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<document id="C39ECACB471C47172223E6C8F033BEA7" ID-CLB-Dataset="5549" ID-DOI="10.11646/zootaxa.4990.2.1" ID-GBIF-Dataset="acda30e5-6f57-43bb-81e6-70d3f96762d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5025886" ID-ZooBank="2C8C9945-A40A-409F-AD90-DE6A792285B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624535705376" checkinUser="plazi" docAuthor="Cherman, Mariana A., Basílio, Daniel S., Mise, Kleber M., Frisch, Johannes, Smith, Andrew B. T. &amp; Almeida, Lúcia M." docDate="2021" docId="03D987EFAE71934796F5FE27FBFA1F66" docLanguage="en" docName="zootaxa.4990.2.1.pdf" docOrigin="Zootaxa 4990 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liogenys genieri Smith &amp; Cherman 2021, new species" docType="treatment" docUuid="828B2892-0A98-453E-8AA0-0EB3B98DDBD3" docUuidSource="ZooBank" docVersion="6" lastPageNumber="209" masterDocId="FFE0FF97AE77934F9662FFAEFF9E1C38" masterDocTitle="Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species" masterLastPageNumber="226" masterPageNumber="201" pageNumber="207" updateTime="1699020235602" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D053F5BC1E5EC77F790820CD0962DC8A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="60F07FABD656B29F8A08391829EBDE09">
<mods:title id="F866AA8ECB5D70F35039B6CAEE39F17E">Liogenys Guerin-Méneville, 1831 (Coleoptera: Scarabaeidae: Melolonthinae Diplotaxini) of northern South America and Central America: taxonomic overview with four new species</mods:title>
</mods:titleInfo>
<mods:name id="7DDAC2DE4995B9877F0B65B9B39ECC64" type="personal">
<mods:role id="E5F044B00E1FAB06809322F488140A1A">
<mods:roleTerm id="89CFF00E5B06B870F5FEECD455D9729A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D6D7D146B2ECB174C4240829CD5F148">Cherman, Mariana A.</mods:namePart>
<mods:affiliation id="51DDCDCF9937BCA7E46F4124205979DB">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil. &amp; Laboratório de Scarabaeoidologia, Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, 78060 - 900, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="FFAF7A274D661455124EC70584E52585" type="personal">
<mods:role id="ECAC185A6FE382831F3A1F69D6F226D4">
<mods:roleTerm id="7F70B2B7ADFFEE669256F7E058FACEFA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3ABE71B25A41098B3A99AA3FC43CC36E">Basílio, Daniel S.</mods:namePart>
<mods:affiliation id="49859F8210674544DA41E0AC5CEDAF1A">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="3BE9D0D98FADB655DEB55348D7D92E8A" type="personal">
<mods:role id="E4D72C676EF4EB6A42E4337614AB72C2">
<mods:roleTerm id="EEF9813D67A4FB3895A3B2C41D4E1AC5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0EFF1A3FF5671742209DF5EBB0360FE0">Mise, Kleber M.</mods:namePart>
<mods:affiliation id="B09A033CF20FC0B657D7AE88D86CC545">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
</mods:name>
<mods:name id="9AA7C256A64954D10CDC885EBBBA179C" type="personal">
<mods:role id="4E5F5ED180EC06B25C0C915DD4512D7C">
<mods:roleTerm id="D48DEC84481284AEEDA9C5BA40B92605">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4311B964203D7763EF9FF3C2B1FD16FF">Frisch, Johannes</mods:namePart>
<mods:affiliation id="844BA8F93731C13B20173D55C547D843">Museum für Naturkunde, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
</mods:name>
<mods:name id="D1243FBC395BE5115C5F970F43775E53" type="personal">
<mods:role id="DDC95E99F9DB43226AAEC7A6D65BF250">
<mods:roleTerm id="F067B51CDE8EEB9220CF93FD408D69CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F8637F0EB2B39E139CC6F6E30A369D2A">Smith, Andrew B. T.</mods:namePart>
<mods:nameIdentifier id="43E8387B7C022E8DFFDF7D97E220BF64" type="ORCID">0000-0002-8059-5133</mods:nameIdentifier>
<mods:affiliation id="A0C8916FCF4EEECAFFDC247AA7DFE697">Research Division, Canadian Museum of Nature, P. O. Box 3443, Station D, Ottawa, Ontario, K 1 P 6 P 4, Canada. asmith @ nature. ca; https: // orcid. org / 0000 - 0002 - 8059 - 5133</mods:affiliation>
<mods:nameIdentifier id="8D0610E3EB12ED83AD16598631080898" type="email">asmith@nature.ca</mods:nameIdentifier>
</mods:name>
<mods:name id="9A88849A8F7DD359FFD51C7A60ACEB65" type="personal">
<mods:role id="50C091FA24DCE6951A318ECC15900D51">
<mods:roleTerm id="A65B7A21427446B3AF38A7D8C8E8552D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A43169A61133C117AB382BD8747D6ECA">Almeida, Lúcia M.</mods:namePart>
<mods:affiliation id="E2177AEC56C3B87B0C5C76997C8714A1">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Caixa Postal 19020, 81531 - 980, Curitiba, Paraná, Brazil.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5543E7DB2EE478E0D6042504E6AF2B79">text</mods:typeOfResource>
