treatments-xml/data/03/D9/87/03D987EFAE7C934296F5FDC5FE261F2D.xml
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<mods:title id="5361792732D7E73D99A5D46F54ECD58E">Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species</mods:title>
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<emphasis id="B904EAEBAE7C934496F5FDC5FE3C1EBD" bold="true" box="[151,418,618,645]" italics="true" pageId="11" pageNumber="212">Liogenys macropelma</emphasis>
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,
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<emphasis id="B904EAEBAE7C934496F5FD7AFEE61ED3" box="[151,376,723,747]" italics="true" pageId="11" pageNumber="212">Liogenys macropelma</emphasis>
<bibRefCitation id="EFE14B08AE7C9344971DFD7AFDB31ED3" author="Bates, H. W." box="[383,557,723,747]" pageId="11" pageNumber="212" pagination="25 - 160" refId="ref17127" refString="Bates, H. W. (1887) Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. In: Salvin, O. &amp; Godman, F. du C. (Eds.), Biologia Centrali-Americana. R. H. Porter, London, pp. 25 - 160." type="book chapter" year="1887">Bates, 1887: 155</bibRefCitation>
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<bibRefCitation id="EFE14B08AE7C93449579FD7DFC7B1ED3" author="Nonfried, A. F." box="[795,997,723,747]" pageId="11" pageNumber="212" pagination="249 - 290" refId="ref18592" refString="Nonfried, A. F. (1892) Verzeichniss der seit 1871 neu beschriebenen Glaphyriden, Melolonthiden und Euchiriden. Berliner Entomologische Zeitschrift, 37, 249 - 290. https: // doi. org / 10.1002 / mmnd. 18920370302" type="journal article" year="1892">Nonfried 1892: 268</bibRefCitation>
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<bibRefCitation id="EFE14B08AE7C9344969FFD5CFE751F32" author="Blackwelder, R. E." box="[253,491,754,778]" pageId="11" pageNumber="212" pagination="189 - 265" refId="ref17187" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous Insects of Mexico, Central America, The West Indies and South America. Part 2. Smithsonian Institution, Washington, D. C., pp. 189 - 265. https: // doi. org / 10.5479 / si. 03629236.185.2" type="book chapter" year="1944">Blackwelder 1944: 227</bibRefCitation>
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<treatmentCitation id="0AD110E8AE7C9344940EFD5CFD621F32" author="Frey, G." box="[620,764,754,778]" page="42" pageId="11" pageNumber="212" year="1969">
<bibRefCitation id="EFE14B08AE7C9344940EFD5CFD621F32" author="Frey, G." box="[620,764,754,778]" pageId="11" pageNumber="212" pagination="38 - 64" refId="ref18071" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey 1969: 42</bibRefCitation>
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<bibRefCitation id="EFE14B08AE7C93449525FD5CFAC91F32" author="United States Department of Agriculture &amp; USDA" box="[839,1367,754,778]" pageId="11" pageNumber="212" pagination="335 - 336" refId="ref18813" refString="United States Department of Agriculture [USDA] (1971) Insects not known to occur in the United States: A Scarab (Liogenys macropelma Bates). Cooperative Economic Insect Report, 21, 335 - 336." type="journal article" year="1971">
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Department of Agriculture 1971: 335
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;
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(natural history);
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<bibRefCitation id="EFE14B08AE7C934497FDFCBFFDD01F11" author="Evans, A. V." box="[415,590,785,809]" pageId="11" pageNumber="212" pagination="1 - 458" refId="ref17920" refString="Evans, A. V. (2003) A checklist of the NewWorld chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211, 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 210</bibRefCitation>
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<bibRefCitation id="EFE14B08AE7C934494B3FCBFFC7C1F11" author="Evans, A. V. &amp; Smith, A. B. T." box="[721,994,784,809]" pageId="11" pageNumber="212" pagination="1 - 344" refId="ref17963" refString="Evans, A. V. &amp; Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) version 3 2009. Papers in Entomology, 1, 1 - 344." type="journal article" year="2009">Evans &amp; Smith 2009: 179</bibRefCitation>
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(checklist); Krajčík 2012: 145 (checklist);
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<bibRefCitation id="EFE14B08AE7C934496A5FC81FE201F7F" author="Cherman, M. A. &amp; Mise, K. M. &amp; Moron, M. A. &amp; Vaz-de-Mello, F. Z. &amp; Almeida, L. M." box="[199,446,815,839]" pageId="11" pageNumber="212" pagination="1 - 120" refId="ref17664" refString="Cherman, M. A., Mise, K. M., Moron, M. A., Vaz-de-Mello, F. Z. &amp; Almeida, L. M. (2017) A taxonomic revision of Liogenys occurring in Brazil with an interactive key and remarks on New World Diplotaxini (Coleoptera, Melolonthidae). ZooKeys, 699, 1 - 120. https: // doi. org / 10.3897 / zookeys. 699.12031" type="journal article" year="2017">
Cherman
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2017: 14
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, 16, 69, 86 (taxonomy); 2019: 18, 64 (taxonomy), 5355 (redescription).
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<treatmentCitation id="0AD110E8AE7C934496F5FCE0FE641F5E" author="Moser, J." box="[151,506,846,870]" page="140" pageId="11" pageNumber="212" year="1921">
<taxonomicName id="4C704D7AAE7C934496F5FCE0FE641F5E" authority="Moser, 1921: 140" authorityName="Moser" authorityPageNumber="140" authorityYear="1921" box="[151,506,846,870]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="gebieni">
<emphasis id="B904EAEBAE7C934496F5FCE0FEDD1F5E" box="[151,323,846,870]" italics="true" pageId="11" pageNumber="212">Liogenys gebieni</emphasis>
<bibRefCitation id="EFE14B08AE7C9344972BFCE0FE641F5E" author="Moser, J." box="[329,506,846,870]" pageId="11" pageNumber="212" pagination="133 - 182" refId="ref18556" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und Sud-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312" type="journal article" year="1921">Moser, 1921: 140</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation id="0AD110E8AE7C93449480FCE0FC531F5E" author="Blackwelder, R. E." box="[738,973,846,870]" page="227" pageId="11" pageNumber="212" year="1944">
<bibRefCitation id="EFE14B08AE7C93449480FCE0FC531F5E" author="Blackwelder, R. E." box="[738,973,846,870]" pageId="11" pageNumber="212" pagination="189 - 265" refId="ref17187" refString="Blackwelder, R. E. (1944) Checklist of the Coleopterous Insects of Mexico, Central America, The West Indies and South America. Part 2. Smithsonian Institution, Washington, D. C., pp. 189 - 265. https: // doi. org / 10.5479 / si. 03629236.185.2" type="book chapter" year="1944">Blackwelder 1944: 227</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="0AD110E8AE7C93449228FCE0FB491F5E" author="Frey, G." box="[1098,1239,846,870]" page="41" pageId="11" pageNumber="212" year="1969">
<bibRefCitation id="EFE14B08AE7C93449228FCE0FB491F5E" author="Frey, G." box="[1098,1239,846,870]" pageId="11" pageNumber="212" pagination="38 - 64" refId="ref18071" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey 1969: 41</bibRefCitation>
</treatmentCitation>
(key);
<treatmentCitation id="0AD110E8AE7C9344937DFCE0FF731FBD" author="Evans, A. V." page="209" pageId="11" pageNumber="212" year="2003">
<bibRefCitation id="EFE14B08AE7C9344937DFCE0FF731FBD" author="Evans, A. V." pageId="11" pageNumber="212" pagination="1 - 458" refId="ref17920" refString="Evans, A. V. (2003) A checklist of the NewWorld chafers (Coleoptera: Scarabaeidae: Melolonthinae). Zootaxa, 211, 1 - 458. https: // doi. org / 10.11646 / zootaxa. 211.1.1" type="journal article" year="2003">Evans 2003: 209</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="0AD110E8AE7C9344970EFCC3FDED1FBD" author="Evans, A. V. &amp; Smith, A. B. T." box="[364,627,876,901]" page="178" pageId="11" pageNumber="212" year="2009">
<bibRefCitation id="EFE14B08AE7C9344970EFCC3FDED1FBD" author="Evans, A. V. &amp; Smith, A. B. T." box="[364,627,876,901]" pageId="11" pageNumber="212" pagination="1 - 344" refId="ref17963" refString="Evans, A. V. &amp; Smith, A. B. T. (2009) An electronic checklist of the New World chafers (Coleoptera: Scarabaeidae: Melolonthinae) version 3 2009. Papers in Entomology, 1, 1 - 344." type="journal article" year="2009">Evans &amp; Smith 2009: 178</bibRefCitation>
</treatmentCitation>
(checklist); Krajčík 2012: 145 (checklist).
