29" pageNumber="108" pagination="329 - 368" refId="ref13244" refString="Bertolani R., Cesari M., Giovannini I., Rebecchi L., Guidetti R., Kaczmarek L. &amp; Pilato G. 2023. The Macrobiotus persimilis-polonicus complex (Eutardigrada, Macrobiotidae), another example of problematic species identification, with the description of four new species. Organisms Diversity &amp; Evolution 23: 329 - 368. https: // doi. org / 10.1007 / s 13127 - 022 - 00599 - z" type="journal article" year="2023">
Bertolani
<emphasis id="B904C229B00F0A6EB292FE7258BEFE6B" box="[641,699,442,468]" italics="true" pageId="29" pageNumber="108">et al.</emphasis>
2023
</bibRefCitation>
), (iv)
<taxonomicName id="4C7065B8B00F0A6EB350FE735957FE47" authority="Kayastha, Mioduchowska, Gawlak, Slugocki, Goncalves Silva &amp; Kaczmarek, 2023" authorityName="Kayastha, Mioduchowska, Gawlak, Slugocki, Goncalves Silva &amp; Kaczmarek" authorityYear="2023" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="29" pageNumber="108" phylum="Tardigrada" rank="species" species="kosmali">
<emphasis id="B904C229B00F0A6EB350FE735E3CFE6B" box="[835,1081,442,468]" italics="true" pageId="29" pageNumber="108">Macrobiotus kosmali</emphasis>
<bibRefCitation id="EFE163CAB00F0A6EB453FE735957FE47" author="Kayastha P. &amp; Mioduchowska M. &amp; Gawlak M. &amp; Slugocki L. &amp; Goncalves Silva A. J. J. &amp; Kaczmarek L." pageId="29" pageNumber="108" pagination="126 - 138" refId="ref14857" refString="Kayastha P., Mioduchowska M. Gawlak M., Slugocki L., Goncalves Silva A. J. J. &amp; Kaczmarek L. 2023. Integrative description of Macrobiotus kosmali sp. nov. (hufelandi group) from the Island of Madeira (Portugal). The European Zoological Journal 90 (1): 126 - 138. https: // doi. org / 10.1080 / 24750263.2022.2163312" type="journal article" year="2023">Kayastha, Mioduchowska, Gawlak, Sługocki, Gonçalves Silva &amp; Kaczmarek, 2023</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C7065B8B00F0A6EB398FE175E27FE48" authorityName="Cuenot" authorityYear="1932" box="[907,1058,477,503]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" isUncertain="true" kingdom="Animalia" order="Parachela" pageId="29" pageNumber="108" phylum="Tardigrada" rank="species" species="recens">
<emphasis id="B904C229B00F0A6EB398FE1759AEFE48" box="[907,939,478,503]" italics="true" pageId="29" pageNumber="108">M.</emphasis>
cf.
<emphasis id="B904C229B00F0A6EB3C4FE175E27FE48" box="[983,1058,478,503]" italics="true" pageId="29" pageNumber="108">recens</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFE163CAB00F0A6EB472FE145F52FE47" author="Kayastha P. &amp; Mioduchowska M. &amp; Gawlak M. &amp; Slugocki L. &amp; Goncalves Silva A. J. J. &amp; Kaczmarek L." box="[1121,1367,477,504]" pageId="29" pageNumber="108" pagination="126 - 138" refId="ref14857" refString="Kayastha P., Mioduchowska M. Gawlak M., Slugocki L., Goncalves Silva A. J. J. &amp; Kaczmarek L. 2023. Integrative description of Macrobiotus kosmali sp. nov. (hufelandi group) from the Island of Madeira (Portugal). The European Zoological Journal 90 (1): 126 - 138. https: // doi. org / 10.1080 / 24750263.2022.2163312" type="journal article" year="2023">
Kayastha
<emphasis id="B904C229B00F0A6EB4C6FE175F15FE48" box="[1237,1296,477,503]" italics="true" pageId="29" pageNumber="108">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>