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<mods:titleInfo id="E493118912488360F2396C7AD8573502">
<mods:title id="CE00ADA04617DA23B20385ED94DA4814">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="05908FB2F735A77E3EBB04731CBCA128">
<mods:date id="37082A57B65F86D2322EF7DB4F40D0B4">2021</mods:date>
<mods:detail id="99C4CBA3E335C13261896CE6439A3EC6" type="pubDate">
<mods:number id="458BA4A0E644110A707013744E640470">2021-06-21</mods:number>
</mods:detail>
<mods:detail id="5C5AC94D1520E9EAE71B0D8F3D0C97CB" type="volume">
<mods:number id="0A24967211B1A86F932DA6C44FED9DC5">4990</mods:number>
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</mods:extent>
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<mods:classification id="FE1EF7C3108C5DB0C5A02672BE62F2BD">journal article</mods:classification>
<mods:identifier id="04572498F5B964A299BA4933498A5463" type="CLB-Dataset">5549</mods:identifier>
<mods:identifier id="4CCD1F49EF736104C4760A09658338B3" type="DOI">10.11646/zootaxa.4990.2.1</mods:identifier>
<mods:identifier id="007507872496227AF14B9234499413B5" type="GBIF-Dataset">acda30e5-6f57-43bb-81e6-70d3f96762d6</mods:identifier>
<mods:identifier id="748F1154D802DC8E8058D6AAE704FEFA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="522F0947C9069CA1E057ED5D7A509592" type="Zenodo-Dep">5025886</mods:identifier>
<mods:identifier id="E5ADF5495DBA7C55ACC9D2E55D56B7AB" type="ZooBank">2C8C9945-A40A-409F-AD90-DE6A792285B3</mods:identifier>
</mods:mods>
<treatment id="03D987EFAE71934796F5FE27FBFA1F66" ID-DOI="http://doi.org/10.5281/zenodo.5025898" ID-GBIF-Taxon="183501233" ID-Zenodo-Dep="5025898" LSID="urn:lsid:zoobank.org:act:828B2892-0A98-453E-8AA0-0EB3B98DDBD3" httpUri="http://treatment.plazi.org/id/03D987EFAE71934796F5FE27FBFA1F66" lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207">
<subSubSection id="C36A6572AE71934996F5FE27FD711D9B" box="[151,751,393,419]" pageId="6" pageNumber="207" type="nomenclature">
<paragraph id="8BCF36F9AE71934996F5FE27FD711D9B" blockId="6.[151,751,393,454]" box="[151,751,393,419]" pageId="6" pageNumber="207">
<heading id="D0878195AE71934996F5FE27FD711D9B" bold="true" box="[151,751,393,419]" fontSize="11" level="1" pageId="6" pageNumber="207" reason="1">
<emphasis id="B904EAEBAE71934996F5FE27FD711D9B" bold="true" box="[151,751,393,419]" pageId="6" pageNumber="207">
<taxonomicName id="4C704D7AAE71934996F5FE27FDCD1D9B" authority="Smith &amp; Cherman" authorityName="Smith &amp; Cherman" authorityYear="2021" box="[151,595,393,419]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="species" species="genieri" status="sp. nov.">
<emphasis id="B904EAEBAE71934996F5FE27FEFF1D9B" bold="true" box="[151,353,393,419]" italics="true" pageId="6" pageNumber="207">Liogenys genieri</emphasis>
Smith &amp; Cherman
</taxonomicName>
,
<taxonomicNameLabel id="A2375790AE719349943DFE27FD711D9B" box="[607,751,393,419]" pageId="6" pageNumber="207" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE71934996F5FE02FE981DFE" box="[151,262,428,454]" pageId="6" pageNumber="207" type="description">
<paragraph id="8BCF36F9AE71934996F5FE02FE981DFE" blockId="6.[151,751,393,454]" box="[151,262,428,454]" pageId="6" pageNumber="207">
<figureCitation id="134B2A7CAE71934996F5FE02FF7A1DFE" box="[151,228,428,454]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@7.[203,1385,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Liogenys genieri Smith &amp; Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) detail of anterolateral margin of pronotum reflexed, D) lateral, E) pygidium, and parameres E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025900" httpUri="https://zenodo.org/record/5025900/files/figure.png" pageId="6" pageNumber="207">Figs. 3</figureCitation>
,
<figureCitation id="134B2A7CAE7193499693FE02FE9C1DFE" box="[241,258,428,454]" captionStart="FIGURE 9" captionStartId="21.[151,250,1756,1780]" captionTargetBox="[184,1431,215,1732]" captionTargetId="figure-17@21.[151,1435,181,1732]" captionTargetPageId="21" captionText="FIGURE 9. Geographical distribution of the Liogenys of A) the northern South America and Central America, overlapping records in B) Panama and in C) Venezuela." figureDoi="http://doi.org/10.5281/zenodo.5025920" httpUri="https://zenodo.org/record/5025920/files/figure.png" pageId="6" pageNumber="207">9</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE71934996F5FE5FFAAD1F19" pageId="6" pageNumber="207" type="materials_examined">
<paragraph id="8BCF36F9AE71934996F5FE5FFB271EC6" blockId="6.[151,1437,497,2016]" pageId="6" pageNumber="207">
<emphasis id="B904EAEBAE71934996F5FE5FFEDA1E33" bold="true" box="[151,324,497,523]" pageId="6" pageNumber="207">Type material.</emphasis>