<emphasis id="B904EAEBAE7C9344924AFCC3FA1F1FBD" bold="true" box="[1064,1409,876,901]" pageId="11" pageNumber="212">New junior subjective synonym.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE7C934496F5FC03FAD21B48" pageId="11" pageNumber="212" type="materials_examined">
<paragraph id="8BCF36F9AE7C934496F5FC03FCCF18DF" blockId="11.[151,1437,941,1904]" pageId="11" pageNumber="212">
<emphasis id="B904EAEBAE7C934496F5FC03FE5D1FFF" bold="true" box="[151,451,941,967]" pageId="11" pageNumber="212">Type material examined</emphasis>
.
<taxonomicName id="4C704D7AAE7C934497B5FC00FD4D1FFF" authorityName="Bates. The" authorityYear="1887" box="[471,723,941,967]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="macropelma">
<emphasis id="B904EAEBAE7C934497B5FC00FD4D1FFF" box="[471,723,941,967]" italics="true" pageId="11" pageNumber="212">Liogenys macropelma</emphasis>
</taxonomicName>
male
<typeStatus id="54CB885BAE7C93449541FC03FC141FFF" box="[803,906,941,967]" pageId="11" pageNumber="212" type="lectotype">lectotype</typeStatus>
(
<collectionCode id="ED61AE3CAE7C934495FCFC03FC651FFF" box="[926,1019,941,967]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="212">MNHN</collectionCode>
): [white typeset] “V. de
<collectingRegion id="49B4F81BAE7C93449346FC03FA1A1FFF" box="[1316,1412,941,967]" country="Panama" name="Chiriqui" pageId="11" pageNumber="212">Chiriqui</collectingRegion>
, /
<quantity id="4C889B1CAE7C934496F5FC7FFE851FD3" box="[151,283,977,1004]" metricMagnitude="2" metricUnit="m" metricValue="6.1341" metricValueMax="12.192" metricValueMin="0.0762" pageId="11" pageNumber="212" unit="ft" value="2012.5" valueMax="4000.0" valueMin="25.0">254000 ft.</quantity>
/ Champion.”, [white handwritten] “
<taxonomicName id="4C704D7AAE7C934494A3FC7FFCB21FD3" box="[705,812,977,1003]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ macropelma / Bates
<specimenCount id="9D76FD70AE7C93449256FC7FFBD51FD3" box="[1076,1099,977,1003]" pageId="11" pageNumber="212" type="male"></specimenCount>
”, [red typeset] “
<typeStatus id="54CB885BAE7C9344936AFC7FFA101FD3" box="[1288,1422,977,1003]" pageId="11" pageNumber="212" type="syntype">SYNTYPE</typeStatus>
”, [white handwritten] “micropygus”, [white, outlined in red, typeset] “
<typeStatus id="54CB885BAE7C934495F0FC5BFBAB1837" box="[914,1077,1013,1039]" pageId="11" pageNumber="212" type="lectotype">LECTOTYPE</typeStatus>
/
<taxonomicName id="4C704D7AAE7C9344922BFC5BFADC1837" authorityName="Bates. The" authorityYear="1887" box="[1097,1346,1013,1039]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="macropelma">Liogenys macropelma</taxonomicName>
/
<bibRefCitation id="EFE14B08AE7C93449335FC58FF51180B" author="Bates, H. W." pageId="11" pageNumber="212" pagination="25 - 160" refId="ref17127" refString="Bates, H. W. (1887) Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. In: Salvin, O. &amp; Godman, F. du C. (Eds.), Biologia Centrali-Americana. R. H. Porter, London, pp. 25 - 160." type="book chapter" year="1887">Bates, 1887</bibRefCitation>
/ Cherman M. A. 2015”. Genitalia mounted. Six males with the same labels as the
<typeStatus id="54CB885BAE7C934492E5FBB7FB6F180B" box="[1159,1265,1049,1075]" pageId="11" pageNumber="212" type="lectotype">lectotype</typeStatus>
, plus the label: [white, outlined in red, typeset] “
<typeStatus id="54CB885BAE7C93449479FB93FC98186F" box="[539,774,1085,1111]" pageId="11" pageNumber="212" type="paralectotype">PARALECTOTYPE</typeStatus>
/
<taxonomicName id="4C704D7AAE7C93449546FB93FBBF186F" authorityName="Bates. The" authorityYear="1887" box="[804,1057,1085,1111]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="macropelma">Liogenys macropelma</taxonomicName>
/
<bibRefCitation id="EFE14B08AE7C93449222FB90FB56186F" author="Bates, H. W." box="[1088,1224,1085,1112]" pageId="11" pageNumber="212" pagination="25 - 160" refId="ref17127" refString="Bates, H. W. (1887) Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. In: Salvin, O. &amp; Godman, F. du C. (Eds.), Biologia Centrali-Americana. R. H. Porter, London, pp. 25 - 160." type="book chapter" year="1887">Bates, 1887</bibRefCitation>
/ Cherman M. A. 2015”. One female with one different label: [white handwritten] “
<taxonomicName id="4C704D7AAE7C93449517FBCFFC7E1843" box="[885,992,1121,1147]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ macropelma / Bates
<specimenCount id="9D76FD70AE7C934492B9FBCFFB711843" box="[1243,1263,1121,1147]" pageId="11" pageNumber="212" type="female"></specimenCount>
”. Six male and
<specimenCount id="9D76FD70AE7C934496F5FB28FE8A18A7" box="[151,276,1157,1184]" pageId="11" pageNumber="212" type="female">one female</specimenCount>
<typeStatus id="54CB885BAE7C9344977FFB2BFE2118A7" box="[285,447,1157,1183]" pageId="11" pageNumber="212" type="paralectotype">paralectotypes</typeStatus>
(
<collectionCode id="ED61AE3CAE7C934497ADFB2BFDB418A7" box="[463,554,1157,1183]" pageId="11" pageNumber="212">BMNH</collectionCode>
): [white, typeset] “V. de
<collectingRegion id="49B4F81BAE7C93449524FB2BFC3818A7" box="[838,934,1157,1183]" country="Panama" name="Chiriqui" pageId="11" pageNumber="212">Chiriqui</collectingRegion>
, /
<quantity id="4C889B1CAE7C934495A1FB2BFBD818A7" box="[963,1094,1157,1184]" metricMagnitude="2" metricUnit="m" metricValue="6.1341" metricValueMax="12.192" metricValueMin="0.0762" pageId="11" pageNumber="212" unit="ft" value="2012.5" valueMax="4000.0" valueMin="25.0">254000 ft.</quantity>
/ Champion.”, [white, typeset and handwritten] “B.C.A.Col.II. (2) /
<taxonomicName id="4C704D7AAE7C9344945CFB07FD3818FB" box="[574,678,1193,1219]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ macropelma”. One male
<typeStatus id="54CB885BAE7C934495B1FB07FBF418FB" box="[979,1130,1193,1219]" pageId="11" pageNumber="212" type="paralectotype">paralectotype</typeStatus>
also has the labels: [round, white with red border] “Type”, [white, typeset] “Sp. figured.”.