<materialsCitation id="3B183CA4AE7193499729FE5FFF601E4B" ID-GBIF-Occurrence="3320554367" collectingDate="1989-06-24" collectingDateMax="1989-06-26" collectingDateMin="1989-06-24" collectionCode="USNM" collectorName="M. Epstein &amp; M. Rodiguez" country="Venezuela" county="Hato" latitude="8.57" location="Calabozo" longLatPrecision="781" longitude="-67.58" municipality="Morades" pageId="6" pageNumber="207" specimenCount="2" stateProvince="Guarico" typeStatus="holotype">
<typeStatus id="54CB885BAE7193499729FE5FFE2D1E33" box="[331,435,497,523]" pageId="6" pageNumber="207" type="holotype">Holotype</typeStatus>
male, labeled: [white, typeset] “
<collectingCountry id="F3677669AE7193499578FE5FFC561E33" box="[794,968,497,523]" name="Venezuela" pageId="6" pageNumber="207">VENEZUELA</collectingCountry>
:
<collectingRegion id="49B4F81BAE71934995B3FE5FFBB01E33" box="[977,1070,497,523]" country="Venezuela" name="Guarico" pageId="6" pageNumber="207">Guarico</collectingRegion>
,
<collectingCounty id="62AE4E75AE719349925BFE5CFBF11E34" box="[1081,1135,498,524]" pageId="6" pageNumber="207">Hato</collectingCounty>
/ Flores
<collectingMunicipality id="6BABAC83AE71934992B3FE5FFAA81E33" box="[1233,1334,497,523]" pageId="6" pageNumber="207">Morades</collectingMunicipality>
,
<quantity id="4C889B1CAE7193499322FE5FFA181E33" box="[1344,1414,497,524]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="6" pageNumber="207" unit="km" value="45.0">45 km</quantity>
S /
<location id="8EAF6022AE71934996C4FDBAFE8D1E16" LSID="urn:lsid:plazi:treatment:03D987EFAE71934796F5FE27FBFA1F66:8EAF6022AE71934996C4FDBAFE8D1E16" box="[166,275,532,558]" country="Venezuela" county="Hato" latitude="8.57" longLatPrecision="781" longitude="-67.58" municipality="Morades" name="Calabozo" pageId="6" pageNumber="207" stateProvince="Guarico">Calabozo</location>
,
<geoCoordinate id="EE44503EAE719349977CFDBAFEFB1E17" box="[286,357,532,559]" degrees="8.57" direction="north" orientation="latitude" pageId="6" pageNumber="207" precision="555" value="8.57">8.57N</geoCoordinate>
,
<geoCoordinate id="EE44503EAE7193499713FDBAFE541E17" box="[369,458,532,559]" degrees="67.58" direction="west" orientation="longitude" pageId="6" pageNumber="207" precision="555" value="-67.58">67.58W</geoCoordinate>
/
<location id="8EAF6022AE71934997BDFDBAFDF21E17" LSID="urn:lsid:plazi:treatment:03D987EFAE71934796F5FE27FBFA1F66:8EAF6022AE71934997BDFDBAFDF21E17" box="[479,620,532,559]" country="Venezuela" county="Hato" latitude="8.57" longLatPrecision="781" longitude="-67.58" municipality="Morades" name="Galry Forest" pageId="6" pageNumber="207" stateProvince="Guarico">Galry Forest</location>
#2, 75 m /
<date id="FFCE1039AE7193499489FDBAFC381E16" box="[747,934,532,558]" pageId="6" pageNumber="207" value="1989-06-24" valueMax="1989-06-26" valueMin="1989-06-24">
<collectingDate id="EF8AE9D1AE7193499489FDBAFC381E16" box="[747,934,532,558]" pageId="6" pageNumber="207" value="1989-06-24" valueMax="1989-06-26" valueMin="1989-06-24">24–26June 1989</collectingDate>
</date>
, uv light /
<collectorName id="2685532FAE719349927FFDBBFB061E16" box="[1053,1176,532,559]" pageId="6" pageNumber="207">M. Epstein</collectorName>
&amp;
<collectorName id="2685532FAE71934992DEFDBBFACD1E16" box="[1212,1363,532,559]" pageId="6" pageNumber="207">M. Rodiguez</collectorName>
”, [red, typeset and handwritten] “
<taxonomicName id="4C704D7AAE71934997A6FD99FDCD1E69" box="[452,595,567,593]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">LIOGENYS</taxonomicName>
/ GENIERI / SMITH &amp; CHERMAN /
<typeStatus id="54CB885BAE7193499242FD99FB241E69" box="[1056,1210,567,593]" pageId="6" pageNumber="207" type="holotype">HOLOTYPE</typeStatus>
”, genitalia mounted (
<collectionCode id="ED61AE3CAE71934996FDFDF7FF681E4B" box="[159,246,601,627]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="207" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B183CA4AE7193499765FDF4FEFB1EC7" ID-GBIF-Occurrence="3320554346" collectingDate="1985-05-03" collectingDateMax="1985-05-10" collectingDateMin="1985-05-03" collectionCode="CMNC, DZUP, USNM" collectorName="Each" country="Venezuela" county="Hato" latitude="8.57" location="Hato Masaguaral" longLatPrecision="781" longitude="-67.58" municipality="One" pageId="6" pageNumber="207" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Guarico" typeStatus="paratype">
<typeStatus id="54CB885BAE7193499765FDF4FEEA1E4C" box="[263,372,602,628]" pageId="6" pageNumber="207" type="paratype">Paratypes</typeStatus>
(8), all bearing the label [yellow, typeset and handwritten] “