</paragraph>
<paragraph id="8BCF36F9AE7C934496A5FB5CFEF719DB" blockId="11.[151,1437,941,1904]" pageId="11" pageNumber="212">
<taxonomicName id="4C704D7AAE7C934496A5FB5CFE161933" authorityName="Moser" authorityYear="1921" box="[199,392,1265,1291]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="species" species="gebieni">
<emphasis id="B904EAEBAE7C934496A5FB5CFE161933" box="[199,392,1265,1291]" italics="true" pageId="11" pageNumber="212">Liogenys gebieni</emphasis>
</taxonomicName>
<materialsCitation id="3B183CA4AE7C934497F3FB5FFEFB19DB" ID-GBIF-Occurrence="3320554330" collectionCode="ZMHB" collectorName="W. Fritsche" location="Norte de Santander" pageId="11" pageNumber="212" specimenCount="5" stateProvince="Norte de Santander" typeStatus="lectotype">
male
<emphasis id="B904EAEBAE7C934497B3FB5FFDA21933" bold="true" box="[465,572,1265,1291]" pageId="11" pageNumber="212">
<typeStatus id="54CB885BAE7C934497B3FB5FFDA21933" box="[465,572,1265,1291]" pageId="11" pageNumber="212" type="lectotype">lectotype</typeStatus>
</emphasis>
, present designation (
<collectionCode id="ED61AE3CAE7C9344955BFB5FFC0F1933" box="[825,913,1265,1291]" pageId="11" pageNumber="212">ZMHB</collectionCode>
): [white typeset] “Columbien / LaGarita [Los Patios,
<collectingRegion id="49B4F81BAE7C9344968BFAB8FE5C1908" box="[233,450,1301,1328]" country="Colombia" name="Norte de Santander" pageId="11" pageNumber="212">Norte de Santander</collectingRegion>
] (Gebirge [mountains]) /
<collectorName id="2685532FAE7C934494B8FAB8FCC71917" box="[730,857,1301,1328]" pageId="11" pageNumber="212">W. Fritsche</collectorName>
/ leg. 1912” [genitalia mounted on the label], [white handwritten] “Isonychus [misspelled
<taxonomicName id="4C704D7AAE7C93449422FA97FD35196C" box="[576,683,1337,1364]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
] / gebieni / Typen Mos.”, [red typeset] “TYPUS”, [white typeset] “
<taxonomicName id="4C704D7AAE7C934496C3FAF3FE92194F" box="[161,268,1373,1399]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">Liogenys</taxonomicName>
/ gebieni / Mos. ”,. [white, outlined in red, typeset and handwritten] “
<taxonomicName id="4C704D7AAE7C93449243FAF3FB2E194F" box="[1057,1200,1373,1399]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">LIOGENYS</taxonomicName>
/ GEBIENI /
<bibRefCitation id="EFE14B08AE7C9344932DFAF0FF5119A3" author="Moser, J." pageId="11" pageNumber="212" pagination="133 - 182" refId="ref18556" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und Sud-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312" type="journal article" year="1921">Moser, 1921</bibRefCitation>
/
<typeStatus id="54CB885BAE7C93449687FA2FFE1B19A3" box="[229,389,1409,1435]" pageId="11" pageNumber="212" type="lectotype">LECTOTYPE</typeStatus>
/ Des. Cherman M. A.”. One female
<typeStatus id="54CB885BAE7C93449545FA2FFC2019A3" box="[807,958,1409,1435]" pageId="11" pageNumber="212" type="paralectotype">paralectotype</typeStatus>
with the same labels as the
<typeStatus id="54CB885BAE7C9344929BFA2FFAFD19A3" box="[1273,1379,1409,1435]" pageId="11" pageNumber="212" type="lectotype">lectotype</typeStatus>
, plus the label: [white, outlined in red, typeset] “
<typeStatus id="54CB885BAE7C93449419FA0BFCF81987" box="[635,870,1445,1471]" pageId="11" pageNumber="212" type="paralectotype">PARALECTOTYPE</typeStatus>
/ Des. Cherman M. A. [handwritten]
<taxonomicName id="4C704D7AAE7C93449372FA0BFA021987" box="[1296,1436,1445,1471]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="212" phylum="Arthropoda" rank="genus">LIOGENYS</taxonomicName>
/ GEBIENI Mos.”
</materialsCitation>
.
</paragraph>
<paragraph id="8BCF36F9AE7C934496A5FA43FD3F1B73" blockId="11.[151,1437,941,1904]" pageId="11" pageNumber="212">
<emphasis id="B904EAEBAE7C934496A5FA43FDAF1A3F" bold="true" box="[199,561,1517,1543]" pageId="11" pageNumber="212">Additional material examined.</emphasis>
<materialsCitation id="3B183CA4AE7C93449455FA40FE921A77" ID-GBIF-Occurrence="3320554376" collectingDate="1976-05-17" collectingDateMax="1981-05" collectingDateMin="1976-05-17" collectionCode="UNSM" collectorName="Ratcliffe" country="Panama" county="Coco Solo" location="Fort Gulich" municipality="Coco Solo Hospital" pageId="11" pageNumber="212" specimenCount="35" specimenCount-female="22" specimenCount-male="13" stateProvince="Colon">
<collectingCountry id="F3677669AE7C93449455FA40FD2C1A30" box="[567,690,1518,1544]" name="Panama" pageId="11" pageNumber="212">PANAMA</collectingCountry>
.
<collectingRegion id="49B4F81BAE7C934494D8FA43FC9D1A3F" box="[698,771,1517,1543]" country="Panama" name="Colon" pageId="11" pageNumber="212">Colón</collectingRegion>
:
<collectingCounty id="62AE4E75AE7C9344956EFA43FC1D1A3F" box="[780,899,1517,1543]" pageId="11" pageNumber="212">Coco Solo</collectingCounty>
,
<date id="FFCE1039AE7C934495EFFA43FB9C1A3F" box="[909,1026,1517,1543]" pageId="11" pageNumber="212" value="1976-05-21">
<collectingDate id="EF8AE9D1AE7C934495EFFA43FB9C1A3F" box="[909,1026,1517,1543]" pageId="11" pageNumber="212" value="1976-05-21">21.V.1976</collectingDate>
</date>
, B.C.
<collectorName id="2685532FAE7C93449224FA43FB341A3F" box="[1094,1194,1517,1543]" pageId="11" pageNumber="212">Ratcliffe</collectorName>
,
<specimenCount id="9D76FD70AE7C934492D6FA43FA871A3F" box="[1204,1305,1517,1543]" pageId="11" pageNumber="212" type="male">10 males</specimenCount>
,
<specimenCount id="9D76FD70AE7C93449341FA43FA021A3F" box="[1315,1436,1517,1543]" pageId="11" pageNumber="212" type="female">20 females</specimenCount>
(
<collectionCode id="ED61AE3CAE7C934496FDF9BFFF681A13" box="[159,246,1553,1579]" country="USA" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="11" pageNumber="212">UNSM</collectionCode>
);
<collectingMunicipality id="6BABAC83AE7C93449768F9BFFE741A13" box="[266,490,1553,1579]" pageId="11" pageNumber="212">Coco Solo Hospital</collectingMunicipality>
,
<date id="FFCE1039AE7C93449796F9BFFDD91A13" box="[500,583,1553,1579]" pageId="11" pageNumber="212" value="1981-05">
<collectingDate id="EF8AE9D1AE7C93449796F9BFFDD91A13" box="[500,583,1553,1579]" pageId="11" pageNumber="212" value="1981-05">V.1981</collectingDate>
</date>
,
<specimenCount id="9D76FD70AE7C93449431F9BFFD371A13" box="[595,681,1553,1579]" pageId="11" pageNumber="212" type="male">2 males</specimenCount>
(
<collectionCode id="ED61AE3CAE7C934494DBF9BFFC8E1A13" box="[697,784,1553,1579]" country="USA" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="11" pageNumber="212">UNSM</collectionCode>
);
<location id="8EAF6022AE7C93449546F9BCFC371A13" LSID="urn:lsid:plazi:treatment:03D987EFAE7C934296F5FDC5FE261F2D:8EAF6022AE7C93449546F9BCFC371A13" box="[804,937,1553,1580]" country="Panama" county="Coco Solo" municipality="Coco Solo Hospital" name="Fort Gulich" pageId="11" pageNumber="212" stateProvince="Colon">Fort Gulich</location>
,
<date id="FFCE1039AE7C934495D7F9BFFBB71A13" box="[949,1065,1553,1579]" pageId="11" pageNumber="212" value="1976-05-17">
<collectingDate id="EF8AE9D1AE7C934495D7F9BFFBB71A13" box="[949,1065,1553,1579]" pageId="11" pageNumber="212" value="1976-05-17">17.V.1976</collectingDate>
</date>
, at lights, B.C.