<taxonomicName id="4C704D7AAE7193499264FDF7FB0B1E4B" box="[1030,1173,601,627]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="207" phylum="Arthropoda" rank="genus">LIOGENYS</taxonomicName>
/ GENIERI / SMITH &amp; CHERMAN /
<typeStatus id="54CB885BAE7193499755FDD2FE541EAE" box="[311,458,636,662]" pageId="6" pageNumber="207" type="paratype">PARATYPE</typeStatus>
”: One male and
<specimenCount id="9D76FD70AE7193499419FDD2FD671EAE" box="[635,761,636,662]" pageId="6" pageNumber="207" type="female">six females</specimenCount>
with the same data as the holotype (
<specimenCount id="9D76FD70AE71934992E5FDD2FB731EAE" box="[1159,1261,636,662]" pageId="6" pageNumber="207" type="male">one male</specimenCount>
and
<specimenCount id="9D76FD70AE7193499340FDD3FA021EAE" box="[1314,1436,636,663]" pageId="6" pageNumber="207" type="female">one female</specimenCount>
at
<collectionCode id="ED61AE3CAE71934996D3FD31FE981E81" box="[177,262,671,697]" pageId="6" pageNumber="207">CMNC</collectionCode>
,
<specimenCount id="9D76FD70AE7193499772FD31FE091E81" box="[272,407,671,697]" pageId="6" pageNumber="207" type="female">two females</specimenCount>
at
<collectionCode id="ED61AE3CAE71934997D4FD31FE611E81" box="[438,511,671,697]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="6" pageNumber="207" type="University or college">DZUP</collectionCode>
,
<specimenCount id="9D76FD70AE719349946BFD31FD031E81" box="[521,669,671,697]" pageId="6" pageNumber="207" type="female">three females</specimenCount>
at
<collectionCode id="ED61AE3CAE71934994DFFD31FC8D1E81" box="[701,787,671,697]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="207" type="Museum">USNM</collectionCode>
).
<collectingMunicipality id="6BABAC83AE7193499546FD31FCCD1E81" box="[804,851,671,697]" pageId="6" pageNumber="207">One</collectingMunicipality>
male with the data: [white, typeset] “
<collectingCountry id="F3677669AE7193499289FD31FA071E81" box="[1259,1433,671,697]" name="Venezuela" pageId="6" pageNumber="207">VENEZUELA</collectingCountry>
:
<collectingRegion id="49B4F81BAE71934996F5FD6FFF6F1EE3" box="[151,241,705,731]" country="Venezuela" name="Guarico" pageId="6" pageNumber="207">Guarico</collectingRegion>
/
<location id="8EAF6022AE7193499768FD6CFE521EE3" LSID="urn:lsid:plazi:treatment:03D987EFAE71934796F5FE27FBFA1F66:8EAF6022AE7193499768FD6CFE521EE3" box="[266,460,705,732]" country="Venezuela" municipality="One" name="Hato Masaguaral" pageId="6" pageNumber="207" stateProvince="Guarico">Hato Masaguaral</location>
/ (
<locationDeviation id="9A7D2FDBAE7193499789FD6FFD271EE4" box="[491,697,705,732]" location="Calabozo" pageId="6" pageNumber="207">
<quantity id="4C889B1CAE7193499789FD6FFDB31EE3" box="[491,557,705,731]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="207" unit="km" value="44.0">44km</quantity>
S Calabozo
</locationDeviation>
) /
<date id="FFCE1039AE71934994BBFD6CFC131EE4" box="[729,909,705,732]" pageId="6" pageNumber="207" value="1985-05-03" valueMax="1985-05-10" valueMin="1985-05-03">
<collectingDate id="EF8AE9D1AE71934994BBFD6CFC131EE4" box="[729,909,705,732]" pageId="6" pageNumber="207" value="1985-05-03" valueMax="1985-05-10" valueMin="1985-05-03">May 3–10 1985</collectingDate>
</date>
/ Menke &amp; Carpenter” (
<collectionCode id="ED61AE3CAE71934992C7FD6FFB621EE3" box="[1189,1276,705,731]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="207" type="Museum">USNM</collectionCode>
).
<collectorName id="2685532FAE7193499372FD6FFAD61EE3" box="[1296,1352,705,731]" pageId="6" pageNumber="207">Each</collectorName>
female
<typeStatus id="54CB885BAE71934996F5FD4BFF691EC7" box="[151,247,741,767]" pageId="6" pageNumber="207" type="paratype">paratype</typeStatus>
of
<collectionCode id="ED61AE3CAE719349977FFD4BFEFB1EC7" box="[285,357,741,767]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="6" pageNumber="207" type="University or college">DZUP</collectionCode>
</materialsCitation>
<materialsCitation id="3B183CA4AE719349970EFD4AFB2B1EC6" ID-GBIF-Occurrence="3320554311" box="[364,1205,739,766]" collectionCode="DZUP" country="Venezuela" pageId="6" pageNumber="207" specimenCode="DZUP / 402748, DZUP / 402749" specimenCount="1" stateProvince="Guarico" typeStatus="paratype">
with the voucher: [white typeset] “
<specimenCode id="DBD69E82AE719349948CFD4AFC381EC6" box="[750,934,740,766]" collectionCode="DZUP" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="6" pageNumber="207" type="University or college">DZUP / 402748</specimenCode>
” and “
<specimenCode id="DBD69E82AE7193499592FD4AFB351EC6" box="[1008,1195,740,766]" collectionCode="DZUP" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="6" pageNumber="207" type="University or college">DZUP / 402749</specimenCode>
”
</materialsCitation>
.