<collectorName id="2685532FAE7C934492BBF9BFFAA31A13" box="[1241,1341,1553,1579]" pageId="11" pageNumber="212">Ratcliffe</collectorName>
,
<specimenCount id="9D76FD70AE7C9344932BF9BFFA061A13" box="[1353,1432,1553,1579]" pageId="11" pageNumber="212" type="male">1 male</specimenCount>
,
<specimenCount id="9D76FD70AE7C934496F5F99BFE921A77" box="[151,268,1589,1615]" pageId="11" pageNumber="212" type="female">2 females</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B183CA4AE7C93449779F99BFA071A77" ID-GBIF-Occurrence="3320554366" box="[283,1433,1589,1616]" collectingDate="1980-04-26" collectingDateMax="1981-04" collectingDateMin="1980-04-26" collectionCode="UNSM" collectorName="Engleman" country="Panama" location="Fort Sherman" pageId="11" pageNumber="212" specimenCount="6" specimenCount-female="1" specimenCount-male="5">
<date id="FFCE1039AE7C93449779F99BFEE91A77" box="[283,375,1589,1615]" pageId="11" pageNumber="212" value="1981-04">
<collectingDate id="EF8AE9D1AE7C93449779F99BFEE91A77" box="[283,375,1589,1615]" pageId="11" pageNumber="212" value="1981-04">IV.1981</collectingDate>
</date>
, at lights,
<specimenCount id="9D76FD70AE7C9344979AF99BFDCD1A77" box="[504,595,1589,1616]" pageId="11" pageNumber="212" type="male">5 males</specimenCount>
(
<collectionCode id="ED61AE3CAE7C93449405F99BFD201A77" box="[615,702,1589,1615]" country="USA" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="11" pageNumber="212">UNSM</collectionCode>
);
<location id="8EAF6022AE7C934494B4F998FCE81A77" LSID="urn:lsid:plazi:treatment:03D987EFAE7C934296F5FDC5FE261F2D:8EAF6022AE7C934494B4F998FCE81A77" box="[726,886,1589,1616]" country="Panama" name="Fort Sherman" pageId="11" pageNumber="212">Fort Sherman</location>
,
<date id="FFCE1039AE7C934495E4F99BFB9B1A77" box="[902,1029,1589,1615]" pageId="11" pageNumber="212" value="1980-04-26">
<collectingDate id="EF8AE9D1AE7C934495E4F99BFB9B1A77" box="[902,1029,1589,1615]" pageId="11" pageNumber="212" value="1980-04-26">26.IV.1980</collectingDate>
</date>
, D.
<collectorName id="2685532FAE7C9344925EF99BFB2E1A77" box="[1084,1200,1589,1615]" pageId="11" pageNumber="212">Engleman</collectorName>
,
<specimenCount id="9D76FD70AE7C934492A2F99BFAB81A77" box="[1216,1318,1589,1615]" pageId="11" pageNumber="212" type="female">1 female</specimenCount>
(
<collectionCode id="ED61AE3CAE7C93449358F99BFA0F1A77" box="[1338,1425,1589,1615]" country="USA" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="11" pageNumber="212">UNSM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B183CA4AE7C934496F5F9F7FB071AE7" ID-GBIF-Occurrence="3320554334" collectingDate="1911-04-27" collectingDateMax="1977-06-19" collectingDateMin="1911-04-27" collectionCode="CMNC, USNM, CNCI" country="Panama" county="Balboa" location="Fort Clayton" municipality="Fort Clayton" pageId="11" pageNumber="212" specimenCount="21" specimenCount-female="4" specimenCount-male="17" stateProvince="Panama">
<collectingRegion id="49B4F81BAE7C934496F5F9F7FF6A1A4B" box="[151,244,1625,1651]" country="Panama" name="Panama" pageId="11" pageNumber="212">Panamá</collectingRegion>
:
<collectingCounty id="62AE4E75AE7C93449762F9F7FECC1A4B" box="[256,338,1625,1651]" pageId="11" pageNumber="212">Balboa</collectingCounty>
,
<date id="FFCE1039AE7C9344973CF9F7FE631A4B" box="[350,509,1625,1651]" pageId="11" pageNumber="212" value="1977-05-16" valueMax="1977-05-17" valueMin="1977-05-16">
<collectingDate id="EF8AE9D1AE7C9344973CF9F7FE631A4B" box="[350,509,1625,1651]" pageId="11" pageNumber="212" value="1977-05-16" valueMax="1977-05-17" valueMin="1977-05-16">1617.V.1977</collectingDate>
</date>
, light,
<collectorName id="2685532FAE7C9344942EF9F7FC9C1A4B" box="[588,770,1625,1651]" pageId="11" pageNumber="212">H.&amp;A. Howden</collectorName>
,
<specimenCount id="9D76FD70AE7C9344956CF9F7FCF61A4B" box="[782,872,1625,1651]" pageId="11" pageNumber="212" type="male">3 males</specimenCount>
,
<specimenCount id="9D76FD70AE7C93449516F9F7FC481A4B" box="[884,982,1625,1651]" pageId="11" pageNumber="212" type="female">1 female</specimenCount>
(
<collectionCode id="ED61AE3CAE7C93449585F9F7FBDE1A4B" box="[999,1088,1625,1651]" pageId="11" pageNumber="212">CMNC</collectionCode>
);
<date id="FFCE1039AE7C93449236F9F7FB4B1A4B" box="[1108,1237,1625,1651]" pageId="11" pageNumber="212" value="1977-06-19">
<collectingDate id="EF8AE9D1AE7C93449236F9F7FB4B1A4B" box="[1108,1237,1625,1651]" pageId="11" pageNumber="212" value="1977-06-19">19.VI.1977</collectingDate>
</date>
,
<collectorName id="2685532FAE7C93449280F9F7FA061A4B" box="[1250,1432,1625,1651]" pageId="11" pageNumber="212">H.&amp;A. Howden</collectorName>
,
<specimenCount id="9D76FD70AE7C934496F5F9D3FF7D1AAF" box="[151,227,1661,1687]" pageId="11" pageNumber="212" type="male">1 male</specimenCount>
(
<collectionCode id="ED61AE3CAE7C93449690F9D3FED51AAF" box="[242,331,1661,1687]" pageId="11" pageNumber="212">CMNC</collectionCode>
); Corazal,
<date id="FFCE1039AE7C934497A6F9D3FDDC1AAF" box="[452,578,1661,1687]" pageId="11" pageNumber="212" value="1911-04-27">
<collectingDate id="EF8AE9D1AE7C934497A6F9D3FDDC1AAF" box="[452,578,1661,1687]" pageId="11" pageNumber="212" value="1911-04-27">27.IV.1911</collectingDate>
</date>
,
<collectorName id="2685532FAE7C9344942EF9D0FD261AAF" box="[588,696,1661,1688]" pageId="11" pageNumber="212">A. Busck</collectorName>
,
<specimenCount id="9D76FD70AE7C934494A1F9D3FC841AAF" box="[707,794,1661,1687]" pageId="11" pageNumber="212" type="male">2 males</specimenCount>
(
<collectionCode id="ED61AE3CAE7C9344954BF9D3FC1E1AAF" box="[809,896,1661,1687]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="212" type="Museum">USNM</collectionCode>
); Curundú,
<date id="FFCE1039AE7C93449267F9D3FBF21AAF" box="[1029,1132,1661,1687]" pageId="11" pageNumber="212" value="1972-05-08">
<collectingDate id="EF8AE9D1AE7C93449267F9D3FBF21AAF" box="[1029,1132,1661,1687]" pageId="11" pageNumber="212" value="1972-05-08">8.V.1972</collectingDate>
</date>
,
<collectorName id="2685532FAE7C9344921AF9D3FADB1AAF" box="[1144,1349,1661,1687]" pageId="11" pageNumber="212">R.&amp;E. Froeschner</collectorName>
,
<specimenCount id="9D76FD70AE7C93449332F9D3FA021AAF" box="[1360,1436,1661,1687]" pageId="11" pageNumber="212" type="male">1 male</specimenCount>
(
<collectionCode id="ED61AE3CAE7C934496FDF90FFF681A83" box="[159,246,1697,1723]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="212" type="Museum">USNM</collectionCode>
); Diablo Heights,
<date id="FFCE1039AE7C934497A0F90FFDB71A83" box="[450,553,1697,1723]" pageId="11" pageNumber="212" value="1971-05-03">
<collectingDate id="EF8AE9D1AE7C934497A0F90FFDB71A83" box="[450,553,1697,1723]" pageId="11" pageNumber="212" value="1971-05-03">3.V.1971</collectingDate>
</date>
,
<collectorName id="2685532FAE7C93449456F90CFD2C1A83" box="[564,690,1697,1724]" pageId="11" pageNumber="212">W.E. Bivin</collectorName>
,