</paragraph>
<paragraph id="8BCF36F9AE71934996A5FCA9FAAD1F19" blockId="6.[151,1437,497,2016]" box="[199,1331,775,801]" pageId="6" pageNumber="207">
<materialsCitation id="3B183CA4AE71934996A5FCA9FAAD1F19" ID-GBIF-Occurrence="3320554312" box="[199,1331,775,801]" collectionCode="USNM, CMNC, DZUP" pageId="6" pageNumber="207" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CB885BAE71934996A5FCA9FEB11F19" box="[199,303,775,801]" pageId="6" pageNumber="207" type="holotype">Holotype</typeStatus>
and
<specimenCount id="9D76FD70AE7193499704FCA9FD981F19" box="[358,518,775,801]" pageId="6" pageNumber="207" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
deposited at
<collectionCode id="ED61AE3CAE71934994F9FCA9FD6E1F19" box="[667,752,775,801]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="6" pageNumber="207" type="Museum">USNM</collectionCode>
. Two
<typeStatus id="54CB885BAE7193499550FCA9FC031F19" box="[818,925,775,801]" pageId="6" pageNumber="207" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED61AE3CAE71934995A2FCA9FB881F19" box="[960,1046,775,801]" pageId="6" pageNumber="207">CMNC</collectionCode>
. Two
<typeStatus id="54CB885BAE719349923AFCA9FB5D1F19" box="[1112,1219,775,801]" pageId="6" pageNumber="207" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED61AE3CAE7193499287FCA9FAB11F19" box="[1253,1327,775,801]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="6" pageNumber="207" type="University or college">DZUP</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE71934996A5FC87FB7D180E" pageId="6" pageNumber="207" type="diagnosis">
<paragraph id="8BCF36F9AE71934996A5FC87FB7D180E" blockId="6.[151,1437,497,2016]" pageId="6" pageNumber="207">
<emphasis id="B904EAEBAE71934996A5FC87FEDC1F7B" bold="true" box="[199,322,809,835]" pageId="6" pageNumber="207">Diagnosis.</emphasis>
Body light brown, elongate, sides almost parallel; elytra light brown; pronotum reddish brown (
<figureCitation id="134B2A7CAE719349930DFC87FF231F5E" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@7.[203,1385,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Liogenys genieri Smith &amp; Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) detail of anterolateral margin of pronotum reflexed, D) lateral, E) pygidium, and parameres E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025900" httpUri="https://zenodo.org/record/5025900/files/figure.png" pageId="6" pageNumber="207">Fig. 3A</figureCitation>
); clypeus quadridentate; clypeal emargination deep, angulate, and narrow; distance between lateral and anterior clypeal teeth approximately equal to basal width of one anterior tooth; angle between lateral and anterior clypeal teeth approximately 90º; pronotal corners rounded (
<figureCitation id="134B2A7CAE719349948CFC3FFCD71F93" box="[750,841,913,939]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@7.[203,1385,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Liogenys genieri Smith &amp; Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) detail of anterolateral margin of pronotum reflexed, D) lateral, E) pygidium, and parameres E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025900" httpUri="https://zenodo.org/record/5025900/files/figure.png" pageId="6" pageNumber="207">Fig. 3B</figureCitation>
); mesotibia subquadrate in cross section; pygidium slightly convex; pygidial disc glabrous, with scattered bristles on apex (
<figureCitation id="134B2A7CAE71934995ACFC1AFBB41FF6" box="[974,1066,948,974]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@7.[203,1385,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Liogenys genieri Smith &amp; Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) detail of anterolateral margin of pronotum reflexed, D) lateral, E) pygidium, and parameres E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025900" httpUri="https://zenodo.org/record/5025900/files/figure.png" pageId="6" pageNumber="207">Fig. 3D</figureCitation>
); in males protarsomere II more than twice the length of protarsomere I in dorsal view (
<figureCitation id="134B2A7CAE7193499567FC79FCC01FC9" box="[773,862,983,1009]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@7.[203,1385,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Liogenys genieri Smith &amp; Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) detail of anterolateral margin of pronotum reflexed, D) lateral, E) pygidium, and parameres E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025900" httpUri="https://zenodo.org/record/5025900/files/figure.png" pageId="6" pageNumber="207">Fig. 3C</figureCitation>
); parameres more than six times the length of their apex; lateral expansion along the parameres only seen subapically; inner margins convergent; apex expanded with backward-facing spine weakly projected, spine length less than one-third length of apex (
<figureCitation id="134B2A7CAE719349921DFBB2FB48180E" box="[1151,1238,1052,1078]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@7.[203,1385,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Liogenys genieri Smith &amp; Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) detail of anterolateral margin of pronotum reflexed, D) lateral, E) pygidium, and parameres E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025900" httpUri="https://zenodo.org/record/5025900/files/figure.png" pageId="6" pageNumber="207">Fig. 3E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE71934796A5FB91FDAD1D25" lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207" type="materials_examined">
<paragraph id="8BCF36F9AE71934796A5FB91FDAD1D25" blockId="6.[151,1437,497,2016]" lastBlockId="8.[151,1437,151,862]" lastPageId="8" lastPageNumber="209" pageId="6" pageNumber="207">
<emphasis id="B904EAEBAE71934996A5FB91FEE11861" bold="true" box="[199,383,1087,1113]" pageId="6" pageNumber="207">
<typeStatus id="54CB885BAE71934996A5FB91FEA81861" box="[199,310,1087,1113]" pageId="6" pageNumber="207" type="holotype">Holotype</typeStatus>
. Male
</emphasis>
. Length:
<quantity id="4C889B1CAE7193499789FB91FDDA1861" box="[491,580,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="6" pageNumber="207" unit="mm" value="9.6">9.6 mm</quantity>
, width:
<quantity id="4C889B1CAE71934994FCFB91FD6A1861" box="[670,756,1087,1113]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="6" pageNumber="207" unit="mm" value="4.6">4.6 mm</quantity>
. Light brown.