<specimenCount id="9D76FD70AE7C934494DCF90FFC921A83" box="[702,780,1697,1723]" pageId="11" pageNumber="212" type="male">1 male</specimenCount>
,
<specimenCount id="9D76FD70AE7C93449575F90FFCE61A83" box="[791,888,1697,1723]" pageId="11" pageNumber="212" type="female">1 female</specimenCount>
(
<collectionCode id="ED61AE3CAE7C934495E5F90FFC401A83" box="[903,990,1697,1723]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="212" type="Museum">USNM</collectionCode>
);
<collectingMunicipality id="6BABAC83AE7C93449593F90CFB1C1A83" box="[1009,1154,1697,1724]" pageId="11" pageNumber="212">Fort Clayton</collectingMunicipality>
,
<date id="FFCE1039AE7C934492EFF90FFA9C1A84" box="[1165,1282,1697,1724]" pageId="11" pageNumber="212" value="1961-05-15">
<collectingDate id="EF8AE9D1AE7C934492EFF90FFA9C1A84" box="[1165,1282,1697,1724]" pageId="11" pageNumber="212" value="1961-05-15">15.V.1961</collectingDate>
</date>
,
<collectorName id="2685532FAE7C9344936CF90FFA061A83" box="[1294,1432,1697,1723]" pageId="11" pageNumber="212">C.E. Yunker</collectorName>
,
<specimenCount id="9D76FD70AE7C934496F5F96BFF711AE7" box="[151,239,1733,1759]" pageId="11" pageNumber="212" type="male">8 males</specimenCount>
,
<specimenCount id="9D76FD70AE7C9344969BF96BFEFA1AE7" box="[249,356,1733,1759]" pageId="11" pageNumber="212" type="female">2 females</specimenCount>
(
<collectionCode id="ED61AE3CAE7C93449713F96BFE241AE7" box="[369,442,1733,1759]" country="Canada" name="Canadian National Collection Insects" pageId="11" pageNumber="212">CNCI</collectionCode>
);
<location id="8EAF6022AE7C934497AEF968FDC21AE7" LSID="urn:lsid:plazi:treatment:03D987EFAE7C934296F5FDC5FE261F2D:8EAF6022AE7C934497AEF968FDC21AE7" box="[460,604,1733,1760]" country="Panama" county="Balboa" municipality="Fort Clayton" name="Fort Clayton" pageId="11" pageNumber="212" stateProvince="Panama">Fort Clayton</location>
,
<date id="FFCE1039AE7C93449404F96BFD261AE7" box="[614,696,1733,1759]" pageId="11" pageNumber="212" value="1961-05">
<collectingDate id="EF8AE9D1AE7C93449404F96BFD261AE7" box="[614,696,1733,1759]" pageId="11" pageNumber="212" value="1961-05">V.1961</collectingDate>
</date>
,
<collectorName id="2685532FAE7C934494A0F96BFCD51AE7" box="[706,843,1733,1759]" pageId="11" pageNumber="212">C.E. Yunker</collectorName>
,
<specimenCount id="9D76FD70AE7C93449537F96BFC001AE7" box="[853,926,1733,1759]" pageId="11" pageNumber="212" type="male">1 male</specimenCount>
(
<collectionCode id="ED61AE3CAE7C934495CEF96BFB9B1AE7" box="[940,1029,1733,1759]" pageId="11" pageNumber="212">CMNC</collectionCode>
);
<collectorName id="2685532FAE7C93449275F968FB071AE7" box="[1047,1177,1733,1760]" pageId="11" pageNumber="212">La Sabanas</collectorName>
</materialsCitation>
,
<materialsCitation id="3B183CA4AE7C934492C1F968FF771B1F" ID-GBIF-Occurrence="3320554364" collectingDate="1911-04-24" collectingDateMax="1927-04" collectingDateMin="1911-04-24" collectionCode="USNM" collectorName="J. Zetek &amp; A. Busck" country="Panama" location="Punta Paitilla" municipality="Paraiso" pageId="11" pageNumber="212" specimenCount="29" specimenCount-female="20" specimenCount-male="9" stateProvince="Panama">
<collectingRegion id="49B4F81BAE7C934492C1F968FAAC1AE7" box="[1187,1330,1733,1760]" country="Panama" name="Panama" pageId="11" pageNumber="212">Panama City</collectingRegion>
,
<date id="FFCE1039AE7C9344935EF96BFA061AE7" box="[1340,1432,1733,1759]" pageId="11" pageNumber="212" value="1927-04">
<collectingDate id="EF8AE9D1AE7C9344935EF96BFA061AE7" box="[1340,1432,1733,1759]" pageId="11" pageNumber="212" value="1927-04">IV.1927</collectingDate>
</date>
, at light,
<collectorName id="2685532FAE7C93449697F944FECC1B3B" box="[245,338,1769,1796]" pageId="11" pageNumber="212">J. Zetek</collectorName>
,
<specimenCount id="9D76FD70AE7C9344973FF944FE291B3B" box="[349,439,1769,1796]" pageId="11" pageNumber="212" type="male">7 males</specimenCount>
,
<specimenCount id="9D76FD70AE7C934497A1F947FDA01B3B" box="[451,574,1769,1795]" pageId="11" pageNumber="212" type="female">19 females</specimenCount>
(
<collectionCode id="ED61AE3CAE7C9344942CF947FD3B1B3B" box="[590,677,1769,1795]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="212" type="Museum">USNM</collectionCode>
);
<collectingMunicipality id="6BABAC83AE7C934494DBF947FC901B3B" box="[697,782,1769,1795]" pageId="11" pageNumber="212">Paraiso</collectingMunicipality>
,
<date id="FFCE1039AE7C93449578F947FC061B3B" box="[794,920,1769,1795]" pageId="11" pageNumber="212" value="1911-04-24">
<collectingDate id="EF8AE9D1AE7C93449578F947FC061B3B" box="[794,920,1769,1795]" pageId="11" pageNumber="212" value="1911-04-24">24.IV.1911</collectingDate>
</date>
,
<collectorName id="2685532FAE7C934495C0F944FB901B3B" box="[930,1038,1769,1796]" pageId="11" pageNumber="212">A. Busck</collectorName>
,
<specimenCount id="9D76FD70AE7C93449278F947FBEA1B3B" box="[1050,1140,1769,1795]" pageId="11" pageNumber="212" type="male">2 males</specimenCount>
,
<specimenCount id="9D76FD70AE7C934492E2F947FB7F1B3B" box="[1152,1249,1769,1795]" pageId="11" pageNumber="212" type="female">1 female</specimenCount>
(
<collectionCode id="ED61AE3CAE7C93449293F947FAD61B3B" box="[1265,1352,1769,1795]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="212" type="Museum">USNM</collectionCode>
);
<location id="8EAF6022AE7C9344933EF944FF771B1F" LSID="urn:lsid:plazi:treatment:03D987EFAE7C934296F5FDC5FE261F2D:8EAF6022AE7C9344933EF944FF771B1F" country="Panama" municipality="Paraiso" name="Punta Paitilla" pageId="11" pageNumber="212" stateProvince="Panama">Punta Paitilla</location>
</materialsCitation>
,
<materialsCitation id="3B183CA4AE7C93449697F8A0FB621B1F" ID-GBIF-Occurrence="3320554342" box="[245,1276,1805,1832]" collectingDate="1970-05-17" collectionCode="CMNC" collectorName="Hespenheide" country="Panama" latitude="8.983334" location="Trivoli Hotel" longLatPrecision="1300" longitude="-79.51667" pageId="11" pageNumber="212" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<collectingRegion id="49B4F81BAE7C93449697F8A0FE1B1B1F" box="[245,389,1805,1832]" country="Panama" name="Panama" pageId="11" pageNumber="212">Panama City</collectingRegion>
,
<geoCoordinate id="EE44503EAE7C934497F3F8A3FE7D1B10" box="[401,483,1805,1832]" degrees="8" direction="north" minutes="59" orientation="latitude" pageId="11" pageNumber="212" precision="925" value="8.983334">8º59N</geoCoordinate>
,
<geoCoordinate id="EE44503EAE7C9344978DF8A3FDCA1B1F" box="[495,596,1805,1831]" degrees="79" direction="west" minutes="31" orientation="longitude" pageId="11" pageNumber="212" precision="925" value="-79.51667">79º31`W</geoCoordinate>
,
<date id="FFCE1039AE7C9344943DF8A3FD4B1B1F" box="[607,725,1805,1831]" pageId="11" pageNumber="212" value="1970-05-17">
<collectingDate id="EF8AE9D1AE7C9344943DF8A3FD4B1B1F" box="[607,725,1805,1831]" pageId="11" pageNumber="212" value="1970-05-17">17.V.1970</collectingDate>
</date>
, H.