<emphasis id="B904EAEBAE71934995FEFB91FC7F1861" box="[924,993,1087,1113]" italics="true" pageId="6" pageNumber="207">Head:</emphasis>
distance between eyes nearly twice the width of one eye; frons equal in length to clypeus; clypeal emargination deep, angulate, and wide; outer sides of anterior teeth parallel; outer margin of anterior teeth shorter than the eye in dorsal view; clypeus convex laterally and strongly produced forming a conspicuous tooth-like projection; distance between lateral and anterior teeth approximately equal to basal width of one anterior tooth, distance between lateral tooth and anterior margin of eye approximately equal to one eye length, angle between anterior and lateral teeth approximately 90º; canthus not exceeding the outer margin of the eye; distal maxillary palpomere, maximum width twice the width at apex; antenna with 10 antennomeres, club lighter in color and longer than funicle.
<emphasis id="B904EAEBAE71934995DDFA9FFB871973" box="[959,1049,1329,1355]" italics="true" pageId="6" pageNumber="207">Thorax:</emphasis>
pronotal anterior margin slightly produced medially, undulate, and flanged; disc glabrous, punctures moderately dense and coarse; anterolateral margin reflexed; pronotal posterior corners rounded; hypomere with long bristles; mesepisternum scaly; anterior portion and sides of metaventrite with bristles; distance between mesocoxae and metacoxae nearly twice the length of the metacoxa; scutellum triangular with sparse, coarse punctures.
<emphasis id="B904EAEBAE71934995C2FA12FC7119EE" box="[928,1007,1468,1494]" italics="true" pageId="6" pageNumber="207">Elytra:</emphasis>
shiny, uniform light brown, lighter in color than the pronotum; elytra near three times the length of pronotum; elytral suture darker than the elytron and weakly elevated; all four elytral ridges weakly defined.
<emphasis id="B904EAEBAE7193499574F9ACFCCB1A23" box="[790,853,1538,1563]" italics="true" pageId="6" pageNumber="207">Legs:</emphasis>
procoxa with abundant, thin and long scales, short scales on outer surface; three protibial teeth, middle and apical teeth equal in size, distance between basal and middle teeth longer than between middle and apical teeth; protibial spur present; mesofemur with a row of long bristles on anterior and posterior margins, mesotibia subquadrate in cross section, surface coarsely sculptured on outer margin, mesotibia with two transverse carinae, the apical carina incomplete; metacoxa with bristles throughout, basal articular lobe of metacoxa produced beyond the outer margin of trochanter; inner margin of metatibia straight, carinate, and produced on apex, apical inner surface with bristles, metatibial surface coarsely sculptured; metatibial transverse carina weakly defined, only present posteriorly; protarsomere II elongate; protarsomeres and mesotarsomeres I–IV enlarged, protarsomeres slightly wider than the mesotarsomeres and two times wider than the metatarsomeres; metatarsomere I slightly shorter than metatarsomere II; claw bifid, symmetrical, outer tooth of a claw longer and as wide as the inner tooth; distance between teeth shorter than the inner tooth.
<emphasis id="B904EAEBAE71934995B9F8F2FBD01B4E" box="[987,1102,1884,1910]" italics="true" pageId="6" pageNumber="207">Abdomen:</emphasis>
disc of ventrites with bristles; propygidium with bristles, pygidium weakly convex, subquadrate, wide; pygidial width approximately equal to distance between spiracles of propygidium; pygidial disc glabrous except for a few bristles on apex, pygidial disc coarsely punctate; pygidial apex oval.