<collectorName id="2685532FAE7C93449560F8A3FC081B1F" box="[770,918,1805,1831]" pageId="11" pageNumber="212">Hespenheide</collectorName>
,
<specimenCount id="9D76FD70AE7C934495C3F8A3FC721B1F" box="[929,1004,1805,1831]" pageId="11" pageNumber="212" type="male">1 male</specimenCount>
(
<collectionCode id="ED61AE3CAE7C93449599F8A3FBCA1B1F" box="[1019,1108,1805,1831]" pageId="11" pageNumber="212">CMNC</collectionCode>
);
<location id="8EAF6022AE7C93449205F8A3FB621B1F" LSID="urn:lsid:plazi:treatment:03D987EFAE7C934296F5FDC5FE261F2D:8EAF6022AE7C93449205F8A3FB621B1F" box="[1127,1276,1805,1831]" country="Panama" latitude="8.983334" longLatPrecision="1300" longitude="-79.51667" name="Trivoli Hotel" pageId="11" pageNumber="212" stateProvince="Panama">Trivoli Hotel</location>
</materialsCitation>
,
<materialsCitation id="3B183CA4AE7C93449365F8A0FD031B73" ID-GBIF-Occurrence="3320554381" collectingDate="1967-04-15" collectionCode="CMNC" collectorName="Hespenheide" country="Panama" location="Panama City" pageId="11" pageNumber="212" specimenCount="1" specimenCount-male="1" stateProvince="Panama">
<collectingRegion id="49B4F81BAE7C93449365F8A0FA061B1F" box="[1287,1432,1805,1832]" country="Panama" name="Panama" pageId="11" pageNumber="212">Panama City</collectingRegion>
,
<date id="FFCE1039AE7C934496F5F89FFE881B74" box="[151,278,1841,1868]" pageId="11" pageNumber="212" value="1967-04-15">
<collectingDate id="EF8AE9D1AE7C934496F5F89FFE881B74" box="[151,278,1841,1868]" pageId="11" pageNumber="212" value="1967-04-15">15.IV.1967</collectingDate>
</date>
, H.
<collectorName id="2685532FAE7C93449721F89FFE491B73" box="[323,471,1841,1867]" pageId="11" pageNumber="212">Hespenheide</collectorName>
,
<specimenCount id="9D76FD70AE7C93449780F89FFDB31B73" box="[482,557,1841,1867]" pageId="11" pageNumber="212" type="male">1 male</specimenCount>
(
<collectionCode id="ED61AE3CAE7C9344945EF89FFD0B1B73" box="[572,661,1841,1867]" pageId="11" pageNumber="212">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCF36F9AE7C934496A5F8FBFAD21B48" blockId="11.[151,1437,941,1904]" box="[199,1356,1877,1904]" pageId="11" pageNumber="212">
<emphasis id="B904EAEBAE7C934496A5F8FBFE601B57" bold="true" box="[199,510,1877,1903]" pageId="11" pageNumber="212">Geographical distribution</emphasis>
.
<collectingCountry id="F3677669AE7C9344946EF8F8FD1F1B48" box="[524,641,1878,1904]" name="Panama" pageId="11" pageNumber="212">PANAMA</collectingCountry>
(
<collectingRegion id="49B4F81BAE7C934494EDF8FBFD6F1B57" box="[655,753,1877,1903]" country="Panama" name="Chiriqui" pageId="11" pageNumber="212">Chiriquí</collectingRegion>
,
<collectingRegion id="49B4F81BAE7C9344949EF8FBFCDD1B57" box="[764,835,1877,1903]" country="Panama" name="Colon" pageId="11" pageNumber="212">Colón</collectingRegion>
,
<collectingCountry id="F3677669AE7C9344952DF8FBFC321B57" box="[847,940,1877,1903]" name="Panama" pageId="11" pageNumber="212">Panamá</collectingCountry>
),
<collectingCountry id="F3677669AE7C934495DDF8FBFBCD1B57" box="[959,1107,1877,1903]" name="Colombia" pageId="11" pageNumber="212">COLOMBIA</collectingCountry>
(
<collectingRegion id="49B4F81BAE7C93449203F8FBFADE1B48" box="[1121,1344,1877,1904]" country="Colombia" name="Norte de Santander" pageId="11" pageNumber="212">Norte de Santander</collectingRegion>
).
</paragraph>
</subSubSection>
<caption id="DF0F6671AE7B934396F5F839FF651BC0" ID-DOI="http://doi.org/10.5281/zenodo.5025910" ID-Zenodo-Dep="5025910" httpUri="https://zenodo.org/record/5025910/files/figure.png" pageId="12" pageNumber="213" startId="12.[151,250,1943,1967]" targetBox="[254,1436,181,1920]" targetPageId="12">
<paragraph id="8BCF36F9AE7B934396F5F839FF651BC0" blockId="12.[151,1437,1943,2040]" pageId="12" pageNumber="213">
<emphasis id="B904EAEBAE7B934396F5F839FE8D1B88" bold="true" box="[151,275,1943,1968]" pageId="12" pageNumber="213">FIGURE 5.</emphasis>
<taxonomicName id="4C704D7AAE7B93439778F836FDD31B88" authority="Moser, 1921" authorityName="Moser" authorityYear="1921" box="[282,589,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="gebieni">
<emphasis id="B904EAEBAE7B93439778F836FE591B88" box="[282,455,1944,1968]" italics="true" pageId="12" pageNumber="213">Liogenys gebieni</emphasis>
<bibRefCitation id="EFE14B08AE7B934397ACF836FDD31B88" author="Moser, J." box="[462,589,1944,1968]" pageId="12" pageNumber="213" pagination="133 - 182" refId="ref18556" refString="Moser, J. (1921) Neue Melolonthiden von Mittel- und Sud-Amerika. Stettiner Entomologische Zeitung, 82, 133 - 182. https: // doi. org / 10.1002 / mmnd. 192119210312" type="journal article" year="1921">Moser, 1921</bibRefCitation>
</taxonomicName>
lectotype views: A) dorsal, B) dorsal of female paralectotype, C) labels, D) lateral, E) pygidium, and dorsal view of F) one paramere compared with G) parameres of the
<taxonomicName id="4C704D7AAE7B93439256F812FA031BEC" authority="Bates, 1887" authorityName="Bates. The" authorityYear="1887" box="[1076,1437,1980,2004]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="213" phylum="Arthropoda" rank="species" species="macropelma">
<emphasis id="B904EAEBAE7B93439256F812FA861BEC" box="[1076,1304,1980,2004]" italics="true" pageId="12" pageNumber="213">Liogenys macropelma</emphasis>
<bibRefCitation id="EFE14B08AE7B93439343F812FA031BEC" author="Bates, H. W." box="[1313,1437,1980,2004]" pageId="12" pageNumber="213" pagination="25 - 160" refId="ref17127" refString="Bates, H. W. (1887) Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia. In: Salvin, O. &amp; Godman, F. du C. (Eds.), Biologia Centrali-Americana. R. H. Porter, London, pp. 25 - 160." type="book chapter" year="1887">Bates, 1887</bibRefCitation>
</taxonomicName>
lectotype.