<emphasis id="B904EAEBAE7193499424F868FD5E1BE7" box="[582,704,1990,2015]" italics="true" pageId="6" pageNumber="207">Parameres</emphasis>
: basal region dorsally narrowed, equal to both sections of the parameres at its midline; parameral split at 2/3 the length of the basal region; inner margins slightly convergent; parameres more than six times the length of their apex; apex harpoon shaped; laterally expanded with backward-facing spine weakly projected straight downward, spine length less than one-third length of apex. Parameres in lateral view straight, not coplanar (
<figureCitation id="134B2A7CAE7F934797B3FEADFDB81D25" box="[465,550,259,285]" captionStart="FIGURE 3" captionStartId="7.[151,250,1943,1967]" captionTargetBox="[203,1385,181,1920]" captionTargetId="figure-17@7.[203,1385,181,1920]" captionTargetPageId="7" captionText="FIGURE 3. Liogenys genieri Smith &amp; Cherman new species male holotype views: A) dorsal, B) clypeus and pronotum, C) detail of anterolateral margin of pronotum reflexed, D) lateral, E) pygidium, and parameres E) dorsal and F) lateral." figureDoi="http://doi.org/10.5281/zenodo.5025900" httpUri="https://zenodo.org/record/5025900/files/figure.png" pageId="8" pageNumber="209">Fig. 3F</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF0F6671AE70934896F5F839FAB41BEC" ID-DOI="http://doi.org/10.5281/zenodo.5025900" ID-Zenodo-Dep="5025900" httpUri="https://zenodo.org/record/5025900/files/figure.png" pageId="7" pageNumber="208" startId="7.[151,250,1943,1967]" targetBox="[203,1385,181,1920]" targetPageId="7">
<paragraph id="8BCF36F9AE70934896F5F839FAB41BEC" blockId="7.[151,1437,1943,2004]" pageId="7" pageNumber="208">
<emphasis id="B904EAEBAE70934896F5F839FE8A1B88" bold="true" box="[151,276,1943,1968]" pageId="7" pageNumber="208">FIGURE 3.</emphasis>
<taxonomicName id="4C704D7AAE709348977FF836FD101B88" authority="Smith &amp; Cherman" authorityName="Smith &amp; Cherman" authorityYear="2021" box="[285,654,1944,1968]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="208" phylum="Arthropoda" rank="species" species="genieri" status="sp. nov.">
<emphasis id="B904EAEBAE709348977FF836FE571B97" box="[285,457,1944,1967]" italics="true" pageId="7" pageNumber="208">Liogenys genieri</emphasis>
Smith &amp; Cherman
</taxonomicName>
<emphasis id="B904EAEBAE70934894F4F836FC8B1B97" bold="true" box="[662,789,1943,1968]" pageId="7" pageNumber="208">
<taxonomicNameLabel id="A2375790AE70934894F4F836FC8B1B97" box="[662,789,1943,1968]" pageId="7" pageNumber="208" rank="species">new species</taxonomicNameLabel>
</emphasis>
male holotype views: A) dorsal, B) clypeus and pronotum, C) detail of anterolateral margin of pronotum reflexed, D) lateral, E) pygidium, and parameres E) dorsal and F) lateral.
</paragraph>
</caption>
<subSubSection id="C36A6572AE7F934796A5FE88FBA41D96" pageId="8" pageNumber="209" type="description">
<paragraph id="8BCF36F9AE7F934796A5FE88FBA41D96" blockId="8.[151,1437,151,862]" pageId="8" pageNumber="209">
<emphasis id="B904EAEBAE7F934796A5FE88FEA61D78" bold="true" box="[199,312,294,320]" pageId="8" pageNumber="209">Variation</emphasis>
. Female length:
<quantity id="4C889B1CAE7F93479463FE89FD371D79" box="[513,681,295,321]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" metricValueMax="1.1199999999999999" metricValueMin="1.02" pageId="8" pageNumber="209" unit="mm" value="10.7" valueMax="11.2" valueMin="10.2">10.2–11.2 mm</quantity>
, width 5.0–
<quantity id="4C889B1CAE7F93479557FE88FC0C1D79" box="[821,914,294,321]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="8" pageNumber="209" unit="mm" value="5.6">5.6 mm</quantity>
. Females as the
<typeStatus id="54CB885BAE7F9347923BFE89FB251D79" box="[1113,1211,295,321]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
except in the size; pronotum wider, with anterolateral margin distinctly reflexed; maximum width of distal maxillary palpomere slightly wider than at apex. Male length:
<quantity id="4C889B1CAE7F93479469FEC1FD0A1DB1" box="[523,660,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="9.6" metricValueMin="9.4" pageId="8" pageNumber="209" unit="mm" value="9.5" valueMax="9.6" valueMin="9.4">9.4–9.6 mm</quantity>
, width 4.0–
<quantity id="4C889B1CAE7F9347957BFEC1FCEF1DB1" box="[793,881,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="8" pageNumber="209" unit="mm" value="4.8">4.8 mm</quantity>
. Males as the
<typeStatus id="54CB885BAE7F93479272FEC1FBEC1DB1" box="[1040,1138,367,393]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
except in the punctures of pronotum slightly less dense; protibial teeth sharper and longer (worn in
<typeStatus id="54CB885BAE7F934795A5FE3DFBB01D96" box="[967,1070,403,430]" pageId="8" pageNumber="209" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE7F934796A5FE19FD621DCD" pageId="8" pageNumber="209" type="etymology">
<paragraph id="8BCF36F9AE7F934796A5FE19FD621DCD" blockId="8.[151,1437,151,862]" pageId="8" pageNumber="209">
<emphasis id="B904EAEBAE7F934796A5FE19FED01DE9" bold="true" box="[199,334,439,465]" pageId="8" pageNumber="209">Etymology.</emphasis>
This species is named after François Génier, scarab taxonomist from the CMNC, in appreciation for his help in photographing specimens for this paper.
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE7F934796A5FE51FCB21E06" pageId="8" pageNumber="209" type="materials_examined">
<paragraph id="8BCF36F9AE7F934796A5FE51FC681E21" blockId="8.[151,1437,151,862]" box="[199,1014,510,537]" pageId="8" pageNumber="209">
<emphasis id="B904EAEBAE7F934796A5FE51FEFE1E20" bold="true" box="[199,352,510,537]" pageId="8" pageNumber="209">Type locality</emphasis>
.