</paragraph>
</caption>
<subSubSection id="C36A6572AE7A934296A5FF39FE261F2D" pageId="13" pageNumber="214" type="discussion">
<paragraph id="8BCF36F9AE7A934296A5FF39FE261F2D" blockId="13.[151,1437,150,789]" pageId="13" pageNumber="214">
<emphasis id="B904EAEBAE7A934296A5FF39FEAA1C89" bold="true" box="[199,308,151,177]" pageId="13" pageNumber="214">Remarks</emphasis>
. After studying the
<typeStatus id="54CB885BAE7A93429445FF39FDC91C89" box="[551,599,151,177]" pageId="13" pageNumber="214">type</typeStatus>
series of
<taxonomicName id="4C704D7AAE7A934294B0FF39FC291C89" authority="(ZMHB)" baseAuthorityName="ZMHB" box="[722,951,151,177]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="gebieni">
<emphasis id="B904EAEBAE7A934294B0FF39FCD91C89" box="[722,839,151,177]" italics="true" pageId="13" pageNumber="214">L. gebieni</emphasis>
(ZMHB)
</taxonomicName>
(
<figureCitation id="134B2A7CAE7A934295A9FF39FBD41C89" box="[971,1098,150,177]" captionStart="FIGURE 5" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[254,1436,181,1920]" captionTargetId="figure-18@12.[254,1436,181,1920]" captionTargetPageId="12" captionText="FIGURE 5. Liogenys gebieni Moser, 1921 lectotype views: A) dorsal, B) dorsal of female paralectotype, C) labels, D) lateral, E) pygidium, and dorsal view of F) one paramere compared with G) parameres of the Liogenys macropelma Bates, 1887 lectotype." figureDoi="http://doi.org/10.5281/zenodo.5025910" httpUri="https://zenodo.org/record/5025910/files/figure.png" pageId="13" pageNumber="214">Fig. 5AC</figureCitation>
), and comparing its external features and male genitalia (
<figureCitation id="134B2A7CAE7A934297BEFF15FDCD1CED" box="[476,595,186,213]" captionStart="FIGURE 5" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[254,1436,181,1920]" captionTargetId="figure-18@12.[254,1436,181,1920]" captionTargetPageId="12" captionText="FIGURE 5. Liogenys gebieni Moser, 1921 lectotype views: A) dorsal, B) dorsal of female paralectotype, C) labels, D) lateral, E) pygidium, and dorsal view of F) one paramere compared with G) parameres of the Liogenys macropelma Bates, 1887 lectotype." figureDoi="http://doi.org/10.5281/zenodo.5025910" httpUri="https://zenodo.org/record/5025910/files/figure.png" pageId="13" pageNumber="214">Fig. 5AF</figureCitation>
) with that of
<taxonomicName id="4C704D7AAE7A93429490FF15FC031CED" authorityName="Bates. The" authorityYear="1887" box="[754,925,187,213]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="macropelma">
<emphasis id="B904EAEBAE7A93429490FF15FC031CED" box="[754,925,187,213]" italics="true" pageId="13" pageNumber="214">L. macropelma</emphasis>
</taxonomicName>
<typeStatus id="54CB885BAE7A934295C4FF15FC481CED" box="[934,982,187,213]" pageId="13" pageNumber="214">type</typeStatus>
series and non-type material (
<figureCitation id="134B2A7CAE7A93429351FF15FA101CED" box="[1331,1422,186,213]" captionStart="FIGURE 5" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[254,1436,181,1920]" captionTargetId="figure-18@12.[254,1436,181,1920]" captionTargetPageId="12" captionText="FIGURE 5. Liogenys gebieni Moser, 1921 lectotype views: A) dorsal, B) dorsal of female paralectotype, C) labels, D) lateral, E) pygidium, and dorsal view of F) one paramere compared with G) parameres of the Liogenys macropelma Bates, 1887 lectotype." figureDoi="http://doi.org/10.5281/zenodo.5025910" httpUri="https://zenodo.org/record/5025910/files/figure.png" pageId="13" pageNumber="214">Fig. 5G</figureCitation>
), we concluded that these specimens belong to the same species. The only difference is the color, which is darker in the
<taxonomicName id="4C704D7AAE7A934296A2FEADFEB11D25" box="[192,303,259,285]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="gebieni">
<emphasis id="B904EAEBAE7A934296A2FEADFEB11D25" box="[192,303,259,285]" italics="true" pageId="13" pageNumber="214">L. gebieni</emphasis>
</taxonomicName>
<typeStatus id="54CB885BAE7A93429754FEADFEEB1D25" box="[310,373,259,285]" pageId="13" pageNumber="214">types</typeStatus>
. Nonetheless, the difference in body color is a common intraspecific variation within
<taxonomicName id="4C704D7AAE7A93429352FEADFA0B1D24" box="[1328,1429,259,284]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B904EAEBAE7A93429352FEADFA0B1D24" box="[1328,1429,259,284]" italics="true" pageId="13" pageNumber="214">Liogenys</emphasis>
</taxonomicName>
. Herein, we propose
<taxonomicName id="4C704D7AAE7A93429719FE89FDA51D79" authority="Moser" authorityName="Moser" authorityYear="1921" box="[379,571,295,321]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="gebieni">
<emphasis id="B904EAEBAE7A93429719FE89FE751D79" box="[379,491,295,321]" italics="true" pageId="13" pageNumber="214">L. gebieni</emphasis>
Moser
</taxonomicName>
as a
<emphasis id="B904EAEBAE7A93429414FE88FC2C1D79" bold="true" box="[630,946,294,321]" pageId="13" pageNumber="214">junior subjective synonym</emphasis>
of
<taxonomicName id="4C704D7AAE7A934295BBFE89FA9D1D79" authority="Bates. The" authorityName="Bates. The" authorityYear="1887" box="[985,1283,295,321]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="macropelma">
<emphasis id="B904EAEBAE7A934295BBFE89FB1D1D79" box="[985,1155,295,321]" italics="true" pageId="13" pageNumber="214">L. macropelma</emphasis>
Bates. The
</taxonomicName>
parameres of each
<taxonomicName id="4C704D7AAE7A934296B0FEE5FEE51D5D" authorityName="Bates. The" authorityYear="1887" box="[210,379,331,357]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="macropelma">
<emphasis id="B904EAEBAE7A934296B0FEE5FEE51D5D" box="[210,379,331,357]" italics="true" pageId="13" pageNumber="214">L. macropelma</emphasis>
</taxonomicName>
and
<taxonomicName id="4C704D7AAE7A934297D3FEE5FDBF1D5D" authorityName="Moser" authorityYear="1921" box="[433,545,331,357]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="gebieni">
<emphasis id="B904EAEBAE7A934297D3FEE5FDBF1D5D" box="[433,545,331,357]" italics="true" pageId="13" pageNumber="214">L. gebieni</emphasis>
</taxonomicName>
drawn by
<bibRefCitation id="EFE14B08AE7A934294FBFEE5FC811D5E" author="Frey, G." box="[665,799,331,358]" pageId="13" pageNumber="214" pagination="38 - 64" refId="ref18071" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
seem to be different. However, there were no specimens labeled as
<taxonomicName id="4C704D7AAE7A9342976CFEC1FEE31DB1" authorityName="Moser" authorityYear="1921" box="[270,381,367,393]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="gebieni">
<emphasis id="B904EAEBAE7A9342976CFEC1FEE31DB1" box="[270,381,367,393]" italics="true" pageId="13" pageNumber="214">L. gebieni</emphasis>