<materialsCitation id="3B183CA4AE7F9347970CFE51FC6C1E21" ID-GBIF-Occurrence="3320554336" box="[366,1010,510,537]" country="Venezuela" latitude="8.57" location="Hato Masaguaral" longLatPrecision="781" longitude="-67.58" pageId="8" pageNumber="209" specimenCount="1" stateProvince="Guarico">
<location id="8EAF6022AE7F9347970CFE51FDAF1E21" LSID="urn:lsid:plazi:treatment:03D987EFAE71934796F5FE27FBFA1F66:8EAF6022AE7F9347970CFE51FDAF1E21" box="[366,561,511,537]" country="Venezuela" latitude="8.57" longLatPrecision="781" longitude="-67.58" name="Hato Masaguaral" pageId="8" pageNumber="209" stateProvince="Guarico">Hato Masaguaral</location>
,
<collectingRegion id="49B4F81BAE7F9347945EFE51FD071E21" box="[572,665,511,537]" country="Venezuela" name="Guarico" pageId="8" pageNumber="209">Guárico</collectingRegion>
,
<collectingCountry id="F3677669AE7F934794C6FE51FC891E21" box="[676,791,511,537]" name="Venezuela" pageId="8" pageNumber="209">Venezuela</collectingCountry>
(
<geoCoordinate id="EE44503EAE7F93479544FE50FCE91E21" box="[806,887,510,537]" degrees="8.57" direction="north" orientation="latitude" pageId="8" pageNumber="209" precision="555" value="8.57">8.57ºN</geoCoordinate>
,
<geoCoordinate id="EE44503EAE7F934795E1FE50FC741E21" box="[899,1002,510,537]" degrees="67.58" direction="west" orientation="longitude" pageId="8" pageNumber="209" precision="555" value="-67.58">67.58ºW</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCF36F9AE7F934796A5FD8CFCB21E06" blockId="8.[151,1437,151,862]" box="[199,812,546,574]" pageId="8" pageNumber="209">
<emphasis id="B904EAEBAE7F934796A5FD8CFD9B1E04" bold="true" box="[199,517,546,572]" pageId="8" pageNumber="209">Geographical distribution.</emphasis>
<collectingCountry id="F3677669AE7F9347946EFD8DFD2F1E05" box="[524,689,547,573]" name="Venezuela" pageId="8" pageNumber="209">VENEZUELA</collectingCountry>
(
<collectingRegion id="49B4F81BAE7F934794DDFD8DFCBE1E06" box="[703,800,547,574]" country="Venezuela" name="Guarico" pageId="8" pageNumber="209">Guárico</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6572AE7F934796A5FDE9FBFA1F66" pageId="8" pageNumber="209" type="discussion">
<paragraph id="8BCF36F9AE7F934796A5FDE9FBFA1F66" blockId="8.[151,1437,151,862]" pageId="8" pageNumber="209">
<emphasis id="B904EAEBAE7F934796A5FDE9FEAA1E59" bold="true" box="[199,308,583,609]" pageId="8" pageNumber="209">Remarks</emphasis>
.
<taxonomicName id="4C704D7AAE7F93479726FDE9FD431E59" authority="Smith &amp; Cherman" authorityName="Smith &amp; Cherman" authorityYear="2021" box="[324,733,583,609]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="genieri" status="sp. nov.">
<emphasis id="B904EAEBAE7F93479726FDE9FD9E1E58" box="[324,512,583,608]" italics="true" pageId="8" pageNumber="209">Liogenys genieri</emphasis>
Smith &amp; Cherman
</taxonomicName>
,
<emphasis id="B904EAEBAE7F9347948BFDE9FCEA1E58" bold="true" box="[745,884,582,609]" pageId="8" pageNumber="209">
<taxonomicNameLabel id="A2375790AE7F9347948BFDE9FCEA1E58" box="[745,884,582,609]" pageId="8" pageNumber="209" rank="species">new species</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="4C704D7AAE7F93479594FDE9FB011E59" authorityName="Cherman" authorityYear="2021" box="[1014,1183,583,609]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="clipeosetosa">
<emphasis id="B904EAEBAE7F93479594FDE9FB011E59" box="[1014,1183,583,609]" italics="true" pageId="8" pageNumber="209">L. clipeosetosa</emphasis>
</taxonomicName>
in the body color and size, shape of the quadridentate clypeus, head and pronotum coarsely punctate, anterior teeth of clypeus long, anterior and lateral teeth forming approximately 90º angle, and posterior corners of pronotum rounded. Males in both species have parameres very long, more than five times the apical length, and apex laterally weakly projected.
<taxonomicName id="4C704D7AAE7F934796F5FD79FECC1EC8" authorityName="Smith &amp; Cherman" authorityYear="2021" box="[151,338,727,752]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="genieri">
<emphasis id="B904EAEBAE7F934796F5FD79FECC1EC8" box="[151,338,727,752]" italics="true" pageId="8" pageNumber="209">Liogenys genieri</emphasis>
</taxonomicName>
differs (
<taxonomicName id="4C704D7AAE7F934797D0FD79FDC71EC9" authorityName="Cherman" authorityYear="2021" box="[434,601,727,753]" class="Insecta" family="Melolonthidae" genus="Liogenys" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="209" phylum="Arthropoda" rank="species" species="clipeosetosa">
<emphasis id="B904EAEBAE7F934797D0FD79FDC71EC9" box="[434,601,727,753]" italics="true" pageId="8" pageNumber="209">L. clipeosetosa</emphasis>
</taxonomicName>
in parenthesis) in the frontoclypeal surface without bristles (with bristles); clypeal emargination angulate, narrower (rounded, wider); pronotum with anterolateral margin reflexed (not as so); apex of pygidium oval (quadrate); males with parameres slightly shorter, lateral expansion along the parameres not visible dorsally and straight in lateral view (lateral expansion entirely visible dorsally).
</paragraph>
</subSubSection>
</treatment>
</document>
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