</taxonomicName>
among Freys material housed at NHMB or USNM, thus, the material used by
<bibRefCitation id="EFE14B08AE7A93429298FEC1FAE11DB2" author="Frey, G." box="[1274,1407,367,394]" pageId="13" pageNumber="214" pagination="38 - 64" refId="ref18071" refString="Frey, G. (1969) Bestimmungstabelle und revision der gattung Liogenys Guerrez (Coleoptera - Melolonthinae - Macrodactylini). Entomologische Arbeiten aus dem Museum G. Frey, 20, 38 - 64." type="journal article" year="1969">Frey (1969)</bibRefCitation>
to illustrate
<taxonomicName id="4C704D7AAE7A93429762FE3DFEF11D95" authorityName="Moser" authorityYear="1921" box="[256,367,403,429]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="gebieni">
<emphasis id="B904EAEBAE7A93429762FE3DFEF11D95" box="[256,367,403,429]" italics="true" pageId="13" pageNumber="214">L. gebieni</emphasis>
</taxonomicName>
parameres remains unknown. According to the
<bibRefCitation id="EFE14B08AE7A934295E5FE3DFA091D95" author="United States Department of Agriculture &amp; USDA" box="[903,1431,403,429]" pageId="13" pageNumber="214" pagination="335 - 336" refId="ref18813" refString="United States Department of Agriculture [USDA] (1971) Insects not known to occur in the United States: A Scarab (Liogenys macropelma Bates). Cooperative Economic Insect Report, 21, 335 - 336." type="journal article" year="1971">
<collectingCountry id="F3677669AE7A934295E5FE3DFB801D95" box="[903,1054,403,429]" name="United States of America" pageId="13" pageNumber="214">United States</collectingCountry>
Department of Agriculture (1971
</bibRefCitation>
,
<bibRefCitation id="EFE14B08AE7A934296F5FE19FF4D1DE9" author="United States Department of Agriculture &amp; USDA" box="[151,211,439,465]" pageId="13" pageNumber="214" pagination="55 - 68" refId="ref18853" refString="United States Department of Agriculture [USDA] (1974) List of intercepted plant pests, 1972 (Pest recorded from July 1, 1971, through June 30, 1972). Hayattsville, Maryland. U. S. G. P. O., Washington, D. C., pp. 55 - 68." type="book chapter" year="1974">1974</bibRefCitation>
),
<taxonomicName id="4C704D7AAE7A93429687FE19FE131DE9" authorityName="Bates. The" authorityYear="1887" box="[229,397,439,465]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="species" species="macropelma">
<emphasis id="B904EAEBAE7A93429687FE19FE131DE9" box="[229,397,439,465]" italics="true" pageId="13" pageNumber="214">L. macropelma</emphasis>
</taxonomicName>
has been intercepted in Florida,
<collectingRegion id="49B4F81BAE7A93429490FE19FCA81DE9" box="[754,822,439,465]" country="United States of America" name="Texas" pageId="13" pageNumber="214">Texas</collectingRegion>
, and
<collectingCountry id="F3677669AE7A9342950CFE19FC6C1DE9" box="[878,1010,439,465]" name="Puerto Rico" pageId="13" pageNumber="214">Puerto Rico</collectingCountry>
(
<collectingCountry id="F3677669AE7A9342959DFE19FA871DEA" box="[1023,1305,439,466]" name="United States of America" pageId="13" pageNumber="214">United States of America</collectingCountry>
) coming by aircraft from
<collectingCountry id="F3677669AE7A93429749FE75FE2E1DCD" box="[299,432,475,501]" name="El Salvador" pageId="13" pageNumber="214">El Salvador</collectingCountry>
,
<collectingCountry id="F3677669AE7A934297D9FE75FD891DCD" box="[443,535,475,501]" name="Panama" pageId="13" pageNumber="214">Panama</collectingCountry>
, and South America (probably
<collectingCountry id="F3677669AE7A93429512FE75FC281DCE" box="[880,950,475,502]" name="Brazil" pageId="13" pageNumber="214">Brazil</collectingCountry>
) several times in a period from 19481971, always between April and May (identification verified based on the illustration in the report). Also, it was recorded being a host of okra plant
<taxonomicName id="4C704D7AAE7A934297A4FD8DFD381E05" box="[454,678,547,573]" class="Magnoliopsida" family="Malvaceae" genus="Hibiscus" kingdom="Plantae" order="Malvales" pageId="13" pageNumber="214" phylum="Tracheophyta" rank="species" species="esculentus">
<emphasis id="B904EAEBAE7A934297A4FD8DFD381E05" box="[454,678,547,573]" italics="true" pageId="13" pageNumber="214">Hibiscus esculentus</emphasis>
</taxonomicName>
L. (
<taxonomicName id="4C704D7AAE7A934294B5FD8DFCCA1E06" box="[727,852,547,574]" class="Magnoliopsida" family="Malvaceae" kingdom="Plantae" order="Malvales" pageId="13" pageNumber="214" phylum="Tracheophyta" rank="family">Malvaceae</taxonomicName>
) coming from
<collectingCountry id="F3677669AE7A93429262FD8DFBE21E05" box="[1024,1148,547,573]" name="Guatemala" pageId="13" pageNumber="214">Guatemala</collectingCountry>
,
<collectingCountry id="F3677669AE7A934292EBFD8DFB7B1E05" box="[1161,1253,547,573]" name="Panama" pageId="13" pageNumber="214">Panama</collectingCountry>
, and
<collectingCountry id="F3677669AE7A93429340FD8DFA061E05" box="[1314,1432,547,573]" name="Venezuela" pageId="13" pageNumber="214">Venezuela</collectingCountry>
, which were intercepted in Florida, California, and
<collectingCountry id="F3677669AE7A934294AAFDE9FCD21E59" box="[712,844,583,609]" name="Puerto Rico" pageId="13" pageNumber="214">Puerto Rico</collectingCountry>
(
<collectingCountry id="F3677669AE7A9342953BFDE9FBE61E5A" box="[857,1144,583,610]" name="United States of America" pageId="13" pageNumber="214">United States of America</collectingCountry>
). Several specimens in the USNM collection had been caught in port interceptions from Houston,
<collectingRegion id="49B4F81BAE7A934295DEFDC5FB9E1EBD" box="[956,1024,619,645]" country="United States of America" name="Texas" pageId="13" pageNumber="214">Texas</collectingRegion>
, originated from
<collectingCountry id="F3677669AE7A934292A1FDC5FA811EBD" box="[1219,1311,619,645]" name="Panama" pageId="13" pageNumber="214">Panama</collectingCountry>
, 1966 with no other explanation.Another specimen in the USNM bears the following data:
<date id="FFCE1039AE7A9342926BFD21FB3C1E91" box="[1033,1186,655,681]" pageId="13" pageNumber="214" value="1962-04-27">27 April 1962</date>
port interception from Miami, Florida originating from “South America”. Based on the poor state of taxonomy of the genus
<taxonomicName id="4C704D7AAE7A93429297FD1DFAC41EF4" box="[1269,1370,691,716]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="214" phylum="Arthropoda" rank="genus">
<emphasis id="B904EAEBAE7A93429297FD1DFAC41EF4" box="[1269,1370,691,716]" italics="true" pageId="13" pageNumber="214">Liogenys</emphasis>
</taxonomicName>
when these port interceptions took place, all identifications should be considered questionable unless they can be verified using voucher specimens.
</paragraph>
</subSubSection>
</treatment>
</document>